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Improved secretion of an Fab fragment by using a modified BiP signal sequence upstream of the heavy chain (Hc) gene. a Antigen-binding activity of an Fab fragment, as detected by enzyme-linked immunosorbent assay (ELISA). The binding activity obtained with the native BiP signal sequence was indicated as 100%. Error bar = 1 SD (n = 3). b Fab fragment (Mw 50,300), as detected by western blotting
