1Section Biology and Evolution of Marine Organisms (BEOM), Stazione Zoologica Anton Dohrn (SZN), Villa Comunale, 80121 Napoli, Italy
2Smithsonian Tropical Research Institute (STRI), Tupper/Naos/Bocas del Toro Labs, Ancón, 0843-03092 Panamá City, Republic of Panama
3US Geological Survey, National Wildlife Health Center, Honolulu Field Station, Honolulu, HI 96850 USA
4Department of Evolutionary Biology Ecology and Environmental Sciences, and Biodiversity Research Institute (IrBIO), University of Barcelona, Faculty of Biology, Av. Diagonal 643, 08028 Barcelona, Catalonia Spain
5Museum of Comparative Zoology & Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge, MA 02138 USA
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