Annotated nomenclator of extant and fossil taxa of the Paludomidae (Caenogastropoda, Cerithioidea)

Abstract Abstract

This nomenclator provides bibliographic details on all names in the family-, genus-, and species-group of the the family Paludomidae. All nomenclaturally available names are discussed including junior homonyms and objective junior synonyms as well as unavailable names such as nomina nuda, infrasubspecific names and, to some extent, also incorrect subsequent spellings. In the family-group a total of 28 names are included in the nomenclator, of which 21 are available and seven unavailable names. Of the available names in the family-group, six are invalid for nomenclatural reasons. In the genus-group a total of 57 names are included in the catalogue. Of the available names in the genus-group, 11 are invalid for nomenclatural reasons. In the species-group a total of 499 names are included, of which 463 are available, but 21 are invalid for nomenclatural reasons. All names are given in their original combination and spelling (mandatory changes are discussed and corrected spellings are provided), along with the reference to the original publication. For each family- and genus-group name, the original classification and the type genus or type species, respectively, are given. For species-group taxa the type locality and type horizon (for fossil taxa) are provided, usually as given in the original publication. A new name, Cleopatraadami nom. nov., is proposed for the fossil Cleopatracylindrica (Adam, 1957), which is a homonym of Cleopatracridlandicylindrica Mandahl-Barth, 1954, and a lectotype for Cleopatradubia Adam, 1959 is designated. A new replacement name Leloupiella nom. nov. is proposed for Stormsia Leloup, 1953 which is a homonym of Stormsia Bourguignat, 1891.

Introduction

The Paludomidae Stoliczka, 1868 is currently understood as representing taxa with a general distribution including most of tropical sub-Saharan Africa and Madagascar as well as South and Southeast Asia (see Neiber and Glaubrecht 2019b who summarize the results of phylogenetic analyses presented by West and Michel 2000; Michel 2004; Wilson et al. 2004; Glaubrecht and Strong 2007; Strong et al. 2011; for a recent compilation on the diversity of family-group names available for this group and recent classifications, see also Bouchet and Rocroi 2005; Bouchet et al. 2017).

Species in this family of freshwater cerithioideans show a high diversity with regard to shell shape and ornamentation. The endemic paludomids of Lake Tanganyika have especially attracted much attention by taxonomists, early biogeographers, and evolutionary biologists because of their thalassoid (Bourguignat 1885) appearance, i.e. their resemblance to various marine gastropod families (Glaubrecht 1996, 2008; see also Glaubrecht 2009, 2011 for reviews). A large number of names have been introduced for Tanganyikan paludomids by the French malacologists J.-R. Bourguignat, the head of the so called ‘Nouvelle École’ (Dance 1970), on the basis of differences of the shell that are often hardly perceptible (Bourguignat 1885, 1888). However, being among the first to work on the gastropod fauna of the lake, several names first introduced by this author are still in use to this day, although the far greater number certainly represent individual variations that do not merit to be accepted and are thus to a large extent a burden today for taxonomists, biogeographers, and evolutionary biologists working on the paludomid fauna of Lake Tanganyika.

Although several synoptical works covering the regional diversity of Paludomidae in Africa or Asia have been published (e.g. Preston 1915b; Pilsbry and Bequaert 1927; Leloup 1953; Brown 1994; West et al. 2003), Asian members of the family are still especially in need of revision. The basic prerequisite of any taxonomic work on a specific group of organisms is the historical perspective of what is already known, or from a nomenclatural point of view, which names have already been proposed for the group in question. However, a comprehensive annotated list of paludomid names is currently unavailable. Therefore, the nomenclator presented here aims at filling this gap by providing information on all names for Recent and fossil family-, genus-, and species-group taxa introduced for Paludomidae. In addition, the present contribution is intended to facilitate future taxonomic work on Paludomidae by providing direct internet links to many important older taxonomic publications following the general outlines of the nomenclators of other groups of freshwater snails presented by Haszprunar (2014) on valvatids and Neubauer (2016) on melanopsids.

Methods

The present nomenclator includes only taxa that are, from the current perspective, members of the Paludomidae Stoliczka, 1868 and with at least some degree of plausibility (based on shell morphology, geographical and/or stratigraphical distribution) assignable to this family. Not included are, following Glaubrecht (1996), taxa referred to the genus Pyrgulifera Meek, 1877 (see Meek 1871: 294) from Cretaceous deposits of North America and Europe, which have often been associated with thalassoid Paludomidae from Lake Tanganyika or that have sometimes been included in the genus Paludomus Swainson, 1840. In the works of Deshayes (1824–1837), Heer (1861), Morelet (1864), Sandberger (1870–1875), Neumayr (1883), Cossmann (1892, 1902), and Spijkerman et al. (2015) several fossils have been described in Paludomus or later attributed to Paludomus, which are not treated here because, in our opinion, they do not belong to the Paludomidae.

Nomenclaturally available as well as unavailable names (e.g., nomina nuda or infrasubspecific names) according to the rules of the fourth edition of the International Code of Zoological Nomenclature (ICZN 1999, hereafter in brief referred to as the “Code”) with subsequent amendments (see http://www.nhm.ac.uk/hosted-sites/iczn/code/index.jsp) are discussed following the general outline of the nomenclators of the Valvatidae Gray, 1840 (Turton and Gray 1840: 79) by Haszprunar (2014) and the Melanopsidae Adams & Adams, 1854 (Adams and Adams 1853–1854: 309) by Neubauer (2016).

The present nomenclator is divided into three parts that cover the family group, the genus group and the species group in the sense of the Code, respectively. All names in these three parts are listed in alphabetical order, except when the same family-, genus- or species-group name was introduced more than once. In this case names are ordered chronological or in cases of simultaneously published names by page precedence. The header of a section dealing with a name is the name of the taxon with mandatory changes as regulated by the Code (Art. 32 and 34). These are presented in the corrected version without mentioning. In cases where names were spelled incorrectly in the original publication, this spelling is given under ‘Original spelling’.

The status of a name that is unavailable is denoted in square brackets behind the taxon’s name. Behind a name that is either an objective junior synonym of an available name in the genus-group, an unjustified emendation or a junior homonym, its status as an objectively invalid name is denoted in square brackets. All remaining names are nomenclaturally available. However, depending on the taxonomic opinion of a researcher, such names may be taxonomically invalid names. A proposal whether a name is currently accepted or not is not made as this is, in our opinion, the task for future revisionary work. The original reference for a name is always provided. An authorship of a name deviating from the authorship of the work in which it was published, is only accepted in cases where clear evidence exists that person(s) other than the author(s) of the work, in which the name was published, were responsible for the name (Code Art. 50.1.1 and 50.1.3). In such cases, the name(s) of the person(s) responsible for the name is/are listed directly behind the name and the author(s) of the work, in which the name was published, is/are cited under ‘Original reference’. Information on type genera and type species, where relevant, are provided including references to the selection of type species (if subsequently designated). Names exclusively based on fossils are marked by a dagger. Information on type localities and type strata (for fossils only) are usually given exactly as provided in the original reference (in quotes); only in few cases also other sources are additionally provided. Additional information is given in square brackets (e.g., country, province). All other nomenclaturally relevant issues are discussed in the ‘Remarks’ under a name; however actions of a First Reviser in the sense of the Code are only discussed in cases where simultaneously published primary homonyms or different spellings of a name in the original publication are involved. Incorrect subsequent spellings are discussed under the respective correctly spelled names whenever these were noticed.

Part 1. Family-group names

In the family-group a total of 28 names are included in the nomenclator, of which 21 are available and seven unavailable names. Of the available names in the family-group, six are invalid for nomenclatural reasons.

Bathanaliidae Ancey, 1906

Original reference.Ancey (1906b: 245–246, 250).

Type genus.Bathanalia Moore, 1898.

Buccinopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Buccinopsidæ Nicolas, 1899.

Remarks. The name is apparently not based on the genus Buccinopsis Conrad, 1857 (see Conrad 1857: 158) but (following Bouchet et al. 2017: 52) introduced as a descriptive name alluding to the resemblance of species from Lake Tanganyika to the Buccinidae Rafinesque, 1815 (see Rafinesque 1815: 145).

Cancellopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Cancellopsidæ Nicolas, 1899.

Remarks. Unavailable because the name is not based on a genus-group name. It was established as a series within Tanganikidae Nourry, 1898. Following Bouchet et al. (2017: 58) the name is interpreted as descriptive alluding to the resemblance of species from Lake Tanganyika to the Cancellariidae Forbes & Hanley, 1851 (see Forbes and Hanley 1851: 360).

Cleopatrinae Pilsbry & Bequaert, 1927

Original reference.Pilsbry and Bequaert (1927: 249).

Original spelling. Cleopatrinæ Pilsbry & Bequaert, 1927.

Type genus.Cleopatra Troschel, 1857.

Remarks. Elevated to family rank by Germain (1933: 30).

Giraudidae Bourguignat, 1885 [Invalid]

Original reference.Bourguignat (1885a: 11, 61).

Original spelling. Giraudidæ Bourguignat, 1885.

Type genus.Giraudia Bourguignat, 1885.

Remarks. Invalid (Code Art. 39) because the type genus Giraudia Bourguignat, 1885 is a junior homonym of Giraudia Foerster, 1868 (Hexapoda, see Foerster 1868: 184); see also the remarks by Bouchet et al. (2017: 113).

Hauttecoeuridae Bourguignat, 1885

Original reference.Bourguignat (1885a: 10, 41).

Original spelling. Hauttecœuridæ Bourguignat, 1885.

Type genus.Hauttecoeuria Bourguignat, 1885.

Remarks. Used as the name of a subfamily (with the suffix –inae) and a tribe (with the suffix –ini) by Ponder and Bouchet (2005: 249); see also the remarks by Bouchet et al. (2017: 120).

Hilacanthidae Bourguignat, 1890 [Invalid]

Original reference.Bourguignat (1890: 125).

Original spelling.Hylacanthidae Bourguignat, 1890.

Type genus.Hilacantha Ancey, 1886.

Remarks. Originally spelled Hylacanthidae based on Hylacantha which is an unjustified emendation of Hilacantha Ancey, 1886 as indicated by the derivation of the name by Bourguignat (1890: 126, footnote on that page). The earlier use of Hylacantha by Bourguignat (1888: 23–24, pl. 9) is here interpreted as an incorrect subsequent spelling as there is no evidence of an intentional change of the name. The name was introduced as a replacement name of Tiphobiidae Bourguignat, 1886 based on Tiphobia Smith, 1880, which is unjustifiedly treated as a homonym of Typhobia Pascoe, 1869 (Hexapoda; see Pascoe 1869: 279) by Bourguignat (1890: 126) (see Bouchet and Rocroi 2005: 89).

Lavigeriidae Thiele, 1925

Original reference.Thiele (1925: 79).

Type genus.Lavigeria Bourguignat, 1888.

Remarks. Used as the name of a subfamily (with the suffix –inae) by Morrison (1954: 358); see also the remarks by Bouchet et al. (2017: 138).

Limnotrochidae Ancey, 1906

Original reference.Ancey (1906b: 245–246, 250).

Type genus.Limnotrochus Smith, 1880.

Littorinidopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Littorinidopsidæ Nicolas, 1899.

Remarks. The name is not based on a genus-group name. It was established as a series within Tanganikidae Nourry, 1898 by Nicolas (1899) as Littorrinidopsidae on p. 519 and as Littorinidopsidae on p. 525. Following Bouchet and Rocroi (2005: 101) and Bouchet et al. (2017: 143; as Littoridinopsidae) the name is interpreted as descriptive alluding to the resemblance of species from Lake Tanganyika to the Littorinidae Children, 1834 (see Children 1834: 110).

Muricopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Muricopsidæ Nicolas, 1899.

Remarks. The name is apparently not based on the genus Muricopsis Bucquoy & Dautzenberg, 1882 (see Bucquoy et al. 1882: 16, 19) but (following Bouchet and Rocroi 2005: 112) introduced as a descriptive name alluding to the resemblance of species from Lake Tanganyika to the Muricidae Rafinesque, 1815 (see Rafinesque 1815: 144).

Nassopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Nassopsidæ Nicolas, 1899.

Remarks. The name is apparently not based on the genus-group name Nassopsis Smith, 1890. It was established as a series within Tanganikidae Nourry, 1898 and following Bouchet and Rocroi (2005: 113) the name is interpreted as descriptive alluding to the resemblance of species from Lake Tanganyika to the ‘Nassaidae’ Swainson, 1835 (= Nassariidae Iredale, 1916) (see Swainson 1835: 18, 20, Iredale 1916: 82); see also the remarks by Bouchet et al. (2017: 160).

Nassopsidae Kesteven, 1903

Original reference.Kesteven (1903: 621, 634).

Type genus.Nassopsis Smith, 1890.

Remarks. Following Bouchet and Rocroi (2005: 113) the name is not accepted to be available from Nicolas (1899). Used as the name of a tribe with ending –ini Bouchet and Strong in Bouchet and Rocroi (2005: 113).

Naticidopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Naticidopsidæ Nicolas, 1899.

Remarks. The name is not based on a genus-group name. It was established as a series within Tanganikidae Nourry, 1898. Following Bouchet and Rocroi (2005: 113) the name is interpreted as descriptive alluding to the resemblance of species from Lake Tanganyika to the Naticidae Guilding, 1834 (see Guilding 1834: 29); see also the remarks by Bouchet et al. (2017: 160–161).

Neritopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Neritopsidæ Nicolas, 1899.

Remarks. The name used by Nicolas (1899) is apparently not based on the genus-group name Neritopsis Grateloup, 1832 (see Grateloup 1832: 125–126) and interpreted as descriptive alluding to the resemblance of species from Lake Tanganyika to the Neritidae Rafinesque, 1815 (see Rafinesque 1815: 144). Not Neritopsidae Gray, 1847 (see Gray 1847: 150).

Paludominae Stoliczka, 1868

Original reference.Stoliczka (1868: 206–207).

Type genus.Paludomus Swainson, 1840.

Remarks. Elevated to family rank by Pilsbry and Bequaert (1927: 248); used as the name of a tribe (with the suffix –eae) by Wenz (1939: 703).

Paramelaniidae Moore, 1898

Original reference.Moore (1898c: 315).

Original spelling.Paramelanidae Moore, 1898.

Type genus.Paramelania Smith, 1881.

Remarks. Stem formation in the original spelling incorrect. Used as the name of a subfamily (with the suffix –inae) by Thiele (1925: 83) and as the name of a tribe (with the suffix –eae) by Thiele (1928: 400).

Philopotamidae Stache, 1889

Original reference.Stache (1889: 107).

Type genus.Philopotamis Layard, 1855.

Remarks. Established originally as a subfamily of Melaniidae Children, 1823 (see Children 1823: 243) despite the use of the suffix –idae (see Bouchet and Rocroi 2005: 131).

Pyramidellopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Pyramidellopsidæ Nicolas, 1899.

Remarks. The name is not based on a genus-group name. It was established as a series within Tanganikidae Nourry, 1898. Following Bouchet and Rocroi (2005: 148) the name is interpreted as descriptive alluding to the resemblance of species from Lake Tanganyika to the Pyramidellidae Gray, 1840 (see Gray 1840: 117, 148); see also the remarks by Bouchet et al. (2017: 211).

Reymondiinae Bandel, 1998

Original reference.Bandel (1998: 273).

Type genus.Reymondia Bourguignat, 1885.

Rissopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Rissopsidæ Nicolas, 1899.

Remarks. The name is not based on a genus-group name. It was established as a series within Tanganikidae Nourry, 1898. Following Bouchet and Rocroi (2005: 152) the name is interpreted as descriptive alluding to the resemblance of species from Lake Tanganyika to the Rissoidae Gray, 1847 (see Gray 1847: 152); see also the remarks by Bouchet et al. (2017: 217).

Rumellidae Ancey, 1906

Original reference.Ancey (1906b: 245–246).

Type genus.Rumella Bourguignat, 1885.

Remarks. Used as the name of a tribe (with the suffix –ini) by Ponder and Bouchet (2005: 249) and Bouchet and Strong in Bouchet and Rocroi (2005: 161).

Spekiidae Ancey, 1906

Original reference.Ancey (1906b: 246).

Type genus.Spekia Bourguignat, 1879.

Remarks. Used as the name of a tribe (with the suffix –ini) by Ponder and Bouchet (2005: 249) and Bouchet and Strong in Bouchet and Rocroi (2005: 153). A name Spekiinae was indicated as intentionally new by Bandel (1998: 265). This represents a subsequent use of Ancey’s name, the indication as new was in error.

Syrnolopsidae Bourguignat, 1890

Original reference.Bourguignat (1890: 139).

Original spelling. Syrnolopsidæ Nicolas, 1899.

Type genus.Syrnolopsis Smith, 1880.

Remarks. Used as the name of a subfamily (with the suffix –inae) by Thiele (1928: 380), as the name of a superfamily (with suffix –oidea) by Golikov and Starborogatov (1987: 27) and as the name of a tribe (with the suffix –ini) by Ponder and Bouchet (2005: 249) and Bouchet and Strong in Bouchet and Rocroi (2005: 166).

Tanganyiciidae Nourry, 1898

Original reference.Nourry (1898: 302).

Original spelling.Tanganyikidae Nourry, 1898.

Type genus.Tanganyikia Bourguignat, 1885.

Remarks. Corrected to Tanganyiciidae Nourry, 1898 under the Code Art. 35.4.1); interpreted here as being based (by inference) on Tanganikia Bourguignat, 1885 which is an unjustified emendation for Tanganyicia Crosse, 1881. The interpretation of Bouchet and Rocroi (2005: 167) is not followed here who attribute the name to Nicolas (1899) (with year 1898) and interpret it as not being based on a genus but rather being descriptive to include all prosobranch gastropods of Lake Tanganyika. Apparently Nicolas (1899) was aware of Nourry’s (1898) introduction of the name Tanganyikidae as evidenced by the statement ‘C’est à ce point de vue que s’était placé M. Nourry, pour mettre en évidence non seulement la nécessité de réunir toutes ces formes Thalassoïdes dans une même famille, mais encore pour faire ressortir la descendance marine de la plupart des gastéropodes du Tanganyika’ (Nicolas 1899: 511 and footnote 1 on that page) and should therefore not be regarded as responsible for the name. A name Tanganyiciinae was indicated as intentionally new by Bandel (1998: 277). This represents a subsequent use of Nourry’s name, the indication as new was in error.

Tiphobiidae Bourguignat, 1886

Original reference.Bourguignat (1886: 143).

Original spelling.Tiphobidae Bourguignat, 1886.

Type genus.Tiphobia Smith, 1880.

Remarks. Stem formation in the original spelling incorrect. Used as the name of a subfamily (with the suffix –inae) by Morrison (1954: 373) and as the name of a tribe (with the suffix –ini) by Ponder and Bouchet (2005: 249) and Bouchet and Strong in Bouchet and Rocroi (2005: 172); see also the remarks by Bouchet et al. (2017: 244). A name Typhobiidae was indicated as an intentionally new name by Moore (1898c: 202) based on Typhobia (as used by Moore 1898a–c), which is an incorrect subsequent spelling of Tiphobia Smith, 1880. This is interpreted to represent a subsequent use of Bourguignat’s name here and therefore the indication as new was in error. Similarly, a name Tiphobiinae was indicated as intentionally new by Bandel (1998: 262). This also represents a subsequent use of Bourguignat’s name, the indication as new was in error.

Trochidopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 519, 525).

Original spelling. Trochidopsidæ Nicolas, 1899.

Remarks. The name is not based on a genus-group name. It was established as a series within Tanganikidae Nourry, 1898 (see Nicolas 1899). Following Bouchet and Rocroi (2005: 176) the name is interpreted as descriptive alluding to the resemblance of species from Lake Tanganyika to the Trochidae Rafinesque, 1815 (see Rafinesque 1815: 143); see also the remarks by Bouchet et al. (2017: 250–251).

Turbinidopsidae Nicolas, 1899 [Unavailable]

Original reference.Nicolas (1899: 525).

Original spelling. Turbinidopsidæ Nicolas, 1899.

Remarks. The name is not based on a genus-group name. It was established as a series within Tanganikidae Nourry, 1898 (see Nicolas 1899). The name is interpreted here as descriptive alluding to the resemblance of species from Lake Tanganyika to the Turbinidae Rafinesque, 1815 (see Rafinesque 1815: 144).

Part 2. Genus-group names

In the genus-group a total of 57 available names are included in the catalogue. Of the available names in the genus-group, 11 are invalid for nomenclatural reasons.

The genus-group name Tanalia Gray, 1847, type species Neritaaculeata Gmelin, 1791 by original designation, which has often been used as a genus-group name in Paludomidae Stoliczka, 1868, is not included in the nomenclator because its type species belongs to the Neritidae Rafinesque, 1815 (Neiber and Glaubrecht 2019a).

Anceya Bourguignat, 1885

Original reference.Bourguignat (1885a: 10, 14–15).

Type species.Anceyagiraudi Bourguignat, 1885, by monotypy.

Remarks. Gender: feminine.

Baizea Bourguignat, 1885

Original reference.Bourguignat (1885a: 10, 33–34).

Type species.Baizeagiraudi Bourguignat, 1885, by monotypy.

Remarks. Gender: feminine.

Bathanalia Moore, 1898

Original reference.Moore (1898a: 451–452, 454–455; 1898b: 165, 174; 1898c: 189, 192, 202–203).

Type species.Bathanaliahowesii Moore, 1898, by monotypy.

Remarks. Gender: feminine. The name Bathanalia was published simultaneously (March 1898) in three separate works by J. E. S. Moore (1998a–c). In Moore (1898a) only the spelling Bathanalia was used. In Moore (1898b) aside from the spelling Bathanalia, also the spellings Bathyanalia (p. 166, 170) and Bathynalia (p. 171), and in Moore (1998c) aside from Bathanalia, also the spelling Batanalia. In a subsequent work by the author, Moore (1898d), only the name Bathanalia was used. Therefore, Moore (1898d) is deemed here as First Reviser under the Code Art. 24.2.4 of the simultaneously published spellings of Moore (1898a–c) and Bathanalia is regarded as the correct original spelling.

Bourguignatia Giraud, 1885 [Invalid]

Original reference.Giraud (1885a: 193–194).

Type species.Bourguignatiaimperialis Giraud, 1885, by monotypy.

Remarks. Gender: feminine. Junior homonym of Bourguignatia Brusina, 1884 (Gastropoda, Neritidae; see Brusina 1884: 72).

Bridouxia Bourguignat, 1885

Original reference.Bourguignat (1885a: 10, 29).

Type species.Bridouxiagiraudi Bourguignat, 1885, by subsequent designation (Pilsbry and Bequaert 1927: 314).

Remarks. Gender: feminine.

Burtonilla Smith, 1904

Original reference.Smith (1904: 97).

Type species.Turbonillaterebriformis Smith, 1890, by monotypy.

Remarks. Gender: feminine.

Bythoceras Moore, 1898

Original reference.Moore (1898a: 451–452; 1898b: 165–166, 171).

Type species.Bythocerasiridescens Moore, 1898, by monotypy.

Remarks. Gender: neuter (Code Art. 30.1.2). The name Bythoceras was published simultaneously (March 1898) in two separate works by J.E.S. Moore (see Moore 1998a–b). In Moore (1898a) aside from the spelling Bythoceras also the spelling Bathoceras was used on p. 455. In Moore (1898b), only the spelling Bythoceras was used. In a subsequent work by the author, Moore (1898d), only the name Bythoceras was used. Therefore, Moore (1898d) is deemed here as First Reviser under the Code Art. 24.2.4 of the simultaneously published spellings of Moore (1898a, b) and Bythoceras is regarded as the correct original spelling.

Cambieria Bourguignat, 1885 [Invalid]

Original reference.Bourguignat (1885a: 42, 78).

Type species.Lithoglyphusrufofilosus Smith, 1880, by subsequent designation (Bourguignat 1890: 82).

Remarks. Gender: feminine. Introduced as a subgenus of Tanganikia Bourguignat, 1885. Objective junior synonym of Tanganyicia Crosse, 1881 (same type species).

Chytra Moore, 1898

Original reference.Moore (1898d: 307).

Type species.Limnotrochuskirkii Smith, 1880, by monotypy.

Remarks. Gender: feminine (Code Art. 30.1.2); from Greek χύτρα (= cooking pot). Chitra as used by Riedel (1993) is an incorrect subsequent spelling.

Cleopatra Troschel, 1857

Original reference. Troschel (1856–1863: 100–101).

Type species.Cyclostomabulimoides Olivier, 1804, by monotypy.

Remarks. Gender: feminine. Introduced as a subgenus of Paludina Férussac, 1812 (see Férussac 1812).

Coodea Bourguignat, 1891

Original reference.Bourguignat and Servain (1891: 165).

Type species.Rissoaponsonbyi Smith, 1889, by monotypy.

Remarks. Gender: feminine. Objective junior synonym of Horea Smith, 1889 (non Horea Bourguignat, 1888) and objective senior synonym of Lechaptoisia Ancey, 1894 (same type species).

Coulboisia Bourguignat, 1888

Original reference.Bourguignat (1888: 40, 78).

Type species.Stanleyagiraudi Bourguignat, 1885, by subsequent designation (Baker 1923: 174).

Remarks. Gender: feminine.

Edgaria Bourguignat, 1888

Original reference.Bourguignat (1888: 34, 78).

Type species.Tiphobianassapaucicostata Smith, 1881, by subsequent designation (Pilsbry and Bequaert 1927: 328).

Remarks. Gender: feminine.

† Ellinoria Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 11, 19, 61–62, 66, 75, fig. 50).

Type species.Ellinoriamichelae Van Damme & Pickford, 2003, by original designation.

Remarks. Gender: feminine.

Ganga Layard, 1855

Original reference.Layard (1855: 88).

Type species.Paludomusdilatata Reeve, 1852, by monotypy.

Remarks. Gender: feminine (Code Art. 30.2.3), from Sinhalese ගඟ (= river). Only the nominal species PaludomusdilatataReeve 1852 was originally included in the new genus Ganga by Layard (1855: 88). The nominal species PaludomusneritoidesReeve 1847 and PaludomusolivaceaReeve 1847 are deemed not to be originally included under the Code Art. 67.2.5, because their assignment to the genus was doubtful as indicated by the question marks behind the names. Hence, PaludomusdilatataReeve 1852 is the type species of Ganga Layard, 1855, by monotypy. Gauga as used by Cossmann (1909: 126) is an incorrect subsequent spelling.

Giraudia Bourguignat, 1885 [Invalid]

Original reference.Bourguignat (1885a: 11, 61–62).

Type species.Giraudiapraeclara Bourguignat, 1885, by subsequent designation (Pilsbry and Bequaert 1927: 311).

Remarks. Gender: feminine. A junior homonym of Giraudia Foerster, 1868 (Hexapoda, see Foerster 1868: 184). Girandia is an incorrect subsequent spelling by Smith (1889).

Hauttecoeuria Bourguignat, 1885

Original reference.Bourguignat (1885a: 10, 46–48).

Original spelling.Hauttecœuria Bourguignat, 1885.

Type species.Hauttecoeuriasoluta Bourguignat, 1885, by subsequent designation (Germain 1908: 37).

Remarks. Gender: feminine.

Hemimitra Swainson, 1840

Original reference.Swainson (1840: 199–200, 202, 340).

Type species.PaludomusretusaSwainson 1840, by monotypy.

Remarks. Gender: feminine. Introduced as a subgenus of PaludomusSwainson 1840. See Neiber and Glaubrecht (2019a) for the identity of the type species.

Heteropoma Benson, 1856 [Invalid]

Original reference.Benson (1856: 494).

Type species.Paludomussulcata Reeve, 1847, by subsequent designation (Neiber and Glaubrecht 2019a).

Remarks. Gender: neuter (Code Art. 30.1.2), from Greek ἕτερο- (= other, different) and πῶμα, neuter (= lid). Objective junior synonym of Philopotamis Layard, 1855 (same type species).

† Heynderycxia Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 11, 19, 59–62, 66–67, 75, 77, figs 49a–d, 54e).

Type species.Heynderycxiacolumellaris Van Damme & Pickford, 2003, by original designation.

Remarks. Gender: feminine.

Hilacantha Ancey, 1886 [Invalid]

Original reference.Ancey (1886: 292–293).

Type species.Tiphobiahorei Smith, 1880, by monotypy for the replaced name Tiphobia Smith, 1880.

Remarks. Gender: feminine (Code Art. 30.1.2), from Greek ὕλη- (= wooden) and ἄκανθα, feminine (= thorn, spine). Unnecessary nom. nov. pro Tiphobia Smith, 1880. Hylacantha (Bourguignat 1888: 23–24, pl. 9) is an incorrect subsequent spelling, Hylacantha (Bourguignat 1890: 126) is an unjustified emendation.

Hirthia Ancey, 1898

Original reference.Ancey (1898: 142).

Type species.Hirthialittorina Ancey, 1898, by subsequent designation (Pilsbry and Bequaert 1927: 328).

Remarks. Gender: feminine. Hirtia as used by Van Damme and Pickford (2003) is an incorrect subsequent spelling.

Horea Bourguignat, 1888

Original reference.Bourguignat (1888: 27–28).

Type species.Horeatanganikana Bourguignat, 1888, by monotypy.

Remarks. Gender: feminine.

Horea Smith, 1889 [Invalid]

Original reference.Smith (1889: 175).

Type species.Rissoaponsonbyi Smith, 1889, by monotypy.

Remarks. Gender: feminine. Established as a subgenus of Rissoa Desmarest, 1814 (Desmarest 1814: 7). Invalid, junior homonym of Horea Bourguignat, 1888. The name Horea was introduced independently by Bourguignat (1888) and Smith (1889) for different taxa in honour of Captain Edward Coode Hore.

Hylacantha Bourguignat, 1890 [Invalid]

Original reference.Bourguignat (1890: 126).

Remarks. Gender: feminine, see Remarks under Hilacantha Ancey, 1886. Unjustified emendation of Hilacantha Ancey, 1886 as evidenced by the derivation of the name by Bourguignat (1890: 126, footnote on that page), see also Martens (1891: 7–8). The earlier use of Hylacantha by Bourguignat (1888: 23–24, pl. 9) is here interpreted as an incorrect subsequent spelling of Hilacantha Ancey, 1886 as there is no evidence of an intentional change of the name. Tylacanæha (Nicolas 1899: 512) is here also regarded as an incorrect subsequent spelling, although hardly recognizable anymore, of Hylacantha Bourguignat, 1890. The work of Nicolas (1899) contains numerous typesetting errors and apparently that author was aware of the name Tiphobia introduced by Smith (1880a) and its replacement name Hilacantha introduced by Ancey (1886) in the emendated form Hylacantha by Bourguignat (1890) as suggested by the statement “L’énumeration se continue par un genre des plus caracteristques, Tiphobia ou Tylacanæha, qui par l’opercule est une Melanidæ, tandis que l’absence d’épiderme l’en sépare, d’eû une grande indecision pour cette espèce” (Nicolas 1899: 512) in combination with the synonymous usage of the names Hylacanthalongirostris and Tiphobialongirostris (Nicolas 1899: 515).

Joubertia Bourguignat, 1888

Original reference.Bourguignat (1888: 32, 78).

Type species.Paramelaniabaizeana Bourguignat, 1885, by subsequent designation (Pilsbry and Bequaert 1927: 324).

Remarks. Gender: feminine.

Lavigeria Bourguignat, 1888

Original reference.Bourguignat (1888: 31, 33–34, 78).

Type species.Tiphobianassagrandis Smith, 1881, by subsequent designation (Pilsbry and Bequaert 1927: 324).

Remarks. Gender: feminine. Lavigieria is an incorrect subsequent spelling by Martens (1891: 9).

Lechaptoisia Ancey, 1894 [Invalid]

Original reference.Ancey (1894: 29).

Type species.Rissoaponsonbyi Smith, 1889, by typification of replaced name.

Remarks. Gender: feminine. Nom. nov. pro Horea Smith, 1889. Objective junior synonym of Coodea Bourguignat, 1891 (same type species).

Leloupiella nom. nov.

http://zoobank.org/17726764-61CF-4026-905E-7149C3B1E6EF

Type species.Giraudiaminima Smith, 1908, by typification of the replaced name.

Remarks. New replacement name for the preoccupied name Stormsia Leloup, 1953 (not Stormsia Bourguignat, 1891). See also Remarks under Stormsia Bourguignat, 1891 and Stormsia Leloup, 1953.

Limnotrochus Smith, 1880

Original reference.Smith (1880b: 425).

Type species.Limnotrochusthomsoni Smith, 1880, by subsequent designation (Pilsbry and Bequaert 1927: 318).

Remarks. Gender: masculine.

Martelia Dautzenberg, 1908

Original reference.Dautzenberg (1908: 328).

Type species.Marteliatanganyicensis Dautzenberg, 1908, by monotypy.

Remarks. Gender: feminine.

Mysorelloides Leloup, 1953

Original reference.Leloup (1953: 25, 87–92, 115, 232, pI. 5, fig. 3, figs 11, 41, 42, 57K, 62, 72CC)

Type species.Bithyniamultisulcata Bourguignat, 1888, by monotypy.

Remarks. Gender: feminine, originally combined with a feminine adjectival species-group name (Code Art. 30.1.4.4).

Nassopsidia Martens, 1897

Original reference.Martens (1897: 208).

Type species.Paramelaniacrassilabris Bourguignat, 1885, by subsequent designation (Pilsbry and Bequaert 1927: 328).

Remarks. Gender: feminine. Introduced as a subgenus of Paramelania Smith, 1881.

Nassopsis Smith, 1890 [Invalid]

Original reference.Smith (1890: 93).

Type species.Tiphobianassagrandis Smith, 1881, by original designation.

Remarks. Gender: feminine (Code Art. 30.1.2), from Latin nassa, feminine (= weel) and Greek ὄψις, feminine (= appearance). Objective junior synonym of Lavigeria Bourguignat, 1888 (same type species).

Odontochasma Tomlin, 1930

Original reference.Tomlin (1930: 23).

Type species.Tanaliastomatodon Benson, 1862, by monotypy for the replaced name Stomatodon Benson, 1862.

Remarks. Gender: neuter (Code Art. 30.1.1). Nom. nov. pro Stomatodon Benson, 1862 (non Stomatodon Seeley, 1861).

Paludomus Swainson, 1840

Original reference.Swainson (1840: 198–201, 210, 340).

Type species.Melaniaconica Griffith & Pidgeon, 1833 in Griffith and Pidgeon (1833–1834) (non Melaniaconica Say, 1821, see Say 1821: 176–177), by subsequent designation (Gray 1847: 155).

Remarks. Gender: feminine (Code Art. 30.1.1), from Latin palus, feminine (= swamp) and domus, feminine (= house). Swainson (1840) also combined the name with feminine adjectival species-group names. Paludimus as used by Kobelt (1909) is an incorrect subsequent spelling.

Paramelania Smith, 1881

Original reference.Smith (1881b: 559).

Type species.Tiphobiadamoni Smith, 1881, by subsequent designation (Bourguignat 1890: 162).

Remarks. Gender: feminine. Introduced as a subgenus of Tiphobia Smith, 1880. The subsequent designation of the same taxon as type species of Paramelania Smith, 1881 by Pilsbry and Bequaert (1927: 320) is invalid. Paramellania as used by Moore (1903) is an incorrect spelling.

Philopotamis Layard, 1855

Original reference.Layard (1855: 88–89).

Type species.Paludomussulcata Reeve, 1847, by subsequent designation (Nevill 1884: 297).

Remarks. Gender: feminine (Code Art. 30.1.3). Cossmann (1909: 126) gave as type species Philopotamisregalis Layard, 1855; this act is invalid.

Ponsonbya Ancey, 1890

Original reference.Ancey (1890: 347).

Type species.Ponsonbyaleucoraphe Ancey, 1890, by monotypy.

Remarks. Gender: feminine.

Potadomoides Leloup, 1953

Original reference.Leloup (1953: 102–105).

Type species.Potadomoidespelseneeri Leloup, 1953, by monotypy.

Remarks. Gender: masculine (Code Art. 30.1.4.4).

Pseudocleopatra Thiele, 1928

Original reference.Thiele (1928: 394).

Type species.Pseudocleopatratogoensis Thiele, 1928, by monotypy.

Remarks. Gender: feminine.

Randabelia Bourguignat, 1888

Original reference.Bourguignat (1888: 31, 78).

Type species.Paramelaniahamyana Bourguignat, 1885, by subsequent designation (Pilsbry and Bequaert 1927: 327).

Remarks. Gender: feminine.

Reymondia Bourguignat, 1885

Original reference.Bourguignat (1885a: 11, 61–62).

Type species.Melaniahorei Smith, 1880, by subsequent designation (Pilsbry and Bequaert 1927: 312).

Remarks. Gender: feminine. Attributed to Giraud, in litt. but J.-R. Bourguignat alone was responsible for the name.

Rivulina Lea & Lea, 1851

Original reference.Lea and Lea (1851: 196–197).

Type species.Melaniamodicella Lea & Lea, 1851, by subsequent designation (Neiber and Glaubrecht 2019a).

Remarks. Gender: feminine.

Rumella Bourguignat, 1885

Original reference.Bourguignat (1885a: 11, 89–90).

Type species.Rumellagiraudi Bourguignat, 1885, by subsequent designation (Baker 1923: 174).

Remarks. Gender: feminine.

Spekia Bourguignat, 1879

Original reference.Bourguignat (1879: 27–28).

Type species.Lithoglyphuszonatus Woodward, 1859, by monotypy.

Remarks. Gender: feminine. Spekea as used by Hoyle (1886: 88) and Martens (1891: 8) are incorrect subsequent spellings.

Stanleya Bourguignat, 1885

Original reference.Bourguignat (1885a: 11, 86–87).

Type species.Lithoglyphusneritinoides Smith, 1880, by original designation.

Remarks. Gender: feminine. The name Stanleyaneritoides as used by Bourguignat (1885, 1888) has to be interpreted as an incorrect subsequent spelling of the name of the nominal taxon Lithoglyphusneritinoides Smith, 1880 and should not be regarded (as e.g. by Ancey 1906b: 254 or Baker 1923: 174) as a newly introduced name by Bourguignat (1888) (although the author attributed the taxon to himself in that publication) because Bourguignat (1885: 87, footnote) explicitly referred to Smith (1880: 426) and Smith (1881: 287) and also used the misspelled name Lithoglyphusneritoides in the synonymy when first using the name Stanleyaneritoides. Therefore, Stanleya is here interpreted as not having been introduced again by Bourguignat (1888) and the fixation of Stanleyaneritoides with author Bourguignat (1888) by Baker (1923: 174) is not valid.

Stomatodon Benson, 1862 [Invalid]

Original reference.Benson (1862: 414).

Type species.Tanaliastomatodon Benson, 1862, by monotypy.

Remarks. Gender: masculine (Code Art. 30.1.2), from Greek στόμα, neuter (= mouth) and ὀδών, masculine (= tooth). The name Stomatodon Benson, 1862 was conditionally proposed by Benson (1862: 414). Benson (1862: 414) stated: “Unfortunately, all the specimens were deficient in the operculum, which, when examined, may possibly authorize its transfer to a new genus, in which case the specific name may fairly be employed to designate it.” And further below on the page and on the next page: “Stomatodon seems partly to supply one of the absent links, inasmuch as its operculum must necessarily be provided with a basal projection, while its constriction is also likely to be unguiculate.” Stomatodon Benson, 1862 is, however, a junior homonym of Stomatodon Seeley, 1861 (see Seeley 1861: 293; Gastropoda, Ringiculidae Philippi, 1853, see Philippi 1853: 190). It has been replaced by Odontochasma Tomlin, 1930 (see Tomlin 1930: 23 and Neiber and Glaubrecht 2019a).

Stormsia Bourguignat, 1891

Original reference.Bourguignat and Servain (1891: 165).

Type species.Syrnolopsiscarinifera Smith, 1889, by monotypy.

Remarks. Gender: feminine. See also Remarks under Stormsia Leloup, 1953.

Stormsia Leloup, 1953 [Invalid]

Original reference.Leloup (1953: 217–219).

Type species.Giraudiaminima Smith, 1908, by monotypy.

Remarks. Gender: feminine. Stormsia Leloup, 1953 is a junior homonym of Stormsia Bourguignat, 1891. Stormsia Bourguignat, 1891 was made available in a note in a biographic and bibliographic work (Servain and Bourguignat 1891) and has been overlooked by most subsequent authors. Although Stormsia Bourguignat, 1891 has, to our knowledge, not been used as an accepted name since its introduction, it had been mentioned by Pilsbry and Bequaert (1927) and Germain (1919b) in the synonymy of Syrnolopis Smith, 1880. Both, Bourguignat and Leloup apparently dedicated their generic names to Émile Pierre Joseph Storms who established a European presence in the Lake Tanganyika region. Leloup (1953) explicitly introduced Stormsia as a new generic name for the single nominal taxon Giraudiaminima Smith, 1908, while Bourguignat in Bourguignat and Servain (1891) had introduced Stormsia for the nominal taxon Syrnolopsiscarinifera Smith, 1889. The name Stormsia with direct reference to Leloup (1953) has been used by Brown (1980, 1994), Bandel (1998, 2006), West et al. (2003), Robertson (2007), Glaubrecht (2008), Wagner et al. (2009), and Neiber and Glaubrecht (2019b). The name Stormsiaminima (Smith, 1908) has been used by Brown and Mandahl-Barth (1987), Glaubrecht (1994, 1996), Baillie and Groombridge (1996), Alin et al. (1999), Irvine and Donohue (1999), Allison et al. (2000), West and Michel (2000), Van Damme and Pickford (2003), Glaubrecht and Strong (2004), Wilson et al. (2004), Levêque et al. (2006), Strong and Glaubrecht (2007), Avanesyan (2008), Strong and Glaubrecht (2008), Fermon (2009), Nicayenzi et al. (2010), and Brown (2014), but the generic name was unfortunately not referenced by these authors and is therefore, although probable, not without doubt attributable to Leloup (1953). Thus, the conditions of the Code (Art 23.9.1.1 and Art. 23.9.1.2) are, in our opinion, not met. The new replacement name Leloupiella nom. nov. is therefore proposed to honour Eugène Leloup’s contributions to the systematics of the malacofauna of Lake Tanganyika; gender: feminine.

Syrnolopsis Smith, 1880

Original reference.Smith (1880b: 217–219).

Type species.Syrnolopsislacustris Smith, 1880, by monotypy.

Remarks. Gender: feminine, see Remarks under Nassopsis Smith, 1890. Syznolopsis (Nicolas 1899: 510) is an incorrect subsequent spelling.

† Taeniodomus Krause, 1897

Original reference. Krause (1887: 209, 213–214).

Type species.Taeniodomusgracilis Krause, 1897, by subsequent designation (Cossmann 1898: 105).

Remarks. Gender: feminine, see Remarks under Paludomus Swainson, 1840.

Tanganikia Bourguignat, 1885 [Invalid]

Original reference.Bourguignat (1885a: 10, 41–42).

Remarks. Gender: feminine. Tanganikia Bourguignat, 1885 is an unjustified emendation of Tanganyicia Crosse, 1881. Tanganikea as used by Martens (1891: 8) is an incorrect subsequent spelling.

Tanganyicia Crosse, 1881

Original reference.Crosse (1881: 123–124).

Type species.Lithoglyphusrufofilosus Smith, 1880, by original designation.

Remarks. Gender: feminine. Tanganyikia as used by Moore (1899) and Germain (1908) are incorrect subsequent spellings.

Tiphobia Smith, 1880

Original reference.Smith (1880a: 348).

Type species.Tiphobiahorei Smith, 1880, by monotypy.

Remarks. Gender: feminine. Typhobia is an incorrect subsequent spelling by Moore (1898a–c) and Thiphobia by Riedel (1993).

Vinundu Michel, 2004

Original reference.Michel (2004: 2, 4).

Type species.Vinunduwestae Michel, 2004, by original designation.

Remarks. Gender: feminine (Code Art. 30.2.2). Vinundu Michel & Todd, 2003 in West et al. (2003: 76–77) is unavailable because the work includes a disclaimer stating “this publication is not deemed to be valid for formal taxonomic/nomenclatural purposes”. The name was also used by Van Damme and Pickford (2003: 2) as a nomen nudum because no diagnosis, definition, or description was given.

Zanguebaria Fischer, 1881

Original reference.Fischer (1880–1887: 2, 4).

Type species.Melaniaamaena Morelet, 1851, by subsequent designation (Pilsbry and Bequaert 1927: 288).

Remarks. Gender: feminine.

Part 3. Species-group names

In the species-group a total of 499 names are included, of which 463 are available but 21 are invalid for nomenclatural reasons.

Paludomusabbreviata Reeve, 1852

Original reference.Reeve (1852: 127).

Original spelling.Paludomusabbreviatus Reeve, 1852.

Type locality. “Ceylon” [Sri Lanka].

Paludomusacuta Reeve, 1852

Original reference.Reeve (1852: 127).

Original spelling.Paludomusacutus Reeve, 1852.

Type locality. “Near Pondicherry” [India, near Puducherry].

† Cleopatraadami nom. nov.

http://zoobank.org/BE605896-D974-47AA-8C40-31CB9AF92C38

Type locality. “Rive du ravin de Nyakasia, 65 à 80 m sur lac” [Uganda, river of Nyakasia Ravine, 65 to 80 m above Lake Albert].

Type stratum. “Pleistocène inférieur, Série de Kaiso” [Lower Pleisocene, Kaiso Series].

Type material. Holotype, Musée royal de l’Afrique centrale, Tervuren MC 2780 (Adam 1957, pl. 4, fig. 1).

Remarks. The species was classified as Cleopatracylindrica (Adam, 1957) by Van Damme and Pickford (2003). This name is a secondary junior homonym of Cleopatracridlandicylindrica Mandahl-Barth, 1954 and because no junior synonyms exist, the new replacement name Cleopatraadami nom. nov. is introduced here (see also Remarks under Viviparuscylindricus Adam, 1957). The replacement name is chosen in honor of William Adam, the former Director of the Malacology Department of the Royal Belgian Institute of Natural Sciences, who was the first to describe the species (Adam 1957).

Anceyaadmirabilis Bourguignat, 1889

Original reference.Bourguignat (1889: 119–120, pl. 7, figs 10, 11).

Type locality. “Entre Mpala et Mlilo” [Democratic Republic of the Congo, between Mpala and Mlilo (= Moliro), Lake Tanganyika].

Paludomusadustus Brot, 1880 [Unavailable]

Original reference.Brot (1880: 25).

Remarks. Attributed to Swainson; published in the synonymy of Paludomusstephanus Benson, 1836; unavailable (Code Art. 11.6).

Melaniaaegyptiaca Reeve, 1860

Original reference.Reeve (1860: pl. 34 fig. 227 and explanation of the plate).

Original spelling.Melania Ægyptiaca Reeve, 1860.

Type locality. “Egypt”.

Bithyniabulimoidesaegyptiaca Frauenfeld, 1864

Original reference.Frauenfeld (1864: 565, 583).

Original spelling.Bythiniabulimoides var. Aegyptiaca Frauenfeld, 1864.

Type locality. “Aegypten” [Egypt].

Remarks. Introduced by Frauenfeld (1864: 583) as a variety of Bithyniabulimoides (Olivier, 1804) and on p. 565 the name is mentioned with some descriptional terms, thus, satisfying the Code (Art. 12.1).

Paludomusaerea Reeve, 1852

Original reference.Reeve (1852: 128).

Original spelling.Melania æreus Reeve, 1852.

Type locality. “Mountain streams of Ceylon” [Sri Lanka].

Remarks.Tanaliaaerens as used by Adams and Adams (1853–1854) is an incorrect subsequent spelling.

Paludomusafricana Martens, 1878

Original reference.Martens (1878: 297, pl. 2, figs 11–13).

Original spelling.PaludomusAfricana Martens, 1878.

Type locality. “Finboni” [Kenya, Finboni, near the coast, between Mombassa and Taita].

Paludomusajanensis Morelet, 1860

Original reference.Morelet (1860b: 110, pl. 6, fig. 10).

Original spelling.PaludomusAjanensis Morelet, 1860.

Type locality. “Dans les eaux saumâtres d’Hafoun (Raz Hhafoun), à trente lieues au sud du cap Guardafui” [Somalia, brackish waters at Hafun, south of Cape Guardafui ; but probably from the Seychelles].

Cleopatraalhiensis Verdcourt, 1957 [Unavailable]

Original reference.Verdcourt (1957: 9).

Remarks. Mentioned under Cleopatraathiensis Verdcourt, 1957 as a manuscript name attributed to H. B. Preston; unavailable (Code Art. 11.6).

Paramelaniaalphonsi Bourguignat, 1890

Original reference.Bourguignat (1890: 217–218).

Original spelling.ParamelaniaAlphonsi Bourguignat, 1890.

Type locality. “Sur la plage de Mlilo” [Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Melaniaamaena Morelet, 1851

Original reference.Morelet (1851a: 220–221).

Original spelling.Melaniaamæna Morelet, 1851.

Type locality. “Palustria ad orientem insulæ Madagascar” [Madagascar, swamps in the south of the island] (Morelet (1851a, b); subsequently stated to be “Zanzibar et des Séchelles” [Tanzania, Zansibar; Seychelles] (Morelet (1860b).

Remarks. Also published as a new species in Morelet (1851b: 192–193, pl. 5, fig. 9), which is considered here as a subsequent use.

Syrnolopsisanceyana Bourguignat, 1885

Original reference.Bourguignat (1885a: 20).

Type locality. “Sur la plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Hylacanthaanceyi Bourguignat, 1891

Original reference.Bourguignat and Servain (1891: 165).

Original spelling.HylacanthaAnceyi Bourguignat, 1891.

Type locality. Not given.

Syrnolopsisanceyi Germain, 1905 [Invalid]

Original reference.Germain (1905a: 259).

Original spelling.SyrnolopsisAnceyi Bourguignat, 1905.

Remarks. Unjustified emendation of Syrnolopsisanceyana Bourguignat, 1885. Germain (1905a) changed the suffix of several species-group names ending in -iana or -ana introduced by Bourguignat (1885, 1888). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Paludomusandersoniana Nevill, 1877

Original reference.Nevill (1877: 35).

Original spelling.PaludomusAndersoniana Nevill, 1877.

Type locality. “Mandalay, Ava, Bhamô, Kabyuet, and Myadoung” [Myanmar, Mandalay, Ava, Bhamô, Kabuyet, Myadoung].

† Cleopatraangulata Williamson, 1979

Original reference.Williamson (1979: 114, 227b, 238b, fig. 65t).

Type locality. “Area 110, east of lake Turkana, north Kenya”.

Type stratum. “Fauna 52, in the lower Member of the Koobi Fora Formation”.

Paludomusannandalei Preston, 1909

Original reference.Preston (1909: 277–278).

Type locality. “Tenmalai, W. Ghats (W. side), Travancore (rocky mountain stream)” [India, Kerala, Thenmala, Western Ghats, western side].

† Cleopatraarambourgi Roger, 1944

Original reference.Roger (1944: 127–128, pl. 1, figs 15–17).

Original spelling.CleopatraArambourgi Roger, 1944.

Type locality. “De la surface du plateau, entre Kalam et Bourillé” [Ethiopia, Omo Plateau].

Type stratum. Not given.

Remarks. Attributed to L. Germain, but J. Roger alone was responsible for the description.

Paramelaniaarenarum Bourguignat, 1888

Original reference.Bourguignat (1888: 39–40, pl. 17, figs 3, 4).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Cleopatraathiensis Verdcourt, 1957

Original reference.Verdcourt (1957: 8–9, fig. 2).

Type locality. “Kenya, Athi Plains”.

† Cleopatraferrugineaattenuata Williamson, 1979

Original reference.Williamson (1979: 227b, 238a, pl. 2, figs 7–9).

Type locality. “Area 202, east of lake Turkana, north Kenya”.

Type stratum. “Fauna 12C, from between the upper and lower tuffs comprising the Suregei tuff complex, basal Koobi Fora Formation”.

Remarks. A potadomid species according to Van Bocxlaer et al. (2008).

Cleopatraaurocincta Martens, 1879

Original reference.Martens (1879: 103)

Type locality. “Bagamojo, gegenüber von Zanzibar” [Tanzania, Bagamoyo].

Paludomusbaccula Reeve, 1852

Original reference.Reeve (1852: 128).

Type locality. “Branch of the Ganges” [India or Bangladesh, Ganges River].

Remarks.Paludomusbacula is an incorrect subsequent spelling by Layard (1855) and Adams and Adams (1853–1854).

Paramelaniabaizeana Bourguignat, 1885

Original reference.Bourguignat (1885a: 74).

Type locality. “Près de Kapampa, sur la côte sud-ouest du lac” [Democratic Republic of the Congo, Kapampa, western shore of Lake Tanganyika].

Anceyabella Pilsbry & Bequaert, 1927

Original reference.Pilsbry and Bequaert (1927: 238, figs 36, 37).

Type locality. “Lake Tanganyika”.

Pseudocleopatrabennikei Mandahl-Barth, 1974

Original reference. Mandahl-Barth (1974: 355–356, fig. 2a).

Type locality. “River Zaire between Mimosa Island and Monkey Island, off Kinshasa” [Democratic Republic of the Congo].

Stomatodonbensoni Brot, 1880

Original reference.Brot (1880: 13–14, pl. 5, fig. 2, 2a, 2b).

Original spelling.StomatodonBensoni Brot, 1880.

Type locality. “Südliches Indien, Travancore (Benson), Malabar (Coll. Mea)” [India, Kerala, Travancore; Malabar].

Cleopatrabequaerti Dautzenberg & Germain, 1914

Original reference.Dautzenberg and Germain (1914: 59–60, pl. 4, figs 1–6).

Original spelling.CleopatraBequaerti Dautzenberg & Germain, 1914.

Type locality. “Kindu, Lualaba, 3° lat. S” [Democratic Republic of the Congo, Kindu, Lualaba River].

Cleopatrabulimoidesbicarinata Pallary, 1924

Original reference.Pallary (1924: 33).

Original spelling.Cleopatrabulimoidesvar.bicarinata Pallary, 1924.

Type locality. “Égypte” [Egypt].

Paludomusbicincta Reeve, 1852

Original reference.Reeve (1852: 129).

Original spelling.Paludomusbicinctus Bourguignat, 1852.

Type locality. “Mountain streams of Ceylon” [Sri Lanka].

Paludomusbifasciatus Adams & Adams, 1854 [Unavailable]

Original reference.Adams and Adams (1853–1854: 340).

Remarks. Attributed to Reeve. Nomen nudum, introduced without description, definition, or indication (Code Art. 12.1).

Paludomusbilineata Troschel, 1857

Original reference. Troschel (1856–1863: 101, pl. 7, fig. 7).

Type locality. Not given.

Original spelling.Paludomusbilineatus Troschel, 1857.

Remarks. Troschel (1857) in Troschel (1856–1863: 101, pl. 7, fig. 7) described and figured a radula under the name Paludomusbilineatus, attributing the name to L. A. Reeve. However, Reeve (1847, 1852) did not introduce the name Paludomusbilineatus, but instead Paludomusbicinctus Reeve, 1852 with a similar meaning. Although the name P.bilineatus is probably the result of a lapsus memoriae, it is deemed available here.

Cleopatrabulimoidesbilirata Germain, 1918

Original reference.Germain (1918: 444–445).

Original spelling.Cleopatrabulimoides variété bilirata Germain, 1918.

Type locality. “Rosières [= Rosaires = Abramat], sur le Nil Bleu” [Sudan, Er Roseires, Blue Nile].

Paludomusbinaluanensis Thiele, 1928

Original reference. Thiele (1828: 392, pl. 8, fig. 17).

Type locality. “Von Binaluan auf Nord-Palawan” [Philippines, north Palawan, Binaluan].

Paludomusblanfordiana Nevill ,1877

Original reference.Nevill (1877: 37).

Original spelling.PaludomusBlanfordiana Nevill, 1877.

Type locality. “Ava” [Myanmar, Ava].

Paludomusborneensis Nevill, 1884

Original reference.Nevill (1884: 292).

Type locality. “Sadong, Sarawak” [Malaysia, Borneo, Sarawak, Sadong Jaya].

Paramelaniabourguignati Bourguignat, 1885

Original reference.Bourguignat (1885a: 73–74).

Type locality. “Plage de Kapampa” [Democratic Republic of the Congo, beach of Kapampa, western shore of Lake Tanganyika].

Remarks. Attributed to V. Giraud by Bourguignat (1885a: 73), but Bourguignat alone was responsible for the description. Therefore, Bourguignat is the author of the name (Code Art. 50.1).

Tiphobiabourguignati Bourguignat, 1886

Original reference.Bourguignat (1886: 148–150, pl. 6, figs 5–7).

Type locality. “Dans l’anse de Kamangu, un peu S.-O. des îlots Kilira Chakabala“ [Democratic Republic of the Congo, Kamangu Bay, southwest of the Kilira Chakabala Islands]].

Remarks. Attributed to L. Joubert by Bourguignat (1886: 148) but Bourguignat alone was responsible for the description. Therefore, Bourguignat is the author of the name (Code Art. 50.1).

Bourguignatiabridouxi Bourguignat, 1888

Original reference.Bourguignat (1888: 29, pl. 12, figs 1–4).

Original spelling.BourguignatiaBridouxi Bourguignat, 1888.

Type locality. “Sur les plages de la côte occidentale” [Democratic Republic of the Congo, western shores of Lake Tanganyika].

Viviparabridouxiana Bourguignat, 1888

Original reference.Bourguignat (1888: 13–14, pl. 4, fig. 2).

Original spelling.ViviparaBridouxiana Bourguignat, 1888.

Type locality. “De la côte nord-occidentale du lac” [Democratic Republic of the Congo, north-western shore of Lake Tanganyika].

Hauttecoeuriabridouxiana Bourguignat, 1888

Original reference.Bourguignat (1888: 21–22, pl. 8, figs 26–28).

Original spelling.Hauttecœuria Bridouxiana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Reymondiabridouxiana Bourguignat, 1888

Original reference.Bourguignat (1888: 27–28, pl. 11, figs 14, 15).

Original spelling.ReymondiaBridouxiana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Viviparabrincatiana Bourguignat, 1888

Original reference.Bourguignat (1888: 13–14, pl. 4, fig. 1).

Original spelling.ViviparaBrincatiana Bourguignat, 1888.

Type locality. “De la côte nord-occidentale du lac” [Democratic Republic of the Congo, north-western shore of Lake Tanganyika].

Hauttecoeuriabrincatiana Bourguignat, 1888

Original reference.Bourguignat (1888: 19, pl. 7, figs 20, 21).

Original spelling.Hauttecœuria Brincatiana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Cleopatrabroecki Putzeys, 1899

Original reference.Putzeys (1899: 60, fig. 16).

Original spelling.CleopatraBroecki Putzeys, 1899.

Type locality. “De la rivière Aruwimi, affluent du Congo” [Democratic Republic of the Cong, Aruwimi River, tributary of Congo River].

Paludomusbroti Issel, 1874

Original reference.Issel (1874: 455–456, pl. 7, figs 19, 20).

Original spelling.PaludomusBroti Issel, 1874.

Type locality. “Sadong, Sarawak” [Malaysia, Borneo, Sarawak, Sadong Jaya].

Cyclostomabulimoides Olivier, 1804

Original reference.Olivier (1804: 39, pl. 31, fig. 6).

Type locality. “Le Kalidje” [Egypt, Le Kalidje near Alexandria].

Paludomusburmanica Nevill, 1877

Original reference.Nevill (1877: 36).

Original spelling.PaludomusBurmanica Nevill, 1877.

Type locality. “Yaylaymaw and Mandalay” [Myanmar, Yaylaymaw and Mandalay].

Hauttecoeuriaburtoni Bourguignat, 1888

Original reference.Bourguignat (1888: 19, pl. 7, figs 1–3).

Original spelling.Hauttecœuria Burtoni Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paramelaniabythiniformis Bourguignat, 1888

Original reference.Bourguignat (1888: 35–36, pl. 15, figs 26, 27).

Original spelling.ParamelaniaBythiniformis Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Rumellacallifera Bourguignat, 1888

Original reference.Bourguignat (1888: 39–40, pl. 17, figs 23–25).

Type locality. “De la côte occidentale” [Democratic Republic of the Congo, Lake Tanganyika, western shore].

Lavigeriacallista Bourguignat, 1888

Original reference.Bourguignat (1888: 33–34, pl. 14, fig. 2).

Type locality. “Sur les rives nord et surtout occidentales du lac” [Democratic Republic of the Congo, northern and western shores of Lake Tanganyika].

Paramelaniacallopleuros Bourguignat, 1885

Original reference.Bourguignat (1885a: 69–70).

Type locality. “Plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Hauttecoeuriacambieri Bourguignat, 1885

Original reference.Bourguignat (1885a: 56–57).

Original spelling.Hauttecœuria cambieri Bourguignat, 1885.

Type locality. “Plage de Mpala” [Democratic Republic of the Congo, beach of Mpala].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 56) but Bourguignat alone was responsible for the description. Therefore, Bourguignat is the author of the name (Code Art. 50.1).

Cleopatracameroni Bourguignat, 1879

Original reference.Bourguignat (1879: 21–22).

Original spelling.CleopatraCameroni Bourguignat, 1879.

Type locality. “Fleuve Kyngani, près de Bagamoyo” [Tanzania, Bagamoyo, Kyngani River].

Spekiacameroni Bourguignat, 1888

Original reference.Bourguignat (1888: 15, pl. 5, figs 13–15).

Original spelling.SpekiaCameroni Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Hauttecœuriacameroni Bourguignat, 1888

Original reference.Bourguignat (1888: 21–22, pl. 8, figs 15–17).

Original spelling.HauttecœuriaCameroni Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paramelaniacameroniana Bourguignat, 1885

Original reference.Bourguignat (1885a: 80).

Type locality. “Sur la plage de Mlilo et sur celle de Kapampa, à l’ouest du lac” [Democratic Republic of the Congo, beach of Mlilo (= Moliro) and beach of Kapampa, Lake Tanganyika].

Edgarianassacamerounensis Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 166).

Original spelling.Edgarianassaf.camerounensis Leloup, 1953.

Remarks. Mentioned in the synonymy of Edgardianassa forme typica; unavailable (Code Art. 11.6)

Cleopatracara Pilsbry & Bequaert, 1927

Original reference.Pilsbry and Bequaert (1927: 294, fig. 53).

Type locality. “Stanleyville” [Democratic Republic of the Congo, Kisangani].

† Pseudocleopatracarinata Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 56–58, 63, 66, fig. 44).

Type locality. “Nkondo locality NK 120, Lake Albert, Uganda”.

Type stratum. “Nkondo Formation, Molluscan Association G3b, Terminal Miocene-Early Pliocene”.

Syrnolopsiscarinifera Smith, 1889

Original reference.Smith (1889: 174).

Type locality. “Lake Tanganyika”.

Cleopatracarinulata Dautzenberg, 1894

Original reference.Dautzenberg (1894: 105–106).

Type locality. “Ambohimarina, province Hova d’Antankara près Diego-Suarez” [Madagascar, Diana, Ambohimarina].

Randabeliacatoxia Bourguignat, 1888

Original reference.Bourguignat (1888: 31, pl. 13, figs 1, 2).

Type locality. “Côtes occidentales et méridionales du lac” [Democratic Republic of the Congo/Zambia, western and southern shores of Lake Tanganyika].

Hauttecoeuriacharmetanti Bourguignat, 1888

Original reference.Bourguignat (1888: 19, pl. 7, figs 15–17).

Original spelling.Hauttecœuria Charmetanti Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusconicacherraensis Nevill, 1884

Original reference.Nevill (1884: 288).

Original spelling.Paludomusconicasubvar.cherraensis Nevill, 1884.

Type locality. “Teria Ghat” [India, Megalhaya, Khasi Hills, Teria Ghat].

Remarks.Nevill (1884) distinguished between ‘subspecies’, ‘var.’, and ‘subvar.’ Not every subvariety was referred to a subspecies or variety, however, but for example on p. 288 subvarieties were directly associated with a species without intermittent subspecies or variety. It seems that Nevill (1884) expressed by using the different terms to what extent a named taxon differed from the nominal species, with subspecies differing considerably, varieties slightly, and subvarieties only very slightly. Therefore, subspecific and varietal names are regarded here as subspecific names in accordance with the Code (Art. 45.6.4). The names denoted by ‘subvar.’ are, however, regarded as infrasubspecific. Made available by Kobelt in Pfeffer and Kobelt (1887: 422) (Code Art. 45.6.4.1).

Paludomuschilinoides Reeve, 1847

Original reference.Reeve (1847: pl. 2, fig. 7a, pl. 3, fig. 7b, c).

Original spelling.PaludomusChilinoides Reeve, 1847.

Type locality. “Bed of the Mahawelle Ganga, near Kandy, Ceylon” [Sri Lanka, Kandy, Mahawelle River].

Paludomusconicachittagongensis Nevill, 1884

Original reference.Nevill (1884: 288).

Original spelling.Paludomusconicasubvar.chittagongensis Nevill, 1884.

Type locality. “Chittagong” [Bangladesh, Chittagong].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884. Made available by Kobelt in Pfeffer and Kobelt (1887: 422) (Code Art. 45.6.4.1).

Paludomuscincta Liu, Zhang & Duan, 1994

Original reference.Liu et al. (1994: 30, 35).

Original spelling.Paludomuscinctus Liu, Zhang & Duan, 1994.

Type locality. “Sinan (27.9°N, 108.2°E), Guizhou Province. On stones or rocks at the bottom of Wujiang River”.

Paludomuscingulata Martens, 1878

Original reference.Martens (1878: pl. 2, figs 14–16).

Type locality. “Finboni” [Kenya, Finboni, near the coast, between Mombassa and Taita].

Remarks. The names Paludomusexarata and Paludomuscingulata were established simultaneously by Martens (1878). On pl. 2 Martens (1878) used the name Paludomuscingulata, but in the text and explanation of the plates the name Paludomusexarata was used. Bourguignat (1885b: 29) used the name in the combination Cleopatraexarata as the valid name of the taxon and remarked in a footnote “Ce nom a été modifié en celui de Cingulata sur l’explication de la planche”. Acting as first reviser, Bourguignat (1885) thus established the precedence of Paludomusexarata Martens, 1878 over Paludomuscingulata Martens, 1878.

† Edgariacingulata Cox, 1939

Original reference.Cox (1939: 247–248, pl. 15, fig. 8).

Type locality. ”The Rungwa Sleeping Sickness Concentration Area, lies about eighty miles to the north-west of the present shore of the lake” [Tanzania, Rungwa].

Type stratum. “Quaternary deposits of Lake Rukwa”.

Paludomusclavata Reeve, 1852

Original reference.Reeve (1852: 129).

Original spelling.Paludomusclavatus Reeve, 1852.

Type locality. “Mountain streams of Ceylon” [Sri Lanka].

Paludinacleopatra Martens, 1860 [Unavailable]

Original reference. Martens (1860: 13).

Original spelling.Paludinabulimoides Olivier, 1804 (cleopatra Troschel)

Remarks. Published as a synonym of Paludinabulimoides (Olivier, 1804) and attributed to Troschel who introduced Cleopatra as a genus-group name in 1857. Treated by Frauenfeld (1864: 588) as an available name, but not in a way that satisfies it to be adopted (Code, Glossary), i.e. not satisfying the Code (Art. 12.1). Therefore, the name was not made available under the Code (Art. 11.6.1) by Frauenfeld (1864).

Spekiacoheni Irvine & Donohue, 1999 [Unavailable]

Original reference.Irvine and Donohue (1999: 11).

Remarks. Introduced without description or definition (Code Art. 13.1.1). Mentioned also by Alin et al. (1999) as Spekia n. sp. “coheni” and by Allison et al. (2000) and Ngereza (2010) as Spekiacoheni.

Paludinacolbeaui Craven, 1880

Original reference.Craven (1880: 216, pl. 22, fig. 5).

Type locality. “Nossi-Bé Island, in a sluggish stream running through coffee-plantations” [Madagascar, Antsiranana, Nosy Be Island].

† Heynderycxiacolumellaris Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 60–61, 67, 77, figs 49a–d, 54c).

Type locality. “Nkondo locality NK 77, Lake Albert, Uganda”.

Type stratum. “Nkondo Formation, Molluscan Association G3b, Terminal Miocene to Early Pliocene”.

Lavigeriacombsa Bourguignat, 1888

Original reference.Bourguignat (1888: 33–34, pl. 14, fig. 7).

Type locality. “Sur les rives nord et surtout occidentales du lac” [Democratic Republic of the Congo, northern and western shores of Lake Tanganyika].

Paludomussulcatacompacta Nevill, 1884

Original reference.Nevill (1884: 299).

Original spelling.Paludomus [Philopotamis] sulcatavar.compacta Nevill, 1884.

Type locality. “Ceylon” [Sri Lanka].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Cleopatracongener Preston, 1913

Original reference.Preston (1913: 59, pl. 4, fig. 6).

Type locality. “Lake Baringo, British East Africa” [Kenya, Lake Baringo].

Melaniaconica Griffith & Pidgeon, 1833 [Invalid]

Original reference.Griffith and Pidgeon (1833–1834: pl. 14, fig. 5).

Type locality. “Ceylon” [Sri Lanka] (Griffith and Pidgeon 1833–1834: 598).

Remarks. Primary junior homonym of Melaniaconica Say, 1821. Although authorship is usually attributed to Gray (see Petit and Coan 2008), the requirements of the Code (Art. 50.1) are not fulfilled in our opinion. The authorship should therefore be attributed to Griffith and Pidgeon.

Paludomusconstricta Reeve, 1852

Original reference.Reeve (1852: 129).

Original spelling.Paludomusconstrictus Reeve, 1852.

Type locality. “Mountain streams of Ceylon” [Sri Lanka].

Paludomussulcatacontracta Nevill, 1884

Original reference.Nevill (1884: 299).

Original spelling.Paludomus [Philopotamis] sulcatavar.contracta Nevill, 1884.

Type locality. “Peradenia [and] Ambegammoa, Ceylon” [Sri Lanka].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Cleopatraconvexa Verdcourt, 1957

Original reference.Verdcourt (1957: 7–8, fig. 1).

Type locality. “Kenya, Turkana Province, Clay pan 7 miles S. E. Lodwar (subfossil)”.

Melaniacoronata von dem Busch, 1842

Original reference.Philippi (1842–1845: 2, pl. 1, figs 5, 6).

Type locality. “Bengalia” [India/Bangladesh].

Remarks. The entire text with descriptions of von dem Busch’s new Melania names was attributed to von dem Busch on p. 4 of Philippi’s (1842–1845) work.

Lavigeriacoronata Bourguignat, 1888

Original reference.Bourguignat (1888: 31, pl. 13, figs 13, 14).

Type locality. “Sur les rives nord et surtout occidentales du lac” [Democratic Republic of the Congo, northern and western shores of Lake Tanganyika].

Bridouxiacostata Bourguignat, 1885

Original reference.Bourguignat (1885a: 31–32).

Type locality. “Environs de Kapampa” [Democratic Republic of the Congo, surroundings of Kapampa, western shore of Lake Tanganyika].

Cleopatramorellicostata Preston, 1905

Original reference.Preston (1905: 300–301, fig. 4).

Original spelling.CleopatraMorellivar.costata Preston, 1905.

Type locality. “Victoria Falls, Zambesi River” [Zimbabwe, Zambesi River, Victoria Falls].

Melaniacrassa von dem Busch, 1842

Original reference.Philippi (1842–1845: 2–3, pl. 1, figs 10, 11).

Type locality. “Bengalia” [India/Bangladesh].

Remarks. The entire text with descriptions of von dem Busch’s new Melania names was attributed to von dem Busch on p. 4 of Philippi’s (1842–1845) work.

† Taeniodomuscrassa Krause, 1897

Original reference.Krause (1897: 214, 219, pl. 12, figs 2, 2a, 3).

Type locality. “Thonmergel vom Sungei Pinoh” [Indonesia, Borneo, Sungai Pinoh].

Type stratum. “α Eocaen (der Sandstein Etage) Verbeek‘s” [Eocene, Verbeek’s sandstone stage].

Paludomuscrassicallosa Rao, 1929

Original reference.Rao (1929: 288–290).

Type locality. “Sankha, a large hill-stream between Kamaing and Mogaung” [Myanmar].

Tiphobiacrassigranulata Smith, 1881

Original reference.Smith (1881b: 560).

Original spelling.Tiphobia (Paramelania) crassigranulata Smith, 1881.

Type locality. “Lake Tanganyika”.

Remarks.Paramelaniacranigranulata as used by Moore (1903) is an incorrect subsequent spelling.

Paramelaniacrassilabris Bourguignat, 1885

Original reference.Bourguignat (1885a: 84–85).

Type locality. “Plage Mlilo” [Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Cleopatracridlandi Mandahl-Barth, 1954

Original reference.Mandahl-Barth (1954: 61–62, fig. 27a, b).

Original spelling.Cleopatracridlandicridlandi Mandahl-Barth, 1954.

Type locality. “Lake Victoria, where it was dredged off Dagusi Island from a depth of 20 to 40 feet” [Uganda].

Paludomuscumingiana Dohrn, 1857

Original reference.Dohrn (1857: 124).

Original spelling.Paludomus Cumingianus Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Remarks. The name Paludomus Cumingii as used by Brot (1862: 21) and Brot (1868: 2) is an incorrect subsequent spelling.

Limnotrochuscyclostoma Bourguignat, 1885

Original reference.Bourguignat (1885a: 60–61).

Type locality. “Plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Paludinacyclostomoides Küster, 1852

Original reference.Küster (1852: 32, pl. 7, figs 6–10).

Type locality. “In Aegypten” [Egypt].

Cleopatracridlandicylindrica Mandahl-Barth, 1954

Original reference.Mandahl-Barth (1954: 62, fig. 27c–d).

Type locality. “From a depth of 60 feet in Buvuma Channel” [Uganda, Buvuma District, Buvuma Channel].

† Viviparuscylindricus Adam, 1957

Original reference.Adam (1957: 35–37, pl. 4, figs 1–5).

Type locality. “Rive du ravin de Nyakasia, 65 à 80 m sur lac” [Uganda, river of Nyakasia Ravine, 65 to 80 m above Lake Albert].

Type stratum. “Pleistocène inférieur, Série de Kaiso” [Lower Pleistocene, Kaiso Series].

Type material. Holotype, Musée royal de l’Afrique centrale, Tervuren MC 2780 (Adam 1957, pl. 4, fig. 1).

Remarks. The species was classified as Cleopatracylindrica (Adam, 1957) by Van Damme and Pickford (2003) in their revision of Albertine Rift Valley freshwater Cerithioidea. By that taxonomic act, the taxon becomes a secondary junior homonym of the nominal taxon Cleopatracridlandicylindrica Mandahl-Barth, 1954 and as no junior synonyms exist, the new replacement name Cleopatraadami nom. nov. is proposed here (see also Remarks under that name above).

Tiphobiadamoni Smith, 1881

Original reference.Smith (1881b: 559–560).

Original spelling.Tiphobia (Paramelania) damoni Smith, 1881.

Type locality. “Lake Tanganyika”.

Pseudocleopatradartevellei Mandahl-Barth, 1973

Original reference.Mandahl-Barth (1973: 280–281, fig. 3d–f, i).

Type locality. “River Congo at Matadi” [Democratic Republic of the Congo, Matadi, Congo River].

Giraudiahoreidautzenbergi Germain, 1905

Original reference.Germain (1905a: 259).

Original spelling.GiraudiaHorei var. Dautzenbergi Germain, 1905.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Marteliadautzenbergi Dupuis, 1924

Original reference.Dupuis (1924: 20–21, 26, fig. 1).

Original spelling.MarteliaDautzenbergi Dupuis, 1924.

Type locality. “Tanganika”…”de la côte occidentale du Lac” [Democratic Republic of the Congo, Lake Tanganyika, western shore].

Cleopatradautzenbergi Pilsbry & Bequaert, 1927

Original reference.Pilsbry and Bequaert (1927: 294–295, fig. 54).

Type locality. “Lovoi River, near Kikondja” [Democratic Republic of the Congo, near Kikondja, Lovoi River].

Paludomusdecussata Reeve, 1852

Original reference.Reeve (1852: 127–128).

Original spelling.Paludomusdecussatus Reeve, 1852.

Type locality. “Ceylon” [Sri Lanka].

Melaniadenticulata Benson, 1836

Original reference.Benson (1836b: 748).

Type locality. “From the north-east frontier of Bengal (Benson 1836a: 350)” [India/Bangladesh].

Remarks. The name was conditionally proposed.

Paludomusdhuma Rao, 1925

Original reference.Rao (1925: 97).

Type locality. “A stream near Thazi Railway Station” [Myanmar].

Lavigeriadiademata Bourguignat, 1888

Original reference.Bourguignat (1888: 31, 34, pl. 13, figs 15–17).

Type locality. “Sur les rives nord et surtout occidentales du lac” [Democratic Republic of the Congo, northern and western shores of Lake Tanganyika].

Paludomusconicadihiriensis Nevill, 1884

Original reference.Nevill (1884: 288).

Original spelling.Paludomusconicasubvar.dihiriensis Nevill, 1884.

Type locality. “Stream on the Dihiri Hill [Brahmaputra watershed]” [India, Assam].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884. Made available by Kobelt in Pfeffer and Kobelt (1887: 422) (Code Art. 45.6.4.1).

Paludomusdilatata Reeve, 1852

Original reference.Reeve (1852: 128–129).

Original spelling.Paludomusdilatatus Reeve, 1852.

Type locality. “Mountain streams of Ceylon” [Sri Lanka].

Paludomusdistinguenda Dohrn, 1857

Original reference.Dohrn (1857: 124).

Original spelling.Paludomusdistinguendus Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Paludomusdromedarius Dohrn, 1857

Original reference.Dohrn (1857: 124).

Type locality. Ceylon [Sri Lanka].

† Cleopatradubia Adam, 1959

Original reference.Adam and Lepersonne (1959: 34–38, pl. 5, figs 1–3, pl. 6, figs 1, 2).

Type locality. “Nyamavi no 68, camp Nyamavi” through selection of lectotype [Democratic Republic of the Congo, Nyamavi, near Lake Albert].

Remarks. The statement by Van Damme and Pickford (2003: 22) that the name Cleopatradubia Adam, 1959 is a nomen nudum is incorrect because W. Adam in Adam and Lepersonne (1959: 34–38) correctly introduced it, although not explicitly selecting a holotype and a type locality. Furthermore, the type selection of Van Damme and Pickford (2003: 49–50) does not constitute a valid lectotype designation in accordance with the Code Art. 74.7.1 because the term lectotype or an exact translation of that term was used not by these authors but only the term type and holotype. To avoid further confusion and nomenclatural instability, we here select the specimen proposed as type/holotype by Van Damme and Pickford (2003), i.e. the specimen figured by Adam (1959: pl. 5, fig. 1a), leg. J. Lepersonne, Lep. 464 III (M.C. 2382) from Nyamavi no 68, camp Nyamavi, as lectotype of Cleopatradubia Adam, 1959.

Hauttecoeuriaduveyieri Germain, 1905 [Invalid]

Original reference.Germain (1905a: 258).

Original spelling.Hauttecœuria Duveyieri Germain, 1905.

Remarks. Unjustified emendation of Hauttecœuriaduveyrieriana Bourguignat, 1885. Germain (1905a) changed the suffix of several species-group names ending in -iana or -ana introduced by Bourguignat (1885, 1888). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Spekiaduveyieri Germain, 1906 [Invalid]

Original reference.Germain (1906: 578).

Original spelling.SpekiaDuveyieri Germain, 1906.

Remarks. Unjustified emendation of Spekiaduveyrieriana Bourguignat, 1885. Germain (1906) changed the suffix of several species-group names ending in -iana or -ana introduced by Bourguignat (1885). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Spekiaduveyrieriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 37–38).

Type locality. “Mpala, dans le Marungu, sur la côte sud-ouest du lac” [Democratic Republic of the Congo, Mpala, Marungu region, at the south-western coast of Lake Tanganika].

Hauttecœuriaduveyrieriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 53).

Type locality. “Plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Paramelaniaduveyrieriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 79).

Type locality. “Plage de Mlilo” [Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 79) but Bourguignat alone was responsible for the description and is therefore the author of the name alone (Code 50.1).

Anceyagiraudiecarinata Leloup, 1953 [Unavailable]

Original spelling.Anceyagiraudivar.ecarinata Leloup, 1953.

Original reference.Leloup (1953: 108, 111–112, fig. 55B).

Remarks. Mentioned in the synonymy of Anceya (Anceya) giraudi as a manuscript name of Dupuis; unavailable (Code Art. 11.6).

Paramelaniaegregia Bourguignat, 1885

Original reference.Bourguignat (1885a: 81).

Type locality. “À Kapampa” [Democratic Republic of the Congo, Kapampa, western shore of Lake Tanganyika].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 81) but Bourguignat alone was responsible for the description and therefore is the author of the name (Code Art. 50.1).

Spekiazonataelata Germain, 1906

Original reference.Germain (1906: 579).

Original spelling.Spekiazonatavar.elata Germain, 1906.

Type locality. “À Mpala, sur la côte ouest, ainsi qu’à Pambété, au sud du lac” as given in Bourguignat (1885a) [Democratic Republic of the Congo, Mpala, western shore of Lake Tanganyika and Zambia, Pambété, southern shore of Lake Tanganyika].

Cleopatrapirothielata Dautzenberg & Germain, 1914

Original reference.Dautzenberg and Germain (1914: 57–58).

Original spelling.CleopatraPirothif.elata Dautzenberg & Germain, 1914.

Type locality. “Stn. 92, Bulongo (Bukama), 9° lat. S” [Democratic Republic of the Congo, Bulongo (Bukama), near Lualaba River].

Spekiazonataelongata Bourguignat, 1888

Original reference.Bourguignat (1888: 14, pl. 4, fig. 23).

Original spelling.Spekiazonatavar.elongata Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paramelaniaelongata Bourguignat, 1888

Original reference.Bourguignat (1888: 37–38, pl. 16, figs 9, 10).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusemini Smith, 1888

Original reference.Smith (1888: 54).

Type locality. “Albert Nyanza” [Democratic Republic of the Congo/Uganda, Lake Albert].

Paludomuserinaceus Reeve, 1852

Original reference.Reeve (1852: 128).

Type locality. “Mountain streams of Ceylon” [Sri Lanka].

Remarks.Tanaliacrinascens is an incorrect subsequent spelling by Layard (1855) and Tanaliacrinacea is an incorrect subsequent spelling by Adams and Adams (1853–1854).

Paludomuserronea Nevill, 1884

Original reference.Nevill (1884: 300).

Original spelling.Paludomus [Philopotamis] erronea Nevill, 1884.

Type locality. “Ceylon [and] Hackalle, Ceylon [at 5,500 ft]” [Sri Lanka].

Paludomuseveretti Smith, 1894

Original reference.Smith (1894: 51).

Type locality. “Batang Lupar district, W. Sarawak, also Gomanton on the N.E. Coast at Sandakan Bay” [Malaysia/Indonesia, Borneo].

Paludomusexarata Martens, 1878

Original reference.Martens (1878: 297–298, pl. 2, figs 14–16).

Type locality. “Finboni” [Kenya, Finboni, near the coast, between Mombassa and Taita].

Hauttecoeuriaeximia Bourguignat, 1885

Original reference.Bourguignat (1885a: 55–56).

Original spelling.Hauttecœuria eximia Bourguignat, 1885.

Type locality. “Plage de Mpala, à l’ouest du lac” [Democratic Republic of the Congo, Mpala, western shore of Lake Tanganyika].

Tanganikiafagoti Germain, 1908

Original reference.Germain (1908: 36).

Original spelling.TanganikiaFagoti Germain, 1908.

Remarks. Unjustified emendation of Tanganikiafagotiana Bourguignat, 1885.

Tanganikiafagotiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 43).

Original spelling.Tanganikia (Tanganikia) fagotiana Bourguignat, 1885.

Type locality. “Dans la partie septentrionale du lac” and “également dans sa partie méridionale […] sur la plage de Pambété” [Democratic Republic of the Congo or Burundi, northern part of Lake Tanganyika; Zambia, Pambété, southern shore of Lake Tanganyika].

Paludinafasciata Kobelt, 1909 [Unavailable]

Original reference.Kobelt (1909: 386).

Type locality. Not given.

Remarks. Published in the synonymy of Cleopatrabulimoides (Olivier, 1804); unavailable (Code Art. 11.6).

Melaniaferruginea Lea & Lea, 1851

Original reference.Lea and Lea (1851: 182).

Type locality. “Zanzibar, East Africa” [Tanzania, Zanzibar].

Paramelaniaflexicosta Martens, 1895

Original reference.Martens (1895: 186–187).

Original spelling.Paramelania (Edgaria) flexicosta Martens, 1895.

Type locality. “Tanganyika” [Lake Tanganyika].

Helixfluviatilis Dillwyn, 1817

Original reference.Dillwyn (1817: 959).

Type locality. “Inhabits fresh water in Coromandel” [India, Coromandel].

Syrnolopsisfoai Mabille, 1901

Original reference.Mabille (1901: 56).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Assimineafoai Mabille, 1901

Original reference.Mabille (1901: 56–57).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Remarks.Assimineafoas is an incorrect subsequent spelling by Pilsbry and Bequaert (1927).

Reymondiafoai Mabille, 1901

Original reference.Mabille (1901: 57).

Original spelling.ReymondiaFoai Mabille, 1901.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paramelaniaformosa Bourguignat, 1888

Original reference.Bourguignat (1888: 35–36, pl. 15, figs 9–11).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusfulgurata Dohrn, 1857

Original reference.Dohrn (1857: 123).

Original spelling.Paludomusfulguratus Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Paludomusfuniculata Reeve, 1847

Original reference.Reeve (1847: pl. 3, fig. 13).

Original spelling.Paludomusfuniculatus Reeve, 1847.

Type locality. “In a mountain stream at Ratnapoora, Ceylon” [Sri Lanka].

Paludomusfutaii Gredler, 1889

Original reference.Gredler (1889: 159–160).

Original spelling.PaludomusFutaii Gredler, 1889.

Type locality. “Im Peho (Kuang-tung)” [China, Guangdong, Peho River].

Cyclostomagaillardotii Bourguignat, 1855

Original reference.Bourguignat (1855: 333–334, pl. 8, figs 5–7).

Original spelling.CyclostomaGaillardotii Bourguignat, 1855.

Type locality. “Les environs de Sayda, en Syrie” [Syria, Sayda].

Paludomusgardneri Reeve, 1847

Original reference.Reeve (1847: pl. 2, fig. 9a, b).

Original spelling.PaludomusGardneri Reeve, 1847.

Type locality. “In a stream at the foot of Adam’s Peak, Ceylon” [Sri Lanka, Adam’s Peak].

Anceyagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 15–16).

Type locality. “Evirons de Mlilo, sur la côte ouest du lac” [Democratic Republic of the Congo, surroundings of Mlilo (= Moliro), western shore of Lake Tanganyika].

Syrnolopsisgiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 20–21).

Type locality. “Plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Bridouxiagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 30).

Type locality. “Plage de Kapampa, au sud-ouest du lac” [Democratic Republic of the Congo, beach of Kapampa, western shore of Lake Tanganyika].

Baizeagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 34–35).

Type locality. “Plage de Kapampa, au sud-ouest du lac, dans la region du Marungu“ [Democratic Republic of the Congo, Marungu region, Kapampa, south-western shore of Lake Tanganyika].

Spekiagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 36).

Type locality. “À Mpala, sur la côte ouest, ainsi qu’à Pambété, au sud du lac” [Democratic Republic of the Congo, Mpala, western shore of Lake Tanganyika and Zambia, Pambété, southern shore of Lake Tanganyika].

Tanganikiagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 44).

Original spelling.Tanganikia (Tanganikia) giraudi Bourguignat, 1885.

Type locality. “Plage de Kapampa, à l’ouest du lac” [Democratic Republic of the Congo, Kapampa, western shore of Lake Tanganyika].

Hauttecoeuriagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 49–50).

Original spelling.Hauttecœuria giraudi Bourguignat, 1885.

Type locality. “Sur la plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Limnotrochusgiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 59–60).

Type locality. “Plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Reymondiagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 65–66).

Type locality. “Plage de Mlilo” [Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Paramelaniagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 82).

Type locality. “Sur la plage de Mlilo” [Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Stanleyagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 88).

Type locality. “Plage de Mlilo” [Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Rumellagiraudi Bourguignat, 1885

Original reference.Bourguignat (1885a: 90–91).

Type locality. “Sur la plage de Mpala” [Democratic Republic of the Congo, Lake Tanganyika, beach of Mpala].

† Pseudocleopatraglaubrechti Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10–11, 57–59, fig. 47).

Type locality. “Kaiso locality A, Lake Albert, Uganda”.

Type stratum. “Kyeoro and Kaiso Village Formations, Molluscan Associations G5a and GX, Late Pliocene”.

Tanganikiaglobosa Bourguignat, 1885

Original reference.Bourguignat (1885a: 46).

Original spelling.Tanganikia (Cambieria) globosa Bourguignat, 1885.

Type locality. “Plage de Kapampa” [Democratic Republic of the Congo, beach of Kapampa, western shore of Lake Tanganyika].

Rumellaglobosa Bourguignat, 1888

Original reference.Bourguignat (1888: 39–40, pl. 17, figs 20–22).

Type locality. “De la côte occidentale” [Democratic Republic of the Congo, western shore of Lake Tanganyika].

Hirthiaglobosa Ancey, 1898

Original reference.Ancey (1898: 144–145, pl. 9, fig. H).

Type locality. “Habitat in lacu Tanganika, ad partem meridionalem et orientalem «Ufipa» dictam” [Tanzania, Ufipa region, Lake Tanganyika, south-eastern shores].

Melaniaglobulosa Griffith & Pidgeon, 1833

Original reference.Griffith and Pidgeon (1833–1834: pl. 14, fig. 6).

Type locality. Not given.

Remarks. See Remarks under Melaniaconica Griffith & Pidgeon, 1833

Paludomusgracilis Brot, 1880 [Unavailable]

Original reference.Brot (1880: 41).

Remarks. Attributed to Parreyss; published in the synonymy of Paludomustanjoriensis Blanford, 1863; unavailable (Code Art. 11.6).

† Taeniodomusgracilis Krause, 1897

Original reference.Krause (1897: 213–214, 219, pl. 12, figs 1, 1a, b).

Type locality. “Thonmergel vom Sungei Pinoh” [Indonesia, Borneo, Sungai Pinoh].

Type stratum. “α Eocaen (der Sandstein Etage) Verbeek‘s” [Eocene, Verbeek’s sandstone stage].

Syrnolopsisgracilis Pilsbry & Bequaert, 1927

Original reference.Pilsbry and Bequaert (1927: 234–235, figs 31a–c).

Type locality. “Lake Tanganyika”.

Paludomusgrandidieri Crosse & Fischer, 1872

Original reference.Crosse and Fischer (1872: 209–210).

Original spelling.PaludomusGrandidieri Crosse & Fischer, 1872.

Type locality. “Habitat in rivulis regionis orientalis insulæ Madagascar dictæ” [Madagascar, rivers in the eastern part of the island].

Syrnolopsisgrandidieri Germain, 1905 [Invalid]

Original reference.Germain (1905a: 259).

Original spelling.SyrnolopsisGrandidieri Germain, 1905.

Remarks. Unjustified emendation of Syrnolopsisgrandidieriana Bourguignat, 1885. Germain (1905a) changed the suffix of several species-group names ending in –iana or –ana introduced by Bourguignat (1885, 1888). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Spekiagrandidieri Germain, 1906 [Invalid]

Original reference.Germain (1906: 579).

Original spelling.SpekiaGrandidieri Germain, 1906.

Remarks. Unjustified emendation of Spekiagrandidieriana Bourguignat, 1885. Germain (1906) changed the suffix of several species-group names ending in –iana or –ana introduced by Bourguignat (1885). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Syrnolopsisgrandidieriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 18–19).

Type locality. “Plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Spekiagrandidieriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 40–41).

Type locality. “Sur les plages du Marungu, sur la côte sud-ouest du lac“ [Democratic Republic of the Congo, beaches of Marungu region, south-western shores of Lake Tanganyika].

Giraudiagrandidieriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 63–64).

Type locality. “Dans le sable d’une plage au nord de Mlilo” [Democratic Republic of the Congo, on a beach north of Mlilo (= Moliro), Lake Tanganyika].

Paramelaniagrandidieriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 70–71).

Type locality. “Aux environs de Pambété” [Zambia, surroundings of Pambété, Lake Tanganyika].

Tiphobianassagrandis Smith, 1881

Original reference.Smith (1881b: 561).

Original spelling.Tiphobia (Paramelania) nassavar.grandis Smith, 1881.

Type locality. “Lake Tanganyika”.

Cleopatraguillemei Bourguignat, 1885

Original reference.Bourguignat (1885c: 6–8).

Original spelling.CleopatraGuillemei Bourguignat, 1885.

Type locality. “Nyanza Oukéwéré, près de la Mission française”[Tanzania, Lake Victoria].

Cleopatraguillemeti Bourguignat, 1888

Original reference.Bourguignat (1888: 13–14, pl. 4, fig. 4).

Original spelling.CleopatraGuillemeti Bourguignat, 1888.

Type locality. “L’embouchure du Malagarazi” [Tanzania, Malagarasi Delta].

Nassopsisguillemei Martel & Dautzenberg, 1899

Original reference.Martel and Dautzenberg (1899: 172–173, pl. 8, figs 12, 13).

Original spelling.NassopsisGuillemei Martel & Dautzenberg, 1899.

Type locality. “Lac Tanganyika” [Lake Tanganyika].

Paramelaniaimperialisguillemei Martel & Dautzenberg, 1899

Original reference.Martel and Dautzenberg (1899: 178–180, pl. 8, fig. 23).

Original spelling.Paramelaniaimperialis var. Guillemei Martel & Dautzenberg, 1899.

Type locality. “Lac Tanganyika” [Lake Tanganyika].

Spekiahamyi Germain, 1906 [Invalid]

Original reference.Germain (1906: 580).

Original spelling.SpekiaHamyi Germain, 1906.

Remarks. Unjustified emendation of Spekiahamyana Bourguignat, 1885. Germain (1906) changed the suffix of several species-group names ending in –iana or –ana introduced by Bourguignat (1885). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Syrnolopsishamyana Bourguignat, 1885

Original reference.Bourguignat (1885a: 17–18).

Type locality. “Sur la plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Spekiahamyana Bourguignat, 1885

Original reference.Bourguignat (1885a: 38–39).

Type locality. “Sur la plage de Mpala” [Democratic Republic of the Congo, Lake Tanganyika, beach of Mpala].

Hauttecoeuriahamyana Bourguignat, 1885

Original reference.Bourguignat (1885a: 48–49).

Original spelling.Hauttecœuria hamyana Bourguignet, 1885.

Type locality. “Sur la plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Paramelaniahamyana Bourguignat, 1885

Original reference.Bourguignat (1885a: 71–72).

Type locality. “De la plage de Pambété, au sud du lac” [Zambia, beach of Pambété, southern shore of Lake Tanganyika].

Paludomushanleyi Dohrn, 1858

Original reference.Dohrn (1858: 535).

Type locality. “Inhabitants of mountain-streams in Ceylon” [Sri Lanka].

Cleopatrahargeri Smith, 1908

Original reference.Smith (1908: 13).

Original spelling.CleopatraHargeri Smith, 1908.

Type locality. “Kalingwisi River, Lake Mweru” [Zambia].

Cleopatrahemmingi Verdcourt, 1956

Original reference.Hemming and Verdcourt (1956: 60–62, figs 3, 4e).

Type locality. “Italian Somaliland, 18 miles W. of Beles Cogani, frequent in the large shallow pools” [Somalia].

Paludomushilberi Gredler, 1886

Original reference.Gredler (1886: 19–20).

Original spelling.Paludomus (?) Hilberi Gredler, 1886.

Type locality. “Aus Hensan, der Provinz Hunan” [China, Hunan].

Cleopatrahirta Dautzenberg & Germain, 1914

Original reference.Dautzenberg and Germain (1914: 59, pl. 4, figs 11–14).

Type locality. “Stn. 49, Nyangwe, Lualaba” [Democratic Republic of the Congo, Nyangwe, Lualaba River].

Tiphobiahorei Smith, 1880

Original reference.Smith (1880a: 348–349, pl. 31, fig. 7).

Type locality. “Lake Tanganyika”.

Melaniahorei Smith, 1880

Original reference.Smith (1880b: 427).

Original spelling.Melania (―?) Horei Smith, 1880.

Type locality. “Lake Tanganyika”.

Bathanaliahowesi Moore, 1898

Original reference.Moore (1898a: 452, fig. 2; 1898c: 192, 203, pl. 12, figs 29, 30).

Original spelling.BathanaliaHowesi Moore, 1898.

Type locality. “Near Mleroes, Tanganyika, at a depth of 950 ft” [Zambia, near Mleroes, southern shore of Lake Tanganyika].

Remarks. The name BathanaliaHowesi was published simultaneously (March 1898) in two separate works by J. E. S. Moore (1998a, c). In Moore (1898a) the name was spelled BathanaliaHowesii, and in Moore (1898b) it was spelled BathanaliaHowesi. Moore (1898d) used BathanaliaHowesii and Bathanalia Howsei, while in another subsequent work by the author, Moore (1898e), only the name BathanaliaHowesi was used. Therefore, Moore (1898e) is deemed here as First Reviser under the Code Art. 24.2.4 of the simultaneously published spellings of Moore (1898a, c) and Bathanaliahowesi Moore, 1998 (including obligate corrections under the Code Art. 32.5.2.5) is regarded as the correct original spelling. Bathanalia Howsei is an incorrect subsequent spelling by Moore (1898d).

Bourguignatiaimperialis Giraud, 1885

Original reference.Giraud (1885: 194).

Type locality. “Région du Marungu, sur la plage de Mpala (côte sud-ouest du Tanganika)“ [Democratic Republic of the Congo, Lake Tanganyika, Marungu region, beach of Mpala, south-western shore of Lake Tanganyika].

Paludomusinflata Brot, 1880

Original reference.Brot (1880: 44–45, pl. 5, fig. 10).

Original spelling.Paludomusinflatus Brot, 1880.

Type locality. “Travancore” and “Amerghat” [India, Kerala; Amerghat].

Paramelaniainfralirata Bourguignat, 1888

Original reference.Bourguignat (1888: 35–36, pl. 15, figs 4, 5).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Bythocerasiridescens Moore, 1898

Original reference.Moore (1898a: 452, fig. 1).

Type locality. “Near Sumbu, Tanganyika, at a depth of 680 ft” [Zambia, near Sumbu, southern part of Lake Tanganyika].

Paludomusisseli Brot, 1880

Original reference.Brot (1880: 31–32, pl. 7, figs 7, 7a, 8).

Original spelling.PaludomusIsseli Brot, 1880.

Type locality. “Sarawak auf Borneo” [Malaysia, Borneo, Sarawak].

Paludomusconicajaintiaca Nevill, 1884

Original reference.Nevill (1884: 289).

Original spelling.Paludomusconicavar.jaintiaca Nevill, 1884.

Type locality. “S. Jaintia Hills” [India, Meghalaya].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Cleopatrajohnstoni Smith, 1893

Original reference.Smith (1893: 637, pl. 59, fig. 9).

Type locality. “Lake Mweru” [Zambia/Democratic Republic of the Congo].

Tiphobiajouberti Bourguignat, 1886

Original reference.Bourguignat (1886: 146–148, 150, pl. 6, figs 11–13).

Type locality. “Sur les plages du Masanzé au N.-O. de la presqu’île Oubouari“ [Democratic Republic of the Congo, north-western part of the Ubwari Peninsula, beaches of Masanzé, Lake Tanganyika].

Cleopatrajouberti Bourguignat, 1888

Original reference.Bourguignat (1888: 13–14, pl. 4, fig. 3).

Original spelling.CleopatraJouberti Bourguignat, 1888.

Type locality. “L’embouchure du Malagarazi” [Tanzania, Malagarasi Delta].

Cambieriajouberti Bourguignat, 1888

Original reference.Bourguignat (1888: 17–18, pl. 6, figs 15, 16).

Original spelling.CambieriaJouberti Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Hauttecoeuriajouberti Bourguignat, 1888

Original reference.Bourguignat (1888: 19, pl. 7, figs 24, 25).

Original spelling.Hauttecœuria Jouberti Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Reymondiajouberti Bourguignat, 1888

Original reference.Bourguignat (1888: 27–28, pl. 11, figs 5, 6).

Original spelling.ReymondiaJouberti Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Bourguignatiajouberti Bourguignat, 1888

Original reference.Bourguignat (1888: 29, pl. 12, figs 5–7).

Original spelling.BourguignatiaJouberti Bourguignat, 1888.

Type locality. “Sur les plages de la côte occidentale” [Democratic Republic of the Congo, western shore of Lake Tanganyika].

Lavigeriajouberti Bourguignat, 1888

Original reference.Bourguignat (1888: 33–34, pl. 14, fig. 4).

Original spelling.LavigeriaJouberti Bourguignat, 1888.

Type locality. “Sur les rives nord et surtout occidentales du lac” [Democratic Republic of the Conga, Lake Tanganyika, northern and western shores].

Rumellajouberti Bourguignat, 1888

Original reference.Bourguignat (1888: 40, pl. 17, figs 29–31).

Original spelling.RumellaJouberti Bourguignat, 1888.

Type locality. “De la côte occidentale” [Democratic Republic of the Congo, western shore of Lake Tanganyika].

Paludomustanjoriensiskadapaensis Nevill, 1884

Original reference.Nevill (1884: 294).

Original spelling.Paludomustanjoriensisvar.kadapaensis Nevill, 1884.

Type locality. “Kadapa District, Madras” [India, Tamil Nadu, Chennai].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

† Cleopatrakaisoensis Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 11, 52–53, fig. 38).

Type locality. “Kaiso Village locality Kaiso A, Lake Albert, Uganda”.

Type stratum. “Kaiso Village Formation, Molluscan Associations GX, Late Pliocene”.

Paludomuskamaingia Rao, 1929

Original reference.Rao (1929: 290–292).

Type locality. “From a small rocky stream near Kamaing” [Myanmar, Kamaing].

Cleopatransendweensiskatangana Pilsbry & Bequaert, 1927

Original reference.Pilsbry and Bequaert (1927: 296, fig. 55b).

Type locality. “Luvua River, between Kiambi and Ankoro” [Democratic Republic of the Congo].

Limnotrochuskirkii Smith, 1880

Original reference.Smith (1880b: 426).

Original spelling.LimnotrochusKirkii Smith, 1880.

Type locality. “Lake Tanganyika”.

Remarks.Limnotrochuskerkii as used by Moore (1903) is an incorrect spelling. The deliberate change by Smith (1881a: 286–287) to Limnotrochuskirki has also to be interpreted as an incorrect subsequent spelling (Code Art. 33.4).

† Potadomoideskisegiensis Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 17, 47–48, 62, fig. 34).

Type locality. “Kisegi locality KI 5, Lake Albert, Uganda”.

Type stratum. “Kakara Formation, Molluscan Association G1, Late Miocene”.

Paludomusconicakopiliensis Nevill, 1884

Original reference.Nevill (1884: 289).

Original spelling.Paludomusconicavar.kopiliensis Nevill, 1884.

Type locality. “Kopili River, North Assam [and] North Assam” [India, Meghalaya/Assam].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Bithyniabulimoideskotschyana Frauenfeld, 1864 [Unavailable]

Original reference.Frauenfeld (1864: 583, 619).

Original spelling.Bythiniabulimoides var. Kotschyana Frauenfeld, 1864.

Remarks. Nomen nudum, name introduced without definition, description, or indication (Code Art. 12.1).

Paludomuskweichowensis Cheng, 1937

Original reference. Cheng (1937: 445–447, figs 1.3, 2.3).

Original spelling.Paludomus (Hemimitra) kweichowensis Cheng, 1937.

Type locality. “At Shih-men-kan, Kweichow Province, China” [China, Guizhou].

Cleopatrakynganica Bourguignat, 1879

Original reference.Bourguignat (1879: 21).

Original spelling.CleopatraKynganica Bourguignat, 1879.

Type locality. “Dans le fleuve Kyngani, près de Bagamoyo” [Tanzania, Bagamoyo, Kyngani River].

Remarks.Cleopatra Kinganica is an incorrect subsequent spelling by Bourguignat (1885b, c).

Paludomuslabiosa Benson, 1856

Original reference.Benson (1856: 495–496).

Type locality. “Hab. in rivulis vallis Tenasserim” [Myanmar, rivers in the Tenasserim Valley].

Paludomuslacunoides Aldrich, 1889

Original reference.Aldrich (1889: 23–24, pl. 3, figs 1, 1a–c).

Type locality. “In the Kusan and Penggiron districts in Southeastern Borneo and from the Kinan and Kiwa Rivers” [Indonesia, Borneo].

Paramelanialacunosa Bourguignat, 1888

Original reference.Bourguignat (1888: 37–38, pl. 16, figs 5, 6).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Syrnolopsislacustris Smith, 1880

Original reference.Smith (1880b: 426–427).

Type locality. “Lake Tanganyika”.

Paludomuslaevis Layard, 1855

Original reference.Layard (1855: 89).

Original spelling.Paludomuslævis Layard, 1855.

Type locality. “Ceylon, in slow-running streams on the northern side of the mountain zone extending into the flat country beyond Anarajahpoora [and] in a paddy field in the south of the island, near the village of Heneratgodde” [Sri Lanka].

Cleopatralangi Pilsbry & Bequaert, 1927

Original reference.Pilsbry and Bequaert (1927: 293–294, figs 51a–d, 52).

Type locality. “Stanleyville” [Democratic Republic of the Congo, Kisangani].

Cleopatralaurenti Bourguignat, 1879

Original reference.Bourguignat (1879: 24).

Original spelling.CleopatraLaurenti Bourguignat, 1879.

Type locality. “Sur les bords du Nil, au-dessus du Caire“ [Egypt, Nile River, upriver of Cairo].

Hauttecoeurialavigeriana Bourguignat, 1888

Original reference.Bourguignat (1888: 19, pl. 7, figs 26–27).

Original spelling.Hauttecœuria Lavigeriana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Giraudialavigeriana Bourguignat, 1888

Original reference.Bourguignat (1888: 27–28, pl. 11, figs 22–24).

Original spelling.GiraudiaLavigeriana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Rumellalavigeriana Bourguignat, 1888

Original reference.Bourguignat (1888: 40, pl. 17, figs 32–34).

Original spelling.RumellaLavigeriana Bourguignat, 1888.

Type locality. “De la côte occidentale” [Democratic Republic of the Congo, western shore of Lake Tanganyika].

Paludomuslayardi Reeve, 1852

Original reference.Reeve (1852: 128).

Original spelling.PaludomusLayardi Reeve, 1852.

Type locality. “Mountain streams of Ceylon” [Sri Lanka].

Lavigerialechaptoisi Ancey, 1898

Original reference.Ancey (1898: 145–146, pl. 9, fig. I).

Original spelling.Lavigeria (?) lechaptoisi Ancey, 1898.

Type locality. “Habitat in ripa meridionali lacus Tanganika «Ufipa» dicta” [Tanzania, Ufipa region, Lake Tanganyika].

Paramelanialedoulxiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 80–81).

Type locality. “Sur la plage Kapampa, à l’ouest du lac” [Democratic Republic of the Congo, Kapampa, western shore of Lake Tanganyika].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 80), but Bourguignat alone was responsible for the description and therefore is the author of the name (Code Art 50.1).

† Viviparuslepersonnei Gautier, 1970

Original reference. Gautier & Charig (1970: 41, 95–96, figs 6a–d).

Original spelling.Viviparus?lepersonnei Gautier, 1970.

Type locality. “Kaiso“ [Uganda, Kaiso].

Type stratum. Kaiso, Site 3.

Cleopatralesnei Germain, 1935

Original reference.Germain (1935: 68–69).

Original spelling.CleopatraLesnei Germain, 1935.

Type locality. “Bas Sangadzé (Zambèse)” [Mozambique, Sofala].

Remarks. Type locality given erroneously; according to Brown and Mandahl-Barth (1989) a Paludomus Swainson, 1840 species.

Paramelanialessepsiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 78–79).

Type locality. “Sur la plage de Mlilo” [Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 78), but Bourguignat alone was responsible for the description and therefore is the author of the name (Code Art 50.1).

Cleopatraletourneuxi Bourguignat, 1879

Original reference.Bourguignat (1879: 19–21).

Original spelling.CleopatraLetourneuxi Bourguignat, 1879.

Type locality. “Fleuve Kyngani, près de Bagamoyo” [Tanzania, Bagamoyo, Kyngani River].

Ponsonbyaleucoraphe Ancey, 1890

Original reference.Ancey (1890: 347).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Hauttecoeurialevesquei Germain, 1905 [Invalid]

Original reference.Germain (1905a: 258).

Original spelling.Hauttecœuria Levesquei Germain, 1905.

Remarks. Unjustified emendation of Hauttecœuria levesquiana Bourguignat, 1888. Germain (1905a) changed the suffix of several species-group names ending in –iana or –ana introduced by Bourguignat (1885, 1888). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Hauttecoeurialevesquiana Bourguignat, 1888

Original reference.Bourguignat (1888: 21–22, pl. 8, figs 9–11).

Original spelling.Hauttecœuria Levesquiana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Cleopatralhotellerii Bourguignat, 1879

Original reference.Bourguignat (1879: 25).

Original spelling.CleopatraLhotellerii Bourguignat, 1879.

Type locality. “Mandara, près d’Alexandrie” [Egypt, Mandara near Alexandria].

† Pseudocleopatralikeae Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 18, 54–55, 59, 63, figs 41a, 42).

Type locality. “Sinda-Mohari Gautier Site 13, Semliki Plain, Congo” [Democratic Republic of the Congo].

Type stratum. “Mohari and Kisegi Formations, Molluscan Association G0 & G1, Late and Middle Miocene”.

Syrnolopsislacustrislilacina Dartevelle & Schwetz, 1948

Original reference.Dartevelle and Schwetz (1948: 30, 37, 39, 52, 59, 74, 85, pl. 1, fig. 14).

Original spelling.Syrnolopsislacustrisvar.lilacina Dartevelle & Schwetz, 1948.

Type locality. “Lac Tanganilsa (?) (Stn. 2065)” [Lake Tanganyika].

Paramelanialimnaea Bourguignat, 1888

Original reference.Bourguignat (1888: 39–40, pl. 17, figs 7, 8).

Original spelling.ParamelaniaLimnæa Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludinalineata Menke, 1828 [Unavailable]

Original reference.Menke (1828: 23).

Remarks. Nomen nudum, introduced without description, definition, or indication (Code Art. 12.1). Only the name Helixtanschaurica Gmelin, 1791 was mentioned. Presented in the same way in Menke (1830: 41).

Edgarialittoralis Bourguignat, 1888

Original reference.Bourguignat (1888: 33–34, pl. 14, figs 14–16).

Type locality. “Çà et là sur tout le pourtour du lac” [Lake Tanganyika].

Hirthialittorina Ancey, 1898

Original reference.Ancey (1898: 142–144, pl. 9, fig. G).

Type locality. “Habitat in lacu Tanganika, ad partem meridionalem et orientalem «Ufipa» dictam” [Tanzania, Ufipa region, south-eastern shore of Lake Tanganyika].

Paramelanialivingstoniana Bourguignat, 1885

Original reference.Bourguignat (1885a: 85–86).

Type locality. “Plages de Pambété et de Kapampa ” [Zambia, beach of Pambété, and Democratic Republic of the Congo, Kapampa, Lake Tanganyika].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 85), but Bourguignat alone was responsible for the description and therefore is the author of the name (Code Art. 50.1).

Hauttecoeurialocardi Germain, 1905 [Invalid]

Original reference.Germain (1905a: 258).

Original spelling.Hauttecœuria Locardi Germain, 1905.

Remarks. Unjustified emendation Hauttecœurialocardiana Bourguignat, 1888. Germain (1905a) changed the suffix of several species-group names ending in –iana or –ana introduced by Bourguignat (1885, 1888). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Paramelanialocardi Germain, 1905 [Invalid]

Original reference.Germain (1905a: 259).

Original spelling.ParamelaniaLocardi Germain, 1905.

Remarks. Unjustified emendation of Paramelanialocardiana Bourguignat, 1885. Germain (1905a) changed the suffix of several species-group names ending in –iana or –ana introduced by Bourguignat (1885, 1888). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Paramelanialocardiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 82–83).

Type locality. “Sur la plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Hauttecœurialocardiana Bourguignat, 1888

Original reference.Bourguignat (1888: 21–22, pl. 8, figs 12–14).

Original spelling.Hauttecœuria Locardiana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Tiphobialongirostris Bourguignat, 1886

Original reference.Bourguignat (1886: 144–146, 150, pl. 6, figs 8–10).

Type locality. “Sur la plage de Kibanga, au sud de la presqu’île Oubouari” [Democratic Republic of the Congo, Kibanga, southern part of the Obwari Peninsula, Lake Tanganyika].

Paludomusloricata Reeve, 1847

Original reference.Reeve (1847: pl. 1, figs 1a–c).

Original spelling.Paludomusloricatus Reeve, 1847.

Type locality. “In rapids flowing from Adam’s Peak, Ceylon” [Sri Lanka, near Adam’s Peak].

Cleopatrabulimoideslutea Pallary, 1909 [Unavailable]

Original reference.Pallary (1909: 64).

Original spelling.Cleopatra bulimoïdes var. ex colore lutea Pallary, 1909.

Remarks. Nomen nudum, introduced without description, definition, or indication (Code Art. 12.1).

Paludomuslutea Adams, 1874

Original reference.Adams (1874: 585–586, pl. 59, figs 5, 5a).

Original spelling.Paludomusluteus Adams, 1874.

Type locality. “Borneo”.

Paludinalutosa Souleyet, 1852

Original reference.Souleyet (1852: 550, pl. 31, figs 28–30).

Type locality. “Cette espèce provient du Gange” [India/Bangladesh, Ganges River].

Paramelaniamabilliana Bourguignat, 1888

Original reference.Bourguignat (1888: 37–38, pl. 16, figs 17–18).

Original spelling.ParamelaniaMabilliana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Hauttecoeuriamacrostoma Bourguignat, 1888

Original reference.Bourguignat (1888: 19, pl. 7, figs 10–11).

Original spelling.Hauttecœuria macrostoma Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusmaculata Lea, 1856

Original reference.Lea (1856: 110).

Type locality. “Ahmednugger, India” [India, Maharashtra].

Paludinamadagascariensis Crosse & Fischer, 1872

Original reference.Crosse and Fischer (1872: 210).

Original spelling.PaludinaMadagascariensis Crosse & Fischer, 1872.

Type locality. “Habitat in rivulis regionis orientalis insulæ Madagascar dictæ” [Madagascar, rivers in the eastern part of the island].

Paludomusmadagascariensis Brot, 1880

Original reference.Brot (1880: 48, pl. 8, fig. 7).

Type locality. “Madagascar” [Madagascar].

Original spelling.PaludomusMadagascariensis Brot, 1880.

Paludomushanleyimajor Nevill, 1884

Original reference.Nevill (1884: 305).

Original spelling.Paludomus [Tanalia] hanleyivar.major Nevill, 1884.

Type locality. “Ceylon” [Sri Lanka].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Paramelaniastanleyanamajor Bourguignat, 1885

Original reference.Bourguignat (1885a: 75).

Original spelling.Paramelaniastanleyanavar.major Bourguignat, 1885.

Type locality. “Plages de Pambété et de Mlilo“ [Zambia, beach of Pambété and Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Syrnolopsisminutamajor Germain, 1905

Original reference.Germain (1905a: 260).

Original spelling.Syrnolopsisminutavar.major Germain, 1905.

Type locality. “Lac Tanganika“ [Lake Tanganyika].

Cleopatrabulimoidesmajor Dautzenberg & Germain, 1914

Original reference.Dautzenberg and Germain (1914: 56–57).

Original spelling.Cleopatrabulimoidesf.major Dautzenberg & Germain, 1914.

Type locality. “Stn. 72, Kibombo, 4° lat. S“ [Democratic Republic of the Congo, Kibombo].

Edgarianassamajor Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 166).

Original spelling.Edgarianassaf.major Leloup, 1953.

Remarks. Mentioned in the synonymy of Edgardianassa forme typica; unavailable (Code Art. 11.6).

Edgarialocardianamajor Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 167).

Original spelling.Edgarialocardianaf.major Leloup, 1953.

Remarks. Mentioned in the synonymy of Edgardianassa forme giraudi; unavailable (Code Art. 11.6).

Paludomustanjoriensismalabarica Nevill, 1884

Original reference.Nevill (1884: 294–295).

Original spelling.Paludomustanjoriensisvar.malabarica Nevill, 1884.

Type locality. “Travancore [and] Pulney Hills” [India, Kerala, Pulney Hills].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Cleopatramangoroensis Ancey, 1890

Original reference.Ancey (1890: 344–345).

Type locality. “Fleuve Mangoro, de Tananarive à la côte orientale de Madagascar, à 700 m” [Madagascar, from Antananarivo to the eastern coast, at 700 m, Mangoro River].

Cleopatramareotica Bourguignat, 1879

Original reference.Bourguignat (1879: 25).

Original spelling.CleopatraMareotica Bourguignat, 1979.

Type locality. “Dans le petit canal de Mustapha, près de Ramlé (Égypte)” [Egypt, Ramle near Alexandria, Mustapha Canal].

Hauttecoeuriamaunoiri Germain, 1905 [Invalid]

Original reference.Germain (1905a: 258).

Original spelling.Hauttecœuria Maunoiri Germain, 1905.

Remarks. Unjustified emendation of Hauttecoeuriamaunoiriana Bourguignat, 1885. Germain (1905a) changed the suffix of several species-group names ending in –iana or –ana introduced by Bourguignat (1885, 1888). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Tanganikiamaunoiriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 44–45).

Original spelling.Tanganikia (Cambieria) maunoiriana Bourguignat, 1885.

Type locality. “Plages de Mpala et de Pambété” [Democratic Republic of the Congo, Lake Tanganyika, beach of Mpala and Zambia, beach of Pambété, southern shore of Lake Tanganyika].

Hauttecoeuriamaunoiriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 55).

Original spelling.Hauttecœuria maunoiriana Bourguignat, 1885.

Type locality. “Sur la plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Paludomusmaurus Reeve, 1852

Original reference.Reeve (1852: 127).

Type locality. “Branch of the Ganges” [India/Bangladesh, Ganges River].

Cleopatrabulimoidesmedia Pallary, 1924

Original reference.Pallary (1924: 33).

Original spelling.Cleopatrabulimoides variété media Pallary, 1924.

Type locality. “Égypte” [Egypt].

Paludomusmelanostoma Hanley & Theobald, 1876

Original reference.Hanley and Theobald (1876: 49, pl. 121, figs 8–9).

Type locality. “Ceylon” [Sri Lanka].

Cremnoconchusmessageri Bavay & Dautzenberg, 1900

Original reference.Bavay and Dautzenberg (1900: 116–117).

Original spelling.CremnoconchusMessageri Bavay & Dautzenberg, 1900.

Type locality. “Rivière Song-Ky-Kong, près de That-Khé” [Vietnam, near Thất Khê, Kỳ Cùng River].

† Ellinoriamichelae Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 61–62, fig. 50).

Type locality. “Nkondo locality NK 34, Lake Albert, Uganda”.

Type stratum. “Nkondo Formation, Molluscan Association G3b, Late Miocene to Early Pliocene”.

Paludomusmicrosculpta Nevill, 1884 [Unavailable]

Original reference.Nevill (1884: 297).

Type locality. “Ceylon” [Sri Lanka].

Remarks. Introduced without description, definition, or indication (Code Art. 12.1).

Paludomusrotundamicrostoma Nevill, 1884

Original reference.Nevill (1884: 295).

Original spelling.Paludomusrotundavar.microstoma Nevill, 1884.

Type locality. “Anamallay Rivers [and] Madura Hills” [India, Tamil Nadu].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Tanganikiasolutamilneedwardsi Germain, 1905 [Invalid]

Original reference.Germain (1905a: 258).

Original spelling.Tanganikia (Hauttecœuria) soluta var. Milne-Edwardsi Germain, 1905.

Remarks. Unjustified emendation of Hauttecœuria milneedwardsiana Bourguignat, 1885. Germain (1905a) changed the suffix of several species-group names ending in –iana or –ana introduced by Bourguignat (1885, 1888). The names thus formed are to be treated as emendations of the original names (Code Art. 33.2.1) and are available and have as unjustified emendations their own author and date (Code Art. 33.2.2).

Hauttecoeuriamilneedwardsiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 50–51).

Original spelling.Hauttecœuriamilne-edwardsiana Bourguignat, 1885.

Type locality. “Sur les plages du sud du lac” [Zambia, southern shores of Lake Tanganyika].

Paramelaniamilneedwardsiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 77–78).

Original spelling.Paramelaniamilne-edwardsiana Bourguignat, 1885.

Type locality. “Plage de Mlilo” [Democratic Republic of the Congo, Mlilo (= Moliro), Lake Tanganyika].

Rumellamilneedwardsiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 91).

Original spelling.Rumellamilne-edwardsiana Bourguignat, 1885.

Type locality. “Plage de Kapampa, au sud-ouest du lac” [Democratic Republic of the Congo, beach of Kapampa, western shore of Lake Tanganyika].

Paludomusminensis Cheng, 1937

Original reference. Cheng (1937: 445–448, figs 1.4, 2.4).

Type locality. “In Min River near Kienway, Szechuan Province, China”.

Giraudiaminima Smith, 1908

Original reference.Smith (1908: 12).

Type locality. “Lake Tanganyika”.

Paludomusregulataminor Nevill, 1884 [Unavailable]

Original reference.Nevill (1884: 291).

Original spelling.Paludomusregulatasubvar.minor Nevill, 1884.

Type locality. “Burma” [Myanmar].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884; not subsequently made available under the Code (Art. 45.6.4.1).

Paludomussulcataminor Nevill, 1884 [Unavailable]

Original reference.Nevill (1884: 299).

Original spelling.Paludomus [Philopotamis] sulcatasubvar.minor Nevill, 1884.

Type locality. “Ceylon” [Sri Lanka].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884. Used by Rao (1929) as the valid name of a taxon, however without definition, description, or indication. Therefore not adopted as the name of a taxon in the sense of the code and thus not made available (Code Art. 45.6.4.1).

Paludomusaculeataminor Nevill, 1884 [Unavailable]

Original reference.Nevill (1884: 302).

Original spelling.Paludomus [Tanalia] aculeatasubvar.minor Nevill, 1884.

Type locality. “Ceylon” [Sri Lanka].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884. Not subsequently made available under the Code (Art. 45.6.4.1).

Paramelaniaduveyrierianaminor Bourguignat, 1885

Original reference.Bourguignat (1885a: 79).

Original spelling.Paramelaniaduveyrierianavar.minor Bourguignat, 1885.

Type locality. “Dans les environs de Pambété” [Zambia, surroundings of Pambété, Lake Tanganyika].

Remarks. Acting as first revisers, we here select the name Paramelaniaduveyrierianavar.minor Bourguignat, 1885 to have the precedence over Paramelanialocardianavar.minor Bourguignat, 1885.

Paramelanialocardianaminor Bourguignat, 1885 [Invalid]

Original reference.Bourguignat (1885a: 83).

Original spelling.Paramelanialocardianavar.minor Bourguignat, 1885.

Type locality. “À Mlilo et aussi à Pambété” [Democratic Republic of the Congo, Mlilo (= Moliro) and Zambia, Pambété, Lake Tanganyika].

Remarks. Invalid, homonym of Paramelaniaduveyrierianavar.minor Bourguignat, 1885 [first reviser action, see under Paramelaniaduveyrierianaminor Bourguignat, 1885].

Reymondiaminor Smith, 1889

Original reference.Smith (1889: 174).

Type locality. “Lake Tanganyika”.

Hauttecoeuriagiraudiminor Bourguignat, 1890

Original reference.Bourguignat (1890: 100).

Original spelling.Hauttecœuria Giraudi var.minor Bourguignat, 1890.

Type locality. “A Kibanga, au sud de la presqu’île Oubouari” [Democratic Republic of the Congo, Kibanga, Obwari Peninsula, Lake Tanganyika].

Reymondiagiraudiminor Bourguignat, 1890 [Invalid]

Original reference.Bourguignat (1890: 155).

Original spelling.ReymondiaGiraudi forme minor Bourguignat, 1890.

Type locality. “Plage au nord de Mlilo; sur une autre plage voisine” [Democratic Republic of the Congo, beach north of Mlilo (= Moliro), Lake Tanganyika].

Remarks. Invalid, junior homonym of Reymondiaminor Smith, 1889.

Bythocerasminor Moore, 1903

Original reference.Moore (1903: 242, 244, fig. 24).

Type locality. “Lake Tanganyika”.

Lavigeriajoubertiminor Germain, 1905 [Unavailable]

Original reference.Germain (1905a: 259).

Original spelling.Lavigeriajoubertivar.minor Germain, 1905.

Type locality. Not given.

Remarks. Nomen nudum, introduced without description, definition, or indication (Code Art. 12.1).

Giraudiabridouxianaminor Ancey, 1906 [Unavailable]

Original reference.Ancey (1906b: 254).

Original spelling.GiraudiaBridouxianaf.minor Ancey, 1906.

Type locality. “Ufipa, M’bwé” [Tanzania, Ufipa region, Lake Tanganyika].

Remarks. Nomen nudum, introduced without description, definition, or indication (Code Art. 12.1).

Cleopatrajohnstoniminor Dautzenberg & Germain, 1914 [Unavailable]

Original reference.Dautzenberg and Germain (1914: 57).

Original spelling.CleopatraJohnstonif.minor Dautzenberg & Germain, 1914.

Type locality. “Stn. 38, riv. Luvua, entre Ankoro et Kiambi“ and “Stn. 79, riv. Luvua, entre Ankoro et Kiambi” [Democratic Republic of the Congo, between Ankoro and Kiambi, Luvua River].

Remarks. Nomen nudum, introduced without description, definition, or indication (Code Art. 12.1).

Paludomusferrugineaminor Dautzenberg & Germain, 1914 [Unavailable]

Original reference.Dautzenberg and Germain (1914: 61).

Original spelling.Paludomus (Zanguebaria) ferrugineaf.minor Dautzenberg & Germain, 1914.

Type locality. “Stn. 36, Lovoi, Kikondja, Katanga“ [Democratic Republic of the Congo, near Kikondja, Lovoi River].

Remarks. Nomen nudum, introduced without description, definition, or indication (Code Art. 12.1).

Cleopatrabulimoidesminor Pallary, 1924

Original reference.Pallary (1924: 33).

Original spelling.Cleopatrabulimoides variété minor Pallary, 1924.

Type locality. “Égypte” … “d’une localité inconnue” [Egypt].

Edgariamilneedwardsianaminor Dartevelle & Schwetz, 1948

Original reference.Dartevelle and Schwetz (1948: 31, 37, 43, 53, 60, 74, 76).

Type locality. “Ufipa” [Tanzania, Ufipa region, Lake Tanganyika], see Ancey (1906b: 263).

Original spelling.Edgariamilne-edwardsianavar.minor Dartevelle & Schwetz, 1948.

Remarks.Dartevelle and Schwetz (1948: 53) remarked “D’autre part, nous avons ulilisé la variété suivante, due à C. F. Ancey et restée manuscrite, mais signalée (2, p. 263): Edgariamilne-edwardsianavar.minor C. F. Ancey”. Ancey (1906b: 263) stated that the specimens are distinguished from the nominate form by their smaller size, thus providing a diagnosis, which is referred to by Dartevelle and Schwetz (1948).

Edgariaegregiaminor Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 166).

Original spelling.Edgariaegregiaf.minor Leloup, 1953.

Remarks. Mentioned in the synonymy of Edgardianassa forme typica; unavailable (Code Art. 11.6).

Edgarianassaminor Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 166).

Original spelling.Edgarianassaf.minor Leloup, 1953.

Remarks. Mentioned in the synonymy of Edgardianassa forme typica; unavailable (Code Art. 11.6).

Edgarialocardianaspinulosaminor Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 167).

Original spelling.Edgarialocardiana var. spinulosaf.minor Leloup, 1953.

Remarks. Mentioned in the synonymy of Edgardianassa forme giraudi; unavailable (Code Art. 11.6).

Syrnolopsisminuta Bourguignat, 1885

Original reference.Bourguignat (1885a: 21–22).

Type locality. “Plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Remarks.Syrnolopsisminima Bourguignat, 1885 as used by Bandel, 1998 is a lapsus for Syrnolopsisminuta Bourguignat, 1885.

Hauttecoeuriaminuta Bourguignat, 1885

Original reference.Bourguignat (1885a: 57–58).

Original spelling.Hauttecœuria minuta Bourguignat, 1885.

Type locality. “Sur la plage de Mpala” [Democratic Republic of the Congo, Lake Tanganyika, beach of Mpala].

Paludomusminutiuscula Gredler, 1885

Original reference.Gredler (1885: 232–233, pl. 6, fig. 8).

Original spelling.Paludomus?minutiusculus Gredler, 1885.

Type locality. “Aus Tschin-chi, einer Stadt 3. Ranges in der Provinz Kuei-tscheu” [China, Guizhou].

Philopotamismisconceptus Nevill, 1884 [Unavailable]

Original reference.Nevill (1884: 299).

Remarks. Mentioned as a manuscript name under Paludomusnigricans Reeve, 1847 by Nevill (1884: 299); unavailable (Code Art. 11.6).

Melaniamodicella Lea & Lea, 1851

Original reference.Lea and Lea (1851: 196–197).

Type locality. “Timor” [Timor].

Hauttecoeuriamoineti Bourguignat, 1888

Original reference.Bourguignat (1888: 19, pl. 7, figs 1–3).

Original spelling.Hauttecœuria Moineti Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Syrnolopsislacustrismolirensis Pilsbry & Bequaert, 1927

Original reference.Pilsbry and Bequaert (1927: 230–231, figs 26h, 27c–d).

Type locality. “Off Moliro, in 70 m and at Moliro; in Lake Tanganyika” [Democratic Republic of the Kongo, off and at Moliro, Lake Tanganyika].

Reymondiamonceti Bourguignat, 1888

Original reference.Bourguignat (1888: 27–28, pl. 11, figs 7–8).

Original spelling.ReymondiaMonceti Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Edgariamonceti Bourguignat, 1888

Original reference.Bourguignat (1888: 33–34, pl. 14, figs 12–13).

Original spelling.EdgariaMonceti Bourguignat, 1888.

Type locality. “Cà et là sur tout le pourtour du lac” [Lake Tanganyika].

Paludomusmonile Hanley & Theobald, 1876

Original reference.Hanley and Theobald (1876: 44, pl. 108, fig. 10).

Type locality. “Southern India”.

Paludomusmoreleti Issel, 1874

Original reference.Issel (1874: 456–457, pl. 7, fig. 21–22).

Original spelling.PaludomusMoreleti Issel, 1874.

Type locality. “Territorio di Sarawak” [Malaysia, Borneo, Sarawak].

Cleopatramorelli Preston, 1905

Original reference.Preston (1905: 300–301, fig. 3).

Original spelling.CleopatraMorelli Preston, 1905.

Type locality. “Just above Victoria Falls, Zambesi River” [Zimbabwe, Zambesi River, above Victoria Falls].

Paramelaniaimperialismpalaensis Martel & Dautzenberg, 1899

Original reference.Martel and Dautzenberg (1899: 180, pl. 8, fig. 24).

Original spelling.Paramelaniaimperialisvar.mpalaensis Martel & Dautzenberg, 1899.

Type locality. “Lac Tanganyika” [Democratic Republic of the Congo, Lake Tanganyika, Mpala].

Cleopatra.mterizensis Melvill & Standen, 1907

Original reference.Melvill and Standen (1907: 5, pl. 1, fig. 2).

Type locality. “Bed Mterize River, a tributary of the Loangwa River” [Zambia].

Syrnolopsisminutamulticarinata Ancey, 1906

Original reference.Ancey (1906b: 267).

Original spelling.Syrnolopsisminutaf.multicarinata Ancey, 1906.

Type locality. “Lac Tanganika, côte orientale” [Democratic Republic of the Congo, western shore of Lake Tanganyika].

Cleopatramultilirata Ancey, 1906

Original reference.Ancey (1906a: 45).

Type locality. “Vinaninony, Madagascar” [Madagascar, Vinaninony].

Bithyniamultisulcata Bourguignat, 1888

Original reference.Bourguignat (1888: 11, pl. 3, figs 7–8).

Original spelling.Bythiniamultisulcata Bourguignat, 1888.

Type locality. “Sur les rives de la presqu’île Oubouari” [Democratic Republic of the Congo, Lake Tanganyika, shores of Obwari Peninsula].

Syrnolopsisminutamulticsulcata Ancey, 1906 [Unavailable]

Original reference.Ancey (1906b: 255).

Original spelling.Syrnolopsisminutaf.multisulcata Ancey, 1906.

Type locality. “Lac Tanganika, côte orientale” [Democratic Republic of the Congo, western shore of Lake Tanganyika].

Remarks. Nomen nudum, introduced without description, definition, or indication (Code Art. 12.1).

† Pseudocleopatramusiimei Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 12–13, 55–58, 63, 66, fig. 43).

Type locality. “Nkondo locality NK 122, Lake Albert, Uganda”.

Type stratum. “Lusso Formation (Biozone L0) (L. Edward), Nkondo Formation (L. Albert), Molluscan Associations G3a, G3b & G3c. Late Miocene to Early Pliocene”.

Cleopatramweruensis Smith, 1893

Original reference.Smith (1893: 637–638, pl. 59, fig. 10).

Type locality. “Lake Mweru” [Democratic Republic of the Congo/Uganda].

Paludomusandersonianamyadoungensis Nevill, 1881

Original reference.Nevill (1881: 160).

Original spelling.PaludomusAndersonianavar.myadoungensis Nevill, 1881.

Type locality. “Myadoung, near the Yunnan” [Myanmar].

Paludomusandersoniananana Nevill, 1881

Original reference.Nevill (1881: 160).

Original spelling.PaludomusAndersonianasubvar.nana Nevill, 1881.

Type locality. “Also from Pegu” [Myanmar].

Remarks. Introduced as a subvariety. However, made available by Rao (1929: 292–293, text fig. 6) (Code Art. 45.6.4.1).

Paludomusconicanana Nevill, 1884 [Invalid]

Original reference.Nevill (1884: 289).

Original spelling.Paludomusconicavar.nana Nevill, 1884.

Type locality. “W. Khasi Hills” [India, Meghalaya].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884. Made available by Kobelt in Pfeffer and Kobelt (1887: 422) (Code Art. 45.6.4.1). Junior homonym of Paludomusandersoniananana Nevill, 1881.

Melanianassa Woodward, 1859

Original reference.Woodward (1859: 349, pl. 47, fig. 4).

Original spelling.Melania (Melanella) nassa Woodward, 1859.

Type locality. “Lake Tanganyika in Central Africa”.

Remarks.Nassopsisnana as used by Moore (1903) is an incorrect spelling.

Paramelanianassatella Bourguignat, 1888

Original reference.Bourguignat (1888: 37–38, pl. 16, figs 3, 4).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paramelanianassatiformis Bourguignat, 1888

Original reference.Bourguignat (1888: 39–40, pl. 17, figs 5, 6).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusnasuta Dohrn, 1857

Original reference.Dohrn (1857: 123–124).

Original spelling.Paludomusnasuta Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Lithoglyphusneritinoides Smith, 1880

Original reference.Smith (1880b: 426).

Type locality. “Lake Tanganyika”.

Remarks.StanleyaNeritoides is an incorrect subsequent spelling by Bourguignat (1885, 1888).

Paludomusneritoides Reeve, 1847

Original reference.Reeve (1847: pl. 1, fig. 3a. b).

Original spelling.PaludomusNeritoides reeve, 1847.

Type locality. “In the bed of a river at Ambegamoa, Ceylon” [Sri Lanka].

Bithyniabulimoidesnigra Frauenfeld, 1864

Original reference.Frauenfeld (1864: 583, 629).

Original spelling.Bythiniabulimoides var. Nigra Frauenfeld, 1864.

Type locality. “Aegypten” [Egypt].

Remarks. Attributed to Caillaud. Introduced by Frauenfeld (1864: 583) as a variety of Bithyniabulimoides (Olivier, 1804), and on p. 629 the name was mentioned with some descriptional terms; thus satisfying the Code (Art. 12.1).

Paludomusnigricans Reeve, 1847

Original reference.Reeve (1847: pl. 2, fig. 6).

Type locality. “Ceylon (in mountain streams at 6,000 feet elevation)” [Sri Lanka].

Paludomusnodulosa Dohrn, 1857

Original reference.Dohrn (1857: 125).

Original spelling.Paludomusnodulosus Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Cleopatrabulimoidesnsendweensis Dupuis & Putzeys, 1901

Original reference.Dupuis and Putzeys (1901: 55–56).

Original spelling.Cleopatrabulimoidesvar.nsendweensis Dupuis & Putzeys, 1901.

Type locality. “Nyangwe, Nsendwe, Lokandu” [Democratic Republic of the Congo, Nyangwe; Nsendwe; Lokandu].

Cleopatranyanzae Mandahl-Barth, 1954

Original reference.Mandahl-Barth (1954: 63, fig. 29a–b).

Type locality. “Kanyakwar wells” [Kenya].

Melaniaobesa Philippi, 1847

Original reference.Philippi (1846–1847: 170, pl. 4, fig. 3).

Original spelling.Melania?obesa Philippi, 1847.

Type locality. “Nova Hollandia?” [Australia; doubtful].

Anceyagiraudiobesa Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 108).

Original spelling.Anceyagiraudi variété obesa Leloup, 1953.

Remarks. Mentioned in the synonymy of Anceya (Anceya) giraudi; unavailable (Code Art. 11.6).

Edgarianassavar.obliqua Ancey, 1906

Original reference.Ancey (1906b: 264).

Type locality. “Sur les rivages de l’Ufipa” [Tanzania, Ufipa region, Lake Tanganyika].

Cleopatraobscura Mandahl-Barth, 1968

Original reference.Mandahl-Barth (1968: 34–35, pl. 1 fig. 8, pl. 7 figs 1–3).

Type locality. ”Kafubu region, Kimilolo River at Keyberg” [Democratic Republic of the Congo, Keyberg Station near Lubumbashi, Kimililo River].

Paramelaniaobtusa Bourguignat, 1888

Original reference.Bourguignat (1888: 35–36, pl. 15, figs 6–8).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusolivacea Reeve, 1847

Original reference.Reeve (1847: pl. 1, fig. 5).

Original spelling.Paludomusolivaceus Reeve, 1847.

Type locality. “Point Palmas, Island of Sumatra (in a muddy stream)” [Indonsesia, Sumatra, Point Palmas; doubtful].

Tanganikiaopalina Bourguignat, 1888

Original reference.Bourguignat (1888: 15, pl. 5, figs 18–19).

Type locality. “Sur les plages de la partie septentrionale du lac“ [Democratic Republic of the Congo/Burundi, northern shores of Lake Tanganyika].

Paludomusornata Benson, 1856

Original reference.Benson (1856: 496).

Type locality. “Hab. in regno Burmanico” [Myanmar].

Tanganikiaovoidea Bourguignat, 1885

Original reference.Bourguignat (1885a: 45–46).

Original spelling.Tanganikia (Cambieria) ovoidea Bourguignat, 1885.

Type locality. “Plage de Kapampa, au sud-ouest du lac” [Democratic Republic of the Congo, beach of Kapampa, western shore of Lake Tanganyika].

Paludomuspalawanica Brot, 1891

Original reference.Brot (1891: 17–18).

Type locality. “In a brook about ten miles from Puerto Princesa in the Island of Palawan, Philippine Archipelago” [Philippines].

Remarks.Brot (1891: 17–18) used both, the spelling P.palawanicus and P.palawanica in the original description. The grammatical gender of the genus-group name Paludomus is feminine. Therefore, Paludomuspalawanica Brot, 1891 is the correct original spelling.

Bithyniabulimoidespallida Frauenfeld, 1864

Original reference.Frauenfeld (1864: 583, 633).

Original spelling.Bythiniabulimoides var. Pallida Frauenfeld, 1864.

Type locality. “Aegypten” [Egypt].

Remarks. Attributed to Caillaud. Introduced by Frauenfeld (1864: 583) as a variety of Bithyniabulimoides (Olivier, 1804), and on p. 633 the name was mentioned with some descriptional terms; thus satisfying the Code (Art. 12.1).

Paludomuspaludinoides Reeve, 1852

Original reference.Reeve (1852: 127).

Original spelling.PaludomusPaludinoides Reeve, 1852.

Type locality. “Sikkim branch of the Ganges” [India/Bangladesh].

Paludomuspalustris Layard, 1855

Original reference.Layard (1855: 89–90).

Type locality. “The grassy margins of a tank at Anarajahpoora” [Sri Lanka].

Paramelaniapalustris Bourguignat, 1888

Original reference.Bourguignat (1888: 36, pl. 15, figs 31, 32).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusparvula Rao, 1929

Original reference.Rao (1929: 293–294).

Type locality. “From an unknown locality in Burma” [Myanmar].

Paludomusparva Layard, 1855

Original reference.Layard (1855: 90).

Original spelling.Paludomusparvus Layard, 1855.

Type locality. “Bombay” [India, Maharashtra, Mumbai].

Tiphobianassapaucicostata Smith, 1881

Original reference.Smith (1881b: 561).

Original spelling.Tiphobia (Paramelania) nassavar.paucicostata Smith, 1881.

Type locality. “Lake Tanganyika”.

Cleopatrapauli Bourguignat, 1885

Original reference.Bourguignat (1885b: 27–28, pl. 1, fig. 3).

Original spelling.CleopatraPauli Bourguignat, 1885.

Type locality. “Bords du fleuve Haouach” [Chad, Haouach River].

Paludomusconicapealiana Nevill, 1884

Original reference.Nevill (1884: 288).

Original spelling.Paludomusconicavar.pealiana Nevill, 1884.

Type locality. “Assam [and] Sibsagar” [India, Assam, Sivasagar].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Paludomusandersonianapeguensis Nevill, 1877

Original reference.Nevill (1877: 35).

Original spelling.PaludomusAndersoniana var. Peguensis Nevill, 1877.

Type locality. “Pegu” [Myanmar].

Potadomoidespelseneeri Leloup, 1953

Original reference.Leloup (1953: 92, 102–105, 115, 132, 136, pl. 3, fig. 6, figs. 43, 47, 57P, 72H, 76).

Type locality. “Dans le delta de la Malagarasi, le long des rives et dans les petites baies, coquilles; par tamisage de la vase et des débris végétaux recueillis par la petite drague dans une anse calme et encombrée de végétations, – 30–40 cm, spécimens vivants” [Tanzania, Malagarasi Delta].

Cleopatrapercarinata Bourguignat, 1885

Original reference.Bourguignat (1885b: 27–28, pl. 1, fig. 1).

Type locality. “Lac Haoussa”.

Lavigeriaperexemia Bourguignat, 1888

Original reference.Bourguignat (1888: 33–34, pl. 14, fig. 3).

Type locality. “Sur les rives nord et surtout occidentales du lac” [Democratic Republic of the Congo, northern and western shores of Lake Tanganyika].

Viviparaamiltonipersolida Issel, 1874

Original reference.Bourguignat (1888: 454–455).

Original spelling.ViviparaHamiltonivar.persolida Issel, 1874.

Type locality. “Sadong, in limpidi ruscelli” [Malaysia, Borneo, Sarawak, Sadong River].

Paludinapetrosa Gould, 1843

Original reference.Gould (1843: 144).

Type locality. “Tavoy” [Myanmar, Dawei].

Paludomusphasianinus Reeve, 1852

Original reference.Reeve (1852: 127).

Original spelling.Paludomusphasianina Reeve, 1852.

Type locality. “Seychelles”.

Paludomuspicta Reeve, 1847

Original reference.Reeve (1847: pl. 2, figs 10a–b).

Original spelling.Paludomuspictus Reeve, 1847.

Type locality. “In a mountain stream at Ratnapoora, Ceylon” [Sri Lanka].

Cleopatrapilula Mandahl-Barth, 1967

Original reference.Mandahl-Barth (1967: 130, pl. 2, fig. 12).

Type locality. “Bushimaie River at Lukuta, Kasai” [Democratic Republic of the Congo, Kasai, Lukuta, Mbuji-Mayi River].

Cleopatrapirothi Jickeli, 1881

Original reference.Jickeli (1881: 338).

Original spelling.CleopatraPirothi Jickeli, 1881.

Type locality. “Nordost-Afrika” [Northeast Africa].

Syrnolopsislacustrisvar.pluricarinata Dartevelle & Schwetz, 1948

Original reference.Dartevelle and Schwetz (1948: 30, 37, 39, 52, 59, 74, 85, pl. 1, fig. 11).

Original spelling.Syrnolopsislacustrisvar.pluricarinata Dartevelle & Schwetz, 1948.

Type locality. “Lac Tanganika, baie de Kasakalawe, profondeur : 10 a 15 m (Stn. 2068)” [Zambia, Bay of Kaskalawe, Lake Tangayika].

Rissoaponsonbyi Smith, 1889

Original reference.Smith (1889: 175).

Original spelling.Rissoa (Horea) Ponsonbyi Smith, 1889.

Type locality. “Lake Tanganyika”.

Cleopatrapoutrini Germain, 1909

Original reference.Germain (1909: 377–378).

Original spelling.CleopatraPoutrini Germain, 1909.

Type locality. ”Dans l’Egueï, à 1,000 kilomètres environ au nord de Fort-Lamy” [Chad, Kanem, Eguei].

Giraudiapraeclara Bourguignat, 1885

Original reference.Bourguignat (1885a: 62–63).

Original spelling.Giraudiapræclara Bourguignat, 1885.

Type locality. “Plage au nord de Mlilo” [Democratic Republic of the Congo, beach north of Mlilo (= Moliro), Lake Tanganyika].

Paramelaniapulchella Bourguignat, 1885

Original reference.Bourguignat (1885a: 86).

Type locality. “Plage de Kapampa” [Democratic Republic of the Congo, beach of Kapampa, western shore of Lake Tanganyika].

Cleopatrabulimoidespulchella Pallary, 1909

Original reference.Pallary (1909: 63).

Original spelling.Cleopatrabulimoïdesvar.pulchella Pallary, 1909.

Type locality. “Égypte” [Egypt].

Remarks. Attributed to Bourguignat in Pallary (1924: 33).

Paludomuspunctata Reeve, 1852

Original reference.Reeve (1852: 127).

Original spelling.Paludomuspunctatus Reeve, 1852.

Type locality. “Mauritius”.

Syrnolopsispupoidea Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 121).

Remarks. Mentioned in the synonymy of Syrnolopsisgracilis as a manuscript name of Dautzenberg & Dupuis; unavailable (Code Art. 11.6).

Hauttecoeuriapusilla Bourguignat, 1888

Original reference.Bourguignat (1888: 22, pl. 8, figs 32–34).

Original spelling.Hauttecœuria pusilla Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomuspustulosa Annandale, 1921

Original reference.Annandale (1921: 563–564, fig. 11A, B).

Type locality. “From one stream in the south part of the Manipur Valley” [India, Manipur].

Reymondiapyramidalis Bourguignat, 1888

Original reference.Bourguignat (1888: 27–28, pl. 11, figs 9–13).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomuspyriformis Dohrn, 1858

Original reference.Dohrn (1858: 536).

Type locality. “Inhabitants of mountain-streams in Ceylon” [Sri Lanka].

Paludomusqianensis Liu, Duan & Zhang, 1994

Original reference.Liu et al. (1994: 30, 34).

Type locality. “Huangguoshu, (25.9°N, 105.6°E), Guizhou Province […] on the stones of the ditch beside river” [China, Guizhou].

Paludomusquadrasi Moellendorff, 1894

Original reference. Quadras & Moellendorff (1894: 130).

Original spelling.Paludomus (Philopotamis) quadrasi Moellendorff, 1894.

Type locality. “Busuanga” [Philippines, Busuanga Island].

Remarks. Authorship attributed to Moellendorff (Code Art. 50.1).

Syrnolopsisquintana Mabille, 1901

Original reference.Mabille (1901: 56).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paramelaniarandabeli Bourguignat, 1888

Original reference.Bourguignat (1888: 37–38, pl. 16, figs 21, 22).

Original spelling.ParamelaniaRandabeli Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusrapaeformis Brot, 1880

Original reference.Brot (1880: 30–31, pl. 5, fig. 10).

Type locality. Not given.

Cleopatraraymondi Bourguignat, 1879

Original reference.Bourguignat (1879: 23–24).

Original spelling.CleopatraRaymondi Bourguignat, 1879.

Type locality. “Lac Ballat dans l’isthme de Suez” [Egypt, Isthmus of Suez].

Tanaliareevei Layard, 1855

Original reference.Layard (1855: 92).

Original spelling.TanaliaReevei Layard, 1855.

Type locality. “The Calloo ganga, Ratnapoora” [Sri Lanka, Ratnapura, Kalu River].

Philopotamisregalis Layard, 1855

Original reference.Layard (1855: 93).

Type locality. “Stream in the Cnia Corle, Western province, Ceylon” [Sri Lanka].

Paludomusregulata Benson, 1856

Original reference.Benson (1856: 496).

Type locality. “Hab. ad Thyet-Myo Burmanorum” [Myanmar, Thayet].

Paludomusreticulata Blanford, 1870

Original reference.Blanford (1870: 9–10, pl. 3, fig. 1).

Type locality. “Hab. in Cachar” [India, Assam, Cachar].

Melaniaretusa Griffith & Pidgeon, 1833

Original reference.Griffith and Pidgeon (1833–1834: pl. 14, fig. 9).

Type locality. Not given.

Remarks. See Remarks under Melaniaconica Griffith & Pidgeon, 1833

Paludomusretusa Swainson, 1840

Original reference.Swainson (1840: 340).

Original spelling.Paludomus (Hemimitra) retusa Swainson, 1840.

Type locality. Not given.

Bridouxiareymondi Bourguignat, 1885

Original reference.Bourguignat (1885a: 32–33).

Type locality. “Plage de Kapampa” [Democratic Republic of the Congo, beach of Kapampa, western shore of Lake Tanganyika].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 32), but Bourguignat alone was responsible for the description and is therefore the author of the name (Code Art. 50.1).

Spekiareymondi Bourguignat, 1885

Original reference.Bourguignat (1885a: 39–40).

Type locality. “Sur la plage de Mpala” [Democratic Republic of the Congo, Lake Tanganyika, beach of Mpala].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 39), but Bourguignat alone was responsible for the description and therefore is the author of the name (Code art. 50.1).

Hauttecoeuriareymondi Bourguignat, 1885

Original reference.Bourguignat (1885a: 54–55).

Original spelling.Hauttecœuria reymondi Bourguignat, 1885.

Type locality. Not given.

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 54), but Bourguignat alone was responsible for the description and therefore is the author of the name (Code Art. 50.1).

Paramelaniareymondi Bourguignat, 1885

Original reference.Bourguignat (1885a: 72–73).

Type locality. “Sur la plage de Kapampa dans le Marungu (côte sud-ouest du lac)“ [Democratic Republic of the Congo, Marungu region, beach of Kapampa, western shore of Lake Tanganyika].

Remarks. The name was attributed to V. Giraud by Bourguignat (1885a: 72), but Bourguignat alone was responsible for the description and therefore is the author of the name (Code Art. 50.1).

Cleopatrabulimoidesrichardi Germain, 1911

Original reference.Germain (1911: 200, pl. 2, figs 5, 6).

Original spelling.Cleopatrabulimoides var. Richardi Germain, 1911.

Type locality. “Intérieur du Tchad, à 40 kilomètres du bord Ouest” [Interior of Lake Chad, 40 km from its western shore] and “Zone sableuse, bord occidental du Tchad” [Lake Chad, western shore].

Paludomusrotunda Blanford, 1870

Original reference.Blanford (1870: 10, pl. 3, fig. 2).

Type locality. “Hab. in regione Travancorica” [India, Kerala].

Stanleyarotunda Smith, 1904

Original reference.Smith (1904: 93).

Type locality. Given as “[Lac Tanganika], côte orientale” [Lake Tanganyika, eastern shore] in Bourguignat (1888) and as “sur les plages entre Oudjiji et l’embouchure du Malagarazi” [Tanzania, Lake Tanganyika, beaches between Ujiji and Malagarasi Delta] in Bourguignat (1890).

Remarks. Name introduced without description, but with reference to descriptions in Bourguignat (1888, 1890).

† Pseudocleopatrarotunda Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 58, 63, 66, fig. 46).

Type locality. “Nkondo locality NK 120, Lake Albert, Uganda”.

Type stratum. “Nkondo Formation, Molluscan Association G3b, Terminal Miocene-Early Pliocene”.

Paludomusrotungensis Preston, 1915

Original reference.Preston (1915a: 539–540).

Type locality. “Tenmalai, Upper Rotung” [India, Arunachal Pradesh].

Paludomusrudis Reeve, 1852

Original reference.Reeve (1852: 126).

Type locality. Not given.

Lavigeriaruellaniana Bourguignat, 1888

Original reference.Bourguignat (1888: 33–34, pl. 14, figs 5, 6).

Original spelling.LavigeriaRuellaniana Bourguignat, 1888.

Type locality. “Sur les rives nord et surtout occidentales du lac” [Democratic Republic of the Congo, northern and western shores of Lake Tanganyika].

Remarks.Lavigeria Ruelliana is an incorrect subsequent spelling by Martel and Dautzenberg (1899).

Anceyarufocincta Smith, 1906

Original reference.Smith (1906: 183–184, pl. 10, fig. 12).

Type locality. “Kirando, towards south end of the east coast, 10 fath” [Tanzania, Kirando, Lake Tanganyika, eastern shore]

Lithoglyphusrufofilosus Smith, 1880

Original reference.Smith (1880b: 426).

Type locality. “Lake Tanganyika”.

Cleopatrapirothirufolirata Germain, 1919

Original reference.Germain (1919a: 119).

Original spelling.Cleopatrapirothi variété rufolirata Germain, 1919.

Type locality. “Rhodésie septentrionale: Lealui, sur le Haut-Zambèse” [Zambia, Lealui].

Cleopatrarugosa Connolly, 1925

Original reference.Connolly (1925: 424–425).

Type locality. “Italian Somaliland; Aggherrar” [Somalia, Aggherrar].

† Edgariarukwaensis Cox, 1939

Original reference.Cox (1939: 246–247, pl. 15, figs 6, 7).

Type locality. “Rungwa Sleeping Sickness Concentration Arealies about, lies about eighty miles to the north-west of the present shore of the lake” [Tanzania, Rungwa].

Type stratum. “Quaternary deposits of Lake Rukwa”.

Paludomusrusiostoma Gredler, 1885

Original reference.Gredler (1885: 231–232, pl. 6, figs 7, 7a).

Type locality. “Aus Tschin-chi, einer Stadt 3. Ranges in der Provinz Kuei-tscheu” [China, Guizhou].

Remarks. Probably a viviparid, see Zilch (1974).

Cleopatraschoutedeni Dautzenberg & Germain, 1914

Original reference.Dautzenberg and Germain (1914: 58–59, pl. 4, figs 15, 16).

Original spelling.CleopatraSchoutedeni Dautzenberg & Germain, 1914.

Type locality. “Stn. 50, Nyangwe, Lualaba” [Democratic Republic of the Congo, Nyangwe, Lualaba River], through the lectotype designation by Glaubrecht and Strong (2007: 387).

Syrnolopsisminutasemilaevis Ancey, 1906

Original reference.Ancey (1906b: 255, 267).

Original spelling.Syrnolopsisminutaf.semilaevis Ancey, 1906.

Type locality. “Lac Tanganika, côte orientale” [Tanzania/Burundi, eastern shore of Lake Tanganyika.

Paludinasenegalensis Morelet, 1860

Original reference.Morelet (1860a: 190).

Original spelling.PaludinaSenegalensis Morelet, 1860.

Type locality. “Habitat circa Podor in paludibus” [Senegal, Podor].

Paramelaniaservainiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 83–84).

Type locality. “Sur la plage Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

Hauttecoeuriaservainiana Bourguignat, 1888

Original reference.Bourguignat (1888: 21–22, pl. 8, figs 18, 19).

Original spelling.Hauttecœuria Servainiana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomussiamensis Blanford, 1903

Original reference.Blanford (1903: 283–284, pl. 8, fig. 3).

Original spelling.PaludomusSiamensis Blanford, 1903.

Type locality. “Siam, in valle superiore Menam fluminis” [Thailand, upper Chao Phraya Valley].

Paludomusconicasibsaugorensis Nevill, 1884

Original reference.Nevill (1884: 288).

Original spelling.Paludomusconicavar.sibsaugorensis Nevill, 1884.

Type locality. “Sibsagar” [India, Assam, Sivasagar].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Paludomusajanensissilhouettensis Nevill, 1884

Original reference.Nevill (1884: 296).

Original spelling.Paludomusajanensisvar.silhouettensis Nevill, 1884.

Type locality. “Silhouette I., Seychelles”.

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Tanaliasimilis Layard, 1855

Original reference.Layard (1855: 92).

Type locality. “A mountain torrent at Kandangamoa, near Ratnapoora” [Sri Lanka, Ratnapura, Kandangamoa].

Hauttecoeuriasingularis Bourguignat, 1885

Original reference.Bourguignat (1885a: 52–53).

Original spelling.Hauttecœuria singularis Bourguignat, 1885.

Type locality. “Près de Pambété” [Zambia, near Pambété, Lake Tanganyika].

Paramelaniasingularis Bourguignat, 1888

Original reference.Bourguignat (1888: 35–36, pl. 15, figs 16, 17).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Paludomusskinneri Dohrn, 1857

Original reference.Dohrn (1857: 124–125).

Original spelling.PaludomusSkinneri Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Paramelaniasmithi Bourguignat, 1888

Original reference.Bourguignat (1888: 37–38, pl. 16, figs 11, 12).

Original spelling.ParamelaniaSmithi Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Cleopatrasmithi Ancey, 1906

Original reference.Ancey (1906a: 45–46).

Original spelling.CleopatraSmithi Ancey, 1906.

Type locality. “River Chozi, which flows into the Chambézi, region of lake Bangwéolo, British Central Africa” [Zambia, region of Lake Bangweulu, Chozi River, tributary of Chambeshi River].

Stanleyasmithiana Bourguignat, 1885

Original reference.Bourguignat (1885a: 88–89).

Type locality. “Plage de Mlilo” [Democratic Republic of the Congo, beach of Mlilo (= Moliro), Lake Tanganyika].

Cleopatrasoleilleti Bourguignat, 1885

Original reference.Bourguignat (1885b: 28, pl. 1, fig. 1).

Original spelling.CleopatraSoleilleti Bourguignat, 1885.

Type locality. “Lac Haoussa” [Ethiopia, Lake Awasa].

Paludomussolida Dohrn, 1857

Original reference.Dohrn (1857: 124).

Original spelling.Paludomussolidus Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Hauttecoeuriasoluta Bourguignat, 1885

Original reference.Bourguignat (1885a: 51–52).

Original spelling.Hauttecœuria soluta Bourguignat, 1885.

Type locality. “Aux environs de Pambété” [Zambia, surroundings of Pambété, Lake Tanganyika].

Paludomussphaerica Dohrn, 1857

Original reference.Dohrn (1857: 124).

Original spelling.Paludomus sphæricus Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Paramelaniaspinulosa Bourguignat, 1885

Original reference.Bourguignat (1885a: 75–76).

Type locality. “Plage de Mlilo” [Democratic Republic of the Congo, beach of Mlilo (= Moliro), Lake Tanganyika].

Paludomusspiralis Reeve, 1847

Original reference.Reeve (1847: pl. 3, fig. 15).

Type locality. “Streams of Ceylon” [Sri Lanka].

Paludomusspurca Adams & Adams, 1854

Original reference.Adams and Adams (1853–1858: pl. 36, fig. 2).

Original spelling.Paludomusspurcus Adams & Adams, 1854.

Type locality. Not given.

Paramelaniastanleyana Bourguignat, 1885

Original reference.Bourguignat (1885a: 75).

Type locality. “Plages de Pambété et de Mlilo” [Zambia, beach of Pambété and Democratic Republic of the Congo, beach of Mlilo (= Moliro), Lake Tanganyika].

Edgarianassastappersi Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 173).

Original spelling.Edgarianassavar.stappersi Leloup, 1953.

Remarks. Mentioned in the synonymy of Edgardianassa forme spinulosa as a manuscript name of Dautzenberg; unavailable (Code Art. 11.6).

Melaniastephanus Benson, 1836

Original reference.Benson (1836b: 747).

Original spelling.MelaniaStephanus Benson, 1836.

Type locality. From the north-east frontier of Bengal (Benson, 1836a: 350).

† Edgariastockleyi Cox, 1939

Original reference.Cox (1939: 246, pl. 15, figs 3–5).

Type locality. “The Rungwa Sleeping Sickness Concentration Area, lies about eighty miles to the north-west of the present shore of the lake [Tanzania, Rungwa].

Type stratum. “Quaternary deposits of Lake Rukwa”.

Tanaliastomatodon Benson, 1862

Original reference.Benson (1862: 414–415).

Original spelling.Tanalia (?) Stomatodon Benson, 1862.

Type locality. “Habitat in aquis dulcibus montium prope Cottyam, regionis Travancoriæ” [India, Kerala, Kottayam].

Paludomusstraeleni Leloup, 1953

Original reference.Leloup (1953: 14–15, 114, 130–131, 133–138, pl. 2, fig. 16, figs. 4, 57T, 70, 72DD, 73A–C).

Type locality. “Récoltes de la Mission hydrobiologique belge. N^o 38 — Moba, chalutage le long de la côte Nord du village, — 20 m, vase noire (I). N^o 43 — Moba, au large, petite drague, vase. N^o 49 — Baie de Toa, — 15 m, petite drague, sable, cailloux (II). N^o 50 — Au large du cap Bwana n’denge, à 1.000 m, le long de la côte, chalutage depuis 3 milles au Nord-Ouest de la Lugumba jusque par le travers de la rivière, — 20–80 m, fonds divers, vase, rochers, coquilles et spécimens vivants. N^o 80 — Dans la baie de Burton, au large de la rivière Mutambala, — 40 m, chalut à panneaux, sable vaseux. N^o 127 — Baie d’Utinta, 45–65 m, drague à herse, sable vaseux (III). N^o 133 — Baie de Katibili, à 500 m de la rive, — 65–70 m, drague à herse, sable (IV). N^o 138 — Baie de Bracone, dans l’île Kavala; — 12 m, petite drague, sable cailloux (V). N^o 143 — Baie au Sud de la Malagarasi, à la pointe Sud du delta et devant la rivière. N^o 246 — Dans la baie de Burton, à 1 mille au large de Baraka, — 30 m, petite drague, sable. N^o 267 — Baie de Nyanza, par le travers, au départ, ± 60 m (VI). N^o 328 — Laguno de Katibili, dans le goulot, — 0,50 m, sable (VII)” and “au large de Kituta, Sud du lac (stn. 2064, — 76 m)” and “dans la baie de Sumbu (stn. 2069, — 20–70 m)” [Lake Tanganyika].

Paludomusstriata Nevill, 1884 [Unavailable]

Original reference.Nevill (1884: 297).

Remarks. Published as a manuscript name under the unavailable name (introduced without description, definition, or indication) Paludomusmicrosculpta; unavailable (Code Art. 11.6).

Paludomusstriatula Nevill, 1884

Original reference.Nevill (1884: 297–298).

Type locality. “Ceylon” [Sri Lanka].

Paludomussubdentata Nevill, 1884

Original reference.Nevill (1884: 300).

Original spelling.Paludomus [Philopotamis] subdentata Nevill, 1884.

Type locality. “Ceylon” [Sri Lanka].

Paludomussubfasciata Martens, 1908

Original reference.Martens and Thiele (1908: 266–267, pl. 5, fig. 2).

Type locality. “Sungei Guleh, Sangkulirang” [Indonesia, Borneo, East Kalimantan].

Paludomusnigricanssubgranulosa Nevill, 1884

Original reference.Nevill (1884: 299).

Original spelling.Paludomus [Philopotamis] nigricansvar.subgranulosa Nevill, 1884.

Type locality. “Ceylon [and] Newera Ellia” [Sri Lanka; Nuwara Eliya].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Paludomusgrandidierisubmutica Crosse & Fischer, 1878

Original reference.Crosse and Fischer (1878: 74, pl. 1, figs 4, 4a).

Original spelling.PaludomusGrandidieri var. β Submutica Crosse & Fischer, 1878.

Type locality. “Madagascar, dans les ruisseaux de la partie orientale de l’île” [Madagascar, eastern part of the island].

Paludomushanleyisubpicta Nevill, 1884

Original reference.Nevill (1884: 305).

Original spelling.Paludomus [Tanalia] hanleyivar.subpicta Nevill, 1884.

Type locality. “Yatteantotte, Ceylon” [Sri Lanka].

Remarks. See Remarks under Paludomusconicacherraensis Nevill, 1884.

Paludomussulcata Reeve, 1847

Original reference.Reeve (1847: pl. 2, fig. 8a, pl. 3, figs 8b–c).

Original spelling.Paludomussulcatus Reeve, 1847.

Type locality. “In a mountain stream at Ratnapoora, Ceylon” [Sri Lanka, Ratnapura].

† Paludomussuraiensis Gurung, 1998

Original reference.Gurung (1998: 19–20).

Type locality. “Surai Khola, western Nepal”.

Type stratum. “Upper part of the Surai Khola Formation, the Dobata Formation

and lower part of the Dhan Khola Formation”.

Remarks. See also Parmar (2013).

Paludomusswainsoni Dohrn, 1857

Original reference.Dohrn (1857: 125).

Original spelling.PaludomusSwainsoni Dohrn, 1857.

Type locality. “Ceylon” [Sri Lanka].

Paramelaniatabulata Sowerby, 1890

Original reference.Sowerby (1890: fig. 8 and explication of plate).

Type locality. “Lake Tanganyika”.

Remarks. See also Verdcourt (2000).

Spekiazonatatanganikana Bourguignat, 1885

Original reference.Bourguignat (1885a: 37).

Original spelling.Spekiazonata var. Tanganikana Bourguignat, 1885.

Type locality. “Plage de Kapampa, dans le Marungu, à l’ouest du lac“ [Democratic Republic of the Congo, Marungu region, beach of Kapampa, western shore of Lake Tanganyika].

Horeatanganikana Bourguignat, 1888

Original reference.Bourguignat (1888: 27–28, pl. 11, figs 28–29).

Original spelling.HoreaTanganikana Bourguignat, 1888.

Type locality. “Lac Tanganika” [Lake Tanganyika].

Reymondiatanganikana Ancey, 1894 [Invalid]

Original reference.Ancey (1894: 28).

Remarks. Unjustified emendation of Reymondiatanganyicensis Smith, 1889.

Melaniatanganyicensis Smith, 1880

Original reference.Smith (1880b: 427).

Type locality. “Lake Tanganyika”.

Reymondiatanganyicensis Smith, 1889

Original reference.Smith (1889: 175).

Type locality. “Lake Tanganyika”.

Marteliatanganyicensis Dautzenberg, 1908

Original reference.Dautzenberg (1908: 329, pl. 4, figs 11–12).

Type locality. “À M’Pala” [Democratic Republic of the Congo, Mpala, Lake Tanganyika].

Hemimitratangi Chen, 1943

Original reference.Chen (1943: 19–20, pl. 6, fig. 2).

Type locality. “Kiang-yang, northern Fukien Province” [China, Fujian].

Paludomustanjorica Blanford, 1880 [Invalid]

Original reference.Blanford (1880: 220).

Remarks. Unjustified emendation of Helixtanschaurica Gmelin, 1791.

Paludomustanjoriensis Blanford, 1863

Original reference.Blanford (1863: 171).

Original spelling.PaludomusTanjoriensis Blanford, 1863.

Type locality. Not given, but by reference to Gmelin (1791: 3655): “Habitat in Coromandel aquis dulcibus” [India, Coromandel]. See also Chemnitz (1786: 174).

Remarks. Intended as an emendation for the unavailable name of the nominal taxon Helixfluviatilistanschauriensis Chemnitz, 1786 (given as tanschauriensis and a reference to Gmelin). Interpreted here not as an emendation but as a newly introduced name by Blanford (1863).

Helixtanschaurica Gmelin, 1791

Original reference.Gmelin (1791: 3655).

Original spelling.Paludomuslanschaurica Gmelin, 1791.

Type locality. “Habitat in Coromandel aquis dulcibus” [India, lives in freshwater in Coromandel]. According to Chemnitz (1786) “bey Tirutschinapalli und Tanschaur auf Coromandel” [India, Tamil Nadu, near Tiruchirappalli and Thanjavur in Coromandel].

Remarks. Spelled as Helixlanschaurica by Gmelin (1791: 3655), which is interpreted as an incorrect original spelling (Code Art. 33.5.1), as the reference to Chemnitz (1786: 175) suggests that Gmelin was aware of the correct etymology of the species epithet. The name Helixlanschaurica has rarely been used as the valid name of a taxon (Rees 1819; Woodarch 1820; Wodarch and Mawe 1822, 1825, 1827, 1831, 1833; Bosc 1802, 1824, 1836; Pfeiffer 1840, 1841) and to our knowledge only once after 1900 (Coan and Petit 2011), whereas Helixtanschaurica is comparatively frequently used (e.g., Gmelin 1792; Frauenfeld 1864 as Helixtanschaurica; Schmidt 1818 as Melaniatanschaurica; Gray 1855 as Paludinatanschaurica; Hanley and Theobald 1876; Preston 1915b; Seshaiya, 1929, 1934; Jacob 1959; Satyamurty 1960; Starobogatov 1970; Subba Rao and Mitra 1979; Brown 1980, 1994; Fretter 1984; Brown and Mandahl-Barth 1989; Daniel et al. 1998; Dillon 2000; Strong and Glaubrecht 2002; Thasun Amarasinghe and Krishnarajah 2009; Strong et al. 2011 as Paludomustanschaurica; Starmühlner 1974, 1977, 1979; Subba Rao 1989 as Paludomustanschauricus) and usually attributed to Gmelin (1791) (although sometimes with a wrong date). Paludomussanschauricus as used by Adams and Adams (1853–1854) is an incorrect subsequent spelling.

Helixfluviatilistanschauriensis Chemnitz, 1786 [Unavailable]

Original reference.Chemnitz (1786: 174, pl. 135, 1243a–b).

Original spelling.HelixfluviatilisTanschauriensis Chemnitz, 1786.

Type locality. “Sie ist bey Tirutschinapalli und bey Tanschaur auf Coromandel in den dortigen Bächen und süssen Wassern gefunden worden” [India, Tamil Nadu, near Tiruchirappalli and Thanjavur in Coromandel].

Remarks. Published in a work rejected by ICZN Direction 1 (ICZN 1987).

Cleopatratchadiensis Germain, 1905

Original reference.Germain (1905b: 328, footnote).

Original spelling.CleopatraTchadiensis Germain, 1905.

Type locality. “Dans le Tchad, notamment aux environs de l’archipel Kouri” [Chad/ Niger/Nigeria/Cameroon, Lake Chad, Kouri Islands].

Remarks.Cleopatrabulimoidesvartschadiensis as used by Kobelt (1909) is an incorrect subsequent spelling.

Paludomustennantii Reeve, 1847

Original reference.Reeve (1847: pl. 3, figs 12a–c).

Original spelling.PaludomusTennantii Reeve, 1847.

Type locality. “In a rocky stream flowing from Adam’s Peak, Ceylon” [Sri Lanka, near Adam’s Peak].

Remarks.Tanaliatennentii in Layard (1855) and Paludomus (Tanalia) aculeatavar.tennenti in Nevill (1884) are incorrect subsequent spellings.

Turbonillaterebriformis Smith, 1890

Original reference.Smith (1890: 95).

Original spelling.Turbonilla?terebriformis Smith, 1890.

Type locality. “Lake Tanganyika”.

Philopotamisthwaitesii Layard, 1855

Original reference.Layard (1855: 93–94).

Original spelling.Paludomus Thwaitsii Layard, 1855.

Type locality. “Weyweldenia, a rocky rivulet [and] at Kandanganoa and in the Calloo ganga” [Sri Lanka].

Remarks.Paludomus (Tanalia) aculeatavar.thwaitesi is an incorrect subsequent spelling by Nevill (1884).

Limnotrochusthomsoni Smith, 1880

Original reference.Smith (1880b: 425).

Original spelling.LimnotrochusThomsoni Smith, 1880.

Type locality. “Lake Tanganyika”.

Paramelaniatiarella Martens, 1895

Original reference.Martens (1895: 187).

Original spelling.Paramelania (Edgaria) tiarella Martens, 1895.

Type locality. “Tanganyika” [Lake Tanganyika].

Paramelaniatimida Bourguignat, 1888

Original reference.Bourguignat (1888: 35–36, pl. 15, figs 24–25).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Pseudocleopatratogoensis Thiele, 1928

Original reference.Thiele (1928: 394–395, 400, 402, textfig. 58, pl. 8, figs 18, 18a).

Type locality. “Fluß Volta bei Apaso (Togo)” [Ghana, Apaso, now inundated by Lake Volta].

Paludomustorrenticola Dohrn, 1858

Original reference.Dohrn (1858: 536).

Original spelling.Paludomus (Tanalia) torrenticola Dohrn, 1858.

Type locality. “Inhabitants of mountain-streams in Ceylon” [Sri Lanka].

Cleopatratrabonjiensis Smith, 1882

Original reference.Smith (1882: 384, pl. 22, figs 10–11).

Type locality. “A small lake at Trabonjy, in the north-west central part of the island” [Madagascar, Trabonjy].

Paludomustravancorica Blanford, 1880

Original reference.Blanford (1880: 219–220, pl. 2, fig. 22).

Type locality. “In Travancor (p. 219) and more specifically: in streams traversing the plains between Trevandrum and the foot of the Aghastyamali hill (p. 220)” [India, Kerala].

Paludomustrifasciata Reeve, 1852

Original reference.Reeve (1852: 126).

Original spelling.Paludomustrifasciatus Reeve, 1852.

Type locality. “Branch of the Ganges” [India/Bangladesh].

Bithyniatrifasciata Frauenfeld, 1864 [Unavailable]

Original reference.Frauenfeld (1864: 583, 655).

Original spelling.Bythiniatrifasciata Frauenfeld, 1864.

Type locality. Not given.

Remarks. Attributed to Parreyss; published in the synonymy of Bithyniabulimoides (Olivier, 1804). Also mentioned by Martens (1865: 203) in the synonymy of Paludina (Cleopatra) bulimoides. Treated as an available name, Paludinatrifasciata, of a taxon by Paetel (1873: 64). However, the name was not adopted, i.e. the unavailable name was not used as the valid name of a taxon in a way which establishes it as a new name with its own authorship and date in accordance with the Code (Art. 12.1), and therefore was not made available under the Code (Art. 11.6.1) by Paetel (1873).

Edgariatrochlearis Leloup, 1953 [Unavailable]

Original reference.Leloup (1953: 166).

Remarks. Mentioned in the synonymy of Edgardianassa forme typica; unavailable (Code Art. 11.6).

Marteliatanganyicensistypica Leloup, 1953 [Invalid]

Original reference.Leloup (1953: 107, 111, 113, 116, 146, pl. 2, fig2, figs 53, 56, 80–81).

Original spelling.Paludomustanganyicensis forme typica Leloup, 1953.

Remarks. Name for the nominal forme.

Bridouxiagirauditypica Leloup, 1953 [Invalid]

Original reference.Leloup (1953: 115, 130, 132, 136, 140–144, pl. 13, figs 6A–B, figs 57V–W, 70, 72J, 75, 78A–B, E).

Original spelling.Bridouxiagiraudi forme typica Leloup, 1953.

Remarks. Name for the nominal forme.

Edgarianassatypica Leloup, 1953 [Invalid]

Original reference.Leloup (1953: 130, 132, 154–158, 164–167, pl. 7, fig. 1, pl. 8, fig. 1, pl. 9, fig. 2, figs 71, 72N, 86).

Original spelling.Edgarianassa forme typica Leloup, 1953.

Remarks. Name for the nominal forme.

Paludomusundata Reeve, 1847

Original reference.Reeve (1847: pl. 1, figs 2a–b).

Original spelling.Paludomusundatus Reeve, 1847.

Type locality. “In rapids flowing from Adam’s Peak, Ceylon” [Sri Lanka, near Adam’s Peak].

Cleopatrapirothiunicarinata Martens, 1886

Original reference.Martens (1886: 126–127).

Original spelling.CleopatraPirothivar.unicarinata Martens, 1886.

Type locality. “Aegypten, Fajum-Becken unweit Adue über den Bats am Bahndamm in 17 Meter Seehöhe” [Egypt, Fayum Basin, close to Adue above the Bats near the train embankment, 17 m. a.s.l.].

Syrnolopsisminutaunicarinata Ancey, 1906

Original reference.Ancey (1906b: 255, 267).

Original spelling.Syrnolopsisminutaf.unicarinata Ancey, 1906.

Type locality. “Lac Tanganika, côte orientale” [Tanzania/Burundi, eastern shore of Lake Tanganyika].

Cleopatrabulimoidesunilirata Germain, 1911

Original reference.Germain (1911: 199, pl. 2, figs 22–24).

Original spelling.Cleopatrabulimoidesvar.unilirata Germain, 1911.

Type locality. “Zone sableuse entre Bosso et l’embouchure de la Komadougou, dans le Tchad” [Between Nigeria, Bosso and Nigeria/Niger, mouth of Komadougou River near Lake Chad], “Kélékorarom, dans le Tchad” [Lake Chad, Kélékorarom Island] and “Intérieur du Tchad, à 30 kilomètres du bord Ouest” [Lake Chad, 30 km from its western shore].

Spekiazonataunisulcata Bourguignat, 1890

Original reference.Bourguignat (1890: 64).

Original spelling.Spekiazonata var. b Unisulcata Bourguignat, 1890.

Type locality. “Plage de Pambété” [Zambia, beach of Pambété, Lake Tanganyika].

† Cleopatravanloockei Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 11, 52–53, fig. 37).

Type locality. “Nkondo locality NK 76, Lake Albert, Uganda”.

Type stratum. “Nkondo Formation, Molluscan Association G3b, Terminal Miocene -Early Pliocene”.

Nassopsisvariabilis Martel & Dautzenberg, 1899

Original reference.Martel and Dautzenberg (1899: 174–175, pl. 8, figs 16–17).

Type locality. “Lac Tanganyika” [Lake Tanganyika].

Paludomusventricosula Nevill, 1884

Original reference.Nevill (1884: 299–300).

Original spelling.Paludomus [Philopotamis] ventriculosa Nevill, 1884.

Type locality. “Ceylon [and] Kandapanne Ella, 4 Corles [and] Hackalle, Ceylon” [Sri Lanka].

Paramelaniavenusta Bourguignat, 1888

Original reference.Bourguignat (1888: 37–38, pl. 16, figs 13–14).

Type locality. “Lac Tanganika” [Lake Tanganyika].

Cleopatraverreauxi Pallary, 1909 [Invalid]

Original reference.Pallary (1909: 64).

Original spelling.CleopatraVerreauxi Pallary, 1909.

Remarks. Unjustified emendation of Bithyniaverrauxiana Bourguignat, 1856; the name as introduced by Bourguignat (1856) was given in the synonymy including a reference to the work (Code Art. 33.2.1).

Bithyniaverreauxiana Bourguignat, 1856

Original reference.Bourguignat (1856: 241–242).

Original spelling.Bithinia Verreauxiana Bourguignat, 1856.

Type locality. “Le Nil, en Égypte” [Egypt, Nile River].

Remarks.Cleopatraverreauxi as used by Kobelt (1909) is apparently an incorrect subsequent spelling, whereas Cleopatraverreauxi Pallary, 1909 is interpreted as an unjustified emendation.

Bithyniavexillata Frauenfeld, 1864 [Unavailable]

Original reference.Frauenfeld (1864: 583, 659).

Original spelling.Bythiniavexillata Frauenfeld, 1864.

Type locality. Not given.

Remarks. Attributed to Parreyss; published in the synonymy of Bithyniabulimoides. Also mentioned by Martens (1865: 203) in the synonymy of Paludina (Cleopatra) bulimoides. Treated as an available name, Paludinavexillata, of a taxon by Paetel (1873: 64). However, the name was not adopted, i.e. the unavailable name was not used as the valid name of a taxon in a way which establishes it as a new name with its own authorship and date in accordance with the Code (Art. 12.1), and therefore was not made available under the Code (Art. 11.6.1) by Paetel (1873).

Bridouxiavilleserriana Bourguignat, 1885

Original reference.Bourguignat (1885a: 30–31).

Type locality. “Sur la plage de Kapampa” [Democratic Republic of the Congo, beach of Kapampa, western shore of Lake Tanganyika].

Tanaliaviolacea Layard, 1855

Original reference.Layard (1855: 92).

Type locality. “A small mountain torrent in a dense forest between Gillymalle and Pallabaddoola, towards Adam’s Peak, Ceylon” [Sri Lanka].

Pseudocleopatravoltana Mandahl-Barth, 1973

Original reference.Mandahl-Barth (1973: 279–281, fig. 3a, h).

Type locality. “River Volta at Daboya” [Ghana, Daboya, River Volta].

Paludomusvondenbuschianus Arifin, 2017 [Unavailable]

Original reference.Arifin (2017: 110, 111).

Remarks. Nomen nudum, introduced without definition or description (Code Art. 13.1.1) or bibliographic reference to a definition or description (Code Art. 13.1.2); also not explicitly indicated as intentionally new (Code Art. 16.1).

† Pseudocleopatrawamalai Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 13, 57, 69–70, 76, 79, fig. 45).

Type locality. “Nkondo locality NK 84, L. Albert, Uganda”.

Type stratum. “Lusso Formation (Biozone I), Upper Nyaburogo and Warwire Formations, Molluscan Associations G4 and G5a, Early to Late Pliocene”.

Cleopatrabulimoideswelwitschi Martens, 1897

Original reference.Martens (1897: 185).

Original spelling.Cleopatrabulimoidesvar.welwitschi Martens, 1897.

Type locality. “Angola, in einem salzhaltigen Bach bei Dungo im Distrikt Pungo-Adongo (am Cuanza)” [Angola, Dungo, Pungo, Cuanza].

Vinunduwestae Michel, 2004

Original reference.Michel (2004: 4–11, figs 1A–D, 2A–I, 3A–C, E, 4B, C, 5A, B, 6C–F, 7A, B, 8A, 9).

Type locality. “Burundi, kilometre marker 29.8 south of Bujumbura, first major rocky shoreline, coordinates –3.640, 29.361 or 3°38'25"S, 29°21'40"E, 20 m water depth. Stromatolite-covered rocks and boulders”.

Remarks. The name was first introduced by West et al. (2003: 77). However, the work includes a disclaimer stating “This publication is not deemed to be valid for formal taxonomic/nomenclatural purposes”.

† Pseudocleopatrawilsoni Van Damme & Pickford, 2003

Original reference.Van Damme and Pickford (2003: 10, 59, fig. 48).

Type locality. “Nkondo locality NK 126, Lake Albert, Uganda”.

Type stratum. “Warwire Formation, Molluscan Association G4, Early Pliocene”.

Paludomusyunnanensis Cheng, 1937

Original reference. Cheng (1937: 445–447, figs 1.2, 2.2).

Original spelling.Paludomus (Hemimitra) yunnanensis Cheng, 1937.

Type locality. “Yangtze River, near Shiikiang, Yunnan Province, China”.

Melaniazanguebarensis Petit de la Saussaye, 1851

Original reference.Petit de la Saussaye (1851: 263–264, pl. 7, fig. 1).

Original spelling.MelaniaZanguebarensis Petit de la Saussaye, 1851.

Type locality. “Zanzibar, côte de Zanguebar (Afrque orientale)” [Tanzania, Zanzibar].

Cleopatrazanguebarica Bourguignat, 1889

Original reference.Bourguignat (1889: 164–165).

Original spelling.CleopatraZanguebarica Bourguignat, 1889.

Type locality. “Cette coquille zanzibarienne parait commune dans les cours d’eau de la vallée du Vouami” [Tanzania, Zanzibar, Vouami Valley].

Melaniazeylanica Lea & Lea, 1851

Original reference.Lea and Lea (1851: 194–195).

Type locality. “Seychelles and Ceylon” [Seychelles; Sri Lanka].

Remarks.Paludomusceylanicus is a subsequent misspelling by Issel (1874).

Meloniazonata von dem Busch, 1842

Original reference.Philippi (1842–1845: 3, pl. 1, fig. 12).

Type locality. “Bengalia” [India/Bangladesh].

Remarks. The entire text with descriptions of von dem Busch’s new Melania names was attributed to von dem Busch on p. 4 of Philippi’s (1842–1845) work.

Cleopatrabroeckizonata Putzeys, 1899

Original reference.Putzeys (1899: 60).

Original spelling.CleopatraBroeckivar.zonata Putzeys, 1899.

Type locality. “De la rivière Aruwimi, affluent du Congo” [Democratic Republic of the Congo, Aruwimi River].

Lithoglyphuszonatus Woodward, 1859

Original reference.Woodward (1859: 349, pl. 47, fig. 3).

Type locality. “Lake Tanganyika in Central Africa”.

Citation

Neiber MT, Glaubrecht M (2019) Annotated nomenclator of extant and fossil taxa of the Paludomidae (Caenogastropoda, Cerithioidea). ZooKeys 850: 1–132. https://doi.org/10.3897/zookeys.850.34238