OLICH: A reference library of DNA barcodes for Nordic lichens

Abstract Abstract

Background

DNA barcodes are increasingly being used for species identification amongst the lichenised fungi. This paper presents a dataset aiming to provide an authoritative DNA barcode sequence library for a wide array of Nordic lichens.

New information

We present 1324 DNA barcode sequences (nrITS) for 507 species in 175 genera and 25 orders. Thirty-eight species are new to GenBank and, for 25 additional species, ITS sequences are here presented for the first time. The dataset covers 20–21% of the Nordic lichenised species. Barcode gap analyses are given and discussed for the three genera Cladonia, Ramalina and Umbilicaria. The new combination Bryobilimbia fissuriseda (Poelt) Timdal, Marthinsen & Rui is proposed for Mycobilimbia fissuriseda and Nordic material of the species, currently referred to as Pseudocyphellaria crocata and Psoroma tenue ssp. boreale, are shown to belong in Pseudocyphellaria citrina and Psoroma cinnamomeum, respectively.

Introduction

Lichens are fungi living in mutualistic symbiosis with green algae and/or cyanobacteria. They are highly polyphyletic and 20–30 independent lichenisation events may have occurred during the evolution of higher Fungi (Schoch et al. 2009, Lücking et al. 2017a). The lichenised fungi currently consists of more than 19,000 species, i.e. about 17% of all known fungal species and 27% of all known ascomycete species (Lücking et al. 2017a). They occur globally in all major terrestrial ecosystems, especially on trees, soil and rock. Although a lichen thallus consists of a symbiotic association between a fungal exhabitant, the mycobiont and one or more photosynthetic algal and/or cyanobacterial inhabitants, the photobionts, the taxonomy and nomenclature follow that of the mycobiont. A lichen thallus may house additional fungi, i.e. parasites, parasymbionts and endophytes (Hawksworth 1982, Richardson 1999, Spribille et al. 2016), making a lichen thallus a complex symbiotic assemblage.

Identifying lichenised fungi has traditionally been based on morphological, anatomical and chemical characters, but DNA-based identification is now increasingly being used. In 2012, the internal transcribed spacer of nuclear ribosomal DNA (ITS) was recommended as the primary fungal DNA barcode marker by the International Fungal Barcoding Consortium (Xu 2016), following studies on its suitability by for example Schoch et al. (2012). For lichenised fungi, studies reporting successful use of the ITS marker for species delimitations include Kelly et al. (2011), Zheng et al. (2011), Parnmen et al. (2013), Leavitt et al. (2014), Magain and Serusiaux (2015) and Divakar et al. (2016). Pino-Bodas et al. (2013), however, found ITS often failing for species delimitations in Cladonia and investigated alternative markers.

In this paper, we present a DNA barcode dataset for Nordic lichens, including 1324 records, comprising 507 species in 175 genera and 25 different orders.

General description

Purpose

This dataset aims to provide an authoritative DNA barcode sequences library for a wide array of Nordic lichens. The library should facilitate DNA-based identification of species for both traditional molecular studies and DNA-metabarcoding studies and should also be a resource for biosystematic research on Nordic lichens.

Additional information

Eighty sequences have been published previously, but are included here as they were produced under the OLICH project, i.e. GenBank IDs KT800002–07 (Westberg et al. 2015), KU687445–60 (McCune et al. 2016), KU873926–32 (Timdal et al. 2016), KY682961 (Pykälä et al. 2018), 38 sequences within the series MG838156–201 (Kistenich et al. 2018), MG968256–61 (Haugan and Timdal 2018) and MK567159–64 (Haugan and Timdal 2019). In addition, 151 sequences (KY266831–981) have been submitted to GenBank previously for an unpublished project.

Results: Thirty-eight species are new to GenBank: Arthonia phaeobaea, Arthopyrenia salicis, Bactrospora corticola, Baeomyces carneus, Buellia arnoldii, B. dives, Candelariella arctica, Cladonia cyanipes, Coccotrema citrinescens, Eopyrenula leucoplaca, Epilichen glauconigellus, E. scabrosus, Gyalecta foveolaris, Lecanora atrosulphurea, L. leptacina, Ochrolechia alaskana, O. alboflavescens, O. gowardii, O. mahluensis, Orphniospora moriopsis, Phaeophyscia kairamoi, Pilophorus dovrensis, Pseudocyphellaria norvegica, Pyrenula occidentalis, Ramalina cuspidata, Rhizocarpon badioatrum, R. bolanderi, R. chioneum, R. jemtlandicum, R. leptolepis, R. rittokense, R. santessonii, R. subgeminatum, Rinodina balanina, R. disjuncta, R. roboris, Stereocaulon capitellatum and S. symphycheilum.

Twenty-five additional species are previously represented in GenBank, but not with ITS sequences: Arthonia didyma, Bryobilimbia fissuriseda, Buellia epigaea, Callome multipartita, Dendrographa latebrarum, Fuscidea mollis, Gyalecta flotowii, G. friesii, G. geoica, Gyrographa gyrocarpa, Halecania alpivaga, Koerberiella wimmeriana, Opegrapha vermicellifera, Pertusaria aspergilla, P. carneopallida, Phlyctis agelaea, Placynthiella oligotropha, Psoronactis dilleniana, Pyrenula laevigata, Ramboldia elabens, Sagiolechia protuberans, Stereocaulon grande, Thelotrema macrosporum, Tremolecia atrata and Varicellaria lactea.

Project description

Title

The name OLICH refers to “contributions from the Oslo lichen herbarium”.

Personnel

Gunnhild Marthinsen (project coordinator), Siri Rui (project engineer), Einar Timdal (taxonomic supervisor), all affiliated to the Natural History Museum, University of Oslo.

Study area description

The Nordic countries, including Svalbard: Norway, mainland (97.0% of the specimens), Sweden (2.0%), Finland (0.5%), Norway, Svalbard (0.5%), Fig. 1.

Figure 1.

Distribution of the OLICH records

Funding

This project is funded by the Norwegian Research Council (RCN 226134) through the Norwegian Barcode of Life project (NorBOL). In addition, the Norwegian Species Information Centre (Artsdatabanken) has funded four projects on lichens under the Norwegian Taxonomy Initiative from which material has been collected and/or sampled (project numbers 70184216, 70184227, 70184233, and 70184237).

Sampling methods

Sampling description

The material was partly freshly collected specimens from the field and partly previously curated specimens in university herbaria. The distribution of specimen age at seqencing is given in Fig. 2. The specimens were initially determined by experts based on morphology and secondary chemistry and the identifications were re-examined at the quality control after sequence acquisition. All collected material were, after sampling, deposited in university herbaria and the entire material is housed in BG (2.1%), H (0.1%), O (94.9%), SAV (0.2%), TRH (1.7%) and TROM (1.1%) (abbreviated according to Index Herbariorum; http://sweetgum.nybg.org/science/ih).

Figure 2.

Distribution of specimen age (years after collecting) at sequencing in the OLICH dataset.

Thin-layer chromatography (TLC) was performed for identification of secondary compounds on selected specimens in accordance with the methods of Culberson (1972), as modified by Menlove (1974) and Culberson and Johnson (1982). Solvent system B' was used for routine examination, A and C for additional confirmation.

Tissue of thallus or preferably apothecia (about 3 mm³) was assembled in 96 well plates obtained from the Canadian Centre for DNA Barcoding (CCDB; http://ccdb.ca). One well was left empty to serve as negative control. The freshly collected material was either sampled in the field or at a field station. In the field, the specimen was first photographed, then tissue was removed with a flame-decontaminated forceps and placed in the well and finally the specimen was collected. Specimens requiring more controlled handling were photographed and sampled either with forceps or scalpel under a binocular lens at a field station. Wells containing fresh material were dried for several days with lids open in a drying chamber containing silica gel. Herbarium material was sampled at the housing institutions under binocular lenses.

Field data was stored on tablets and GPSs. Tissue sampling data and photography data were transferred directly to standard BOLD spreadsheets, whereas the collection data passed through herbarium databases for obtaining institution storing, voucher ID (= sample ID), uniform nomenclature and taxonomical hierarchy and contact information. Material sampled in herbaria used a similar pipeline, although simpler, as collection data was already available.

The plates were sent to CCDB for DNA extraction (following Ivanova et al. 2011) and sequencing of the ITS region (following Kuzmina and Ivanova 2011). Primers used were ITS1-F or ITS5 (forward) and ITS4 (reverse) (Gardes and Bruns 1993, White et al. 1990). Trace files and sequences were uploaded to BOLD (Ratnasingham and Hebert 2007). The sequences were edited at CCDB, but occasionally inspected and re-edited at OLICH.

Quality control

All OLICH sequences were subjected to a local BLASTn search against a database consisting of all lichen ITS sequences in GenBank (downloaded 26-06-2018) and all unpublished lichen ITS sequences were produced at various projects at our own institution, including the OLICH dataset. We inspected automatically generated distance-based neighbour-joining trees for all genera represented in the OLICH dataset, i.e. we merged all OLICH sequences with all GenBank ITS sequences from the same genus before running ClustalW2 (http://www.clustal.org/clustal2) in batch mode (one run for each genus) and saving the trees. For selected cases (Figs. 5–15), phylogenetic analyses were performed through the following pipeline: selecting relevant additional sequences for comparison from the GenBank download; aligning the dataset using the online version of MAFFT (https://mafft.cbrc.jp/alignment/server); trimming ends and manually editing the alignment using BioEdit 7.1.3.0 (Hall 1999); analysing the alignment by the maximum likelihood method using RAxML v. 8.2.10 (Stamatakis 2014) by applying the “ML + rapid bootstrap” method with 100 bootstrap repeats and the GTRGAMMA substitution model; and editing the best-scoring tree using TreeGraph2 (Stöver and Müller 2010).

To further check that the sequences and species determinations were correct and also to check whether species delimitation, based on traditional characters, coincides with ITS species delimitation, we performed barcode gap analyses. The three genera in our dataset with most sampled species were chosen for these analyses; Cladonia, Ramalina and Umbilicaria. Sequences were aligned separately for each genus using the ClustalW algorithm implemented in MEGA7 (Kumar et al. 2016), with additional manual editing. Pairwise genetic distances between all specimens within each of the three genera were calculated in MEGA7, using the K2P substitution model. The distances were summarised in a bar plot, based on a histogram of all pairwise distances. For each species pair within the genera, we plotted the maximum intraspecific distance against the minimum distance to nearest neighbour-species.

Geographic coverage

Description

The Nordic countries, including Svalbard (Fig. 1).

Coordinates

58.02°N and 78.94°N Latitude; 5.06°E and 31.08°E Longitude.

Taxonomic coverage

Description

The OLICH dataset comprises 1324 records of 507 species belonging to 25 orders, 8 classes and 2 phyla of fungi (Table 1). The Nordic lichen flora comprises some 2400–2500 lichenised species (2387 are listed in Santesson et al. (2004)), i.e. the dataset comprises about 20–21% of the currently known Nordic species. Several taxonomic groups are still poorly understood, however, making the production of authoritative DNA-barcodes for many species premature due to uncertain taxonomy and/or lack of fresh material. The dataset is hence biased towards the more well-known groups, mainly the macrolichens in the orders Lecanorales (233 spp.), Peltigerales (62 spp.) and Caliciales (37 spp.) of the class Lecanoromycetes (Ascomycota). The taxonomy and nomenclature follows Nordin et al. (2019) unless commented on under Observations below.

Table 1.

List of specimens included in the OLICH dataset (sequence length >200bp, 1324 records, 507 species). Thin Layer Chromotagrophy (TLC, Chemistry-column) was only done in species complexes where chemical data is relevant for the identification of the species or where there are chemical strains within the species.

Species	Order	Voucher	Sample ID	Process ID	GenBank ID	Chemistry
Acarospora glaucocarpa	Acarosporales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201295	OLICH3852-17	MK811746
Acarospora molybdina	Acarosporales	NO, Finnmark, 2014, E. Timdal	O-L-195556	OLICH1514-14	KY266941
Acarospora wahlenbergii	Acarosporales	NO, Hedmark, 2013, E. Timdal	O-L-184511	OLICH737-13	MK811773
Acarospora wahlenbergii	Acarosporales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184363	OLICH742-13	MK812641
Acarospora wahlenbergii	Acarosporales	NO, Telemark, 2013, E. Timdal	O-L-184419	OLICH870-13	MK811926
Acolium inquinans	Caliciales	NO, Nordland, 2012, J.T. Klepsland	O-L-186246	OLICH2077-15	MK812481
Acolium inquinans	Caliciales	NO, Buskerud, 2012, T.H. Hofton	O-L-194494	OLICH2080-15	MK811701
Acolium inquinans	Caliciales	NO, Aust-Agder, 2014, J.T. Klepsland	O-L-200169	OLICH3168-16	MK812233
Acolium inquinans	Caliciales	NO, Hedmark, 2014, R. Haugan	O-L-207991	OLICH3827-17	MK811747
Alectoria nigricans	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195780	OLICH1652-14	MK812557
Alectoria nigricans	Lecanorales	NO, Finnmark, 2014, M. Westberg	O-L-195858	OLICH1703-14	KY266967
Alectoria nigricans	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195886	OLICH1731-14	KY266922
Alectoria nigricans	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195950	OLICH1795-14	KY266831
Alectoria nigricans	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196314	OLICH2244-15	MK812235
Alectoria nigricans	Lecanorales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200895	OLICH2548-16	MK812358
Alectoria ochroleuca	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195746	OLICH1618-14	KY266936
Alectoria ochroleuca	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195883	OLICH1728-14	KY266959
Alectoria ochroleuca	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-195823	OLICH2195-15	MK811939
Alectoria ochroleuca	Lecanorales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200894	OLICH2547-16	MK812226
Alectoria ochroleuca	Lecanorales	NO, Oppland, 2012, K.A. Lye	O-L-206884	OLICH4129-17	MK812278
Alectoria sarmentosa	Lecanorales	NO, Nord-Trøndelag, 2012, J.T. Klepsland	O-L-186255	OLICH1441-14	MK812505
Alectoria sarmentosa	Lecanorales	NO, Sogn og Fjordane, 2009, T.H. Hofton	O-L-186870	OLICH1452-14	MK812223
Alectoria sarmentosa	Lecanorales	NO, Østfold, 2013, B.P. Løfall & O. Wergeland Krog	O-L-187364	OLICH1460-14	MK812403
Alectoria sarmentosa	Lecanorales	NO, Sogn og Fjordane, 2014, K.J. Grimstad & O. Olsen	O-L-193185	OLICH1478-14	MK812717
Alectoria sarmentosa	Lecanorales	NO, Sør-Trøndelag, 2014, S. Vatne	O-L-200758	OLICH3273-16	MK812509
Allantoparmelia alpicola	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195748	OLICH1620-14	KY266874
Allantoparmelia alpicola	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195850	OLICH1695-14	KY266926
Allantoparmelia alpicola	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195899	OLICH1744-14	KY266946
Allantoparmelia alpicola	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196324	OLICH2254-15	MK811991
Allantoparmelia alpicola	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201391	OLICH4173-17	MK811876
Anaptychia ciliaris	Caliciales	NO, Aust-Agder, 2012, J.T. Klepsland	O-L-186030	OLICH1419-14	MK812701
Anaptychia ciliaris	Caliciales	NO, Hedmark, 2011, T.H. Hofton	O-L-194359	OLICH1483-14	MK812099
Anaptychia ciliaris	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195591	OLICH1532-14	MK812249
Anaptychia ciliaris	Caliciales	NO, Finnmark, 2014, G. Thor	O-L-195736	OLICH1608-14	KY266912
Anaptychia ciliaris	Caliciales	NO, Akershus, 2014, E. Timdal	O-L-199982	OLICH1956-15	MK812295
Anaptychia ciliaris	Caliciales	NO, Oslo, 2014, E. Timdal	O-L-198929	OLICH2482-16	MK812296
Anaptychia runcinata	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195767	OLICH1639-14	MK812680
Anaptychia runcinata	Caliciales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196019	OLICH1864-14	MK811742
Anaptychia runcinata	Caliciales	NO, Østfold, 2016, S. Rui & E. Timdal	O-L-208017	OLICH4189-17	MK811645
Arctocetraria andrejevii	Lecanorales	NO, Finnmark, 2012, H. Henriksen	O-L-180951	OLICH869-13	MK811992	norrangiformic acid (minor), rangiformic acid (major)
Arctoparmelia centrifuga	Lecanorales	NO, Troms, 2014, R. Haugan	O-L-198774	OLICH2399-16	MK812567
Arctoparmelia centrifuga	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200879	OLICH2536-16	MK812513
Arctoparmelia centrifuga	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184727	OLICH942-13	MK812611
Arctoparmelia incurva	Lecanorales	NO, Finnmark, 2014, T. Tønsberg	O-L-195717	OLICH1589-14	KY266852
Arctoparmelia incurva	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-195832	OLICH2204-15	MK811976
Arthonia didyma	Arthoniales	NO, Nord-Trøndelag, 2013, J.T. Klepsland	O-L-197934	OLICH4022-17	MK812348
Arthonia phaeobaea	Arthoniales	NO, Sør-Trøndelag, 2013, J.T. Klepsland	O-L-198129	OLICH3187-16	MK812215
Arthopyrenia salicis	Pleosporales	NO, Nord-Trøndelag, 2016, J.T. Klepsland	O-L-206752	OLICH3471-17	MK812299
Arthopyrenia salicis	Pleosporales	NO, Møre og Romsdal, 2016, J.T. Klepsland	O-L-206774	OLICH3493-17	MK812520
Asahinea chrysantha	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195754	OLICH1626-14	KY266872
Bacidia bagliettoana	Lecanorales	NO, Akershus, 2009, R. Haugan	O-L-161668	OLICH282-13	MG838180
Bacidia bagliettoana	Lecanorales	NO, Troms, 2009, J.T. Klepsland	O-L-164908	OLICH284-13	MG838178
Bacidia subincompta	Lecanorales	NO, Finnmark, 2011, E. Timdal	O-L-170623	OLICH198-13	MG838157	no lichen substances
Bacidia subincompta	Lecanorales	NO, Sogn og Fjordane, 2013, J.T. Klepsland	O-L-197862	OLICH2086-15	MG838186
Bacidia subincompta	Lecanorales	NO, Østfold, 2001, B.P. Løfall	O-L-105331	OLICH310-13	MG838165
Bacidia subincompta	Lecanorales	NO, Oslo, 2014, J.T. Klepsland	O-L-200148	OLICH3180-16	MG838175
Bacidia subincompta	Lecanorales	NO, Nordland, 2015, J.T. Klepsland	O-L-206520	OLICH3420-17	MG838176
Bacidia vermifera	Lecanorales	NO, Troms, 2011, J.T. Klepsland	O-L-177162	OLICH264-13	MG838160
Bacidia vermifera	Lecanorales	NO, Troms, 2009, J.T. Klepsland	O-L-165007	OLICH299-13	MG838169
Bactrospora corticola	Arthoniales	NO, Nord-Trøndelag, 2010, H. Holien	TRH-L-13774	OLICH1208-14	MK811961
Bactrospora corticola	Arthoniales	NO, Nord-Trøndelag, 2009, H. Holien	TRH-L-12953	OLICH1209-14	MK811741
Bactrospora corticola	Arthoniales	NO, Nord-Trøndelag, 2009, H. Holien	TRH-L-12974	OLICH1210-14	MK811643
Bactrospora corticola	Arthoniales	NO, Sør-Trøndelag, 2010, G. Gaarder	TRH-L-651772	OLICH1211-14	MK811979
Bactrospora corticola	Arthoniales	NO, Sør-Trøndelag, 2014, S. Vatne	O-L-200761	OLICH2438-16	MK812695
Baeomyces carneus	Baeomycetales	NO, Østfold, 2003, B. P. Løfall	O-L-125189	OLICH4208-17	MK811898
Baeomyces carneus	Baeomycetales	NO, Østfold, 2010, J.T. Klepsland	O-L-174982	OLICH4421-17	MK812113
Baeomyces placophyllus	Baeomycetales	NO, Telemark, 2014, E. Timdal	O-L-196267	OLICH1942-15	MK812574
Baeomyces placophyllus	Baeomycetales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196313	OLICH2243-15	MK812640
Baeomyces rufus	Baeomycetales	NO, Telemark, 2014, S. Rui & E. Timdal	O-L-194160	OLICH1481-14	MK811923
Baeomyces rufus	Baeomycetales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200015	OLICH1989-15	MK812462
Baeomyces rufus	Baeomycetales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201257	OLICH2765-16	MK812004
Baeomyces rufus	Baeomycetales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201258	OLICH2766-16	MK812643
Baeomyces rufus	Baeomycetales	NO, Aust-Agder, 2014, J.T. Klepsland	O-L-200581	OLICH3271-16	MK812401
Bilimbia lobulata	Lecanorales	NO, Nordland, 2001, B.P. Løfall	O-L-105772	OLICH469-13	MK812451
Bilimbia lobulata	Lecanorales	NO, Troms, 2009, J.T. Klepsland	O-L-164992	OLICH470-13	MK812712
Bilimbia lobulata	Lecanorales	NO, Nordland, 2000, S. Rui & E. Timdal	O-L-51771	OLICH471-13	MK812122
Bilimbia lobulata	Lecanorales	NO, Oppland, 1995, R. Haugan & E. Timdal	O-L-19936	OLICH630-13	MK812573
Bilimbia sabuletorum	Lecanorales	NO, Buskerud, 2008, J.T. Klepsland	O-L-158388	OLICH206-13	MK812338
Bilimbia sabuletorum	Lecanorales	NO, Telemark, 2010, S. Rui & E. Timdal	O-L-163841	OLICH207-13	MK812048	no lichen substances
Bilimbia sabuletorum	Lecanorales	NO, Oslo, 2009, J.T. Klepsland	O-L-164642	OLICH208-13	MK812294
Bilimbia sabuletorum	Lecanorales	NO, Aust-Agder, 2010, J.T. Klepsland	O-L-175077	OLICH209-13	MK812254
Bilimbia sabuletorum	Lecanorales	NO, Hedmark, 2004, R. Haugan	O-L-131717	OLICH632-13	MK811845
Blastenia ammiospila	Teloschistales	NO, Finnmark, 2014, U. Arup	O-L-195889	OLICH1734-14	KY266873
Blastenia crenularia	Teloschistales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196018	OLICH1863-14	MK811924
Blastenia crenularia	Teloschistales	NO, Nordland, 2014, J.T. Klepsland	O-L-200301	OLICH3173-16	MK811921
Blastenia crenularia	Teloschistales	NO, Akershus, 2013, S. Rui & E. Timdal	O-L-203051	OLICH3289-16	MK812322
Brianaria lutulata	Lecanorales	NO, Aust-Agder, 2015, J.T. Klepsland	O-L-206489	OLICH3390-17	MK812659
Brianaria tuberculata	Lecanorales	NO, Aust-Agder, 2015, J.T. Klepsland	O-L-206488	OLICH3389-17	MK811824
Brodoa atrofusca	Lecanorales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200891	OLICH2544-16	MK811833
Brodoa atrofusca	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184308	OLICH725-13	MK811829	atranorin (major), physodic acid (minor), protocetraric acid (major)
Brodoa intestiniformis	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-190610	OLICH1474-14	MK811819
Brodoa intestiniformis	Lecanorales	NO, Hordaland, 2014, E. Timdal	O-L-195806	OLICH1678-14	MK811842
Brodoa intestiniformis	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-195819	OLICH2191-15	MK812115
Brodoa intestiniformis	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184709	OLICH928-13	MK812552
Brodoa oroarctica	Lecanorales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200939	OLICH2660-16	MK811789
Brodoa oroarctica	Lecanorales	NO, Hedmark, 2017, E. Timdal	O-L-207794	OLICH4155-17	MK812012
Brodoa oroarctica	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184297	OLICH726-13	MK812600
Bryobilimbia diapensiae	Lecideales	NO, Finnmark, 2011, E. Timdal	O-L-170679	OLICH1167-14	MK812130
Bryobilimbia diapensiae	Lecideales	NO, Nordland, 2012, R. Haugan	O-L-182225	OLICH1168-14	MK811983
Bryobilimbia diapensiae	Lecideales	NO, Sør-Trøndelag, 2004, R. Haugan	O-L-142025	OLICH144-11	MK812681
Bryobilimbia diapensiae	Lecideales	NO, Troms, 2003, R. Haugan	O-L-141675	OLICH145-11	MK812677
Bryobilimbia diapensiae	Lecideales	NO, Finnmark, 2014, S. Rui	O-L-195960	OLICH1805-14	KY266915
Bryobilimbia fissuriseda	Lecideales	NO, Oppland, 2005, E. Timdal	O-L-148945	OLICH627-13	MK811695
Bryobilimbia fissuriseda	Lecideales	NO, Oppland, 2006, R. Haugan	O-L-145176	OLICH628-13	MK812621
Bryobilimbia fissuriseda	Lecideales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184390	OLICH647-13	MK811896
Bryobilimbia hypnorum	Lecideales	NO, Nordland, 2014, J.T. Klepsland	O-L-200331	OLICH3196-16	MK812370
Bryobilimbia hypnorum	Lecideales	NO, Aust-Agder, 2014, J.T. Klepsland	O-L-200584	OLICH3198-16	MK811910
Bryocaulon divergens	Lecanorales	NO, Hedmark, 2013, R. Haugan	O-L-190313	OLICH1472-14	MK812080
Bryocaulon divergens	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195982	OLICH1827-14	KY266917
Bryocaulon divergens	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196360	OLICH1919-15	MK812069
Bryocaulon divergens	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196317	OLICH2247-15	MK811658
Bryoplaca sinapisperma	Teloschistales	NO, Troms, 2014, J.T. Klepsland	O-L-200396	OLICH4055-17	MK812160
Bryoria bicolor	Lecanorales	NO, Buskerud, 2009, S. Svantesson	O-L-157404	OLICH1369-14	MK811780
Bryoria bicolor	Lecanorales	NO, Rogaland, 2011, J.T. Klepsland	O-L-176874	OLICH1396-14	MK812469
Bryoria bicolor	Lecanorales	NO, Hedmark, 2012, R. Haugan	O-L-179764	OLICH1400-14	MK811677
Bryoria bicolor	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183745	OLICH1403-14	MK812128
Bryoria bicolor	Lecanorales	NO, Hedmark, 2012, R. Haugan	O-L-183997	OLICH1404-14	MK812174
Bryoria capillaris	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-199996	OLICH1970-15	MK812045	alectorialic acid, barbatolic acid
Bryoria capillaris	Lecanorales	SE, Lule Lappmark, 2014, E. Timdal	O-L-195685	OLICH2008-15	MK812229	alectorialic acid, barbatolic acid
Bryoria capillaris	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-195816	OLICH2188-15	MK812443	alectorialic acid, barbatolic acid
Bryoria capillaris	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184687	OLICH871-13	MK812425	alectorialic acid, barbatolic acid, fumarprotocetraric acid
Bryoria fremontii	Lecanorales	FI, Perä-Pohjanmaa, 2014, E. Timdal	O-L-195488	OLICH1498-14	MK812707
Bryoria fremontii	Lecanorales	SE, Härjedalen, 2014, E. Timdal	O-L-195688	OLICH2011-15	MK812076
Bryoria fremontii	Lecanorales	NO, Hedmark, 2014, R. Haugan	O-L-207993	OLICH3830-17	MK812563
Bryoria furcellata	Lecanorales	FI, Perä-Pohjanmaa, 2014, E. Timdal	O-L-195487	OLICH1497-14	MK812435
Bryoria furcellata	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-195834	OLICH2206-15	MK811793
Bryoria furcellata	Lecanorales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200527	OLICH2618-16	MK812653
Bryoria furcellata	Lecanorales	NO, Hedmark, 2014, R. Haugan	O-L-207994	OLICH3831-17	MK812397
Bryoria implexa	Lecanorales	FI, Perä-Pohjanmaa, 2014, E. Timdal	O-L-195485	OLICH1495-14	MK812092	fumarprotocetraric acid
Bryoria implexa	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195887	OLICH1732-14	KY266878	no lichen substances
Bryoria implexa	Lecanorales	SE, Torne Lappmark, 2014, E. Timdal	O-L-195665	OLICH2004-15	MK812090	no lichen substances
Bryoria implexa	Lecanorales	SE, Lule Lappmark, 2014, E. Timdal	O-L-195686	OLICH2009-15	MK812159	no lichen substances
Bryoria implexa	Lecanorales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-197823	OLICH2087-15	MK812714	fumarprotocetraric acid
Bryoria implexa	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196325	OLICH2255-15	MK812347	fumarprotocetraric acid
Bryoria implexa	Lecanorales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200539	OLICH2604-16	MK812375	fumarprotocetraric acid
Bryoria implexa	Lecanorales	NO, Hedmark, 2017, E. Timdal	O-L-207753	OLICH4156-17	MK811675	no lichen substances
Bryoria implexa	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184688	OLICH872-13	MK812231	fumarprotocetraric acid
Bryoria nadvornikiana	Lecanorales	NO, Sogn og Fjordane, 2008, G. Gaarder	O-L-155318	OLICH1367-14	MK812079
Bryoria nadvornikiana	Lecanorales	NO, Hedmark, 2012, R. Haugan	O-L-183994	OLICH2172-15	MK811678
Bryoria nadvornikiana	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-176834	OLICH2173-15	MK812465
Bryoria nadvornikiana	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-198885	OLICH2397-16	MK812326
Bryoria nadvornikiana	Lecanorales	NO, Hedmark, 2017, E. Timdal	O-L-207754	OLICH4157-17	MK812417	alectorialic acid (major), barbatolic acid (minor), fumarprotocetraric acid (minor)
Bryoria nadvornikiana	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184689	OLICH873-13	MK812390
Bryoria nitidula	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195715	OLICH1587-14	KY266934	fumarprotocetraric acid (major), protocetraric acid (minor)
Bryoria simplicior	Lecanorales	FI, Inarin Lappi, 2014, E. Timdal	O-L-195506	OLICH1505-14	MK812146
Bryoria simplicior	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-195825	OLICH2197-15	MK812415
Bryoria simplicior	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200936	OLICH2562-16	MK811887
Bryoria smithii	Lecanorales	NO, Vest-Agder, 2011, J.T. Klepsland	O-L-177044	OLICH2165-15	MK811714
Bryoria smithii	Lecanorales	NO, Rogaland, 2011, J.T. Klepsland	O-L-176881	OLICH2166-15	MK812250
Bryoria tenuis	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196345	OLICH1905-15	MK812591
Buellia arnoldii	Caliciales	NO, Aust-Agder, 2015, H. Holien	TRH-L-16501	OLICH3670-17	MK811634
Buellia badia	Caliciales	NO, Vestfold, 2015, E. Timdal	O-L-198941	OLICH2498-16	MK812426
Buellia disciformis	Caliciales	NO, Troms, 2014, J.T. Klepsland	O-L-200381	OLICH4052-17	MK812587
Buellia dives	Caliciales	NO, Nord-Trøndelag, 2015, H. Holien	TRH-L-16419	OLICH3631-17	MK811893
Buellia epigaea	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195523	OLICH1507-14	KY266900
Buellia erubescens	Caliciales	NO, Nord-Trøndelag, 2015, U. Nordin	O-L-201275	OLICH3229-16	MK811955
Bunodophoron melanocarpum	Lecanorales	NO, Rogaland, 2011, H. Fjeldstad	O-L-177411	OLICH1399-14	MK812059
Bunodophoron melanocarpum	Lecanorales	NO, Hordaland, 2011, J.T. Klepsland	O-L-177097	OLICH2161-15	MK812620
Bunodophoron melanocarpum	Lecanorales	NO, Sogn og Fjordane, 2010, J.T. Klepsland	O-L-175004	OLICH2163-15	MK811949
Calicium adspersum	Caliciales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200822	OLICH2410-16	MK812068
Calicium pinicola	Caliciales	NO, Oppland, 2015, R. Haugan	O-L-204191	OLICH4087-17	MK811674
Calicium viride	Caliciales	NO, Hedmark, 2017, S. Rui & E. Timdal	O-L-207800	OLICH4161-17	MK812371
Callome multipartita	Peltigerales	NO, Buskerud, 2013, S. Reiso & T. Høitomt	O-L-194256	OLICH2054-15	MK811803
Calogaya biatorina	Teloschistales	NO, Sogn og Fjordane, 2013, J.T. Klepsland	O-L-197877	OLICH2459-16	MK812288
Calogaya decipiens	Teloschistales	NO, Oslo, 2014, R. Haugan et al.	O-L-196061	OLICH1877-14	MK812143
Calogaya decipiens	Teloschistales	NO, Hedmark, 1998, R. Haugan	O-L-55723	OLICH4389-17	MK812258
Calogaya decipiens	Teloschistales	NO, Akershus, 2010, R. Haugan	O-L-167011	OLICH4390-17	MK811994
Caloplaca chlorina	Teloschistales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-198079	OLICH4423-17	MK811786
Caloplaca chlorina	Teloschistales	NO, Telemark, 2014, J.T. Klepsland	O-L-200720	OLICH4425-17	MK811988
Caloplaca demissa	Teloschistales	NO, Rogaland, 2010, J.T. Klepsland	O-L-200085	OLICH3270-16	MK812156
Caloplaca demissa	Teloschistales	NO, Sogn og Fjordane, 2010, S. Vatne	O-L-169301	OLICH3953-17	MK812032
Caloplaca monacensis	Teloschistales	NO, Buskerud, 1998, H. Bratli	O-L-39356	OLICH4422-17	MK811632
Calvitimela armeniaca	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195741	OLICH1613-14	KY266899	alectorialic acid
Calvitimela livida	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195842	OLICH1687-14	KY266947	atranorin, stictic acid
Calvitimela melaleuca	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195711	OLICH1583-14	KY266838	alectorialic acid, atranorin
Calvitimela melaleuca	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195915	OLICH1760-14	KY266897	alectorialic acid, norstictic acid
Candelaria concolor	Candelariales	NO, Rogaland, 2015, T. Tønsberg & E. Nygaard	BG-L-98168	OLICH3699-17	MK812350
Candelariella arctica	Candelariales	NO, Finnmark, 2014, J.T. Klepsland	O-L-200509	OLICH3171-16	MK812015
Candelariella aurella	Candelariales	NO, Troms, 1998, R. Armstrong	TROM_L_50030	OLICH3532-17	MK812178
Candelariella coralliza	Candelariales	NO, Akershus, 2014, E. Timdal	O-L-199992	OLICH1966-15	MK812334
Candelariella coralliza	Candelariales	NO, Buskerud, 2014, E. Timdal	O-L-195833	OLICH2205-15	MK812024
Candelariella coralliza	Candelariales	NO, Aust-Agder, 2004, J.T. Klepsland	O-L-186001	OLICH2460-16	MK811884
Candelariella coralliza	Candelariales	NO, Telemark, 2015, E. Timdal & S. Rui	O-L-200875	OLICH3836-17	MK812646
Candelariella placodizans	Candelariales	NO, Troms, 2003, J.W. Bjerke	TROM_L_521176	OLICH3583-17	MK812262
Candelariella placodizans	Candelariales	NO, Troms, 2009, J.T. Klepsland	O-L-164896	OLICH3949-17	MK811687
Candelariella placodizans	Candelariales	NO, Nordland, 2000, E. Timdal	O-L-51849	OLICH3950-17	MK811709
Candelariella placodizans	Candelariales	NO, Oppland, 2009, R. Haugan	O-L-159822	OLICH3951-17	MK812369
Candelariella placodizans	Candelariales	NO, Telemark, 2008, J.T. Klepsland	O-L-164600	OLICH3952-17	MK812523
Candelariella vitellina	Candelariales	NO, Nord-Trøndelag, 1999, H. Bratli	O-L-41044	OLICH3941-17	MK811713
Candelariella vitellina	Candelariales	NO, Vest-Agder, 2011, H. Bratli	O-L-175901	OLICH3943-17	MK811697
Candelariella vitellina	Candelariales	NO, Oslo, 2010, R. Haugan	O-L-166987	OLICH3948-17	MK812582
Cetraria aculeata	Lecanorales	NO, Finnmark, 2014, M. Bendiksby	O-L-195862	OLICH1707-14	KY266883
Cetraria aculeata	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196315	OLICH2245-15	MK811967
Cetraria aculeata	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196032	OLICH3808-17	MK811881
Cetraria aculeata	Lecanorales	NO, Østfold, 2013, K.A. Lye	O-L-194757	OLICH4090-17	MK811694
Cetraria aculeata	Lecanorales	NO, Vest-Agder, 2011, H. Bratli	O-L-175907	OLICH4091-17	MK811732
Cetraria ericetorum	Lecanorales	NO, Hedmark, 2013, R. Haugan	O-L-190276	OLICH1469-14	MK811667
Cetraria ericetorum	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195750	OLICH1622-14	KY266868
Cetraria ericetorum	Lecanorales	NO, Buskerud, 2015, E. Timdal & S. Rui	O-L-200904	OLICH3839-17	MK811635
Cetraria ericetorum	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184702	OLICH884-13	MK811886
Cetraria islandica	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-200003	OLICH1977-15	MK812195
Cetraria islandica	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-195818	OLICH2190-15	MK812355
Cetraria islandica	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196304	OLICH2234-15	MK812550
Cetraria islandica	Lecanorales	NO, Sør-Trøndelag, 2017, E. Timdal	O-L-208279	OLICH4164-17	MK812647
Cetraria islandica	Lecanorales	NO, Sør-Trøndelag, 2017, E. Timdal	O-L-208278	OLICH4165-17	MK812026
Cetraria islandica	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184703	OLICH885-13	MK812121
Cetraria muricata	Lecanorales	NO, Hordaland, 2014, E. Timdal	O-L-195799	OLICH1671-14	MK812154
Cetraria muricata	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-195815	OLICH2187-15	MK812716
Cetraria nigricans	Lecanorales	NO, Troms, 2012, J.T. Klepsland	O-L-186197	OLICH1436-14	MK812225
Cetraria nigricans	Lecanorales	NO, Finnmark, 2011, E. Timdal	O-L-170662	OLICH2159-15	MK812613
Cetraria odontella	Lecanorales	NO, Buskerud, 2012, T.H. Hofton	O-L-194457	OLICH1485-14	MK812629
Cetraria sepincola	Lecanorales	NO, Hedmark, 2013, R. Haugan	O-L-190020	OLICH1463-14	MK812418
Cetraria sepincola	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200881	OLICH2538-16	MK812133
Cetraria sepincola	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184705	OLICH886-13	MK811790
Cetrariella commixta	Lecanorales	NO, Finnmark, 2014, M. Westberg	O-L-195926	OLICH1771-14	KY266843
Cetrariella delisei	Lecanorales	NO, Oppland, 2013, R. Haugan	O-L-190044	OLICH1465-14	MK812542
Cetrariella delisei	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195559	OLICH1517-14	KY266903
Cetrariella delisei	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195763	OLICH1635-14	KY266858
Cetrariella delisei	Lecanorales	NO, Finnmark, 2014, M. Westberg	O-L-195882	OLICH1727-14	KY266955
Cetrariella delisei	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195946	OLICH1791-14	KY266892
Cetrariella delisei	Lecanorales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200893	OLICH2546-16	MK811762
Cetrariella fastigiata	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195560	OLICH1518-14	KY266875
Cetrariella fastigiata	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195762	OLICH1634-14	KY266870
Cetrariella fastigiata	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195985	OLICH1830-14	KY266890
Cetrariella fastigiata	Lecanorales	NO, Finnmark, 2011, E. Timdal	O-L-170481	OLICH2154-15	MK811637
Cetrariella fastigiata	Lecanorales	NO, Troms, 2011, J.T. Klepsland	O-L-177156	OLICH2155-15	MK812493
Cetrelia cetrarioides	Lecanorales	NO, Telemark, 2005, S. Reiso	O-L-141329	OLICH1354-14	MK811823	anziaic acid (minor), atranorin (major), perlatolic acid (major), 4-O-methylolivetoric acid (minor)
Cetrelia cetrarioides	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-176836	OLICH1395-14	MK812086	anziaic acid (minor), atranorin (major), perlatolic acid (major), 4-O-methylolivetoric acid (minor)
Cetrelia cetrarioides	Lecanorales	NO, Sogn og Fjordane, 2009, T.H. Hofton	O-L-186891	OLICH1453-14	MK812180	anziaic acid (minor), atranorin (major), perlatolic acid (major), 4-O-methylolivetoric acid (minor)
Cetrelia cetrarioides	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200022	OLICH2470-16	MK812062	anziaic acid (minor), atranorin (major), perlatolic acid (major), 4-O-methylolivetoric acid (minor)
Cetrelia cetrarioides	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201355	OLICH2740-16	MK812302	anziaic acid (minor), atranorin (major), perlatolic acid (major), 4-O-methylolivetoric acid (minor)
Cetrelia cetrarioides	Lecanorales	NO, Buskerud, 2016, M. Bendiksby & E. Timdal	O-L-208029	OLICH4257-17	MK812008	anziaic acid (minor), atranorin (major), perlatolic acid (major), 4-O-methylolivetoric acid (minor)
Cetrelia monachorum	Lecanorales	NO, Buskerud, 2012, T.H. Hofton	O-L-194448	OLICH4095-17	MK811729	anziaic acid (minor), atranorin (major), imbricaric acid (major), 4-O-denmethylimbricaric acid (minor)
Chaenotheca chlorella	Coniocybales	NO, Sør-Trøndelag, 2014, S. Vatne	O-L-200765	OLICH2447-16	MK812013
Chaenotheca ferruginea	Coniocybales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200838	OLICH2426-16	MK812287
Chaenotheca gracilenta	Coniocybales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201318	OLICH2790-16	MK812452
Chaenotheca gracillima	Coniocybales	NO, Sør-Trøndelag, 2014, S. Vatne	O-L-200773	OLICH2446-16	MK812119
Cladonia albonigra	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195659	OLICH1999-15	MK812147
Cladonia amaurocraea	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195776	OLICH1648-14	MK811717
Cladonia amaurocraea	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195877	OLICH1722-14	KY266961
Cladonia amaurocraea	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196364	OLICH1923-15	MK812471
Cladonia amaurocraea	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196316	OLICH2246-15	MK811885
Cladonia arbuscula	Lecanorales	NO, Finnmark, 2014, A. Kosuthova	O-L-195866	OLICH1711-14	KY266933
Cladonia arbuscula	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196358	OLICH1917-15	MK812466
Cladonia arbuscula	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-196268	OLICH1943-15	MK811810
Cladonia arbuscula	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196301	OLICH2231-15	MK812036
Cladonia arbuscula	Lecanorales	NO, Vestfold, 2015, E. Timdal	O-L-198942	OLICH2497-16	MK811763
Cladonia bellidiflora	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195740	OLICH1612-14	KY266902
Cladonia bellidiflora	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195881	OLICH1726-14	KY266913
Cladonia bellidiflora	Lecanorales	NO, Finnmark, 2014, A. Kosuthova	O-L-195906	OLICH1751-14	KY266845
Cladonia bellidiflora	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195958	OLICH1803-14	KY266921
Cladonia caespiticia	Lecanorales	NO, Østfold, 2012, J.T. Klepsland	O-L-186036	OLICH1420-14	MK811962
Cladonia caespiticia	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200916	OLICH2576-16	MK812605
Cladonia caespiticia	Lecanorales	NO, Hordaland, 2011, J.T. Klepsland	O-L-183605	OLICH4081-17	MK811712
Cladonia cariosa	Lecanorales	NO, Buskerud, 2012, E.W. Hanssen et al.	O-L-186370	OLICH1443-14	MK811826
Cladonia cariosa	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184343	OLICH743-13	MK812622
Cladonia carneola	Lecanorales	NO, Finnmark, 2014, A. Kosuthova	O-L-195963	OLICH1808-14	KY266855
Cladonia cenotea	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196307	OLICH2237-15	MK811749
Cladonia cenotea	Lecanorales	NO, Hedmark, 2014, R. Haugan	O-L-198892	OLICH2393-16	MK812468
Cladonia cenotea	Lecanorales	NO, Hedmark, 2014, R. Haugan	O-L-207986	OLICH3824-17	MK812541
Cladonia cenotea	Lecanorales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201300	OLICH3855-17	MK812267
Cladonia cenotea	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184714	OLICH932-13	MK812699
Cladonia chlorophaea	Lecanorales	NO, Buskerud, 2016, S. Rui & E. Timdal	O-L-208034	OLICH4262-17	MK812242
Cladonia coccifera	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195895	OLICH1740-14	KY266935
Cladonia coniocraea	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196310	OLICH2240-15	MK812094
Cladonia coniocraea	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200908	OLICH2569-16	MK812138
Cladonia cornuta	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196260	OLICH2267-15	MK812497
Cladonia cornuta	Lecanorales	NO, Hedmark, 2015, E. Timdal	O-L-200857	OLICH2508-16	MK812414
Cladonia cornuta	Lecanorales	NO, Hedmark, 2016, E. Timdal	O-L-201516	OLICH3979-17	MK811731
Cladonia cornuta	Lecanorales	NO, Telemark, 2017, E. Timdal	O-L-208059	OLICH4167-17	MK812047
Cladonia cornuta	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184715	OLICH933-13	MK812277
Cladonia crispata	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195880	OLICH1725-14	KY266865
Cladonia crispata	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-195824	OLICH2196-15	MK811863
Cladonia crispata	Lecanorales	NO, Telemark, 2016, S. Rui & E. Timdal	O-L-205005	OLICH3995-17	MK811728
Cladonia crispata	Lecanorales	NO, Vest-Agder, 2016, S. Rui & E. Timdal	O-L-204984	OLICH3996-17	MK812514
Cladonia crispata	Lecanorales	NO, Telemark, 2017, E. Timdal	O-L-208058	OLICH4275-17	MK811707
Cladonia cyanipes	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195758	OLICH1630-14	KY266889
Cladonia cyanipes	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195981	OLICH1826-14	KY266977
Cladonia cyathomorpha	Lecanorales	NO, Vest-Agder, 2012, J.T. Klepsland	O-L-186087	OLICH1430-14	MK811836
Cladonia cyathomorpha	Lecanorales	NO, Aust-Agder, 2010, J.T. Klepsland	O-L-175272	OLICH2147-15	MK811648
Cladonia decorticata	Lecanorales	SE, Torne Lappmark, 2014, E. Timdal	O-L-195679	OLICH2006-15	MK811738
Cladonia ecmocyna	Lecanorales	NO, Hordaland, 2014, E. Timdal	O-L-195793	OLICH1665-14	MK812344
Cladonia ecmocyna	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195956	OLICH1801-14	KY266972
Cladonia ecmocyna	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200933	OLICH2559-16	MK812085
Cladonia fimbriata	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195743	OLICH1615-14	KY266953
Cladonia fimbriata	Lecanorales	NO, Vest-Agder, 2011, J.T. Klepsland	O-L-183619	OLICH2145-15	MK812488	fumarprotocetraric acid
Cladonia fimbriata	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200909	OLICH2570-16	MK811629
Cladonia furcata	Lecanorales	NO, Buskerud, 2011, K.A. Lye & T. Høitomt	O-L-194749	OLICH1492-14	MK811985
Cladonia furcata	Lecanorales	NO, Finnmark, 2014, T. Tønsberg	O-L-195964	OLICH1809-14	KY266930
Cladonia furcata	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196298	OLICH2228-15	MK812624
Cladonia furcata	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196308	OLICH2238-15	MK811830
Cladonia furcata	Lecanorales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201273	OLICH3847-17	MK812251
Cladonia gracilis	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195764	OLICH1636-14	KY266906
Cladonia gracilis	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196359	OLICH1918-15	MK812362
Cladonia gracilis	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196321	OLICH2251-15	MK811813
Cladonia gracilis	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196262	OLICH2269-15	MK811638
Cladonia gracilis	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196047	OLICH3802-17	MK812342
Cladonia grayi	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195757	OLICH1629-14	KY266978
Cladonia krogiana	Lecanorales	NO, Telemark, 2003, B.P. Løfall & A. Ognedal	O-L-127158	OLICH1337-14	MK811907	barbatic acid, chlorovinetorin
Cladonia krogiana	Lecanorales	NO, Akershus, 2004, B.P. Løfall	O-L-129021	OLICH1339-14	MK811663	barbatic acid, chlorovinetorin
Cladonia krogiana	Lecanorales	NO, Akershus, 2012, J.T. Klepsland	O-L-186068	OLICH2136-15	MK812057
Cladonia macrophylla	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-200002	OLICH1976-15	MK811782
Cladonia macrophylla	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200021	OLICH2469-16	MK812683
Cladonia macrophyllodes	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196379	OLICH1938-15	MK812614
Cladonia maxima	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-195829	OLICH2201-15	MK811693
Cladonia maxima	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184361	OLICH692-13	MK812518
Cladonia mitis	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184706	OLICH887-13	MK811894
Cladonia parasitica	Lecanorales	NO, Hedmark, 2014, R. Haugan	O-L-207988	OLICH3825-17	MK811639
Cladonia parasitica	Lecanorales	NO, Hedmark, 2016, E. Timdal	O-L-201525	OLICH3989-17	MK812470
Cladonia phyllophora	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196375	OLICH1934-15	MK812644
Cladonia phyllophora	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196320	OLICH2250-15	MK812551
Cladonia pleurota	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195959	OLICH1804-14	KY266968	usnic acid, zeorin
Cladonia pulvinata	Lecanorales	NO, Sør-Trøndelag, 2015, H. Holien	TRH-L-16327	OLICH2920-16	MK811846	fumarprotocetraric acid, psoromic acid
Cladonia pyxidata	Lecanorales	NO, Østfold, 2012, J.T. Klepsland	O-L-186049	OLICH1425-14	MK812571	fumarprotocetraric acid
Cladonia pyxidata	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195914	OLICH1759-14	KY266877	fumarprotocetraric acid (major), protocetraric acid (minor)
Cladonia pyxidata	Lecanorales	NO, Oslo, 2014, S. Rui & E. Timdal	O-L-196073	OLICH1889-14	MK812676
Cladonia rangiferina	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195755	OLICH1627-14	KY266884
Cladonia rangiferina	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195774	OLICH1646-14	MK811970
Cladonia rangiferina	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196302	OLICH2232-15	MK812460
Cladonia rangiferina	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196037	OLICH3806-17	MK811708
Cladonia rangiferina	Lecanorales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201298	OLICH3854-17	MK812260
Cladonia rangiformis	Lecanorales	NO, Telemark, 2012, S. Reiso	O-L-194209	OLICH2062-15	MK812118	atranorin
Cladonia rangiformis	Lecanorales	NO, Rogaland, 2013, J.T. Klepsland	O-L-198309	OLICH2351-15	MK811656
Cladonia rei	Lecanorales	NO, Telemark, 2014, S. Rui & E. Timdal	O-L-194142	OLICH1479-14	MK811774
Cladonia rei	Lecanorales	NO, Oslo, 2014, J.T. Klepsland	O-L-200262b	OLICH4086-17	MK812389	fumarprotocetraric acid, homosekikaic acid
Cladonia squamosa	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184733	OLICH1415-14	MK812303
Cladonia squamosa	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195646	OLICH1558-14	MK811958
Cladonia squamosa	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195909	OLICH1754-14	KY266973
Cladonia squamosa	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196309	OLICH2239-15	MK812360
Cladonia stellaris	Lecanorales	NO, Finnmark, 2014, A. Kosuthova	O-L-195905	OLICH1750-14	KY266887
Cladonia stellaris	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196263	OLICH2270-15	MK812506
Cladonia stellaris	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184707	OLICH926-13	MK812280
Cladonia straminea	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195976	OLICH1821-14	KY266880
Cladonia strepsilis	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-200000	OLICH1974-15	MK812544
Cladonia strepsilis	Lecanorales	NO, Østfold, 2017, S. Rui & E. Timdal	O-L-208340	OLICH4270-17	MK812494
Cladonia stygia	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-196269	OLICH1944-15	MK812684
Cladonia stygia	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196303	OLICH2233-15	MK812337
Cladonia stygia	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184708	OLICH927-13	MK812669
Cladonia subcervicornis	Lecanorales	NO, Rogaland, 2013, S. Vatne	O-L-194122	OLICH2072-15	MK812018
Cladonia subcervicornis	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196028	OLICH3794-17	MK812661
Cladonia subcervicornis	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196053	OLICH3811-17	MK812155
Cladonia subcervicornis	Lecanorales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-204985	OLICH3998-17	MK811825
Cladonia subcervicornis	Lecanorales	NO, Vest-Agder, 2011, J.T. Klepsland	O-L-176962	OLICH4113-17	MK812379
Cladonia subfurcata	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195987	OLICH1832-14	KY266923
Cladonia subfurcata	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196319	OLICH2249-15	MK812649
Cladonia subfurcata	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200934	OLICH2560-16	MK812597
Cladonia subulata	Lecanorales	NO, Sør-Trøndelag, 2013, R. Haugan	O-L-190281	OLICH1471-14	MK812581
Cladonia subulata	Lecanorales	FI, Inarin Lappi, 2014, E. Timdal	O-L-195490	OLICH1500-14	MK811811
Cladonia subulata	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196299	OLICH2229-15	MK812279
Cladonia subulata	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200914	OLICH2574-16	MK811787
Cladonia sulphurina	Lecanorales	NO, Finnmark, 2014, A. Kosuthova	O-L-195911	OLICH1756-14	KY266846
Cladonia symphycarpa	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195597	OLICH1535-14	MK811971	atranorin
Cladonia turgida	Lecanorales	NO, Akershus, 2014, S. Rui & E. Timdal	O-L-199980	OLICH1954-15	MK811657
Cladonia turgida	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200932	OLICH2558-16	MK812027
Cladonia uncialis	Lecanorales	NO, Østfold, 2013, R. Haugan	O-L-190182	OLICH1467-14	MK812589
Cladonia uncialis	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195939	OLICH1784-14	KY266974
Cladonia verticillata	Lecanorales	NO, Hordaland, 2014, E. Timdal	O-L-195797	OLICH1669-14	MK812461
Cladonia verticillata	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196312	OLICH2242-15	MK811806
Cladonia verticillata	Lecanorales	NO, Sør-Trøndelag, 2015, H. Holien	TRH-L-16326	OLICH2921-16	MK812385	fumarprotocetraric acid
Cladonia zopfii	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196044	OLICH3804-17	MK811805
Coccotrema citrinescens	Pertusariales	NO, Nordland, 2015, T. Tønsberg & A. Botnen	BG-L-99172	OLICH3741-17	MK811814
Coccotrema citrinescens	Pertusariales	NO, Nordland, 2015, T. Tønsberg	BG-L-98897	OLICH3754-17	MK811895
Coccotrema citrinescens	Pertusariales	NO, Møre og Romsdal, 2015, T. Tønsberg & Z. Palice	BG-L-99069	OLICH3774-17	MK812329
Coccotrema citrinescens	Pertusariales	NO, Nordland, 2013, J.T. Klepsland	O-L-198020	OLICH4212-17	MK812183
Coccotrema citrinescens	Pertusariales	NO, Sør-Trøndelag, 2011, R. Haugan	O-L-173864	OLICH4403-17	MK812186	stictic acid
Cornicularia normoerica	Lecanorales	NO, Aust-Agder, 2014, J.T. Klepsland	O-L-200754	OLICH2603-16	MK812042
Cornicularia normoerica	Lecanorales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201280	OLICH3850-17	MK811980
Cystocoleus ebeneus	Capnodiales	NO, Sør-Trøndelag, 1998, R. Haugan	O-L-55770	OLICH3942-17	MK811795
Dactylina ramulosa	Lecanorales	NO, Troms, 2012, J.T. Klepsland	O-L-186199	OLICH1438-14	MK812632
Dendrographa latebrarum	Arthoniales	NO, Møre og Romsdal, 2016, H. Holien	TRH-L-16972	OLICH3682-17	MK811843
Dermatocarpon bachmannii	Verrucariales	NO, Østfold, 2000, B. P. Løfall et al.	O-L-78257	OLICH2297-15	MK812540
Dermatocarpon bachmannii	Verrucariales	NO, Østfold, 2005, B. P. Løfall	O-L-147654	OLICH2305-15	MK812444
Dermatocarpon deminuens	Verrucariales	NO, Østfold, 2000, B. P. Løfall	O-L-78264	OLICH2298-15	MK812502
Dermatocarpon deminuens	Verrucariales	NO, Østfold, 1998, B. P. Løfall	O-L-36303	OLICH2299-15	MK812476
Dermatocarpon deminuens	Verrucariales	NO, Østfold, 2000, B. P. Løfall et al.	O-L-78258	OLICH2304-15	MK812075
Dermatocarpon luridum	Verrucariales	NO, Buskerud, 2009, T.H. Hofton	O-L-187021	OLICH1455-14	MK812236
Dermatocarpon meiophyllizium	Verrucariales	NO, Østfold, 2006, B. P. Løfall	O-L-147725	OLICH2296-15	MK812273
Dermatocarpon meiophyllizium	Verrucariales	NO, Hedmark, 2006, R. Haugan	O-L-145251	OLICH2300-15	MK812522
Dermatocarpon meiophyllizium	Verrucariales	NO, Østfold, 2001, B. P. Løfall	O-L-105979	OLICH2306-15	MK811820
Dermatocarpon miniatum	Verrucariales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196009	OLICH1854-14	MK812245
Dibaeis baeomyces	Pertusariales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200009	OLICH1983-15	MK812560
Dibaeis baeomyces	Pertusariales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201296	OLICH2796-16	MK812219
Diploschistes scruposus	Ostropales	NO, Finnmark, 2014, S. Fernandez	O-L-195913	OLICH1758-14	KY266920
Diploschistes scruposus	Ostropales	NO, Aust-Agder, 2013, J.T. Klepsland	O-L-198319	OLICH2350-15	MK811671
Diplotomma alboatrum	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195577	OLICH1527-14	MK812220
Eopyrenula leucoplaca	Vezdaeales	NO, Telemark, 2016, J.T. Klepsland	O-L-206771	OLICH3490-17	MK811948
Epilichen glauconigellus	Rhizocarpales	NO, Oppland, 2012, S. Rui & E. Timdal	O-L-179907	OLICH1100-14	MK812566
Epilichen glauconigellus	Rhizocarpales	NO, Troms, 2003, R. Haugan	O-L-141634	OLICH1101-14	MK811705
Epilichen scabrosus	Rhizocarpales	NO, Hedmark, 2012, S. Rui & E. Timdal	O-L-179955	OLICH1102-14	MK812558
Epilichen scabrosus	Rhizocarpales	NO, Troms, 2009, J.T. Klepsland	O-L-164936	OLICH832-13	MK812386
Evernia divaricata	Lecanorales	NO, Buskerud, 2011, T.H. Hofton	O-L-194431	OLICH1484-14	MK812536
Evernia divaricata	Lecanorales	NO, Oppland, 2012, T.H. Hofton	O-L-194526	OLICH2315-15	MK811891
Evernia mesomorpha	Lecanorales	NO, Hedmark, 2012, T.H. Hofton	O-L-194511	OLICH1489-14	MK812555
Evernia mesomorpha	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200008	OLICH1982-15	MK811689
Evernia mesomorpha	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-176825	OLICH2308-15	MK811679
Evernia mesomorpha	Lecanorales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200526	OLICH2614-16	MK812464
Evernia prunastri	Lecanorales	NO, Oslo, 2014, S. Rui & E. Timdal	O-L-196076	OLICH1892-14	MK811993
Evernia prunastri	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-199988	OLICH1962-15	MK812247
Evernia prunastri	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196292	OLICH2222-15	MK812058
Evernia prunastri	Lecanorales	NO, Hedmark, 2015, E. Timdal	O-L-200856	OLICH2507-16	MK811838
Evernia prunastri	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200923	OLICH2531-16	MK811722
Flavocetraria cucullata	Lecanorales	NO, Østfold, 2013, B.P. Løfall	O-L-187362	OLICH1459-14	MK812129
Flavocetraria cucullata	Lecanorales	NO, Finnmark, 2014, S. Rui	O-L-195752	OLICH1624-14	KY266924
Flavocetraria cucullata	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195778	OLICH1650-14	MK811878
Flavocetraria cucullata	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196338	OLICH2276-15	MK812660
Flavocetraria cucullata	Lecanorales	NO, Svalbard, 2015, S. Rui	O-L-203288	OLICH2707-16	MK812458
Flavocetraria cucullata	Lecanorales	NO, Buskerud, 2015, E. Timdal & S. Rui	O-L-200903	OLICH3838-17	MK812490
Flavocetraria cucullata	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184721	OLICH938-13	MK812603
Flavocetraria nivalis	Lecanorales	NO, Østfold, 2013, B.P. Løfall	O-L-187361	OLICH1458-14	MK812252
Flavocetraria nivalis	Lecanorales	NO, Finnmark, 2014, S. Rui	O-L-195723	OLICH1595-14	KY266896
Flavocetraria nivalis	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195777	OLICH1649-14	MK811972
Flavocetraria nivalis	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196311	OLICH2241-15	MK811652
Flavocetraria nivalis	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196339	OLICH2277-15	MK812419
Flavocetraria nivalis	Lecanorales	NO, Svalbard, 2015, S. Rui	O-L-203283	OLICH2702-16	MK811681
Flavocetraria nivalis	Lecanorales	NO, Hedmark, 2014, R. Haugan	O-L-207989	OLICH3829-17	MK812064
Flavocetraria nivalis	Lecanorales	NO, Buskerud, 2015, E. Timdal & S. Rui	O-L-200902	OLICH3837-17	MK812356
Flavoparmelia soredians	Lecanorales	NO, Rogaland, 2013, E. Nygaard	BG-L-98015	OLICH2437-16	MK812200
Flavoplaca citrina	Teloschistales	NO, Nord-Trøndelag, 2015, V. Alstrup	O-L-201322	OLICH2845-16	MK811952
Flavoplaca citrina	Teloschistales	NO, Sør-Trøndelag, 2015, R. Haugan	O-L-204037	OLICH3891-17	MK812194
Flavoplaca flavocitrina	Teloschistales	NO, Hedmark, 2015, R. Haugan	O-L-204199	OLICH3887-17	MK811821
Flavoplaca marina	Teloschistales	NO, Rogaland, 2010, J.T. Klepsland	O-L-200087	OLICH3206-16	MK811686
Flavoplaca marina	Teloschistales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200616	OLICH3207-16	MK811757
Frutidella caesioatra	Lecanorales	NO, Troms, 2014, J.T. Klepsland	O-L-200359	OLICH3257-16	MK812163
Frutidella caesioatra	Lecanorales	NO, Sogn og Fjordane, 2010, E. Timdal	O-L-163395	OLICH387-13	MK811625
Frutidella caesioatra	Lecanorales	NO, Nordland, 2010, E. Timdal	O-L-163746	OLICH388-13	MK812446
Frutidella caesioatra	Lecanorales	NO, Nordland, 2010, E. Timdal	O-L-163752	OLICH390-13	MK812346
Frutidella caesioatra	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184399	OLICH657-13	MK812561
Fuscidea mollis	Umbilicariales	NO, Finnmark, 2014, M. Westberg	O-L-195707	OLICH1579-14	KY266980	divaricatic acid
Fuscidea mollis	Umbilicariales	NO, Finnmark, 2014, E. Timdal	O-L-195917	OLICH1762-14	KY266834	divaricatic acid
Fuscopannaria ahlneri	Peltigerales	NO, Nordland, 2008, T.H. Hofton	O-L-186679	OLICH2464-16	MK812382
Fuscopannaria confusa	Peltigerales	NO, Nordland, 2013, J.T. Klepsland	O-L-198017	OLICH2465-16	MK812003
Fuscopannaria ignobilis	Peltigerales	NO, Nord-Trøndelag, 2015, E. Timdal et al.	O-L-201286	OLICH2675-16	MK811940
Fuscopannaria ignobilis	Peltigerales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201281	OLICH2778-16	MK812023
Fuscopannaria ignobilis	Peltigerales	NO, Sør-Trøndelag, 2015, H. Holien	TRH-L-16696	OLICH3629-17	MK812459
Fuscopannaria ignobilis	Peltigerales	NO, Sør-Trøndelag, 2012, H. Fjeldstad	O-L-203080	OLICH4062-17	MK812103
Fuscopannaria mediterranea	Peltigerales	NO, Sogn og Fjordane, 2013, J.T. Klepsland	O-L-197869	OLICH2096-15	MK812182
Fuscopannaria mediterranea	Peltigerales	NO, Hordaland, 2013, B. Nordén & J.B. Jordal	O-L-199530	OLICH3166-16	MK811995
Fuscopannaria praetermissa	Peltigerales	NO, Telemark, 2014, E. Timdal	O-L-196265	OLICH1940-15	MK812190
Fuscopannaria praetermissa	Peltigerales	NO, Nordland, 2013, J.T. Klepsland	O-L-198031	OLICH2097-15	MK811734
Fuscopannaria praetermissa	Peltigerales	NO, Finnmark, 2003, E. Timdal	O-L-123194	OLICH3883-17	MK812710
Fuscopannaria praetermissa	Peltigerales	NO, Troms, 2009, J.T. Klepsland	O-L-174932	OLICH3884-17	MK811837
Gyalecta flotowii	Ostropales	NO, Hordaland, 2013, J.B. Jordal	O-L-199472	OLICH3165-16	MK811791
Gyalecta flotowii	Ostropales	NO, Sogn og Fjordane, 2013, H. Fjeldstad	O-L-203030	OLICH4063-17	MK811730
Gyalecta foveolaris	Ostropales	NO, Finnmark, 2014, T.B. Wheeler	O-L-195735	OLICH1607-14	KY266833
Gyalecta friesii	Ostropales	NO, Sør-Trøndelag, 2014, S. Vatne	O-L-200781	OLICH2442-16	MK812083
Gyalecta geoica	Ostropales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200544	OLICH3164-16	MK811956
Gyalecta jenensis	Ostropales	NO, Finnmark, 2014, M. Westberg	O-L-195708	OLICH1580-14	KY266916
Gyalecta jenensis	Ostropales	NO, Finnmark, 2014, A. Millanes	O-L-195860	OLICH1705-14	KY266964
Gyalecta jenensis	Ostropales	NO, Nordland, 2013, J.T. Klepsland	O-L-198022	OLICH4034-17	MK811684
Gyalecta ulmi	Ostropales	NO, Sør-Trøndelag, 2014, S. Vatne	O-L-200778	OLICH2443-16	MK812657
Gyalolechia bracteata	Teloschistales	NO, Finnmark, 2011, E. Timdal	O-L-170861	OLICH3215-16	MK811756
Gyalolechia flavorubescens	Teloschistales	NO, Telemark, 2014, E. Timdal	O-L-195786	OLICH1658-14	MK811715
Gyrographa gyrocarpa	Arthoniales	NO, Nord-Trøndelag, 2015, H. Holien	TRH-L-16879	OLICH3651-17	MK812164
Gyrographa gyrocarpa	Arthoniales	NO, Aust-Agder, 2015, J.T. Klepsland	O-L-204523	OLICH4234-17	MK812125
Haematomma ochroleucum	Lecanorales	NO, Hordaland, 2015, T. Tønsberg	BG-L-99143	OLICH3704-17	MK812424
Haematomma ochroleucum	Lecanorales	NO, Nordland, 2015, T. Tønsberg	BG-L-99181	OLICH3749-17	MK812361
Haematomma ochroleucum	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196048	OLICH3817-17	MK812222
Halecania alpivaga	Leprocaulales	NO, Finnmark, 2014, M. Westberg	O-L-195847	OLICH1692-14	KY266832
Hertelidea botryosa	Lecanorales	NO, Hedmark, 2016, E. Timdal	O-L-201530	OLICH3988-17	MK811999
Hertelidea botryosa	Lecanorales	NO, Sør-Trøndelag, 2007, E. Timdal	O-L-158480	OLICH415-13	MK812335
Heterodermia speciosa	Caliciales	NO, Buskerud, 2009, T.H. Hofton	O-L-186983	OLICH1454-14	MK812396
Heterodermia speciosa	Caliciales	NO, Oppland, 2011, J.T. Klepsland	O-L-176833	OLICH2287-15	MK812055
Heterodermia speciosa	Caliciales	NO, Oppland, 2011, J.T. Klepsland	O-L-176831	OLICH2288-15	MK812697
Heterodermia speciosa	Caliciales	NO, Oppland, 2011, J.T. Klepsland	O-L-176827	OLICH2290-15	MK812504
Heterodermia speciosa	Caliciales	NO, Oppland, 2011, T. Høitomt et al.	O-L-176666	OLICH2291-15	MK812537
Heterodermia speciosa	Caliciales	NO, Oppland, 2011, J.T. Klepsland	O-L-176840	OLICH2292-15	MK812549
Heterodermia speciosa	Caliciales	NO, Oppland, 2011, J.T. Klepsland	O-L-176824	OLICH2293-15	MK811755
Heterodermia speciosa	Caliciales	NO, Hedmark, 2011, R. Haugan	O-L-173228	OLICH2294-15	MK812405
Hypocenomyce scalaris	Umbilicariales	SE, Dalarna, 2014, E. Timdal	O-L-195692	OLICH2014-15	MK812269
Hypocenomyce scalaris	Umbilicariales	NO, Telemark, 2013, J.T. Klepsland	O-L-198107	OLICH2352-15	MK811723
Hypocenomyce scalaris	Umbilicariales	NO, Telemark, 2013, J.T. Klepsland	O-L-198111	OLICH2366-15	MK812170
Hypocenomyce scalaris	Umbilicariales	NO, Hedmark, 2016, E. Timdal	O-L-201526	OLICH3987-17	MK811809
Hypocenomyce scalaris	Umbilicariales	SE, Jämtland, 2017, E. Timdal	O-L-208182	OLICH4285-17	MK812211
Hypogymnia austerodes	Lecanorales	NO, Hedmark, 2013, R. Haugan	O-L-190094	OLICH1466-14	MK812650
Hypogymnia austerodes	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196349	OLICH1908-15	MK812410
Hypogymnia austerodes	Lecanorales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200537	OLICH2613-16	MK812531
Hypogymnia bitteri	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184690	OLICH874-13	MK811649
Hypogymnia farinacea	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196258	OLICH2265-15	MK811676
Hypogymnia farinacea	Lecanorales	NO, Hedmark, 2010, R. Haugan	O-L-183979	OLICH2279-15	MK812081
Hypogymnia farinacea	Lecanorales	NO, Hedmark, 2010, R. Haugan	O-L-183983	OLICH2280-15	MK812607
Hypogymnia farinacea	Lecanorales	NO, Hedmark, 2010, R. Haugan	O-L-183984	OLICH2281-15	MK812314
Hypogymnia farinacea	Lecanorales	NO, Hedmark, 2011, R. Haugan	O-L-174340	OLICH2283-15	MK812440
Hypogymnia farinacea	Lecanorales	NO, Hedmark, 2010, R. Haugan	O-L-183985	OLICH2285-15	MK812305
Hypogymnia farinacea	Lecanorales	NO, Buskerud, 2013, J.T. Klepsland	O-L-198103	OLICH2369-15	MK812704
Hypogymnia farinacea	Lecanorales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201310	OLICH2785-16	MK811721
Hypogymnia hultenii	Lecanorales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201260	OLICH2767-16	MK811640
Hypogymnia incurvoides	Lecanorales	NO, Rogaland, 2012, J.T. Klepsland	O-L-186120	OLICH1433-14	MK811919	atranorin, physodalic acid, physodic acid, protocetraric acid (minor)
Hypogymnia incurvoides	Lecanorales	NO, Rogaland, 2012, J.T. Klepsland	O-L-186139	OLICH1435-14	MK812392	atranorin, physodalic acid, physodic acid, protocetraric acid (minor)
Hypogymnia incurvoides	Lecanorales	NO, Nord-Trøndelag, 2014, J.T. Klepsland	O-L-200733	OLICH2620-16	MK812655	atranorin, physodalic acid, physodic acid
Hypogymnia incurvoides	Lecanorales	NO, Hordaland, 2014, J.T. Klepsland	O-L-200683	OLICH3161-16	MK812638	atranorin, physodalic acid, physodic acid
Hypogymnia physodes	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195888	OLICH1733-14	KY266854
Hypogymnia physodes	Lecanorales	NO, Finnmark, 2014, M. Westberg	O-L-195921	OLICH1766-14	KY266919
Hypogymnia physodes	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200883	OLICH2565-16	MK811818
Hypogymnia physodes	Lecanorales	NO, Hordaland, 2014, J.T. Klepsland	O-L-200741	OLICH2621-16	MK812700	3-hydroxyphysodic acid, atranorin, physodalic acid, physodic acid
Hypogymnia physodes	Lecanorales	NO, Nord-Trøndelag, 2014, J.T. Klepsland	O-L-200731	OLICH3158-16	MK812007	3-hydroxyphysodic acid, atranorin, physodalic acid, physodic acid
Hypogymnia physodes	Lecanorales	NO, Nordland, 2014, J.T. Klepsland	O-L-200736	OLICH3159-16	MK812416	3-hydroxyphysodic acid, atranorin, physodalic acid, physodic acid
Hypogymnia physodes	Lecanorales	NO, Akershus, 2014, J.T. Klepsland	O-L-200255	OLICH3160-16	MK812648
Hypogymnia physodes	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184691	OLICH875-13	MK812139
Hypogymnia tubulosa	Lecanorales	NO, Telemark, 2014, S. Rui & E. Timdal	O-L-194145	OLICH1480-14	MK811733
Hypogymnia tubulosa	Lecanorales	NO, Rogaland, 2015, T. Tønsberg & E. Nygaard	BG-L-98167	OLICH3698-17	MK812120
Hypogymnia tubulosa	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184692	OLICH876-13	MK811664
Hypogymnia vittata	Lecanorales	NO, Buskerud, 2008, G. Gaarder	O-L-155319	OLICH1368-14	MK811984
Hypogymnia vittata	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195648	OLICH1560-14	MK812063
Hypogymnia vittata	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200917	OLICH2577-16	MK812228
Hypogymnia vittata	Lecanorales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200536	OLICH2612-16	MK812445	atranorin, physodic acid, vittatolic acid
Hypogymnia vittata	Lecanorales	NO, Akershus, 2017, S. Rui & E. Timdal	O-L-206863	OLICH4108-17	MK812679
Hypotrachyna afrorevoluta	Lecanorales	NO, Rogaland, 2011, J.T. Klepsland	O-L-176869	OLICH2182-15	MK812595
Hypotrachyna afrorevoluta	Lecanorales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-205001	OLICH3991-17	MK811750
Hypotrachyna laevigata	Lecanorales	NO, Rogaland, 2010, J.T. Klepsland	O-L-175129	OLICH1391-14	MK812534
Hypotrachyna laevigata	Lecanorales	NO, Rogaland, 2011, J.T. Klepsland	O-L-176890	OLICH1397-14	MK812711
Hypotrachyna revoluta	Lecanorales	NO, Rogaland, 2005, J.T. Klepsland	O-L-141145	OLICH1352-14	MK812078
Hypotrachyna revoluta	Lecanorales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-197837	OLICH2099-15	MK812441
Hypotrachyna revoluta	Lecanorales	NO, Rogaland, 2011, T. Høitomt	O-L-176684	OLICH2183-15	MK812141
Hypotrachyna revoluta	Lecanorales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-205002	OLICH3990-17	MK812580
Hypotrachyna sinuosa	Lecanorales	NO, Rogaland, 2016, T. Tønsberg	BG-L-99501	OLICH3758-17	MK812199
Icmadophila ericetorum	Pertusariales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184734	OLICH1416-14	MK812230
Icmadophila ericetorum	Pertusariales	NO, Finnmark, 2014, E. Timdal	O-L-195557	OLICH1515-14	KY266881
Icmadophila ericetorum	Pertusariales	NO, Finnmark, 2014, H. Holien	O-L-195756	OLICH1628-14	KY266943
Icmadophila ericetorum	Pertusariales	NO, Finnmark, 2014, T. Tønsberg	O-L-195874	OLICH1719-14	KY266876
Icmadophila ericetorum	Pertusariales	NO, Nord-Trøndelag, 2015, H. Holien	TRH-L-16544	OLICH3627-17	MK811879
Imshaugia aleurites	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-196257	OLICH2264-15	MK812672
Japewia subaurifera	Lecanorales	NO, Nord-Trøndelag, 2012, T. Tønsberg	BG-L-94252	OLICH577-13	MK812331
Japewia subaurifera	Lecanorales	NO, Finnmark, 2012, T. Tønsberg	BG-L-94707	OLICH578-13	MK812698
Japewia subaurifera	Lecanorales	NO, Nordland, 2012, T. Tønsberg	BG-L-94659	OLICH596-13	MK812623
Japewia tornoensis	Lecanorales	NO, Buskerud, 2009, J.T. Klepsland	O-L-165163	OLICH357-13	MK812546
Japewia tornoensis	Lecanorales	NO, Buskerud, 2011, J.T. Klepsland	O-L-176857	OLICH358-13	MK812668
Koerberiella wimmeriana	Lecideales	NO, Hordaland, 2010, E. Timdal	O-L-163472	OLICH414-13	MK812168
Lasallia pustulata	Umbilicariales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-204990	OLICH4001-17	MK811849
Lecanactis abietina	Arthoniales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200836	OLICH2424-16	MK812703
Lecanactis abietina	Arthoniales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201261	OLICH2768-16	MK811953
Lecania aipospila	Lecanorales	NO, Finnmark, 2014, J.T. Klepsland	O-L-200506	OLICH3254-16	MK812577
Lecania aipospila	Lecanorales	NO, Finnmark, 2003, E. Timdal	O-L-123172	OLICH3962-17	MK811636
Lecania erysibe	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183713	OLICH3956-17	MK812191
Lecanora atrosulphurea	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195558	OLICH1516-14	KY266931
Lecanora intricata	Lecanorales	NO, Finnmark, 2014, U. Arup	O-L-195841	OLICH1686-14	KY266891
Lecanora intricata	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195876	OLICH1721-14	KY266975
Lecanora leptacina	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195896	OLICH1741-14	KY266950
Lecidea auriculata	Lecideales	NO, Finnmark, 2014, E. Timdal	O-L-195916	OLICH1761-14	KY266956	no lichen substances
Lecidea leucothallina	Lecideales	NO, Oppland, 2013, T. Tønsberg	BG-L-96170	OLICH1065-14	MG968256	pannarin (thallus)
Lecidea leucothallina	Lecideales	NO, Nordland, 2010, E. Timdal	O-L-163621	OLICH179-11	MG968259	pannarin (apothecium), no lichen substances (thallus)
Lecidea leucothallina	Lecideales	NO, Nordland, 2010, E. Timdal	O-L-163673	OLICH180-11	MG968258	pannarin (thallus)
Lecidea leucothallina	Lecideales	NO, Møre og Romsdal, 2010, R. Haugan	O-L-165815	OLICH181-11	MG968260	no lichen substances (thallus)
Lecidea toensbergii	Lecideales	NO, Oppland, 2013, T. Tønsberg	BG-L-95936	OLICH1010-13	MG968261	pannarin (thallus)
Lecidea toensbergii	Lecideales	NO, Møre og Romsdal, 2010, R. Haugan	O-L-165806	OLICH182-11	MG968257	pannarin (thallus)
Leprocaulon quisquiliare	Leprocaulales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200884	OLICH2566-16	MK812636
Leprocaulon quisquiliare	Leprocaulales	NO, Akershus, 2015, S. Rui & E. Timdal	O-L-200816	OLICH2587-16	MK811899
Leprocaulon quisquiliare	Leprocaulales	NO, Østfold, 2017, S. Rui & E. Timdal	O-L-208334	OLICH4265-17	MK812206
Leproplaca cirrochroa	Teloschistales	NO, Oppland, 2011, J.T. Klepsland	O-L-183714	OLICH4395-17	MK812049
Leproplaca cirrochroa	Teloschistales	NO, Telemark, 2009, R. Haugan & S. Reiso	O-L-160423	OLICH4427-17	MK812423
Leproplaca cirrochroa	Teloschistales	NO, Nord-Trøndelag, 1998, R. Haugan, R. Hollas	O-L-44774	OLICH4428-17	MK812709
Leproplaca obliterans	Teloschistales	NO, Akershus, 2013, J.T. Klepsland	O-L-197812	OLICH4426-17	MK812503
Leptogium cochleatum	Peltigerales	NO, Hordaland, 2013, B. Nordén & J.B. Jordal	O-L-199489	OLICH3149-16	MK812473
Leptogium cochleatum	Peltigerales	NO, Hordaland, 2013, B. Nordén & J.B. Jordal	O-L-199545	OLICH3150-16	MK811946
Leptogium saturninum	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195592	OLICH1533-14	MK812662
Leptogium saturninum	Peltigerales	NO, Hordaland, 2013, B. Nordén & J.B. Jordal	O-L-199516	OLICH3250-16	MK812179
Leptogium saturninum	Peltigerales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196046	OLICH3800-17	MK811641
Letharia vulpina	Lecanorales	NO, Hedmark, 2012, R. Haugan	O-L-184032	OLICH1405-14	MK812266	unknown acid, vulpinic acid
Letharia vulpina	Lecanorales	NO, Hedmark, 2012, R. Haugan	O-L-184029	OLICH2179-15	MK812109
Letharia vulpina	Lecanorales	NO, Hedmark, 2011, B. Skinderhaug	O-L-169053	OLICH2180-15	MK811718
Letharia vulpina	Lecanorales	NO, Hedmark, 2010, R. Haugan	O-L-165378	OLICH2181-15	MK811860
Letharia vulpina	Lecanorales	NO, Hedmark, 2014, R. Haugan	O-L-198890	OLICH2395-16	MK811758
Lichenomphalia alpina	Agaricales	NO, Finnmark, 2014, H. Holien	O-L-195732	OLICH1604-14	KY266895
Lichenomphalia velutina	Agaricales	NO, Oslo, 2014, E. Timdal	O-L-200004	OLICH1978-15	MK811822
Lobaria amplissima	Peltigerales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196021	OLICH1866-14	MK812432
Lobaria amplissima	Peltigerales	NO, Sør-Trøndelag, 2010, J.T. Klepsland	O-L-174993	OLICH2176-15	MK812486
Lobaria amplissima	Peltigerales	NO, Sør-Trøndelag, 2013, J.T. Klepsland	O-L-198062	OLICH2326-15	MK811832
Lobaria amplissima	Peltigerales	NO, Buskerud, 2011, T.H. Hofton	O-L-194379	OLICH2327-15	MK811802
Lobaria amplissima	Peltigerales	NO, Sogn og Fjordane, 2010, T.H. Hofton	O-L-187118	OLICH2328-15	MK812689
Lobaria amplissima	Peltigerales	NO, Sør-Trøndelag, 2011, J.T. Klepsland	O-L-176912	OLICH2329-15	MK812498
Lobaria amplissima	Peltigerales	NO, Nord-Trøndelag, 2012, J.T. Klepsland	O-L-186329	OLICH2330-15	MK811725
Lobaria amplissima	Peltigerales	NO, Nord-Trøndelag, 2013, J.T. Klepsland	O-L-198158	OLICH2387-16	MK812569
Lobaria amplissima	Peltigerales	NO, Nordland, 2003, A. Granmo & L. Molster	TROM_L_46394	OLICH3586-17	MK811901
Lobaria linita	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195728	OLICH1600-14	KY266856
Lobaria linita	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195892	OLICH1737-14	KY266932
Lobaria pulmonaria	Peltigerales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196024	OLICH1869-14	MK811957
Lobaria pulmonaria	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196356	OLICH1915-15	MK811682
Lobaria pulmonaria	Peltigerales	NO, Østfold, 2013, O. Wergeland Krog	O-L-193007	OLICH2335-15	MK812447
Lobaria pulmonaria	Peltigerales	NO, Østfold, 2012, O. Wergeland Krog et al.	O-L-193009	OLICH2336-15	MK812330
Lobaria pulmonaria	Peltigerales	NO, Østfold, 2013, O. Wergeland Krog	O-L-193011	OLICH2337-15	MK812533
Lobaria pulmonaria	Peltigerales	NO, Hedmark, 2013, R. Haugan	O-L-190366	OLICH2338-15	MK812202
Lobaria pulmonaria	Peltigerales	NO, Buskerud, 2011, T.H. Hofton & S. Reiso	O-L-194285	OLICH2339-15	MK812005
Lobaria pulmonaria	Peltigerales	NO, Buskerud, 2012, T.H. Hofton	O-L-194456	OLICH2340-15	MK812631
Lobaria pulmonaria	Peltigerales	NO, Vest-Agder, 2011, J.T. Klepsland	O-L-177093	OLICH2346-15	MK812031
Lobaria pulmonaria	Peltigerales	NO, Hedmark, 2012, R. Haugan	O-L-183966	OLICH2347-15	MK812039
Lobaria pulmonaria	Peltigerales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200863	OLICH2515-16	MK811954
Lobaria pulmonaria	Peltigerales	NO, Troms, 2005, A. Granmo	TROM_L_50934	OLICH3536-17	MK811932
Lobaria scrobiculata	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196350	OLICH1909-15	MK812570
Lobaria scrobiculata	Peltigerales	NO, Østfold, 2013, B.P. Løfall & O. Wergeland Krog	O-L-187365	OLICH2341-15	MK812073
Lobaria scrobiculata	Peltigerales	NO, Hedmark, 2013, R. Haugan	O-L-190098	OLICH2342-15	MK812310
Lobaria scrobiculata	Peltigerales	NO, Buskerud, 2012, T.H. Hofton	O-L-194516	OLICH2343-15	MK812214
Lobaria scrobiculata	Peltigerales	NO, Rogaland, 2012, J.T. Klepsland	O-L-186129	OLICH2344-15	MK811651
Lobaria scrobiculata	Peltigerales	NO, Nord-Trøndelag, 2010, H. Bratli	O-L-168086	OLICH2345-15	MK812309
Lobaria scrobiculata	Peltigerales	NO, Troms, 2005, A. Granmo	TROM_L_50935	OLICH3537-17	MK812074
Lobaria scrobiculata	Peltigerales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196051	OLICH3810-17	MK811855
Lobaria virens	Peltigerales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201271	OLICH2774-16	MK811660
Lobaria virens	Peltigerales	NO, Nordland, 2014, J.T. Klepsland	O-L-200346	OLICH3249-16	MK811736
Lobaria virens	Peltigerales	NO, Vestfold, 2003, T.H. Hofton et al.	O-L-128546	OLICH4103-17	MK812046
Lobothallia alphoplaca	Pertusariales	NO, Finnmark, 2014, J.T. Klepsland	O-L-200411	OLICH2658-16	MK812484
Lobothallia melanaspis	Pertusariales	SE, Torne Lappmark, 2014, E. Timdal	O-L-195680	OLICH2007-15	MK811796
Lobothallia melanaspis	Pertusariales	NO, Hedmark, 2014, R. Haugan	O-L-198879	OLICH2390-16	MK811944
Loxospora elatina	Sarrameanales	NO, Nord-Trøndelag, 2014, J.T. Klepsland	O-L-200270	OLICH2657-16	MK812411
Massalongia carnosa	Peltigerales	NO, Aust-Agder, 2013, J.T. Klepsland	O-L-198320	OLICH2371-15	MK812187
Massalongia carnosa	Peltigerales	NO, Buskerud, 2011, E. Timdal	O-L-175425	OLICH3285-16	MK811716
Megalaria grossa	Lecanorales	NO, Akershus, 2010, J.T. Klepsland	O-L-174947	OLICH1036-13	MK811839
Megalaria grossa	Lecanorales	NO, Sør-Trøndelag, 2005, E. Timdal	O-L-149362	OLICH1039-13	MK812151
Megalaria grossa	Lecanorales	NO, Nordland, 2014, J.T. Klepsland	O-L-200308	OLICH2656-16	MK812492
Megalaria pulverea	Lecanorales	NO, Sogn og Fjordane, 2009, J.T. Klepsland	O-L-165033	OLICH1033-13	MK812535
Megalaria pulverea	Lecanorales	NO, Sør-Trøndelag, 2009, R. Haugan	O-L-167026	OLICH1034-13	MK812297
Megalaria pulverea	Lecanorales	NO, Nord-Trøndelag, 2008, J.T. Klepsland	O-L-158214	OLICH1035-13	MK812205
Megalaria pulverea	Lecanorales	NO, Hordaland, 2011, J.T. Klepsland	O-L-176983	OLICH1317-14	MK812565
Megalaria pulverea	Lecanorales	NO, Nordland, 2013, J.T. Klepsland	O-L-197986	OLICH2104-15	MK812321
Melanelia agnata	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196376	OLICH1935-15	MK812394
Melanelia hepatizon	Lecanorales	NO, Hordaland, 2014, E. Timdal	O-L-195807	OLICH1679-14	MK812512
Melanelia hepatizon	Lecanorales	NO, Finnmark, 2014, M. Westberg	O-L-195864	OLICH1709-14	KY266879
Melanelia hepatizon	Lecanorales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201254	OLICH2762-16	MK812070
Melanelia hepatizon	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184723	OLICH939-13	MK812188
Melanelia stygia	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184736	OLICH1418-14	MK812608
Melanelia stygia	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196377	OLICH1936-15	MK812312
Melanelixia fuliginosa	Lecanorales	NO, Vestfold, 2015, E. Timdal	O-L-198943	OLICH2499-16	MK812485
Melanelixia fuliginosa	Lecanorales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200866	OLICH2518-16	MK812619
Melanelixia fuliginosa	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196050	OLICH3818-17	MK812455
Melanelixia glabratula	Lecanorales	NO, Hordaland, 2011, J.T. Klepsland	O-L-176965	OLICH4122-17	MK812674
Melanelixia subargentifera	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-200810	OLICH2475-16	MK812517
Melanelixia subargentifera	Lecanorales	NO, Hedmark, 2015, E. Timdal	O-L-200854	OLICH2505-16	MK812491
Melanelixia subargentifera	Lecanorales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200867	OLICH2519-16	MK811964
Melanelixia subargentifera	Lecanorales	NO, Hedmark, 2014, R. Haugan	O-L-207985	OLICH3820-17	MK812324
Melanelixia subaurifera	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-195997	OLICH1842-14	MK811631
Melanelixia subaurifera	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-199993	OLICH1967-15	MK811942
Melanelixia subaurifera	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-198930	OLICH2483-16	MK812688
Melanohalea exasperata	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196008	OLICH1853-14	MK812395
Melanohalea exasperata	Lecanorales	NO, Telemark, 2015, E. Timdal	O-L-208004	OLICH3878-17	MK812307
Melanohalea exasperata	Lecanorales	NO, Sogn og Fjordane, 2004, A. Breili	O-L-151072	OLICH4123-17	MK812184
Melanohalea infumata	Lecanorales	NO, Troms, 2003, E. Timdal	O-L-123135	OLICH4126-17	MK812651
Melanohalea olivacea	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195933	OLICH1778-14	KY266962
Melanohalea olivacea	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-195826	OLICH2198-15	MK812054
Melanohalea olivacea	Lecanorales	NO, Telemark, 2015, E. Timdal	O-L-208005	OLICH3879-17	MK812340
Melanohalea septentrionalis	Lecanorales	NO, Akershus, 2017, S. Rui & E. Timdal	O-L-206865	OLICH4110-17	MK812433
Menegazzia subsimilis	Lecanorales	NO, Rogaland, 2007, E. Timdal	O-L-149092	OLICH864-13	MK812421
Menegazzia subsimilis	Lecanorales	NO, Rogaland, 2012, J.T. Klepsland	O-L-186134	OLICH944-13	MK812020
Menegazzia terebrata	Lecanorales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200861	OLICH2513-16	MK812093
Menegazzia terebrata	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200924	OLICH2532-16	MK812635
Menegazzia terebrata	Lecanorales	NO, Hordaland, 2013, T. Tønsberg	BG-L-95692	OLICH858-13	MK812654
Menegazzia terebrata	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-176828	OLICH861-13	MK812025
Menegazzia terebrata	Lecanorales	NO, Rogaland, 2012, J.T. Klepsland	O-L-186132	OLICH949-13	MK812145
Micarea botryoides	Lecanorales	NO, Rogaland, 2013, J.I. Johnsen	BG-L-95780	OLICH1006-13	MK812289
Micarea denigrata	Lecanorales	NO, Oppland, 2011, R. Haugan & A.M. Haugan	O-L-174438	OLICH1013-13	MK811696
Micarea misella	Lecanorales	NO, Nord-Trøndelag, 2013, H. Holien	TRH-L-15146	OLICH990-13	MK812399
Micarea paratropa	Lecanorales	NO, Sør-Trøndelag, 2010, R. Haugan	O-L-165711	OLICH101-11	MK812318	no lichen substances
Montanelia disjuncta	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195590	OLICH1531-14	MK811852
Montanelia disjuncta	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196357	OLICH1916-15	MK811711
Montanelia disjuncta	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-198675	OLICH2404-16	KY266910
Montanelia panniformis	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195904	OLICH1749-14	KY266847
Montanelia panniformis	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200016	OLICH1990-15	MK812407
Montanelia panniformis	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200878	OLICH2535-16	MK812634
Montanelia panniformis	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201338	OLICH2722-16	MK812263
Montanelia sorediata	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-195791	OLICH1663-14	MK811963
Montanelia sorediata	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195658	OLICH1998-15	MK811965
Montanelia sorediata	Lecanorales	NO, Buskerud, 2016, M. Stapnes Dahl et al.	O-L-204941	OLICH4115-17	MK811977
Montanelia tominii	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201344	OLICH2728-16	MK812373
Mycobilimbia berengeriana	Lecideales	NO, Aust-Agder, 2011, J.T. Klepsland	O-L-177070	OLICH1113-14	MK812040
Mycobilimbia berengeriana	Lecideales	NO, Oppland, 2011, J.T. Klepsland	O-L-176838	OLICH1114-14	MK812343
Mycobilimbia berengeriana	Lecideales	NO, Finnmark, 2011, E. Timdal	O-L-170554	OLICH1117-14	MK811751	no lichen substances
Mycobilimbia berengeriana	Lecideales	NO, Aust-Agder, 2009, J.T. Klepsland	O-L-167344	OLICH1170-14	MK811880
Mycobilimbia berengeriana	Lecideales	NO, Finnmark, 2014, E. Llop Vallverdu	O-L-195696	OLICH1568-14	KY266871
Mycobilimbia berengeriana	Lecideales	NO, Sør-Trøndelag, 2004, R. Haugan	O-L-142068	OLICH629-13	MK811889
Mycobilimbia carneoalbida	Lecideales	NO, Telemark, 2010, J.T. Klepsland	O-L-175294	OLICH1109-14	MK812353
Mycobilimbia carneoalbida	Lecideales	NO, Aust-Agder, 2005, J.T. Klepsland	O-L-151121	OLICH1120-14	MK811646
Mycobilimbia carneoalbida	Lecideales	NO, Troms, 2009, J.T. Klepsland	O-L-164904	OLICH1121-14	MK812149
Mycobilimbia carneoalbida	Lecideales	NO, Oppland, 2011, R. Haugan	O-L-174298	OLICH1122-14	MK812290
Mycobilimbia carneoalbida	Lecideales	NO, Hedmark, 2011, R. Haugan	O-L-174282	OLICH1123-14	MK812691
Mycobilimbia carneoalbida	Lecideales	NO, Telemark, 2007, E. Timdal	O-L-149057	OLICH1124-14	MK812639
Mycobilimbia epixanthoides	Lecideales	NO, Hedmark, 2006, J.T. Klepsland	O-L-164598	OLICH1105-14	MK812105
Mycobilimbia epixanthoides	Lecideales	NO, Aust-Agder, 2006, J.T. Klepsland	O-L-151124	OLICH1106-14	MK812388
Mycobilimbia epixanthoides	Lecideales	NO, Troms, 2007, J.T. Klepsland	O-L-154903	OLICH1107-14	MK812645
Mycobilimbia epixanthoides	Lecideales	NO, Telemark, 2007, E. Timdal	O-L-149055	OLICH1108-14	MK812402	no lichen substances
Mycobilimbia epixanthoides	Lecideales	NO, Akershus, 2000, R. Haugan	O-L-160303	OLICH1112-14	MK811772
Mycobilimbia pilularis	Lecideales	NO, Vest-Agder, 2009, J.T. Klepsland	O-L-164694	OLICH1110-14	MK812275
Mycobilimbia pilularis	Lecideales	NO, Møre og Romsdal, 2012, B. Nordén & J.B. Jordal	O-L-180158	OLICH1111-14	MK812666
Mycobilimbia pilularis	Lecideales	NO, Hordaland, 2008, J.B. Jordal	O-L-176020	OLICH1125-14	MK812525
Mycobilimbia pilularis	Lecideales	NO, Sogn og Fjordane, 2009, A. Breili	O-L-159489	OLICH1126-14	MK812665
Mycobilimbia tetramera	Lecideales	NO, Finnmark, 2011, E. Timdal	O-L-170621	OLICH1119-14	MK812422	no lichen substances
Mycobilimbia tetramera	Lecideales	NO, Nordland, 2011, R. Haugan	O-L-174132	OLICH1127-14	MK812216	no lichen substances
Mycobilimbia tetramera	Lecideales	NO, Aust-Agder, 2006, J.T. Klepsland	O-L-183493	OLICH1128-14	MK811935
Mycobilimbia tetramera	Lecideales	NO, Finnmark, 2014, E. Llop Vallverdu	O-L-195846	OLICH1691-14	MK812124
Mycobilimbia tetramera	Lecideales	NO, Akershus, 2014, J.T. Klepsland	O-L-200258	OLICH3147-16	MK812192
Mycobilimbia tetramera	Lecideales	NO, Akershus, 2009, J.T. Klepsland	O-L-164634	OLICH631-13	MK812158
Mycobilimbia tetramera	Lecideales	NO, Hedmark, 2008, R. Haugan	O-L-160476	OLICH633-13	MK812208
Mycoblastus sanguinarius	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195768	OLICH1640-14	MK812482
Mycoblastus sanguinarius	Lecanorales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201263	OLICH2769-16	MK811859
Mycoblastus sanguinarius	Lecanorales	NO, Hedmark, 2010, E. Timdal	O-L-163524	OLICH391-13	MK811665
Mycoblastus sanguinarius	Lecanorales	NO, Telemark, 2005, E. Timdal	O-L-138597	OLICH410-13	MK812197
Mycoblastus sanguinarius	Lecanorales	NO, Oppland, 2006, A. Breili	O-L-150999	OLICH617-13	MK812135
Myriolecis straminea	Lecanorales	NO, Finnmark, 2014, M. Westberg	O-L-195870	OLICH1715-14	KY266929
Myriospora smaragdula	Acarosporales	NO, Finnmark, 2014, E. Timdal	O-L-195609	OLICH1543-14	MK811918
Nephroma arcticum	Peltigerales	NO, Troms, 2014, E. Timdal	O-L-195654	OLICH1566-14	MK811882
Nephroma arcticum	Peltigerales	NO, Finnmark, 2014, S. Rui	O-L-195724	OLICH1596-14	KY266888
Nephroma arcticum	Peltigerales	NO, Finnmark, 2014, A. Millanes	O-L-195873	OLICH1718-14	KY266963
Nephroma arcticum	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196368	OLICH1927-15	MK812091
Nephroma arcticum	Peltigerales	NO, Telemark, 2014, E. Timdal	O-L-196272	OLICH1945-15	MK812336
Nephroma arcticum	Peltigerales	NO, Nord-Trøndelag, 2014, Z. Fackovcova et al.	SAV-36	OLICH3073-16	MK812204
Nephroma arcticum	Peltigerales	NO, Nord-Trøndelag, 2014, Z. Fackovcova et al.	SAV-37	OLICH3074-16	MK812592
Nephroma arcticum	Peltigerales	NO, Finnmark, 2008, A. Granmo & G. Mathiassen	TROM_L_46162	OLICH3570-17	MK811792
Nephroma arcticum	Peltigerales	NO, Hedmark, 2014, R. Haugan	O-L-207992	OLICH3828-17	MK812028
Nephroma bellum	Peltigerales	NO, Nord-Trøndelag, 2010, H. Bratli	O-L-168043	OLICH2027-15	MK811672	triterpenoids
Nephroma bellum	Peltigerales	NO, Sogn og Fjordane, 2012, O. Roen	O-L-181644	OLICH2028-15	MK812308	triterpenoids
Nephroma bellum	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196348	OLICH2373-15	MK812483	triterpenoids
Nephroma expallidum	Peltigerales	NO, Finnmark, 2014, M. Westberg	O-L-195701	OLICH1573-14	KY266905
Nephroma expallidum	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195891	OLICH1736-14	KY266844
Nephroma expallidum	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196346	OLICH1906-15	MK812408
Nephroma expallidum	Peltigerales	NO, Finnmark, 2011, E. Timdal	O-L-170548	OLICH2365-15	MK811945
Nephroma expallidum	Peltigerales	NO, Svalbard, 2015, S. Rui	O-L-203292	OLICH2712-16	MK812041
Nephroma helveticum	Peltigerales	NO, Buskerud, 2013, T.H. Hofton	O-L-196491	OLICH2036-15	KT800007	triterpenoids
Nephroma helveticum	Peltigerales	NO, Oppland, 2011, J.T. Klepsland	O-L-181601	OLICH2037-15	KT800006	triterpenoides (T4, T1)
Nephroma laevigatum	Peltigerales	NO, Nord-Trøndelag, 2010, H. Bratli	O-L-168088	OLICH2025-15	MK812687
Nephroma laevigatum	Peltigerales	NO, Sogn og Fjordane, 2010, J.T. Klepsland	O-L-175024	OLICH2038-15	MK812108	pigments, triterpenoid
Nephroma laevigatum	Peltigerales	NO, Nord-Trøndelag, 2015, E. Timdal et al.	O-L-201288	OLICH2676-16	MK812261
Nephroma laevigatum	Peltigerales	NO, Troms, 2010, K. Wannebo Nilsen	TROM_L_48759	OLICH3531-17	MK811868
Nephroma laevigatum	Peltigerales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196055	OLICH3812-17	MK811969
Nephroma laevigatum	Peltigerales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-204997	OLICH4002-17	MK812002
Nephroma parile	Peltigerales	NO, Finnmark, 2014, T. Tønsberg	O-L-195875	OLICH1720-14	KY266927	triterpenoids
Nephroma resupinatum	Peltigerales	NO, Hedmark, 2013, R. Haugan	O-L-190528	OLICH2020-15	MK811650
Nephroma resupinatum	Peltigerales	NO, Sogn og Fjordane, 2010, E. Timdal	O-L-163401	OLICH2022-15	MK812367
Nephroma resupinatum	Peltigerales	NO, Troms, 2011, J.T. Klepsland	O-L-177168	OLICH2023-15	MK812612
Nephroma resupinatum	Peltigerales	NO, Nord-Trøndelag, 2011, J.T. Klepsland	O-L-183598	OLICH2024-15	MK811912	no lichen substances
Nephroma resupinatum	Peltigerales	NO, Troms, 2005, A. Granmo	TROM_L_50937	OLICH3538-17	MK812301
Nephroma resupinatum	Peltigerales	NO, Finnmark, 2008, A. Granmo & G. Mathiassen	TROM_L_48235	OLICH3588-17	MK812553
Nephroma tangeriense	Peltigerales	NO, Hordaland, 2012, J.T. Klepsland	O-L-186150	OLICH2018-15	MK811986	pigments, triterpenoides (T3, T4, T6)
Nephroma tangeriense	Peltigerales	NO, Rogaland, 2010, J.T. Klepsland	O-L-181602	OLICH2019-15	MK812084	pigments, triterpenoides (T2, T3, T6)
Nephroma tangeriense	Peltigerales	NO, Vest-Agder, 2011, J.T. Klepsland	O-L-183599	OLICH2039-15	MK812530	pigments, triterpenoids
Nephroma tangeriense	Peltigerales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-197824	OLICH2105-15	MK812579	pigments
Nephroma tangeriense	Peltigerales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-197826	OLICH2106-15	MK812593
Nevesia sampaiana	Peltigerales	NO, Sogn og Fjordane, 2009, H. Fjeldstad	O-L-203003	OLICH3277-16	MK812089
Normandina pulchella	Verrucariales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-204993	OLICH4003-17	MK812131
Ochrolechia alaskana	Pertusariales	NO, Finnmark, 2008, A. Granmo & G. Mathiassen	TROM_L_60552	OLICH3582-17	MK811874
Ochrolechia alboflavescens	Pertusariales	NO, Nord-Trøndelag, 2015, F. Jonsson	O-L-201276	OLICH3230-16	MK812244
Ochrolechia androgyna	Pertusariales	NO, Nord-Trøndelag, 2015, H. Holien	TRH-L-16947	OLICH3684-17	MK812264
Ochrolechia androgyna	Pertusariales	NO, Oppland, 2015, R. Haugan	O-L-203919	OLICH3885-17	MK811897	atranorin, gyrophoric acid, lecanoric acid, unknown acid
Ochrolechia androgyna	Pertusariales	NO, Oppland, 2015, E. Timdal	O-L-201397	OLICH4178-17	MK812682	gyrophoric acid
Ochrolechia frigida	Pertusariales	NO, Finnmark, 2014, A. Launis	O-L-195945	OLICH1790-14	KY266952
Ochrolechia gowardii	Pertusariales	NO, Nord-Trøndelag, 2014, J.T. Klepsland	O-L-200283	OLICH3146-16	MK812101
Ochrolechia gowardii	Pertusariales	NO, Nord-Trøndelag, 2012, J.T. Klepsland	O-L-200100	OLICH3225-16	MK812175
Ochrolechia mahluensis	Pertusariales	NO, Finnmark, 2014, H. Holien	O-L-195912	OLICH1757-14	KY266861
Ochrolechia microstictoides	Pertusariales	NO, Aust-Agder, 2014, J.T. Klepsland	O-L-200753	OLICH3145-16	MK812077
Ochrolechia parella	Pertusariales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196015	OLICH1860-14	MK811779
Ochrolechia parella	Pertusariales	NO, Nordland, 2014, J.T. Klepsland	O-L-200300	OLICH3144-16	MK811835
Ochrolechia subviridis	Pertusariales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200638	OLICH2655-16	MK812495	gyrophoric acid
Ochrolechia szatalaensis	Pertusariales	NO, Troms, 2011, J.T. Klepsland	O-L-200097	OLICH2653-16	MK811817	murolic acid (complex), variolaric acid
Ochrolechia szatalaensis	Pertusariales	NO, Hordaland, 2013, B. Nordén & J.B. Jordal	O-L-199529	OLICH2654-16	MK812256
Ochrolechia szatalaensis	Pertusariales	NO, Nord-Trøndelag, 2014, J.T. Klepsland	O-L-200284	OLICH3143-16	MK811903
Ochrolechia szatalaensis	Pertusariales	NO, Finnmark, 2011, J.T. Klepsland	O-L-200096	OLICH3227-16	MK811865	murolic acid (complex), variolaric acid
Ochrolechia tartarea	Pertusariales	NO, Finnmark, 2014, H. Holien	O-L-195951	OLICH1796-14	KY266863
Ochrolechia tartarea	Pertusariales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196041	OLICH3795-17	MK812341
Ochrolechia upsaliensis	Pertusariales	NO, Finnmark, 2014, H. Holien	O-L-195967	OLICH1812-14	KY266960
Ochrolechia upsaliensis	Pertusariales	NO, Oppland, 2013, J.T. Klepsland	O-L-198354	OLICH2376-16	MK811642
Opegrapha vermicellifera	Arthoniales	NO, Hordaland, 2016, J.T. Klepsland	O-L-206757	OLICH3476-17	MK811737
Ophioparma lapponica	Umbilicariales	NO, Finnmark, 2014, T.B. Wheeler	O-L-195838	OLICH1683-14	KY266901
Ophioparma ventosa	Umbilicariales	FI, Inarin Lappi, 2014, E. Timdal	O-L-195498	OLICH1503-14	MK812137
Ophioparma ventosa	Umbilicariales	NO, Finnmark, 2014, E. Timdal	O-L-195543	OLICH1512-14	KY266864
Ophioparma ventosa	Umbilicariales	NO, Finnmark, 2014, E. Timdal	O-L-195574	OLICH1525-14	MK812332
Ophioparma ventosa	Umbilicariales	NO, Oppland, 2015, E. Timdal	O-L-201417	OLICH3307-16	MK811904
Ophioparma ventosa	Umbilicariales	NO, Troms, 2000, K. Lerfall	TROM_L_565529	OLICH3550-17	MK811740
Ophioparma ventosa	Umbilicariales	NO, Hedmark, 2013, E. Timdal	O-L-184520	OLICH701-13	MK811771
Orphniospora moriopsis	Umbilicariales	NO, Finnmark, 2014, H. Holien	O-L-195716	OLICH1588-14	KY266869
Pannaria conoplea	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196369	OLICH1928-15	MK811685
Pannaria conoplea	Peltigerales	NO, Sogn og Fjordane, 2012, T.H. Hofton & T. Høitomt	O-L-194474	OLICH2379-16	MK812100
Pannaria conoplea	Peltigerales	NO, Hedmark, 2011, R. Haugan	O-L-173219	OLICH2380-16	MK812203
Pannaria conoplea	Peltigerales	NO, Hedmark, 2011, R. Haugan	O-L-173231	OLICH2382-16	MK812521
Pannaria conoplea	Peltigerales	NO, Hedmark, 2011, R. Haugan	O-L-173220	OLICH2383-16	MK811788
Pannaria conoplea	Peltigerales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200927	OLICH2553-16	MK812604
Pannaria conoplea	Peltigerales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201274	OLICH2776-16	MK812480
Pannaria hookeri	Peltigerales	NO, Nordland, 2013, J.T. Klepsland	O-L-198018	OLICH3882-17	MK812291
Pannaria rubiginosa	Peltigerales	NO, Nordland, 2004, A. Elvebakk	TROM_L_1340509	OLICH3563-17	MK812061
Pannaria rubiginosa	Peltigerales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201270	OLICH3846-17	MK812317
Parmelia ernstiae	Lecanorales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201279	OLICH2777-16	MK567160	atranorin, lichesterinic acid, lobaric acid, protolichesterinic acid, salazinic acid
Parmelia ernstiae	Lecanorales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201277	OLICH3848-17	MK567162	atranorin, lichesterinic acid, lobaric acid, protolichesterinic acid, salazinic acid
Parmelia fraudans	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196374	OLICH1933-15	MK812511
Parmelia omphalodes	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195947	OLICH1792-14	KY266894
Parmelia saxatilis	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-195994	OLICH1839-14	MK567163	atranorin, lobaric acid, salazinic acid
Parmelia saxatilis	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184724	OLICH940-13	MK567164	atranorin, lobaric acid, salazinic acid
Parmelia serrana	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-195995	OLICH1840-14	MK567161	atranorin, lichesterinic acid, lobaric acid, protolichesterinic acid, salazinic acid
Parmelia serrana	Lecanorales	NO, Vestfold, 2016, S. Rui & E. Timdal	O-L-205007	OLICH4004-17	MK567159	atranorin, lichesterinic acid, lobaric acid, protolichesterinic acid, salazinic acid
Parmelia sulcata	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-195789	OLICH1661-14	MK811938
Parmelia sulcata	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196000	OLICH1845-14	MK812088
Parmelia sulcata	Lecanorales	NO, Oslo, 2014, M. Bendiksby et al.	O-L-196057	OLICH1873-14	MK812065
Parmelia sulcata	Lecanorales	NO, Akershus, 2014, S. Rui & E. Timdal	O-L-196331	OLICH2261-15	MK812499
Parmelia sulcata	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184694	OLICH878-13	MK812475
Parmeliella parvula	Peltigerales	NO, Nord-Trøndelag, 2013, J.T. Klepsland	O-L-197955	OLICH2107-15	MK812304
Parmeliella triptophylla	Peltigerales	NO, Telemark, 2015, E. Timdal	O-L-207999	OLICH3873-17	MK812457
Parmelina pastillifera	Lecanorales	NO, Rogaland, 2013, J.T. Klepsland	O-L-198298	OLICH2377-16	MK812633
Parmelina pastillifera	Lecanorales	NO, Hordaland, 2006, H. Fjeldstad & G. Gaarder	O-L-202999	OLICH3231-16	MK812400
Parmelina pastillifera	Lecanorales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-205003	OLICH4005-17	MK812564
Parmelina pastillifera	Lecanorales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-204989	OLICH4006-17	MK812554
Parmelina pastillifera	Lecanorales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-205000	OLICH4007-17	MK812365
Parmelina tiliacea	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-199984	OLICH1958-15	MK812323
Parmelina tiliacea	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196287	OLICH2217-15	MK812693
Parmelina tiliacea	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-198924	OLICH2477-16	MK811628
Parmelina tiliacea	Lecanorales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200873	OLICH2525-16	MK812259
Parmelina tiliacea	Lecanorales	NO, Akershus, 2015, S. Rui & E. Timdal	O-L-200818	OLICH2589-16	MK812562
Parmelina tiliacea	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201334	OLICH2718-16	MK811862
Parmelina tiliacea	Lecanorales	NO, Oslo, 2016, S. Rui & E. Timdal	O-L-203344	OLICH3314-16	MK812472
Parmelina tiliacea	Lecanorales	NO, Rogaland, 2015, T. Tønsberg & E. Nygaard	BG-L-98117	OLICH3696-17	MK812477
Parmeliopsis ambigua	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195902	OLICH1747-14	KY266965
Parmeliopsis ambigua	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195934	OLICH1779-14	KY266851
Parmeliopsis ambigua	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195990	OLICH1835-14	KY266850
Parmeliopsis ambigua	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184695	OLICH879-13	MK811727
Parmeliopsis hyperopta	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195989	OLICH1834-14	KY266853
Parmeliopsis hyperopta	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184696	OLICH880-13	MK812515
Parmotrema crinitum	Lecanorales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-197835	OLICH2109-15	MK812284
Parmotrema crinitum	Lecanorales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-197840	OLICH2110-15	MK811794
Parmotrema perlatum	Lecanorales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200600	OLICH2651-16	MK812450
Parmotrema perlatum	Lecanorales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200643	OLICH4358-17	MK812630
Pectenia cyanoloma	Peltigerales	NO, Nord-Trøndelag, 2013, H. Holien	O-L-193006	OLICH1477-14	MK811927
Pectenia cyanoloma	Peltigerales	NO, Sogn og Fjordane, 2017, O. Olsen	O-L-207808	OLICH4320-17	MK812029
Peltigera aphthosa	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195726	OLICH1598-14	KY266857
Peltigera aphthosa	Peltigerales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200017	OLICH1991-15	MK811726
Peltigera aphthosa	Peltigerales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184749	OLICH2048-15	MK812596
Peltigera aphthosa	Peltigerales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200534	OLICH2611-16	MK811760
Peltigera aphthosa	Peltigerales	NO, Telemark, 2015, E. Timdal	O-L-208000	OLICH3874-17	MK811699
Peltigera aphthosa	Peltigerales	NO, Buskerud, 2016, M. Bendiksby & E. Timdal	O-L-208028	OLICH4256-17	MK811816
Peltigera britannica	Peltigerales	NO, Nord-Trøndelag, 2015, E. Timdal et al.	O-L-201329	OLICH2689-16	MK812626
Peltigera canina	Peltigerales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200554	OLICH2610-16	MK811916
Peltigera canina	Peltigerales	NO, Østfold, 2012, K.A. Lye	O-L-206918	OLICH4134-17	MK812107
Peltigera canina	Peltigerales	NO, Østfold, 2012, K.A. Lye	O-L-206873	OLICH4135-17	MK811752
Peltigera collina	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196371	OLICH1930-15	MK812439
Peltigera collina	Peltigerales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200865	OLICH2517-16	MK811975
Peltigera collina	Peltigerales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200926	OLICH2540-16	MK812019
Peltigera collina	Peltigerales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201268	OLICH3845-17	MK811743
Peltigera degenii	Peltigerales	NO, Telemark, 2014, E. Timdal	O-L-196276	OLICH1950-15	MK812238
Peltigera degenii	Peltigerales	NO, Akershus, 2014, S. Rui & E. Timdal	O-L-196335	OLICH2273-15	MK812333
Peltigera degenii	Peltigerales	NO, Buskerud, 2016, S. Rui & E. Timdal	O-L-208031	OLICH4259-17	MK812618
Peltigera didactyla	Peltigerales	NO, Telemark, 2015, E. Timdal	O-L-208003	OLICH3877-17	MK811844
Peltigera elisabethae	Peltigerales	NO, Buskerud, 2014, E. Timdal	O-L-196071	OLICH1887-14	MK811869
Peltigera extenuata	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196343	OLICH1903-15	MK811873
Peltigera extenuata	Peltigerales	NO, Oslo, 2014, E. Timdal	O-L-200007	OLICH1981-15	MK811908
Peltigera extenuata	Peltigerales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200018	OLICH1992-15	MK811654
Peltigera horizontalis	Peltigerales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184750	OLICH2083-15	MK811925
Peltigera horizontalis	Peltigerales	NO, Hedmark, 2014, R. Haugan	O-L-198894	OLICH2391-16	MK812678
Peltigera horizontalis	Peltigerales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200862	OLICH2514-16	MK811673
Peltigera horizontalis	Peltigerales	NO, Østfold, 2016, S. Rui & E. Timdal	O-L-208021	OLICH4193-17	MK812656
Peltigera hymenina	Peltigerales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200019	OLICH1993-15	MK812274
Peltigera hymenina	Peltigerales	NO, Oslo, 2014, E. Timdal	O-L-198933	OLICH2485-16	MK811966
Peltigera hymenina	Peltigerales	NO, Vestfold, 2015, E. Timdal	O-L-198944	OLICH2495-16	MK812144
Peltigera hymenina	Peltigerales	NO, Nord-Trøndelag, 2015, E. Timdal et al.	O-L-201328	OLICH2688-16	MK811735
Peltigera lepidophora	Peltigerales	NO, Telemark, 2015, E. Timdal & S. Rui	O-L-200874	OLICH3835-17	MK812313
Peltigera lepidophora	Peltigerales	NO, Sør-Trøndelag, 2013, E. Timdal	O-L-184530	OLICH732-13	MK812625
Peltigera leucophlebia	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196342	OLICH1902-15	MK812292
Peltigera leucophlebia	Peltigerales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200020	OLICH1994-15	MK811981
Peltigera leucophlebia	Peltigerales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200929	OLICH2555-16	MK811680
Peltigera leucophlebia	Peltigerales	NO, Nord-Trøndelag, 2015, E. Timdal et al.	O-L-201315	OLICH2684-16	MK812123
Peltigera leucophlebia	Peltigerales	NO, Svalbard, 2015, S. Rui	O-L-203280	OLICH2699-16	MK811857
Peltigera leucophlebia	Peltigerales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196030	OLICH3807-17	MK812241
Peltigera leucophlebia	Peltigerales	NO, Oppland, 2015, E. Timdal	O-L-201400	OLICH4181-17	MK812102
Peltigera malacea	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195725	OLICH1597-14	KY266940
Peltigera malacea	Peltigerales	NO, Finnmark, 2014, R. Haugan	O-L-195848	OLICH1693-14	KY266836
Peltigera malacea	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196362	OLICH1921-15	MK811739
Peltigera malacea	Peltigerales	NO, Oslo, 2014, E. Timdal	O-L-200006	OLICH1980-15	MK812014
Peltigera malacea	Peltigerales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184751	OLICH2085-15	MK811998
Peltigera malacea	Peltigerales	NO, Nord-Trøndelag, 2015, E. Timdal et al.	O-L-201331	OLICH2690-16	MK811990
Peltigera membranacea	Peltigerales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-195991	OLICH1836-14	MK811987
Peltigera membranacea	Peltigerales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200024	OLICH2472-16	MK812374
Peltigera neckeri	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195753	OLICH1625-14	KY266937
Peltigera neckeri	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195932	OLICH1777-14	MK811778
Peltigera neckeri	Peltigerales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200910	OLICH2582-16	MK811968
Peltigera neocanina	Peltigerales	NO, Buskerud, 2014, R. Haugan & E. Timdal	O-L-196062	OLICH1878-14	MK812413
Peltigera neopolydactyla	Peltigerales	NO, Telemark, 2014, E. Timdal	O-L-196266	OLICH1941-15	MK812243
Peltigera neopolydactyla	Peltigerales	NO, Telemark, 2014, E. Timdal	O-L-196277	OLICH1946-15	MK812022
Peltigera neopolydactyla	Peltigerales	NO, Oslo, 2014, E. Timdal	O-L-199994	OLICH1968-15	MK812548
Peltigera neopolydactyla	Peltigerales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184753	OLICH2084-15	MK811653
Peltigera neopolydactyla	Peltigerales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200025	OLICH2473-16	MK812286
Peltigera neopolydactyla	Peltigerales	NO, Nord-Trøndelag, 2015, E. Timdal et al.	O-L-201332	OLICH2691-16	MK812172
Peltigera polydactylon	Peltigerales	NO, Telemark, 2014, E. Timdal	O-L-196264	OLICH1939-15	MK811815
Peltigera polydactylon	Peltigerales	NO, Telemark, 2014, E. Timdal	O-L-195820	OLICH2192-15	MK812112
Peltigera polydactylon	Peltigerales	NO, Buskerud, 2016, S. Rui & E. Timdal	O-L-206899	OLICH4136-17	MK812583
Peltigera praetextata	Peltigerales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184752	OLICH2044-15	MK811692
Peltigera praetextata	Peltigerales	NO, Oslo, 2014, E. Timdal	O-L-195837	OLICH2209-15	MK811856
Peltigera praetextata	Peltigerales	NO, Akershus, 2009, T. Starholm	O-L-201769	OLICH3880-17	MK812051
Peltigera praetextata	Peltigerales	NO, Østfold, 2016, S. Rui & E. Timdal	O-L-208022	OLICH4194-17	MK812578
Peltigera retifoveata	Peltigerales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200533	OLICH2605-16	MK811996
Peltigera rufescens	Peltigerales	NO, Oslo, 2014, S. Rui & E. Timdal	O-L-196072	OLICH1888-14	MK812663
Peltigera rufescens	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196341	OLICH1901-15	MK811858
Peltigera rufescens	Peltigerales	NO, Akershus, 2014, E. Timdal	O-L-199989	OLICH1963-15	MK812224
Peltigera rufescens	Peltigerales	NO, Oppland, 2015, E. Timdal	O-L-201347	OLICH2733-16	MK812696
Peltigera scabrosa	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195729	OLICH1601-14	KY266948
Peltigera scabrosa	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195765	OLICH1637-14	KY266970
Peltigera scabrosa	Peltigerales	NO, Telemark, 2015, E. Timdal	O-L-207997	OLICH3871-17	MK812044
Peltigera scabrosella	Peltigerales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201255	OLICH2763-16	MK812406
Peltigera scabrosella	Peltigerales	NO, Telemark, 2015, E. Timdal	O-L-207998	OLICH3872-17	MK812463
Peltula euploca	Lichinales	NO, Oppland, 2013, J.T. Klepsland	O-L-197859	OLICH2111-15	MK811928
Pertusaria alpina	Pertusariales	NO, Nordland, 2014, J.T. Klepsland	O-L-200312	OLICH3141-16	MK811905
Pertusaria alpina	Pertusariales	NO, Nord-Trøndelag, 2013, J.T. Klepsland	O-L-197933	OLICH4038-17	MK811785
Pertusaria aspergilla	Pertusariales	NO, Sør-Trøndelag, 2015, H. Holien	TRH-L-16775	OLICH3686-17	MK812282
Pertusaria aspergilla	Pertusariales	NO, Nordland, 2015, T. Tønsberg	BG-L-99158	OLICH3716-17	MK812384
Pertusaria aspergilla	Pertusariales	NO, Østfold, 2015, R. Haugan	O-L-204101	OLICH3929-17	MK812496	fumarprotocetraric acid
Pertusaria carneopallida	Pertusariales	NO, Nordland, 2013, J.T. Klepsland	O-L-198006	OLICH4039-17	MK812218
Pertusaria excludens	Pertusariales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-203909	OLICH3927-17	MK811670	norstictic acid
Pertusaria excludens	Pertusariales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-203523	OLICH3935-17	MK812675	norstictic acid
Pertusaria flavicans	Pertusariales	NO, Møre og Romsdal, 2015, T. Tønsberg	BG-L-99071	OLICH3777-17	MK812409
Pertusaria flavida	Pertusariales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200860	OLICH2512-16	MK811913
Pertusaria leioplaca	Pertusariales	NO, Troms, 2000, S. Werth	TROM_L_565683	OLICH3551-17	MK811922
Pertusaria pertusa	Pertusariales	NO, Vestfold, 2015, E. Timdal	O-L-198948	OLICH2502-16	MK812037
Phaeophyscia endococcina	Caliciales	NO, Finnmark, 2014, R. Haugan	O-L-198738	OLICH2400-16	MK812217
Phaeophyscia kairamoi	Caliciales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196355	OLICH1914-15	MK811724
Phaeophyscia orbicularis	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195734	OLICH1606-14	KY266882
Phaeophyscia orbicularis	Caliciales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196004	OLICH1849-14	MK811854
Phaeophyscia orbicularis	Caliciales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196291	OLICH2221-15	MK811807
Phaeophyscia orbicularis	Caliciales	NO, Akershus, 2014, S. Rui & E. Timdal	O-L-196330	OLICH2259-15	MK812383
Phaeophyscia orbicularis	Caliciales	NO, Hedmark, 2014, R. Haugan	O-L-207984	OLICH3819-17	MK812363
Phaeophyscia sciastra	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195721	OLICH1593-14	KY266837
Phaeophyscia sciastra	Caliciales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196352	OLICH1911-15	MK812372
Phlyctis agelaea	Ostropales	NO, Hordaland, 2015, J.T. Klepsland	O-L-206746	OLICH3465-17	MK812009
Phlyctis argena	Ostropales	NO, Nordland, 2015, T. Tønsberg & A. Botnen	BG-L-98987	OLICH3760-17	MK811759
Phlyctis argena	Ostropales	NO, Nordland, 2015, T. Tønsberg & A. Botnen	BG-L-99194	OLICH3761-17	MK811753
Phlyctis argena	Ostropales	NO, Buskerud, 2014, T. Tønsberg	BG-L-99532	OLICH3786-17	MK811872
Physcia adscendens	Caliciales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196005	OLICH1850-14	MK811630
Physcia adscendens	Caliciales	NO, Vestfold, 2015, E. Timdal	O-L-198947	OLICH2501-16	MK812201
Physcia aipolia	Caliciales	NO, Telemark, 2014, E. Timdal	O-L-195783	OLICH1655-14	MK812601
Physcia aipolia	Caliciales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-195992	OLICH1837-14	MK811770
Physcia aipolia	Caliciales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200890	OLICH2528-16	MK811989
Physcia aipolia	Caliciales	NO, Oslo, 2014, J.T. Klepsland	O-L-200142	OLICH2609-16	MK812087
Physcia aipolia	Caliciales	NO, Østfold, 2012, K.A. Lye	O-L-206860	OLICH4138-17	MK811719
Physcia alnophila	Caliciales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201269	OLICH2773-16	MK812315
Physcia caesia	Caliciales	NO, Telemark, 2014, E. Timdal	O-L-195792	OLICH1664-14	MK811883
Physcia caesia	Caliciales	NO, Oslo, 2014, M. Bendiksby et al.	O-L-196078	OLICH1894-14	MK812056
Physcia caesia	Caliciales	NO, Akershus, 2014, S. Rui & E. Timdal	O-L-196333	OLICH2263-15	MK812325
Physcia caesia	Caliciales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196035	OLICH3791-17	MK812524
Physcia dimidiata	Caliciales	NO, Buskerud, 2015, E. Timdal	O-L-201418	OLICH3299-16	MK812556
Physcia dubia	Caliciales	NO, Oppland, 2013, J.T. Klepsland	O-L-197852	OLICH2113-15	MK812017
Physcia dubia	Caliciales	NO, Oppland, 2015, E. Timdal	O-L-201373	OLICH2748-16	MK811700
Physcia magnussonii	Caliciales	NO, Sogn og Fjordane, 2013, J.T. Klepsland	O-L-197861	OLICH2114-15	MK811875
Physcia phaea	Caliciales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196373	OLICH1932-15	MK811892
Physcia stellaris	Caliciales	NO, Troms, 2014, E. Timdal	O-L-195769	OLICH1641-14	MK811861
Physcia stellaris	Caliciales	NO, Østfold, 2012, K.A. Lye	O-L-206903	OLICH4137-17	MK812381
Physcia tenella	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195561	OLICH1519-14	KY266867
Physcia tenella	Caliciales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196007	OLICH1852-14	MK811920
Physcia tenella	Caliciales	NO, Oslo, 2014, M. Bendiksby et al.	O-L-196079	OLICH1895-14	MK811798
Physcia tenella	Caliciales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196294	OLICH2224-15	MK811848
Physcia tenella	Caliciales	NO, Rogaland, 2013, J.T. Klepsland	O-L-198307	OLICH2357-15	MK811950
Physcia tenella	Caliciales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196034	OLICH3798-17	MK812588
Physconia distorta	Caliciales	NO, Akershus, 2014, E. Timdal	O-L-199983	OLICH1957-15	MK811745
Physconia distorta	Caliciales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196290	OLICH2220-15	MK812071
Physconia enteroxantha	Caliciales	NO, Oslo, 2014, R. Haugan et al.	O-L-196058	OLICH1874-14	MK811840
Physconia enteroxantha	Caliciales	NO, Akershus, 2014, E. Timdal	O-L-199986	OLICH1960-15	MK811812
Physconia enteroxantha	Caliciales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196293	OLICH2223-15	MK811936
Physconia enteroxantha	Caliciales	NO, Akershus, 2014, S. Rui & E. Timdal	O-L-196328	OLICH2258-15	MK812142
Physconia muscigena	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195722	OLICH1594-14	KY266849
Physconia muscigena	Caliciales	NO, Finnmark, 2014, E. Timdal	O-L-195920	OLICH1765-14	KY266925
Physconia muscigena	Caliciales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196344	OLICH1904-15	MK811781
Pilophorus dovrensis	Lecanorales	NO, Troms, 2003, E. Timdal	O-L-123083	OLICH211-13	MK812599
Placynthiella oligotropha	Trapeliales	NO, Oppland, 2012, E. Timdal	O-L-182032	OLICH1176-14	MK811853
Placynthiella oligotropha	Trapeliales	NO, Oppland, 2011, M. Bendiksby et al.	O-L-175738	OLICH1177-14	MK812378
Platismatia glauca	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195649	OLICH1561-14	MK812715
Platismatia glauca	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-195993	OLICH1838-14	MK812351
Platismatia glauca	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200882	OLICH2564-16	MK812456
Platismatia glauca	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184697	OLICH881-13	MK812210
Platismatia norvegica	Lecanorales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201267	OLICH2772-16	MK812616
Platismatia norvegica	Lecanorales	NO, Akershus, 2017, S. Rui & E. Timdal	O-L-206864	OLICH4109-17	MK812705
Pleopsidium chlorophanum	Acarosporales	NO, Troms, 2014, E. Timdal	O-L-195643	OLICH1555-14	MK811850
Pleopsidium chlorophanum	Acarosporales	NO, Finnmark, 2014, E. Timdal	O-L-195940	OLICH1785-14	KY266860
Pleopsidium chlorophanum	Acarosporales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201252	OLICH2759-16	MK812117
Pleopsidium chlorophanum	Acarosporales	NO, Sør-Trøndelag, 2017, E. Timdal	O-L-208244	OLICH4322-17	MK812354
Pleopsidium chlorophanum	Acarosporales	NO, Telemark, 2013, E. Timdal	O-L-184414	OLICH695-13	MK812213
Pleopsidium flavum	Acarosporales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200545	OLICH2624-16	MK811960
Pleopsidium flavum	Acarosporales	NO, Telemark, 2013, E. Timdal	O-L-184415	OLICH696-13	MK812038
Pleopsidium flavum	Acarosporales	NO, Hordaland, 2013, E. Timdal	O-L-184439	OLICH697-13	MK811877
Pleurosticta acetabulum	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-199985	OLICH1959-15	MK812572
Pleurosticta acetabulum	Lecanorales	NO, Vestfold, 2015, E. Timdal	O-L-198935	OLICH2488-16	MK812152
Pleurosticta acetabulum	Lecanorales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200869	OLICH2521-16	MK811775
Pleurosticta acetabulum	Lecanorales	NO, Rogaland, 2015, T. Tønsberg & E. Nygaard	BG-L-98170	OLICH3700-17	MK811799
Pleurosticta acetabulum	Lecanorales	NO, Østfold, 2002, B. P. Løfall	O-L-108808	OLICH4128-17	MK812479
Polycauliona candelaria	Teloschistales	NO, Akershus, 2014, E. Timdal	O-L-199981	OLICH1955-15	MK811915
Polycauliona candelaria	Teloschistales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196326	OLICH2256-15	MK812066
Polycauliona candelaria	Teloschistales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196340	OLICH2278-15	MK811959
Polychidium muscicola	Peltigerales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201293	OLICH2795-16	MK812283
Protoblastenia rupestris	Lecanorales	NO, Finnmark, 2014, A. Millanes	O-L-195859	OLICH1704-14	KY266981
Protomicarea limosa	Lecanorales	NO, Hedmark, 2012, R. Haugan	O-L-182289	OLICH1084-14	MK811666
Protomicarea limosa	Lecanorales	NO, Hedmark, 2004, R. Haugan	O-L-131761	OLICH1085-14	MK812532
Protomicarea limosa	Lecanorales	NO, Nord-Trøndelag, 2009, J.T. Klepsland	O-L-165102	OLICH1086-14	MK812181
Protomicarea limosa	Lecanorales	NO, Oppland, 2012, E. Timdal	O-L-182031	OLICH1140-14	MK812033
Protomicarea limosa	Lecanorales	NO, Nordland, 2010, T. Tønsberg	BG-L-92492	OLICH601-13	MK812586
Protomicarea limosa	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184281	OLICH821-13	MK812176
Protopannaria pezizoides	Peltigerales	NO, Finnmark, 2014, A. Guillermo	O-L-195697	OLICH1569-14	KY266840
Protopannaria pezizoides	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195890	OLICH1735-14	KY266909
Protopannaria pezizoides	Peltigerales	NO, Finnmark, 2014, H. Holien	O-L-195970	OLICH1815-14	KY266886
Protopannaria pezizoides	Peltigerales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196347	OLICH1907-15	MK811767
Protopannaria pezizoides	Peltigerales	NO, Nord-Trøndelag, 2013, J.T. Klepsland	O-L-197950	OLICH2115-15	MK812467
Protopannaria pezizoides	Peltigerales	NO, Svalbard, 2015, S. Rui	O-L-203297	OLICH2706-16	MK811974
Protoparmelia badia	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195568	OLICH1522-14	KY266939
Protoparmelia badia	Lecanorales	NO, Finnmark, 2014, A. Launis	O-L-195839	OLICH1684-14	KY266945
Pseudephebe minuscula	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196378	OLICH1937-15	MK811997
Pseudephebe minuscula	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184305	OLICH736-13	MK812067
Pseudephebe pubescens	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195619	OLICH1548-14	MK812357
Pseudephebe pubescens	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195849	OLICH1694-14	KY266938
Pseudephebe pubescens	Lecanorales	NO, Finnmark, 2014, R. Haugan	O-L-195986	OLICH1831-14	KY266914
Pseudephebe pubescens	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184396	OLICH694-13	MK812150
Pseudephebe pubescens	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184306	OLICH735-13	MK811888
Pseudephebe pubescens	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184725	OLICH941-13	MK812652
Pseudevernia furfuracea	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-195999	OLICH1844-14	MK812319
Pseudevernia furfuracea	Lecanorales	NO, Hedmark, 2016, E. Timdal	O-L-201522	OLICH3984-17	MK812161
Pseudevernia furfuracea	Lecanorales	NO, Aust-Agder, 2016, S. Rui & E. Timdal	O-L-204977	OLICH4008-17	MK812508
Pseudevernia furfuracea	Lecanorales	NO, Telemark, 2016, S. Rui & E. Timdal	O-L-205006	OLICH4009-17	MK811934
Pseudevernia furfuracea	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184698	OLICH882-13	MK812628
Pseudocyphellaria citrina	Peltigerales	NO, Nord-Trøndelag, 2013, J.T. Klepsland	O-L-197920	OLICH2116-15	MK812391
Pseudocyphellaria citrina	Peltigerales	NO, Sør-Trøndelag, 2013, J.T. Klepsland	O-L-198050	OLICH2117-15	MK811890
Pseudocyphellaria citrina	Peltigerales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201266	OLICH2771-16	MK812527
Pseudocyphellaria norvegica	Peltigerales	NO, Hordaland, 2006, H. Fjeldstad	O-L-203000	OLICH3242-16	MK812692
Psilolechia clavulifera	Lecanorales	NO, Akershus, 2009, J.T. Klepsland	O-L-164653	OLICH1081-14	MK811831
Psilolechia leprosa	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195611	OLICH1544-14	MK812516
Psilolechia leprosa	Lecanorales	NO, Aust-Agder, 2015, J.T. Klepsland	O-L-204517	OLICH4229-17	MK811951
Psilolechia leprosa	Lecanorales	NO, Aust-Agder, 2015, J.T. Klepsland	O-L-204519	OLICH4251-17	MK812436
Psilolechia lucida	Lecanorales	SE, Närke, 2013, M. Bendiksby et al.	O-L-184487	OLICH665-13	MK811768
Psora decipiens	Lecanorales	NO, Oppland, 2013, J.T. Klepsland	O-L-198348	OLICH2358-15	MK812030
Psora decipiens	Lecanorales	NO, Troms, 2014, J.T. Klepsland	O-L-200362	OLICH2646-16	MK811804
Psora globifera	Lecanorales	NO, Nord-Trøndelag, 2011, J.T. Klepsland	O-L-177145	OLICH1319-14	KU873929	no lichen substances
Psora globifera	Lecanorales	NO, Sør-Trøndelag, 2012, R. Hjelmstad	O-L-184143	OLICH1320-14	KU873932	no lichen substances
Psora globifera	Lecanorales	NO, Buskerud, 2014, J.T. Klepsland	O-L-200597	OLICH2645-16	MK812110
Psora globifera	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183774	OLICH782-13	KU873928	no lichen substances
Psora globifera	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184327	OLICH820-13	KU873930	no lichen substances
Psora rubiformis	Lecanorales	NO, Oppland, 2012, H. Bratli & R. Haugan	O-L-183466	OLICH1082-14	MK811688
Psora rubiformis	Lecanorales	NO, Nord-Trøndelag, 2011, J.T. Klepsland	O-L-177142	OLICH1321-14	MK812240
Psora rubiformis	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195525	OLICH1508-14	KY266957
Psora rubiformis	Lecanorales	NO, Hordaland, 2013, E. Timdal	O-L-184422	OLICH648-13	MK811706
Psora rubiformis	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184359	OLICH649-13	MK812000
Psora vallesiaca	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184392	OLICH650-13	KU873926	norstictic acid
Psora vallesiaca	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183778	OLICH780-13	KU873927	norstictic acid
Psora vallesiaca	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183760	OLICH781-13	KU873931	norstictic acid
Psorinia conglomerata	Lecanorales	NO, Oppland, 2012, H. Bratli & R. Haugan	O-L-182192	OLICH1061-14	MK812001	stictic acid
Psoroma cinnamomeum	Peltigerales	NO, Sør-Trøndelag, 2013, E. Timdal	O-L-184538	OLICH702-13	MK811978
Psoroma hypnorum	Peltigerales	NO, Finnmark, 2014, E. Timdal	O-L-195857	OLICH1702-14	KY266971
Psoroma hypnorum	Peltigerales	NO, Sør-Trøndelag, 2017, E. Timdal	O-L-208202	OLICH4325-17	MK811800
Psoroma hypnorum	Peltigerales	NO, Sør-Trøndelag, 2013, E. Timdal	O-L-184537	OLICH703-13	MK811764
Psoronactis dilleniana	Arthoniales	NO, Nord-Trøndelag, 2015, H. Holien	TRH-L-16932	OLICH3649-17	MK812257
Punctelia stictica	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201333	OLICH2698-16	MK812448
Punctelia subrudecta	Lecanorales	NO, Rogaland, 2015, T. Tønsberg	BG-L-98118	OLICH3697-17	MK812035
Punctelia subrudecta	Lecanorales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200662	OLICH4106-17	MK812237
Pycnothelia papillaria	Lecanorales	NO, Aust-Agder, 2006, J.T. Klepsland	O-L-151129	OLICH1364-14	MK811765	atranorin, lichesterinic acid
Pycnothelia papillaria	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-200001	OLICH1975-15	MK812272
Pycnothelia papillaria	Lecanorales	NO, Nord-Trøndelag, 2015, M. Bendiksby et al.	O-L-201299	OLICH2797-16	MK811947
Pyrenula laevigata	Pyrenulales	NO, Hordaland, 2016, J.T. Klepsland	O-L-206758	OLICH3477-17	MK812685
Pyrenula laevigata	Pyrenulales	NO, Møre og Romsdal, 2016, J.T. Klepsland	O-L-206773	OLICH3492-17	MK812185
Pyrenula occidentalis	Pyrenulales	NO, Møre og Romsdal, 2016, J.T. Klepsland	O-L-206777	OLICH3496-17	MK811633
Racodium rupestre	Capnodiales	NO, Rogaland, 2013, J.T. Klepsland	O-L-198464	OLICH3945-17	MK812095
Ramalina capitata	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200012	OLICH1986-15	MK811917
Ramalina capitata	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-195831	OLICH2203-15	MK811626
Ramalina capitata	Lecanorales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200896	OLICH2549-16	MK812528
Ramalina capitata	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184378	OLICH693-13	MK812398
Ramalina cuspidata	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196016	OLICH1861-14	MK812006	usnic acid
Ramalina cuspidata	Lecanorales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-197829	OLICH2118-15	MK811797
Ramalina farinacea	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196002	OLICH1847-14	MK812207	protocetraric acid, usnic acid
Ramalina farinacea	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-199987	OLICH1961-15	MK812169	protocetraric acid, usnic acid
Ramalina farinacea	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196286	OLICH2216-15	MK812449	protocetraric acid , usnic acid
Ramalina fastigiata	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196289	OLICH2219-15	MK811906
Ramalina fastigiata	Lecanorales	NO, Vestfold, 2014, J.T. Klepsland	O-L-200570	OLICH2644-16	MK811841
Ramalina fastigiata	Lecanorales	NO, Østfold, 2016, S. Rui & E. Timdal	O-L-208026	OLICH4197-17	MK812196
Ramalina fraxinea	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196288	OLICH2218-15	MK812590
Ramalina fraxinea	Lecanorales	NO, Vestfold, 2014, J.T. Klepsland	O-L-200571	OLICH4105-17	MK812489
Ramalina fraxinea	Lecanorales	NO, Akershus, 2012, K.A. Lye	O-L-206861	OLICH4140-17	MK811900
Ramalina fraxinea	Lecanorales	NO, Østfold, 2016, S. Rui & E. Timdal	O-L-208025	OLICH4196-17	MK812162
Ramalina pollinaria	Lecanorales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196372	OLICH1931-15	MK812166
Ramalina pollinaria	Lecanorales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200850	OLICH2516-16	MK812114
Ramalina pollinaria	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200901	OLICH2552-16	MK811866
Ramalina polymorpha	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195737	OLICH1609-14	KY266885
Ramalina polymorpha	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195928	OLICH1773-14	MK811690
Ramalina polymorpha	Lecanorales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-200011	OLICH1985-15	MK811661
Ramalina polymorpha	Lecanorales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200898	OLICH2551-16	MK812673
Ramalina siliquosa	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196010	OLICH1855-14	MK811720	protocetraric acid (minor), salazinic acid (major), usnic acid (trace)
Ramalina subfarinacea	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196284	OLICH2214-15	MK812104	norstictic acid (major), usnic acid (major)
Ramalina subfarinacea	Lecanorales	NO, Nordland, 2014, J.T. Klepsland	O-L-200339	OLICH2643-16	MK812227	norstictic acid (major), usnic acid (major)
Ramboldia elabens	Lecanorales	NO, Sør-Trøndelag, 2007, E. Timdal	O-L-158501	OLICH411-13	MK812637
Ramboldia elabens	Lecanorales	NO, Hedmark, 2008, R. Haugan	O-L-160473	OLICH412-13	MK812132
Ramboldia elabens	Lecanorales	NO, Oppland, 2004, T.H. Hofton	O-L-136707	OLICH413-13	MK812702
Ramboldia elabens	Lecanorales	NO, Nord-Trøndelag, 2006, J.T. Klepsland	O-L-183511	OLICH472-13	MK811668
Ramboldia elabens	Lecanorales	NO, Finnmark, 2011, E. Timdal	O-L-170692	OLICH473-13	MK811870
Ramboldia elabens	Lecanorales	NO, Nordland, 2011, T. Tønsberg	BG-L-95299	OLICH580-13	MK812420
Rhizocarpon badioatrum	Rhizocarpales	NO, Oppland, 2010, E. Timdal	O-L-163824	OLICH002-11	KU687450	no lichen substances
Rhizocarpon badioatrum	Rhizocarpales	NO, Buskerud, 2010, E. Timdal	O-L-163839	OLICH003-11	KU687453	no lichen substances
Rhizocarpon bolanderi	Rhizocarpales	NO, Aust-Agder, 2010, E. Timdal	O-L-163373	OLICH023-11	KU687451	norstictic acid (trace), stictic acid (major)
Rhizocarpon chioneum	Rhizocarpales	NO, Finnmark, 2014, E. Timdal	O-L-195535	OLICH1510-14	KY266951
Rhizocarpon copelandii	Rhizocarpales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184329	OLICH766-13	KU687447
Rhizocarpon copelandii	Rhizocarpales	NO, Hedmark, 2013, E. Timdal	O-L-184521	OLICH790-13	KU687456	norstictic acid
Rhizocarpon copelandii	Rhizocarpales	NO, Oppland, 2012, S. Rui & E. Timdal	O-L-180002	OLICH837-13	KU687455	norstictic acid
Rhizocarpon geminatum	Rhizocarpales	NO, Finnmark, 2014, E. Timdal	O-L-195564	OLICH1520-14	KY266908
Rhizocarpon jemtlandicum	Rhizocarpales	NO, Telemark, 2010, E. Timdal	O-L-163364	OLICH015-11	KU687445	norstictic acid (trace), stictic acid (major)
Rhizocarpon jemtlandicum	Rhizocarpales	NO, Nordland, 2010, E. Timdal	O-L-163756	OLICH019-11	KU687446	norstictic acid (minor), stictic acid (major)
Rhizocarpon jemtlandicum	Rhizocarpales	NO, Finnmark, 2011, E. Timdal	O-L-170452	OLICH803-13	KU687460	stictic acid
Rhizocarpon leptolepis	Rhizocarpales	FI, Inarin Lappi, 2014, E. Timdal	O-L-195500	OLICH1504-14	KU687449
Rhizocarpon pycnocarpoides	Rhizocarpales	NO, Nordland, 2012, R. Haugan	O-L-183808	OLICH511-13	KT800004	no lichen substances
Rhizocarpon pycnocarpoides	Rhizocarpales	NO, Nord-Trøndelag, 2012, R. Haugan	O-L-183810	OLICH512-13	KT800005	no lichen substances
Rhizocarpon pycnocarpoides	Rhizocarpales	NO, Oppland, 2012, S. Rui & E. Timdal	O-L-179903	OLICH513-13	KT800002
Rhizocarpon pycnocarpoides	Rhizocarpales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184267	OLICH531-13	KT800003	no lichen substances
Rhizocarpon rittokense	Rhizocarpales	NO, Finnmark, 2014, M. Westberg	O-L-195695	OLICH1567-14	KU687458
Rhizocarpon rittokense	Rhizocarpales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184293	OLICH542-13	KU687459
Rhizocarpon santessonii	Rhizocarpales	NO, Oppland, 2011, R. Haugan & E. Timdal	O-L-169805	OLICH432-13	KU687454
Rhizocarpon subgeminatum	Rhizocarpales	NO, Hedmark, 2010, R. Haugan	O-L-166501	OLICH033-11	KU687457	barbatic acid
Rhizocarpon subgeminatum	Rhizocarpales	NO, Sør-Trøndelag, 2010, R. Haugan	O-L-165477	OLICH034-11	KU687452	barbatic acid
Rhizocarpon suomiense	Rhizocarpales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184330	OLICH769-13	KU687448
Rhizoplaca melanophthalma	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195929	OLICH1774-14	MK812478	unknown substances, usnic acid
Rhizoplaca melanophthalma	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201366	OLICH2743-16	MK811669
Rinodina balanina	Caliciales	NO, Finnmark, 2014, T.B. Wheeler	O-L-195705	OLICH1577-14	KY266842
Rinodina disjuncta	Caliciales	NO, Nord-Trøndelag, 2014, H. Holien	TRH-L-15387	OLICH3675-17	MK812529
Rinodina disjuncta	Caliciales	NO, Nord-Trøndelag, 2011, H. Holien	TRH-L-14116	OLICH3676-17	MK812438
Rinodina mniaraea	Caliciales	NO, Svalbard, 2001, A. Elvebakk	TROM_L_565871	OLICH3552-17	MK812098
Rinodina roboris	Caliciales	NO, Rogaland, 2016, J.T. Klepsland	O-L-206765	OLICH3484-17	MK811851
Romjularia lurida	Lecideales	NO, Akershus, 2010, E. Timdal	O-L-169118	OLICH150-11	MK811769
Ropalospora lugubris	Umbilicariales	NO, Finnmark, 2014, E. Timdal	O-L-195537	OLICH1511-14	KY266893
Ropalospora lugubris	Umbilicariales	NO, Nordland, 2015, T. Tønsberg & A. Botnen	BG-L-98792	OLICH3732-17	MK812617
Ropalospora lugubris	Umbilicariales	NO, Finnmark, 2011, E. Timdal	O-L-170519	OLICH3969-17	MK812431
Ropalospora lugubris	Umbilicariales	NO, Sør-Trøndelag, 2011, R. Haugan	O-L-168439	OLICH3970-17	MK812670
Ropalospora lugubris	Umbilicariales	NO, Sogn og Fjordane, 2011, R. Haugan	O-L-173455	OLICH3971-17	MK812610
Rusavskia elegans	Teloschistales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196351	OLICH1910-15	MK812165
Rusavskia elegans	Teloschistales	NO, Oppland, 2013, J.T. Klepsland	O-L-200120	OLICH2630-16	MK812212
Rusavskia sorediata	Teloschistales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196353	OLICH1912-15	MK811744
Sagiolechia protuberans	Ostropales	NO, Finnmark, 2014, M. Westberg	O-L-195714	OLICH1586-14	KY266966
Sagiolechia protuberans	Ostropales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201297	OLICH3853-17	MK811691
Sclerophora peronella	Coniocybales	SE, Södermanland, 2015, S. Kistenich & E. Timdal	O-L-200846	OLICH2432-16	MK812193
Scoliciosporum umbrinum	Lecanorales	NO, Oslo, 2014, J.T. Klepsland	O-L-200123	OLICH3322-16	MK812255
Scoliciosporum umbrinum	Lecanorales	NO, Akershus, 2012, E. Timdal	O-L-177770	OLICH822-13	MK812271
Siphula ceratites	Pertusariales	NO, Finnmark, 2014, E. Timdal	O-L-195919	OLICH1764-14	KY266949
Siphula ceratites	Pertusariales	NO, Nord-Trøndelag, 2015, E. Timdal et al.	O-L-201291	OLICH2679-16	MK812106
Siphula ceratites	Pertusariales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196042	OLICH3790-17	MK812034
Solorina saccata	Peltigerales	NO, Buskerud, 2014, E. Timdal	O-L-203070	OLICH3286-16	MK812538
Sphaerophorus fragilis	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195650	OLICH1562-14	MK812306
Sphaerophorus fragilis	Lecanorales	NO, Hordaland, 2014, E. Timdal	O-L-195800	OLICH1672-14	MK811867
Sphaerophorus fragilis	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195977	OLICH1822-14	KY266958
Sphaerophorus fragilis	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196043	OLICH3799-17	MK812393
Sphaerophorus globosus	Lecanorales	NO, Troms, 2014, E. Timdal	O-L-195651	OLICH1563-14	MK811784	sphaerophorin
Sphaerophorus globosus	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195855	OLICH1700-14	KY266841	sphaerophorin
Sphaerophorus globosus	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195918	OLICH1763-14	KY266859	sphaerophorin, squamatic acid
Sphaerophorus globosus	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195949	OLICH1794-14	KY266898
Sphaerophorus globosus	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-199995	OLICH1969-15	MK812694	sphaerophorin, thamnolic acid, unknowns
Sphaerophorus globosus	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200918	OLICH2578-16	MK812575	sphaerophorin
Sphaerophorus globosus	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196029	OLICH3793-17	MK811761	sphaerophorin, squamatic acid
Sphaerophorus globosus	Lecanorales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201265	OLICH3844-17	MK812327	sphaerophorin, thamnolic acid
Staurolemma omphalarioides	Peltigerales	NO, Nordland, 2013, J.T. Klepsland	O-L-197994	OLICH2467-16	MK811827
Staurolemma omphalarioides	Peltigerales	NO, Nordland, 2014, J.T. Klepsland	O-L-200306	OLICH2640-16	MK811929
Stereocaulon alpinum	Lecanorales	NO, Telemark, 2006, E. Timdal	O-L-149404	OLICH925-13	MK812253	atranorin, lobaric acid
Stereocaulon capitellatum	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201401	OLICH4182-17	MK812320	anziaic acid, atranorin, miriquidic acid, perlatolic acid
Stereocaulon coniophyllum	Lecanorales	NO, Sogn og Fjordane, 2010, S. Vatne	O-L-169267	OLICH713-13	MK811930	lobaric acid, atranorin
Stereocaulon cumulatum	Lecanorales	NO, Finnmark, 2014, M. Svensson	O-L-195924	OLICH1769-14	KY266904
Stereocaulon cumulatum	Lecanorales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200899	OLICH2529-16	MK812171
Stereocaulon cumulatum	Lecanorales	NO, Hedmark, 2013, E. Timdal	O-L-184515	OLICH674-13	MK812474
Stereocaulon cumulatum	Lecanorales	NO, Oppland, 2012, E. Timdal	O-L-182037	OLICH711-13	MK812671
Stereocaulon grande	Lecanorales	NO, Troms, 2003, E. Timdal	O-L-122991	OLICH917-13	MK812157	atranorin, bourgeanic acid, lobaric acid
Stereocaulon pileatum	Lecanorales	NO, Aust-Agder, 2016, S. Rui & E. Timdal	O-L-204980	OLICH4012-17	MK812053
Stereocaulon pileatum	Lecanorales	NO, Østfold, 2002, B.P. Løfall & A. Often	O-L-113318	OLICH908-13	MK812510	atranorin, lobaric acid
Stereocaulon pileatum	Lecanorales	NO, Oslo, 2011, H. Bratli & R. Haugan	O-L-174391	OLICH909-13	MK811864	atranorin, lobaric acid
Stereocaulon rivulorum	Lecanorales	NO, Troms, 2003, E. Timdal	O-L-127342	OLICH899-13	MK812658	anziaic acid, atranorin, perlatolic acid
Stereocaulon rivulorum	Lecanorales	NO, Finnmark, 2003, E. Timdal	O-L-123189	OLICH901-13	MK811662	anziaic acid, atranorin, perlatolic acid
Stereocaulon symphycheilum	Lecanorales	NO, Oppland, 2012, H. Bratli	O-L-183927	OLICH706-13	MK811941	lobaric acid
Stereocaulon symphycheilum	Lecanorales	NO, Buskerud, 2012, S. Rui & E. Timdal	O-L-184640	OLICH730-13	MK812328	atranorin, lobaric acid
Stereocaulon symphycheilum	Lecanorales	NO, Sør-Trøndelag, 2009, E. Timdal	O-L-158545	OLICH895-13	MK812189	atranorin, lobaric acid
Sticta fuliginoides	Peltigerales	NO, Rogaland, 2012, B. Nordén & J.B. Jordal	O-L-199414	OLICH2638-16	MK811703
Sticta sylvatica	Peltigerales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200925	OLICH2539-16	MK811909
Tephromela atra	Lecanorales	NO, Finnmark, 2014, A. Millanes	O-L-195861	OLICH1706-14	KY266944
Tephromela atra	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196054	OLICH3815-17	MK812606
Tephromela atra	Lecanorales	NO, Telemark, 2009, R. Haugan & S. Reiso	O-L-160434	OLICH399-13	MK812380
Tephromela grumosa	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-190787	OLICH2594-16	MK812127	atranorin, lichesterinic acid
Tetramelas chloroleucus	Caliciales	NO, Sør-Trøndelag, 2014, H. Holien	TRH-L-15560	OLICH3601-17	MK811644
Tetramelas chloroleucus	Caliciales	NO, Nordland, 2013, J.T. Klepsland	O-L-198004	OLICH4032-17	MK812300
Thamnolia vermicularis	Pertusariales	NO, Finnmark, 2014, E. Timdal	O-L-195710	OLICH1582-14	KY266979
Thamnolia vermicularis	Pertusariales	NO, Troms, 2014, E. Timdal	O-L-195773	OLICH1645-14	MK812339
Thamnolia vermicularis	Pertusariales	NO, Finnmark, 2014, H. Holien	O-L-195948	OLICH1793-14	KY266969
Thamnolia vermicularis	Pertusariales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196363	OLICH1922-15	MK811783
Thamnolia vermicularis	Pertusariales	NO, Buskerud, 2015, E. Timdal & S. Rui	O-L-200905	OLICH3840-17	MK811627
Thelotrema lepadinum	Ostropales	NO, Hordaland, 2013, B. Nordén & J.B. Jordal	O-L-199496	OLICH3321-16	MK812434
Thelotrema macrosporum	Ostropales	NO, Hordaland, 2013, B. Nordén & J.B. Jordal	O-L-199493	OLICH3320-16	MK812111
Tholurna dissimilis	Caliciales	NO, Buskerud, 2014, E. Timdal	O-L-195822	OLICH2194-15	MK812234
Tholurna dissimilis	Caliciales	NO, Nord-Trøndelag, 2015, H. Holien	TRH-L-16038	OLICH2760-16	MK811683
Timdalia intricata	Acarosporales	NO, Telemark, 2013, E. Timdal	O-L-194731	OLICH3244-16	MK812713
Timdalia intricata	Acarosporales	NO, Oppland, 2015, E. Timdal	O-L-201379	OLICH4168-17	MK812246
Timdalia intricata	Acarosporales	NO, Hedmark, 2017, E. Timdal	O-L-207764	OLICH4349-17	MK811937	psoromic acid
Timdalia intricata	Acarosporales	SE, Jämtland, 2017, E. Timdal	O-L-208174	OLICH4350-17	MK811871	psoromic acid
Timdalia intricata	Acarosporales	SE, Härjedalen, 2017, E. Timdal	O-L-208190	OLICH4351-17	MK812454	psoromic acid
Toensbergia leucococca	Rhizocarpales	NO, Nordland, 2012, T. Tønsberg	BG-L-94650	OLICH592-13	MK812615
Toninia alutacea	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195931	OLICH1776-14	MG838194
Toninia alutacea	Lecanorales	NO, Buskerud, 2014, J.T. Klepsland	O-L-200591	OLICH2636-16	MG838164
Toninia alutacea	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184350	OLICH551-13	MG838166
Toninia alutacea	Lecanorales	NO, Troms, 2011, J.T. Klepsland	O-L-177188	OLICH561-13	MG838197
Toninia alutacea	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183755	OLICH783-13	MG838156
Toninia aromatica	Lecanorales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200557	OLICH3317-16	MG838189
Toninia aromatica	Lecanorales	NO, Troms, 2009, J.T. Klepsland	O-L-167368	OLICH560-13	MG838184
Toninia aromatica	Lecanorales	NO, Nordland, 2011, R. Haugan	O-L-174042	OLICH614-13	MG838161
Toninia candida	Lecanorales	NO, Buskerud, 2014, E. Timdal	O-L-196069	OLICH1885-14	MG838163
Toninia candida	Lecanorales	NO, Buskerud, 2011, K.A. Lye & T. Høitomt	O-L-177883	OLICH557-13	MG838185
Toninia coelestina	Lecanorales	NO, Nordland, 2010, J.T. Klepsland	O-L-175215	OLICH285-13	MG838190
Toninia nordlandica	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195601	OLICH1537-14	MG838198
Toninia nordlandica	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184351	OLICH552-13	MG838159
Toninia opuntioides	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195603	OLICH1539-14	MG838192
Toninia opuntioides	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183630	OLICH559-13	MG838168	triterpenoid (minor)
Toninia opuntioides	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184387	OLICH635-13	MG838172	triterpenoid (possible trace)
Toninia opuntioides	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183757	OLICH784-13	MG838158
Toninia rosulata	Lecanorales	NO, Nord-Trøndelag, 2011, J.T. Klepsland	O-L-177141	OLICH1323-14	MG838195
Toninia sedifolia	Lecanorales	NO, Finnmark, 2014, E. Timdal	O-L-195599	OLICH1536-14	MG838193
Toninia sedifolia	Lecanorales	NO, Aust-Agder, 2010, J.T. Klepsland	O-L-183585	OLICH558-13	MG838201	no lichen substances
Toninia sedifolia	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184381	OLICH637-13	MG838196
Toninia sedifolia	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184332	OLICH817-13	MG838162
Toninia squalida	Lecanorales	NO, Oppland, 2011, R. Haugan & E. Timdal	O-L-169774	OLICH553-13	MG838183
Toninia squalida	Lecanorales	NO, Oppland, 2012, H. Bratli & R. Haugan	O-L-182301	OLICH554-13	MG838182
Toninia squalida	Lecanorales	NO, Hordaland, 2013, E. Timdal	O-L-184426	OLICH638-13	MG838181
Toninia squalida	Lecanorales	NO, Sør-Trøndelag, 2013, E. Timdal	O-L-184539	OLICH639-13	MG838200
Toninia taurica	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184333	OLICH546-13	MG838174
Toninia verrucarioides	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184389	OLICH641-13	MG838167
Toninia verrucarioides	Lecanorales	NO, Oppland, 2011, J.T. Klepsland	O-L-183717	OLICH850-13	MG838179
Trapelia coarctata	Trapeliales	NO, Oslo, 2014, J.T. Klepsland	O-L-200174	OLICH3316-16	MK812584
Trapelia coarctata	Trapeliales	NO, Hedmark, 2012, R. Haugan et al.	O-L-182063	OLICH562-13	MK812526
Trapelia coarctata	Trapeliales	NO, Telemark, 2012, E. Timdal	O-L-179924	OLICH563-13	MK812177
Trapelia corticola	Trapeliales	NO, Nord-Trøndelag, 2011, H. Holien	TRH-L-14083	OLICH993-13	MK811911
Trapelia placodioides	Trapeliales	NO, Nord-Trøndelag, 2012, R. Haugan	O-L-183857	OLICH1324-14	MK812664
Trapelia placodioides	Trapeliales	NO, Telemark, 1996, H. Bratli & E. Timdal	O-L-20869	OLICH3976-17	MK812430
Trapelia placodioides	Trapeliales	NO, Hedmark, 2015, R. Haugan	O-L-204154	OLICH4366-17	MK812690	gyrophoric acid, lecanoric acid
Trapelia placodioides	Trapeliales	NO, Oslo, 2014, J.T. Klepsland	O-L-200245	OLICH4453-17	MK812050
Trapelia placodioides	Trapeliales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184337	OLICH548-13	MK812359
Trapelia placodioides	Trapeliales	NO, Oppland, 2012, S. Rui & E. Timdal	O-L-179918	OLICH569-13	MK812043
Trapeliopsis flexuosa	Trapeliales	NO, Nord-Trøndelag, 2013, H. Holien	TRH-L-15151	OLICH994-13	MK811801
Trapeliopsis granulosa	Trapeliales	NO, Sør-Trøndelag, 2007, E. Timdal	O-L-158481	OLICH565-13	MK812609
Trapeliopsis granulosa	Trapeliales	NO, Oslo, 2011, R. Haugan	O-L-177342	OLICH566-13	MK811943
Trapeliopsis granulosa	Trapeliales	NO, Troms, 2011, J.T. Klepsland	O-L-177175	OLICH567-13	MK812708
Trapeliopsis granulosa	Trapeliales	NO, Finnmark, 2011, E. Timdal	O-L-170698	OLICH568-13	MK812198
Trapeliopsis granulosa	Trapeliales	NO, Nord-Trøndelag, 2013, H. Holien	TRH-L-15150	OLICH995-13	MK812377
Trapeliopsis pseudogranulosa	Trapeliales	NO, Telemark, 2015, E. Timdal	O-L-208001	OLICH3875-17	MK812568
Trapeliopsis pseudogranulosa	Trapeliales	NO, Rogaland, 2011, J.T. Klepsland	O-L-183692	OLICH852-13	MK812270
Tremolecia atrata	Hymeneliales	NO, Sør-Trøndelag, 2012, M. Bendiksby et al.	O-L-179677	OLICH4017-17	MK812298
Tremolecia atrata	Hymeneliales	NO, Finnmark, 2011, E. Timdal	O-L-170578	OLICH606-13	MK811647
Tremolecia atrata	Hymeneliales	NO, Finnmark, 2011, E. Timdal	O-L-170776	OLICH607-13	MK812097
Tuckermanopsis chlorophylla	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196001	OLICH1846-14	MK811914
Tuckermanopsis chlorophylla	Lecanorales	NO, Akershus, 2014, S. Rui & E. Timdal	O-L-196329	OLICH2260-15	MK812387
Umbilicaria arctica	Umbilicariales	NO, Finnmark, 2014, E. Timdal	O-L-195898	OLICH1743-14	KY266911
Umbilicaria arctica	Umbilicariales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200892	OLICH2545-16	MK812428
Umbilicaria arctica	Umbilicariales	NO, Aust-Agder, 2016, S. Rui & E. Timdal	O-L-204983	OLICH4016-17	MK812268
Umbilicaria cinereorufescens	Umbilicariales	NO, Buskerud, 2014, E. Timdal	O-L-195830	OLICH2202-15	MK811655
Umbilicaria cinereorufescens	Umbilicariales	NO, Oppland, 2015, E. Timdal	O-L-201349	OLICH2734-16	MK812545
Umbilicaria crustulosa	Umbilicariales	NO, Hordaland, 2014, E. Timdal	O-L-195804	OLICH1676-14	MK811776
Umbilicaria crustulosa	Umbilicariales	NO, Buskerud, 2015, S. Rui & E. Timdal	O-L-200941	OLICH2662-16	MK812060
Umbilicaria cylindrica	Umbilicariales	NO, Hordaland, 2014, E. Timdal	O-L-195808	OLICH1680-14	MK811748
Umbilicaria cylindrica	Umbilicariales	NO, Finnmark, 2014, T. Tønsberg	O-L-195872	OLICH1717-14	KY266907
Umbilicaria cylindrica	Umbilicariales	NO, Finnmark, 2014, H. Holien	O-L-195957	OLICH1802-14	KY266835
Umbilicaria cylindrica	Umbilicariales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196354	OLICH1913-15	MK812072
Umbilicaria cylindrica	Umbilicariales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184711	OLICH929-13	MK812153
Umbilicaria decussata	Umbilicariales	NO, Sør-Trøndelag, 2015, M. Stapnes Dahl et al.	O-L-201307	OLICH2692-16	MK812293
Umbilicaria decussata	Umbilicariales	NO, Oppland, 2015, E. Timdal	O-L-201367	OLICH2744-16	MK812686
Umbilicaria dendrophora	Umbilicariales	NO, Sør-Trøndelag, 2013, E. Timdal	O-L-184529	OLICH731-13	MK812276
Umbilicaria deusta	Umbilicariales	NO, Finnmark, 2014, E. Timdal	O-L-195766	OLICH1638-14	KY266862
Umbilicaria deusta	Umbilicariales	NO, Telemark, 2014, E. Timdal	O-L-195790	OLICH1662-14	MK812011
Umbilicaria deusta	Umbilicariales	NO, Vestfold, 2015, E. Timdal	O-L-198940	OLICH2493-16	MK812096
Umbilicaria deusta	Umbilicariales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184712	OLICH930-13	MK812412
Umbilicaria havaasii	Umbilicariales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200930	OLICH2556-16	MK812376
Umbilicaria hirsuta	Umbilicariales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196282	OLICH2212-15	MK812437
Umbilicaria hirsuta	Umbilicariales	NO, Vestfold, 2015, E. Timdal	O-L-198937	OLICH2490-16	MK811754
Umbilicaria hirsuta	Umbilicariales	NO, Oppland, 2015, E. Timdal	O-L-201350	OLICH2735-16	MK812136
Umbilicaria hyperborea	Umbilicariales	NO, Hordaland, 2014, E. Timdal	O-L-195803	OLICH1675-14	MK812311
Umbilicaria hyperborea	Umbilicariales	NO, Finnmark, 2014, R. Haugan	O-L-195879	OLICH1724-14	KY266918
Umbilicaria hyperborea	Umbilicariales	NO, Finnmark, 2014, R. Haugan	O-L-195925	OLICH1770-14	KY266976
Umbilicaria hyperborea	Umbilicariales	NO, Finnmark, 2014, E. Timdal	O-L-195971	OLICH1816-14	KY266928
Umbilicaria hyperborea	Umbilicariales	NO, Troms, 2014, E. Timdal	O-L-195656	OLICH1996-15	MK812016
Umbilicaria hyperborea	Umbilicariales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184713	OLICH931-13	MK811931
Umbilicaria lyngei	Umbilicariales	NO, Sør-Trøndelag, 2000, B. P. Løfall	O-L-78093	OLICH4074-17	MK812368	no lichen substances
Umbilicaria lyngei	Umbilicariales	NO, Oppland, 2009, R. Haugan	O-L-159920	OLICH4075-17	MK812559
Umbilicaria lyngei	Umbilicariales	NO, Sør-Trøndelag, 2014, J.T. Klepsland	O-L-200558b	OLICH4096-17	MK811698
Umbilicaria nylanderiana	Umbilicariales	NO, Hedmark, 2015, M. Stapnes Dahl et al.	O-L-201305	OLICH2670-16	MK812167
Umbilicaria nylanderiana	Umbilicariales	NO, Hedmark, 2016, E. Timdal	O-L-201508	OLICH3983-17	MK812209
Umbilicaria nylanderiana	Umbilicariales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184729	OLICH950-13	MK812316
Umbilicaria polyrrhiza	Umbilicariales	NO, Oslo, 2014, E. Timdal	O-L-199999	OLICH1973-15	MK812126
Umbilicaria polyrrhiza	Umbilicariales	NO, Østfold, 2013, R. Braathen & E.W. Hanssen	O-L-201768	OLICH2595-16	MK812352
Umbilicaria polyrrhiza	Umbilicariales	NO, Østfold, 2017, S. Rui & E. Timdal	O-L-208345	OLICH4384-17	MK812239
Umbilicaria proboscidea	Umbilicariales	NO, Finnmark, 2014, H. Holien	O-L-195719	OLICH1591-14	KY266839
Umbilicaria proboscidea	Umbilicariales	NO, Troms, 2014, E. Timdal	O-L-195772	OLICH1644-14	MK811704
Umbilicaria proboscidea	Umbilicariales	NO, Finnmark, 2014, R. Haugan	O-L-195923	OLICH1768-14	KY266848
Umbilicaria proboscidea	Umbilicariales	NO, Finnmark, 2014, A. Millanes	O-L-195936	OLICH1781-14	KY266866
Umbilicaria proboscidea	Umbilicariales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196361	OLICH1920-15	MK812366
Umbilicaria proboscidea	Umbilicariales	NO, Buskerud, 2014, S. Rui & E. Timdal	O-L-196322	OLICH2252-15	MK812345
Umbilicaria rigida	Umbilicariales	SE, Torne Lappmark, 2014, E. Timdal	O-L-195668	OLICH2005-15	MK812082
Umbilicaria rigida	Umbilicariales	NO, Møre og Romsdal, 2015, R. Haugan	O-L-203950	OLICH4077-17	MK811777
Umbilicaria spodochroa	Umbilicariales	NO, Vestfold, 2015, E. Timdal	O-L-198938	OLICH2491-16	MK812519
Umbilicaria torrefacta	Umbilicariales	NO, Troms, 2014, E. Timdal	O-L-195634	OLICH1553-14	MK812116
Umbilicaria torrefacta	Umbilicariales	NO, Finnmark, 2014, E. Timdal	O-L-195843	OLICH1688-14	KY266942
Umbilicaria torrefacta	Umbilicariales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200900	OLICH2530-16	MK812602
Umbilicaria vellea	Umbilicariales	NO, Telemark, 2014, E. Timdal	O-L-196275	OLICH1949-15	MK811710
Umbilicaria virginis	Umbilicariales	NO, Møre og Romsdal, 2015, R. Haugan	O-L-203963	OLICH3162-16	MK812576
Usnea dasypoga	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184699	OLICH1411-14	MK812140
Usnea dasypoga	Lecanorales	NO, Telemark, 2014, E. Timdal	O-L-196273	OLICH1947-15	MK812173
Usnea dasypoga	Lecanorales	NO, Oslo, 2014, E. Timdal	O-L-200005	OLICH1979-15	MK812052
Usnea flammea	Lecanorales	NO, Vest-Agder, 2013, J.T. Klepsland	O-L-197827	OLICH2127-15	MK811847	constictic acid, stictic acid, usnic acid
Usnea fragilescens	Lecanorales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200604	OLICH2634-16	MK812021
Usnea subfloridana	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184700	OLICH1412-14	MK811828
Usnea subfloridana	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200814	OLICH2585-16	MK811766
Usnea subfloridana	Lecanorales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200728	OLICH2623-16	MK812453	thamnolic acid, usnic acid
Usnea wasmuthii	Lecanorales	NO, Sør-Trøndelag, 2013, J.T. Klepsland	O-L-198061	OLICH2131-15	MK812232	barbatic acid, salazinic acid, usnic acid
Usnea wasmuthii	Lecanorales	NO, Rogaland, 2013, J.T. Klepsland	O-L-197890	OLICH2132-15	MK812501	barbatic acid, usnic acid
Varicellaria lactea	Pertusariales	NO, Oppland, 2015, R. Haugan	O-L-204064	OLICH4380-17	MK812539	lecanoric acid, variolaric acid
Variospora flavescens	Teloschistales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200626	OLICH4054-17	MK812667
Variospora thallincola	Teloschistales	NO, Rogaland, 2014, J.T. Klepsland	O-L-200631	OLICH4056-17	MK812547
Verrucaria tenebrosa	Verrucariales	No, Nord-Trøndelag, 2015, J. Pykälä 48333	H-WG2_2130	OLICH2807-16	KY682961
Vulpicida juniperinus	Lecanorales	NO, Oppland, 2013, M. Bendiksby et al.	O-L-184298	OLICH1406-14	MK812148
Vulpicida juniperinus	Lecanorales	NO, Nord-Trøndelag, 2015, R. Haugan	O-L-201311	OLICH3857-17	MK812404
Vulpicida juniperinus	Lecanorales	NO, Sør-Trøndelag, 2017, E. Timdal	O-L-208203	OLICH4356-17	MK812500
Vulpicida pinastri	Lecanorales	NO, Finnmark, 2014, H. Holien	O-L-195944	OLICH1789-14	KY266954
Vulpicida pinastri	Lecanorales	NO, Telemark, 2015, S. Rui & E. Timdal	O-L-200937	OLICH2563-16	MK811933
Vulpicida pinastri	Lecanorales	NO, Buskerud, 2013, S. Rui & E. Timdal	O-L-184701	OLICH883-13	MK812364	pinastric acid, vulpinic acid
Xanthomendoza borealis	Teloschistales	NO, Sør-Trøndelag, 2014, E. Timdal	O-L-196336	OLICH2274-15	MK812442
Xanthomendoza borealis	Teloschistales	NO, Oppland, 2013, J.T. Klepsland	O-L-198333	OLICH2359-15	MK811659
Xanthomendoza fulva	Teloschistales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196295	OLICH2225-15	MK812265
Xanthomendoza fulva	Teloschistales	NO, Hedmark, 2006, B. P. Løfall et al.	O-L-147714	OLICH4440-17	MK812487
Xanthomendoza fulva	Teloschistales	NO, Sogn og Fjordane, 2012, B. Nordén & J.B. Jordal	O-L-199397	OLICH4442-17	MK811973
Xanthoparmelia conspersa	Lecanorales	NO, Telemark, 2014, S. Rui & E. Timdal	O-L-196278	OLICH1951-15	MK812598
Xanthoparmelia conspersa	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196296	OLICH2226-15	MK812585
Xanthoparmelia conspersa	Lecanorales	NO, Rogaland, 2016, S. Rui & E. Timdal	O-L-204999	OLICH3992-17	MK812248
Xanthoparmelia loxodes	Lecanorales	NO, Akershus, 2014, E. Timdal	O-L-199990	OLICH1964-15	MK811808
Xanthoparmelia loxodes	Lecanorales	NO, Vestfold, 2014, S. Rui & E. Timdal	O-L-196281	OLICH2211-15	MK811902
Xanthoparmelia loxodes	Lecanorales	NO, Aust-Agder, 2013, J.T. Klepsland	O-L-198317	OLICH2375-16	MK811982
Xanthoparmelia loxodes	Lecanorales	NO, Østfold, 2012, K.A. Lye	O-L-206876	OLICH4133-17	MK812427
Xanthoparmelia loxodes	Lecanorales	NO, Østfold, 2016, S. Rui & E. Timdal	O-L-208018	OLICH4190-17	MK812543
Xanthoparmelia loxodes	Lecanorales	NO, Østfold, 2017, S. Rui & E. Timdal	O-L-208343	OLICH4272-17	MK812627
Xanthoparmelia protomatrae	Lecanorales	NO, Aust-Agder, 2015, H. Riddervold	O-L-201574	OLICH3243-16	MK812594
Xanthoparmelia pulla	Lecanorales	NO, Sør-Trøndelag, 2014, R. Haugan	O-L-196012	OLICH1857-14	MK812429
Xanthoparmelia pulla	Lecanorales	NO, Østfold, 2016, S. Rui & E. Timdal	O-L-208019	OLICH4191-17	MK812706
Xanthoparmelia pulla	Lecanorales	NO, Østfold, 2017, S. Rui & E. Timdal	O-L-208344	OLICH4273-17	MK812285
Xanthoparmelia stenophylla	Lecanorales	NO, Østfold, 2017, S. Rui & E. Timdal	O-L-208338	OLICH4268-17	MK811834
Xanthoparmelia tinctina	Lecanorales	SE, Bohuslän, 2012, E. Timdal	O-L-184585	OLICH3233-16	MK812507
Xanthoparmelia tinctina	Lecanorales	NO, Østfold, 2017, S. Rui & E. Timdal	O-L-208333	OLICH4264-17	MK812642
Xanthoparmelia verruculifera	Lecanorales	NO, Oppland, 2015, E. Timdal	O-L-201346	OLICH2731-16	MK812281
Xanthoria parietina	Teloschistales	NO, Telemark, 2014, E. Timdal	O-L-195782	OLICH1654-14	MK812349
Xanthoria parietina	Teloschistales	NO, Oslo, 2014, M. Bendiksby et al.	O-L-196081	OLICH1897-14	MK812134
Xanthoria parietina	Teloschistales	NO, Østfold, 2012, K.A. Lye	O-L-206852	OLICH4143-17	MK811702
Xylographa parallela	Trapeliales	NO, Finnmark, 2011, E. Timdal	O-L-170700	OLICH3294-16	MK812221	no lichen substances
Xylographa vitiligo	Trapeliales	NO, Nord-Trøndelag, 2014, J.T. Klepsland	O-L-200273	OLICH3315-16	MK812010

Temporal coverage

Notes

The sampled material was collected in the period 1995–2017, with 90% being after 2010 and with a peak of 532 specimens in 2014 (Fig. 3).

Figure 3.

Distribution of year of collection of the specimens in the OLICH dataset

Usage rights

Use license

Open Data Commons Attribution License

Data resources

Data package title

DNA barcodes of Nordic lichens

Resource link

dx.doi.org/10.5883/DS-OLICH

Number of data sets

1

Data set 1.

Data set name

DS-OLICH

Data format

dwc, xml, tsv, fasta, trace

Number of columns

41

Download URL

http://www.boldsystems.org/index.php/Public_SearchTerms?query=DS-OLICH

Description

The OLICH dataset can be downloaded from the Public Data Portal of BOLD (www.boldsystems.org) in different formats (data as dwc, xml or tsv, sequences as fasta and trace files as trace). Alternatively, BOLD users can log in and access the dataset via the Workbench platform of BOLD. All records are also searchable within BOLD using the search function of the database.

Depending on future funding, the OLICH project will continue sequencing Nordic lichens for the BOLD database, with the ultimate (although unrealistic) goal of complete coverage. Hence, the dataset will be updated in the future with new sequences and information. The version of the dataset at the time of writing the manuscript is included as Suppl. materials 1, 2, 3, 4 in the form of two text files for record information as downloaded from BOLD, one text file with the collecting and identification data in Darwin Core Standard format (downloaded from BOLD and formatted by the authors) and of a fasta file containing all sequences as downloaded from BOLD.

It should be noted that, as the BOLD database is not compliant with the Darwin Core Standard format, the Darwin Core formatted file (dwc) that can be downloaded from BOLD is not strictly Darwin Core formatted. For a proper Darwin Core formatted file, see Suppl. material 3.

All records have images and geographical coordinates in BOLD.

Column labels below follow the labels downloaded in the tsv format. Columns with no content in our dataset are left out in the list below.

Data set 1.

Column label	Column description
processid	Unique identifier for the sample
sampleid	Unique identifier for the voucher specimen, i.e. museum catalogue number in either lichen herbarium or DNA bank at the Natural History Museum, University of Oslo
recordID	Entry number in the database
catalognum	Same as sampleid; Unique identifier for the voucher specimen, i.e. museum catalogue number in either lichen herbarium or DNA bank at the Natural History Museum, University of Oslo
institution_storing	The full name of the institution where specimen is stored
phylum_taxID	Taxonomic identifier of level Phylum
phylum_name	Phylum name
class_taxID	Taxonomic identifier of level Class
class_name	Class name
order_taxID	Taxonomic identifier of level Order
order_name	Order name
family_taxID	Taxonomic identifier of level Family
family_name	Family name
genus_taxID	Taxonomic identifier of level Genus
genus_name	Genus name
species_taxID	Taxonomic identifier of level Species
species_name	Species name
identification_provided_by	Name of primary individual repsonsible for providing taxonomic identification of the specimen
identification_method	The method(s) used to identify the specimen
identification_reference	Authorship of species name
voucher_status	Status of the specimen in an accessioning process
tissue_type	A brief description of the type of tissue analysed
collectors	Name(s) of the individual(s) responsible for collecting the sample in the field
habitat	A description of the habitat
Notes	Notes regarding species identification of the specimen or other note concerning the specimen.
lat	The geographical latitude (in decimal degrees) of the geographic centre of a location
long	The geographical longitude (in decimal degrees) of the geographic centre of a location
coord_accuracy	An estimation of the uncertainty of the coordinates given in the lat and long columns (metres)
elev	Elevation of sampling site. Measured in metres relative to sea level.
country	Name of country in which the organism was collected
province_state	Name of the province (in Norway "fylke") in which the organism was collected
region	Name of municipality in which the organism was collected
exactsite	Name of exact locality where the organism was collected
image_ids	Entry number in the database
image_urls	URL(s) from which the image(s) of the organism(s) can be accessed
media_descriptors	Term depicting, in this case, side of organism photographed, i.e. dorsal
captions	Short description of part of organism photographed
copyright_years	The year of the licence declaration for the image
copyright_licences	Type of licence associated with the image
copyright_institutions	The primary licence holder's institutional affiliation
photographers	The individual responsible for photographing

Additional information

Summary statistics and genetic distances

The dataset contains on average 2.6 records per species, ranging from one (184 species) to 12 (one species) records per species. Average sequence length was 541 bp (including sequences that did not include the whole ITS region due to unreadable trace files). The shortest sequence in the dataset was an incomplete sequence, 229 bp long. The species with the shortest complete ITS region was Haematomma ocroloeucum with 481 bp. The longest ITS region was 856 bp from Leptogium cochleatum. However, the two sequences of the latter species did not align with the others at the start of the sequence and it is possible that introns in the SSU rDNA are included.

We calculated intra- and interspecific genetic distances for the three genera with most sampled species and performed barcode gap analyses on those. Distances and key statistics are found in Table 2.

Table 2.

Key statistics and genetic distances for the three genera with the most sampled species; Ramalina, Umbilicaria and Cladonia.

Genus	Ramalina	Umbilicaria	Cladonia
Number of species in genus in this dataset	9	19	47
Number of records	26	52	122
Average number of records per species	2.9	2.7	2.6
Average intraspecific distances	0–1.3%	0–2.3%	0–3.4%
Interspecific distances	0.4–8.2%	2.5–12.1%	0.2–15.6%

In Ramalina, there was an overlap between intra- and interspecific distances (Table 2, Fig. 4a). The overlap is caused by the one species pair with higher maximum intraspecific distance than the distance to the nearest neighbour (Fig. 4b); i.e. R. farinacea had an intraspecific distance of 0.9% and a distance of 0.4% to R. subfarinacea. If this species pair is excluded from the distance-dataset, a barcode gap will occur at 2% (Fig. 4a).

Genetic distances within the three lichen genera with most sampled species in the OLICH dataset.

Figure 4a.

Frequency distribution of intraspecific distances (blue) and interspecific distances (orange). A gap between intra- and interspecific distances illustrates the barcode gap in the genus.

Figure 4b.

Scatterplots representing maximum intraspecific distances versus minimum distance to closest neighbour for all species with more than one specimen per species, in each of the three genera.

Figure 4c.

Frequency distribution of intraspecific distances (blue) and interspecific distances (orange). A gap between intra- and interspecific distances illustrates the barcode gap in the genus.

Figure 4d.

Scatterplots representing maximum intraspecific distances versus minimum distance to closest neighbour for all species with more than one specimen per species, in each of the three genera.

Figure 4e.

Frequency distribution of intraspecific distances (blue) and interspecific distances (orange). A gap between intra- and interspecific distances illustrates the barcode gap in the genus.

Figure 4f.

Scatterplots representing maximum intraspecific distances versus minimum distance to closest neighbour for all species with more than one specimen per species, in each of the three genera.

For Umbilicaria, there were no species pairs with higher maximum intraspecific distance than the distance to the nearest neighbour, but one species had the same maximum intraspecific distance as the distance to the nearest neighbour, namely U. proboscidea whose closest neighbour was U. virginis (Fig. 4d). There is thus no barcode gap in our Umbilicaria dataset, as the intra- and interspecific distances overlaps around 3% (Fig. 4c).

Whereas Ramalina and Umbilicaria, despite the above-mentioned issues, show a tendency for displaying a barcode gap, Cladonia does not show the same pattern. There is no gap, but rather overlapping, smooth curves of intra- and interspecific distances in the barplot (Table 2, Fig. 4e). Most of the species pairs that disrupt the pattern are complexes where species are known to be sometimes difficult to distinguish with traditional characters and these are found below the line in Fig. 4f: C. coniocraea/gracilis, C. maxima/ecmocyna and C. cenotea/subfurcata/crispata/squamosa.

ITS has previously been shown to give poor resolution amongst Cladonia species (Fontaine et al. 2010, Kotelko and Piercey-Normore 2010, Pino-Bodas et al. 2013) and a lack of barcode gap has been demonstrated (Kelly et al. 2011). This probably has to do with their high morphological plasticity (Pino-Bodas et al. 2013) which may have led to erroneous identifications/taxonomy, but there might also be evolutionary explanations, such as incomplete lineage sorting (van Velzen et al. 2012).

Observations

From the phylogenetically and BLASTn based quality control, some specific taxonomic observations were made that are listed below.

Alectoria sarmentosa (Ach.) Ach. ssp. vexillifera (Nyl.) D.Hawksw.: This subspecies is currently accepted in the Nordic flora (Velmala and Myllys 2011, Nordin et al. 2019). It differs morphologically from spp. sarmentosa in forming a more prostrate thallus with broader and more flattened main branches and, ecologically, in growing on the ground or on rocks mainly in coastal and alpine areas. Subspecies sarmentosa grows mainly on trunks and branches of conifers in boreal forests. Our sequence of spp. vexillifera (OLICH1478) is nested within our four sequences of spp. sarmentosa, however, which does not support the recognition of the subspecies (Fig. 5).

Figure 5.

Maximum likelihood tree based on the OLICH sequences of Alectoria sarmentosa.

Numbers above branches and thick lines indicate bootstrap support ≥ 70%. Bold font indicates focus taxon and grey font the outgroup. The two-letter ISO country codes are included. This applies for all phylogram figures (Figs 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15).

Brodoa oroarctica (Krog) Goward: The species differs from B. atrofusca (Schaer.) Goward in the shape of the thallus and lobes and in the lack of physodic acid in most of the thallus except for in some lobe tips (Krog 1974). The former species is circumpolar arctic-alpine, with southernmost occurrences in Europe in the southern Scandinavian mountains, whereas the latter is a European species known mainly from the Alps and Pyrenées and with scattered occurrences in Northern Europe. Our four barcode sequences of B. oroarctica are however nested within the four sequences of B. atrofusca (Fig. 6), and the complex seems to be in need of revision.

Figure 6.

Maximum likelihood tree of Brodoa atrofusca and B. oroarctica based on all available ITS sequences.

Bryobilimbia fissuriseda (Poelt) Timdal, Marthinsen & Rui, comb. nov.

Mycobank: MB 830330

Basionym: Lecidea fissuriseda Poelt, Mitteilungen der Botanischen Staatssammlung München 4: 181, 1961. ≡ Mycobilimbia fissuriseda (Poelt) Poelt & Hafellner, Herzogia 8: 56, 1989. Type: Switzerland, Berner Oberland, Gipfelgrader Niesen, Nordhänge, alt. 2300 m, 1955, Poelt B55189 (M!, holotype).

This rarely collected species was described from the Alps in the large, collective genus Lecidea Ach. (Poelt 1961). It was later transferred to the segregate genus Mycobilimbia Rehm by Poelt & Hafellner (in Hafellner 1989). Partly due to the presence of characteristic blue, K+ green granules in the hymenium and the hypothecium, Poelt (1961) and Timdal (1992) assumed it was closely related to M. hypnorum (Lib.) Kalb & Hafellner (syn. Lecidea fusca (Schaer.) Th. Fr.). Fryday et al. (2014), based on a molecular study, excluded M. hypnorum from Mycobilimbia and placed it, with four additional species, in the new genus Bryobilimbia Fryday et al. Mycobilimbia fissuriseda was not included in that study, but our phylogenetic tree, based on the DNA barcodes (Fig. 7), shows that it is, in fact, closely related to B. diapensiae and B. hypnorum and it is hence here transferred to Bryobilimbia.

Figure 7.

Maximum likelihood tree of Bryobilimbia and Mycobilimbia based on all available ITS sequences. The presumedly related genera Clauzadea and Romjularia are included and Farnoldia and Lecidea are used as the outgroup.

Bryoria capillaris (Ach.) Brodo & D.Hawksw.: The species is still accepted in the Nordic checklist (Nordin et al. 2019), despite several recent molecular phylogenies (Myllys et al. 2011, Velmala et al. 2014, Myllys et al. 2016, Boluda et al. 2019) showing it nested within B. implexa (Hoffm.) Brodo & D. Hawksw. The OLICH dataset shows the same topography (Fig. 8): B. capillaris nested within B. implexa. However, all our specimens of the B. capillaris/implexa complex were identified by morphology prior to TLC analysis and the correlation holds: all pale specimens (B. capillaris) contain alectorialic acid (plus accessories) and barbatolic acid, whereas no dark specimens (B. implexa) do. Fumarprotocetraric and protocetraric acids occur variably in both species. Either the DNA barcode marker and the other markers used in recent phylogenetic studies do not distinguish between two species which may be recognised by the correlation of thallus colour and secondary chemistry or the two "species" are conspecific and their morphological/chemical difference is caused by other factors. For example, the presence/absence of an additional symbiont (basidiomycete yeast) were found likely to trigger the production of vulpinic acid in the cortex of Bryoria fremontii/tortuosa by Spribille et al. (2016).

Figure 8.

Maximum likelihood tree of Bryoria capillaris and B. implexa based on the OLICH sequences.

Cetrelia cetrarioides (Duby) W.L. Culb. & C.F. Culb. and C. monachorum (Zahlbr.) W.L. Culb. & C.F. Culb.: The three chemotypes of Cetrelia olivetorum (Nyl.) W.L. Culb. & C.F. Culb. s. lat., currently recognised in the Nordic countries (Bjelland et al. 1997, Obermayer and Mayrhofer 2007, Nordin et al. 2019), were recently shown to represent three distinct species in the phylogenetic analysis by Mark et al. (2019). The diagnostic, main compounds of those species are imbricaric acid in C. monachorum, olivetoric acid in C. olivetorum s. str. and perlatolic acid in C. cetrarioides (Mark et al. 2019). The seven specimens sequenced in the OLICH project fell into the C. cetrarioides and C. monachorum clades (Fig. 9) and the TLC analyses of the sequenced specimens showed the chemotypes were in accordance with the species identifications (Table 1).

Figure 9.

Maximum likelihood tree of the OLICH sequences of Cetrelia, with selected ITS sequences from Mark et al. (2019).

Hypotrachyna afrorevoluta (Krog & Swinscow) Krog & Swinscow: The species is morphologically and chemically quite similar to H. revoluta (Flörke) Hale and the OLICH sequences (Fig. 10) confirm the presence of the two species in the Nordic countries. See Elix and Thell (2011) for the subtile morphological differences between the species.

Figure 10.

Maximum likelihood tree of the OLICH sequences of Hypotrachyna afrorevolulta and H. revoluta, with all available ITS sequences in GenBank.

Nephroma tangeriense (Maheu & A.Gillet) Zahlbr.: The species was recently reported as new to the Nordic countries (Klepsland 2013) from two collections from Rogaland county. The OLICH project has sequenced one of those (OLICH2019), as well as four additional collections from Hordaland and Vest-Agder counties and confirms its identity and distinction from N. laevigatum Ach. (Fig. 11).

Figure 11.

Maximum likelihood tree based on the OLICH sequences of Nephroma laevigatum and N. tangeriense, with selected ITS sequences from GenBank.

Peltigera “neocanina”: BLASTn search revealed the identity of OLICH1878, which was originally identified as P. canina (L.) Willd., as P. “neocanina”. This is still an invalid name despite the taxon, which must be regarded as a morphologically cryptic species, has been known since the phylogenetic study of Peltigera by Miadlikowska and Lutzoni (2000). Jüriado et al. (2017) showed its presence in Estonia and Magain et al. (2018) in Norway.

Pseudocyphellaria citrina (Gyeln.) Lücking, Moncada & S. Stenroos: The Pseudocyphellaria species with yellow soralia, occurring in the Nordic countries, is currently named P. crocata (L.) Vain. (Nordin et al. 2019). Lücking et al. (2017b) showed that P. crocata was heterogeneous and split it into 13 species. European material was not included in that study and it remained unknown which species occurs in our area. The sequences obtained from three Norwegian specimens in the OLICH project all belong in the P. citrina clade of Lücking et al. (2017b) (Fig. 12).

Figure 12.

Maximum likelihood tree of the Pseudocyphellaria crocata complex with the OLICH sequences added to those of Lücking et al. (2017b).

Psoroma cinnamomeum Malme: The specimen from which OLICH702 was generated was originally identified as Psoroma tenue Henssen var. boreale Henssen, but BLASTn search against GenBank indicated its identity as P. cinnamomeum. A phylogenetic analysis, based on most of the ITS sequences of Psoroma in GenBank (including those used by Park et al. 2018), supported this identification as OLICH702 and came out in a strongly supported clade with the six other specimens of P. cinnamomeum, two of which are from southernmost Chile and four from Svalbard (Fig. 13). The clade was the sister of the strongly supported clade of the two specimens of P. tenue in GenBank, both from Antarctica. Both P. cinnamomeum and P. tenue were described from Tierra del Fuego, var. boreale from USA, Colorado (cf. Henssen and Renner 1981). OLICH702 contains porphyrilic acid and two unknown compounds (apparently related to porphyrilic acid due to the colour and fluorescence on the chromatograms) and hence fits the current concept of P. tenue var. boreale (cf. Jørgensen 2007). Unless molecular studies show that there are two morphologically very similar species in the Northern Hemisphere, both containing porphyrilic acid, we believe P. cinnamomeum is the correct name for the taxon currently known as P. tenue var. boreale.

Figure 13.

Maximum likelihood tree of Psoroma based on all available ITS sequences.

Ramalina capitata (Ach.) Nyl.: The species has often been included in R. polymorpha (Lilj.) Ach. in the past (e.g. Krog and James 1977), but is now usually accepted as distinct (e.g. Nimis et al. 2018, Nordin et al. 2019). In our phylogeny (Fig. 14), Nordic and western European material of the two species are mostly well separated, but the four sequences from Turkey in GenBank are deviant and the complex seems to be in need of further study.

Figure 14.

Maximum likelihood tree of the Ramalina capitata/polymorpha complex, based on all available ITS sequences.

Ramalina subfarinacea (Cromb.) Nyl.: Our two sequences of this species, as well as the two ITS sequences available in GenBank, are nested in the clade of Ramalina farinacea (L.) Ach. in our phylogeny (Fig. 15). According to Krog and James (1977), there are subtle morphological differences between the species, but apparently the major distinguishing feature is the ecology: mainly on coastal rock (R. subfarinacea) vs. mainly on bark of deciduous, inland trees (R. farinacea). More study is needed in order to evaluate the taxonomic status of R. subfarinacea.

Figure 15.

Maximum likelihood tree of the Ramalina farinacea/subfarinacea complex, based on all available ITS sequences.

Sticta fuliginoides Magain & Sérus.: BLASTn search of the sequence of an originally identified Sticta fuliginosa (Hoffm.) Ach. (OLICH2638) showed 99.4–100% similarity with the five sequences of S. fuliginoides and 90.0–90.2% similarity with the 9 sequences of S. fuliginosa in the phylogeny of Magain and Serusiaux (2015). This confirms the presence of S. fuliginoides in the Nordic countries, as recently reported by Ekman et al. (2019), based on morphological characters.

Supplementary Material

Supplementary material 1

OLICH library - Processing details

Gunnhild Marthinsen, Siri Rui, Einar Timdal

Data type: Record information - processing details on ITS sequences

Brief description: The file includes information about all records in BOLD for the OLICH library, i.e. the DS-OLICH dataset, at time of writing. It contains data on ITS sequences. The data are as downloaded from BOLD without further processing.

File: oo_302451.txt

Supplementary material 2

OLICH library - Specimen details

Gunnhild Marthinsen, Siri Rui, Einar Timdal

Data type: Record information - specimen data

Brief description: The file includes information about all records in BOLD for the OLICH library, i.e. the DS-OLICH dataset, at time of writing. It contains collecting and identification data. The data are as downloaded from BOLD, without further processing.

File: oo_302452.txt

Supplementary material 3

OLICH library - Specimen details - Darwin Core Standard

Gunnhild Marthinsen, Siri Rui, Einar Timdal

Data type: Record information - specimen data in Darwin Core Standard format

Brief description: The file includes information about all records in BOLD for the OLICH library, i.e. the DS-OLICH dataset, at time of writing. It contains collecting and identification data. The data are downloaded from BOLD and further formatted to the DC standard by the authors.

File: oo_302453.txt

Supplementary material 4

OLICH library - DNA sequences

Gunnhild Marthinsen, Siri Rui, Einar Timdal

Data type: Genomic data, DNA sequences

Brief description: ITS sequences in fasta format. Each sequence is identified by the BOLD ProcessID and species name, separated by pipe. The data are as downloaded from BOLD.

File: oo_301034.fas