Abstract Abstract
A synopsis of the genera Coleus Lour, Equilabium A.J.Paton, Mwany. & Culham and Plectranthus L’Hér. (Lamiaceae, Tribe Ocimeae, Subtribe Plecranthinae) is presented. Generic delimitation follows a recently published molecular phylogeny which identified Coleus as the sister of the remaining genera of Subtribe Plectranthinae; Plectranthus as sister to Tetradenia Benth. and Thorncroftia N.E.Br., and a separate phylogenetically distinct genus Equilabium comprising species previously placed in Plectranthus. In this treatment, 294 species of Coleus, 42 of Equilabium, and 72 of Plectranthus are recognized. All but one of the combinations in Equilabium are new as only the genus and type species have been previously published. Two-hundred and twelve names are changed to combinations in Coleus from Plectranthus, Pycnostachys Hook. and Anisochilus Benth.
Keywords: Anisochilus , Equilabium , generic delimitation, nomenclature, Ocimeae, Plectranthinae , Pycnostachys , Solenostemon
Introduction
Plectranthus L’Hér. is a widely used horticultural and medicinal plant genus (van Jaarsveld 2007, Lukhoba et al. 2006, Rice et al. 2011). It belongs in tribe Ocimeae, subtribe Plectranthinae, an Old World tropical group of around 450 species. Current taxonomic treatments generally follow Harley et al. (2004) by including the formerly recognized Coleus Lour. and Solenostemon Thonn. within Plectranthus. However, within horticultural literature, Solenostemon is still frequently recognized (e.g. RHS (2017), Shepherd and Maybry (2016)). The commonly cultivated plant shown in Fig. 1 is thus variously known as Plectranthus scutellarioides (L.) R.Br or Solenostemon scutellarioides (L.) Codd., but “coleus” is frequently used as a common name for this species, increasing the confusion.
Figure 1.
A Display of cultivated Coleus scutellarioides (L.) Benth. in the Real Jardin Botánico, Madrid. B Close up of the cultivar ‘Campfire’, winner of the Royal Horticultural Society’s peoples choice award at Wisley 2017 (Photo: A.Paton).
Recent phylogenetic work has shown that the circumscription of Plectranthus needs to be viewed in the context of related genera in Tribe Ocimeae, subtribe Plectranthinae. Paton et al. (2018) demonstrate that a clade containing the type of Coleus (C. amboinicus Lour. ≡ P. amboinicus (Lour.) Spreng.) is sister to the other genera of the Plectranthinae, and the type species of Plectranthus, P. fruticosus L’Hér., belongs in a clade which is sister to Thorncroftia N.E.Br and Tetradenia Benth. A further clade containing species that were previously placed in Plectranthus was recognized by Paton et al. (2018) as Equilabium Mwany., A.J.Paton & Culham (Fig. 2).
Figure 2.
Summarised phylogeny of the Plectranthinae showing relationships of recognised genera following Paton et al. (2018): the numbers in triangles represent species numbers though the numbers of species in the two subclades of Coleus cannot be estimated (see discussion); dates of branching are given in black.
The aim of this paper is to provide the necessary nomenclatural changes consistent with the recognition of Plectranthus, Coleus and Equilabium as distinct genera following phylogenetic results provided by Paton et al. (2018).
Taxonomic history
Plectranthus was described by L’Héritier in 1788 (L’Héritier 1788) and Coleus by Loureiro in 1790 (de Loureiro 1790). In previous global reviews of the Lamiaceae, e.g. Bentham (1832, 1848, 1876) and Briquet (1897), Coleus was treated as distinct from Plectranthus by having the 4 stamens fused together rather than free to their base. Morton (1962) recognized that the character of the fused stamens was more plastic than previously thought and merged Coleus into Plectranthus, as first suggested by Brown (1810). However, Morton also placed weight on calyx characteristics leading him to recognize several genera based on calyx morphology including Solenostemon, Isodictyophorus A.Chev, Leocus A.Chev., Englerastrum Briq. and Neohyptis J.K.Morton. The merging of Coleus into Plectranthus has been followed by most authors since Morton’s account, for example, Keng (1978), covering Malesian species, Blake (1971, Australia), Codd (1985, southern Africa), Hedge et al. (1998, Madagascar), Suddee et al. (2004, Indochina), Forster (2011, Australia) and Paton et al. (2009, 2013, eastern and southern tropical Africa). Furthermore, although Codd (1985) and Morton (2006) also maintained Solenostemon, Keng (1978), Hedge et al. (1998) and subsequent authors recognised Solenostemon as synonymous with Plectranthus.
Despite the agreed taxonomic status by the above authors, Cramer (1978) maintained that Coleus is a separate genus from Plectranthus, as did Wu and Huang (1977)and Li and Hedge (1994) for Chinese species. Cramer’s decisions were in part based upon Sri Lankan Coleus that have imbricate bracts (“floral leaves”) and a decurrent upper calyx lip in addition to fused stamens. Cramer also suggested Plectranthus could have a 3-lobed upper calyx lip, but that is misleading since he was considering species of Isodon (Schrad. ex Benth.) Spach as part of Plectranthus: Isodon is now recognized in a separate subtribe and is very distinct from the Plectranthinae (Ryding 1993, Harley et al. 2004, Zhong et al. 2010).
A common theme in the treatments listed above is that, since Briquet (1897), all have been based on regional accounts of relatively small numbers of species. Morton (1962) considered 28 species of the Coleus/Plectranthus complex and Cramer (1978), 11 species. Morton recognized the problems in this narrow regional approach noting, “hence the genus Plectranthus must for the moment be a repository for Coleus or Plectranthus-like species which cannot satisfactorily be placed in other genera” and Cramer considered his account to be “ by no means conclusive, least of all authoritative”. Paton in Harley et al. (2004) thus took a broader approach, considering characters used to delimit allied genera in the context of the Plectranthinae as a whole. In Paton’s treatment, Coleus and the small segregate genera recognized by Morton were merged into Plectranthus, making Plectranthuss.l. morphologically recognizable from other genera and an easily communicable concept. However, this was a conservative, morphologically based account and did not tackle how best to circumscribe taxa within the large Plectranthuss.l.
Paton et al. (2004) provided a preliminary test of the broad concept of Plectranthus by presenting a phylogeny of tribe Ocimeae, demonstrating that the Plectranthinae was a monophyletic group which could be generally diagnosed by having stamens adnate and contiguous at the base of the anterior corolla lip. The presence of fused stamens, the original defining character of Coleus, was shown to be homoplasious by mapping it onto this phylogeny. However, the study identified two main clades within the Plectranthinae: one containing the type of Coleus and members of Solenostemon, Leocus A.Chev., Pycnostachys Hook. and Anisochilus Benth.; and the other containing the type species of Plectranthus and members of Alvesia Welw., Aeollanthus C. Mart. ex Spreng, Capitanopsis S.Moore, Tetradenia and Thorncroftia.
The phylogenetic study by Paton et al. (2018) increased the sampling of the Plectranthinae and added another plastid marker to increase tree resolution, and similarly found that Plectranthuss. l. was polyphyletic (see summary of analysis in Fig. 2). As a result of this work, Paton et al. (2018) suggested recognition of Coleus for the clade containing the Coleus type, Plectranthuss.s. for the clade sister to Thorncroftia and Tetradenia, and a new genus, Equilabium, for the clade formerly containing Plectranthus species mainly from tropical Africa, but with one in India. Paton et al. (2018) pointed out that these clades were morphologically diagnosable by characters that had been used to define informal morphological groups by Paton et al. (2009, 2013). These characters are the position of attachment of the pedicel to the calyx, the relative lengths of the corolla lobes and the shape of the corolla tube (Fig. 3). The phylogenetic results indicate that three genera recognized by Harley et al. (2004), (Pycnostachys, Anisochilus and Leocus) are nested within Coleus, and this close relationship is corroborated by species of intermediate morphology between Coleus and the former taxa Pycnostachys (Paton et al. 2009, 2013), Anisochilus (Suddee et al. 2014), and Leocus (Pollard and Paton 2009). Paton et al. (2018) also showed that Solenostemon is not monophyletic, and that recognition of Solenostemon would render Coleus paraphyletic by its exclusion, therefore we argue that Solenostemon should not be recognised.
Figure 3.
Calyx and corolla morphology. Calyces in Coleus with pedicel attached excentrically above midline of calyx: 1C. betonicifolius2C. platyphyllus showing the rounded lateral calyx lobe in some species of the formally recognized Solenostemon. Calyx in Equilabium with pedicel attached ± at centre of calyx base with the lateral lobe held midway between the anterior and posterior lobes: 3E. megafolium4E. masukensis. Calyces in Plectranthus with pedicel attached ± at centre of calyx base with the lateral lobe closer to the anterior than the posterior lobes: 5P. alboviolaceus. Corolla shape in Coleus showing unequal anterior and posterior corolla lips: 6C. schizophyllus7C. hockii. Corolla shape of Equilabium showing equal anterior and posterior corolla lips: 8P. parvus10P. laxiflorus. Corolla shape of Plectranthus showing equal anterior and posterior corolla lips and gibbous-based corolla: 9P. strangulatus;. Drawn by Emmanuel Papadopoulos. Reproduced from Flora of Tropical East Africa (Paton et al. 2009).
Methods
The phylogenetic study presented by Paton et al. (2018) was based on a representative sample of species of the Plectranthinae. Recent floristic accounts and the World Checklist Of Selected Plant Families were consulted (Govaerts et al. 2019, Paton et al. 2009, 2013, Suddee et al. 2004, Codd 1985, Hedge et al. 1998, Forster 1992, 1994, 2011 and references within). IPNI (www.ipni.org) was consulted to trace any recently described species of Coleus or Plectranthus and the protologues of these were examined. A survey of herbarium collections (AAU, BK, BKF, BM, BOL; BR, C, CALI, COI, EA, ETH, FT, G, HBG, K, L, LE, LMA, LISC, LMU, MEL, MH, MO, NHT, P, PRE, RNG, S, STU, TCD, UPS, W, WAG; Abbreviations following Thiers (2017)), was undertaken also using JSTOR Global Plants (https://plants.jstor.org/) to access type specimens not otherwise seen. Cited types have been seen unless otherwise stated. The attribution of species to the genera Coleus, Plectranthus and Equilabium was guided by the characters given in Paton et al. (2018) and in the key at the beginning of the taxonomic account below.
Results and discussion
The changes in generic delimitation proposed by Paton et al. (2018) have a large impact on nomenclature. An alternative nomenclatural treatment would be to consider the monophyletic Plectranthinae to be components of a single genus, Plectranthus. Most recent taxonomic papers have included Coleus within Plectranthus. However, taking 2005 as a starting point, following publication of Harley et al. (2004) as the most comprehensive baseline taxonomy for the Lamiaceae, 15,700 papers cite Coleus in the title, but only 9320 cite Plectranthus (source Google Scholar). This suggests that Coleus is still a broadly used concept. Tetradenia with mainly dioecious, ± actinomorphic flowers is part of the sister group of the Plectranthus in the strict sense. Thus there are no easily observed morphological apomorphies that could be used to easily diagnose an enlarged Plectranthus; whereas the generic delimitation proposed in Paton et al. (2018), results in monophyletic genera which are relatively easy to diagnose morphologically.
In this treatment, 294 species of Coleus, 42 of Equilabium, and 72 Plectranthus are recognized. All the combinations in Equilabium are new as only the genus and type species E. laxiflorum have been previously published (Paton et al. 2018). 212 names need to be changed to combinations in Coleus from combinations in Plectranthus, Pycnostachys and Anisochilus. However, 82 of these combinations in Coleus already exist, reflecting previous confusion. No new names are necessary for species in Plectranthus.
Of the characters cited by Cramer (1978) as supporting the delimitation of Coleus and Plectranthus, the presence of a decurrent calyx tube is variable and found in Coleus, Plectranthus and Equilabium. Imbricate bracts do only occur in Coleus, but this character is only obvious in some species. Generally the species are very easy to attribute to Coleus, Plectranthus or Equilabium using the pedicel attachment, corolla lobe length and tube shape characters described in the key below. The presence of persistent bracts and slightly flattened nutlets also supports the division of Coleus (often cauducus bracts and slightly flattened nutlets), from Plectranthus and Equilabium (usually persistent bracts and ovoid nutlets), though those characters have more exceptions than those characters used in the key. Both Equilabium and Plectranthus have rarely more than 3-flowered cymes, whereas Coleus can have one to very many flowers per cyme. Although species can be misplaced in some instances due to atypical corolla structure, the asymetrical pedicel attachment at the base of the calyx gives the most clear indication of the species’ placement in Coleus. Generic identification can be harder with small-flowered Coleus species that show asymetrical pedicel attachment but have more or less equal corolla lips; however, the number of these cases is small. In Equilabium the upper lip can be slightly shorter than the lower lip in some cases but at the extreme it is about 0.8 of the length of the lower lip, whereas in Coleus the upper lip is usually much shorter. The southern African Coleus zombensis (Baker) A.J.Paton, C. gibbosus A.J.Paton (Plectranthus rehmannii Gürke) and Coleus sessilifolius (A.J.Paton) A.J.Paton can also provide confusion as the corolla is basally gibbous, similar to that of many species of Plectranthus; however, they have a Coleus-like calyx and the corolla tube is sigmoid rather than straight as in true Plectranthus.
We present the list of species in alphabetical order, rather than according to the previously proposed infrageneric classification by Codd (1975), which is unworkable. Although the smaller infrageneric groups he recognized in the infrageneric classification of Plectranthuss.l. of Southern Africa are natural groups of related Southern African species, Codd’s classification does not cover the extent of character variation and combinations seen outside the Flora of Southern Africa area, making it difficult to apply globally. As pointed out by Paton et al. (2018), groups previously thought to be easily delimited by morphological characters can be paraphyletic. For example, subgenus Calceolanthus Codd was delimited by Codd by having a dense ring of hairs in the calyx throat, but the phylogeny presented by Paton et al. (2018) suggests that the clade containing these species also contains species which lack this character. Larger infrageneric groupings recognized by Codd contain a mixture of lineages, for example, Codd’s section Plectranthus while containing mostly species placed in Plectranthus here, also includes P. dolichopodus Briq. (here placed in Coleus) and P. laxiflorus Benth. and P. petiolaris Benth. (here both are placed in Equilabium).
Further study is required to provide a more robust formal infrageneric classification of Coleus and Plectranthus, although within Coleus there seems to be a continuity of form making further morphological diagnoses of clades difficult. There are two well supported clades within Coleus (clades A and B sensuPaton et al. (2018)), but no clear morphological apomorphies to diagnose these clades have been identified. Further phylogenetic work is underway by the authors: fifty species are now described for Australia and a current phylogenetic analysis focusing on these species is being prepared from a comprehensively sampled data set (T.C. Wilson, unpubl.); and an analysis of species in India is also being prepared (K. Smitha, unpubl.).
Taxonomic account
A key to the genera of the Plectranthinae is provided and followed by treaments of the three genera now containing species of Plectranthus s.l. sensuHarley et al. (2004): Coleus, Equilabium and Plectranthus. New combinations of Capitanopsis and the generic protologue of Equilabium can be found in Paton et al. (2018). No changes to other genera of the Plectranthinae are proposed. The generic descriptions are followed by a listing of the species in alphabetical order with details of synonymy, types and distribution. The account ends with a list of unplaced names, followed by a list of excluded names now placed in other genera. Readers should also consult the online treatment in the World Checklist of Selected Plant Families (Govaerts et al. 2019, www.kew.org/wcsp) as this is continually updated and includes TDWG level 3 distributions and Raunkiaer life form descriptors. A short description of the Plectranthinae and key to the genera recognized is provided below.
Subtribe Plectranthinae Endlicher, Gen. Pl.: 609. 1838.
Herbs or shrubs. Inflorescence an indeterminate thyrse with cymes sessile or pedunculate, ebracteolate, rarely with minute persistent or deciduous bracteoles; bracts persistent or deciduous, rarely coloured. Calyx usually with a single-lobed posterior lip and a 4-lobed anterior lip, sometimes actinomorphic and regularly 5-lobed. Corolla strongly 2-lipped, rarely actinmorphic or subactinomorphic (Tetradenia), anterior lip concave to cucullate, more rarely flat. Stamens declinate, rarely spreading (Tetradenia), exserted, anterior and posterior attached adjacently at corolla throat (separated in Tetradenia). Disc usually 4-lobed, lobes alternating with nutlets. Nutlets with small inconspicuous areole.
Key to the genera of the Plectranthinae
| 1 | Corolla actinomorphic, or almost so, stamens spreading not declinate | Tetradenia |
| – | Corolla strongly zygomorphic, stamens declinate | 2 |
| 2 | Lateral lobes of the corolla deflexed towards the anterior lobe | Thorncroftia |
| – | Lateral lobes of the corolla ascending towards the median lobes of the posterior lip or horizontal and held between the anterior lobe and median lobes of the posterior lip | 3 |
| 3 | Posterior lip of corolla truncate, lateral lobes usually horizontal, short, held between the anterior lobe and median lobes of the posterior lip | Capitanopsis |
| – | Posterior lip of corolla not truncate, diverging from anterior lip of the corolla, with lateral lobes usually ascending and close to median lobes of posterior lip | 4 |
| 4 | Calyx basally circumscissile; tube distally dorsiventrally flattened | Aeollanthus |
| – | Calyx very rarely circumscissile; tube not dorsiventrally flattened | 5 |
| 5 | Calyx 3-lobed (1 upper, 2 lower), expanded and membranous in fruit | Alvesia |
| – | Calyx 5-lobed (1 upper, 4 lower, or actinomorphic), rarely expanded and membranous in fruit | 6 |
| 6 | Calyx with pedicel attached asymmetrically to the base of the calyx tube, opposite the posterior lip; corolla with upper (posterior) lip shorter than lower (anterior), rarely subequally lobed (Fig. 3) | Coleus |
| – | Calyx with pedicel attached symmetrically to the base of the calyx tube, not opposite the upper lip, corolla lobes more or less equal in length (Fig. 3) | 7 |
| 7 | Calyx throat truncate with lateral lobes of the calyx held midway between the uppermost and lowermost lobes; corolla tube usually sigmoid, parallel-sided at base, dilating towards throat (Fig. 3) | Equilabium |
| – | Calyx throat oblique with lateral lobes of the calyx held closer to the lowermost pair of lobes than the upper lobe; corolla tube straight or curved downwards, usually gibbous, saccate or shortly spurred at base (Fig. 3) | Plectranthus |
Synopsis of genera and species of Coleus, Equilabium and Plectranthus
Genus Coleus
Coleus
Lour., Fl. Cochinch.: 372. 1790
14400339EBB05DDDB662E2861BE2D048
Figure 4.
Species of Coleus 1. AColeus anamudianus (Photo: K. Smitha) BC. amboinicus (Photo: Neil Crouch) CC. barbatus (Photo: Neil Crouch) DC. bracteatus (Photo: Witawat Keiwbang) EC. harmandii (Photo: Thamarat Phutthai).
Figure 5.
Species of Coleus 2. AColeus maculosus subsp. lanatus (Photo: Vincent Droissart) BC. mirabilis (Photo: David Goyder) CC. porphyranthus (Photo: Neil Crouch) DC. orthodontus (Photo: Bart Wursten) EC. tenuicaulis (Photo: Ernst van Jaarsveld) FC. strobilifer (Photo: Naiyana Tetsana).
Pycnostachys Hook., Exot. Fl. 3: t. 202. 1826. synon. nov. Type: P. coerulea W.J.Hooker (here recognized under Coleus stenostachys (Baker) A.J.Paton, non Coleus coeruleus Gürke).
Solenostemon Thonn. in C.F.Schumacher, Beskr. Guin. Pl.: 271. 1827. Type: Solenostemon ocymoides Schumach. & Thonn (here recognized under Coleus monostachyus (P. Beauv.) A.J.Paton).
Anisochilus Wall. ex Benth., Edwards’s Bot. Reg. 15: t. 1300. 1830. synon. nov. Type: Anisochilus carnosus (L.f.) Wall. ex Benth. (here recognized under Coleus strobilifer (Roxb.) A.J.Paton, non Coleus carnosus Hassk.).
Mitsa Chapel. ex Benth. nom. inval, Labiat. Gen. Spec.: 52. 1832.
Echinostachys E.Mey., Comm. Pl. Afr. Austr.: 243. 1838. synon. nov. Type: Echinostachys reticulata E.Mey. (here recognized under Coleus kirkii (Baker) A.J.Paton, non Coleus reticulatus A.Chev.).
Saccostoma Wall. ex Voigt, Hort. Suburb. Calcutt.: 449. 1845. Type: S. urticifolius Voigt. (See unplaced names).
Majana Rumph. ex Kuntze, nom. superfl. Revis. Gen. Pl. 2: 523. 1891.
Englerastrum Briq., Bot. Jahrb. Syst. 19: 178. 1894. Type: E. schweinfurthii Briq. (here recognized under Coleus rhodesianum (N.E.Br.) A.J.Paton).
Neomuellera Briq., Bot. Jahrb. Syst. 19: 180. 1894. Type: N. welwitschii Briquet (here recognized under Coleus fredericii G.Taylor).
Capitanya Schweinf. ex Gürke, Bot. Jahrb. Syst. 21: 105. 1895. Type: Capitanya ostostegioides Gürke (here recognized as Coleus otostegioides (Gürke) A.J.Paton).
Burnatastrum Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4. 3A: 358. 1897. Type: Burnatastrum lanceolatum (Bojer ex Benth.) Briq. (here recognized as Coleus lanceolatus (Bojer ex Benth.) A.J.Paton).
Stiptanthus Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4. 3A: 352. 1897., synon. nov. Type: Stiptanthus polystachyus (Benth.) Briq. (here recognized as Coleus polystachys (Benth.) A.J.Paton).
Symphostemon Hiern, Cat. Afr. Pl. 1: 867. 1900. Type: S. insolitus (C. H. Wright) Hiern (here recognized as Coleus insolitus (C.H.Wright) Robyns & Lebrun).
Leocus A.Chev., J. Bot. (Morot) 22: 125. 1909. Type: L. lyratus Chevalier (here recognized as Coleus lyratus (A.Chev.) Roberty).
Isodictyophorus Briq., Mém. Soc. Bot. France 8: 285. 1917. Type: I. chevalieri Briquet (here recognized as Coleus reticulatus A.Chev.).
Briquetastrum Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 102. 1929. Type: B. africanum (Baker) W. Robyns et Lebrun (here recognized under Coleus engleri (Briq.) A.J.Paton, non C. africanum Benth.).
Holostylon Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 103. 1929. Type designated here: Holostylon gracilipedicellatum Robyns & Lebrun (here recognized as Coleus gracilipedicellatum (Robyns & Lebrun) A.J.Paton).
Ascocarydion G.Taylor, J. Bot. 69 (suppl. 2): 162. 1931. Type: Ascocarydion mirabile (Briq.) G.Taylor (here recognized as Coleus mirabilis Briquet).
Neohyptis J.K.Morton, J. Linn. Soc., Bot. 58: 258. 1962. Type: Neohyptis paniculata (Baker) J.K.Morton (here recognized under Coleus guerkei (Briq.) A.J.Paton, non Coleus paniculatus Benth.).
Rabdosiella Codd, Bothalia 15: 9. 1984. Type: R. calycina (Bentham) Codd (here recognized as Coleus calycinus (Benth.) A.J.Paton).
Coleus sect Solenostemonoides Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3A): 359. 1897. Type as below.
Calchas P.V.Heath, Calyx 5: 160. 1997. Type: Solenostemon latifolius (Hochst. ex Benth) J.K.Morton, designated by Codd (1975), here recognized under Coleus bojeri Benth.
Type species.
C. amboinicus Lour.
Description.
Annual or perennial herbs or shrubs, sometimes succulent, sometimes with a fleshy or tuberous rootstock. Leaves opposite, sometimes whorled. Inflorescence thyrsoid, strongly condensed to lax, with cymes sessile or pedunculate, sometimes with long cincinnate branches, 1– many-flowered; bracts subtending cymes usually not persistent beyond flowering, sometimes erect forming a coma at apex of flowering stem. Flowers sessile or pedicellate, sometimes spirally arranged on inflorescence axis. Calyx funnel-shaped to tubular, straight or curved, often ventrally gibbous, with pedicel attaching asymmetrically at calyx base opposite the posterior lip, two lipped to regularly 5-lobed, rarely circumscissile at base (C. coeruleus); throat glabrous, pubescent or bearded, open or closed by scales or converging lips; if regularly lobed the lobes triangular, lanceolate or spinescent; if two lipped, posterior lip 1-lobed, larger or similar size to other lobes, decurrent or not; anterior lip 4-lobed with lobes variously shaped; lateral lobes lanceolate, triangular to oblong or rounded; held midway between the posterior lobe and median lobes of the anterior lip; median lobes lanceolate to triangular, sometimes separated by a sinus or sometimes fused at base for a longer distance than the lateral and median lobes. Corolla two-lipped with posterior lip shorter than anterior lip, rarely subequal; tube sigmoid to almost straight, narrow and parallel-sided at base, expanding towards throat, very rarely gibbous, rarely annulate; posterior lip 4-lobed, erect or ascending; anterior lip horizontal cucullate to concave. Stamens 4; filaments fused together or free, held within the anterior lip or exserted. Style bifid with subulate lobes, rarely entire. Nutlets ovoid, often dorso-ventrally flattened.
294 species in the Old World tropics and subtropics; often cultivated elsewhere.
Coleus abyssinicus
(Fresen.) A.J.Paton comb. nov.
9AEC70AA05F255F3A4AA30A3A74B7C82
urn:lsid:ipni.org:names:77200982-1
Pycnostachys abyssinica Fresen., Flora 21: 608. 1838. Type: Ethiopia, Sidamo, Rüppell s.n. (holotype: FR? Possibly lost, see Ryding 2006).
Distribution.
Ethiopia.
Coleus acariformis
(P.I.Forst.) P.I.Forst. comb. nov.
A11A3E9217B05708BB7A025596FF5D52
urn:lsid:ipni.org:names:77200983-1
Plectranthus acariformis P.I.Forst., Austrobaileya 9: 279. 2014. Type: Australia, Queensland, Moreton District, Noosa National Park, Mt Emu, 5 Feb. 2014, P.I.Forster & G.Leiper PIF40678 (holotype: BRI; isotypes: K, MEL, NSW, US).
Distribution.
Australia: Queensland.
Coleus achankoviliensis
Smitha & A.J. Paton nom. nov.
4242CECD833E595B97557996DADCBA4E
urn:lsid:ipni.org:names:77200984-1
Anisochilus petraeus J.Mathew & Yohannan, Taiwania 62(2): 144. 2017, non Coleus petraeus (Backer ex Adelb.) A.J.Paton. Type: India, Kollam district, 1.5 km from Achankovil town, Achankovil, J.Mathew 4527 (holotype TBGT; isotype MSSRF).
Distribution.
S. India.
Coleus actites
(P.I.Forst.) P.I.Forst. comb. nov.
2BCF6342FF635FAF8C033388B51E1D38
urn:lsid:ipni.org:names:77200985-1
Plectranthus actites P.I.Forst., Austrobaileya 4: 163. 1994. Type: Australia, Queensland, Leichhardt District, Anvil Peak, Hodgson Range, NE of Clermont, 28 Aug. 1990, P.I.Forster PIF7255 (holotype: BRI).
Distribution.
Australia: Queensland.
Coleus adenanthus
(Dalzell & A.Gibson) A.J.Paton comb. nov.
69E8D465BA125F75BC80A8D8986E7678
urn:lsid:ipni.org:names:77200986-1
Anisochilus adenanthus Dalzell & A.Gibson, Bombay Fl.: 206. 1861. Type: India, the Concan, near Dharwar, Bababoodan hills, Bombay Herbarium of the late N. A. Dalzell. Presented by Mrs Dalzell, April 1878, Dalzell s.n. (holotype: K).
Anisochilus verticillatus Hook.f., Fl. Brit. India 4: 629. 1885. Type: India, the Concan, near Dharwar, Bababoodan hills, Bombay Herbarium of the late N. A. Dalzell. Presented by Mrs Dalzell, April 1878, Dalzell s.n. (lectotype: K, designated by Suddee and Paton 2009).
Distribution.
S. India.
Coleus adenophorus
(Gürke) A.J.Paton comb. nov.
2AC191EE7BB65D059391BDB5D9D9762C
urn:lsid:ipni.org:names:77200987-1
Plectranthus adenophorus Gürke, Bot. Jahrb. Syst. 30: 398. 1901.
Englerastrum adenophorum (Gürke) T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 75. 1924. Type: Tanzania, Njombe District: Kipengere, Goetze 960 (holotype: B destroyed).
Distribution.
SW. Tanzania.
Coleus aegyptiacus
(Forssk.) A.J.Paton comb. nov.
64C528D1CC695DC7A62ADED8568FEA39
urn:lsid:ipni.org:names:77200990-1
Ocimum aegyptiacum Forssk., Fl. Aegypt.-Arab.: cxv, 109. 1775.
Plectranthus aegyptiacus (Forssk.) C.Chr., Dansk Bot. Ark. 4(3): 21. 1922.
Plectranthus crassifolius Vahl, Symb. Bot. 1: 44. 1790., nom. illeg.
Germanea crassifolia Poir. in J.B.A.M.de Lamarck, Encycl., suppl. 2: 764. 1812., nom. superfl.
Coleus zatarhendi Benth., Labiat. Gen. Spec.: 50. 1832., nom. superfl.
Plectranthus zatarhendi E.A.Bruce, Bull. Misc. Inform. Kew 1935: 590. 1935., nom. superfl.
Ocimum zatarhendi Forssk., Fl. Aegypt.-Arab.: 109. 1775., nom. inval.
Majana zatarhendi (Forssk.) Kuntze, Revis. Gen. Pl. 2: 524. 1891, nom. inval. Type: Yemen, Beit el Fakih (Bayt al Faqih), Forsskål s.n. (type: not seen, believed destroyed); “Arabia”, Herb. Forsskål s.n. (neotype: KIEL; C, K (photos), designated by Ryding and Paton (2001)).
Coleus tenuiflorus Vatke, Linnaea 43: 92. 1881.
Plectranthus tenuiflorus (Vatke) Agnew, Upland Kenya Wild Fl.: 636. 1974. Type: Kenya, Kitui, Hildebrandt 2742 (lectotype: CORD, designated by Paton et al. (2009)).
Coleus camporum Gürke, Bot. Jahrb. Syst. 19: 214. 1894. Type: Tanzania, Pangani District: Pangani R. ford, Volkens 485 (holotype: B, destroyed; isotype: BM, G).
Coleus ghindanus Schweinf. ex Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 435. 1900. Type. Eritrea, Ghinda valley, Schweinfurth & Riva 2149 (holotype: K; isotypes: BR, FT, P).
Distribution.
Eritrea to Tanzania, SW. Arabian Peninsula.
Coleus affinis
(Gürke) A.J.Paton comb. nov.
5338F22FA1865A27B73B40B15B2436DB
urn:lsid:ipni.org:names:77200991-1
Pycnostachys affinis Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 345. 1895.Type: Tanzania, Mwanza, Stuhlmann 4693 (holotype: B, destroyed).
Pycnostachys speciosa Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 345. 1895., non Coleus speciosus Baker f. Type: Kenya/Tanzania, east shore of Lake Victoria, Fischer 499 (lectotype: B, destroyed, designated by Bruce 1940).
Pycnostachys dawei N.E.Br., Gard. Chron., ser. 3, 41: 18. 1907. Type: cultivated from seed sent from Uganda, Dawe s.n. 1906 (holotype: K).
Distribution.
Rwanda to E. Trop. Africa.
Coleus albicalyx
(Suddee) Suddee comb. nov.
8DD2E1F132B05104997CFDB75DA408F9
urn:lsid:ipni.org:names:77200993-1
Plectranthus albicalyx Suddee, Kew Bull. 59: 401. 2004. Type: Thailand, Kanchanaburi, Salak Phrae W.S., Khao Tab Tao, 480 m, 29 Sept. 1998, Suddee et al. 868 (holotype: BKF; isotypes: K, TCD).
Distribution.
Thailand.
Coleus aliciae
(Codd) A.J.Paton comb. nov.
633F827EFAA658FD815B1D407E12624F
urn:lsid:ipni.org:names:77200995-1
Plectranthus madagascariensis var. aliciae Codd, Bothalia 11: 404. 1975.
Plectranthus aliciae (Codd) van Jaarsv. & T.J.Edwards, Bothalia 27: 5. 1997. Type: South Africa, Transkei, Butterworth, Kentani, Pegler 909 (holotype: PRE).
Distribution.
South Africa: E. Cape Prov. to S. KwaZulu-Natal.
Coleus alloplectus
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
CA72711D3DAB58BFABB6D43BCF9CE1FB
urn:lsid:ipni.org:names:77200997-1
Plectranthus alloplectus S.T.Blake, Contr. Queensland Herb. 9: 29. 1971. Type: Australia, Queensland, Mt Greville, 21 Apr. 1962, S.T.Blake 21703 (holotype: BRI).
Distribution.
Australia: New South Wales, Queensland.
Coleus alpinus
Vatke, Linnaea 37: 322. 1872
94B8D2A9E17C57AFA7D91905AEDB080F
Plectranthus alpinus (Vatke) Ryding, Bot. Jahrb. Syst. 121: 147. 1999. Type: Ethiopia, Edda Jesus near Dawra Tabor (Debra Tabor), 25 Sept. 1863, Schimper s.n. (lectotype: BM, designated by Ryding (1999); isolectotype JE).
Coleus assurgens Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 428. 1900.
Plectranthus assurgens (Baker) J.K.Morton, J. Linn. Soc., Bot. 58: 267. 1962. Type: Ethiopia, Begemder, 1863- 8, Schimper s.n. (holotype: K, (K000431867)).
Coleus schoensis Gürke, Bot. Jahrb. Syst. 38: 170. 1906. Type: Ethiopia, Addis Abeba Highlands, Sept. 1900, Ellenbeck 1576 (holotype: B destroyed).
Coleus gracilentus S.Moore, J. Linn. Soc., Bot. 38: 273. 1908. Type: Uganda, E Ruwenzori, Wollaston s.n. (holotype: BM).
Coleus microtrichus Chiov., Nuovo Giorn. Bot. Ital., n.s., 36: 368. 1929. Type: Ethiopia, Arussi, Lake Zuai basin, Abruzzi 37 (holotype: TO; isotype: C).
Coleus lebrunii Robyns, Bull. Jard. Bot. État Bruxelles 17: 73. 1943. Type: DRC, Ruwenzori, Butagu valley, Robyns 4579 (holotype: BR; isotype: K).
Coleus wittei Robyns, Bull. Jard. Bot. État Bruxelles 17: 74. 1943. Type: DRC, Kamatembe, R. Bishkishaki, de Witte 1551 (holotype: BR).
Distribution.
Nigeria to Ethiopia and south to Malawi.
Coleus altanmouiensis
(T.C.Wilson, P.I.Forst., & M.A.M.Renner) T.C.Wilson & P.I.Forst. comb. nov.
9A51C6E3315451788577987807681589
urn:lsid:ipni.org:names:77200999-1
Plectranthus altanmouiensis T.C.Wilson, P.I.Forst., & M.A.M.Renner, Telopea 21: 84. 2018. Type: Cape Melville National Park, Cape York, Cook District, Queensland, 14 Aug 2015, T.C. Wilson 610 and M.A.M. Renner (holotype: NSW 985152; isotype: BRI).
Distribution.
Australia: Queensland.
Coleus amboinicus
Lour., Fl. Cochinch.: 372. 1790
9B7F1879D0D655A1B9FC63FF9377E089
Plectranthus amboinicus (Lour.) Spreng., Syst. Veg. 2: 690. 1825.
Majana amboinica (Lour.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Rumphius plate in Herb. Amb. 5, t. 102/2. 1750 (lectotype: illustration, designated by Cramer (1978); epitype, designated by Suddee et al. (2004) and based on the choice of a representative specimen selected by Launert (1968): Thailand, Pai District: Mae Hong Son, 25 May 1921, Kerr s.n. (epitype: BM).
Ocimum vaalae Forssk., Fl. Aegypt.-Arab.: 111. 1775., nom. rej.
Coleus vaalae (Forssk.) Deflers, Rev. Egypt: 423. 1894.
Coleus aromaticus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830. Type: India, Patna, Buchanan-Hamilton Wall. Cat. 2730B (syntypes: K-W (K001116968, K001116969).
Coleus crassifolius Benth. in N.Wallich, Pl. Asiat. Rar. 2: 15. 1830. Type: India, Dindygul, Herb. Wight s.n. in Wall. Cat. 2731 (syntypes: BM, K (K000820149, K (K000820151); K-W (K001116970).
Coleus suganda Blanco, Fl. Filip.: 483. 1837.
Majana suganda (Blanco) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Philippines, Luzon, Maragondong, Cavite Province, Merrill: Species Blancoanae No. 129 (neotype, designated by Suddee et al. (2004): K). Blanco’s specimens have not been traced and are believed destroyed.
Coleus carnosus Hassk., Flora 25(2 Beibl.): 25. 1842.
Majana carnosa (Hassk.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Coleus suborbicularis Zoll. & Moritzi, Syst. Verz.: 4. 1846., nom. illeg. Type: Indonesia, Zollinger 1950 (holotype: G-DC; isotypes: A, P, S).
Coleus amboinicus var. violaceus Gürke, Bot. Jahrb. Syst. 19: 210. 1894. Type: Tanzania, Moshi District: Lake Chala, Volkens 321 (syntypes: B destroyed, BM, K) & Pangani R. ford, Volkens 487 (syntypes: B, destroyed, BM, K).
Coleus subfrutectosus Summerh., Bull. Misc. Inform. Kew 1928: 392. 1928. Type: Seychelles, Mahé, Botanical Station, Dupont 204 (holotype: K).
Distribution.
Kenya to South Africa, Arabian Peninsula, India.
Coleus amicorum
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
C4477FB4D2D051DA94433283C66335F7
urn:lsid:ipni.org:names:77201000-1
Plectranthus amicorum S.T.Blake, Contr. Queensland Herb. 9: 26. 1971. Type: Australia, Queensland, Tinaroo Range, 21 Aug. 1963, S.T.Blake 22094 (holotype: BRI).
Distribution.
Australia: Queensland.
Coleus amiculatus
(T.C.Wilson, P.I.Forst. & M.A.M.Renner) T.C.Wilson & P.I.Forst. comb. nov.
1280233D5A75588AA868AA4F21F0A44C
urn:lsid:ipni.org:names:77201001-1
Plectranthus amiculatus T.C.Wilson, P.I.Forst. & M.A.M.Renner, Telopea 21: 81. 2018. Type: Along Pajinka Road, Cook District, Queensland, 8 Aug 2015, T.C. Wilson 595 & M.A.M. Renner (holotype: NSW (NSW985146); isotype: BRI).
Distribution.
Australia: Queensland.
Coleus amoenus
(P.I.Forst.) P.I.Forst. comb. nov.
BDE0CF7E84B05E3D800815F16A3443F5
urn:lsid:ipni.org:names:77201002-1
Plectranthus amoenus P.I.Forst., Austrobaileya 4: 656. 1997. Type: Australia, Queensland, Cook District, Baldy Mt., Atherton, Jun. 1991, P.I.Forster PIF8488 (holotype: BRI; isotypes: CNS, MEL).
Distribution.
Australia: Queensland.
Coleus anamudianus
(Smitha & Sunojk.) Smitha comb. nov.
129CD01A824D5810A87FE38218EFDCE4
urn:lsid:ipni.org:names:77201003-1
Plectranthus anamudianus Smitha & Sunojk., Phytotaxa 284(1): 52. 2016. Type: India, Kerala, Idukki district, in valley, wet rocky surfaces, Rajamala, Smitha & Sunojkumar 135412 (holotype: CALI; isotype MH).
Distribution.
S. India.
Coleus angolensis
(G.Taylor) A.J.Paton comb. nov.
61481BF0F4B85C41B0480E968422DAAC
urn:lsid:ipni.org:names:77201004-1
Pycnostachys angolensis G.Taylor, J. Bot. 69 (suppl. 2): 164. 1931. Type: Angola, Casima, Anha, Gossweiler 1734 (holotype: BM).
Distribution.
Angola.
Coleus angulatus
(Hedge) A.J.Paton & Phillipson comb. nov.
EA63D4B6F7B1530C9F2AEE5E14784CFE
urn:lsid:ipni.org:names:77201005-1
Plectranthus angulatus Hedge, Fl. Madag. 175: 211. 1998. Type: Madagascar, Bekopaka, May 1964, Bosser 19616 (holotype: P; isotypes: E, MO, TAN).
Distribution.
Madagascar.
Coleus apoensis
Elmer, Leafl. Philipp. Bot. 7: 2694. 1915
06FC86CC9CED5587B63A7E0C9208C19F
Plectranthus apoensis (Elmer) H.Keng, Gard. Bull. Singapore 24: 147. 1969. Type: Phillipines, Mindanao, Davao distr., Todaya, Mount Apo, Sept. 1909, A.D.E. Elmer 11601 (holotype PNH (not seen); isotypes: GH, K).
Distribution.
Philippines (Mindanao).
Coleus apreptus
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
C6356DCBB1465AD1970E366C039A5104
urn:lsid:ipni.org:names:77201006-1
Plectranthus apreptus S.T.Blake, Contr. Queensland Herb. 9: 47. 1971. Type: Australia, Queensland, Freshwater Creek, near Cairns, 17 May 1962, S.T.Blake 21730 (holotype: BRI).
Distribution.
Australia: Queensland.
Coleus apricus
(P.I.Forst.) P.I.Forst. comb. nov.
AD3BDD541DEC5231A44A3EC861EBA00D
urn:lsid:ipni.org:names:77201007-1
Plectranthus apricus P.I.Forst., Austrobaileya 4: 165. 1994. Type: Australia, Queensland, Cook District. Turtle Rock Area, 22 Jan. 1991, P.I. Forster PIF12831 & A.R.Bean (holotype: BRI, isotypes: CNS, K, MEL).
Distribution.
Queensland.
Coleus arabicus
Benth. in A.P. de Candolle, Prodr. 12: 79. 1848
3879FB0F480F51E6B4B3C1C667A62980
Majana arabica (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Arabia, Botta s.n. (holotype: P).
Plectranthus asirensis J.R.I.Wood, Kew Bull. 37: 601. 1983. Type: Saudi Arabia, Taif, Botta s.n. (holotype: P; isotype K).
Teucrium mazziarii Heldr. ex Nyman, Consp. Fl. Eur.: 567. 1881, nom. inval.
Distribution.
Arabian Peninsula.
Coleus arenicola
(P.I.Forst.) P.I.Forst. comb. nov.
9C6656551A725A9D9FC845F67121CCB2
urn:lsid:ipni.org:names:77201008-1
Plectranthus arenicola P.I.Forst., Muelleria 7: 375. 1991. Type: Australia, Queensland, Plant cultivated at St Lucia, Brisbane from material the same collection as P.I.Forster 5456 from Moreton Telegraph Station, near Kennedy Hill, 22 Oct. 1989, P.I.Forster PIF5835 (holotype: BRI; isotypes: CNS, K, MEL).
Distribution.
Australia: Queensland.
Coleus argentatus
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
48B8F571C0BF5A29AC4EEDD40C309AF9
urn:lsid:ipni.org:names:77201009-1
Plectranthus argentatus S.T.Blake, Contr. Queensland Herb. 9: 27. 1971. Type: Australia, Queensland, Mt Roberts, McPherson Range, 5 May 1955, S.T.Blake 19803 (holotype: BRI).
Distribution.
Australia: Queensland to NE. New South Wales. Widely cultivated (or hybrids from it).
Coleus argenteus
(Gamble) A.J.Paton comb. nov.
C109F0C89FC154B29578E07A2ADE6287
urn:lsid:ipni.org:names:77201010-1
Anisochilus argenteus Gamble, Fl. Madras: 1127. 1924. Type: India, Madras, Pulney Hills, Kodaikanal Ghat new road, 29 Dec. 1898, Bourne 1441C (lectotype: K; isolectotype: K, designated by Suddee & Paton (2009).
Distribution.
S. India.
Coleus argentifolius
(Ryding) A.J.Paton comb. nov.
099168CBFEAA5872B2D50D2F4B6FEC2D
urn:lsid:ipni.org:names:77201011-1
Plectranthus argentifolius Ryding, Kew Bull. 54: 122. 1999. Type: Solalia, Sanaag Region, Erigavo Escarpment, Popov 1218 (holotype: K; isotype: ETH).
Distribution.
N. Somalia, Yemen.
Coleus articulatus
(I.M.Johnst.) A.J.Paton comb. nov.
4A0A3296CA17545A9FBD470010AA3552
urn:lsid:ipni.org:names:77201012-1
Symphostemon articulatus I.M.Johnst., Contr. Gray Herb., n.s., 73: 38. 1924. Type: Angola, east of Cuanza (Coanza) R., Curtis 309 (holotype: GH).
Plectranthus hockii De Wild., Repert. Spec. Nov. Regni Veg. 11: 542. 1913.
Plectranthus katangensis De Wild., Ann. Mus. Congo Belge, Bot., sér. 4, 2: 135. 1913., nom. superfl.
Holostylon katangense (De Wild.) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 105. 1929., nom. superfl.
Ocimum hockii (De Wild.) Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 368. 1928., non Coleus hockii De Wild. Type: DRC, Upper Katanga, Dilemba, Hock s.n. (syntype: BR, K) & Plateau de Shinkwari (Manika), Hock s.n. (syntype: BR).
Distribution.
SW. Tanzania, DRC, Malawi, Zambia, Angola.
Coleus ater
A.J.Paton nom. nov.
A02D111B37595FB38F6474D45F1A32F8
urn:lsid:ipni.org:names:77201014-1
Anisochilus wightii Hook.f., Fl. Brit. India 4: 628. 1885., non Coleus wightii Benth. Type: India, Deccan Peninsula, Anamallay Mts, July 1851, Herb. Wight 2132/1 (holotype: K; isotypes: C, L).
Distribution.
S. India.
Coleus auriglandulosus
(A.J.Paton) A.J.Paton comb. nov.
F4AC65F6AC7F5D84BAB5E216E51999B1
urn:lsid:ipni.org:names:77201015-1
Plectranthus auriglandulosus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 307. 2009. Type: Kenya, Tana R. District: Tana River National Primate Reserve, near Makere, Luke et al. TPR 605 (holotype, K; isotype: EA).
Distribution.
SE. Kenya.
Coleus australis
(R.Br.) A.J.Paton comb. nov.
9FAFC92A22F95381BA495DA75604993E
urn:lsid:ipni.org:names:77201016-1
Plectranthus australis R.Br., Prodr. Fl. Nov. Holl.: 506. 1810.
Moschosma australe (R.Br.) Benth., Labiat. Gen. Spec.: 708. 1835.
Moschosma brownii Heynh., Alph. Aufz. Gew.: 421. 1847.
Plectranthus parviflorus var. australis (R.Br.) Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 357. 1895.
Germanea australis (R.Br.) Britten, Bot. Cook Voy. 3: 75. 1905. Type: New South Wales, Port Jackson, 17 May, 1802, R. Brown (lectotype: BM; isolectotypes: BRI, K, designated by Blake (1971)).
Plectranthus australis var. vulgaris Domin, Biblioth. Bot. 89: 564. 1928., nom. inval.
Plectranthus parviflorus Willd., Hort. Berol.: t. 65. 1806., non Coleus parviflorus Benth. Type: cultivated in Berlin, unknown origin, Wildenow Herbarium 11078 (holotype: B).
Plectranthus paniculatus Jacq., Fragm. Bot.: 62.tab 91. 1806., non Coleus paniculatus Benth. Type: Cultivated in Vienna. Illustration cited above.(lectotype: illustration designated here).
Plectranthus parviflorus Spreng. in L.V.F.Henckel von Donnersmarck, Adumbr. Pl. Hort. Hal.: 17. 1806., nom. illeg.
Germanea parviflora Poir. in J.B.A.M.de Lamarck, Encycl., suppl. 2: 764. 1811.
Majana parviflora (Poir.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Plectranthus sieberi Benth., Labiat. Gen. Spec.: 710. 1835. Type: Australia, “In Nova Hollandia tropica Sieber” (holotype: K).
Plectranthus parviflorus var. elatior Benth. in A.P.de Candolle, Prodr. 12: 67. 1848. Type: Northern Territory, Port Essington, Armstrong (holotype: K).
Plectranthus parviflorus var. major F.M.Bailey, Queensland Agric. J. 28: 199. 1912., nom. nud.
Plectranthus parviflorus var. minor F.M.Bailey, Queensland Agric. J. 28: 199. 1912., nom. nud.
Plectranthus australis f. densiflora Domin, Biblioth. Bot. 89: 564. 1928. Type: Queensland, Tamborin Mtn., Domin s.n. (holotype: PR).
Distribution.
Australia, Lesser Sunda Is. to Pacific.
Coleus autranii
Briq., Bull. Herb. Boissier 2: 129. 1894
465B3DE94EF95525A706FA1CB6DEFB41
Solenostemon autranii (Briq.) J.K.Morton, Novon 8: 266. 1998.
Plectranthus autranii (Briq.) Erhardt, Götz & Seybold, Grosse Zander 2: 1825. 2008.
Calchas autranii (Briq.) P.V.Heath, Calyx 5: 160. 1996. Type: Ethiopia, no precise locality, 1853, Schimper 693 (holotype: G; isotype: K).
Coleus silvaticus Gürke, Bot. Jahrb. Syst. 19: 219. 1894.
Solenostemon silvaticus (Gürke) Agnew, Upland Kenya Wild Fl.: 640. 1974. Types: Tanzania, Lushoto District: Usambara, Holst 3704 & 3708 (syntypes: B, destroyed); neotype: Tanzania, W Usambara Mts, Shagai Forest, near Sunga, Drummond & Hemsley 2593 (neotype: K, designated by Paton et al. (2009)).
Plectranthus tysonii Gürke, Bot. Jahrb. Syst. 26: 77. 1898. Type: South Africa, Umzimkulu, Clydesdale, Tyson 1295 (isotypes: K, SAM (numberered 1295 SAM herbarium number and cited by Gürke), PRE (PRE numbered Hb. Tyson 2769 and cross referenced to 1295 as cited in Codd (1985)).
Coleus odoratus Gürke, Bot. Jahrb. Syst. 38: 167. 1906. Type: Tanzania, W Usambara Mts, Sakare, Engler 939 (syntype: B, destroyed; photo K); & Engler 1006 (syntype: B destroyed).
Coleus latidens S.Moore, J. Linn. Soc., Bot. 38: 273. 1908.
Calchas latidens (S.Moore) P.V.Heath, Calyx 6: 51. 1999. Type: Uganda, E Ruwenzori, Wollaston s.n. (holotype: BM).
Coleus ostinii Chiov., Ann. Bot. (Rome) 9: 125. 1911. Type: Ethiopia, Dembia, Angareb to Gondar and Fenter, Chiovenda 1450 (syntype: FT); Cococc above Gondar, Chiovenda 1854 (syntype: FT).
Coleus darfurensis R.D.Good, J. Bot. 62: 138. 1924.
Solenostemon darfurensis (R.D.Good) B.Mathew, Kew Bull. 25: 258. 1971. Type: Sudan, Jebel Mara, Lynes 4c (holotype: BM; fragment K).
Coleus uliginosus T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 29. 1930. Type: Kenya, Meru, R.E. & T.C.E. Fries 1762 (syntype: B destroyed; isosyntype: K).
Coleus ascendens Gilli, Ann. Naturhist. Mus. Wien 77: 32. 1973. Type: Tanzania, Morogoro District: Uluguru Mts, Chenzema (Tchenzema), Gilli 456 (holotype: W).
Distribution.
Ethiopia, Central Africa, East Africa to South Africa.
Coleus barbatus
(Andrews) Benth. ex G.Don in J.C.Loudon, Hort. Brit.: 483. 1830
A0C4A89A257156DE835504673BC22D8C
Plectranthus barbatus Andrews, Bot. Repos. 10: t. 594. 1810. Type: Illustration of cultivated material in Bot. Rep. 9, t. 594. 1809., (lectotype: illustration cited above; Eritrea, Dekemehare, Colville 47 (epitype: K, designated by Ryding (1999)).
Coleus forskalaei auct., non Briq.
Distribution.
Eritrea to East and Central Africa, Arabian Peninsula, Indian Subcontinent to SC. China. Widely cultivated.
Notes.
The name Coleus forskalaei or forskohlii has been often used incorrectly for this species.
Coleus barbatus var. barbatus
DEA7D2C4AD125CCC9290E4A500A15B82
Ocimum asperum Roth, Nov. Pl. Sp.: 268. 1821.
Plectranthus asper (Roth) Spreng., Syst. Veg. 2: 690. 1825.
Orthosiphon asperus (Roth) Benth. ex Sweet, Hort. Brit., ed. 2: 413. 1830. Type: India, Mysore, Nundidroog, Heyne s.n. in Wall. Cat. 2728C (holotype: B destroyed; isotype: K-W).
Plectranthus comosus Sims, Bot. Mag. 49: t. 2318. 1822. Type: illustration of plant originally from Nepal, (lectotype: illustration cited above).
Coleus huberi Regel syn. nov., Trudy Imp. S.-Peterburgsk. Bot. Sada 7: 543. 1881. Type: Russia, cultivated in St Petersburg from seed collected in Ethiopia, (type not found in LE).
Coleus penzigii Dammann ex Baker, Gard. Chron., ser. 3, 14: 616. 1893. Type: cultivated at Kew, received from Hort. Dammann, probably originating from material collected by Penzig in “the mountains of Abyssinia” (holotype: K).
Coleus vestitus Baker, Bull. Misc. Inform. Kew 1895: 224. 1895. Type: Somalia, Golis Range, E. Cole s.n. (syntype: K); Lort-Phillips (syntype: K).
Plectranthus pseudobarbatus J.K.Morton, Novon 8: 265. 1998. Type: Somalia, Togdheer Region, Golis Range, Cole s.n. (lectotype: K; designated by Ryding (1999)).
Coleus forskohlii var. adoensis Briq., Annuaire Conserv. Jard. Bot. Genève 2: 235. 1898. Type: Ethiopia, Adua, 1 Oct. 1837, Schimper 333 (holotype: not seen in G; isotypes: BR, K, MPU, WAG).
Coleus speciosus Baker f., J. Bot. 1899: 64. 1899. Type: Somalia, Woqooyi Galbeed/Togdheer Regions, “Wagga Mountains”, Lort-Phillips s.n. (holotype: BM; isotype K).
Coleus coerulescens Gürke, Bot. Jahrb. Syst. 38: 169. 1906.
Plectranthus coerulescens (Gürke) R.H.Willemse, Kew Bull. 40: 95. 1985. Type: Ethiopia, Harar, Ellenbeck 743 (holotype: B, destroyed; isotype: K).
Coleus adolfi-friderici Perkins in G.W.J.Mildbraed (ed.), Wiss. Erg. Deut. Zentr.-Afr. Exped., Bot. 2: 550. 1913. Type: Rwanda, Mohasi, Büschen to Buganza, Mildbraed 671 (holotype: B, destroyed).
Distribution.
Eritrea to N. Tanzania, Arabian Peninsula, Indian Subcontinent to SC. China.
Coleus barbatus var. grandis
(L.H.Cramer) A.J.Paton comb. nov.
E878A6A3237858CEAE6A8360E5636DD8
urn:lsid:ipni.org:names:77201018-1
Coleus grandis L.H.Cramer, Kew Bull. 32: 556. 1978.
Plectranthus grandis (L.H.Cramer) R.H.Willemse, Blumea 25: 509. 1979.
Plectranthus barbatus var. grandis (L.H.Cramer) Lukhoba & A.J.Paton, Kew Bull. 58: 915. 2003. publ. 2004. Type: Sri Lanka, Sita Eleiya, Cramer 3869 (holotype: PDA; isotypes: K, US).
Coleus kilimandschari Gürke, Abh. Königl. Akad. Wiss. Berlin 1891: 359. 1892.
Plectranthus kilimandschari (Gürke) H.I.Maass in R.Mansfeld, Verz. Landwirtsch. Gärtn. Kulturpfl. 3: 1136. 1986. Types: Tanzania, Kilimanjaro, Marangu, Meyer 377 (syntype: B, destroyed) & forest nearby, 2400 m, Meyer 116 (syntype: B, destroyed). Marangu, Volkens 427 (neotype, designated by Paton et al. (2009): K; isoneotype: G).
Distribution.
NE. & E. Trop. Africa to DRC.
Coleus bariensis
(Ryding) A.J.Paton comb. nov.
92399E0187595E0DB39288E1841E2E98
urn:lsid:ipni.org:names:77201019-1
Plectranthus bariensis Ryding, Kew Bull. 54: 123. 1999. Type: Somalia, Bari Region, Escarpment of Bunder Murrayha, Buraha Dhaxi, Thulin & Warfa 5286 (holotype: UPS).
Distribution.
N. Somalia.
Coleus batesii
(Baker) A.J.Paton comb. nov.
BD309928DFFF5422B0CDBC3B24D3BDED
urn:lsid:ipni.org:names:77201020-1
Pycnostachys batesii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 386. 1900. Type: Cameroon, Efulen, Bates 372 (holotype: K.; isotype: BM).
Distribution.
Cameroon to SW. Uganda.
Coleus batianoffii
(P.I.Forst.) P.I.Forst. comb. nov.
E9555CAC4FE85C80BD4CBF70E42211E5
urn:lsid:ipni.org:names:77201021-1
Plectranthus batianoffii P.I.Forst., Austrobaileya 7: 707. 2008. Type: Australia, Queensland, Cook District, Palfrey Island, 23 July 1990, G.N.Batianoff 12118 (holotype: BRI).
Distribution.
Australia: Queensland.
Coleus bellus
(P.I.Forst.) P.I.Forst. comb. nov.
06A284A032415856B3E925F135E6EB65
urn:lsid:ipni.org:names:77201402-1
Plectranthus bellus P.I.Forst., Austrobaileya 8: 388. 2011. Type: Australia, Queensland, Cook District, Daintree National Park, Adeline Creek headwaters, 18 May 1999, P.I.Forster PIF24610 & R.Booth (holotype: BRI; isotype, MEL).
Distribution.
Australia: Queensland.
Coleus betonicifolius
(Baker) A.J.Paton comb. nov.
07D7C747F48F5F44B0DC406999B4B9DC
urn:lsid:ipni.org:names:77201022-1
Plectranthus betonicifolius Baker, Bull. Misc. Inform. Kew 1895: 72. 1895. Types: Zambia, Fwambo, Carson 64 & 79 (syntypes: K).
Elsholtzia carsonii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 450. 1900. Type: Zambia, Fwambo, S of Lake Tanganyika, 1890, Carson s.n. (holotype: K).
Coleus baumii Gürke in O. Warburg (ed.), Kunene-Sambesi Exped.: 357. 1903. Type: Angola, Cuito, Baum 544 (holotype: B, destroyed ; isotypes: G, K, W).
Coleus kasomenensis De Wild., Repert. Spec. Nov. Regni Veg. 11: 514. 1913. Types: DRC, Upper Katanga, Kasomenia, Kassner 2555 (syntype: BR; isosyntype: K) & near Lubumbashi (Elizsabethville), Homblé 246 (syntype: BR) & Welgelegen, Bequaert 382 & 564 (syntypes: BR).
Distribution.
Tanzania, DRC, Zambia and Angola.
Notes.
The West African C. lyratus (A.Chev.) Roberty has been removed from synonymy and recognized here as a separate species, differing from Paton et al. (2013).
Coleus bifidus
(A.J.Paton) A.J.Paton comb. nov.
180A6BB5B85C5744849E5D345B16CB86
urn:lsid:ipni.org:names:77201023-1
Plectranthus bifidus A.J.Paton, Fl. Zambes. 8(8): 235. 2013. Type: Zambia, Solwezi, Apr. 1960, Robinson 3520 (holotype: K; isotype: SRGH).
Distribution.
Zambia.
Coleus bishopianus
(Gamble) Smitha & A.J.Paton comb. nov.
82583CD31FFB5BE694526C19B9C2F285
urn:lsid:ipni.org:names:77201025-1
Plectranthus bishopianus Gamble, Bull. Misc. Inform. Kew 265. 1924. Type: India. Tamil Nadu: Pulneys, shola forest at Pillar Rocks, 20 May 1901, A.G. Bourne and E.G. Bourne 1398 (lectotype: K, designated by Matthew (1993)).
Distribution.
S. India.
Coleus bipartitus
(P.I.Forst.) P.I.Forst. comb. nov.
8B04245AEAC85415A52D20D1775947C6
urn:lsid:ipni.org:names:77201026-1
Plectranthus bipartitus P.I.Forst., Austrobaileya 9: 208. 2014. Type: Australia, Queensland, Cook District, Hann Tableland National Park, west of Mareeba, 8 April 2013, P.I.Forster PIF39595 (holotype: BRI; isotypes: CNS, K, MEL).
Distribution.
Australia: Queensland.
Coleus blakei
(P.I.Forst.) P.I.Forst. comb. nov.
EF2A5A6B074153B38E3771B463906444
urn:lsid:ipni.org:names:77201027-1
Plectranthus blakei P.I.Forst., Muelleria 7: 417. 1992. Type: Australia, Queensland, Leichardt District, Mimosa Creek, Blackdown Tableland National Park, 16 May 1984, P.I.Forster PIF1782 (holotype: BRI; isotypes: K, MEL, NSW).
Distribution.
Australia: Queensland.
Coleus bojeri
Benth., Labiat. Gen. Spec.: 52. 1832
6720DA33BF6B54D1A089DF1E92CE22BB
Solenostemon bojeri (Benth.) Guillaumet & A.Cornet, Adansonia, n.s., 15: 525. 1976.
Calchas bojeri (Benth.) P.V.Heath, Calyx 5: 160. 1997.
Plectranthus bojeri (Benth.) Hedge, Fl. Madag. 175: 218. 1998. Type: Madagascar, Emirna, Betani-mena, Bojer s.n. (lectotype: P, designated by Hedge et al. 1998).
Mitsa maculata Chapel. ex Benth., Labiat. Gen. Spec.: 52. 1832. nom. inval.
Plectranthus maculatus Bojer ex Benth., Labiat. Gen. Spec.: 52. 1832. nom. inval.
Coleus latifolius Hochst. ex Benth. in A.P.de Candolle, Prodr. 12: 74. 1848.
Majana latifolia (Hochst. ex Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Solenostemon latifolius (Hochst. ex Benth.) J.K.Morton, J. Linn. Soc., Bot. 58: 271. 1962. Types: Ethiopia, Gapdia, 17 Sept. 1838, Schimper 825 (syntypes: BM, E, K, G, GH, LG, STU, W) & Gafta, 18 Sept. 1838, Schimper 1228 (syntypes: BM, K, G, LG, W).
Coleus latifolius var. elatior Vatke, Linnaea 37: 322. 1872. Type: Ethiopia, Tigray, NNW of Axum, Bellitschen, 8 Sept. 1862, Schimper 378 (syntype: BM, JE).
Coleus micranthus Maxim., Trudy Imp. S.-Peterburgsk. Bot. Sada 3(1): 108. 1875. Type: seed from Ethiopia, 1872, Schimper (holotype: LE).
Plectranthus cymosus Baker, J. Linn. Soc., Bot. 21: 434. 1885.
Solenostemon cymosus (Baker) Guillaumet & A.Cornet, Adansonia, n.s., 15: 524. 1976., nom. illeg. Type: Central Madagascar, Baron 2250 (holotype: K; isotype: P).
Coleus bernieri Briq., Bull. Herb. Boissier 2: 128. 1894.
Solenostemon bernieri (Briq.) Guillaumet & A.Cornet, Adansonia, n.s., 15: 525. 1976.
Calchas bernieri (Briq.) P.V.Heath, Calyx 5: 160. 1997. Type: Madagascar, North of Madagascar, Bernier 147 (holotype: G).
Coleus gracilifolius Briq., Bull. Herb. Boissier 2: 127. 1894.
Solenostemon gracilifolius (Briq.) Guillaumet & A.Cornet, Adansonia, n.s., 15: 525. 1976. Type: Madagascar, Fito (Ambanivoulu), Goudot s.n. 1832 (holotype: G).
Coleus schweinfurthii Briq., Bot. Jahrb. Syst. 19: 181. 1894., nom. illeg., non C. schweinfurthii Vatke.
Coleus briquetii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 441. 1900., nom. superfl.
Calchas briquetii (Baker) P.V.Heath, Calyx 5: 160. 1997. nom. illeg.
Plectranthus bongensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 410. 1900. Type: Sudan, Bongo, Jur (Gir), Schweinfurth 2490 (holotype: K; isotype: PRE).
Coleus dewevrei Briq., Bull. Soc. Roy. Bot. Belgique 37: 71. 1899.
Calchas dewevrei (Briq.) P.V.Heath, Calyx 6: 51. 1999. Type: DRC, Dewèvre 1092a (holotype: BR).
Coleus latifolius var. madiensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 437. 1900. Type: Uganda, West Nile District: Madi, Grant in Speke & Grant s.n. (holotype: K).
Coleus heterotrichus Briq., Bull. Soc. Roy. Bot. Belgique 40: 40. 1901. Type: DRC, Bolobo, Demeuse 1155 (holotype: G; isotype: Z).
Coleus copiosiflorus Briq., Mém. Soc. Bot. France 8: 288. 1917.
Calchas copiosiflorus (Briq.) P.V.Heath, Calyx 5: 160. 1997. Types: Republic Central Africa, Haute Ombella, Kaga Do, 26 Oct. 1902, A. Chevalier 5967 (syntype: P); Mali, Koulikoro, 6–14 Oct. 1899, A. Chevalier s.n. (syntype, P).
Coleus phymatodes Briq., Mém. Soc. Bot. France 8: 289. 1917. Type: Mali, Koulikoro, 6–14 Oct 1899, A. Chevalier s.n. (holotype, P).
Coleus claessensii De Wild., Bol. Soc. Ibér. Ci. Nat. 19: 117. 1920.
Calchas claessensii (De Wild.) P.V.Heath, Calyx 5: 160. 1997. Types: DRC, Mobwassa, Claessens 741 (syntype: BR) & Ekuta on the Lua, Sapin s.n. (syntype: BR).
Coleus homblei De Wild., Contr. Fl. Katanga: 174. 1921. Types: DRC, Lubumbashi (Elisabethville), Homblé 210 (syntype: BR) & Tshisinka, Homblé 1259 (syntype: BR.).
Coleus ringoetii De Wild., Contr. Fl. Katanga: 174. 1921. Type: DRC, Shinsenda, Ringoet 546 (holotype: BR).
Coleus termetophilus De Wild., Contr. Fl. Katanga: 175. 1921. Type: DRC, Lubumbashi (Elisabethville), Homblé 220 (holotype: BR; isotype: Z).
Coleus platostomoides Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 362. 1928.
Solenostemon platostomoides (Robyns & Lebrun) Troupin, Bull. Jard. Bot. Natl. Belg. 55: 299. 1985.
Calchas platostomoides (Robyns & Lebrun) P.V.Heath, Calyx 6: 51. 1999. Type: DRC, Lake Kivu, Saké Bay, Kuteruzi, Robyns 2482 (holotype: BR; isotype: K).
Coleus quarrei Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 3. 1928. Type: DRC, Lubumbashi (Elisabethville), Kibembe Farm, Quarré 608 (holotype: BR).
Coleus collinus Lebrun & L.Touss., Bull. Jard. Bot. État Bruxelles 17: 81. 1943.
Solenostemon collinus (Lebrun & L.Touss.) Troupin, Bull. Jard. Bot. Natl. Belg. 55: 299. 1985.
Calchas collinus (Lebrun & L.Touss.) P.V.Heath, Calyx 5: 160. 1997. Type: Congo, Kivu, Rutshuru, Nov. 1937, Lebrun 8232 (holotype: BR).
Distribution.
Eastern, Central & Southern Africa, Madagascar.
Coleus botryosus
(A.J.Paton) A.J.Paton comb. nov.
0ED09930E15C522FB5095E2705A2679D
urn:lsid:ipni.org:names:77201028-1
Plectranthus botryosus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 335. 2009. Type: Tanzania, Songea District: just E of R. Gembabili ± 3 km SW of Kitai, Milne-Redhead & Taylor 9735 (holotype:K, sheet 2; isotypes EA, K sheet 1).
Distribution.
S. Tanzania.
Coleus bourneae
(Gamble) Smitha & A.J.Paton comb. nov.
3A95EF3A890957CA809594B273F760C2
urn:lsid:ipni.org:names:77201029-1
Plectranthus bourneae Gamble, Bull. Misc. Inform. Kew 1924: 264. 1924. Type: India. Tamil Nadu: Pulneys, Coelogyne rock, Poombarai road 16 May 1901, A.G. Bourne & E.G. Bourne 2028 (lectotype: K, designated by Smitha and Sunojkumar 2018).
Distribution.
S. India.
Coleus bracteatus
Dunn, Notes Roy. Bot. Gard. Edinburgh 8: 158. 1913
372C6C047F805D8FB1B7002D2EF7A263
Plectranthus bracteatus (Dunn) Suddee, Kew Bull. 59: 399. 2004. Type: China, Yunnan, Puerh, 1350 m, Henry 13498 (lectotype: K, designated by Suddee et al. 2004; isolectotype: NY).
Plectranthus saphinensis Phuong, Bot. Zhurn. (Moscow & Leningrad) 67: 1131. 1982. Type: Vietnam, Sa Phin, Prov. Ha Tuyen, 15 Sept. 1977, Phuong 294B (holotype: HN; isotype: HN).
Distribution.
China: Yunnan to Indo-China.
Coleus brazzavillensis
A.Chev., Veg. Ut. Afr. Trop. Franç. 1: 124. 1905
F704D6AF54785946AE429782510F1C65
Coleus brazzavillensis A.Chev., Veg. Ut. Afr. Trop. Franç. 1: 124. 1905. Type: Congo, Brazzaville, Mission du Chari-Lac Chad, Chevalier 11154 (holotype: P).
Distribution.
Republic of Congo (Brazzaville).
Coleus buchananii
(Baker) Brenan, Mem. New York Bot. Gard. 9: 43. 1954
1C61C2F2F05152CFB4E004879409A3AC
Plectranthus buchananii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 402. 1900. Type: Malawi, Shire Highlands, Nakajumbu, Buchanan 365 (holotype: K).
Coleus scaposus C.H.Wright, Bull. Misc. Inform. Kew 1906: 167. 1906. Type: Malawi, Namasi, Cameron 60 (holotype: K).
Coleus hockii De Wild., Repert. Spec. Nov. Regni Veg. 11: 514. 1913. Type: DRC, Upper Katanga, valley of small Luembe, Hock s.n. (holotype: BR).
Distribution.
Tanzania to S. Trop. Africa.
Coleus burorum
Chiov., Fl. Somala 1: 278. 1929
05337C56FC0C5830B41CB5F6EE851C7F
Plectranthus burorum (Chiov.) J.K.Morton, Novon 8: 265. 1998. Type: Somalia, Somalia Meridionale: Boscaglia di Bur Budulca, 16 Mar.1924, N.Puccioni & G. Stefanini 287 (323) (holotype: FT).
Coleus cicatricosus Hutch. & E.A.Bruce, Bull. Misc. Inform. Kew 1941: 180. 1941.
Plectranthus cicatricosus (Hutch. & E.A.Bruce) J.R.I.Wood, Handb. Yemen Fl.: 249. 1997. Type: Somalia, Buramo, 29 Jan. 1933, J.B.Gillett 4880 (holotype: K; isotype: FT).
Distribution.
Ethiopia to Somalia, Yemen.
Coleus caillei
A.Chev. ex Hutch. & Dalziel, Fl. W. Trop. Afr. 2: 291. 1931
9E743C948B1658ED986CF651885063C1
Leocus caillei (A.Chev. ex Hutch. & Dalziel) J.K.Morton, J. Linn. Soc., Bot. 58: 270. 1962. Plectranthus caillei (A.Chev. ex Hutch. & Dalziel) B.J.Pollard, Kew Bull. 64: 260. 2009. Type: Guinea, Longuery, Caille in Herb Chevalier 14684 (holotype: P).
Distribution.
W. Trop. Africa to Cameroon.
Coleus calaminthoides
(Baker) A.J.Paton comb. nov.
2655747311655931BC1ACF1D8C49BE2F
urn:lsid:ipni.org:names:77201030-1
Solenostemon calaminthoides Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 421. 1900. Type: Gabon, Gabon River, Mann (holotype, K).
Distribution.
Cameroon, Gabon, DRC.
Coleus calcicola
Murata, Acta Phytotax. Geobot. 24: 105. 1970
171C3785F94751A8B7866BAEECC24095
Plectranthus chiangdaoensis Suddee, Kew Bull. 59: 404. 2004., non Plectranthus calcicola Hand.-Mazz. Type: Thailand, Chiang Mai, Doi Chiang Dao, 1500 m, 15 Sept. 1967, Shimizu & Hutoh T-10153 (holotype: KYO; isotype: BKF, KYO, TI).
Distribution.
N. Thailand (Doi Chiang Dao).
Coleus caldericola
(P.I.Forst.) P.I.Forst. comb. nov.
FA2AD3B2D94550C5A71EE4E0259C941A
urn:lsid:ipni.org:names:77201031-1
Plectranthus caldericola P.I.Forst., Austrobaileya 8: 392. 2011. Type: Australia, New South Wales, The Pinnacle, 13 km W of Mt. Warning, 6 Apr. 1994, P.I. Forster & G. Leiper PIF15093 (holotype: BRI; isotypes: MEL, NSW).
Distribution.
Australia: New South Wales.
Coleus calycinus
(Benth.) A.J.Paton comb. nov.
39211FE122C652EDAF707C982DE5FC10
urn:lsid:ipni.org:names:77201032-1
Plectranthus calycinus Benth. in E.H.F.Meyer, Comm. Pl. Afr. Austr.: 230. 1838.
Rabdosia calycina (Benth.) Codd, Bothalia 11: 117. 1973.
Rabdosiella calycina (Benth.) Codd, Bothalia 15: 10. 1984.
Isodon calycinus (Benth.) H.W.Li, J. Arnold Arbor. 69: 293. 1988. Type: South Africa, Eastern Cape, between St Johns and Umsikaba rivers, 1837, Drège 3584 (lectotype: K designated by Codd (1975); isolectotypes: MO, P).
Plectranthus pyramidatus Gürke, Bull. Herb. Boissier 6: 552. 1898. Type: South Africa, Mpumalanga, Houtbosch, s.d., Rehmann 6179 (holotype: Z).
Plectranthus pachystachyus Briq., Bull. Herb. Boissier, sér. 2, 3: 1003. 1903.
Plectranthus calycinus var. pachystachyus (Briq.) T.Cooke in W.H.Harvey & auct. suc. (eds.), Fl. Cap. 5(1): 271. 1910. Type: South Africa, KwaZulu-Natal, Umkomaas, iii.1892, Medley Wood 4621 (holotype: G; isotype: K).
Distribution.
S. Mozambique to South Africa.
Coleus cambodianus
(Murata) A.J.Paton comb. nov.
7100517EFCCE55ABBD49F5EA3E0342F4
urn:lsid:ipni.org:names:77201033-1
Anisochilus cambodianus Murata, Acta Phytotax. Geobot. 28: 26. 1977. Type: Cambodia, Kampot Province, Poporkvil, c. 1000 m alt, 4 Dec. 1964, Kira, Hozumi, Yoda & Kokawa 292 (holotype: KYO).
Distribution.
SE. Thailand to Cambodia.
Coleus caninus
(Roth) Vatke, Linnaea 37: 318. 1872
55D51E895FC157F39EE833A49306C890
Plectranthus caninus Roth, Nov. Pl. Sp.: 279. 1821.
Majana canina (Roth) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: India, Heyne s.n. in Wallich Cat. 2729C (holotype: B, destroyed; isotypes: BM, K(000820142), K-W (K001116966), P).
Distribution.
Eritrea to South Africa, India to Myanmar.
Coleus caninus subsp. caninus
958378A0302C5130971E1B1C50E83921
Plectranthus caninus subsp. caninus , A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 345. 2009. Type: as above.
Ocimum monadelphum R.Br. ex Roth, Nov. Pl. Sp.: 267. 1821.
Coleus heynei Benth., Labiat. Gen. Spec.: 50. 1832. Type: India, Heyne s.n. (holotype: B, destroyed; neotype,: India, Herb. Ham. in Wall. Cat. 2729B (neotype: K-W (K001116965), designated by Paton et al. 2009; isoneotype: K).
Coleus spicatus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 15. 1830.
Majana spicata (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: India, Herb. Wight in Wall. Cat. 2729A (syntypes: BM, K (K000820139), K (K000820141), K-W(K001116964), G-DC).
Plectranthus monadelphus Roxb., Fl. Ind. ed. 1832, 3: 22. 1832. Type: Roxburgh’s illustration No. 1459 (lectotype: K, designated by Suddee et al. 2004).
Coleus spicatus var. rondinella Spreng., Gartenflora 45: 358. fig. 62. 1896. Type: cultivated, material from Eritrea, illustration cited above (lectotype: illustration designated by Paton et al. 2009).
Coleus pachyphyllus Gürke, Bot. Jahrb. Syst. 38: 168. 1906. Types: Ethiopia, Galla Highlands, Ginea, Ellenbeck 1951 (syntype: B, destroyed) & Korkora in Borana, Ellenbeck 2233 (syntype: B, destroyed); neotype: Somalia, Juba R. north of Gelib, Mooney 7672 (neotype: K, designated by Ryding (2000)).
Distribution.
Eritrea to N. Tanzania, India to Myanmar.
Coleus caninus subsp. flavovirens
(Gürke) A.J.Paton comb. nov.
6EB6C1FBF7A054119ED4E89C2F3734E1
urn:lsid:ipni.org:names:77201034-1
Coleus flavovirens Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 347. 1895.
Plectranthus caninus subsp. flavovirens (Gürke) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 345. 2009. Type: Tanzania, Lake Chala (Dschallasee), 1893, Volkens 1771 (holotype: B destroyed; isotype: BR).
Coleus omahekensis Dinter, Repert. Spec. Nov. Regni Veg. Beih. 53: 123. 1928. Type: Namibia, Etemba and Otjikuara, Dinter 3265, two localities same number (syntypes: not seen.).
Distribution.
Ethiopia to South Africa.
Coleus capitatus
A.J.Paton nom. nov.
A3BCA52620A65CADBDACB3FF7E66C57C
urn:lsid:ipni.org:names:77201035-1
Pycnostachys parvifolia Baker, Bull. Misc. Inform. Kew 1895: 72. 1895., non Coleus parviflorus Benth. Type: Zambia, Northern Province, Fwambo, Carson 103 (lectotype: K), designated by Bruce (1940).
Distribution.
W. & S. Tanzania to Zambia.
Coleus carnosifolius
(Hemsl.) Dunn, Notes Roy. Bot. Gard. Edinburgh 8: 158. 1913
FD068B26C6F2543884C10261F88E70D4
Plectranthus carnosifolius Hemsl., J. Linn. Soc., Bot. 26: 270. 1890. Type: China, Guangdong Prov., Aug. 1887, C. Ford 107 (holotype: K).
Distribution.
SE. China.
Coleus cataractarum
(B.J.Pollard) A.J.Paton comb. nov.
096AB5E0BA9751D090F7A459D5706E1F
urn:lsid:ipni.org:names:77201036-1
Plectranthus cataractarum B.J.Pollard, Kew Bull. 56: 976. 2001. Type: Mt Cameroon, 25 Nov. 1993, Cheek 5563 (holotype: K; isotypes MA, MO, SCA, WAG, YA).
Distribution.
Bioko, Cameroon.
Coleus caudatus
(S.Moore) E.Downes & I.Darbysh. Blumea 62: 172. 2017
4A53C8CD57C55A09B85771BE3652AD43
Plectranthus caudatus S.Moore, J. Linn. Soc., Bot. 40: 176. 1911. Type: Zimbabwe, Chimanimani, 26 Nov. 1906, Swynnerton 2010 (holotype: BM; isotype: K).
Distribution.
Mozambique and Zimbabwe.
Coleus celsus
A.J.Paton nom. nov.
A6E315F9B4D2554B9C721EC9A3A18129
urn:lsid:ipni.org:names:77201037-1
Solenostemon robustus Hiern, Cat. Afr. Pl. 1: 864. 1900., non Coleus robustus (Hook.f.) A.J.Paton.
Holostylon robustum (Hiern) G.Taylor, J. Bot. 69(suppl. 2): 161. 1931.
Plectranthus robustus (Hiern) A.J.Paton in Fl. Zambesiaca 8,8: 234. 2013. Type: Angola, Pungo Andongo, from Lombe to Condo, March 1857, Welwitsch 5538 (holotype: LISU; isotypes: BM, G).
Distribution.
Angola.
Coleus centraliafricanus
A.J.Paton nom. nov.
4832340EA6655E74900C3F2D05D26E65
urn:lsid:ipni.org:names:77201038-1
Pycnostachys chevalieri Briq., Mém. Soc. Bot. France 8: 193. 1912., non Coleus chevalieri Briq. Type: Central African Republic, Dar Banda du Kaga Djé au Kaga Pongouru, A.Chevalier 6561 (holotype: P).
Distribution.
Central African Republic.
Coleus chevalieri
Briq., Mém. Soc. Bot. France 8: 287. 1917
DBFD322069A856E78E69DEBABF1F63CC
Coleus chevalieri Briq., Mém. Soc. Bot. France 8: 287. 1917. Type: Central African Republic, Krebedjén (Fort Sibut), vallée de la moyenne Tomi, 8 Oct.1902, Chevalier 5662 (holotype: P).
Distribution.
Central African Republic.
Coleus ciliatus
(Bramley) A.J.Paton comb. nov.
58CE1E4E8F935B5593888B58D9DE7383
urn:lsid:ipni.org:names:77201039-1
Pycnostachys ciliata Bramley, Kew Bull. 60: 587. 2005 publ. 2006. Type: Tanzania, Mbeya Dist., Songwe Valley, c. 2 km N of Mbeya–Tunduma road, fl. 25 Mar, 1988, Bidgood, Mwasumbi & Vollesen 693 (holotype: K; isotypes: C, DSM, NHT).
Distribution.
Tanzania to S. Trop. Africa.
Coleus circinnatus
(Hedge) A.J.Paton & Phillipson comb. nov.
DE971F71FBE15E4A91AD3565270FC902
urn:lsid:ipni.org:names:77201041-1
Plectranthus circinnatus Hedge, Fl. Madag. 175: 228. 1998. Type: Madagascar, Prov. de Fianarantsoa, Massive d’Itremo, sur RN 35, 42 Km W. of Ambatofinandrahana, Clement, Phillipson & Rafamantanantsoa 2045 (holotype: E; isotypes MO, P).
Distribution.
Madagascar.
Coleus coeruleus
Gürke, Bot. Jahrb. Syst. 19: 217. 1894
480C6863ABF7585AB9F77AE5E80DCE78
Calchas coeruleus (Gürke) P.V.Heath, Calyx 5: 160. 1997.
Plectranthus coeruleus (Gürke) Agnew, Upland Kenya Wild Fl.: 638. 1974. Type: Tanzania, Lushoto District: Kwa Mshusa, Holst 8895 (holotype: B, destroyed; isotypes: G, K, Z).
Distribution.
Kenya to N. Malawi.
Coleus comosus
Hochst. ex Gürke, Bot. Jahrb. Syst. 19: 212. 1894
8F58D936CAE8524DABAE12638E0AD341
Plectranthus ornatus Codd, Bothalia 11: 393. 1975., non Plectranthus comosus Sims. Type: Ethiopia, Dschenausa, 1 June 1840, Schimper 1328 (holotype: B, destroyed; isotypes: BM, FI, G, P).
Distribution.
NE and E. Africa, naturalized in South Africa.
Coleus confertiflorus
(A.J.Paton) A.J.Paton comb. nov.
142FF14BAF895192A6E094584FF3272F
urn:lsid:ipni.org:names:77201042-1
Plectranthus confertiflorus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 303. 2009. Type: Tanzania, Pare District: Mkomazi Game Reserve, Kisima Hill, Abdallah, Mboya & Vollesen 96/129 (holotype: K; isotypes: C, NHT, P).
Distribution.
N. Tanzania.
Coleus congensis
(Gürke) A.J.Paton comb. nov.
F281CE61DB5056AA87A4F650517DB8C7
urn:lsid:ipni.org:names:77201043-1
Pycnostachys congensis Gürke, Bull. Herb. Boissier 4: 819. 1896. Types: DRC, Samba, Deschamps 28 (syntype: B); Lusambo (syntype: B).
Distribution.
DRC, Zambia.
Coleus congestus
(R.Br.) A.J.Paton comb. nov.
A91F48E4D45C5789956E59247975B8B6
urn:lsid:ipni.org:names:77201044-1
Plectranthus congestus R.Br., Prodr. Fl. Nov. Holl.: 506. 1810. Type: Australia, Queensland, Endeavour River, 1770, Banks & Solander (holotype: BM).
Distribution.
N. Australia, Lesser Sunda Is., New Guinea.
Coleus conglomeratus
(T.C.E.Fr.) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 105. 1929
2AF18428B8555321B9D5F218CB76A0A5
Englerastrum conglomeratum T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 72. 1924.
Plectranthus conglomeratus (T.C.E.Fr.) Hutch. & Dandy, Bull. Misc. Inform. Kew 1926:481. 1926. Type: Togo (Benin), Sokode, Kersting 93 (holotype: B, destroyed; fragment K).
Plectranthus harrisii J.K.Morton, J. Linn. Soc., Bot. 58: 267. 1962. Type: Ghana, between Toga Mt. and Maliato, Togo Plateau, 24 Dec. 1959, J.K.Morton A3830 (holotype: GC; isotypes: K, WAG). synon. nov.
Distribution.
Tropical West Africa.
Coleus crassus
(N.E.Br.) Culham comb. nov.
588D33AAB9C356D08B1C5D7BBF8D0655
urn:lsid:ipni.org:names:77201045-1
Plectranthus crassus N.E.Br., Gard. Chron., ser. 3, 35: 21. 1904. Type: Plant cultivated at Kew in 1899 from material collected by Mahon 393 (holotype: K).
Distribution.
Malawi (Mt. Mulanje).
Coleus cremnus
(B.J.Conn) A.J.Paton comb. nov.
49BA082BC63B518F9A2030F13BD71A6D
urn:lsid:ipni.org:names:77201046-1
Plectranthus cremnus B.J.Conn, Telopea 4: 643. 1992. Type: Australia, New South Wales, Sawtell, 20 Feb. 1990, B.J. Conn 3478 (holotype: NSW; isotype: MEL).
Distribution.
Australia, New South Wales.
Coleus cucullatus
(A.J.Paton) A.J.Paton comb. nov.
0AB5EC2C9B3353BBA188E5AAAA24BB49
urn:lsid:ipni.org:names:77201047-1
Plectranthus cucullatus A.J.Paton – Fl. Zambes. 8(8): 261. 2013. Type: Mozambique, Ribáuè Dist., Ribáuè mountains, Oct. 1931, Gomes & Sousa 754 (holotype: K).
Distribution.
Mozambique.
Coleus cuneatus
Baker f., J. Bot. 1899: 64. 1899
85A96E683BE151D7A1C97FB0F530C5D2
Plectranthus cuneatus (Baker f.) Ryding, Kew Bull. 54: 124. 1999. Type: Somalia, Woqooyi Galbeed/ Togdher Regions, Wagga Mt, Lort-Phillips s.n. (holotype: BM).
Distribution.
S. Djibouti to N. Somalia.
Coleus cyanophyllus
(P.I.Forst.) P.I.Forst. comb. nov.
D64AFEB8C2C553FD97B40976CD9C48EE
urn:lsid:ipni.org:names:77201048-1
Plectranthus cyanophyllus P.I.Forst., Austrobaileya 4: 166. 1994. Type: Australia, Queensland, North Kennedy District. Mt Abbot, 50 km W of Bowen, 27 Oct. 1992, A.R.Bean 5188 (holotype: BRI; isotypes: CNS, K, MEL).
Distribution.
Australia: Queensland.
Coleus cylindraceus
(Hochst. ex Benth.) A.J.Paton comb. nov.
4972B3B7929350BCB4A7E284BD43A776
urn:lsid:ipni.org:names:77201050-1
Plectranthus cylindraceus Hochst. ex Benth. in A.P.de Candolle, Prodr. 12: 60. 1848.
Germanea cylindracea (Hochst. ex Benth.) Hiern, Cat. Afr. Pl. 1: 861. 1900.
Burnatastrum cylindraceum (Hochst. ex Benth.) P.V.Heath, Calyx 4: 175. 2001. Type: Ethiopia, near Gapdia, 29 Nov.1838, Schimper 1113 (holotype: K; isotypes: BM, G, P,W).
Plectranthus montanus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 17. 1830., non Coleus montanus Hochst. ex Ces. Type: India, Deccan Peninsula, exact locality unknown (“Peninsula India Orientalis”), Herb. Wight in Wall. Cat. 2747B (lectotype: K ((K000820120), designated by Suddee et al. (2004); isolectotype: K-W (K001117007).
Plectranthus marrubioides Hochst. ex Benth. in A.P.de Candolle, Prodr. 12: 60. 1848. Type: Ethiopia, near Jaja (Dschadscha), Jan. 1844, Schimper 1925 (holotype: K; isotypes: BM,G, K, P, W).
Geniosporum lasiostachyum Briq., Bot. Jahrb. Syst. 19: 164. 1894. Type: Angola, Huíla, Mar. 1879, Welwitsch 5489 (syntypes: K, BM, LISC).
Plectranthus fischeri Gürke, Bot. Jahrb. Syst. 19: 200. 1894. Types: Tanzania, Kilimanjaro (Kilimandcharo), Ugweno, Volkens 518 (syntype: B, destroyed); Masai Highlands (Massaihochland), Fischer ser.1 77 (syntype: B, destroyed; isosyntype: HBG) & Fischer ser.2 501 (syntype: B, destroyed).
Plectranthus moschosmoides Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 414. 1900. Type: Angola, Huíla, Mar.1879, Welwitsch 5489 (holotype: BM; isotypes: K, LISC).
Geniosporum fissum S.Moore, J. Bot. 39: 263. 1901. Type: Kenya, Northern Frontier District: Dadaro, 1110 m, Delamere s.n. (holotype: BM).
Plectranthus densiflorus T.Cooke, Bull. Misc. Inform. Kew 1909: 378. 1909. Type: South Africa, KwaZulu-Natal, near the Mooi R., Medley-Wood 4475 (holotype: K; isotypes: GRA, NH, SAM).
Plectranthus villosus T.Cooke, Bull. Misc. Inform. Kew 1909: 378. 1909. Type: South Africa, KwaZulu-Natal, Entumeni, Medley-Wood 3955 (holotype: K; isotype NH).
Plectranthus glomeratus R.A.Dyer, Fl. Pl. South Africa 24: t. 946. 1944. nom. superfl. Based on the above.
Plectranthus spiciformis R.A.Dyer, Fl. Pl. South Africa 24: t. 946. 1944. Type: South Africa, Gauteng, Hammanskraal, Mogg 27138 (holotype: PRE).
Coleus subbaraoi Kumari & Malathi, Fl. Visakhapatnam Distr. 2: 68. 2008. Type: India, Andhra Pradesh, Visakhapatnam District, Kappakonda, Subba Rao 29637 (holotype: CAL; isotype: MH).
Distribution.
Trop. & South Africa, Arabian Peninsula, S. India
Coleus daviesii
E.A.Bruce, Bull. Misc. Inform. Kew 1933: 477. 1933
B9746B8F1DA75C3DA78A8C0E97120CE6
Plectranthus daviesii (E.A.Bruce) B.Mathew, Kew Bull. 31: 174. 1976. Type: Tanzania, Rungwe, R.M. Davies R4 (holotype: K; isotype: SRGH).
Distribution.
SW. Tanzania to N. Malawi.
Coleus decimus
(A.J.Paton) A.J.Paton comb. nov.
B506598FF6335B12847D055381077A5A
urn:lsid:ipni.org:names:77201051-1
Plectranthus decimus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 324. 2009. Type: Zambia, Mbala (Abercorn) District: Kawimbe, Richards 7288 (holotype: K).
Distribution.
W. Tanzania, Zambia to Angola.
Coleus decurrens
Gürke, Bot. Jahrb. Syst. 19: 215. 1894
33E9FFC428FE524380A115BD22BABE7C
Plectranthus decurrens (Gürke) J.K.Morton, J. Linn. Soc., Bot. 58: 267. 1962. Type: Cameroon, Buea, Preuss 948 (holotype: B, destroyed; isotypes: COI, HBG).
Coleus elatus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 427. 1900. Types: Bioko (Fernando Po), Mann 584 (syntype: K); Gabon, Crystal Mts, Mann s.n. 1661 (synype: K).
Coleus toroensis S.Moore, J. Bot. 45: 95. 1907. Type: Uganda, Toro District: Dura (Durro) R., Bagshawe 1043 (holotype: BM).
Coleus variifolius De Wild., Bol. Soc. Ibér. Ci. Nat. 19: 124. 1920. Type: DRC, between Buta and Bima, Seret 96 (lectotype: BR, designated by Paton et al. 2009).
Distribution.
Nigeria to Uganda and Angola.
Coleus deflexifolius
(Baker) A.J.Paton comb. nov.
B85CAB4059A85AD8B2FD0E2F9AB613DB
urn:lsid:ipni.org:names:77201052-1
Pycnostachys deflexifolia Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 381. 1900. Type: Kenya, Naivasha District: near Lake Elmenteita, Scott Elliot 6756 (holoype: K).
Distribution.
E. Trop. Africa.
Coleus defoliatus
(Hochst. ex Benth.) A.J.Paton comb. nov.
320AF1016F115895952D72C326A890AB
urn:lsid:ipni.org:names:77201053-1
Plectranthus defoliatus Hochst. ex Benth. in A.P.de Candolle, Prodr. 12: 60. 1848.
Isodictyophorus defoliatus (Hochst. ex Benth.) Agnew, Upland Kenya Wild Fl.: 640. 1974. Type: Ethiopia, Jomara (Dschomara), 26 Dec. 1839, Schimper 847 (holotype: K; isotypes BR, FI, G, MPU, TUB, P, W).
Coleus floribundus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 438. 1900. Type: Ethiopia, 1 Jan. 1853, Schimper 21 (holotype: K; isotype: BR, G).
Distribution.
Eritrea to S. Trop. Africa.
Coleus densus
(N.E.Br.) A.J.Paton comb. nov.
7A7C9B98C00653639F74EEA8F02506D1
urn:lsid:ipni.org:names:77201054-1
Plectranthus densus N.E.Br., Bull. Misc. Inform. Kew 1894: 12. 1894. Type: Tanzania, N of Lake Malawi (Nyassa), Thomson s.n. (holotype: K).
Plectranthus primulinus Baker, Bull. Misc. Inform. Kew 1895: 292. 1895.
Coleus primulinus (Baker) A.Chev., Rev. Int. Bot. Appl. Agric. Trop. 33: 340. 1953. Type: Zambia, Mwero Plateau, 1894, Carson 36 (holotype: K).
Distribution.
SW. Tanzania to S. Trop. Africa.
Coleus descampsii
(Briq.) A.J.Paton comb. nov.
B105996976755454A4E8A7A1B2F68656
urn:lsid:ipni.org:names:77201055-1
Pycnostachys descampsii Briq., Bull. Soc. Roy. Bot. Belgique 37: 63. 1899. Type: DRC, Katanga, Lufonzo (Lufogo) R., Mar. 1896, Descamps s.n. (holotype: BR).
Pycnostachys linifolia Gürke, Bot. Jahrb. Syst. 30: 397. 1901. Type: Tanzania, Njombe District: Ubena, near Liangira, Goetze 798 (holotype: B, destroyed; isotypes: BR, K, fragment).
Pycnostachys lavanduloides Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 68. 1921. Type: Tanzania, Iringa District, Uhehe, Udzungwa Mts (Utschungwe-Berge), 1600 m, Prince 28a (holotype: B, destroyed).
Pycnostachys pallide-caerulea Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 67. 1921. Type: Cameroon, Kumbo, Mt Bansso, Oct. 1909, Ledermann 5738 (holotype: B, destroyed, fragment K).
Distribution.
Cameroon to W. Tanzania.
Coleus dewildemanianus
(Robyns & Lebrun) A.J.Paton comb. nov.
470CABD8BA515CC192EAA589717BF9BE
urn:lsid:ipni.org:names:77201057-1
Pycnostachys dewildemaniana Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 352. 1928. Type: DRC, Katanga, Munama, Quarré 1143 (holotype: BR; isotype: K).
Distribution.
Tanzania to S. Trop. Africa.
Coleus dichotomus
(A.J.Paton) A.J.Paton comb. nov.
8B19A58B077959BA87D510E03113B1A4
urn:lsid:ipni.org:names:77201058-1
Plectranthus dichotomus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 317. 2009. Type: Tanzania, Kilosa District: Ukaguru, Mamiwa Forest Reserve, E of Ikwamba peak, Mabberley 1465 (holotype: K).
Distribution.
Tanzania (Ukaguru Mts.).
Coleus dinteri
(Briq.) A.J.Paton comb. nov.
B4E95EB2E5005CC8BB06A538552B2E54
urn:lsid:ipni.org:names:77201059-1
Plectranthus dinteri Briq., Bull. Herb. Boissier, sér. 2, 3: 1070. 1903. Type: Namibia, Waterberg, Dinter 336 (holotype: Z).
Distribution.
Namibia, South Africa: Limpopo (W. Waterberg).
Coleus dissitiflorus
Gürke, Bot. Jahrb. Syst. 19: 217. 1894
8D26032E0593504B9B3CF9AFF0DC62BE
Plectranthus dissitiflorus (Gürke) J.K.Morton, J. Linn. Soc., Bot. 58: 267. 1962. Type: Cameroon, between Buea and Mimbia, 9 Oct. 1891, Preuss 1055 (holotype: B, destroyed; isotype: COI).
Distribution.
WC. Trop. Africa.
Coleus divaricatus
A.J.Paton nom. nov.
CECF71B11E535E7AABD1524ECC52FB90
urn:lsid:ipni.org:names:77201060-1
Anisochilus paniculatus Benth. in A.P.de Candolle, Prodr. 12: 82. 1848, non Coleus paniculatus Benth. Type: Sri Lanka, unknown locality, Walker 52 (holotype: K).
Distribution.
S. India.
Coleus diversus
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
A2CECD00A26B539184FE9DA43E61EFF9
urn:lsid:ipni.org:names:77201061-1
Plectranthus diversus S.T.Blake, Contr. Queensland Herb. 9: 20. 1971. Type: Australia, Queensland, Spring Creek near Bowen (cultivated at The Gap), Brisbane., Oct. 1963, S.T.Blake 22128 (holotype: BRI).
Distribution.
Australia: Queensland.
Coleus dolichopodus
(Briq.) A.J.Paton comb. nov.
208B4B0C67DC592CBA4442A9BE09CC02
urn:lsid:ipni.org:names:77201062-1
Plectranthus dolichopodus Briq., Bull. Herb. Boissier, sér. 2, 3: 1069. 1903. Type: South Africa, KwaZulu-Natal, Karkloof, n.d., Rehmann 7383 (holotype: Z).
Distribution.
Malawi to South Africa.
Coleus dumicola
(P.I.Forst.) P.I.Forst. comb. nov.
2E1E5F9C0A2E5BB59A06BDDA07AA1B71
urn:lsid:ipni.org:names:77201063-1
Plectranthus dumicola P.I.Forst., Austrobaileya 4: 168. 1994. Type: Australia, Queensland, Cook District, 31 km along road to Leo Creek Mine, McIlwraith Range, 8 June 1992, P.I.Forster PIF10268 & M.C.Tucker & G. Kenning (holotype: BRI; isotypes: CNS, K, MEL).
Distribution.
Australia: Queensland.
Coleus dysophylloides
(Benth.) A.J.Paton comb. nov.
7FC25DFE906C582BA735FECB2B029B9E
urn:lsid:ipni.org:names:77201064-1
Anisochilus dysophylloides Benth. in N.Wallich, Pl. Asiat. Rar. 2: 19. 1830. Type: India, Madras, Nilghiri Hills, Herb. Wight in Wall. Cat. 2756 (lectotype: K ((K000674673)); isolectotypes: E, G-DC,K-W(K001117033), designated Suddee and Paton 2009).
Anisochilus sericeus Benth. in A.P.de Candolle, Prodr. 12: 82. 1848. Type: India, Deccan Peninsula, Courtallum, Herb. Wight 2515 (holotype: K: (K000674767)); isotype K (K000674768).
Anisochilus albidus Wight, Icon. Pl. Ind. Orient. 4: t. 1436. 1849. Type: Madras, Nilghiri Hills, About Coonoor and Kaitie, ex Herb. Wight, Wight Ic. t. 1436 (holotype: K (K000674776)).
Anisochilus purpureus Wight, Icon. Pl. Ind. Orient. 4: t. 1435. 1849.
Anisochilus dysophylloides var. purpureus (Wight) Gamble, Fl. Madras 2: 1128. 1924. Type: Madras, Nilghiri Hills, About Coonoor, Ex Herb. Wight Propr., Wight Ic. t. 1435 (holotype: K (K000674774)).
Distribution.
SW. India (Shervarayan Hills).
Coleus efoliatus
De Wild., Contr. Fl. Katanga: 173. 1921
CDE32CD6DE0750C6AF6CD4C40C2556EA
Plectranthus efoliatus (De Wild.) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 289. 2009. Type: DRC, Welgelegen, Bequaert 486 (lectotype: BR, designated by Paton et al. (2009)).
Plectranthus leviculus N.E.Br., Bull. Misc. Inform. Kew 1921: 296. 1921. Type: DRC, Lubumbashi (Elisabethville), F.A. Rogers 26211 (holotype: K).
Englerastrum hjalmarii T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 71. 1924.
Plectranthus hjalmarii (T.C.E.Fr.) Hutch. & Dandy, Bull. Misc. Inform. Kew 1926: 481. 1926.
Coleus hjalmarii (T.C.E.Fr.) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 106. 1929. Type: Tanzania, Rungwe District: Kyimbila, Stoltz 926 (holotype: B, destroyed; isotypes: G, K, W).
Englerastrum kassneri T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 70. 1924.
Plectranthus kassneri (T.C.E.Fr.) Hutch. & Dandy, Bull. Misc. Inform. Kew 1926: 481. 1926.
Coleus kassneri (T.C.E.Fr.) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 106. 1929. Type: DRC, Mt Morumbe, Kassner 2951 (holotype: B, destroyed; isotypes: BM, K).
Distribution.
Rwanda to S. Trop. Africa.
Coleus elliotii
(S.Moore) A.J.Paton comb. nov.
61AA789513C75193B6BB911AC5F4BAC5
urn:lsid:ipni.org:names:77201065-1
Pycnostachys elliotii S.Moore, J. Linn. Soc., Bot. 38: 275. 1908. Type: Uganda, Ruwenzori E, 2750 m, 10 Feb. 1906, Wollaston s.n. (holotype: BM).
Pycnostachys mildbraedii Perkins in G.W.J.Mildbraed (ed.), Wiss. Erg. Deut. Zentr.-Afr. Exped., Bot. 2: 548. 1913. Type: Uganda, Toro District/DRC, Ruwenzori, Butahu (Butagu) Valley, Friedrich 2537 (holotype: B, destroyed).
Pycnostachys cinerascens Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 352. 1928.
Pycnostachys bequaertii De Wild., Pl. Bequaert. 4: 393. 1928., nom. illeg., non Pycnostachys bequaertii De Wild. (1921). Type: DRC, Ruwenzori Lamia Valley, Bequaert 4287 (holotype: BR).
Pycnostachys butaguensis De Wild., Pl. Bequaert. 4: 389. 1928. Type: DRC, Ruwenzori, Butahu (Butagu) Valley, Bequaert 3715 (holotype: BR).
Distribution.
DRC to Uganda (Ruwenzori Mts.).
Coleus elongatus
Trimen, J. Bot. 27: 165. 1889
0E8205F5EE6B5AA4BAA35D61E6FAAE80
Englerastrum elongatum (Trimen) Alston, Bull. Misc. Inform. Kew 1926: 298. 1926.
Plectranthus elongatus (Trimen) R.H.Willemse, Blumea 25: 510. 1979. Type: Sri Lanka, Ritgala, July 1887, Trimen s.n. (holotype: PDA; isotype: K).
Distribution.
Sri Lanka.
Coleus eminii
(Gürke) A.J.Paton comb. nov.
8B7A91141F3B525A9B7A9D9249301998
urn:lsid:ipni.org:names:77201066-1
Pycnostachys eminii Gürke, Bot. Jahrb. Syst. 22: 145. 1895. Type: Tanzania, Kanessa, west of Lake Victoria, Kanessa, 14 Nov. 1890, Stuhlmann 943 (lectotype: B, destroyed, designated by Bruce (1940).
Pycnostachys ruwenzoriensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 384. 1900. Type: Uganda, Ruwenzori, Scott Elliot 7621 (holotype: K).
Pycnostachys rotundatodentata De Wild., Pl. Bequaert. 4: 391. 1928. Type: DRC, Ruwenzori, Kisuki, Bequaert 4701 (lectotype: BR, designated by Bramley in Paton et al. (2009)).
Distribution.
Cameroon to Ethiopia and NW. Tanzania.
Coleus engleri
(Briq.) A.J.Paton comb. nov.
26F6E93B029656428668DCF0EBB27EDD
urn:lsid:ipni.org:names:77201067-1
Anisochilus engleri Briq., Bot. Jahrb. Syst. 19: 190. 1894. Type: DRC, between Nyangwe and Kibundo, Pogge 1019 (lectotype: K, designated by Paton et al. (2009)).
Anisochilus africanus Baker, J. Linn. Soc., Bot. 30: 94. 1894., non Coleus africanus Benth.
Briquetastrum africanum (Baker) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 102. 1929.
Coleus africanus (Baker) Roberty, Bull. Inst. Fondam. Afrique Noire, Sér. A, Sci. Nat. 16: 330. 1954. nom. illeg.
Leocus africanus (Baker) J.K.Morton, J. Linn. Soc., Bot. 58: 270. 1962.
Plectranthus africanus (Baker) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 308. 2009. Type: Sierra Leone, Freetown, Scott Elliot 5033 (holotype: K).
Distribution.
W. Trop. Africa to Uganda and NE. Angola.
Notes.
Coleus africanus Benth., Labiat. Gen. Spec.: 54. 1832., is superfluous being based on Solenostemon ocymoides Schumach. & Thonn. in C.F.Schumacher, Beskr. Guin. Pl.: 271 (1827).
Coleus erici-rosenii
(R.E.Fr.) A.J.Paton comb. nov.
61DF0748851E5A3FB7BE2E398836BC3A
urn:lsid:ipni.org:names:77201068-1
Pycnostachys erici-rosenii R.E.Fr., Wiss. Erg. Schwed. Rhod.-Kongo Exped. 1: 281. 1916. Type: DRC, Ninagongo, 2000 m, R.E. Fries 1588 (holotype: UPS).
Pycnostachys albidoviolacea De Wild., Pl. Bequaert. 4: 400. 1928. Type: DRC, Kivu, Mukule, Bequaert 5889 (lectotype: BR, designated by Bramley in Paton et al. (2009)).
Pycnostachys robynsii De Wild., Pl. Bequaert. 4: 398. 1928. Type: DRC/Burundi, Busiga, Robyns 2356 (holotype: BR; isotype: K).
Distribution.
E. DRC to Uganda.
Coleus esculentus
(N.E.Br.) G.Taylor, J. Bot. 69 (suppl. 2): 158. 1931
54E36E9A093D5EE08E793F6A484BAE6F
Plectranthus esculentus N.E.Br., Bull. Misc. Inform. Kew 1894: 12. 1894. Type: cultivated at K from material sent by J. Medley Wood from KwaZulu-Natal, 1893 (lectotype: K, designated by Codd (1975); isolectotype: BOL).
Plectranthus floribundus N.E.Br., Bull. Misc. Inform. Kew 1894: 12. 1894.
Englerastrum floribundum (N.E.Br.) T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 73. 1924.
Coleus floribundus (N.E.Br.) Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 359. 1928. nom. illeg., non C. floribundus Baker. Type: South Africa, KwaZulu-Natal, Inanda, Medley Wood 646 (lectotype: K; isolectotype: PRE designated by Robyns & Lebrun (1928)).
Plectranthus floribundus var. longipes N.E.Br., Bull. Misc. Inform. Kew 1894: 13. 1894.
Coleus floribundus var. longipes (N.E.Br.) Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 360. 1928. Type: South Africa, valley of Umzingwani R., Baines s.n. (lectotype: K, designated by Robyns & Lebrun (1928)).
Coleus coppinii Heckel, Rev. Cultures Colon. 8: 166. 1900. Type and protologue not seen.
Coleus dazo A.Chev., Agric. Prat. Pays Chauds 4: 104. 1905. Type: tubers from Central African Republic, upper Oubangui, fragment at K sent from Oubangui, 1904 (syntype: K).
Coleus langouassiensis A.Chev., Veg. Ut. Afr. Trop. Franç. 1: 127. 1905. Type: not seen.
Distribution.
Trop. & South Africa.
Coleus eungellaensis
P.I.Forst. & A.J.Paton nom. nov.
5779B0CE91555BC9B6B0E27545D49B54
urn:lsid:ipni.org:names:77201068-1
Plectranthus graniticola P.I.Forst., Austrobaileya 3: 732. 1992., non C. graniticola (A.Chev.) A.J.Paton. Type: Australia, Queensland, South Kennedy District, Clarke Range, Eungella National Park, 23 Apr. 1991, P.I.Forster PIF8056 & W.J.F. Mc Donald (holotype: BRI; isotypes: CNS, K, MEL).
Distribution.
Australia: Queensland.
Coleus excelsus
(P.I.Forst.) P.I.Forst. comb. nov.
F229C1C981A25A0CB4946AE6C5039EC2
urn:lsid:ipni.org:names:77201070-1
Plectranthus excelsus P.I.Forst., Austrobaileya 4: 172. 1994. Type: Australia, Queensland, Cook District, Garroway Hill, 11 July 1993, P.I.Forster PIF13543 & M.C.Tucker (holotype: BRI, isotypes: A, AD, CBG, CNS, K, L, MEL).
Distribution.
Australia: Queensland.
Coleus exilis
(A.J.Paton) A.J.Paton comb. nov.
0584ECAA793A554D87711286582FE26C
urn:lsid:ipni.org:names:77201071-1
Plectranthus exilis A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 350. 2009. Type: Tanzania, Njombe District: Ruhudje, Lupembe, Schlieben 185 (holotype, K; isotypes: B, K, LISC, MO, PRE).
Distribution.
SW. Tanzania.
Coleus fasciculatus
(P.I.Forst.) P.I.Forst. comb. nov.
1CFFB3D7258059C4968371B242E569DE
urn:lsid:ipni.org:names:77201072-1
Plectranthus fasciculatus P.I.Forst., Haseltonia 6: 14. 1998 publ. 1999. Type: Australia, Queensland, Cook District, Mt Windsor Tableland, State Forest 144, 24 July 1993, P.I.Forster PIF13709, G. Sankowsky & M.C. Tucker (holotype: BRI; isotype: CNS).
Distribution.
Australia: Queensland.
Coleus ferricola
Phillipson, O. Hooper & A.J. Paton, Kew. Bull. 74, 24: 4. 2019
EDBEF0D8CB8F55B8922C2C847DFA9B17
Coleus ferricola Phillipson, O. Hooper & A.J. Paton, Kew. Bull. 74, 24: 4. 2019. Type: Guinea (Conakry), Nzérékoré, Lola, Guinée Forestière, Nimba Mountains, SMFG iron ore mine concession. East side of Pierre Richaud., Phillipson 6324 (holotype MO; isotypes BR, K, P, SERG, WAG).
Coleus foetidus
(Benth.) A.J.Paton comb. nov.
97552498D3D45B15BF9045B04F15D45E
urn:lsid:ipni.org:names:77201073-1
Plectranthus foetidus Benth., Labiat. Gen. Spec.: 35. 1832.
Ocimum foetidum Banks ex Benth., Labiat. Gen. Spec.: 35. 1832. nom. nud. Type: Australia, Queensland, Endeavour River, 1770, Banks & Solander, (holotype: BM).
Distribuion.
Australia: Queensland.
Coleus foliatus
(A.J.Paton) A.J.Paton comb. nov.
CB3171494ACD57189A62EA00435DA2FF
urn:lsid:ipni.org:names:77201074-1
Plectranthus foliatus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 284. 2009. Type: Tanzania, Sumbawanga District: Tatanda, Mbaa Hill, Bidgood et al. 3459 (holotype: K; isotypes: C, DSM, K, NHT).
Distribution.
SW. Tanzania to N. Zambia.
Coleus forsteri
(Benth.) A.J.Paton comb. nov.
E7809D5F088F5AE69AB07091BF11E9A8
urn:lsid:ipni.org:names:77201075-1
Plectranthus forsteri Benth., Labiat. Gen. Spec.: 38. 1832. Type: Vanuatu, Tanna, J.R. & G. Forster (holotype, BM).
Distribution.
SW. Pacific.
Coleus fragrantissimus
(P.I.Forst.) P.I.Forst. comb. nov.
249C6C344DBA57F8B85A2A83AAF778C8
urn:lsid:ipni.org:names:77201076-1
Plectranthus fragrantissimus P.I.Forst., Austrobaileya 8: 394. 2011. Type: Australia, Queensland, Port Curtis District, State Forest 316, Kroombit Tops, 16 Feb. 1995, P.I.Forster PIF16246 (holotype: BRI; MEL, NE, istotypes).
Distribution.
Australia: Queensland.
Coleus fredericii
G.Taylor, J. Bot. 69 (Suppl. 2): 159. 1931
9A99019536FE5110B6F025FDD743880A
Neomuellera welwitschii Briq., Bot. Jahrb. Syst. 19: 180. 1894., non Coleus welwitschii Briq.
Plectranthus welwitschii (Briq.) Codd, Fl. Pl. Africa 42: t. 1646. 1972. Type: Angola, Pungo Andongo, April 1857, Welwitsch 5544 (isotypes: BM, C, K).
Distribution.
Angola.
Coleus fruticosus
Wight ex Benth. in A.P.de Candolle, Prodr. 12: 78. 1848
C06F0E8DD3785B7BBEA4910F6227AD64
Plectranthus fruticosus (Wight ex Benth.) Hook.f., Fl. Brit. India 4: 623. 1885, nom. illeg.
Plectranthus deccanicus Briq., Annuaire Conserv. Jard. Bot. Genève 2: 234. 1898. Type: India, Pulney Hills, 1837, Wight 2514 (holotype: K).
Distribution.
SW. India
Coleus galeatus
(Vahl) Benth., Labiat. Gen. Spec.: 56. 1832
02FFA3ACC72D5D12B682AABEE71CCD3F
Germanea galeata (Vahl) Poir. in J.B.A.M.de Lamarck, Encycl. 2: 763. 1788.
Majana galeata (Vahl) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Plectranthus galeatus Vahl, Symb. Bot. 1: 43. 1790. Type: Indonesia, Java, Vahl s.n. (holotype: C).
Ocimum plectranthus J.F.Gmel., Syst. Nat. ed. 13(bis): 917. 1792. Type: not seen.
Plectranthus bicolor Blume, Bijdr. Fl. Ned. Ind.: 837. 1826.
Coleus bicolor (Blume) Benth., Labiat. Gen. Spec.: 55. 1832.
Majana bicolor (Blume) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Indonesia, Java, Blume 2037. (lectotype: L, designated by Keng (1969)).
Plectranthus macrophyllus Blume, Bijdr. Fl. Ned. Ind.: 835. 1826.
Coleus macrophyllus (Blume) Benth., Labiat. Gen. Spec.: 55. 1832.
Majana macrophylla (Blume) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Indonesia, Java, Blume 1453 (syntype: L).
Plectranthus leschenaultii Benth., Labiat. Gen. Spec.: 34. 1832. Type: Java, Leschenault (holotype: P).
Coleus macrostachys Benth., Labiat. Gen. Spec.: 57. 1832. Type: Indonesia, Java, Commerson s.n. (lectotype: P, designated here).
Coleus macropus Miq., Fl. Ned. Ind. 2: 956. 1858. Type: Indonesia, ?Java, Horsfield s.n. (holotype: U; isotypes: BM, K).
Coleus puberulus Miq., Fl. Ned. Ind. 2: 955. 1858.
Majana puberula (Miq.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Indonesia, Sumatra, Singalang, Teysmann s.n. (isotype: U).
Coleus remotiflorus Miq., Fl. Ned. Ind. 2: 954. 1858. Type: Indonesia, Java in Soerakerta, Horsfield Lab. 112 (holotype: U; isotypes: BM, K).
Coleus spectabilis Miq., Fl. Ned. Ind. 2: 955. 1858. Type: Indonesia, Java, in Blam.-bagan, Horsfield s.n. (holotype: U; isotypes: BM, K).
Distribution.
W. Malesia to Lesser Sunda Is. (Bali).
Coleus gamblei
(Smitha & Sunojk.) Smitha comb. nov.
D8E0E09BA1A45F77A94BBE5324652092
urn:lsid:ipni.org:names:77201077-1
Plectranthus gamblei Smitha & Sunojk., Nordic J. Bot. 36 (8): 4. 2018. Type: India. Tamil Nadu, Coonoor, Aug 1883, Gamble 12263 (holotype: K).
Distribution.
S. India.
Coleus garckeanus
Vatke, Linnaea 37: 323. 1872
710092C92DF35CCFAF16672B5772B2CA
Plectranthus garckeanus (Vatke) J.K.Morton, Novon 8: 265. 1998. Type: Ethiopia, Begember in valley of river Repp, 1 July 1863, Schimper 1193 (holotype: B, destroyed; isotypes: BM, E, JE, K).
Plectranthus salviiflorus Chiov., Malpighia 34: 516. 1937. Type: Ethiopia, Boyi near Kardobi and Abeba Kolla, Taschdjian 188 (syntype: FT); 124 (syntype: FT).
Distribution.
Ethiopia.
Coleus geminatus
(P.I.Forst.) P.I.Forst. comb. nov.
EF1959025B775D3E964DEFED08E6F80B
urn:lsid:ipni.org:names:77200982-1
Plectranthus geminatus P.I.Forst., Austrobaileya 9: 285. 2014. Type: Australia, Queensland, Moreton District, Lamington National Park, Cainbable Falls, 27 Feb. 2014. P.I.Forster PIF40722 & G.Leiper (holotype: BRI; isotypes: K, MEL, NSW).
Distribution.
Australia: Queensland.
Coleus gibbosus
A.J.Paton nom. nov.
11699B4C21205A1B9D16CC37121543E9
urn:lsid:ipni.org:names:77201079-1
Plectranthus rehmannii Gürke, Bull. Herb. Boissier 6: 553. 1898., non Coleus rehmanii Briq. Type: South Africa, Natal, Karkloof, Rehmann 7359 (holotype: Z; isotype: K).
Distribution.
South Africa: KwaZulu-Natal.
Coleus gigantifolius
(Suddee) Suddee comb. nov.
439BC34F8615530593FC5A675683EA3C
urn:lsid:ipni.org:names:77201080-1
Plectranthus gigantifolius Suddee, Kew Bull. 59: 401 (2004). Type: Thailand, Kanchanaburi, Thung Yai Naresuarn Wildlife Sanctuary, Ban Saneh Pawng, Lai Wo Subdistr., 225 m, 7 Oct. 1993, Maxwell 93-1156 (holotype: CMU; isotypus: L).
Distribution.
SW. Thailand.
Coleus gillettii
(J.K.Morton) A.J.Paton comb. nov.
D8B4F73840AD5BEB8FB2D946C6E11099
urn:lsid:ipni.org:names:77201081-1
Plectranthus gillettii J.K.Morton, Kew Bull. 53: 998. 1998. publ. 1999. Type: Kenya, Northern Frontier District: Moyale, Gillett 13721 (holotype: K; isotypes: EA, LMA).
Distribution.
S. Ethiopia, S. Somalia, N. Kenya.
Coleus glabriflorus
(P.I.Forst.) P.I.Forst. comb. nov.
8EE40C1352355672ABF463F2682C897E
urn:lsid:ipni.org:names:77201082-1
Plectranthus glabriflorus P.I.Forst., Austrobaileya 4: 174. 1994. Type: Australia, Queensland, North Kennedy District, Herberton, 20 Jan. 1993, P.I.Forster PIF12814 & A.R.Bean (holotype: BRI; isotype: CNS).
Distribution.
Australia: Queensland.
Coleus globosus
(Ryding) A.J.Paton comb. nov.
2999EEC71FDE5BE98C3B70A4ACE5F3D2
urn:lsid:ipni.org:names:77201083-1
Plectranthus globosus Ryding, Bull. Jard. Bot. Natl. Belg. 66: 101. 1997. Type: DRC, Shaba, Kundulungu Plateau, 20 Mar. 1971, Lisowski 23152 (holotype: POZG, photo C).
Distribution.
DRC.
Coleus goetzenii
(Gürke) A.J.Paton comb. nov.
7BDADD094C8F5D0EA4379AB00F8F0AE5
urn:lsid:ipni.org:names:77201084-1
Pycnostachys goetzenii Gürke, Durch Afr., reimpr.: 8. 1896). Type: Rwanda, Virunga Mts, Goetzen 98 (holotype: B, destroyed).
Pycnostachys vulcanicola Lebrun & L.Touss., Bull. Jard. Bot. État Bruxelles 17: 71. 1943. Type: DRC, Virunga Range, Lebrun 5006 (holotype: BR; isotype: K).
Distribution.
EC. Trop. Africa (Virunga Mts.).
Coleus gossweileri
A.J.Paton nom. nov.
33490339B59255FFB08DD672BE48A875
urn:lsid:ipni.org:names:77201085-1
Pycnostachys gracilis R.D.Good, J. Bot. 69 (suppl. 2): 164. 1931., non Coleus gracilis Gürke. Type: Angola, R. Cuartiri, Gossweiler 4035 (holotype: BM).
Distribution.
Angola.
Coleus gracillimus
(T.C.E.Fr.) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 106. 1929
AC0FC4CE129C5FA8929DC4A2F1B2FCCE
Englerastrum gracillimum T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 69. 1924.
Plectranthus gracillimus (T.C.E.Fr.) Hutch. & Dandy, Bull. Misc. Inform. Kew 1926: 481. 1926. Type: DRC, Mt Corva, Deschamps s.n. (holotype: B, destroyed; isotypes: UPS, K, fragment).
Englerastrum schlechteri T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 70. 1924.
Plectranthus schlechteri (T.C.E.Fr.) Hutch. & Dandy, Bull. Misc. Inform. Kew 1926: 481. 1926.
Coleus schlechteri (T.C.E.Fr.) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 106. 1929. Types: Mozambique, Dondo near Beira, Schlechter 12240 (syntype: B, destroyed, isosyntype: PRE, K, fragment) & Tanzania, Rungwe District: Kyimbila, Mulinda, Stoltz 1440 (syntype: B, destroyed, isosyntypes: K, W).
Plectranthus micranthus Chiov., Boll. Soc. Bot. Ital. 1924: 44. 1924, nom. illeg., non Spreng.
Englerastrum diffusum Alston, Bull. Misc. Inform. Kew 1926: 298. 1926.
Plectranthus tenuis Hutch. & Dandy, Bull. Misc. Inform. Kew 1926: 481. 1926.
Coleus diffusus (Alston) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 106. 1929. Type: Nigeria: Nupe, Barter s.n. (syntype: K); Nigeria, Muri Province, weed in arable land, Lamb 72 (syntype, K).
Distribution.
Widespread in Trop. Africa.
Coleus gracilipedicellatum
(Robyns & Lebrun) A.J.Paton comb. nov.
7F92193B7D7A5194BB16E97B00058191
urn:lsid:ipni.org:names:77201086-1
Holostylon gracilipedicellatum Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 103. 1929. Type: DRC, Katanga, Pweto to Moba (Baudouinville), between Kayabala and Lungulungu, 29 Apr. 1926, Robyns 2196 (holotype: BR; isotypes: BM, K).
Plectranthus baumii Gürke in O. Warburg (ed.), Kunene-Sambesi Exped.: 356. 1903, non Coleus baumii Gürke.
Holostylon baumii (Gürke) G.Taylor, J. Bot. 69(suppl. 2): 161. 1931. Type: Angola, Kubango, Massaca, 19 Oct. 1899, Baum 283 (holotype: B, destroyed, isotypes: BM, E (as 238), K, W).
Distribution.
Southern DRC to Botswana.
Coleus gracilis
Gürke, Bot. Jahrb. Syst. 38: 170. 1906
33C3E683198D538E844020BA52535672
Plectranthus puberulentus J.K.Morton, Novon 8: 265. 1998. Types: Ethiopia, Sagan, Riva 1564 (syntype: B, destroyed; isosyntype: FT), Harar, Ellenbeck 965 (syntype: B, destroyed).
Distribution.
Ethiopia to Tanzania.
Coleus graminifolius
(Perkins) A.J.Paton comb. nov.
404C010CBFCE5437A7D350C5518A845E
urn:lsid:ipni.org:names:77201087-1
Pycnostachys graminifolia Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 66. 1921. Type: Tanzania, Dodoma District: Kilimatinde, Prittwitz 118 (holotype: B, destroyed; isotype: UPS, K, fragment).
Distribution.
Kenya.
Coleus grandicalyx
E.A.Bruce, Hooker’s Icon. Pl. 34: t. 3391 1939
AE0F113F15085EC5B198D5D09FD0AC8D
Plectranthus grandicalyx (E.A.Bruce) J.K.Morton, Novon 8: 265. 1998. Type: Sudan, Imatong Mts, Lomul, A.S.Thomas 1935 (holotype: K).
Distribution.
S. Sudan to N. Uganda.
Coleus grandidentatus
(Gürke) A.J.Paton comb. nov.
376BBE47C37451FAA3B56C01FD325163
urn:lsid:ipni.org:names:77201088-1
Plectranthus grandidentatus Gürke, Bull. Herb. Boissier 6: 554. 1898. Type: South Africa, Cape, East Griqualand, Emyembe Mt, April 1885, Tyson 1517 (lectotype: K, designated by Codd (1975)).
Distribution.
S. Zimbabwe to South Africa.
Coleus graniticola
(A.Chev.) A.J.Paton comb. nov.
854BEE4A6DB2519FB28E4B92EB262F7A
urn:lsid:ipni.org:names:77201089-1
Solenostemon graniticola A.Chev., J. Bot. (Morot) 22: 121. 1909.
Solenostemon monostachyus var. graniticola (A.Chev.) Brenan, Kew Bull. 5: 231. 1950.
Plectranthus saxicola B.J.Pollard & A.J.Paton, Kew Bull. 61: 229. 2006. Type: Ivory Coast, Mt Nienokue, 20 km NE of Fort Binger, Chevalier 19469 (holotype: P; isotype: K).
Distribution.
W. & WC. Trop. Africa.
Coleus gratus
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
E5B4CEF2E41853A280B207A18282D171
urn:lsid:ipni.org:names:77201090-1
Plectranthus gratus S.T.Blake, Contr. Queensland Herb. 9: 49. 1971. Type: Australia, Queensland, Walsh’s Pyramid (Cultivated at The Gap), 10 Jan. 1960, S.T. Blake 21192 (holotype: BRI).
Distribution.
Australia: NE. Queensland.
Coleus graveolens
(R.Br.) A.J.Paton comb. nov.
7B378354F4445F0C82192A5E8E6BC82D
urn:lsid:ipni.org:names:77201091-1
Plectranthus graveolens R.Br., Prodr. Fl. Nov. Holl.: 506. 1810.
Plectranthus parviflorus var. graveolens (R.Br.) Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 357. 1895.
Plectranthus australis var. graveolens (R.Br.) Domin, Biblioth. Bot. 89: 565. 1928. Type: Australia, Queensland, Port Clinton, 22. Aug. 1802, R.Brown (holotype: BM).
Plectranthus australis f. eximius Domin, Biblioth. Bot. 89: 565. 1928. Type: not seen.
Distribution.
E. Australia, Lord Howe I.
Coleus guerkei
(Briq.) A.J.Paton comb. nov.
07091383DD765B4CA75F84DC4D5F07DB
urn:lsid:ipni.org:names:77201092-1
Hyptis baumii Gürke in O. Warburg (ed.), Kunene-Sambesi Exped.: 354. 1903., non Coleus baumii Gürke.
Plectranthus guerkei Briq., Annuaire Conserv. Jard. Bot. Genève 7–8: 323. 1904.
Germanea guerkei Briq., Annuaire Conserv. Jard. Bot. Genève 7–8: 323. 1904., nom. altern. Type: Angola, Cuito (Kuito), Baum 789 (isotypes: K, W).
Geniosporum paniculatum Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 351. 1900., non Coleus paniculatus Benth.
Neohyptis paniculata (Baker) J.K.Morton, J. Linn. Soc., Bot. 58: 259. 1962. Type: Angola, Pungo Andongo, near Catete, Welwitsch 5528 (syntypes: BM, K).
Geniosporum paniculatum var. debile Hiern, Cat. Afr. Pl. 1: 853. 1900. Type: Angola, Pungo Andongo, Casalala near Pedra Songue, Welwitsch 5527 (holotype: LISU; isotype: BM).
Hyptis quadrialata A.Chev., Explor. Bot. Afrique Occ. Franç. 1: 522. 1920., nom. nud.
Geniosporum borzianum Chiov. Ann. Bot. (Rome) 10: 402. 1912. Type: Ethiopia. Amhara-Dembià, Mt. Incedubà near Gondar, Chiovenda 2266 (holotype: FT), synon. nov.
Distribution.
Widespread in Trop. & South Africa.
Coleus gymnostomus
Gürke, Bot. Jahrb. Syst. 19: 212. 1894
3203B26ABCC6500995F8418C8B49DF05
Plectranthus gymnostomus (Gürke) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 337. 2009. Type: East Africa, without precise locality, Fischer 333 (holotype: B, destroyed; isotype: K, photo.).
Distribution.
Tanzania.
Coleus habrophyllus
(P.I.Forst.) P.I.Forst. comb. nov.
431DEBE8A5E85641B0857E8D44640C62
urn:lsid:ipni.org:names:77201093-1
Plectranthus habrophyllus P.I.Forst., Austrobaileya 4: 175. 1994. Type: Australia, Queensland, Moreton District, Ormeau, CSR land near water tower, 12 Nov. 1992, P.I.Forster PIF12391 & G.Leiper (holotype: BRI, isotypes: K, MEL).
Distribution.
Australia: Queensland.
Coleus hadiensis
(Forssk.) A.J.Paton comb. nov.
CB1026D886145A4CB550FA4C13D58EF9
urn:lsid:ipni.org:names:77201104-1
Ocimum hadiense Forssk., Fl. Aegypt.-Arab.: 109. 1775.
Plectranthus forskalaei Vahl, Symb. Bot. 1: 44. 1790., nom. illeg.
Germanea forskalaei Poir. in J.B.A.M.de Lamarck, Encycl., suppl. 2: 764. 1812., nom. superfl.
Majana forskalaei (Poir.) Kuntze, Revis. Gen. Pl. 2: 524. 1891., nom. superfl.
Plectranthus hadiensis (Forssk.) Schweinf. ex Sprenger, Wiener Ill. Gart.-Zeitung 19: 2. 1894.
Coleus forskalaei Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 359. 1897., nom. superfl. Type: Yemen, Hadiyah ((Hadi)) Mts, Forsskål s.n (C, holotype).
Plectranthus zeylanicus Benth., Labiat. Gen. Spec.: 36. 1832.
Coleus zeylanicus (Benth.) L.H.Cramer, Kew Bull. 32: 560. 1978. Type: Ceylon, Macrae s.n. (holotype: K).
Plectranthus tomentosus Benth. in E.H.F.Meyer, Comm. Pl. Afr. Austr.: 229. 1838. Plectranthus zatarhendi var. tomentosus (Benth.) Codd, Bothalia 11: 399. 1975.
Plectranthus hadiensis var. tomentosus (Benth.) Codd, Fl. S. Afr. 28: 153. 1985. Type: South Africa, Durban (Port Natal), Drège s.n. (holotype: K).
Coleus personatus Lem., Hort. Universel 6: 128. 1844. Type: not seen. Material at G under this name, labelled Hort Paris Aug. 1845, is P. hadiensis.
Plectranthus madagascariensis var. ramosior Benth. in A.P.de Candolle, Prodr. 12: 68. 1849.
Plectranthus ramosior (Benth.) van Jaarsv., Aloe 43: 46. 2006. Type: South Africa, Gauteng, Magaliesberg (Macalisberg), Burke s.n. (holotype: BM).
Coleus schweinfurthii Vatke, Linnaea 37: 323. 1872. Type: Ethiopia, Bellaka, Sept. 1854, Schimper s.n. (Type: destroyed B).
Plectranthus paucicrenatus Franch., Sert. Somal.: 56. 1882. Type: Somalia, Northern Somalia, Révoil s.n. (holotype: P).
Coleus rupestris Hochst. ex Briq., Bull. Herb. Boissier 2: 131. 1894., nom. nud.
Plectranthus cyaneus Gürke, Bot. Jahrb. Syst. 19: 208. 1894. Type: Tanzania, Lushoto District: Mashewa (Mascheua), Holst 8850 (syntype: B, destroyed; isosyntype: K) & Handei, near Kwa Mshusa, Holst 8984 (syntype: B, destroyed; isosyntype: JE, K, W).
Plectranthus woodii Gürke, Bot. Jahrb. Syst. 26: 76. 1898.
Plectranthus zatarhendi var. woodii (Gürke) Codd, Bothalia 11: 401. 1975.
Plectranthus hadiensis var. woodii (Gürke) Codd, Fl. S. Afr. 28: 154. 1985. Type: South Africa, KwaZulu-Natal, Ipolweni, Medley Wood s.n. (lectotype: GRA, designated by Codd (1975)).
Germanea horrida Hiern, Cat. Afr. Pl. 1: 863. 1900.
Plectranthus horridus (Hiern) Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 416. 1900. Types: Angola, Pungo Andongo, R. Cuanza near Condo, Welwitsch 5537 (syntypes: BM, K, LISU.); Huila, Morro de Monino, Welwitsch 5613 (syntypes: LISU, K).
Plectranthus rupestris Vatke ex Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 409. 1900.
Coleus rupestris Hochst. ex Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 409. 1900., nom. inval. (based on Schimper 2172 as below) Type: Ethiopia, near Bellaka, Sept. 1854, Schimper 2172 (lectotype: K; isolectotypes: BM, W, designated by Paton et al. 2009).
Plectranthus draconis Briq., Bull. Herb. Boissier, sér. 2, 3: 1071. 1903. Type: South Africa, KwaZulu-Natal, Biggarsberg, Rehmann 7092 (holotype: Z).
Plectranthus hararensis Gürke, Bot. Jahrb. Syst. 36: 132. 1905. Type: Ethiopia, Harar, Bachufer, Ellenbeck 846 (holotype: B, destroyed; K, fragment).
Plectranthus erlangeri Gürke, Bot. Jahrb. Syst. 38: 166. 1906. Type: Ethiopia, Abuelkasim, Ellenbeck 1396 (holotype: B, destroyed but see fig. in protologue).
Plectranthus pachyphyllus Gürke ex T.Cooke in W.H.Harvey & auct. suc. (eds.), Fl. Cap. 5(1): 285. 1910. Type: South Africa, Natal, Inchanga, Rehmann 7878 (holotype: Z).
Plectranthus petrensis S.Moore, J. Linn. Soc., Bot. 40: 175. 1911. Type: Zimbabwe, Chimanimani, Swynnerton 2018 (holotype: BM; isotype: K).
Plectranthus fragrans Lebrun & L.Touss., Bull. Jard. Bot. État Bruxelles 17: 70. 1943. Type: DRC, Katanda, Lebrun 7618 (holotype: BR; isotype: K).
Distribution.
Widespread, Egypt to South Africa, Arabian Peninsula, Maldives, Sri Lanka.
Notes.
Heckenhauer et al. (2019) suggest the Sri Lankan C. zeylanicus may represent a separete species or subspecies but further work is required.
Coleus hallii
(J.K.Morton) A.J.Paton comb.nov.
7DFA8723CBD65071AAF3428E4FC2765E
urn:lsid:ipni.org:names:77201105-1
Plectranthus hallii J.K.Morton, J. Linn. Soc., Bot. 58: 267. 1962. Type: Ghana, Gambaga Scarp, 14 Nov. 1959, J.K.Morton A3791, (holotype: GC; isotype: K).
Distribution.
Ghana.
Coleus harmandii
(Doan ex Suddee & A.J.Paton) A.J.Paton comb. nov.
3FEF7B50FF6854B1B09D9DB861E7268C
urn:lsid:ipni.org:names:77201106-1
Anisochilus harmandii Doan ex Suddee & A.J.Paton, Kew Bull. 59: 384. 2004. Type: Cambodia, Mlu Prey, Jan. 1876, Harmand 326 (holotype: P; isotype: A).
Distribution.
Indo-China.
Coleus helferi
(Hook.f.) A.J.Paton comb. nov.
30CE589FB87E5247B11A3B92CCA13D3C
urn:lsid:ipni.org:names:77201107-1
Plectranthus helferi Hook.f., Fl. Brit. India 4: 623. 1885. Type: Burma, Tenasserim, Helfer 4061 (holotype: K).
Plectranthus fulvescens Prain, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 66: 521. 1897. Type: Burma, Attran, Brandis 811 (holotype: CAL; isotype: K),
Coleus fulvescens Kurz, Mss in Herb. Calcutta, nom. inval., in protologue discussion above.
Distribution.
Myanmar to N. Thailand.
Coleus hereroensis
(Engl.) A.J.Paton comb. nov.
0A03D34E51075D769F82C049F4F0D441
urn:lsid:ipni.org:names:77201108-1
Plectranthus hereroensis Engl., Bot. Jahrb. Syst. 10: 267. 1888.
Burnatastrum hereroensis (Engl.) P.V.Heath, Calyx 4: 175. 2001. Type: Namibia, Hereroland, Kaiser Wilhelmsberg near Okahandja, May 1886, Marloth 1350 (holotype: B, destroyed; isotypes: G, M, PRE, SAM).
Coleus aconitiflorus Welw. ex Hiern, Cat. Afr. Pl. 1: 866. 1900. Type: Angola, Huíla, Catumba, Mar.1879, Welwitsch 5495 (holotype: LISU; isotypes: G, K).
Plectranthus matabelensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 417. 1900. Type: Zimbabwe, Matabeleland, Shashe R., 27.Mar.1875, Holub 1403–1406 (holotype: K).
Neomuellera damarensis S.Moore, J. Bot. 1901: 265. 1901. Type: Namibia, Damaraland, 1879, Een s.n. (holotype: BM).
Germanea myriantha Briq., Bull. Herb. Boissier, sér. 2, 3: 1001. 1903.
Plectranthus myrianthus Briq., Bull. Herb. Boissier, sér. 2, 3: 1001. 1903.
Coleus myrianthus (Briq.) Brenan, Mem. New York Bot. Gard. 9: 43. 1954. Type: South Africa, Witwatersrand, Apr.1895, Hutton 877 (holotype: Z: isotypes: GRA, K, PRE).
Coleus matopensis S.Moore, J. Bot. 45: 96. 1907. Type: Zimbabwe, Matobo Hills, Eyles 1024 (holotype: BM; isotype: GRA).
Coleus polyanthus S.Moore, J. Bot. 45: 96. 1907. Type: Zimbabwe, Matobo Hills, Nov. 1903, Eyles 1012 (holotype: BM).
Coleus gazensis S.Moore, J. Linn. Soc., Bot. 40: 178. 1911. Type: Zimbabwe, Chimanimani (Melsetter), 28 June1907, Swynnerton 1998 (holotype: BM).
Plectranthus otaviensis Dinter, Repert. Spec. Nov. Regni Veg. Beih. 53: 116. 1928. nom. inval. Based on Namibia, Otavi, Dinter 5699 (B, destroyed, PRE, SAM).
Plectranthus aurifer Dinter ex Launert, Mitt. Bot. Staatssamml. München 2: 312. 1957. Type: Namibia, Grootfontein, Nossib, 22 Apr.1934, Dinter 7367 (holotype: M, isotypes: BOL, K).
Distribution.
Southern tropical & South Africa.
Coleus hijazensis
(Abdel Khaik) A.J.Paton comb. nov.
88D2EEAB5059561189D863D836E3FB01
urn:lsid:ipni.org:names:77201109-1
Plectranthus hijazensis Abdel Khalik, Turkish J. Bot. 40(5): 507. 2016. Type: Saudi Arabia, Al Baha Province, Saad Medhass, Abdel Khalik & Howladar s.n (holotype: UQU; isotypes: E, K, SHG).
Distribution.
SW. Arabian Peninsula
Coleus humulopsis
(B.J.Pollard) A.J.Paton comb. nov.
281D85B75F2D5047926D46F6DC179F7B
urn:lsid:ipni.org:names:77201110-1
Leocus membranaceus J.K.Morton, J. Linn. Soc., Bot. 58: 270. 1962., non Coleus membranaceus Briq.
Plectranthus humulopsis B.J.Pollard, Kew Bull. 64: 260. 2009. Type: Ghana, Kintampo, Morton A2585 (holotype: GC; isotypes: FHI, IFAN, K, WAG).
Distribution.
W. Trop. Africa to Cameroon.
Coleus hymalis
(J.R.I.Wood) A.J.Paton comb. nov.
914F5E6F05B8501980BA7DA6ABF23067
urn:lsid:ipni.org:names:77201111-1
Plectranthus hyemalis J.R.I.Wood, Kew Bull. 39: 133. 1984. Type: Yemen, between Musalla & Saqool, Ahkum, Hujariyah, 2 Feb. 1977, J.R.I. Wood 1532 (holotype: K; isotypes BM, E).
Distribution.
Yemen.
Coleus idukkianus
(J.Mathew, Yohannan & B.J.Conn) Smitha comb. nov.
D369DECF8A3959D8A030CD58BDF3E6CA
urn:lsid:ipni.org:names:77201112-1
Plectranthus idukkianus J.Mathew, Yohannan & B.J.Conn, Telopea 20: 183. 2017. Type: India: Kerala: Idukki District, 10 km away from Kuttikkanam, Panchalimedu Hills, 14 Dec. 2015, J. Mathew 4821 (holotype: TBGT; isotype: MSSRF).
Distribution.
India, endemic to the Western Ghats.
Coleus igniarioides
(Ryding) A.J.Paton comb. nov.
02D1F1DAFE2B5DB9B8DBB6350480C080
urn:lsid:ipni.org:names:77201113-1
Plectranthus igniarioides Ryding, Novon 15: 361. 2005. Type: Somalia, Sanaag, 43 km from Ceel Afweyn, on road to Ceerigaabo, M. Thulin 10768 (holotype: UPS; isotype: K).
Distribution.
Somalia.
Coleus igniarius
Schweinf., Beitr. Fl. Aethiop.: 121. 1867
D453AD03D9C55572A1796C04655FD83F
Plectranthus igniarius (Schweinf.) Agnew, Upland Kenya Wild Fl.: 638. 1974. Types: Ethiopia, Dehli-Dikeno, 1854, Schimper 527 (holotype: B, destroyed; neotype: Ethiopia, Gondar Region, Dehli Dikeno, July 1853, Schimper 1623 (neotype: P, designated by Ryding (2001)).
Plectranthus malinvaldii Briq., Bull. Herb. Boissier 2: 125. 1894.
Coleus malinvaldii (Briq.) Briq., Annuaire Conserv. Jard. Bot. Genève 2: 240. 1898. Type: Ethiopia, Gondar Region, Dehli Dikeno, 22 Oct. 1854, Schimper 529 (lectotype: G-DC; isolectotype: G, FT, P; K, photo., designated by Ryding (2001)).
Coleus degasparisianus Buscal. & Muschl., Bot. Jahrb. Syst. 49: 487. 1913. Types: Mozambique, Mbusi (but in reality from Mahio in Haddes Valley, Eritrea), von Aosta (holotype: B, destroyed; neotype: Eritrea, Arrot Valley, south of Aidereso, Schweinfurth & Riva 1398 (neotype: FT; BR, G, P, S, isoneo. designated by Ryding (2001)).
Coleus guidottii Chiov., Atti Reale Accad. Italia, Mem. Cl. Sci. Fis. 11: 54. 1940. Type: Eritrea, Habab, Cub Cub near Nacfa, Guidotti 776 (holotype: FT).
Plectranthus igniarius subsp. grandicalyx Beentje, Kenya Trees, Shrubs & Lianas: 633. 1994, nom. inval. Based on: Kenya, Mt Kulal, Verdcourt 2269 (single collection at K or EA not specified).
Distribution.
NE. Trop. Africa to N. Kenya, Arabian. Peninsula.
Coleus ignotus
(A.J.Paton) A.J.Paton comb. nov.
F16EF7422E5551A0B22DA5ED6A1BDDF7
urn:lsid:ipni.org:names:77201114-1
Plectranthus ignotus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 278. 2009. Type: Tanzania, Ufipa District: Mbisi (Mbizi) Mts, Bidgood, Mbago & Vollesen 2544 (holotype: K; isotypes: C, DSM, NHT).
Distribution.
SW. Tanzania.
Coleus inflatus
Benth., Labiat. Gen. Spec.: 58. 1832
C14D1923EEE3559783C819B175165C62
Majana inflata (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Plectranthus inflatus (Benth.) R.H.Willemse, Blumea 25: 509. 1979. Type: Ceylon, Macrae s.n., in Hb. Benth. (holotype: K (K000820110)).
Coleus benthamianus Arn., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18: 354. 1836.
Majana benthamiana (Arn.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Ceylon, Graham s.n. (type: not seen).
Distribution.
Sri Lanka.
Coleus inselbergii
(B.J.Pollard & A.J.Paton) A.J.Paton comb. nov.
58AE5A6B8D3F5B33A26840F64E710ABA
urn:lsid:ipni.org:names:77201115-1
Plectranthus inselbergi B.J.Pollard & A.J.Paton, Kew Bull. 61: 225. 2006. Type: Equatorial Guinea, Dalle Rocheuse (inselberg), 3 km S of Asok, 13 Jan. 1999, LeJoly 99/282 (holotype: BRLU; isotype: K).
Distribution.
Equatorial Guinea to NW. Gabon.
Coleus insignis
(Hook. f.) A.J.Paton comb. nov.
4A11BB56FAAB56DEAE489530D78D6C47
urn:lsid:ipni.org:names:77201116-1
Plectranthus insignis Hook.f., J. Proc. Linn. Soc., Bot. 7: 210. 1864. Type: Cameroons Upper Guinea Cameroon Mountain, 7000 ft., Mann 1257 (holotype: K).
Distribution.
Cameroon, Gabon, Central African Rep.
Coleus insolitus
(C.H.Wright) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 94. 1929
891946BC47495B868C4804504C943CD1
Symphostemon insolitus (C.H.Wright) Hiern, Cat. Afr. Pl. 1: 867. 1900.
Plectranthus insolitus C.H.Wright, J. Linn. Soc., Bot. 34: 275. 1900. Type: Angola, Welwitsch 5593 (holotype: BM).
Distribution.
Angola.
Coleus insularis
(P.I.Forst.) P.I.Forst. comb. nov.
7F73D2314FFE58F18754B53FAA2244D0
urn:lsid:ipni.org:names:77201117-1
Plectranthus insularis P.I.Forst., Austrobaileya 8: 398. 2011. Type: Australia, Queensland, Darling Downs District, Mingimarny State Forest 131, Pine hill, 20 km S of Millmerran, 28 Apr. 1995, P.I.Forster PIF16468 & S.J.Figg (holotype: BRI; isotype: MEL).
Distribution.
Australia: Queensland.
Coleus intraterraneus
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
3B82D6CCFB575BCF9F28993BD84DC276
urn:lsid:ipni.org:names:77201118-1
Plectranthus intraterraneus S.T.Blake, Contr. Queensland Herb. 9: 33. 1971. Type: Australia, Northern Territory, Palm Valley W of Alice Springs (Cultivated at The Gap), 1958, S.T.Blake 20193 (holotype: BRI).
Distribution.
Australia: Queensland, Northern Territory, South Australia, Western Australia.
Coleus kanneliyensis
L.H.Cramer & S. Balas., Kew Bull. 32: 558. 1978
DFB3730B8CEF5FB6AB9C990864D8DF85
Plectranthus kanneliyensis (L.H.Cramer & S. Balas.) R.H.Willemse, Blumea 25: 510. 1979. Type: Sri Lanka, Kanneliya, Cramer 4317 (holotype: PDA; isotypes: E, K, NVB, US).
Distribution.
Sri Lanka.
Coleus kanyakumariensis
(Shinoj & Sunojk.) Smitha comb. nov.
4489B05666C0538EAA3A8CF2A579BDBF
urn:lsid:ipni.org:names:77201119-1
Anisochilus kanyakumariensis Shinoj & Sunojk. Phytotaxa 333(1): 100. 2018. Type: India, Kanyakumari district, Maruthwamala, K.Shinoj & P.Sunojkumar CU138400a (holotype: CALI; isotypes (CALI, MH).
Distribution.
S. India.
Coleus kapatensis
R.E.Fr., Wiss. Erg. Schwed. Rhod.-Kongo Exped. 1: 283. 1916
2D706495C0B55F3E91ED51855EFDEA24
Plectranthus kapatensis (R.E.Fr.) J.K.Morton, Novon 8: 265. 1998. Type: Zambia, Bangweulu (Bangwelo), Kamindas on Kapata Peninsula, R.E.Fries 871 (isotype: K).
Distribution.
S. Tanzania to S. Trop. Africa.
Coleus kirkii
(Baker) A.J.Paton comb. nov.
9B71E3D57D615A00926719E90CEFE222
urn:lsid:ipni.org:names:77201120-1
Pycnostachys kirkii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 381. 1900. Type: Malawi, Manganja Hills, Kirk s.n. (lectotype; K, designated by Bramley in Paton et al. (2009)).
Echinostachys reticulata E.Mey., Comm. Pl. Afr. Austr.: 243. 1838., non Coleus reticulatus A.Chev.
Pycnostachys reticulata (E.Mey.) Benth. in A.P.de Candolle, Prodr. 12: 83. 1848. Type: South Africa, KwaZulu-Natal, Umlazi Valley, near Durban, Drège s.n. (holotype: K; isotype: PRE).
Pycnostachys reticulata var. angustifolia Benth. in A.P.de Candolle, Prodr. 12: 83. 1848. Types: South Africa, KwaZulu-Natal, Durban, 1840, Krauss 329 (syntype: K); Gauteng, Magaliesburg, n.d., Burke s.n. (syntype: K).
Pycnostachys uliginosa Gürke, Bot. Jahrb. Syst. 30: 396. 1901. Type: Tanzania, Njombe District: Ubena, Luhigi stream, Goetze 806 (holotype: B, destroyed; K, fragment).
Pycnostachys holophylla Briq., Bull. Herb. Boissier, sér. 2, 3: 1000. 1903. Type: South Africa, Gauteng, Johannesburg, n.d., E.S.C.A. Herb. 347 (holotype: G).
Pycnostachys purpurascens Briq., Bull. Herb. Boissier, sér. 2, 3: 998. 1903. Type: South Africa, Witwatersrand, no locality, April.1895, Hutton 878 (holotype: Z).
Pycnostachys schlechteri Briq., Bull. Herb. Boissier, sér. 2, 3: 999. 1903. Type: South Africa, Eastern Cape, Mt Frere, 24 Jan.1895, Schlechter 6406 (holotype Z; isotypes: GRA, PRE).
Distribution.
Nigeria to Cameroon, Tanzania to South Africa.
Coleus kivuensis
Lebrun & L.Touss., Bull. Jard. Bot. État Bruxelles 17: 72. 1943
6D0E0A6D54835FCCB2D3E95CDC93E42D
Plectranthus kivuensis (Lebrun & L.Touss.) R.H.Willemse, Kew Bull. 40: 96. 1985. Type: DRC, Kivu, Rutshuru, Lebrun 9031 (holotype BR; isotype: K).
Distribution.
Eritrea to N. Tanzania.
Coleus klossii
(S.Moore) P.I.Forst. & T.C.Wilson comb. nov.
7504C1349C835A80A8DD3E53B16A10BE
urn:lsid:ipni.org:names:77201121-1
Plectranthus klossii S.Moore, Trans. Linn. Soc. London, Bot. 9: 137. 1916. Type: Indonesia, Papua, Utakwa River to Mt Carstensz, C. Boden-Kloss (holotype: BM (BM000950283); isotype: K).
Plectranthus klossii var. major S.Moore, Trans. Linn. Soc. London, Bot. 9: 137. 1916. Type: Indonesia, Papua, Utakwa River to Mt Carstenz, C. Boden-Kloss (holotype: BM (BM000950282)).
Coleus koualensis
A.Chev. ex Hutch. & Dalziel, Fl. W. Trop. Afr. 2: 290. 1931
776D773C74575AB0BA5A627D308F0AE5
Solenostemon koualensis (A.Chev. ex Hutch. & Dalziel) J.K.Morton, J. Linn. Soc., Bot. 58: 270. 1962.
Plectranthus koualensis (A.Chev. ex Hutch. & Dalziel) B.J.Pollard & A.J.Paton, Kew Bull. 61: 228. 2006. Type: Ivory Coast, Upper Sassandra, Mt Dourou, Fleury in Chevalier 21715 (holotype: P).
Distribution.
Ivory Coast (Mt. Dourou).
Coleus koulikoroensis
A.J.Paton nom. nov.
74B0C809881751AD9EBC214ED06BA9D2
urn:lsid:ipni.org:names:77201122-1
Solenostemon chevalieri Briq., Mém. Soc. Bot. France 8: 286. 1917., non Coleus chevalieri Briq.
Plectranthus chevalieri (Briq.) B.J.Pollard & A.J.Paton, Kew Bull. 61: 229. 2006. Type: Mali, Koulikoro, Chevalier 2779 (holotype: P).
Coleus nigericus A.Chev., Explor. Bot. Afrique Occ. Franç. 1: 519. 1920, nom. nud.
Distribution.
W. Trop. Africa.
Coleus kunstleri
(Prain) A.J.Paton comb. nov.
61FFD9DF75CA55BC83D4C8459DC64513
urn:lsid:ipni.org:names:77201123-1
Plectranthus kunstleri Prain, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 66: 521. 1897. Type: Malaysia, Perak, Kwala Dipoug (Kuala Dipang), Kunstler, 8240 (holotype: CAL).
Distribution.
Peninsula Malaysia (Perak).
Coleus lactiflorus
Vatke, Linnaea 43: 89. 1881
A5C1F299ED0D5F5B8990586DD4784552
Plectranthus lactiflorus (Vatke) Agnew, Upland Kenya Wild Fl.: 637. 1974. Type: Kenya, Teita District: Ndi Hill (N’Di), Hildebrandt 2522 (holotype: B, destroyed; neotype: Kenya, Teita District, Mbololo Forest, Taita Hills Expedition 1042 (neotype: K, isoneotype: EA, designated by Paton et al. (2009)).
Distribution.
Ethiopia to N. & NW. Tanzania.
Coleus laetus
(P.I.Forst.) P.I.Forst. comb. nov.
AE0B8A9A1660515E99848975FFF4C489
urn:lsid:ipni.org:names:77201124-1
Plectranthus laetus P.I.Forst., Austrobaileya 9: 431. 2015. Type: Australia, Queensland, Cook District, Orchid Creek Station, 30 April 2014, P.I.Forster PIF41138 & S.L.Thompson (holotype: BRI; isotypes: CNS, MEL, NSW).
Distribution.
Australia: Queensland.
Coleus lageniocalyx
Briq., Mém. Soc. Bot. France 8: 292. 1917
79AE9C2F796F5CF694F2770FDB523E4D
Coleus lageniocalyx Briq., Mém. Soc. Bot. France 8: 292. 1917. Type: Central African Republic, Bassinde Haute Ombella, Kaga Do, near Diouma, 26 Oct.1902, Chevalier 5923 (holotype: P).
Distribution.
Central African Republic
Coleus lanceolatus
(Bojer ex Benth.) A.J.Paton & Phillipson comb. nov.
5BD843FB12B252D29F89E39C72152526
urn:lsid:ipni.org:names:77201125-1
Plectranthus lanceolatus Bojer ex Benth., Labiat. Gen. Spec.: 40. 1832.
Burnatastrum lanceolatum (Bojer ex Benth.) Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 358. 1897. Type: Madagascar, Bojer s.n. (holotype: P; isotype: K).
Plectranthus lavanduloides Baker, J. Linn. Soc., Bot. 20: 230. 1883.
Burnatastrum lavanduloides (Baker) Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 358. 1897. Type: Madagascar, Central Madagascar, Baron 978 (holotype: K).
Plectranthus burnatii Briq., Bull. Herb. Boissier 2: 124. 1894. Type: Madagascar, around Antananarive, Goudot s.n. (holotype: G; isotype: G).
Distribution.
Madagascar.
Coleus lancifolius
(Bramley) A.J.Paton comb. nov.
4A37F371476B5E73832C771295243A37
urn:lsid:ipni.org:names:77201126-1
Pycnostachys lancifolia Bramley, Kew Bull. 60: 590. 2005 publ. 2006. Type: Tanzania, Kigoma District: Lubalisi village to Ntakatta, Bidgood, Leliyo & Vollesen 4621 (holotype: K; isotypes: C, DSM, NHT, P).
Distribution.
W. Tanzania.
Coleus lanuginosus
Hochst. ex Benth. in A.P.de Candolle, Prodr. 12: 79. 1848
25A6CF4494D157E19A3B8A9695A06DBD
Majana lanuginosa (Hochst. ex Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Plectranthus lanuginosus (Hochst. ex Benth.) Agnew, Upland Kenya Wild Fl.: 638. 1974. Type: Ethiopia, Adua, 4 Oct. 1842, Schimper 1915 (lectotype: K; isolectotypes: BM, FT, KIEL, UPS, W; designated by Ryding (2000)).
Coleus albidus Vatke, Linnaea 37: 321. 1872. Type: Ethiopia, Mettaro, 1 Oct. 1862, Schimper 347 (holotype: B, destroyed; isotypes: JE, PRE, Z).
Coleus schimperi Vatke, Linnaea 37: 320. 1872.
Coleus barbatus var. schimperi (Vatke) Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 430. 1900.
Plectranthus semayatensis Cufod., Bull. Jard. Bot. État Bruxelles 33(Suppl.): 838. 1963., non P. schimperi Vatke. Type: Ethiopia, Semajada (Semayata), 12 Oct. 1862, Schimper 618 (holotype: B, destroyed; isotypes: BM, G, JE, K).
Coleus montanus Hochst. ex Ces., Rendiconto Reale Accad. Sci. Fis. 12: 264–266. 1879. Type: Ethiopia, Schimper 2460 (holotype: RO, Erbario Cesati) synon. nov.
Coleus trichophorus Briq., Bull. Herb. Boissier 2: 130. 1894. Type: Ethiopia, Aman-eski, 5 Nov. 1854, Schimper 342 (holotype; G; isotype: P).
Coleus maculatus Gürke, Bot. Jahrb. Syst. 19: 210. 1894. Type: Tanzania, Moshi District: Marangu, Volkens 429 (holotype: B, destroyed; isotypes: BM, G).
Coleus maranguensis Gürke, Bot. Jahrb. Syst. 19: 216. 1894. Type: Tanzania, Moshi District: Marangu, Volkens 630 (holotype: B, destroyed; isotype: BM).
Coleus gomphophyllus Baker, Bull. Misc. Inform. Kew 1895: 225. 1895. Type: Somalia, Golis Range, Lort Phillips s.n. (holotype, K).
Coleus schweinfurthii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 432. 1900., nom. illeg., non C. schweinfurthii Vatke.
Coleus sodalium Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 526. 1900. Type: Eritrea, Mogad (Magod) Valley, Schweinfurth & Riva 1810 (holotype: K; isotype: Z).
Coleus somalensis S.Moore, J. Bot. 1901: 265. 1901. Type: Somalia, Gan Liban, Donaldson-Smith s.n. (holotype: BM).
Coleus gallaensis Gürke, Bot. Jahrb. Syst. 38: 169. 1906. Type: Ethiopia, Belana, Ellenbeck 374 (syntype: B, destroyed); Gara Mulata, Ellenbeck 576 (syntype: B, destroyed; neotype: Ethiopia, Harerge Region, Gara Mulata, Gillett 5348 (neotype: K, designated by Ryding (2000)).
Coleus helenae Buscal. & Muschl., Bot. Jahrb. Syst. 49: 487. 1913. Type: Zambia, Lake Bangweulu (Banguelo-See), but in reality from Eritrea, Mai Gogol, near Segeneti, von Aosta 862 (holotype: B, destroyed; neotype: Eritrea, Sageneti, Schweinfurth & Riva 1184 (neotype: FT; isoneotype: BR, K, P; designated by Ryding (2001)).
Distribution.
Eritrea to N. Tanzania, SW. Arabian Peninsula. Naturalized in Zimbabwe.
Coleus lasianthus
Gürke, Bot. Jahrb. Syst. 19: 212. 1894
6A4CF6922F2350408A81D5F944D3E10F
Plectranthus lasianthus (Gürke) Vollesen, Opera Bot. 59: 84. 1980. Types: Tanzania, Kondoa District: Irangi, Fischer 507 (holotype: B, destroyed; neotype selected here: Tanzania, Same District: Mkomazi Game Reserve, Ngurunga Dam, Abdallah, Mboya & Vollesen 96/109, (neotype: K; isoneotype: NHT, designated by Paton in Paton et al. (2009)).
Coleus decumbens Gürke, Bot. Jahrb. Syst. 19: 211. 1894. Types: Tanzania, Moshi District: Lake Chala (Dschallasee), Volkens 327 (syntype: B, destroyed; isosyntype; K) & Kenya, Kwale District: Duruma, Txamtei, Hildebrandt 2320 (syntype: B, destroyed).
Coleus tetensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 431. 1900.
Plectranthus tetensis (Baker) Agnew, Upland Kenya Wild Fl.: 635. 1974. Type: Mozambique, Lower Zambesi, near Tete, Kirk s.n. (holotype: K).
Coleus menyharthii Briq., Bull. Herb. Boissier, sér. 2, 3: 1074. 1903. Type: Mozambique, Mefídze (Mfidzi) R., towards Katacha, Menyharth 1185 (isotype: Z).
Coleus vagatus E.A.Bruce, Bothalia 6: 227. 1951.
Plectranthus vagatus (E.A.Bruce) Codd in Ross, Fl. Natal: 305. 1971, nom. inval. Type: South Africa, Nelspruit District: Kruger National Park, 2.4 km E of Skukuza, Codd 5489 (holotype: PRE; isotype: K).
Plectranthus erubescens Hedge, Fl. Madag. 175: 216. 1998. Type: Madagascar, Beza Mahafaly Reserve, Phillipson 1833 (holotype: E; isotypes: K, MO, P).
Distribution.
Ethiopia to South Africa, Madagascar.
Coleus lateriticola
(A.Chev.) Phillipson, O. Hooper & A.J. Paton, Kew. Bull. 74, 24: 9. 2019, non C. lateriticola A. Chev. Explor. Bot. Afrique Occ. Franc. 1: 519. 1920, nom. nud.
E1C074129E025731994ADDA93737ACF4
Solenostemon lateriticola A.Chev., J. Bot. (Morot) 22: 121. 1909.
Solenostemon monostachyus subsp. lateriticola (A.Chev.) J.K.Morton, J. Linn. Soc., Bot. 58: 271. 1962.
Plectranthus monostachyus subsp. lateriticola (A.Chev.) B.J.Pollard, Kew Bull. 56: 980. 2001. Type: Guinea, Fonta-Djalon, entre Irebeleya et Timbo, Chevalier 18300 (holotype: P, photo K).
Distribution.
Guinea, Ivory Coast, Sierra Leone.
Coleus laxus
(T.C.Wilson & P.I.Forst) T.C.Wilson & P.I.Forst comb. nov.
8F19CDAAE4535415B648A80A9652C4CF
urn:lsid:ipni.org:names:77201128-1
Plectranthus laxus T.C.Wilson & P.I.Forst., Australian Systematic Botany 31: 440. 2018. Type: Queensland: Cook: Mount Cook walking track, before National Park, 7 April 2016, P.I. Forster PIF45043 & K.R. McDonald (holotype: BRI; isotype: NSW).
Distribution.
Australia: Queensland.
Coleus leiperi
(P.I.Forst.) P.I.Forst. comb. nov.
328C05D97E6953559A1B1D061487F73F
urn:lsid:ipni.org:names:77201129-1
Plectranthus leiperi P.I.Forst., Austrobaileya 4: 177. 1994. Type: Australia, Queensland, Moreton District, Diana’s Bath area, D’Aguilar Range, 9 km E of Somerset Dam, 19 Jun. 1993, P.I.Forster PIF13348 (holotype: BRI, isotype: MEL).
Distribution.
Australia: Queensland.
Coleus linearifolius
(J.K.Morton) A.J.Paton comb. nov.
4970BE69CD6B58EAA2739142666B39C5
urn:lsid:ipni.org:names:77201130-1
Plectranthus linearifolius (J.K.Morton) B.J.Pollard & A.J.Paton, Kew Bull. 61: 229. 2006.
Solenostemon linearifolius J.K.Morton, J. Linn. Soc., Bot. 58: 271. 1962. Type: Guinea, Fiendiou, Pobeguin 1920 (holotype: IFAN; isotype P).
Distribution.
Guinea.
Coleus livingstonei
A.J.Paton nom. nov.
958FCD53B6D25F30B4F6B5426E5F2657
urn:lsid:ipni.org:names:77201131-1
Pycnostachys urticifolia Hook., Bot. Mag. 89: t. 5365. 1863.
Coleus urticifolius (Hook.) Roberty, Bull. Inst. Fondam. Afrique Noire, Sér. A, Sci. Nat. 16: 330. 1954, nom. illeg., non Coleus urticifolius Benth. Type: Malawi, ex Hort. Kew, seeds sent from Mt Zomba by Livingstone (holotype: K).
Pycnostachys pubescens Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 345 (1895), non Coleus pubescens Merr.
Pycnostachys urticifolia var. pubescens (Gürke) Gürke, Bot. Jahrb. Syst. 22: 146 (1895). Types: Mozambique, Gorungosa, Carvalho s.n. (syntype: B, destroyed; isosyntype:, COI); Malawi, without precise locality, Buchanan s.n. (syntype: B, destroyed).
Distribution.
Tanzania to South Africa.
Coleus longipetiolatus
Gürke, Bot. Jahrb. Syst. 19: 214. 1894
3C3F76A4C12A5697AD4D42D747DD34A1
Coleus longipetiolatus Gürke, Bot. Jahrb. Syst. 19: 214. 1894. Type: Tanzania, Usambara Mts, Kwa Mshusa, 15.Aug.1893, Holst 9076 (holotype: B destroyed; isotypes: COI, G, K, M, P, W).
Coleus leptophyllus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 427. 1900.
Plectranthus leptophyllus (Baker) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 284. 2009. Type: Kenya, Ribe (Ribi) to Galla country, Wakefield s.n. (holotype: K).
Coleus swynnertonii S.Moore, J. Linn. Soc., Bot. 40: 177. 1911.
Plectranthus chiridensis Codd, Fl. Pl. Africa 48: t. 1887. 1984. Type: Zimbabwe, Chirinda, Swynnerton 349 (holotype: BM; isotype K).
Distribution.
E. & S. Trop. Africa.
Coleus lyratus
(A.Chev.) Roberty, Bull. Inst. Fondam. Afrique Noire, Sér. A, Sci. Nat. 16: 330. 1954
5A119CAA57A95D6FB58EF4CBD99A7729
Leocus lyratus A.Chev., J. Bot. (Morot) 22: 126. 1909. Type: Guinea, Fouta-Djalon, Mt Tinka, near Dalaba, Chevalier s.n. (holotype: P).
Distribution.
Sierra Leone, Guinea.
Notes.
Paton et al. (2013) considered this as a synonym of P. betonicifolius, but the west African C. lyratus is a larger plant with retrorse, rather than antrorse stem hairs.
Coleus macranthus
Merr., Philipp. J. Sci. 1(suppl.): 234. 1906
351CFE2C23BC5E978C34183126313B5D
Plectranthus merrillii H.Keng, Fl. Males. 8: 391. 1978. Types: Phillippines, Luzon, Mt Data, Merrill 4502, 4483, 4505 (isosyntypes: K).
Distribution.
Philippines (N. Luzon).
Coleus maculosus
(Lam.) A.J.Paton comb. nov.
FA000AE425CD58E0AABD549EE20E2611
urn:lsid:ipni.org:names:77201133-1
Galeopsis maculosa Lam., Encycl. 2: 601. 1788.
Germanea maculosa Lam., Encycl. 2: 691. 1788.
Plectranthus maculosus (Lam.) Salisb., Prodr. Stirp. Chap. Allerton: 88. 1796. Type: cultivated, from Africa (holotype: P-Lam).
Ocimum punctatum L.f., suppl. Pl.: 275. 1782, non Coleus punctatus Baker.
Plectranthus punctatus (L.f.) L’Hér., Stirp. Nov.: 87. 1788.
Germanea punctata (L.f.) Poir. in J.B.A.M.de Lamarck, Encycl. 2: 763. 1788.
Coleus praetermissus Bullock & Killick, Taxon 6: 239. 1957.
Plectranthus punctatus L’Hér. subsp. punctatus . Fl. Trop. E. Afr., Lamiac.: 350. 2009. Type: “Abyssinia”, cultivated from seed sent by Bruce at Hort. Kew, 1774, (neotype: BM; isoneotype; G; designated by Hedge et al. (1998)).
Distribution.
Widespread. Trop. Africa, Madagascar.
Coleus maculosus subsp. maculosus
8425A5CDD44050BC8E1B56D132352BF9
Coleus glandulosus Hook.f., J. Proc. Linn. Soc., Bot. 7: 211. 1864.
Majana glandulosa (Hook.f.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Types: Cameroon, Cameroon Mt, Mann 1301 & 1988 (syntypes: K).
Coleus doba Hochst. ex Chiov., Ann. Bot. (Rome) 9: 126. 1911. Types: Ethiopia, Tigre, near Adua, 1852, Schimper 2040 (syntype: P; isosyntype: TUB); Semien, Dabarek, Chiovenda 969 (syntype: FT); Dembia, Gondar, Chiovenda 1500 (syntype: P).
Coleus serrulatus Robyns, Bull. Jard. Bot. État Bruxelles 17: 78. 1943.
Plectranthus serrulatus (Robyns) Troupin & Ayob., Bull. Jard. Bot. Natl. Belg. 55: 299. 1985. Type: DRC, R. Susa, de Witte 2214 (isosyntype: K).
Distribution.
Cameroon, to Eritrea and Tanzania, Madagascar.
Coleus maculosus subsp. edulis
(Vatke) A.J.Paton comb. nov.
F812ECA72D2859EBB4D4EA3ACBE99D63
urn:lsid:ipni.org:names:77201134-1
Coleus edulis Vatke, Linnaea 37: 319. 1872.
Plectranthus edulis (Vatke) Agnew, Upland Kenya Wild Fl.: 640. 1974.
Plectranthus punctatus subsp. edulis (Vatke) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 350. 2009. Type: Ethiopia, near Gaffat, 1863, Schimper 1212 (isotype: K, Z, mixed with C. barbatus).
Coleus tuberosus A.Rich., Tent. Fl. Abyss. 2: 185. 1850., nom. illeg., non C. tuberosus (Blume) Benth.
Majana richardiana Kuntze, Revis. Gen. Pl. 2: 524. 1891., nom. illeg., based on above.
Coleus palustris Vatke, Linnaea 37: 319. 1872. Type: Ethiopia, Daschan Meda, 15 Sept. 1863, Schimper 1186 (holotype: B, destroyed; isotypes: BM, JE, K, PRE, W, Z).
Coleus rivularis Vatke, Linnaea 37: 320. 1872. Type: Ethiopia, Gaffat, 25 Sept. 1863, Schimper 1172 (holotype: B, destroyed; isotypes: BM, JE, K, PRE, US, W, Z).
Coleus aquaticus Gürke, Bot. Jahrb. Syst. 19: 218. 1894. Types: Tanzania, Kilimanjaro, Marangu, Volkens 583 (syntype: B, destroyed); Mawenzi, Volkens 860 (syntype: B, destroyed; isosyntype: E).
Coleus clivicola S.Moore, J. Linn. Soc., Bot. 38: 274. 1908. Type: Uganda, E Ruwenzori, Wollaston s.n. (holotype: BM).
Coleus fimbriatus Lebrun & L.Touss., Bull. Jard. Bot. État Bruxelles 17: 79. 1943.
Plectranthus fimbriatus (Lebrun & L.Touss.) Troupin & Ayob., Bull. Jard. Bot. Natl. Belg. 55: 299. 1985. Type: DRC, Mt Mushumangabo, Lebrun 7163 (holotype: BR).
Distribution.
Ethiopia, Kenya, Uganda Tanzania, Burundi, Rwanda and DRC.
Coleus maculosus subsp. lanatus
(J.K.Morton) A.J.Paton comb. nov.
C24583B1586F57AEBFD093BA2407955D
urn:lsid:ipni.org:names:77201135-1
Plectranthus punctatus subsp. lanatus J.K.Morton, J. Linn. Soc., Bot. 58: 268. 1962. Type: Cameroon, Bamenda, near Bambui, Bamenda, Adams GC11322 (holotype: GC).
Distribution.
Cameroon.
Notes.
Paton et al. (2009, 2013) considered this as a synonym of subsp. edulis, but the prominent indumentum and disjunction warrant separate recognition.
Coleus madagascariensis
(Pers.) A.Chev., Rev. Int. Bot. Appl. Agric. Trop. 33: 338. 1953
46D6BBCF9C7E593D8AAB9FAC18A7FDED
Ocimum madagascariense Pers., Syn. Pl. 2: 135. 1806.
Plectranthus madagascariensis (Pers.) Benth., Labiat. Gen. Spec.: 37. 1832. Type: Mauritius or Reunion, n.d., Commerson 266 (holotype: P-JU 5706).
Plectranthus hirtus Benth., Labiat. Gen. Spec.: 38. 1832. Type: South Africa, Cape, n.d., Masson s.n. (holotype: BM).
Plectranthus villosus Sieber ex Benth., Labiat. Gen. Spec.: 38. 1832. Type: Mauritus, Sieber, Fl. Maurit. Exsic. 152 (holotype: K; isotype: P).
Plectranthus mauritianus Bojer, Hortus Maurit.: 254. 1837., nom. inval.
Plectranthus pubescens Willd. ex Benth., Linnaea 11: 344. 1837., nom. inval.
Ocimum auricula Forssk. ex Benth. in A.P.de Candolle, Prodr. 12: 43, 68. 1848., nom. inval.
Distribution.
Mozambique to South Africa, Mascarenes.
Coleus malabaricus
Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830
AD94126416F85E06B8C4B299E29434D1
Plectranthus malabaricus (Benth.) R.H.Willemse, Blumea 25: 509. 1979. Type: India, Travancore, Wall. Num. List 2735A, in Hb. Madr. (holotype: K-W (K001116979)).
Coleus macraei Benth., Labiat. Gen. Spec.: 58. 1832. Type: Ceylon, Macrae s.n., (holotype: K(K000820112)).
Coleus ovatus Benth., Labiat. Gen. Spec.: 57. 1832. Type: India, in Coromandelia, Wright, s.n. (holotype: K (K000820121)).
Coleus leptostachys Benth. in A.P.de Candolle, Prodr. 12: 77. 1848.
Plectranthus malabaricus var. leptostachys (Benth.) R.H.Willemse, Blumea 25: 510. 1979. Type: Ceylon, Walker s.n., (holotype: K (K000820114)).
Coleus walkeri Benth. in A.P.de Candolle, Prodr. 12: 77. 1848. Type: Ceylon, Walker s.n., in Hb, Benth. (holotype: K (K000820113)).
Distribution.
S. India, Sri Lanka.
Coleus mannii
Hook.f., J. Proc. Linn. Soc., Bot. 7: 211. 1864
BC2C6A350429562BBFB7D1196C422EFB
Majana mannii (Hook.f.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Solenostemon mannii (Hook.f.) Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 422. 1900.
Plectranthus occidentalis B.J.Pollard, Kew Bull. 60: 146. 2005. Type: Mt Cameroon, Mann 1967 (holotype: K).
Solenostemon cymosus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 422. 1900. Type: Cameroon, Mt. Cameroon, Mann 1251 (holotype: K).
Coleus giorgii De Wild., Bol. Soc. Ibér. Ci. Nat. 19: 120. 1920.
Solenostemon giorgii (De Wild.) Champl., Syst. Geogr. Pl. 69: 16. 1999. Type: DRC, Likimi, De Giorgi 1511 (BR: lectotype, designated by Champluvier & Dowsett-Lemaire (1999)). synon. nov.
Distribution.
W. Trop. Africa to DRC and Sudan.
Coleus marrubatus
(J.K.Morton) A.J.Paton comb. nov.
C74111B00143578EB3D31943FDF9BD77
urn:lsid:ipni.org:names:77201136-1
Plectranthus marrubatus J.K.Morton, Kew Bull. 53: 997. 1998 publ. 1999. Type: Ethiopia, Mega, Gillett 14206 (holotype: K; isotype: FT).
Distribution.
S. Ethiopia to N. Kenya.
Coleus megacalyx
(A.J.Paton) A.J.Paton comb. nov.
3F2F63242D1C5B7FAE5752C2EB80DC49
urn:lsid:ipni.org:names:77201137-1
Plectranthus megacalyx A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 316. 2009. Type: Tanzania, Rungwe District: Ngozi, Geilinger 2884 (holotype: K).
Distribution.
SW. Tanzania to E. Zambia.
Coleus megadontus
(P.I.Forst.) P.I.Forst. comb. nov.
038BD6CD3587541B91565E2855625A96
urn:lsid:ipni.org:names:77201138-1
Plectranthus megadontus P.I.Forst., Austrobaileya 4: 181. 1994. Type: Australia, Queensland, Cook District, Rocky River Scrub, eastern fall of McIlwraith Range, 16 June 1992, P.I.Forster PIF10637A, M.C.Tucker & G.Kenning (holotype: BRI; isotypes: CNS, K, L, MEL).
Distribution.
Australia: Queensland.
Coleus melleri
(Baker) A.J.Paton & Phillipson comb. nov.
AF9DC9C2C6F054C085C30AC1CEEE072A
urn:lsid:ipni.org:names:77201145-1
Plectranthus melleri Baker, J. Bot. 20: 243. 1882. Type: Madagascar, between Toamasina (Tamatave) and Antananarivo, July–August 1862, Meller s.n. (holotype: K).
Plectranthus luteus Gürke, Bot. Jahrb. Syst. 28: 468. 1900.
Coleus luteus (Gürke) Staner, Bull. Agric. Congo Belge 25: 426. 1934. Type: Tanzania, Morogoro District: SE Uluguru Mts, Stuhlmann 8790 (holotype: B, destroyed).
Coleus entebbensis S.Moore, J. Bot. 44: 89. 1906. Type: Uganda, Mengo District: Entebbe, Bagshawe 701 (holotype: BM).
Distribution.
Liberia, Gabon, Uganda to S. Trop. Africa, Madagascar.
Coleus meyeri
(Gürke) A.J.Paton comb. nov.
7F9AA388322752AA808E30D59E924BE8
urn:lsid:ipni.org:names:77201139-1
Pycnostachys meyeri Gürke, Abh. Königl. Akad. Wiss. Berlin 1891: 362. 1892. Type: Tanzania, Kilimanjaro, Rua stream, Meyer 279 (holotype: B, destroyed).
Pycnostachys volkensii Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 344. 1895. Type: Tanzania, Kilimanjaro, Mawenzi, Volkens 823 (lectotype: K, designated by Bramley in Paton et al. (2009); isolectotype: BM).
Pycnostachys oblongifolia Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 385. 1900. Type: Uganda, Toro District: Ruimi (Wimi) Valley, Scott Elliot 7883 (lectotype: K, designated by Bruce (1940)).
Pycnostachys bowalensis A.Chev., J. Bot. (Morot) 22: 126. 1909. Type: Guinea, Conakry, Fouta-Djalon, Kala, Diaguissa, Oct. 1907, A. Chevalier 18853 (holotype: P; isotype: K).
Pycnostachys longiacuminata Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 76. 1921. Type: Ethiopia, Sidamo, Lake Abassa, (?Lake Margharita), Dec. 1900, Ellenbeck 1728a (holotype: B, destroyed).
Pycnostachys longibracteata De Wild., Pl. Bequaert. 4: 388. 1928. Type: DRC, Ruwenzori, Lanuri Valley, Bequaert 4490 (holotype: BR; K fragment).
Pycnostachys ovoideoconica De Wild., Pl. Bequaert. 4: 396. 1928. Type: DRC, Mukule–Mokoto, Bequaert 6325 (holotype: BR; K, fragment).
Distribution.
W. Trop. Africa to Ethiopia and Tanzania.
Coleus minor
(J.K.Morton) A.J.Paton comb. nov.
2C3E937AADB3541BA7F2DC6E102A95A0
urn:lsid:ipni.org:names:77201140-1
Solenostemon minor J.K.Morton, J. Linn. Soc., Bot. 58: 271. 1962. Type: Nigeria, West Kagoro Hills, Jos Plateau, Killick 15 (holotype: K).
Plectranthus decumbens Hook.f., J. Proc. Linn. Soc., Bot. 7: 210. 1864., non Coleus decumbens Gürke.
Solenostemon decumbens (Hook.f.) Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 421. 1900.
Calchas decumbens (Hook.f.) P.V.Heath, Calyx 5: 160. 1997. Type: Cameroon, Mt Cameroon, Mann 2002 (holotype: K).
Distribution.
WC. Trop. Africa.
Coleus minutiflorus
(Ryding) A.J.Paton comb. nov.
0C2FB604406E501CA49D75B20F75DB05
urn:lsid:ipni.org:names:77201141-1
Plectranthus minutiflorus Ryding, Nordic J. Bot. 20: 43. 2000. Type: Ethiopia, Sidamo, ± 35 km from the main road Yabelo–Mega, on road to Arero, Friis, Bidgood, L. & W. Malaku & Gashaw 8374 (holotype: C; isotypes: ETH, K).
Distribution.
Ethiopia to N. Kenya.
Coleus minutus
(P.I.Forst.) P.I.Forst. comb. nov.
18223A189CC65C0F9CB76AEFF6FAC9F0
urn:lsid:ipni.org:names:77201142-1
Plectranthus minutus P.I.Forst., Austrobaileya 3: 734. 1992. Type: Australia, Queensland, Cook District, Mount Muligan, c 40 km NW of Dimbulah, c. 0.5 km SE of Dam on top of mountain, 17 April 1985, J.R.Clarkson 5902 (holotype: BRI; isotypes: CNS, MEL).
Distribution.
Australia: Queensland.
Coleus mirabilis
Briq., Bot. Jahrb. Syst. 19: 183. 1894
60BAB9AC8F24557F81D9F0BEE60CA738
Coleus mirabilis var. hypisodontus Briq., Bot. Jahrb. Syst. 19: 184. 1894., nom. inval.
Ascocarydion mirabile (Briq.) G.Taylor, J. Bot. 69(suppl. 2): 162. 1931.
Burnatastrum mirabile (Briq.) P.V.Heath, Calyx 4: 175. 2001.
Plectranthus mirabilis (Briq.) Launert, Mitt. Bot. Staatssamml. München 7: 299. 1968. Type: Angola, Malanje (Malandsche), Mar. 1880, Mechow 489 (lectotype: Z isolectotype: W; designated by Codd (1975)).
Coleus mirabilis var. buchnerianus Briq., Bot. Jahrb. Syst. 19: 184. 1894. Types: Angola, Moma, N of Malanje (Malandsche), Buchner 81, 82, 83, 84 & 85 (syntypes: B, destroyed).
Coleus mirabilis var. mecjowianus Briq., Bot. Jahrb. Syst. 19: 184. 1894. Type: Angola, Lumsuneja stream, between Malanje (Malandsche) and Cuango (Quango) rivers, June1880, Mechow s.n. (isotype: Z).
Coleus mirabilis var. poggeanus Briq., Bot. Jahrb. Syst. 19: 184. 1894. Type: Congo, R.Lulua, 9°30'S, Pogge 350 (holotype: B, destroyed).
Coleus leucophyllus Baker, Bull. Misc. Inform. Kew 1895: 292. 1895. Type: Zambia, near Mwero, W of Lake Tanganyika, 1894, Carson 26 (holotype: K).
Distribution.
S. DRC to NE. Namibia.
Coleus mirus
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
90CA9B1F6A835041A5447A6886480D6F
urn:lsid:ipni.org:names:77201143-1
Plectranthus mirus S.T.Blake, Contr. Queensland Herb. 9: 22. 1971. Type: Australia, Queensland, White Cliff Point NW of Cairns, 21 May 1962, S.T.Blake 21791 (holotype: BRI).
Distribution.
Australia: Queensland.
Coleus mitis
(R.A.Clement) A.J.Paton comb. nov.
866971CB54135D5187712EA321F0E187
urn:lsid:ipni.org:names:77201144-1
Anisochilus mitis R.A.Clement, Edinburgh J. Bot. 50: 35. 1993. Type: Bhutan, Khoma, open spaces, 1219 m, 21 July 1949, Ludlow, Sherriff & Hicks 20927 (holotype: BM).
Distribution.
E. Himalaya.
Coleus modestus
(Baker) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 106. 1929
921DB52C900C5A4EB562F84B44B407A1
Plectranthus modestus Baker, Bull. Misc. Inform. Kew 1895: 72. 1895.
Englerastrum modestum (Baker) T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 76. 1924. Type: Zambia, Mbala District: Tanganyika Plateau, 1889, Carson s.n. (holotype: K).
Distribution.
SW. Tanzania to Zambia.
Coleus mollis
Benth. in A.P.de Candolle, Prodr. 12: 77. 1848
211CCC4C4F5452D4A06F0CADFF1CA049
Plectranthus crameri R.H.Willemse, Blumea 25: 510. 1979. Type: Ceylon, Walker s.n., in Hb. Benth. (lectotype: K (K000820116), designated by Cramer (1978)).
Distribution.
Sri Lanka.
Coleus monostachyus
(P.Beauv.) A.J.Paton comb. nov.
745CBD14B860517AA90EBDC219920D59
urn:lsid:ipni.org:names:77201146-1
Ocimum monostachyum P.Beauv., Fl. Oware 2: 60. 1818.
Plectranthus palisotii Benth., Labiat. Gen. Spec.: 39. 1832., nom. illeg.
Solenostemon monostachyus (P.Beauv.) Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 359. 1897.
Solenostemon ocymoides var. monostachyus (P.Beauv.) Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 421. 1900.
Plectranthus monostachyus (P.Beauv.) B.J.Pollard, Kew Bull. 56: 980. 2001. Type: Benin, Palisot de Beauvois s.n. (holotype: G).
Coleus monostachyus
(P.Beauv.) A.J.Paton subsp. monostachyus
A2842C217AF15A2EAD8B73B80EDA21AC
Solenostemon ocymoides Schumach. & Thonn. in C.F.Schumacher, Beskr. Guin. Pl.: 271. 1827.
Coleus africanus Benth., Labiat. Gen. Spec.: 54. 1832., nom. illeg.
Solenostemon africanus Briq., Bot. Jahrb. Syst. 19: 181. 1894. Type: “Guinea”, Thonning 92 (lectotype: C, designated by Pollard & Paton (2001); Isert & Thonning herbarium, microf. 102: III. 7).
Solenostemon monostachyus var. amplifrons Briq., Bull. Herb. Boissier, sér. 2, 6: 826. 1906. Type: DRC, Bangala, 26 May 1888, Hens Ser. C 33 (holotype: Z).
Solenostemon godefroyae N.E.Br., Bot. Mag. 139: t. 8511. 1913. Type: illustration cited above (lectotype: illustration designated here).
Coleus godfroyae Godfroy-Lebeuf, Cat Pl. Nouv.: 2 1903. Type: not seen. Protologue of S. godefoyae suggests the name C. godfroy exists (with drawing), but this manuscript has not been seen.
Distribution.
W. Trop. Africa to S. Chad and Cabinda.
Coleus monostachyus
(P.Beauv.) A.J.Paton subsp. marrubiifolius (Brenan) A.J.Paton comb. nov.
573CB6F3E3F15EFF97D8343A400B0849
urn:lsid:ipni.org:names:77201147-1
Solenostemon monostachyus var. marrubiifolius Brenan, Kew Bull. 5: 231. 1950.
Solenostemon monostachyus subsp. marrubiifolius (Brenan) J.K.Morton, J. Linn. Soc., Bot. 58: 272. 1962.
Plectranthus monostachyus subsp. marrubiifolius (Brenan) B.J.Pollard, Kew Bull. 56: 980. 2001. Type: Nigeria, Ondo Province, Carter’s Peak, Idanre Hills, Keay FHI 22585 (holotype: K).
Solenostemon gouanensis A.Chev., Explor. Bot. Afrique Occ. Franç. 1: 517. 1920., nom. nud.
Distribution.
W. & WC. Trop. Africa.
Coleus monostachyus
(P.Beauv.) A.J.Paton subsp. perennis (J.K. Morton) A.J.Paton comb. nov.
EDD774B6B219534FB89E817061419B18
urn:lsid:ipni.org:names:77201148-1
Solenostemon monostachyus (P.Beauv.) Briq subsp. perennis J.K.Morton, J. Linn. Soc., Bot. 58: 272. 1962.
Plectranthus monostachyus (P.Beauv.) Briq subsp. perennis (J.K.Morton) B.J.Pollard, Kew Bull. 56: 981. 2001. Type: Ghana, Northern Province, Damongo Scarp, Morton A1242 (holotype: GC, not seen).
Distribution.
Sierra Leone, Ivory Coast, Ghana and Nigeria.
Coleus mutabilis
(Codd) A.J.Paton comb. nov.
0C2D11AC139154ECB01F183EE93ACC9A
urn:lsid:ipni.org:names:77201149-1
Plectranthus mutabilis Codd, Bothalia 11: 404. 1975. Type: South Africa, Transvaal, Pietersburgh District, farm Bulbul, eastern end of Blouberg, Codd 7953 (holotype: PRE).
Distribution.
South Africa: Northern Prov.
Coleus myrianthellus
Briq., Mém. Soc. Bot. France 8: 290. 1917
2B8FB18BBCC258FCA6241317665AA0E5
Coleus myrianthellus Briq., Mém. Soc. Bot. France 8: 290. 1917. Type: Central African Republic, Gribingui, bassin de la Moyenne Kododo, 29 Nov. 1902, Chevalier 6454 (holotype: P).
Distribution.
Central African Republic.
Notes.
Related to C. splendidus. A revision of this group is required.
Coleus namuliensis
E.Downes & I.Darbysh., Blumea 62: 169. 2017
C220D1D74EAB535992E934978CB7195B
Coleus namuliensis E.Downes & I.Darbysh., Blumea 62: 169. 2017. Type: Mozambique, Zambezia Province, Mt Namuli, small peak 5 km west of highest peak, Harris 237 (holotype: K (K000962083); isotype K(K000962089), LMA).
Distribution
. Mozambique.
Coleus neochilus
(Schltr.) Codd, Bothalia 7: 432. 1961
73DC9EA214325C039147129FD25C2F61
Plectranthus neochilus Schltr., J. Bot. 34: 394. 1896. Type: South Africa, Mpumalanga, Rimero Creek near Baberton, 1890, Galpin 968 (holotype: K; isotypes: BOL, GRA, NH).
Coleus schinzii Gürke, Bull. Herb. Boissier 6: 555. 1898. Type: Namibia, Ovamboland, Tsumeb, Schinz 56 (holotype: Z).
Coleus palliolatus S.Moore, J. Bot. 38: 464. 1900. Type: Zimbabwe, Bulawayo, Jan.1898, Rand 144 (holotype: BM).
Distribution.
S. Trop. & South Africa.
Coleus nepetifolius
(Baker) A.J.Paton comb. nov.
613E0B752BBD582AAC93863A4E9F116B
urn:lsid:ipni.org:names:77201150-1
Pycnostachys nepetifolia Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 383. 1900. Type: Kenya, Nandi/North Kavirondo District: Nandi Range, Scott Elliot 6471 (holotype: K; isotype: BM).
Distribution.
Kenya.
Coleus niamniamensis
(Gürke) A.J.Paton comb. nov.
165F40FF8A9A5F03ADB887561BDA7A3D
urn:lsid:ipni.org:names:77201151-1
Pycnostachys niamniamensis Gürke, Bot. Jahrb. Syst. 22: 145. 1895. Type: Sudan, Niamniam, east of R. Huuh, Schweinfurth 3750a (holotype: B, destroyed; isotype: UPS, K, fragment).
Pycnostachys decussata Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 382. 1900. Type: Kenya, ‘Kavirondo’, Scott Elliot 7155 (holotype: K).
Pycnostachys petherickii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 383. 1900. Type: Sudan, White Nile, Petherick s.n. (holotype: K).
Pycnostachys lindblomii T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 24. 1930. Type: Kenya, North Kavirondo District: Elgon, Kitosh, Lindblom s.n. (holotype: S; K, fragment).
Distribution.
Sudan to Kenya.
Coleus nigericus
(Alston) A.J.Paton comb. nov.
084225E8AF97531AA639D48B734630B0
urn:lsid:ipni.org:names:77201152-1
Englerastrum nigericum Alston, Bull. Misc. Inform. Kew 1926: 298. 1926. Type: Nigeria, Northern Nigeria, Katagum District, Dalziel 109 (holotype: K).
Plectranthus denudatus A.Chev. ex Hutch. & Dalziel, Fl. W. Trop. Afr. 2: 288. 1931.
Coleus denudatus (A.Chev. ex Hutch. & Dalziel) Robyns, Mem. Inst. Roy. Colon. Belge, Sect. Sci. Nat. (4°): 50. 1936. Type: Senegal, Diola (Casamance), Flory-Feygan, Chevalier 2815 (holotype: P).
Distribution.
W. Trop. Africa.
Notes.
Non C. nigericus A.Chev., nom. nud.
Coleus nitidus
(P.I.Forst.) P.I.Forst. comb. nov.
769DF8D0DB07503CB8F5C0F6BCD1CF67
urn:lsid:ipni.org:names:77201153-1
Plectranthus nitidus P.I.Forst., Austrobaileya 3: 736. 1992. Type: Australia, New South Wales, cultivated at Byron Bay, from plant collected at Terania Creek, Nightcap Range, P. Hardwick s.n. (holotype: BRI; isotypes: CNS, K, L, MEL, NSW).
Distribution.
E. Australia.
Coleus niveus
(Hiern) A.J.Paton comb. nov.
9CDF0B6134B95023B81AB795012D2D21
urn:lsid:ipni.org:names:77201154-1
Solenostemon niveus Hiern, Cat. Afr. Pl. 1: 864. 1900 (August). Type: Angola, Bumbo, Sierra de Xella, Welwitsch 5619 (holotype: LISU; isotype: BM).
Coleus orbicularis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 437. 1900. (December). Type: Angola, Benguella Prov., Welwitsch 5619, (holotype: K).
Distribution.
Angola.
Coleus nyikensis
Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 439. 1900
8A8084F5BF975E638D75BF627E1F921A
Plectranthus malawiensis B.Mathew, Kew Bull. 31: 174. 1976. Type: Malawi, Nyika Plateau, July 1896, Whyte s.n. (holotype: K).
Distribution.
SE. Kenya to N. Zambia.
Coleus omissus
(P.I.Forst) P.I.Forst. comb. nov.
882A9F36D6D854D2A4D296F546A1FB0C
urn:lsid:ipni.org:names:77201155-1
Plectranthus omissus P.I.Forst., Austrobaileya 3: 731. 1992. Type: Australia, Queensland, Wide Bay District, Mudlow Gap, Beauty Spot 11, Timber reserve 26, 8 km N of Kilkivan, 21 Feb. 1989, P.I. Forster PIF4977 (holotype: BRI; isotypes: CANB, K, MO).
Distribution.
Australia: Queensland.
Coleus orthodontus
(Gürke) A.J.Paton comb. nov.
FD831479F1425B89B8A29B4F200643BF
urn:lsid:ipni.org:names:77201156-1
Pycnostachys orthodonta Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 345. 1895. Type: Mozambique, Manica, near Catandica (Gouveia), 1884, Carvalho s.n. (holotype: B, destroyed; isotypes: COI, K frag.).
Pycnostachys cyanea Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 345. 1895. Type: Tanzania, Mpwapwa, Tubugwe, Stuhlmann 213 (lectotype: B, destroyed; designated by Bramley in Paton et al. (2009)).
Pycnostachys leptophylla Baker, Bull. Misc. Inform. Kew 1898: 161. 1898. Type: Malawi, Shire Highlands, 1891, Buchanan 873 (lectotype: K; designated by Bramley in Paton et al. (2009)).
Pycnostachys hanningtonii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 384. 1900. Type: Tanzania, Dodoma Dist., Ugogo, n.d., Hannington s.n. (holotype: K).
Pycnostachys bussei Gürke, Bot. Jahrb. Syst. 36: 131. 1905. Type: Tanzania, Lindi Dist., near Nguya, n.d., Busse 2834 (holotype: B, destroyed; isotypes: BR, K frag.).
Distribution.
Tanzania to S. Trop. Africa.
Coleus otostegioides
(Gürke) A.J.Paton comb. nov.
38BDAF7F870B595D996D24C2D0ADB186
urn:lsid:ipni.org:names:77201157-1
Capitanya otostegioides Gürke, Bot. Jahrb. Syst. 21: 106. 1895.
Plectranthus otostegioides (Gürke) Ryding, Kew Bull. 54: 126. 1999. Types: Eritrea, upper Lebka valley, Heuglin s.n. (syntype: B, destroyed) & Müller-Prosko-Capitany s.n. (syntype: B, destroyed) & Tanzania, Kilimanjaro, Volkens 492 (syntype: B, destroyed); neotype: Tanzania, Arusha District: Ngaserai Plain, Richards & Arasululu 26472 (neotype: K, designated by Ryding (1999)).
Distribution.
NE. Trop. Africa to N. Tanzania.
Coleus pallidus
(Wall.) A.J.Paton comb. nov.
4D7DA40F660D5F87974BC9D9120D2E61
urn:lsid:ipni.org:names:77201158-1
Anisochilus pallidus Wall., Pl. Asiat. Rar. 2: 18. 1830. Type: Burma, Taong Dong, 26 Nov. 1826, Wallich Cat. 2754, (lectotype: K (K000674759); isolectotypes: G-DC, K-W (K001117031), designated by Suddee and Paton (2009)).
Distribution.
Nepal to SC. China and Indo-China.
Coleus paniculatus
Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830
83E0C0F7F07E5E25841EB3389DA34F55
Majana paniculata (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Solenostemon paniculatus (Benth.) Guillaumet & A.Cornet, Adansonia, n.s., 15: 525. 1976. Type: India, Dindygul, Herb. Wight in Wall. Cat. 2734 (syntypes: K (K000820126); K-W(K001116978).
Coleus glabratus Benth., Labiat. Gen. Spec.: 58. 1832.
Plectranthus glabratus (Benth.) Alston in H.Trimen, Handb. Fl. Ceylon 6(suppl.): 236. 1931. Type: India, Madras, Wight s.n. ((holotype: K (K000820127)).
Coleus wightii Benth., Labiat. Gen. Spec.: 58. 1832. Type: India, Nilgiris, Wight s.n. (holotype: K (K000820125)).
Plectranthus coleoides Benth. in A.P.de Candolle, Prodr. 12: 64. 1848. Type: India, Nilgiris, Perrottet s.n. (syntypes: 2 shts. G-DC; isosyntype: K (K000820123), P (P00720715).
Plectranthus bernardii Doan in H.Lecomte, Fl. Indo-Chine 4: 949. 1936 nom. inval. no Latin descr.
Distribution.
S. India to S. Indo-China.
Coleus parishii
(Hook.f.) A.J.Paton comb. nov.
C987EA16264F5219B0372FEA6A0595D2
urn:lsid:ipni.org:names:77201159-1
Plectranthus parishii Hook.f., Fl. Brit. India 4: 622. 1885. Type: Burma, Tenasserim, Moulmein, Parish 444 (holotype: K).
Calamintha esquirolii H.Lév., Repert. Spec. Nov. Regni Veg. 8: 450. 1910.
Coleus esquirolii (H.Lév.) Dunn, Notes Roy. Bot. Gard. Edinburgh 8: 158. 1913. Type: China, Kweichow, Shin- gny-hien, chemin pierreux de Sy-Koua-Tong, Nov. 1906, Esquirol 1058 (holotype: E).
Coleus mucosus Hayata, J. Coll. Sci. Imp. Univ. Tokyo 30(1): 225. 1911. Types: Kodensho, Oct. 1906, Kawakami & Mori 1755 (TI: syntype); Kashito, Aug. 1905, Nakahara 302 (TI: syntype); Banchoryo, Oct. 1905, Nakahara 582, Oct. 1905 (TI: syntype); Kodensho, Oct. 1905, Nagasawa 764 (TI: syntype)
Distribution.
Myanmar to SC. China and Taiwan.
Notes.
See Suddee et al. (2004) for discussion of C. mucosus.
Coleus parvicalyx
(A.J.Paton) A.J.Paton comb.nov.
654403345AC451A79106F2F673D554F9
urn:lsid:ipni.org:names:77201160-1
Plectranthus parvicalyx A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 316. 2009. Type: Tanzania, Rungwe District: Kiejo Volcano, Hepper, Field & Mhoro 5389 (holotype: K).
Distribution.
SW. Tanzania.
Coleus penicillatus
(A.J.Paton) A.J.Paton comb. nov.
309EED11F8CA595D9DEFF1464F9CA9C5
urn:lsid:ipni.org:names:77201161-1
Plectranthus penicillatus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 325. 2009. Type: Zambia, Mbuzi–Kaluza, Greenway & Trapnell 5627 (holotype: K).
Distribution.
S. Tanzania to E. Zambia.
Coleus pentheri
Gürke, Ann. K. K. Naturhist. Hofmus. 20: 48. 1905
CAFDFA700761557F83C18D557F2CD874
Plectranthus pentheri (Gürke) van Jaarsv. & T.J.Edwards, Bothalia 27: 5. 1997. Type: South Africa, Mpumalanga, Rimero Creek near Baberton, 1890, Galpin 968 (holotype: K; isotypes: BOL, GRA, NH).
Distribution.
SW. Zimbabwe to South Africa.
Coleus perrieri
(Hedge) A.J.Paton & Phillipson comb. nov.
EB1F1536B9E15629A8BB5DB500E70364
urn:lsid:ipni.org:names:77201162-1
Plectranthus perrieri Hedge, Fl. Madag. 175: 199. 1998. Type: Madagascar, Ampandrandava, Seyrig 637 (holotype: P).
Distribution.
Madagascar.
Coleus persoonii
Benth., Labiat. Gen. Spec.: 55. 1832
C1D7573655165365B03267CCEF60E0BD
Ocimum paniculatum Pers., Syn. Pl. 2: 135. 1806, non Coleus paniculatus Benth.
Solenostemon paniculatus (Pers.) Guillaumet & Cornet, Adansonia n.s., 15: 525. 1976
Plectranthus persoonii (Benth.) Hedge, Fl. Madag. 175: 222. 1998. Type: Madagascar, Commerson s.n. (holotype: P).
Coleus goudotii Briq., Bull. Herb. Boissier 2: 126. 1894.
Solenostemon goudotii (Briq.) Guillaumet & A.Cornet, Adansonia, n.s., 15: 525. 1976. Type: Madagascar, Goudot s.n. (G, holotype).
Distribution.
Madagascar.
Coleus petiolatissimus
Briq., Mém. Soc. Bot. France 8: 291. 1917
37784DC39BC854AC8E63993C1D842142
Coleus petiolatissimus Briq., Mém. Soc. Bot. France 8: 291. 1917. Type: Central African Republic (Bangui), 16 Aug. 1902, Chevalier 5214 bis. (holotype: P).
Distribution.
Central African Republic.
Notes.
Related to P. bojeri. Complex in need of revision.
Coleus petraeus
(Backer ex Adelb.) A.J.Paton comb. nov.
46AFA218EC5654B58610F3849FAA9305
urn:lsid:ipni.org:names:77201163-1
Plectranthus petraeus Backer ex Adelb., Reinwardtia 3: 152. 1954. Type: Java, Idjin Plateau, Sempolm, Backer 36387 (holotype: L).
Distribution.
E. Java.
Coleus petricola
(J.Mathew & B.J.Conn) A.J.Paton comb. nov.
249D7D2F54A550A1B05818CADAED43CA
urn:lsid:ipni.org:names:77201164-1
Plectranthus petricola J.Mathew & B.J.Conn, Telopea 20: 179. 2017. Type: India: Kerala: Idukki District, 10 km away from Udumbanchola, Chemmannar Hills, 20 Feb 2010, J. Mathew 4417 (holotype: TBGT; isotype: MSSRF).
Distribution.
India, endemic to Western Ghats.
Coleus phulangkaensis
(Suddee, Suphuntee & Saengrit) Suddee comb. nov.
385942F0C5265C21A5F391A81ECEC515
urn:lsid:ipni.org:names:77201165-1
Plectranthus phulangkaensis Suddee, Suphuntee & Saengrit, Thai Forest Bull., Bot. 42: 6. 2014 (Dec 2014) Type: Thailand. Nakhon Phanom, Ban Phaeng District, Phu Langka National Park, 9 Oct. 2013, Suddee, Puudjaa, Rueangruea, Kiewbang, Hemrat & Pansamrong 4588 (holotype: BKF; isotypes: BKF).
Distribution.
Thailand.
Coleus plantagineus
(Hook.f.) A.J.Paton comb. nov.
8675B0F033865921B928AB0193DDCC63
urn:lsid:ipni.org:names:77201166-1
Anisochilus plantagineus Hook.f., Fl. Brit. India 4: 628. 1885. Type: India, Deccan Peninsula, Mysore, Bababoodan Hills, Dalzell s.n. (lectotype: K(K000674770); isolectotype K(K000674771), designated by Suddee & Paton (2009).
Distribution.
SW. India.
Coleus platyphyllus
(A.J.Paton) A.J.Paton comb. nov.
536ECA77945C5EA587F771753CE0189F
urn:lsid:ipni.org:names:77201167-1
Plectranthus platyphyllus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 331. 2009. Type: Tanzania, Iringa District: Mafinga (Sao Hill) to Madibira road, Richards 17354 (holotype: K; isotype: EA).
Distribution.
SW. Tanzania.
Coleus pobeguinii
Hutch. & Dalziel, Fl. W. Trop. Afr. 2: 291. 1931
99B20744A9955CEEBA70A198C51B49A0
Leocus pobeguinii (Hutch. & Dalziel) J.K.Morton, J. Linn. Soc., Bot. 58: 270. 1962.
Plectranthus pobeguinii (Hutch. & Dalziel) B.J.Pollard, Kew Bull. 64: 260. 2009. Type: Guinea (Conakry), Kouroussa, Pobéguin 556 (P, lectotype, designated by Pollard and Paton (2009)).
Distribution.
W. Trop. Africa.
Coleus polystachys
(Benth.) A.J.Paton comb. nov.
1A2BFD0526605AEE9EFD7DF93FB3A233
urn:lsid:ipni.org:names:77201168-1
Anisochilus polystachys Benth. in N.Wallich, Pl. Asiat. Rar. 2: 19. 1830.
Stiptanthus polystachyus (Benth.) Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 352. 1897. Type: Nepal, Hetoundah (Hetauda), 14 Dec. 1820, Wall. Cat. 2755 (lectotype: K (K000674760); isolectotypes: BM, E, G-DC, K (K000674761), K-W(K001117032), designated Suddee & Paton (2009)).
Distribution.
Nepal to Bangladesh.
Coleus porcatus
(van Jaarsv. & P.J.D.Winter) A.J.Paton comb. nov.
74A21B1A193F5CEB818D5E9869603E3F
urn:lsid:ipni.org:names:77201169-1
Plectranthus porcatus van Jaarsv. & P.J.D.Winter, Bothalia 35: 170. 2005. Type: South Africa, Sekukuniland, N. Leolo Range, Farm de Kamp, P.J.D. Winter 6725 (holotype: PRE; isotype: NBG).
Distribution.
South Africa: Limpopo (Leolo Mts.).
Coleus porphyranthus
(T.J.Edwards & N.R.Crouch) A.J.Paton comb. nov.
1FD5B1637A8958D08A1AAEAE1EDAC29D
urn:lsid:ipni.org:names:77201170-1
Plectranthus porphyranthus T.J.Edwards & N.R.Crouch, Kew Bull. 55: 459. 2000. Type: Zimbabwe, Masvingo Dist., Richmond Farm, Harare road, 4 km from Masvingo, Dec. 1997, Crouch 800 (holotype: NU; isotypes: E, K, PRE, SRGH).
Distribution.
S. Zimbabwe.
Coleus prittwitzii
(Perkins) A.J.Paton comb. nov.
7736BD6436AB5CE7A81B3B01A8F55A57
urn:lsid:ipni.org:names:77201171-1
Pycnostachys prittwitzii Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 68. 1921. Type: Tanzania, Iringa District: Ndembera flood plain, near Gominyi, Prittwitz & Gaffron 28 (holotype: B, destroyed; K, fragment).
Distribution.
SW. Tanzania to N. Zambia.
Coleus prostratus
(Gürke) A.J.Paton comb. nov.
2C7B73AF8B1F50C78C99EB79C76CEDFC
urn:lsid:ipni.org:names:77201172-1
Plectranthus prostratus Gürke, Bot. Jahrb. Syst. 19: 206. 1894. Types: Tanzania, Pangani, Volkens 484 (syntype: B, destroyed; isosyntype: G) & North Mara District: Ukira, Fischer 497 (syntype: B, destroyed).
Plectranthus quadridentatus Schweinf. ex Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 409. 1900. Type: Eritrea, Mount Alam Kale, NW of Aidereso, Schweinfurth & Riva 2086 (holotype: K).
Plectranthus margeritae Buscal. & Muschl., Bot. Jahrb. Syst. 49: 485. 1913. Type: Neumann Camp (Kenya, but in reality based on material from E or NE Africa and Yemen), von Aosta 1688 (holotype: B, destroyed); neotype: Eritrea, Nefasit, base of Mt Bizen, Ryding et al. 2095 (neotype: C; isoneotypes: ASMU, ETH, UPS, designated by Ryding (2001)).
Plectranthus ugandensis Lye, Norweg. J. Bot. 20: 57. 1973, nom. illeg., non P. ugandensis S.Moore.
Distribution.
Yemen, Eritrea to Tanzania.
Coleus psammophilus
(Codd) A.J.Paton comb. nov.
6860B521635E59898D92E46AD0FBF19A
urn:lsid:ipni.org:names:77201173-1
Plectranthus psammophilus Codd, Bothalia 11: 405. 1975. Type: South Africa, KwaZulu-Natal, Makatini Flats, Jan.1964, Strey 5779 (holotype: PRE; isotypes: BR, K, M, NH).
Distribution.
Mozambique to N. KwaZulu-Natal.
Coleus pseudospeciosus
(Buscal. & Muschl.) A.J.Paton comb. nov.
0042D4643D8F57B99382B0387ACE977D
urn:lsid:ipni.org:names:77201174-1
Pycnostachys pseudospeciosa Buscal. & Muschl., Bot. Jahrb. Syst. 49: 486. 1913. Type: Zambia, Lake Bangweulu, n.d, D’Aosta 1002 (holotype: B, destroyed; lectotype: K, designated by Ryding (2001)).
Pycnostachys ballotoides Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 72. 1921. Type: Congo, Katanga, Mt Senga, Kassner 2930a (holotype: B, destroyed).
Pycnostachys mausaensis Gürke, Notes Fl. Katanga 7: 47. 1921. Type: Congo, Katanga, Mt Senga, 31.May 1908, Kassner 2920a (holotype: BR; isotype: K).
Distribution.
S. DRC to N. Zambia.
Coleus pulchellus
(P.I.Forst.) P.I.Forst. comb. nov.
632E1D9275CC5B9596B11668367122DB
urn:lsid:ipni.org:names:77201175-1
Plectranthus pulchellus P.I.Forst., Austrobaileya 4: 182. 1994. Type: Australia, Queensland, Cook District, Wasp Gully, Glennie Tableland, 5 Jul. 1991, M.C.Tucker (holotype: BRI (AQ509442); isotype: CNS).
Distribution.
Australia: Queensland.
Coleus recurvata
(Ryding) A.J.Paton comb. nov.
404D2ED423055256B0AC15A252078488
urn:lsid:ipni.org:names:77201176-1
Pycnostachys recurvata Ryding, Novon 9: 101. 1999. Type: Ethiopia, 60km W of Ambo along Lekemti (Nekemte) road, W.J.de Wilde & de Wilde-Duyfjes 8754 (holotype: C; isotypes: BR, ETH, K, UPS, WAG).
Distribution.
Ethiopia.
Coleus repens
Gürke, Bot. Jahrb. Syst. 19: 213. 1894
EEF94661CEA75915B03B9DFBEA736214
Solenostemon repens (Gürke) J.K.Morton, J. Linn. Soc., Bot. 58: 272. 1962.
Plectranthus epilithicus B.J.Pollard, Kew Bull. 60: 145. 2005. Type: Kamerun, im Urwald westlich von Buea, Preuss 949 (holotype: B, destroyed); (neotype: Cameroon, Mt Kupe, Nyasoso, nature trail above the Government High School, Pollard 83 (neotype: K; isoneotypes KUPE, WAG YA, designated by Pollard (2005)).
Coleus carnosus A.Chev., J. Bot. (Morot) 22: 125. 1909., nom. illeg.
Coleus carnosus var. lamiifolius A.Chev., Explor. Bot. Afrique Occ. Franç. 1: 518. 1920. nom. illeg.
Distribution.
W. & WC. Trop. Africa.
Coleus reticulatus
A.Chev., Mém. Soc. Bot. France 8: 200. 1912
B0BA64E970E25BAB90819DBD93CB90C3
Isodictyophorus reticulatus (A.Chev.) J.K.Morton, J. Linn. Soc., Bot. 58: 272. 1962.
Plectranthus reticulatus (A.Chev.) B.J.Pollard, Kew Bull. 67: 49. 2012. Type: Guinée Francaise, Cercle de Faranah (Faranna), dans la brousse très marécageuse et au bord des petits marigots, entre Kaba et Tonfili, 11 Jan.1909, Chevalier 20393 (holotype: P; isotype: K).
Isodictyophorus chevalieri Briq., Mém. Soc. Bot. France 8: 285. 1917. Type: Mali, Sedwhi toTimbuktu, 11–12 Feb. 1900, Chevalier s.n. (holotype: P).
Coleus casamancicus A.Chev. ex Hutch. & Dalziel, Fl. W. Trop. Afr. 2: 291. 1931. Type: Senegal, Casamance from Farancounda to Tambanaba, Sedhiou, 12 Feb. 1900, Chevalier 2777b (holotype: P).
Distribution.
W. Trop. Africa.
Coleus rhodesianum
(N.E.Br.) A.J.Paton comb. nov.
DD75E338E5755E8E9F519E7E158C1D8F
urn:lsid:ipni.org:names:77201177-1
Englerastrum rhodesianum N.E.Br., Bull. Misc. Inform. Kew 1922: 31. 1922. Types: Zambia, Mumbwa, Macauly 637 (syntype: K); Livingstone, Rogers 7205 (syntype: not seen).
Englerastrum schweinfurthii Briq., Bot. Jahrb. Syst. 19: 178. 1894, non Coleus schweinfurthii Vatke.
Coleus englerastrum Roberty, Bull. Inst. Fondam. Afrique Noire, Sér. A, Sci. Nat. 16: 330. 1954. Type: Sudan, headwaters of Ghasal, Addai, Schweinfurth 2532 (isotypes: K, WU).
Englerastrum djalonense A.Chev., J. Bot. (Morot) 22: 126. 1909, non Coleus djalonensis A.Chev.
Plectranthus djalonensis (A.Chev.) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 286. 2009. Type: Guinea, between Timbo and Kouria, Sept. 1907, Chevalier s.n. (holotype: P; isotype: K).
Plectranthus piliferus Chiov., Boll. Soc. Bot. Ital. 1924: 44. 1924. Type: Angola, Mossamedes, Huambo, Mazzocchi-Alemanni (holotype: FT).
Englerastrum hutchinsonianum Alston, Bull. Misc. Inform. Kew 1926: 298. 1926. Type: Nigeria, Lokoja, Mt Patti, Dalziel 105 (holotype: K).
Englerastrum schlechteri Alston, Bull. Misc. Inform. Kew 1926: 297. 1926., nom. illeg., non. T.C.E.Fr.
Distribution.
Trop. Africa to Caprivi Strip.
Coleus robustus
(Hook.f.) A.J.Paton comb. nov.
C438E6E172C75142B98288AF2C4A4059
urn:lsid:ipni.org:names:77201178-1
Anisochilus robustus Hook.f., Fl. Brit. India 4: 629. 1885. Type: India, Deccan Peninsula, Courtallum, Herb. Wight 625 (holotype: K (K000674766); isotype: E).
Anisochilus henryi K.Ravik. & V.Lakshm., Rheedea 9: 72. 1999. Type: India, Tamil Nadu, Theni Distr., near Varaiyaatumottai peak, Venniar Estate, Pachakumatchi hills, 1800 m, 28 May 1989, Lakshmanan & Ravikumar 91078 (holotype: CAL; isotypes: K, MH).
Distribution.
S. India.
Coleus rotundifolius
(Poir.) A.Chev. & Perrot, Veg. Ut. Afr. Trop. Franç. 1: 101. 1905
C1DDEE0493485C4EA4BE13BF7D7F7480
Germanea rotundifolia Poir. in J.B.A.M.de Lamarck, Encycl. 2: 763. 1788.
Plectranthus rotundifolius (Poir.) Spreng., Syst. Veg. 2: 690. 1825.
Solenostemon rotundifolius (Poir.) J.K.Morton, J. Linn. Soc., Bot. 58: 272. 1962. Type: Mauritius (Ile de France), Commerson s.n. (holotype: P; isotype: FI).
Nepeta madagascariensis Lam., Encycl. 1: 712. 1785, non Coleus madagascariensis (Benth.) A.Chev. Type: illustration in Rheede, Hortus Mal. 11: 49 t.25. 1692.(lectotype: illustration designated here).
Plectranthus ternatus Sims, Bot. Mag. 51: t. 2460. 1824.
Coleus ternatus (Sims) A.Chev., Veg. Ut. Afr. Trop. Franç. 1: 109. 1905. Type: specimen of cultivated plant illustrated in Bot. Mag. t. 2460 (lectotype: illustration designated here).
Plectranthus tuberosus Blume, Bijdr. Fl. Ned. Ind.: 838. 1826.
Coleus tuberosus (Blume) Benth., Labiat. Gen. Spec.: 59. 1832.
Majana tuberosa (Blume) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Indonesia, Java, Blume s.n. (holotype: L: isotypes: NY, P).
Coleus rugosus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 15. 1830. Type: India, Madras, Palamcotta, Herb. Madras, with ‘Dracocephalum rugosum’ mss., Wall. Cat. 2760 (holotype: K (no barcode.); isotype: K-W(K001117037).
Coleus parviflorus Benth. in A.P.de Candolle, Prodr. 12: 72. 1848.
Calchas parviflorus (Benth.) P.V.Heath, Calyx 6: 51. 1999. Type: South India, Quilon and Panpanassum, Wight 2512 (holotype: K(K000674660); isotype: E).
Coleus dysentericus Baker, Bull. Misc. Inform. Kew 1894: 10. 1894. Type: Niger, Zomba region, Barter 846 (holotype: K).
Coleus salagensis Gürke, Bot. Jahrb. Syst. 19: 220. 1894. Type: Benin, between Paratau and station Bismarkburg, Kling 199, Buttner 94, 690 (syntypes: not seen).
Plectranthus coppinii Heckel, Rev. Cultures Colon. 8: 164. 1900. Type and protologue not seen.
Coleus rehmannii Briq., Bull. Herb. Boissier, sér. 2, 3: 1075. 1903. Type: South Africa, Transvaal, Houtbosch, Rehmann 6156 (holotype: Z).
Coleus pallidiflorus A.Chev., J. Bot. (Morot) 22: 124. 1909. Type: Guinea, Conakry, Fouta Djalon, Dalaba, Nov. 1907, Chevalier s.n. (holotype: P). Locality of Chevalier 18234 in P and annotated as type, does not match protologue locality.
Coleus rotundifolius var. nigra A.Chev., Explor. Bot. Afrique Occ. Franç. 1: 520. 1920. Type: Ivory Coast, Haute sassandra, Pays Dyolas, village de Man, 13 May 1909, Chevalier 21532 (holotype: P).
Distribution.
Tropical Asia, cultivated elsewhere.
Coleus ruandensis
(De Wild.) A.J.Paton comb. nov.
42FCB74BBB115E3EB75DE01A807419AD
urn:lsid:ipni.org:names:77201179-1
Pycnostachys ruandensis De Wild., Pl. Bequaert. 4: 401. 1928. Type: Rwanda, between Kirinda and Lubengera, Robyns 2449 (holotype: BR).
Pycnostachys clinodon Mildbr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 405. 1932. Type: Tanzania, Njombe District: Lupembe, Schlieben 1053 (holotype: B, destroyed; isotypes: BM, BR, K, fragment).
Distribution.
Uganda, Burundi, Rwanda, DRC and Malawi.
Coleus rutenbergianus
(Vatke) A.J.Paton & Phillipson comb. nov.
7378B3F125235356A80C65B7C5BC3F2C
urn:lsid:ipni.org:names:77201180-1
Plectranthus rutenbergianus Vatke, Abh. Naturwiss. Vereins Bremen 9: 135. 1885.
Solenostemon rutenbergianus (Vatke) Guillaumet & A.Cornet, Adansonia, n.s., 15: 524. 1976. Type: Madagascar, Nahe des Itasi Sees, 19 Dec. 1877, Rütenberg s.n. (holotype: B, destroyed).
Distribution.
Madagascar.
Coleus sahyadricus
(Smitha & Sunojk.) Smitha comb. nov.
76495AE96C2D5B39BC66558171CF2DBB
urn:lsid:ipni.org:names:77201181-1
Plectranthus sahyadricus Smitha & Sunojk., Phytotaxa 345 (2): 166. 2018. Type: India. Kerala: Idukki district, Munnar, Lockhart Gap view point, 21 Sept. 2015, Smitha & Sunojkumar 135456 (holotype: CALI).
Distribution.
S. India.
Coleus sallyae
(A.J.Paton) A.J.Paton comb. nov.
E645952803BB5756B89D3A793B25D78F
urn:lsid:ipni.org:names:77201182-1
Plectranthus sallyae A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 275. 2009. Type: Tanzania, Ufipa District: Nkansi, 15 km on Kipili–Namanyere road, Bidgood, Mbago & Vollesen 2661 (holotype: K(K000431951); isotypes: K (K000431952), BR, C, DSM, EA, NHT, UPS).
Distribution.
SW. Tanzania.
Coleus sanguineus
(Britten) A.J.Paton comb. nov.
ECB32B811A395F9383FC7415F3C13A6B
urn:lsid:ipni.org:names:77201183-1
Plectranthus sanguineus Britten, Trans. Linn. Soc. London, Bot. 4: 36. 1894. Type: Malawi, Mulanje, Dec.1891, Whyte 46 (holotype: BM; isotypes: G, K).
Distribution.
S. Trop. Africa.
Coleus saxorum
(J.Mathew, Yohannan & B.J.Conn) Smitha comb. nov.
10B63389BA06539E8D96D911DA3445A2
urn:lsid:ipni.org:names:77201184-1
Plectranthus saxorum J.Mathew, Yohannan & B.J.Conn, Telopea 20: 187. 2017. Type: India, Kerala, Kozhikode District, Vellarimala, 10 km away from Muthappanpuzha, 30 Jan. 2015, J.Mathew 4911 (holotype: TBGT; isotype, MSSRF).
Distribution.
India, endemic to Western Ghats.
Coleus scaber
(Benth.) A.J.Paton comb. nov.
EE8FFEB91EFB5C6E9F45A767E06D2970
urn:lsid:ipni.org:names:77201185-1
Anisochilus scaber Benth. in A.P.de Candolle, Prodr. 12: 81. 1848. Type: India, Deccan Peninsula, Courtallum, Herb. Wight 2520 (holotype: K (K000674680); isotypes E, K (K000674681).
Distribution.
S. India.
Coleus scandens
Gürke, Abh. Königl. Akad. Wiss. Berlin 1894: 51. 1894
5C800E84FBF55277B4B36E2186AE72A9
Englerastrum scandens (Gürke) Alston, Bull. Misc. Inform. Kew 1926: 299. 1926.
Plectranthus scandens (Gürke) R.H.Willemse, Blumea 25: 510. 1979. Types: Tanzania, Lushoto District: Msinga to Kwa Mshusa, Holst 9119 (syntype: B, destroyed; isosyntype: K) & 9120a (syntype: B, destroyed; isosyntypes: G, JE, K, W).
Distribution.
Kenya to N. Tanzania.
Coleus scebeli
Chiov., Fl. Somala 1: 2. 1929
A5A2C1D2498B5431B4160756513B7188
Plectranthus scebeli (Chiov.) Ryding, Kew Bull. 54: 125. 1999. Type: Somalia, Shabeellaha Dhexe Rguion, between Ganale and Audegle, Puccioni & Stafanini 48 (holotype: FT).
Distribution.
S. Somalia.
Coleus schizophyllus
(Baker) A.J.Paton comb. nov.
96CB99B4F30A5B7086A96E4936FBFF88
urn:lsid:ipni.org:names:77201186-1
Plectranthus schizophyllus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 414. 1900.
Calchas schizophyllus (Baker) P.V.Heath, Calyx 6: 51. 1999. Type: Malawi, between Khondowe and Karonga, Whyte 376 (holotype: K).
Distribution.
SW. Tanzania to N. Malawi.
Coleus schliebenii
(Mildr.) A.J.Paton comb. nov.
C3D39DAEB79B595F88AC6348BCD48278
urn:lsid:ipni.org:names:77201187-1
Pycnostachys schliebenii Mildbr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 405. 1932. Type: Tanzania, Njombe District: Lupembe, Schlieben 713 (holotype: B, destroyed: isotypes: BR, Z; K, fragment).
Distribution.
Tanzania to Zambia.
Coleus scruposus
A.J.Paton nom. nov.
B1F52070805B56418AC1CB9700751EDF
urn:lsid:ipni.org:names:77201188-1
Pycnostachys kassneri De Wild., Contr. Fl. Katanga: 172 1921., non Coleus kassneri Robyns & Lebrun. Type: DRC, Katanga, West Kundelungu, Kassner 2794 (holotype: BR; isotypes: BM, K).
Pycnostachys carigensis Gürke, Ann. Mus. Congo Belge, Bot., sér. 4, 2: 135. 1913., nom. nud.
Distribution.
Tanzania to Zambia.
Coleus scutellarioides
(L.) Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830
14B2B4EE11B954E0B716C471EFCD2253
Ocimum scutellarioides L., Sp. Pl. ed. 2: 833. 1763.
Plectranthus scutellarioides (L.) R.Br., Prodr. Fl. Nov. Holl.: 506. 1810.
Plectranthus aromaticus Roxb., Hort. Bengal.: 45. 1814.
Majana scutellariodes (L.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Solenostemon scutellarioides (L.) Codd, Bothalia 11: 439. 1975.
Calchas scutellarioides (L.) P.V.Heath, Calyx 6: 51. 1999. Type: Majana (alba et rubra) Rumphius, Herb. Amboin. 5: 291, t. 101. 1747, (lectotype: illustration designated by Merrill (Interpr. Rumph. Herb. Amboin.: 460 1917).
Polypodium ovatum Burm. f., Fl. Ind.: 233. 1768. Type: Java, Unknown collector (holotype: G).
Germanea nudiflora Poir. in J.B.A.M.de Lamarck, Encycl. 2: 763. 1788.
Plectranthus nudiflorus (Poir.) Willd., Sp. Pl. 3: 168. 1800. Type: Amboina, Willdenow Herbarium 11079.1 & 11079.2 (syntypes: B-W).
Ocimum peltatum Schweigg. ex Schrank, Denkschr. Königl.-Baier. Bot. Ges. Regensburg 2: 55. 1822. Type: Cultivated from seed collected by Martius in Brazil (type: not seen).
Plectranthus ingratus Blume, Bijdr. Fl. Ned. Ind.: 836. 1826.
Coleus ingratus (Blume) Benth., Labiat. Gen. Spec.: 53. 1832.
Coleus scutellarioides var. ingratus (Blume) Miq., Fl. Ned. Ind. 2: 950. 1858. Type: Indonesia, Java, G. Parang, Blume 992 (lectotype: L, designated by Keng (1969)).
Plectranthus laciniatus Blume, Bijdr. Fl. Ned. Ind.: 838. 1826.
Coleus laciniatus (Blume) Benth., Labiat. Gen. Spec.: 56. 1832.
Coleus scutellarioides var. laciniatus (Blume) Miq., Fl. Ned. Ind. 2: 950. 1858. Type: Indonesia, Java, Blume s.n. (lectotype: L, designated by Keng (1969)).
Plectranthus scutellarioides Blume, Bijdr. Fl. Ned. Ind.: 837. 1826., nom. illeg.
Coleus blumei Benth., Labiat. Gen. Spec.: 56. 1832.
Coleus scutellarioides var. blumei (Benth.) Miq., Fl. Ned. Ind. 2: 950. 1858.
Majana scutellarioides var. blumei (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Solenostemon blumei (Benth.) M.Gómez, Bol. Secr. Agric. Comerc. Trab., Cuba 22: 127. 1914.
Plectranthus blumei (Benth.) Launert, Mitt. Bot. Staatssamml. München 7: 301. 1968. Type: Indonesia, Java, Blume s.n. (lectotype: L, designated by Suddee et al. (2004)).
Coleus atropurpureus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830.
Majana scutellarioides var. atropurpureus (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Calchas atropurpureus (Benth.) P.V.Heath, Calyx 5: 160. 1997. Type: Singapore, Wall. Cat. 2733A (syntypes: K (no barcode), K-W (K001116976)).
Coleus atropurpureus var. ramosus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830. Type: Malaysia, Penang, Wall. Cat. 2733B (syntypes: G-DC(microfiche), K-W (K001116976).
Coleus acuminatus Benth., Linnaea 6: 81. 1831.
Majana acuminata (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Calchas acuminatus (Benth.) P.V.Heath, Calyx 5: 160. 1996. Type: Philippines, Island of Samar, 1841, Cuming 1683 (neotype: K (K000897809), neotype designated by Suddee et al. 2004; isoneotypes: BM, K (K000897810), P, TCD).
Coleus atropurpureus var. densiflorus Benth., Labiat. Gen. Spec.: 54. 1832. Type: Philippines, Manila, 1830, Chamisso s.n. (syntype: K (K000897808).
Coleus grandifolius Benth., Labiat. Gen. Spec.: 53. 1832.
Majana grandifolia (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Coleus scutellarioides var. grandifolius (Benth.) Keng, Gard. Bull. Singapore 24: 58. 1969. Type: Timor, Guichenot s.n. (syntype P; isosyntypes: K, L, G-DC).
Coleus multiflorus Benth., Labiat. Gen. Spec.: 55. 1832.
Majana multiflora (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Philippines, Manila, Perrottet s.n. (holotype: P).
Coleus secundiflorus Benth., Labiat. Gen. Spec.: 55. 1832.
Majana secundiflora (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Timor, Guichenot s.n. (holotype: P).
Coleus grandifolius Blanco, Fl. Filip.: 482. 1837., nom. illeg.
Coleus blancoi Benth. in A.P.de Candolle, Prodr. 12: 79. 1848.
Majana blancoi (Benth.) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Phillippines, Blanco, s.n. (type: not seen).
Coleus pumilus Blanco, Fl. Filip.: 482. 1837.
Majana pumila (Blanco) Kuntze, Revis. Gen. Pl. 2: 524. 1891. Type: Philippines, Barrio of Pineda, Pasig, Rizal Province, Luzon, Oct. 1914, Merrill: Species Blancoanae No. 190 (neotype: K, designated by Suddee et al. (2004)).
Coleus atropurpureus var. javanicus Benth. in A.P.de Candolle, Prodr. 12: 74. 1848. Type: Indonesia, Java, Zollinger 138 (syntypes: G-DC, K).
Coleus blumei var. pectinatus C.Morren, Belgique Hort. 6: 99. t 18. 1856. Type: Cultivated in Liege, illustration cited above (lectotype: illustration designated here).
Coleus scutellarioides var. celebicus Miq., Fl. Ned. Ind. 2: 950. 1858. Type: Indonesia, Celebes, Reinwardt s.n. (type: not seen).
Coleus scutellarioides var. gracilis Miq., Fl. Ned. Ind. 2: 950. 1858. Type: Indonesia, Ceram, Bangka, Miquel (type: not seen).
Perilla nankinensis Wender., Linnaea 29: 726. 1859, nom. illeg., non P. nankinensis (Lour.) Decne.
Coleus verschaffeltii Lem., Ill. Hort. 8: t. 293. 1861. Type: based on cultivated plants from Java,illustration cited above (lectotype: illustration, designated by Suddee et al. 2004).
Coleus marmoratus W.Bull, Proc. Roy. Hort. Soc. London 4: 133. 1864.
Coleus verschaffeltii var. marmoratus (W.Bull) André, Pl. Feuill. Ornem.: 144. 1866. Type: not seen.
Coleus gibsonii Verl., Veitch & Sons in Gard. Chron. 1866: 432 (12 May 1866). Rev. Hort. (Paris) 37: 279. 1866 (July 1866). Type: Cultivated in London, sent by John Veitch from New Caledonia. (type: not seen).
Coleus veitchii Dombrain, Fl. Mag. (London) 6: t. 345. 1867. Type: Cultivated in London, sent by John Veitch from New Caledonia, illustration cited above (lectotype: illustration, designated here).
Coleus batemannii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.
Coleus bausei T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.
Coleus berkeleyi T.Moore, Gard. Chron. 1868: 376. 1868. Type: not seen.
Coleus clarkii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.
Coleus dixii T.Moore, Gard. Chron. 1868: 376. 1868. Type: not seen.
Coleus marshallii T.Moore, Gard. Chron. 1868: 376. 1868.
Coleus blumei var. marshallii (T.Moore) Rothsch., Pl. Feuill. Col., 2: 74. t. 37. 1869. Type: Illustration cited above (lectotype: illustration, designated here).
Coleus murrayi T.Moore, Gard. Chron. 1868: 377. 1868.
Coleus blumei var. murrayi (T.Moore) Rothsch., Pl. Feuill. Col., 2: 74 t. 37. 1869. Type: Illustration cited above (lectotype: illustration, designated here).
Coleus reveesii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.
Coleus ruckeri T.Moore, Gard. Chron. 1868: 376. 1868. Type: not seen.
Coleus saundersii T.Moore, Gard. Chron. 1868: 376. 1868. Type: not seen.
Coleus scottii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.
Coleus wilsonii T.Moore, Gard. Chron. 1868: 377. 1868. Type: not seen.
Coleus telfordii McPhail ex H.Laurentius, Nursery Cat. (H.Laurentius) 40: 3. 1868.
Coleus blumei var. telfordii (McPhail ex H.Laurentius) Rothsch., Pl. Feuill. Col., 2: 74 t. 37. 1869. Type: Illustration cited above (lectotype: illustration, designated here).
Coleus scutellarioides var. angustifolia Benth., Fl. Austral. 5: 80. 1870.
Calchas scutellarioides var. angustifolia (Benth.) P.V.Heath, Calyx 6: 51. 1999. Type: Australia, Table Hill, Victoria River & Macadam Range, 1855, F. Mueller s.n. (syntype: K); Port Essington, Armstrong 244 (syntype: K).
Coleus scutellarioides var. laxa Benth., Fl. Austral. 5: 80. 1870. Types: Australia, Roe River, York Sound, NW coast, A.Cunningham (syntype: K); Roebuck Bay NW coast, Martin s.n. (syntype: not seen); Arnhems land, Mackinley, R.Brown (syntype: MEL).
Coleus scutellarioides var. limnophila Benth., Fl. Austral. 5: 80. 1870.
Calchas scutellarioides var. limnophila (Benth.) P.V.Heath, Calyx 6: 51. 1999. Type: Australia, Nicholson & MacArthur Rivers, F. Mueller (syntypes: K).
Coleus tryonii auct., Fl. Mag. (London), n.s., 1: t. 34. 1872. Type: cultivated in UK, Leicester, garden of Captain Tryon (lectotype: illustration designated here).
Coleus verschaffeltii var. splendens auct., Garden (London 1871–1927) 5: 54. 1874. Type: not seen.
Coleus multicolor J.H.Veitch ex Kellock, Gard. Chron., n.s., 7: 571. 1877. Type: not seen.
Coleus hendersonii Regel, Gartenflora 28: 244. 1879. Type: not seen.
Coleus × eureka auct., Gard. Chron., ser. 3, 5: 808. 1889. nom. nud.
Coleus gaudichaudii Briq., Annuaire Conserv. Jard. Bot. Genève 2: 237. 1898. Type: Philippines, fle de Luzon pres de Manille, Dec. 1836, Gaudichaud s.n. (holotype: G-DEL).
Coleus igolotorum Briq., Annuaire Conserv. Jard. Bot. Genève 2: 236. 1898. Type: Philippines, fle de Luzon, dans le pays des Igolotes, Callery 48 (holotype: G-DEL; isotype: P).
Coleus savannicola K.Schum. in K.M.Schumann & C.A.G.Lauterbach, Fl. Schutzgeb. Südsee: 529. 1900. Type: not seen.
Coleus formosanus Hayata, J. Coll. Sci. Imp. Univ. Tokyo 22: 320. 1906. Type: Kotosho, 21 Nov. 1899, Miyake s.n. (holotype: TI).
Coleus macranthus var. crispipilus Merr., Philipp. J. Sci. 1(suppl.): 235. 1906.
Coleus crispipilus (Merr.) Merr., Philipp. J. Sci., C 5: 382. 1910.
Calchas crispipilus (Merr.) P.V.Heath, Calyx 5: 160. 1996.
Coleus scutellarioides var. crispipilus (Merr.) H.Keng, Gard. Bull. Singapore 24: 56. 1969.
Calchas scutellarioides var. crispipilus (Merr.) P.V.Heath, Calyx 6: 51. 1999. Type: Philippines, Suyoc to Panai, Province of Benguet, Luzon, 2200 m, Nov. 1905, Merrill 4780 (holotype: PNH; isotypes: BM, K, NY).
Coleus pubescens Merr., Philipp. J. Sci., C 3: 432. 1909. Type: Philippines, Babuyan Island, 17 June 1907, Eugenio Zenix, Bureau of Science 3892 (holotype: PNH; isotype: BM).
Coleus zschokkei Merr., Philipp. J. Sci., C 5: 382. 1910. Type: Philippines, Mt Pulog, Prov. of Bengnet, Luzon, May 1909, Merrill 6529 (holotype: PNH; isotypes: K, NY, P).
Coleus integrifolius Elmer, Leafl. Philipp. Bot. 7: 2696. 1915.
Coleus scutellarioides var. integrifolius (Elmer) Keng, Gard. Bull. Singapore 24: 58. 1969. Type: Philippines, Mindanao, Cabadbaran (Mt Urdaneta), Prov. of Agusan, Island of Mindanao, August 1912, Elmer 13627 (holotype: PNH; isotypes: A, BM, K, L, NY, P).
Coleus gibbsiae S.Moore in L.S.Gibbs, Fl. Arfak Mts.: 178. 1917.
Coleus scutellarioides var. gibbsiae (S.Moore) Keng, Gard. Bull. Singapore 24: 58. 1969. Type: Indonesia, New Guinea, Arfak Mts, 2100 m, Gibbs 5909 (holotype: BM; isotypes: K, L).
Coleus rehneltianus A.Berger, Bot. Jahrb. Syst. 54 (5, Beibl. 120): 197. 1917. Type: not seen.
Coleus hybridus Cobeau, Arch. Bot. (Forlì) 4: 35. 1928. Type: not seen.
Distribution.
Trop. & Subtrop. Asia to N. Australia. Widely cultivated as an ornamental.
Coleus seretii
De Wild., Bol. Soc. Ibér. Ci. Nat. 19: 122. 1920
4D40E3C7F0795D6FADD56FB1E0F10A4F
Calchas seretii (De Wild.) P.V.Heath, Calyx 6: 51. 1999.
Plectranthus seretii (De Wild.) Vollesen, Opera Bot. 59: 84. 1980. Type: DRC, Buta to Bima (Bali), Seret 68 (syntype: BR) & Suronga Forest, Seret 420 (wrongly cited as 120 in protologue) (syntype: BR; isosyntype: K).
Distribution.
Ethiopia, DRC, Tanzania, NW. Madagascar.
Coleus sessilifolius
(A.J.Paton) A.J.Paton comb. nov.
6BC35D9710C6580EB7A5A98CE86D6A59
urn:lsid:ipni.org:names:77201190-1
Plectranthus sessilifolius A.J. Paton, Fl. Zambes. 8(8): 281. 2013. Type: Zimbabwe, Mutare Dist., Himalaya Mts, 5 Mar.1954, Wild 4505 (holotype: K; isotypes: LISC, MO, SRGH).
Distribution.
Zimbabwe, Mozambique.
Coleus shirensis
Gürke, Bot. Jahrb. Syst. 19: 216. 1894
90C0FCF670C058DCB40ADA22EAD39426
Solenostemon shirensis (Gürke) Codd, Bothalia 11: 440. 1975.
Calchas shirensis (Gürke) P.V.Heath, Calyx 6: 51. 1999.
Plectranthus shirensis (Gürke) A.J.Paton, SABONET Rep. Ser. 31: 189. 2005. Type: Malawi, 1891, Buchanan 376 (lectotype: K; isolectotype: E, designated by Mathew (1976)).
Coleus punctatus Baker, Bull. Misc. Inform. Kew 1895: 291. 1895. Type: Zambia, Mweru Plateau, W of Lake Tanganyika, 1894, Carson 25 (holotype: K).
Solenostemon zambesiacus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 421. 1900. Types: Malawi, Blantyre, 1887, Last s.n. (syntype: K); ?Mozambique, “Zambesia”, between Shibisa and Tshinmuzo, Nov.1859, Kirk s.n. (syntype: K).
Distribution.
SW. & S. Tanzania to S. Trop. Africa.
Coleus shoolamudianus
(Sunil & Naveen Kum.) Smitha & A.J. Paton comb. nov.
0A55301B80C05BCAB6DADF8807C5D15A
urn:lsid:ipni.org:names:77201191-1
Anisochilus shoolamudianus Sunil & Naveen Kum. Webbia 70(2): 217. 2015. Type: India, Ernakulam District, Edamalayar Forest Range, Shoolamudi, Sunil & Naveen Kumar 6918 (holotype: MH; isotype CALI).
Coleus sigmoideus
(A.J.Paton) A.J.Paton comb. nov.
377EB9F4B94D51668924FBD7F2BBB624
urn:lsid:ipni.org:names:77201192-1
Plectranthus sigmoideus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 333. 2009. Type: Zambia, track opposite turning to Mbala (Abercorn) Club, Richards 4353 (holotype: K).
Distribution.
SW. Tanzania to Zambia.
Coleus socotranus
(A.R.Sm.) A.J.Paton comb. nov.
5BA72BB59DCC5EF09274DF75DF63504D
urn:lsid:ipni.org:names:77201193-1
Plectranthus socotranus A.R.Sm., Hooker’s Icon. Pl. 37: t. 3692. 1971. Type: Socotra, Adhoh de Melhoh, Hagghiher Mts, 22 Apr. 1967, Smith & Lavranos 499 (holotype: K; isotype FI).
Distribution.
Socotra.
Coleus sparsiflorus
Elmer, Leafl. Philipp. Bot. 7: 2699 1915
B057F35FE03C5712A90E177938C3C5F6
Plectranthus sparsiflorus (Elmer) H.Keng, Fl. Males. 8: 391. 1978. Type: Philippines, Mindanao, Mt. Apo, Sept. 1912, A.D.E.Elmer 11646 (isotypes: G, L).
Distribution.
Philippines.
Coleus magnificus
P.I.Forst. & A.J.Paton nom. nov.
FFADA3A679B6554C963C125FF5CAF5CC
urn:lsid:ipni.org:names:77201303-1
Plectranthus spectabilis S.T.Blake, Contr. Queensland Herb. 9: 50. 1971, non Coleus spectabilis Miq. Type: Australia, Queensland, Macalister Range near Hartley Creek C NW of Cairns (Cultivated at The Gap), Apr. 1959, S.T.Blake 20535 (holotype: BRI).
Distribution.
Australia: Queensland.
Coleus sphaerocephalus
(Baker) A.J.Paton comb. nov.
B72ACBC3885C5E04900E5655624EB3A4
urn:lsid:ipni.org:names:77201194-1
Pycnostachys sphaerocephala Baker, Bull. Misc. Inform. Kew 1898: 162. 1898. Type: Malawi, Nyika Plateau, 1800–2100 m, Whyte 139 (holotype: K).
Pycnostachys whytei Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 383. 1900. Type: Malawi, Chitipa (Fort Hill), Whyte s.n. (holotype: K).
Pycnostachys perkinsii E.A.Bruce, Bull. Misc. Inform. Kew 1939: 587. 1940.
Pycnostachys kaessneri Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 73. 1921., nom. illeg. Type: DRC, Katanga, Muhila (Mugila) Mts, western slope, Kassner 2991a (holotype: B, destroyed; isotypes: BR, K).
Distribution.
Tanzania to Zambia.
Coleus splendens
(P.I.Forst.) P.I.Forst. comb. nov.
0B0DD26941745327BBA08DA5B6747517
urn:lsid:ipni.org:names:77201302-1
Plectranthus splendens P.I.Forst., Austrobaileya 9(2): 211. 2014. Type: Australia, Queensland, Cook District, Hann Tableland National Park, 8 April 2013, P.I.Forster PIF39587 (holotype: BRI; isotypes: CNS, K, MEL).
Distribution.
Australia: Queensland.
Coleus splendidus
A.Chev., J. Bot. (Morot) 22: 122. 1909
1186C275898D54939D4400C1D7AD04D4
Coleus splendidus A.Chev., J. Bot. (Morot) 22: 122. 1909. Type: Guinea, Timbo, 16 September 1907, A. Chevalier s.n. (holotype: P).
Coleus djalonensis A.Chev., J. Bot. (Morot) 22: 123. 1909. Type: Guinea, between Kouria and Bouroukountou, 12 Sept. 1907, A. Chevalier s.n. (holotype: P).
Coleus peulhorum var. violacea A.Chev., J. Bot. (Morot) 22: 124. 1909. Type: Guinea: Fouta Djallon, entre Diaguissa et Boulwel, Chevalier 18606bis (holotype: P; isotype: P).
Distribution.
Guinea.
Coleus stachyoides
(Oliv.) E.A.Bruce, Bull. Misc. Inform. Kew 1934: 306. 1934
EB866CACEF0E50598A37449E22D0E3D3
Plectranthus stachyoides Oliv., Trans. Linn. Soc. London 29: 136. 1875. Type: Uganda, West Nile District: Madi, Grant 732 (holotype: K).
Plectranthus cylindrostachys Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 356. 1928. Type: Burundi, Irubura, Akanguru valley, Robyns 2403 (holotype: BR; isotype: K).
Distribution.
Central African Rep. to E. Trop. Africa.
Coleus steenisii
(H.Keng) A.J.Paton comb. nov.
A71407183F835BEAACB03DC78D69B68C
urn:lsid:ipni.org:names:77201304-1
Plectranthus steenisii H.Keng, Gard. Bull. Singapore 24: 145. 1969. Type: Java, res. Pasoeroean, G. Ardjoeno, rd bet. Sawahan Bakal and Sawaham Ardjoeno, 5 June 1935, van Steenis 7111 (holotype: L).
Distribution.
E. Java.
Coleus stenostachys
(Baker) A.J.Paton & Phillipson comb. nov.
B511EF53132E5233A12766A7A75DF1DF
urn:lsid:ipni.org:names:77201305-1
Pycnostachys stenostachys Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 380. 1900. Type: Uganda, Bunyoro District: sides of Nile, 1862, Grant in Speke & Grant s.n. (holotype: K).
Pycnostachys coerulea Hook., Exot. Fl. 3: t. 202. 1826, non Coleus coeruleus Gürke. Type: cultivated from seeds sent from Madagascar, collected by Bojer & Helsinger (holotype: K; isotype: M).
Pycnostachys micrantha Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 345. 1895, non Coleus micranthus Maxim. Types: Tanzania, Bukoba District: Karagwe, Stuhlmann 1720 & Mpororo, Stuhlmann 2056 & Bukoba, Stuhlmann 3276, 3718 & 3953 (syntypes: B, destroyed).
Pycnostachys brevipetiolata De Wild., Pl. Bequaert. 4: 394. 1928. Type: DRC, Kivu, Rutshuru R., Kaisafu, Bequaert 5972 (holotype: BR).
Distribution.
Ethiopia to South Africa, Madagascar.
Coleus strictipes
(G.Taylor) A.J.Paton comb. nov.
A263CCA35F23507A972BC69242401565
urn:lsid:ipni.org:names:77201306-1
Plectranthus strictipes (G.Taylor) A.J.Paton, Fl. Zambes. 8(8): 234. 2013.
Holostylon strictipes G.Taylor in J. Bot. 69, suppl.2: 161. 1931. Types: Angola, Cahaingalamush near Mulanje, 1903, Gossweiler 1038 (syntype: BM, K, P); Gola Luige, Malanje, Nov. 1922, Gossweiler 8848 (syntype: BM, K).
Distribution.
Angola.
Coleus strobilifer
(Roxb.) A.J.Paton comb. nov.
272D5F557CAA5B1E88D134E89AEFE1FD
urn:lsid:ipni.org:names:77201307-1
Plectranthus strobilifer Roxb., Hort. Bengal.: 45. 1814. Type: Katu-Kurka, Illustration in Rheede, Hort. Malab. 10: 179., t.90. 1690, (lectotype: illustration, designated by Suddee and Paton 2009).
Lavandula carnosa L.f., suppl. Pl.: 273. 1782 non Coleus carnosus Hassk.
Plectranthus carnosus (L.f.) Sm. in A.Rees, Cycl. 27: n.° 7. 1814.
Anisochilus carnosus (L.f.) ex Benth., Edwards’s Bot. Reg. 15: t. 1300. 1830. Type: India, Koenig s.n., Linnean Herbarium 727.7 (lectotype LINN).
Plectranthus dubius Spreng., Syst. Veg. 2: 691. 1825., nom. illeg.
Anisochilus carnosus var. purpurascens Benth. in N.Wallich, Pl. Asiat. Rar. 2: 18. 1830. Type: Burma, inm onte Taong Dong Ava, 24 Nov. 1826, Wall. Cat. 2753A (lectotype: K (K000674685); isolectotypes G-DC, K-W (K001117024); designated by Suddee & Paton (2009)).
Anisochilus glaber Schrad., Index Seminum (GOET) 1833: 1. 1833.
Anisochilus carnosus var. glaber (Schrad.) Benth., Labiat. Gen. Spec.: 711. 1835. Type: India, W Himalaya, to 2400 m, Edgworth 14 (neotype: K(K000674684), designated by Suddee et al. (2004)).
Anisochilus carnosus var. villosior Benth. in A.P.de Candolle, Prodr. 12: 81. 1848. Type: Deccan Peninsula. Herb. Wight 2516 (K (K000674688), lectotype designated Suddee and Paton 2009).
Anisochilus carnosus var. viridis Benth. in A.P.de Candolle, Prodr. 12: 81. 1848. Type: India, Deccan Peninsula, Herb. Wight 2521, (holotype: K (K000674686).
Anisochilus crassus Benth. in A.P.de Candolle, Prodr. 12: 81. 1848. Type: Deccan Peninsula, Herb. Wight. 2517 (lectotype: K (K000674682); isolectotype: E; designated Suddee and Paton 2009).
Anisochilus eriocephalus Benth. in A.P.de Candolle, Prodr. 12: 81. 1848.
Anisochilus carnosus var. eriocephalus (Benth.) T.Cooke, Fl. Bombay 2: 450. 1908. Type: Deccan Peninsula, Bellary, Herb. Wight 2518 (lectotype: K (K000674676); isolectotypes: E, K (K000674678, K000674677), designated Suddee & Paton (2009)).
Anisochilus decussatus Dalzell & A.Gibson, Bombay Fl.: 206. 1861. Type: India, Concan, on the Highest Ghauts opposite Bombay, Dalzell s.n. (holotype: K).
Anisochilus rupestris Wight ex Hook.f., Fl. Brit. India 4: 627. 1885. Type: India Deccan Peninsula, Herb. Wight 2521, (holotype: K (K000674686).
Distribution.
Indian Subcontinent to Indo-China.
Coleus stuhlmannii
(Gürke) A.J.Paton comb. nov.
A0B533D857285625B122B7AD4AF5A32F
urn:lsid:ipni.org:names:77201308-1
Pycnostachys stuhlmannii Gürke in H.G.A.Engler, Pflanzenw. Ost-Afrikas, C: 345. 1895. Type: Tanzania, Bukoba District: Karagwe, Ngaramo, Stuhlmann 1630 (holotype: B, destroyed; K, fragment).
Pycnostachys remotifolia Baker, Bull. Misc. Inform. Kew 1898: 161. 1898. Type: Malawi, Chitipa (Fort Hill), July 1896, Whyte s.n. (holotype: K).
Pycnostachys bequaertii De Wild., Contr. Fl. Katanga: 171. 1921. Type: DRC, Katanga, Shinsenda, Bequaert 425 (lectotype: BR; designated by Bramley in Paton et al. (2009)).
Pycnostachys longifolia De Wild., Contr. Fl. Katanga: 172. 1921. Type: DRC, Katanga, Welgelegen, Bequaert 562 (holotype: BR; isotype: K, fragment).
Distribution.
Kenya to S. Trop. Africa.
Coleus suaveolens
(S.T.Blake) P.I.Forst. & T.C.Wilson comb. nov.
76B5C28DF61F5377A90B6A51215CBD41
urn:lsid:ipni.org:names:77201309-1
Plectranthus suaveolens S.T.Blake, Contr. Queensland Herb. 9: 30. 1971. Type: Australia, Queensland, Cultivated at The Gap, Brisbane. Grown from transplants collected at Queen Mary’s Falls, by J.A. Gresty, Jan. 1959, S.T.Blake 20506 (holotype: BRI).
Distribution.
E. Australia.
Coleus subspicatus
(Hochst.) Walp., Repert. Bot. Syst. 6: 658. 1847
072BC123B9CE57228BA8CDB3E0991DDC
Plectranthus subspicatus Hochst., Flora 28: 67. 1845. Type: South Africa, Cape, Uitenhage, Krauss 1112 (syntypes: TUB (TUB008736, TUB008735).
Plectranthus spicatus E.Mey., Comm. Pl. Afr. Austr.: 230. 1838.
Burnatastrum spicatum (E.Mey.) Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 358. 1897, non Coleus spicatus Benth. Type: South Africa, Cape, Bathurst Division, Glenfilling, 1836, Drège 4731b (lectotype: K; isolectotypes: MO, P, S; designated by Codd (1975)).
Distribution.
S. Mozambique to South Africa.
Coleus succulentus
Pax, Bot. Jahrb. Syst. 39: 646. 1907
6F423DC3EB2850C8B1B8943A026BB8EF
Coleus succulentus Pax, Bot. Jahrb. Syst. 39: 646. 1907. Types. Ethiopia, Shoa, Schankora, Rosen s.n. (holotype: WRSL, destroyed); neotype: Ethiopia, Zuquala Mt, Ryding & Puff 1657 (neotype: UPS; isoneotype: ETH ; designated by Ryding (2000)).
Plectranthus pseudomarrubioides R.H.Willemse, Kew Bull. 40: 93. 1985. Type: Ethiopia, Debre Libanos, de Wilde 8656 (holotype: WAG).
Distribution.
Ethiopia to N. Tanzania, Arabian Peninsula.
Coleus suffruticosus
(Wight) A.J.Paton comb. nov.
5F88994DDCF85F759667346DD7C26DEC
urn:lsid:ipni.org:names:77201310-1
Anisochilus suffruticosus Wight, Icon. Pl. Ind. Orient. 4 (1): t. 1437. 1849. Type: India, Deccan Peninsula, Western Ghats, Nilghiri Hills, western slopes at Sisparah, Ex Herb. Wight Propr, Wight 1437 (holotype: K).
Distribution.
India.
Coleus sylvestris
(Gürke) A.J.Paton & Phillipson comb. nov.
1BFE2CDB139F5BF8B5F5F08402264ACD
urn:lsid:ipni.org:names:77201311-1
Plectranthus sylvestris Gürke, Bot. Jahrb. Syst. 19: 205. 1894. Type: Tanzania, Kilimanjaro, Rifinika Hill on Mawenzi, 14 Sept. 1893, Volkens 965 (holotype: B, destroyed; isotype: BM, G).
Coleus ruwenzoriensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 427. 1900. Types: Uganda, Ruwenzori, Scott Elliot 7556 & 7798 (syntypes: BM).
Plectranthus chiradzulensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 412. 1900. Types: Malawi, Mt Chiradzulu summit, Meller s.n. (syntype: K (K000430788); Mt Mulanje, Whyte s.n. (syntype: K (K000430787).
Coleus ferrugineus Robyns, Bull. Jard. Bot. État Bruxelles 17: 77. 1943.
Plectranthus ferrugineus (Robyns) Troupin & Ayob. in G.M.D.Troupin, Fl. Rwanda 3: 336. 1985., nom. inval. Type: DRC, Karisimbi, Lebrun 5005 (holotype: BR; isotype: K).
Coleus subulatus Robyns, Bull. Jard. Bot. État Bruxelles 17: 76. 1943. Type: DRC, Kivu, Tschamugussa, de Witte 1854 (holotype: BR).
Plectranthus bosseri Hedge, Fl. Madag. 175: 214. 1998. Type: Madagascar, Bosser 17953 (holotype: P).
Distribution.
Trop. Africa, Madagascar.
Coleus tenuicaulis
Hook.f., J. Proc. Linn. Soc., Bot. 7: 211. 1864
1285716E274F592195D345AD3A6E2B38
Majana tenuicaulis (Hook.f.) Kuntze, Revis. Gen. Pl. 2: 524. 1891.
Plectranthus tenuicaulis (Hook.f.) J.K.Morton, J. Linn. Soc., Bot. 58: 268. 1962. Type: Cameroon, Mt Cameroon, Dec.1862, Mann 1939 (holotype: K).
Coleus membranaceus Briq., Bot. Jahrb. Syst. 19: 182. 1894. Type: Angola, Cissacala (Sacalla?) in Cuango (Quango) region, Jan.1881, Mechow 554 (isotype: W).
Coleus montanus Gürke, Bot. Jahrb. Syst. 19: 218. 1894., nom. illeg., non Coleus montanus Hochst. ex Ces.
Plectranthus herbaceus Briq., Bot. Jahrb. Syst. 19: 179. 1894.
Germanea herbacea (Briq.) Hiern, Cat. Afr. Pl. 1: 861. 1900.
Coleus herbaceus (Briq.) G.Taylor, J. Bot. 69 (suppl. 2): 158. 1931. Type: Angola, Huíla, Lopolo, Mar. 1879, Welwitsch 5506 (syntypes: BM, G, K).
Plectranthus minimus Gürke, Bot. Jahrb. Syst. 19: 205. 1894. Type: Cameroon, Buea, Preuss 1019 (holotype: B, destroyed).
Coleus cunenensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 443. 1900. Type: Angola, near Rio Cunene, Sept. 1883, Johnston s.n. (holotype: K).
Germanea andongensis Hiern, Cat. Afr. Pl. 1: 862. 1900.
Plectranthus andongensis (Hiern) K.Schum., Just’s Bot. Jahresber. 28(1): 486. 1902.
Coleus andongensis (Hiern) G.Taylor, J. Bot. 69 (suppl. 2): 158. 1931. Type: Angola, Pungo Andongo, 1856–57, Welwitsch 5543 (holotype: LISU, isotype: BM).
Germanea concinna Hiern, Cat. Afr. Pl. 1: 861. 1900.
Plectranthus concinnus (Hiern) K.Schum., Just’s Bot. Jahresber. 28(1): 486. 1902. Type: Angola, Pungo Andongo, Mata de Pungo, Apr. 1857, Welwitsch 5533 (holotype: LISU; isotype: K).
Germanea concinna var. albiflora Hiern, Cat. Afr. Pl. 1: 862. 1900. Type: Angola, Pungo Andongo, between Luxilo and Cazela, May 1857, Welwitsch 5586 (holotype: LISU; isotypes: BM, G).
Germanea concinna var. caerulea Hiern, Cat. Afr. Pl. 1: 862. 1900. Type: Angola, Pungo Andongo, Nov. 1856, Welwitsch 5521 (holotype: LISU; isotype: BM).
Coleus peulhorum A.Chev., J. Bot. (Morot) 22: 123. 1909.
Plectranthus peulhorum (A.Chev.) J.K.Morton, J. Linn. Soc., Bot. 58: 268. 1962. Type: Guinea, Fouta Djalon, 13 Sept. 1907, Chevalier s.n. (holotype: P).
Coleus drymophilus G.Taylor, J. Bot. 69 (Suppl. 2): 160. 1931. Type: Angola, Cassuango, Cuiriri, n.d., Gossweiler 4056 (holotype: BM; isotype: K).
Distribution.
W. Trop. Africa to Cameroon, SW. Tanzania to S. Trop. Africa.
Coleus tetradenifolius
(A.J.Paton) A.J.Paton comb. nov.
10BE129CED205F47A330B12D8D441253
urn:lsid:ipni.org:names:77201312-1
Plectranthus tetradenifolius A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 304. 2009. Type: Uganda, Karamoja District: Mt Moroto, Tweedie 2665 (holotype: K).
Distribution.
Cameroon, S. Sudan to E. Trop. Africa.
Coleus tetragonus
(Gürke) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 106. 1929
58E659BF3E5C5624931C582A11334C4D
Plectranthus tetragonus Gürke, Bot. Jahrb. Syst. 19: 209. 1894.
Englerastrum tetragonum (Gürke) T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 73. 1924. Type: Tanzania, Lushoto District: Usambara Mts, Mashewa (Mascheua), Holst 3573 (holotype: B, destroyed; isotype: K, W, Z).
Plectranthus melanocarpus Gürke, Bot. Jahrb. Syst. 19: 210. 1894.
Englerastrum melanocarpum (Gürke) T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 71. 1924.
Coleus melanocarpus (Gürke) Robyns & Lebrun, Ann. Soc. Sci. Bruxelles, Sér. B 49: 106. 1929. Type: Tanzania, Masai steppe, Fischer 511 (holotype: B, destroyed; isotype: K, fragment).
Plectranthus biflorus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 402. 1900.
Englerastrum biflorum (Baker) T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 9: 73. 1924. Type: Malawi, between Khondowe and Karonga, Whyte s.n. (holotype: K).
Plectranthus dekindtianus De Wild., Ann. Mus. Congo Belge, Bot., sér. 4, 2: 135. 1913. Type: DRC, Bugege, Hock s.n. (holotype: BR).
Distribution.
Kenya to South Africa.
Coleus thalassoscopicus
(P.I.Forst.) P.I.Forst. comb. nov.
B5403512B4155FA7B9CFCBAD2BB41F8E
urn:lsid:ipni.org:names:77201313-1
Plectranthus thalassoscopicus P.I.Forst., Austrobaileya 4: 653. 1997. Type: Australia, Queensland, Cook District, North Bell Peak, Malbon Thompson Range, 12 Nov. 1995, P.I. Forster PIF18048, R.L. Jago & R. Jensen (holotype: BRI; isotypes: CNS, K, MEL).
Distribution.
Australia: Queensland (Cook).
Coleus thyrsoideus
Baker, Bot. Mag. 125: t. 7672 1899
AF3F0A1B10AF5D4685DEAD10D3D2CF22
Plectranthus thyrsoideus (Baker) B.Mathew, Kew Bull. 31: 174. 1976. Type: Plant cultivated at Kew 3 Jan.1899; seed from herbarium specimen from N of Lake Malawi collected by Whyte (holotype: K).
Distribution.
S. Trop. Africa.
Coleus togoensis
(Perkins) A.J.Paton comb. nov.
A3979C7F1E7E574584D74876EADE1CEC
urn:lsid:ipni.org:names:77201314-1
Pycnostachys togoensis Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 69. 1921. Type: Benin (Togo), between Kete and the Volta, Jan. 1895, Baumann 404 (B, holotype, destroyed; K, fragment).
Pycnostachys schweinfurthii Briq., Bot. Jahrb. Syst. 19: 191. 1894, non Coleus schweinfurthii Vatke. Type: Sudan, Equatoria, Dokuto (Dukuttu), Schweinfurth 2770 (holotype: B, destroyed; isotype: K).
Distribution.
W. Trop. Africa to Tanzania.
Coleus tomentifolius
(Suddee) Suddee comb. nov.
F0696565C5FF52D7A6EBFB30BE8C313F
urn:lsid:ipni.org:names:77201315-1
Plectranthus tomentifolius Suddee, Kew Bull. 59: 393. 2004. Type: Thailand, Takua Thung, Nov. 1896, Curtis s.n. (holotype: SING).
Distribution.
Peninsula Thailand.
Coleus torrenticola
(P.I.Forst.) P.I.Forst. comb. nov.
07BE50F80EFB586E80614B09C36D8935
urn:lsid:ipni.org:names:77201316-1
Plectranthus torrenticola P.I.Forst., Austrobaileya 3: 729. 1992. Type: Australia, Moreton District, Pooles Dam, Cooloolabin, State Forest Drive, 20 Feb. 1991, P.I. Forster PIF7797 & P.R. Sharpe (holotype: BRI; isotypes: CANB, K, MEL, NSW).
Distribution.
Australia: Queensland.
Coleus triangularis
(A.J.Paton) A.J.Paton comb. nov.
DDE3A327533A5614B2BC0B96A185B09C
urn:lsid:ipni.org:names:77201317-1
Plectranthus triangularis A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 313. 2009. Type: Kenya, Taita District: Ngangao Forest, Luke et al. 5497 (holotype: K; isotype: EA).
Distribution.
SE. Kenya to E. Tanzania.
Coleus trullatus
(A.J.Paton) A.J.Paton comb. nov.
9F9147F91CA35923A4BF21D918A374EE
urn:lsid:ipni.org:names:77201318-1
Plectranthus trullatus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 315. 2009. Type: Tanzania, Iringa District: Udzungwa Mts, Sanje path above loggers’ camp, Bridson 641 (holotype: K).
Distribution.
Tanzania.
Coleus umbrosus
Vatke, Linnaea 43: 91. 1881
EF77B007B5C55CF5B4D34E4E23068093
Pycnostachys umbrosa (Vatke) Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 8: 77. 1921. Type: Kenya, Teita District: Ndara, Hildebrandt 2424 (holotype: B, destroyed).
Coleus tricholobus Gürke, Abh. Königl. Akad. Wiss. Berlin 1894: 62. 1894. Type: Tanzania, Lushoto District: W Usambara Mts, Mlalo, Holst 455 (holotype: B, destroyed; K, fragment).
Distribution.
Kenya to N. Tanzania.
Coleus unguentarius
(Codd) A.J.Paton comb. nov.
B4DDD00694D75810A0DA47D3AA4BB4FE
urn:lsid:ipni.org:names:77201319-1
Plectranthus unguentarius Codd, Bothalia 11: 387. 1975. Type: Namibia, Kaokoveld, 17 km S. of Kaoko Otavi on road to Sesfontein, 21 April 1957, De Winter & Leistner 5595 (holotype: PRE).
Distribution.
Namibia.
Coleus urticifolius
Benth. in A.P.de Candolle, Prodr. 12: 78. 1848
051C320AA02A5C01B93FF5BCBDD1F64A
Plectranthus urticifolius (Benth.) Hook.f., Fl. Brit. India 4: 622. 1885., nom. illeg.
Plectranthus beddomei Raizada, Indian Forester 84: 503. 1958. Type: India, Courtallum, R.Wight 622 (holotype: K (K000820175); isotype: K(K000820174)).
Distribution.
S. India.
Coleus velutinus
(Trimen) A.J.Paton comb. nov.
BCABFB600AA75429B5162E5574086D89
urn:lsid:ipni.org:names:77201320-1
Anisochilus velutinus Trimen, Handb. Fl. Ceylon 3: 377. 1895. Type: Sri Lanka, Gunner’s Quoin, in the Batticaloa Distr, CP 3573 (holotype: PDA; isotypes: BM, K).
Anisochilus suffruticosus sensu Thwaites, Enum. Pl. Zeyl.: 238. 1860., non Anisochilus suffruticosus Wight; non Coleus suffrutescens (Wight) A.J.Paton.
Distribution.
Sri Lanka.
Coleus venteri
(van Jaarsv. & L.Hankey) A.J.Paton comb. nov.
6290E4BF872457A0BBC26DE2498A10FD
urn:lsid:ipni.org:names:77201321-1
Plectranthus venteri van Jaarsv. & L.Hankey, Aloe 34: 40. 1997. Type: South Africa, Northern Prov., Tzaneen, Geeneinde, Schoonoord, Sekekuneland, Venter 13626 (holotype: PRE).
Distribution.
S. Zimbabwe to Limpopo (Leolo Mts.).
Coleus ventosus
(P.I.Forst.) P.I.Forst. comb. nov.
E312E82D99FF575C985F32B6CB30D6E0
urn:lsid:ipni.org:names:77201322-1
Plectranthus ventosus P.I.Forst., Austrobaileya 9: 435. 2015. Type: Australia, Queensland, Cook District; ex situ cultivation from Melville Peak, Cape Melville National Park, 10 April 2015, H.B.Hines & C.J.Hoskin CM4 (holotype: BRI).
Distribution.
Australia: Queensland.
Coleus venustus
(P.I.Forst.) P.I.Forst. comb. nov.
B91E751F1D9F5B168195B09DC8355B96
urn:lsid:ipni.org:names:77201323-1
Plectranthus venustus P.I.Forst., Austrobaileya 8: 401. 2011. Type: Queensland, Cook District, Kennedy Hill Gorge, 21 June 1989, P.I.Forster PIF5388 & M.C.Tucker (holotype: BRI; isotypes: CNS, MEL).
Distribution.
Australia: Queensland.
Coleus verticillatus
(Baker) A.J.Paton comb. nov.
AC5129ADA2D65CE4884FDEACCE6C5C86
urn:lsid:ipni.org:names:77201324-1
Pycnostachys verticillata Baker, Bull. Misc. Inform. Kew 1895: 71. 1895. Type: Zambia, Northern Province, Fwambo, Carson 38 (holotype: K).
Distribution.
W. Tanzania to Zambia.
Coleus vettiveroides
K.C.Jacob, J. Bombay Nat. Hist. Soc. 42: 320. 1941
648DE2AB4C705DEB8104DA942019FB73
Plectranthus vettiveroides (K.C.Jacob) N.P.Singh & B.D.Sharma, J. Bombay Nat. Hist. Soc. 79: 712. 1982. publ. 1983. Type: India, Coimbatore, Madras Herbarium No. 85676. (holotype: MH).
Coleus osmirrhizon Elliot, Fl. Andhrica: 105. 1859. nom. inval.
Distribution.
S. India.
Coleus veyretiae
(Guillaumet & A.Cornet) A.J.Paton & Phillipson comb. nov.
E0C40A1E42F25782BE8B8CF4B54F4BA3
urn:lsid:ipni.org:names:77201325-1
Solenostemon veyretiae Guillaumet & A.Cornet, Adansonia, n.s., 15: 526. 1976.
Plectranthus veyretiae (Guillaumet & A.Cornet) Hedge, Fl. Madag. 175: 226. 1998. Type: Madagascar, Mahajanga, Sofia, Veyret 1154 (holotype: P; isotypes: K, TAN).
Distribution.
Madagascar.
Coleus villosus
(Forssk.) A.J.Paton comb. nov.
F83D5F8E70235CA99B9BF538890D59E9
urn:lsid:ipni.org:names:77201326-1
Ocimum villosum Forssk., Fl. Aegypt.-Arab.: cxv, 369. 1775.
Ocimum β zatarhendi Forssk. Fl. Aegypt.-Arab.: 110. 1775.
Plectranthus villosus (Forssk.) C, Chr.,, Dansk Bot. Arkiv. 4, 21. 1922, nom. illeg., non P. villosus Sieber ex Benth.
Plectranthus arabicus E.A.Bruce, Bull. Misc. Inform. Kew 1935: 324. 1935., non Coleus arabicus Benth. Type: Yemen, between Dorebat and Taaes (Taiz), Forsskal (holotype C, destroyed). Neotype: Saudi Arabia, Wolledje, Gebel Melhan Schweinfurth 690 (neotype: K; isoneotype: BM, P; designated by Wood (1983)).
Distribution.
SW. Arabian Peninsula.
Coleus wallamanensis
(T.C.Wilson & P.I.Forst.) T.C.Wilson & P.I.Forst. comb. nov.
9879B6AE966A5D06BF1C551B4CB454FC
urn:lsid:ipni.org:names:77201328-1
Plectranthus wallamanensis T.C.Wilson & P.I.Forst. Australian Systematic Botany, 31: 443. 2018. Type: Australia, Queensland:,North Kennedy District, Girringun National Park, rock fall at the base of Wallaman Falls, 9 May 2014, T.C.Wilson 521 & M.A.M.Renner (holotype: NSW 852570; isotypes: BRI, CANB).
Distribution.
Australia: Queensland.
Coleus welwitschii
Briq., Bot. Jahrb. Syst. 19: 185. 1895
F747977754985EB7A80BB4B5B8D9683F
Coleus welwitschii Briq., Bot. Jahrb. Syst. 19: 185. 1895. Type. Angola, Welwitsch 5589 (isosyntypes: BM, K) & Angola, Pungo Andongo, Mechow 75 (not seen).
Coleus marquesii Briq., Annuaire Conserv. Jard. Bot. Genève 2: 239. 1898. Type: Angola, Cuango and Cuillo, Marques 191 (holotype: G).
Coleus dupuisii Briq., Bull. Soc. Roy. Bot. Belgique 37: 70. 1899.
Plectranthus dupuisii (Briq.) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 329. 2009. Type: DRC, Kasai, Mayumbe, Dupuis s.n. (holotype: BR).
Coleus eetveldeanus Briq., Bull. Soc. Roy. Bot. Belgique 37: 73. 1899. Type: Congo, Lake Tanganyika, Mtowa, Deschamps s.n. (holotype: BR).
Coleus betonicoides Baker ex Hiern, Cat. Afr. Pl. 1: 866. 1900.
Calchas betonicoides (Baker ex Hiern) P.V.Heath, Calyx 5: 160. 1997. Type: Angola, Huila, Lopollo, Ferrão da Sola, Welwitsch 5612 (holotype: LISU; isotypes: BM, G, K).
Plectranthus microphyllus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 410. 1900., nom. illeg.
Plectranthus porpeodon Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 525. 1900.
Solenostemon porpeodon (Baker) J.K.Morton, Novon 8: 266. 1998. Type: Tanzania, Tanga District: Magila, Kirk s.n. (holotype: K).
Coleus newtonii Briq., Bull. Herb. Boissier, sér. 2, 6: 826. 1906. Type: Angola, Mossamedes, Huila, Newton 105 (holotype: Z).
Coleus delpierrei De Wild., Bol. Soc. Ibér. Ci. Nat. 19: 119. 1920. Types: DRC, Vankerkhovenville, Delpierre s.n. (syntype: BR) & Niangara, Delpierre s.n. (syntype: BR).
Coleus laurentii De Wild., Bol. Soc. Ibér. Ci. Nat. 19: 121. 1920. Type: DRC, Gombe, E. & M. Laurent s.n. (holotype: BR).
Coleus thyrsiflorus Lebrun & L.Touss. in J.-P.A.Lebrun, Expl. Parc Nat. Kagera 1: 120. 1948.
Solenostemon thyrsiflorus (Lebrun & L.Touss.) Vollesen, Opera Bot. 59: 85. 1980. Type: DRC, Kagera, Nyakayaga, Lebrun 9447 (holotype: BR; isotype: K).
Distribution.
Ethiopia to S. Trop. Africa.
Coleus xanthanthus
C.Y.Wu & Y.C.Huang, Acta Phytotax. Sin. 10: 241. 1965
1EEC6C5E1E955A4C8E1D850A00530C68
Coleus xanthanthus C.Y.Wu & Y.C.Huang, Acta Phytotax. Sin. 10: 241. 1965. Type: China, Yunnan, Mon-lun, 15 Sept. 1959, S.K.Pei (holotype: HK).
Distribution.
China: Yunnan.
Coleus xerophilus
(Codd) A.J.Paton comb. nov.
D68970B37A78550FB00E0D6E8E7ECB4B
urn:lsid:ipni.org:names:77201329-1
Plectranthus xerophilus Codd, Bothalia 11: 282. 1974. Type: South Africa, Transvaal, Lydenburg, district, near Marone, Codd & Dyer 7729 (holotype: PRE).
Distribution.
South Afica: Northern Prov.
Coleus xylopodus
(Lukhoba & A.J.Paton) A.J.Paton comb. nov.
B8AD155863D75B509AE9F1CE9EF761DD
urn:lsid:ipni.org:names:77201330-1
Plectranthus xylopodus Lukhoba & A.J.Paton, Kew Bull. 55: 960. 2000. Type: Kenya, Trans-Nzoia District: S Cherangani, Symes 293 (holotype: EA; isotype: K).
Ocimum cinereum R.Br. in H.Salt, Voy. Abyss.: 64. 1814., nom. nud.
Distribution.
S. Ethiopia to Uganda.
Coleus yemenensis
A.J.Paton nom. nov.
97F42180348553CF9D5F47C8E4D1B24B
urn:lsid:ipni.org:names:77201331-1
Plectranthus ovatus Benth., Labiat. Gen. Spec.: 709 1835., non Coleus ovatus Benth.
Ocimum gratissimum Forssk., Fl. Aegypt.-Arab.: 110. 1775., nom. illeg. Type: Yemen, July 1763, Forsskal 350 (holotype: C).
Distribution.
Arabian Peninsula.
Coleus zombensis
(Baker) Mwany. comb. nov.
6D246A427E1D574FAA093B8C87C5D451
urn:lsid:ipni.org:names:77201332-1
Plectranthus zombensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 402. 1900. Type: Malawi, summit of Mt Zomba, 1881, Buchanan 395 (holotype: K).
Plectranthus lastii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 403. 1900. Type: Malawi, near Blantyre, Jan. 1887, Last s.n. (holotype: K).
Distribution.
S. Malawi.
Genus Equilabium
Equilabium
Mwany., A.J. Paton & Culham, Bot. J. Linn. Soc. 188(4): 367. 2018
8C5FF8F2207654C2B8BFE28B2052CE1D
Figure 6.
Species of Equilabium 1. AEquilabium dolomiticum (Photo: Neil Crouch) BE. dissectum var. dissectum (Photo: Montfort Mwanyambo) CE. gracilum (Photo: Bart Wursten) DE. laxiflorum (Photo: Bart Wursten).
Figure 7.
Species of Equilabium 2. AEquilabium laxiflorum (Photo: Bart Wursten) BE. petiolare (Photo: Neil Crouch) CE. subincisum (Photo: K. Smitha) DE. stenophyllum (Photo: Bart Wursten).
Type species.
Equilabium laxiflorum (Benth.) Mwany., A.J.Paton & Culham.
Description.
Annual or perennial herbs or soft wooded shrubs, rarely woody shrubs, sometimes with a persistent woody or fleshy rootstock. Leaves opposite. Inflorescence thrysoid, usually lax; cymes sessile or pedunculate, 1–3(–5, very rarely –7); bracts subtending cymes persistent. Flowers pedicelate. Calyx funnel-shaped, straight with pedicel attaching symmetrically at the calyx base, not opposite posterior lip, two-lipped; throat glabrous within, open; posterior lip 1-lobed, usually broader than anterior lobes, shortly decurrent on tube or not; anterior lip 4-lobed; lobes triangular or lancolate; lateral lobes broader or equal to median lobes and held midway between the posterior lobe and the median lobes of the anterior lip; median lobes triangular to lanceolate. Corolla two-lipped with lips equal in length; tube sigmoid, sometimes strongly so, narrow and parallel-sided at base and expanding towards throat; posterior lip 4-lobed erect or ascending; anterior lobe horizontal, cucullate, sometimes frilled at apex. Stamens 4; filaments not fused together, held within the anterior lip. Style bifid with lobes subulate. Nutlets ovoid.
42 species mainly in Tropical Africa but including two in India.
Equilabium acaule
(Brummitt & Seyani) Mwany., Culham & A.J.Paton comb. nov.
DBEFD0556EEC554496DD7906D37AB609
urn:lsid:ipni.org:names:77201333-1
Plectranthus acaulis Brummitt & Seyani, Kew Bull. 42: 695. 1987. Type: Malawi, Nyika Plateau, 8. Dec. 1965, Banda 760 (holotype: K; isotype: MAL).
Distribution.
Malawi to Zambia (Nyika Plateau).
Equilabium agnewii
(Lukhoba & A.J.Paton) Mwany. & A.J.Paton comb. nov.
711E1CB515EE5F11BF9DBD2C91073DC0
urn:lsid:ipni.org:names:77201334-1
Plectranthus agnewii Lukhoba & A.J.Paton, Kew Bull. 55: 957. 2000. Type: Kenya, Nakuru National Park, W of Lake, Gillett 20847 (holotype: EA; isotype: K).
Distribution.
E. Tropical Africa.
Equilabium annuum
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
B17C544FD85E55CBB4CF9E643DE16EB1
urn:lsid:ipni.org:names:77201335-1
Plectranthus annuus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 270. 2009. Type: Tanzania, Masasi District: Ndanda Mission, Bidgood, Abdallah & Vollesen 2077 (holotype: K (sheet 2); isotypes: BR, C, DSM, EA, K (sheet 1), NHT).
Distribution.
S. Tanzania to Mozambique.
Equilabium caespitosum
(Lukhoba & A.J.Paton) Mwany. & A.J.Paton comb. nov.
FCECDABED747591683C62F135B5556D1
urn:lsid:ipni.org:names:77201336-1
Plectranthus caespitosus Lukhoba & A.J.Paton, Kew Bull. 58: 910. 2003, publ. 2004. Type: Kenya, E Mt Elgon, Irwin 286 (holotype: EA; isotype: K).
Distribution.
Kenya to N. Tanzania.
Equilabium candelabriforme
(Launert) Mwany. & A.J.Paton comb. nov.
6E4C9769421D5D12B182A90B7CB9173C
urn:lsid:ipni.org:names:77201337-1
Plectranthus candelabriformis Launert, Mitt. Bot. Staatssamml. München 7: 300. 1968. Type: Namibia, 16 km E of Runtu, 7 Mar. 1958, Merxmüller & Giess 1912 (holotype: M).
Distribution.
Tanzania to N. Namibia.
Equilabium cinereum
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
62AF822C3642523092FC576092051359
urn:lsid:ipni.org:names:77201338-1
Plectranthus cinereus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 247. 2009. Type: Tanzania, Mbulu District: Lake Manyara National Park, Ndala R., Greenway & Kanuri 11151 (holotype: K; isotype: EA).
Distribution.
Kenya to Tanzania.
Equilabium dissectum
(Brenan) Mwany. & A.J.Paton comb. nov.
D7F6EFB0A06153D8A6C6FA07DD5F2375
urn:lsid:ipni.org:names:77201339-1
Plectranthus dissectus Brenan, Mem. New York Bot. Gard. 9: 42. 1954. Type: Malawi, Zomba Plateau, 31 May 1946, Brass 16124 (holotype: K; isotypes: MO, PRE).
Distribution.
S. Malawi.
Equilabium dissectum var. dissectum
01ECF23866825ABAA37246622976C5BF
Plectranthus dissectus Brenan var. dissectus A.J.Paton, Fl. Zambes. 8(8): 224. 2013.
Equilabium dissectum
var. chambense (A.J.Paton) Mwany. & A.J. Paton comb. nov.
024134DB7F2F53F09BBF00EA097FF789
urn:lsid:ipni.org:names:77201340-1
Plectranthus dissectus Brenan var. chambensis A.J.Paton, Fl. Zambes. 8(8): 224. 2013. Type: Malawi, Mulanje, Chambe basin, 4 May1989, Goyder, Pope & Radcliffe-Smith 3205 (holotype: K).
Distribution.
S. Malawi.
Equilabium dolomiticum
(Codd) Mwany. & A.J.Paton comb. nov.
9739644D0B795AD09898B116CF07BD78
urn:lsid:ipni.org:names:77201341-1
Plectranthus dolomiticus Codd, Bothalia 15: 142. 1984. Type: South Africa, Transvaal, Farm Ostend, 7 km NW of Penge Mine, 10 Dec. 1982, Van Jaarsveld 7052 (holotype, PRE).
Distribution.
Zimbabwe to Limpopo.
Equilabium equisetiforme
(E.A.Bruce) Mwany. & A.J.Paton comb. nov.
06B2D414014253FD80DC35C51A96AD4F
urn:lsid:ipni.org:names:77201342-1
Coleus equisetiformis E.A. Bruce in Bull. Misc. Inform., Kew 1935: 285. 1935.
Plectranthus equisetiformis (E.A. Bruce) Launert in Mitt. Bot. Staatssamml. München 7: 299. 1968. Type: Tanzania, Mpwapwa, Kiboriani Mts, 19 Apr. 1932, Burtt 3892 (holotype: K).
Distribution.
Tanzania to N. Zambia.
Equilabium flaccidum
(Vatke) Mwany. & A.J.Paton comb. nov.
F3D6F235C07A5022A211C89BEE1DDAC5
urn:lsid:ipni.org:names:77201343-1
Coleus flaccidus Vatke in Linnaea 43: 9. 1881.
Plectranthus flaccidus (Vatke) Gürke, Bot. Jahrb. Syst. 19: 206. 1894. Type: Tanzania, Bagamoyo District: Ruvu [Kingani] R., Hildebrandt 1264 (holotype: B, destroyed; isotype: K).
Plectranthus fragilis Baker in Fl. Trop. Afr. 5: 407. 1900. Type: Zanzibar, Kirk s.n. (holotype: K).
Distribution.
S. Somalia to Mozambique, Comoros.
Equilabium glandulosum
(Hook. f.) Mwany. & A.J.Paton comb. nov.
A703241C3C485415A4CE37BB3D68C4B7
urn:lsid:ipni.org:names:77201344-1
Plectranthus glandulosus Hook.f., J. Proc. Linn. Soc., Bot. 6: 17. 1862. Type: Bioko [Fernando Po], Clarence Peak, Mann 318 (holotype: K).
Plectranthus hylophilus Gürke, Bot. Jahrb. Syst. 19: 283. 1894. Type: Cameroon, Mann’s Spring, Preuss 815 (holotype B, destroyed; isotype: K).
Plectranthus laxiflorus var. genuinus Briq., Bot. Jahrb. Syst. 19: 180. 1894., nom. inval.
Germanea laxiflora var. genuina (Briq.) Hiern, Cat. Afr. Pl. 1: 861. 1900., nom. inval. Based on: Angola, Pungo Andongo, Welwitsch 5545 (BM, COI, K, PRE).
Plectranthus phryxotrichus Briq., Bull. Soc. Roy. Bot. Belgique 37: 66. 1899. Type: Congo-Kinshasa, Dewèvre 862 a (holotype: BR).
Plectranthus urticoides Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 412. 1900. Type: Angola, Pungo Andongo, Welwitsch 5545 (holotype: LISU; isotype: K).
Plectranthus almamii A.Chev., J. Bot. (Morot) 22: 120. 1909. Type: Guinea, Conakry, Diagussa, around Diagussa and Dalaba, Oct. 1907, A.Chevalier 18609 (holotype: P; isotype: P).
Distribution.
Widespread in Trop. Africa.
Equilabium goetzei
(Gürke) Mwany. & A.J.Paton comb. nov.
F75E54B95476502EBD582F0E162ADA91
urn:lsid:ipni.org:names:77201345-1
Plectranthus goetzei Gürke, Bot. Jahrb. Syst. 28: 469. 1900. Type: Tanzania, Iringa Dist., Rungemba, Mar. 1899, Goetze 724 (holotype: B, destroyed; isotype: BM).
Plectranthus salubenii Brummitt & Seyani, Kew Bull. 42: 690. 1987. Type: Malawi, Katumbi Escarpment, 41.5 km N of Rumphi, 26 Dec. 1970, Pawek 4155 (holotype: K).
Distribution.
SW. Tanzania to Zambia.
Equilabium gracile
(Suess.) Mwany. & A.J.Paton comb. nov.
71157292C0E05E7E8EA668EA79AAC2BF
urn:lsid:ipni.org:names:77201346-1
Plectranthus gracilis Suess., Trans. Rhodesia Sci. Assoc. 43: 52. 1951. Type: Zimbabwe, Marondera (Marandellas), 26 Feb. 1943, Dehn 748 (holotype: M).
Plectranthus zernyi Gilli, Ann. Naturhist. Mus. Wien 77: 39. 1973. Type: Tanzania, Matengo Highlands, WSW from Songea, Ugano, 2 Mar. 1936, Zerny 483 (holotype: W; isotype: K).
Plectranthus seyanii A.J.Paton & Brummitt, Kew Bull. 46: 523. 1991. Type: Malawi, Shire Valley, 16 km W of Zomba, 12 Mar. 1977, Brummitt & Patel 14821 (holotype: K; isotypes: MAL, SRGH, WAG).
Distribution.
S. Tanzania to S. Trop. Africa.
Equilabium intrusum
(Briq.) Mwany. & A.J.Paton comb. nov.
2C07F7D6CE5E56F28D206E137B98AD2D
urn:lsid:ipni.org:names:77201347-1
Plectranthus intrusus Briq., Bull. Herb. Boissier, sér. 2, 1: 834. 1901. Type: DRC, N’Dembo, ann. 1898, J.Gillet s.n. (holotype: BR).
Distribution.
DRC.
Equilabium janthinothryx
(Lebrun & L. Touss) Mwany. & A.J.Paton comb. nov.
72B8E98639D65ABBB0F64932FEB0BC18
urn:lsid:ipni.org:names:77201348-1
Plectranthus janthinothryx Lebrun & L.Touss., Bull. Jard. Bot. État Bruxelles 17: 69. 1943. Type: DRC, Rwindi, Lebrun 7930 (holotype: BR, isotype: K).
Distribution.
DRC and Uganda.
Equilabium jebel-marrae
(Wickens & B.Mathew) A.J.Paton comb. nov.
C3DD58E53F365EAD88019E33E239E5C2
urn:lsid:ipni.org:names:77201349-1
Plectranthus jebel-marrae Wickens & B.Mathew, Kew Bull. 25: 255. 1971. Type: Sudan, Darfur Province: Kulme, Sept. 1921. Lynes 589 (holotype: BM).
Distribution.
Sudan.
Equilabium kamerunense
(Gürke) Mwany. & A.J.Paton comb. nov.
C140C58CE4195CB28EDB5C39BD95B8DB
urn:lsid:ipni.org:names:77201350-1
Plectranthus kamerunensis Gürke, Bot. Jahrb. Syst. 19: 202. 1894. Type: Cameroon, West Buea, Preuss 1039 (holotype: B, destroyed).
Plectranthus mannii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 408. 1900. Type: Cameroon, Mt Cameroon, Mann 1947 (holotype: K).
Plectranthus keniensis S.Moore, J. Bot. 1901: 264. 1901. Type: Kenya, Mt Kenya, Mackinder s.n. (holotype: BM).
Plectranthus sangerawensis Gürke, Bot. Jahrb. Syst. 36: 134. 1905. Types: Tanzania, E Usambara Mts, Sangerawe, Mongu, Engler 872 & Amani, Warnecke in Herb. Amani 481 (syntypes: B, destroyed).
Plectranthus aberdaricus T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 27. 1930. Type: Kenya, Aberdares, R.E. & T.C.E. Fries 2784 (isotype: BM).
Plectranthus sangerawensis var. glabrior T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 25. 1930. Types: Kenya, W Mt Kenya, R.E. & T.C.E. Fries 514, 518, 843 & Aberdares, R.E. & T.C.E. Fries 2443a (syntypes: UPS; isosyntype of 518: K).
Plectranthus vicinus T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 27. 1930. Type: Kenya, Meru, R.E. & T.C.E. Fries 1580 (isotypes: K, MO).
Distribution.
Nigeria to Cameroon, E. Trop. Africa.
Equilabium laxiflorum
(Benth.) Mwany. & A.J.Paton, Bot. J. Linn. Soc. 188(4): 367. 2018
54923CB65CF15BE69A019D3FC57CB97E
Plectranthus laxiflorus Benth. in E.H.F.Meyer, Comm. Pl. Afr. Austr.: 228. 1838.
Germanea laxiflora (Benth.) Hiern, Cat. Afr. Pl. 1: 861. 1900.
Coleus laxiflorus (Benth.) Roberty, Bull. Inst. Fondam. Afrique Noire, Sér. A, Sci. Nat. 16: 331. 1954. Type: South Africa, KwaZulu-Natal, between Umzimkulu and Umkomaas Rivers, Drège 3586 (lectotype: K; designated by Codd (1975)).
Plectranthus albus Gürke, Bot. Jahrb. Syst. 19: 202. 1894. Type: Tanzania, Moshi District: Marangu, Volkens 744 (holotype: B, destroyed, isotype: BM).
Plectranthus violaceus Gürke, Bot. Jahrb. Syst. 19: 201. 1894. Type: Tanzania, Lushoto District: Usambara Mts, near Lutindi, Holst 3317 (holotype: B, destroyed; isotypes: K, W).
Plectranthus johnstonii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 411. 1900. Type: Tanzania, Kilimanjaro, H.H. Johnston 69 (holotype: K).
Plectranthus kondowensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 417. 1900. Type: Malawi, between Khondowe and Karonga, 1896, Whyte s.n. (holotype: K).
Plectranthus triflorus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 417. 1900. Type: Tanzania, Kilimanjaro, Thomson s.n. (holotype: K).
Plectranthus neumannii Gürke, Bot. Jahrb. Syst. 36: 131. 1905. Types: Ethiopia, Gofa, Neumann 168 & 179 (syntype: B, destroyed); neotype: Ethiopia, Gamo Gofa, Busa Forest, near Gidole. Ryding et al. 1630 (neotype: UPS; isoneotype: ETH designated by Ryding (2000)).
Plectranthus lilacinus Gürke, Bot. Jahrb. Syst. 41: 315. 1908. Types: Tanzania, Kilimanjaro, Engler 1797 (syntype: B, destroyed) & Moshi, Uhlig 137 (isosyntype: EA).
Coleus keniensis Standl., Smithsonian Misc. Collect. 68(5): 14. 1917. Type: Kenya, W of Mt Kenya, Mearns 1334 (holotype: US; isotype: BM).
Plectranthus fraternus T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 26. 1930. Type: Kenya W Mt Kenya, between Cole’s Mill and Forest Station, R.E. & T.C.E. Fries 911 (lectotype UPS; isolectotype K, MO; designated by Paton et al. (2009)).
Distribution.
Ethiopia to South Africa.
Equilabium longipes
(Baker) Mwany. & A.J.Paton comb. nov.
7BA213878CC85B499CA7F0C43DE07A02
urn:lsid:ipni.org:names:77201351-1
Plectranthus longipes Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 406. 1900. Type: Eritrea, Ghinda, Schweinfurth 178 (holotype: K, isotype: G).
Plectranthus amaniensis Gürke, Bot. Jahrb. Syst. 36: 134. 1905. Types: Tanzania, Lushoto District: E Usambara Mts, between Amani & Kwambugo, Engler 783 & same vicinity, Warnecke in Herb. Amani 413a (syntypes: B, destroyed).
Plectranthus ellenbeckii Gürke, Bot. Jahrb. Syst. 36: 132. 1905. Types: Ethiopia, Gallaland, Belana, Ellenbeck 334 (holotype: B, destroyed); neotype: Ethiopia, Harerge Region, Awalla Valley, Jansen 1952 (neotype: WAG, designated by Ryding (2000)).
Plectranthus panganensis Gürke, Bot. Jahrb. Syst. 41: 314. 1908. Type: Kenya, Kwale District: Samburu, Kassner 483 (lectotype: BM, designated by Paton et al. (2009)).
Plectranthus emanueli Buscal. & Muschl., Bot. Jahrb. Syst. 49: 484. 1913. Types: Kenya, Guaso Nyiro valley, but probably from elsewhere, von Aosta 1558 (holotype: B, destroyed); neotype: Eritrea, Nefasit, Mt Bizen, Ryding et al. 2034 (neotype: UPS; isoneotype: ASMU, ETH, designated by Ryding (2001)).
Plectranthus buraensis S.Moore, J. Bot. 54: 290. 1916. Type: Kenya, Teita District: Bura, A. Buchanan s.n. (holotype: BM).
Distribution.
Eritrea to Rwanda and Tanzania.
Equilabium mafiense
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
7EE497E22892528A888DAA6EF5E2DCB7
urn:lsid:ipni.org:names:77201352-1
Plectranthus mafiensis A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 267. 2009. Type: Tanzania, Mafia I., Kikutani, Greenway 5218 (holotype: EA; isotype: K).
Distribution.
Tanzania (Mafia I.).
Equlabium masukense
(Baker) Mwany. & A.J.Paton comb. nov.
61265E75F03C5B41835E9BFF5DE29846
urn:lsid:ipni.org:names:77201353-1
Plectranthus masukensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 405. 1900. Type: Malawi, Masuku (Masuka) Plateau, Whyte 299 (holotype: K).
Distribution.
Kenya to N. Zambia.
Equilabium masukense
(Baker) Mwany. & A.J.Paton var. masukense.
994A102254425A6FA4188A5A2BAC63E3
Orthosiphon nyasicus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 373. 1900.
Plectranthus nyasicus (Baker) M.R.Ashby, J. Bot. 76: 47. 1938. Type: Malawi, “north Nyassa”, 1896, Whyte s.n. (holotype: K).
Plectranthus brevipes Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 406. 1900. Type: Malawi, Nyika Plateau, Whyte s.n. (lectotype: K (K000430762), designated by Paton et al. (2009)).
Distriution.
Kenya to N. Zambia.
Equilabium masukense
(Baker) Mwany. & A.J.Paton var. dissectum (A.J.Paton) A.J.Paton comb. nov.
0BF42AA7235C5C8082C70DC80EF549F8
urn:lsid:ipni.org:names:77201354-1
Plectranthus masukensis var. dissectus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 264. 2009. Type: Tanzania, Ufipa District: Sumbawanga, Tatanda Mission, Bidgood, Mbago & Vollesen 2383 (holotype: K; isotype: DSM).
Distribution.
W. Tanzania.
Equilabium megafolium
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
9DF04E568C2857D39FCCDCE23DAFD1B1
urn:lsid:ipni.org:names:77201355-1
Plectranthus megafolius A.J.Paton, Fl. Zambes. 8(8): 217. 2013.
Plectranthus macrophyllus A.J.Paton, nom. illegit. in F.T.E.A., Lamiaceae: 262. 2009, non Blume, Bijdr. Fl. Ned. Ind. 14: 835, 1826. Type: Tanzania, Pare Dist., Mkomazi Game Reserve, 6 km on Kisiwani–Mnazi road, Abdallah & Vollesen 95/181 (holotype: K; isotypes: C, NHT, P).
Equilabium molle
(Aiton) Mwany. & A.J.Paton comb. nov.
9DF371E39DCE5B3284D8FC33A536FF72
urn:lsid:ipni.org:names:77201356-1
Ocimum molle Aiton, Hort. Kew. 2: 322. 1789.
Plectranthus mollis (Aiton) Spreng., Syst. Veg. 2: 690. 1825.
Plectranthus secundus Roxb., Fl. Ind. ed. 1832, 3: 20. 1832. Nom. superfl. based on Ocimum molle Aiton. Type: Cultivated, native of the East Indies. Introduced by Sir Joseph Banks in 1781 (holotype: BM).
Ocimum maypurense Roth, Nov. Pl. Sp.: 271. 1821.
Plectranthus maypurensis (Roth) Spreng., Syst. Veg. 2: 691. 1825. Type: India, Heyne s.n. in Wall. Cat. 2736C (holotype B, destroyed; isotypes: K(K000820180), K-W (K001116983)).
Plectranthus incanus Link, Enum. Hort. Berol. Alt. 2: 120. 1822. Type: Cultivated, (B, destroyed).
Plectranthus divaricatus Weinm., Syll. Pl. Nov. 1: 68. 1824. Type: Cultivated (not seen).
Ocimum cordifolium Hamilton in D.Don Prodr. Fl. Nepal.: 116. 1825. nom inval.
Plectranthus cordifolius Buch.-Ham. ex D.Don, Prodr. Fl. Nepal.: 116. 1825. Type: Nepal, Wall. Cat. 2736A (holotype: not at BM; neotype: K (K001116984), isoneotype: K (K000820186); K-W (K001116982) designated by Suddee et al. (2004)).
Distribution.
Indian Subcontinent to N. Myanmar.
Equilabium orbiculare
(Gürke) Mwany. & A.J.Paton comb. nov.
C81B943509F352D0B2CCFB050559B1D5
urn:lsid:ipni.org:names:77201357-1
Plectranthus orbicularis Gürke, Bot. Jahrb. Syst. 19: 203. 1894. Types: Tanzania, Usambara Mts, Upanga swamp, Holst 4159 (syntype: B, destroyed) & Zanzibar, Stuhlmann 826 (syntype: B, destroyed; isosyntype: HBG).
Plectranthus sphaerophyllus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 410. 1900. Types: Tanzania, Uzaramo District: Dar es Salaam, Kirk s.n. (syntype: K) & Zanzibar, Kirk s.n. (syntype: K).
Distribution.
E. Tanzania (incl. Zanzibar, Pemba).
Equilabium parvum
(Oliv.) Mwany. & A.J.Paton comb. nov.
53C4FB3DD1B95FBBA4C671C55866031B
urn:lsid:ipni.org:names:77201358-1
Plectranthus parvus Oliv., Trans. Linn. Soc. London, Bot. 2: 347. 1887. Type: Tanzania, Kilimanjaro, H.H. Johnston s.n. (holotype: K).
Distribution.
Uganda to N. Zambia.
Equilabium pauciflorum
(Baker) Mwany. & A.J.Paton comb. nov.
D6AA0B7F25CA56C58DB89D04C38923A9
urn:lsid:ipni.org:names:77201359-1
Plectranthus pauciflorus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 405. 1900. Type: Tanzania, Shinyanga District: Msalala [Misilala], Hannington s.n. (holotype: K).
Plectranthus ugandensis S.Moore, J. Linn. Soc., Bot. 37: 200. 1903. Type: Uganda, Kigezi District: Rukiga [Ruchigga], Bagshawe 532 (holotype: BM).
Plectranthus auriculatus Robyns & Lebrun, Rev. Zool. Bot. Africaines 16: 353. 1928. Type: DRC, Kivu, Saké, Katouzi, Robyns 2486 (holotype: BR, isotype: K).
Distribution.
DRC, East Africa, Zambia.
Equilabium petiolare
(Benth.) Mwany. & A.J.Paton comb. nov.
6D0182B438575D419A2D9E96B7745E59
urn:lsid:ipni.org:names:77201360-1
Plectranthus petiolaris Benth. in E.H.F.Meyer, Comm. Pl. Afr. Austr.: 228. 1838. Type: South Africa, Eastern Cape, between Umtata and Umzimvubu Rivers, 1836, Drège 4773b (K lectotype), lectotypified by Codd (1975).
Plectranthus kuntzei Gürke in C.E.O.Kuntze, Revis. Gen. Pl. 3(2): 260. 1898. Type: South Africa, KwaZulu-Natal, Clairmont, 18 Oct. 1913, Kuntze s.n. (holotype: NY; isotype: K).
Distribution.
S. Mozambique to South Africa.
Equilabium pinetorum
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
124D4EC046BE586DB88E1E160CB65CA4
urn:lsid:ipni.org:names:77201361-1
Plectranthus pinetorum A.J.Paton, Fl. Zambes. 8(8): 224. 2013. Type: Malawi, Mulanje, Sombani basin, 15 Feb. 1992, Goyder & Paton 3660 (holotype: K).
Distribution.
Malawi and E. Zimbabwe.
Equilabium pulcherissimum
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
DB68459AD03655CFB1AAD5793BA44E92
urn:lsid:ipni.org:names:77201362-1
Plectranthus pulcherissimus A.J.Paton, Fl. Zambes. 8(8): 214. 2013. Type: Zambia, Mwinilunga Dist., Kalene Hill, 5 km N of bridge over Zambezi R. on Kalene Hill–Jimbe Bridge road, 2 Mar.1995, Harder, Zimba, Luwiika & Nawa 2854 (holotype: K; isotype: MO).
Distribution.
DRC and Zambia.
Equilabium pubescens
(Baker) Mwany. & A.J.Paton comb. nov.
5149930812B15D4C9A2C6426F1F94458
urn:lsid:ipni.org:names:77201363-1
Plectranthus pubescens Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 416. 1900. Type: Malawi, Mt Malosa, Nov. 1896, Whyte s.n. (holotype: K).
Plectranthus manganjensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 406. 1900. Types: Malawi, Manganja Highlands, iv.1859, Kirk (syntype: K); Malawi, Mt Zomba, Dec. 1896, Whyte (syntype: K).
Plectranthus nyikensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 416. 1900. Type: Malawi, Nyika Plateau, n.d., Whyte 162 (holotype: K).
Distribution.
SW. Tanzania to Mozambique.
Equilabium radiatum
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
81D643A89B0C53FCA0A031C2BCED61D9
urn:lsid:ipni.org:names:77201364-1
Plectranthus radiatus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 277. 2009. Type: Tanzania, Tunduru District: 13 km NW of Tunduru, Milne-Redhead & Taylor 7842 (holotype: K).
Distribution.
S. Tanzania.
Equilabium rungwense
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
7A75AC6C99DD59BF85FDCA71375FB5E9
urn:lsid:ipni.org:names:77201365-1
Plectranthus rungwensis A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 252. 2009. Type: Tanzania, Rungwe District: NW slope of Mt Rungwe on N slope of Mwashitu Hill, above Sinihi stream, Mwasumbi 16222 (holotype: K; isotypes: DSM, MO).
Distribution.
Tanzania (Mt. Rungwe).
Equilabium scopulicola
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
4D8921D21E9157C5BF6F8A5E3C5F6FFC
urn:lsid:ipni.org:names:77201366-1
Plectranthus scopulicola A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 259. 2009. Type: Tanzania, Lushoto District: World’s View, 1.5 km W of Gologolo, Drummond & Hemsley 2846 (holotype: K (K000431939); isotypes: BR, EA, K).
Distribution.
Tanzania (W. Usambara Mts.).
Equilabium selukwense
(N.E.Br.) Mwany. & A.J.Paton comb. nov.
BDE4D3B48263581990487C15E9D17A92
urn:lsid:ipni.org:names:77201367-1
Plectranthus selukwensis N.E.Br., Bull. Misc. Inform. Kew 1906: 167. 1906. Type: Zimbabwe, Shurugwi (Selukwe), Nov. 1899, Cecil 123 (holotype: K).
Distribution.
Zambia to Zimbabwe.
Equilabium spananthum
(A.J.Paton, Friis & Sebsebe) A.J.Paton comb. nov.
D4FEC0C2BCB25907AD1801521DD26F5E
urn:lsid:ipni.org:names:77201368-1
Plectranthus spananthus A.J.Paton, Friis & Sebsebe, Webbia 73(2): 216. 2018. Type: Ethiopia, Somali Regional State, Afder Zone, El Kere Woreda (in Bale floristic province of the Flora of Ethiopia and Eritrea), on road from Imi to El Kere, 10 km N of El Kere, I. Friis et al. 15539 (holotype: ETH; isotypes: C, K).
Distribution.
Ethiopia.
Equilabium stenophyllum
(Baker) Mwany. & A.J.Paton comb. nov.
4552F6258D4E5A258D88423A09F10A08
urn:lsid:ipni.org:names:77201369-1
Plectranthus stenophyllus Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 402. 1900. Type: Malawi, Shire Highlands, near Blantyre, 1887, Last s.n. (holotype: K).
Distribution.
S. Tanzania to S. Trop. Africa.
Equilabium stenosiphon
(Baker) Mwany. & A.J.Paton comb. nov.
4B3F6768EE0A5B2BA73945E72B95889F
urn:lsid:ipni.org:names:77201370-1
Plectranthus stenosiphon Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 415. 1900. Type: Malawi, R. Shire, 1863, Kirk s.n. (lectotype: K designated by Paton et al. (2013)).
Plectranthus albocaeruleus N.E.Br., Bull. Misc. Inform. Kew 1901: 130. 1901. Type: Malawi, Zomba, plant cultivated at Kew raised from seed sent June1898, Mahon s.n. (holotype: K).
Distribution.
S. Malawi to C. Mozambique and Zimbabwe.
Equilabium stoltzii
(Gilli) Mwany. & A.J.Paton comb. nov.
CBEE084E29A45FCA839C5418F41650A2
urn:lsid:ipni.org:names:77201371-1
Plectranthus stolzii Gilli, Ann. Naturhist. Mus. Wien 77: 40. 1973. Type: Tanzania, Rungwe Dist., Kyimbila, 6 Oct. 1911, Stolz 914 (holotype: W; isotypes: B, C, G, K, LD, M, STU, Z).
Distribution.
SW. Tanzania to N. Malawi.
Equilabium subincisum
(Benth.) Mwany., Smitha & A.J.Paton comb. nov.
5761B1226F4158AA8522D813BC0BA200
urn:lsid:ipni.org:names:77201372-1
Plectranthus subincisus Benth. in N.Wallich, Pl. Asiat. Rar. 2: 16. 1830. Type: India, Tamil Nadu: Courtallum, 1829, Wallich, Cat. No. 2737 (holotype: K (K000820179); isotype: K-W (K001116985)).
Distribution.
S. India, Sri Lanka. Occurrence in Sri Lanka is questionable and it is likely to be extinct there (Cramer 1981).
Equilabium vesiculare
(A.J.Paton) Mwany. & A.J.Paton comb. nov.
6ED98D9990CB5794A9D290B562F1B904
urn:lsid:ipni.org:names:77201373-1
Plectranthus vesicularis A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 272. 2009. Type: Tanzania, Kilwa Dist., Kilunda, 6.5 km (4 mile), 24 Apr. 1971, Ludanga in MRC 1301 (holotype: K; isotype: EA).
Distribution.
Tanzania to N. Mozambique.
Equilabium viphyense
(Brummitt & Seyani) Mwany., Culham & A.J.Paton comb. nov.
A1C1B94C1D9E5FE6AC0503741AC320A6
urn:lsid:ipni.org:names:77201374-1
Plectranthus viphyensis Brummitt & Seyani, Kew Bull. 42: 692. 1987. Type: Malawi, Mzimba Dist., Viphya, 11 Jan. 1970, Pawek 3338 (holotype: K).
Distribution.
Tanzania to S. Trop. Africa.
Equilabium viphyense subsp. viphyense
227AF21629CF55F985D7676706B08504
Distribution.
Tanzania to S. Trop. Africa.
Equilabium viphyense subsp. zebrarum
(Brummitt & Seyani) Mwany. & A.J.Paton comb. nov.
4685FE2AB0BC5BF38F445A419C331D6B
urn:lsid:ipni.org:names:77201375-1
Plectranthus zebrarum Brummitt & Seyani, Kew Bull. 42: 694. 1987.
Plectranthus viphyensis subsp. zebrarum (Brummitt & Seyani) A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 276. 2009. Type: Malawi, Nyika Plateau, Chelinda, below No.1 dam, 27 Feb.1982, Brummitt, Polhill & Banda 16140 (holotype: K).
Distribution.
E. Zambia to N. Malawi.
Equilabium wollastonii
(S.Moore) Mwany. & A.J. Paton comb. nov.
5B90F2FD180351A4ACD87516501BCE91
urn:lsid:ipni.org:names:77201378-1
Plectranthus wollastonii S.Moore, J. Linn. Soc., Bot. 38: 272. 1908. Type: Uganda, E Ruwenzori, Wollaston s.n. (holotype: BM).
Distribution.
EC. Trop. Africa to S. Kenya.
Genus Plectranthus
Plectranthus
L’Hér., Stirp. Nov.: 84, verso 1788 nom. cons.
DC50FA9D169359FE96C017901EA3915C
Figure 8.
Species of Plectranthus 1. APlectranthus ambiguus (Photo: Neil Crouch) BP. brevicaulis (Photo: Jean Bosser) CP. brevimentus (Photo: Neil Crouch) DP. ciliatus (Photo: Neil Crouch) EP. fruticosus (Photo: Neil Crouch) FP. ecklonii (Photo: Neil Crouch).
Figure 9.
Species of Plectranthus 2. APlectranthus papilionaceus (Photo: Patrick Ranirison) BP. purpuratus subsp. montanus (Photo: Neil Crouch) CP. saccatus var. longitubus (Photo RBG Kew) DP. vestitus (Photo: Patrice Antilahimena).
Germanea Lam., Encycl. 2: 690. 1788. Lectotype species: G. urticifolia Lam. (here placed under Plectranthus fruticosus L’Hér.) designated by Heath (2001).
Lectotype species.
Plectranthus fruticosus L’ Hér. (typ. cons.).
Description.
Perennial, rarely annual herbs or soft-wooded shrubs, sometimes succulent; sometimes with a tuberous base. Leaves opposite. Inflorescence thrysoid, usually lax; cymes sessile, 1–3-flowered; bracts subtending cymes persistent. Flowers pedicelate. Calyx funnel-shaped straight, with pedicel attaching symmetrically at calyx base, not opposite posterior lip, two-lipped; throat glabrous within, open; posterior lip 1-lobed, broader than lobes of anterior lip, shortly decurrent on tube or not; anterior lip 4-lobed; lobes lanceolate; lateral lobes broader at base than medial lobes and held closer to the median lobes than the posterior lip; median lobes usually fused for a greater distance than the lateral and median lobe. Corolla two-lipped with lips equal in length; tube straight or gently curved downwards, often gibbous, saccate or almost spurred a base, more rarely dilating gradually towards throat; posterior lip 4-lobed erect, often becoming reflexed; anterior lip cucullate or concave straight or becoming reflexed. Stamens 4, very rarely only 2 fertile (P. zuluensis); filaments not fused together (anterior stamen and adjacent posterior stamen sometimes fused in P. alboviolaceus), held within the anterior lip or becoming free from it and sometimes coiling downwards away from style after anthesis. Style bifid with lobes subulate. Nutlets ovoid.
72 species in Southern and Tropical Africa and Madagascar and including one in Sri Lanka.
Plectranthus alboviolaceus
Gürke, Bot. Jahrb. Syst. 30: 397. 1901
90DF24CCA8025E028E74FB311E9A5233
Plectranthus alboviolaceus Gürke, Bot. Jahrb. Syst. 30: 397. 1901. Type: Tanzania, Rungwe District: Kalengalenga Mts, Goetze 1140 (isotype: E).
Plectranthus rhomboideus Gürke, Bot. Jahrb. Syst. 36: 135. 1905. Type: Tanzania, Kilimanjaro, Engler 1775 (holotype: B, destroyed).
Distribution.
Ethiopia to S. Trop. Africa.
Plectranthus ambiguus
(Bolus) Codd, Bothalia 8: 159. 1964
C79E57EDEC3A5512890A8D5CCD9F0C4D
Orthosiphon ambiguus Bolus, J. Linn. Soc., Bot. 18: 394. 1881. Type: South Africa, Eastern Cape, near Grahamstown, MacOwen (holotype: BOL; isotype: SAM).
Plectranthus coloratus E.Mey., Comm. Pl. Afr. Austr.: 228. 1838., nom. illeg., non. P. coloratus D.Don.
Plectranthus dregei Codd, Fl. Pl. Africa 32: t. 1244. 1957. Type: South Africa, between Umgazana and Umzimvubu rivers, 19 May 1832, Drège a (syntypes: BM, HBG, K (K000430874), MO, P(P00450719, P00450717), S); between Umizimkulu and Umkomass Rivers, 1 May 1832, Drège b (syntype: P(00450718).
Distribution.
South Africa: E. Cape Prov. to N. KwaZulu-Natal.
Plectranthus amplexicaulis
Hedge, Fl. Madag. 175: 203. 1998
8F409B028D2451E6A82881F73F0F6416
Plectranthus amplexicaulis Hedge, Fl. Madag. 175: 203. 1998. Type: Madagascar, Andrambovato, à l’Est de Fianarantsoa, H. Humbert & R.P.R. Capuron 28517 (holotype: P).
Distribution.
Madagascar.
Plectranthus antongilicus
Hedge, Fl. Madag. 175: 160. 1998
D721D77392865430A269FEE22F4E9998
Plectranthus antongilicus Hedge, Fl. Madag. 175: 160. 1998. Type: Madagascar, around Rantabe, baie d’Antongil, Perrier de la Bâthie 10475 (holotype: P; isotypes: MO, TAN).
Distribution.
Madagascar.
Plectranthus asymmetricus
A.J.Paton, Fl. Zambes. 8(8): 207. 2013
E6ED26F06E495D3B84C2A711F06EB6B5
Plectranthus asymmetricus A.J.Paton,, Fl. Zambes. 8(8): 207. 2013. Type: Zambia, Mbala Dist., Chinakila, Jembele Forest, 13 Jan.1965, Richards 19510 (holotype: K).
Distribution.
Zambia.
Plectranthus atroviolaceus
Hedge, Fl. Madag. 175: 156. 1998
B0C0907ED63E56DCB9CB22901E2AC4B1
Plectranthus atroviolaceus Hedge, Fl. Madag. 175: 156. 1998. Type: Madagascar, Haute vallée de la Rienana (bassin du Matitanana). 2e voyage. Humbert 3464 (holotype: P; isotype: E).
Distribution.
Madagascar.
Plectranthus betamponus
Hedge, Fl. Madag. 175: 175. 1998
3A46E4C489EA50F4BE8A1EC3F61C4AFB
Plectranthus betamponus Hedge, Fl. Madag. 175: 175. 1998. Type: Madagascar, Reserve Integrale #1 Betampona, approximately 3 km N of Fotsimavo, Schatz, Goldblatt, Rakotozafy & Randrianasolo 2682 (holotype: MO; isotype: P).
Distribution.
Madagascar.
Plectranthus bracteolatus
A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 245. 2009
55890083BE2851839E9F6E69986CE951
Plectranthus bracteolatus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 245. 2009. Type: Tanzania, Morogoro District: Uluguru Mts, Schlieben 4215 (holotype: K; isotypes: K, MO, PRE, SRGH).
Distribution.
Tanzania.
Notes.
More work is needed to be confident about the generic placement of this species. A preliminary investigation suggests a possible relationship to Isodon or a clade basal to the Ocimeae and Plectranthinae.
Plectranthus brevicaulis
(Baker) Hedge, Fl. Madag. 175: 154. 1998
B2A6C9D41B9D5941B4E516371EDAE9BF
Orthosiphon brevicaulis Baker, J. Linn. Soc., Bot. 21: 433. 1885. Type: Madagascar, central, Baron 2656 (holotype: K; isotype: P).
Distribution.
Madagascar.
Plectranthus brevimentus
T.J.Edwards, Bothalia 35: 149. 2005
AD608C3D5CB45BDB95CF310B0CB47359
Plectranthus brevimentus T.J.Edwards, Bothalia 35: 149. 2005. Type: South Africa, Eastern Cape; Lupatana River Gorge; ca 1km from sea, T.J.Edwards, D.G.A. Styles, N.Crouch, N.D.U.Bellstedt, C.J.Potgieter 3210 (holotype: NU).
Distribution.
South Africa: Eastern Cape Prov.
Plectranthus canescens
Benth ., Labiat. Gen. Spec.: 33. 1832
EE1E951BCA7757FCBE9E31623E004102
Plectranthus canescens Benth., Labiat. Gen. Spec.: 33. 1832. Type: Madagascar, Bojer s.n. (lectotype: K, designated by Hedge et al. (1998); isolectotype: P).
Distribution.
Madagascar.
Plectranthus capuronii
Hedge, Fl. Madag. 175: 164. 1998
3BE845847FB3515D8EF6E7E0803EE699
Plectranthus capuronii Hedge, Fl. Madag. 175: 164. 1998. Type: Madagascar, montagnes au nord de Mangindrano jusqu’aux sommets d’Ambohimirahavavy, Humbert & Capuron 25183 (holotype: P; isotypes: B, BR, K, MA, MO, NY, TAN, WAG).
Distribution.
Madagascar.
Plectranthus chimanimanensis
S.Moore, J. Linn. Soc., Bot. 40: 174. 1911
F36FF1DB2C7F5F668D6D1027DEB39E0A
Plectranthus chimanimanensis S.Moore, J. Linn. Soc., Bot. 40: 174. 1911. Type: Mozambique, Gazaland, 26 Sept. 1906, Swynnerton 2019 (holotype: BM; isotype: K).
Distribution.
E. Zimbabwe to W. Mozambique.
Plectranthus ciliatus
E.Mey., Comm. Pl. Afr. Austr.: 227. 1838
A95ECB15D0715D92B45150C066D485AB
Plectranthus ciliatus E.Mey., Comm. Pl. Afr. Austr.: 227. 1838. Type: South Africa, Natal, Omsamvubo [Umzimvubu] Drège 4777 (lectotype: K (K000430860), designated by Codd (1975)).
Plectranthus natalensis Gürke, Bull. Herb. Boissier 6: 552. 1898. Type: South Africa, Natal, Camperdown, Rehmann 7701 (holotype: Z).
Distribution.
South Africa and Swaziland.
Plectranthus clementiae
Hedge, Fl. Madag. 175: 168. 1998
4220B892D6985DE888173A13BBB27E6D
Plectranthus clementiae Hedge, Fl. Madag. 175: 168. 1998. Type: Madagascar:, prov. de Tuléar, près de la réserve intégrale d’Andohahela, parcelle 1, 33km sur la route vers Ranomafana, à partir de la RN 13, Clement, Phillipson & Rafamantanantsoa 2097 (holotype: E; isotypes: MO, P).
Distribution.
Madagascar.
Plectranthus cordatus
A.J.Paton & Phillipson nom. nov.
7CE40C2AD9C85E3FB8B9E33CD71484A9
urn:lsid:ipni.org:names:77201380-1
Plectranthus cardiophyllus Hedge, Fl. Madagasc. 175: 172. 1998, nom. illeg., non. P. cardiophyllus Hemsl. Type: Madagascar, Mt Vatovavy, bassin du Mananjary, Perrier de la Bâthie 10441 (holotype: P).
Distribution.
Madagascar.
Plectranthus decaryi
Hedge, Fl. Madag. 175: 182. 1998
C2D592DC3E6C5CC5B8C3B8D9F55DB80C
Plectranthus decaryi Hedge, Fl. Madag. 175: 182. 1998. Type: Madagascar, Mt Maromizaha, près d’Analamazaotra, Perrier de la Bâthie 16027 (holotype: P).
Distribution.
Madagascar.
Plectranthus delicatissimus
Hedge, Fl. Madag. 175: 197. 1998
59FF5F48B0125E8FB6A4E83D201F5C2B
Plectranthus delicatissimus Hedge, Fl. Madag. 175: 197. 1998. Type: Madagascar, prov. d’Antsiranana, réserve spéciale 5, Manongarivo, sud-est d’Ankaramy et à 1–3 km au sud d’Ambalafary, Schatz 3204 (holotype: MO; isotype: E).
Distribution.
Madagascar.
Plectranthus ecklonii
Benth. in A.P.de Candolle, Prodr. 12: 64. 1848
B7C5AE235E845828A850ACDC40FF6EFE
Plectranthus ecklonii Benth. in A.P.de Candolle, Prodr. 12: 64. 1848. Type: South Africa, Cape, slopes of Katberg, Ecklon s.n. (holotype: K).
Distribution.
South Africa.
Plectranthus elegans
Britten, Trans. Linn. Soc. London, Bot. 4: 36. 1894
DC7F2DF9BE1056B09DF0808466F4CA1A
Plectranthus elegans Britten, Trans. Linn. Soc. London, Bot. 4: 36. 1894. Type: Malawi, Mulanje, 1891, Whyte s.n. (holotype: BM).
Plectranthus mahonii (Baker) N.E.Br. ex Hook.f., Bot. Mag. 128: t. 7818. 1902.
Coleus mahonii Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 434. 1900. Type: Cultivated plant, raised from seed sent from Malawi by Mahon 19 Dec.1899 (holotype: K).
Distribution.
S. Malawi.
Plectranthus elegantulus
Briq., Bull. Herb. Boissier, sér. 2, 3: 1005 1903
3E00E09BF993518AABCB79D2D10BF06D
Germanea elegantula Briq., Bull. Herb. Boissier, sér. 2, 3: 1005. 1903. Type: South Africa, Natal, Karkloof, Rehmann 7368 (holotype: Z).
Distribution.
South Africa: KwaZulu-Natal.
Plectranthus ellipticus
Hedge, Fl. Madag. 175: 186. 1998
8004E0874F9C533BB7C95CA3F173AF93
Plectranthus ellipticus Hedge, Fl. Madag. 175: 186. 1998. Type: Madagascar, haute vallée de la Rienana, Hubert 3638 (holotype: P; isotype: E).
Distribution.
Madagascar.
Plectranthus emirnensis
(Baker) Hedge, Fl. Madag. 175: 187. 1998
6E28066F49F9567B80A126E88FF5AB67
Orthosiphon emirnensis Baker, J. Linn. Soc., Bot. 21: 433. 1885. Type: Madagascar, centre, Baron 1056 (lectotype: K, designated by Hedge et al. (1998)).
Orthosiphon hildebrandtii Vatke, Abh. Naturwiss. Vereins Bremen 9: 134. 1885. nom. superfl. based on Baron 1506.
Ocimum hildebrandtii (Vatke) Briq. in H.G.A.Engler & K.A.E.Prantl, Nat. Pflanzenfam. 4(3a): 372. 1897., nom. illeg.
Ocimum siphonanthum Briq., Bull. Herb. Boissier 2: 121. 1894. Type: Madagascar, Betsileo, near Ankafina, Hildebrandt 3947 (holotype: G, not seen; isotype: P).
Distribution.
Madagascar.
Plectranthus ernstii
Codd, Fl. Pl. Africa 47: t. 1855. 1982
4C6C98C3C449549786A168637E9471FB
Plectranthus ernstii Codd, Fl. Pl. Africa 47: t. 1855. 1982. Type: South Africa, Kwa Zulu Natal, Oribi Gorge, Fairacres Farm, van Jaarsveld 2196; Cultivated in Pretoria sub Garden 24529 in PRE 57984 (holotype: PRE; isotype: NBG).
Distribution.
South Africa: E. Cape Prov. to S. KwaZulu-Natal.
Plectranthus forsythii
Hedge, Fl. Madag. 175: 177. 1998
E92677DABD305E89862BB9F6D00FD4A0
Plectranthus forsythii Hedge, Fl. Madag. 175: 177. 1998. Type: Madagascar, entre Itandraka et Ambohimitombo, Forsyth Major 727 (holotype: K, isotype: BM).
Distribution.
Madagascar.
Plectranthus fruticosus
L’Hér., Stirp. Nov.: 85. 1788
5CE226FDF1E65551809D8EAF6E4E7037
Plectranthus fruticosus L’Hér., Stirp. Nov.: 85. 1788. Type: cultivated, illustration t.41 of L’Hér. Stirp. Nov. fasc. 4. 1788 (lectotype: Illustration, designated by Codd (1975)).
Germanea urticifolia Lam., Encycl. 2: 691. 1788.
Plectranthus urticifolius (Lam.) Salisb., Prodr. Stirp. Chap. Allerton: 88. 1796. Type: Cultivated, illustration, Tabl. Encycl. 3: 514. 1819 (lectotype: Illustration, designated by Codd (1975)).
Plectranthus galpinii Schltr., J. Bot. 1896: 393. 1896. Type: South Africa, Transvaal, Barberton, Rimers Creek, Galpin 939 (holotype: B, destroyed; isotype: PRE).
Plectranthus arthropodus Briq., Bull. Herb. Boissier, sér. 2, 3: 1073. 1903. Type: South Africa, Transvaal, Houtbosch, Rehman 6151 (holotype: Z).
Plectranthus charianthus Briq., Bull. Herb. Boissier, sér. 2, 6: 824. 1906. Type: South Africa, Transvaal, Houtbosch, Rehman 6157 (holotype: Z).
Plectranthus peglerae T.Cooke, Bull. Misc. Inform. Kew 1909: 378. 1909. Type: South Africa, Eastern Cape, Kentani, Pegler 377 (holotype: K; isotypes: GRA, PRE).
Plectranthus behrii Compton, J. S. African Bot. 11: 122. 1945. Type: South Africa, Pondoland, Lusikisiki, Behr, sub NBG 1252/31 (holotype: SAM; isotype: PRE).
Distribution.
S. Mozambique to South Africa.
Plectranthus gardneri
Thwaites, Enum. Pl. Zeyl.: 237. 1860
8744230AA3735097BFBDC2020EB999D7
Plectranthus subincisus Benth. var. gardneri (Thwaites) Hook. f. in Flora of British India 4: 622. 1885. Type: Sri Lanka, Newera Ellis, Gardner in Thwaites CP16 (holotype: PDA; isotype: K).
Distribution.
Sri Lanka.
Plectranthus gibbosus
Hedge, Fl. Madag. 175: 200. 1998
3BDA0BCB85765915B8C02D02DC893E67
Plectranthus gibbosus Hedge, Fl. Madag. 175: 200. 1998. Type: Madagascar, Fenerive, Decary 3922 (holotype: P(P00541407); isotype: P(P00541408)).
Distribution.
Madagascar.
Plectranthus grallatus
Briq., Bull. Herb. Boissier, sér. 2, 3: 1004. 1903
E077B642F1B7507088B3D12513BB6944
Germanea grallata Briq. loc. cit as alternative name). Type: South Africa, Natal, Mt Frere, Jan. 1895, Schlechter 6415 (holotype: Z; isotype: GRA).
Plectranthus transvaalensis Briq., Bull. Herb. Boissier, sér. 2, 3: 1005. 1903.
Germanea transvaalensis Briq., loc. cit as alternative name. Type: South Africa, Transvaal, Houtbosch, Rehmann 6154 (holotype: Z).
Plectranthus krookii Gürke, Ann. K. K. Naturhist. Hofmus. 20: 48. 1905. Type: South Africa, Griqualand East, R. Umzinklawa, Krook in Herb Penther 1698 (holotype: W).
Plectranthus praetervisus Briq., Bull. Herb. Boissier, sér. 2, 6: 825. 1906. Type: South Africa, Natal, Drakensberg, Mountain Prospect, Rehmann 6965 (holotype: Z).
Plectranthus cooperi T.Cooke, Bull. Misc. Inform. Kew 1909: 377. 1909. Type: South Africa: Orange River Colony, Cooper, 2982 (lectotype: K, designated here).
Distribution.
South Africa, Lesotho.
Plectranthus grandibracteatus
Hedge, Fl. Madag. 175: 152. 1998
60359C3234275274BBF18113A3C6BF7B
Plectranthus grandibracteatus Hedge, Fl. Madag. 175: 152. 1998. Type: Madagascar, vallée de l’Androranga, affluent de la Bemarivo aux env. Antongondriha, mont Anjenabe, Mubolt & Cuperon 24144 (holotype: P).
Distribution.
Madagascar.
Plectranthus guruensis
A.J.Paton, Fl. Zambes. 8(8): 205. 2013
D52BB1DCBF75510D934BC29346BEA99A
Plectranthus guruensis A.J.Paton, Fl. Zambes. 8(8): 205. 2013. Type: Mozambique, Gurué Dist., Rio Malema, Marope, 22 km from Gurué, 1 Aug. 1979, de Koning 7530 (holotype: LMU; isotype: LISC).
Distribution.
Mozambique.
Plectranthus hexaphyllus
Baker, J. Linn. Soc., Bot. 20: 231. 1883
4A80CFB1121D5D90A98B27EDE05F2AEA
Plectranthus hexaphyllus Baker, J. Linn. Soc., Bot. 20: 231. 1883. Type: Madagascar, central, Baron 1799 (holotype: K; isotype: P).
Distribution.
Madagascar.
Plectranthus hilliardiae
Codd, Bothalia 11: 282. 1974
FCDC7BB161E959E28AA62D083E9E8491
Plectranthus hilliardiae Codd, Bothalia 11: 282. 1974. Type: South Africa, Natal, near Umtamvuna River, Hilliard & Burtt 6767 (holotype: PRE; isotype: NU).
Distribution.
South Africa: E. Cape Prov. to KwaZulu-Natal.
Plectranthus hilliardiae subsp. hilliardiae
.
B7387B2005985832A93956A282EB625A
Distribution.
South Africa: KwaZulu-Natal.
Plectranthus hilliardiae subsp. australis
van Jaarsv. & A.E.van Wyk, Bothalia 31: 44. 2001
AD31483CE57F57958CD5D0F86B542CE2
Plectranthus hilliardiae subsp. australis van Jaarsv. & A.E.van Wyk, Bothalia 31: 44. 2001. Type: South Africa, Eastern cape, Mbotyi forest, van Jarsveld, Bellsredt, Dekker, van Wyk & Adams 16345 (holotype: NBG).
Distribution.
South Africa: E. Cape Prov.
Plectranthus hirsutus
Hedge, Fl. Madag. 175: 158. 1998
781F5654845A5D76A5DAF6ED216B0B17
Plectranthus hirsutus Hedge, Fl. Madag. 175: 158. 1998. Type: Madagascar, Mont Kalabenono, Perrier de la Bâthie 10397 (holotype: P).
Distribution.
Madagascar.
Plectranthus hoslundioides
Scott-Elliot, J. Linn. Soc., Bot. 29: 44. 1891
A3DDFF22D3C552D9B3AF53067EEA4BB6
Plectranthus hoslundioides Scott-Elliot, J. Linn. Soc., Bot. 29: 44. 1891. Type: Madagascar, near Fort Dauphin, Scott- Elliot 2645 (holotype: K; isotype: P).
Distribution.
Madagascar.
Plectranthus humbertii
Hedge, Fl. Madag. 175: 194. 1998
5A5E859D6BD35FB2BC5D0643A135C849
Plectranthus humbertii Hedge, Fl. Madag. 175: 194. 1998. Type: Madagascar, partie occidentale du massif du Marojejy, de la vallée de l’Ambatoharanana au bassin supérieur de l’Antsahaberoka, Humbert & Saboureau 31311 (holotype: P; isotype: P).
Distribution.
Madagascar.
Plectranthus incrassatus
Hedge, Edinburgh J. Bot. 60: 246. 2003
363A992C8C1D5DD399B648456C76E33A
Plectranthus incrassatus Hedge, Edinburgh J. Bot. 60: 246. 2003.Type: Madagascar: massif de l’Anjanaharibe, à l’ouest d’Andapa, basin de la Lokoho, Humbert, Capuron & Cours 24774 (holotype: P; isotype: K).
Plectranthus pachyphyllus Hedge, Fl. Madag. 175: 176. 1998., nom. illeg., non. P. pachyphyllus Gürke.
Distribution.
Madagascar.
Plectranthus laurifolius
Hedge, Fl. Madag. 175: 170. 1998
9F24D977866D51F986D9CABA663F8278
Plectranthus laurifolius Hedge, Fl. Madag. 175: 170. 1998. Type: Madagascar, vallée de la Lokoho, mont Ambodilaitra, au nord d’Andranomiforitra, Humbert 23277 (holotype: P; isotype: MO).
Distribution.
Madagascar.
Plectranthus linearis
Hedge, Fl. Madag. 175: 191. 1998
690F09F9994C53869C28B1F5FAA28396
Plectranthus linearis Hedge, Fl. Madag. 175: 191. 1998. Type: Madagascar, pentes occidentales du massif du Marojejy, bassin de la Lokoho, à l’est d’Ambalamanasy, Humbert & Capuron 22069 (holotype: P; isotypes: MO, P).
Distribution.
Madagascar.
Plectranthus longiflorus
Benth., Labiat. Gen. Spec.: 33. 1832
13B72BA4DB2B5CF5938E07926F38108E
Plectranthus longiflorus Benth., Labiat. Gen. Spec.: 33. 1832.Type: Madagascar, forêt de Beforona, Bojer s.n. (holotype: K; isotype: P).
Distribution.
Madagascar.
Plectranthus longipetiolatus
Hedge, Fl. Madag. 175: 147. 1998
68D41BCE5EF257EEA1A0C8DA1824BAE5
Plectranthus longipetiolatus Hedge, Fl. Madag. 175: 147. 1998. Type: Madagascar, massif de Beampingaratra, du col de Bevava au sommet de Bekoho, Humbert 6432 (holotype: P; isotypes: B, E, K, MO, WAG).
Distribution.
Madagascar.
Plectranthus lucidus
(Benth.) van Jaarsv. & T.J.Edwards, Bothalia 27: 5. 1997
B4F06025D2C35A8B86DA0294BE2A84E8
Plectranthus strigosus var. lucidus Benth. in A.P.de Candolle, Prodr. 12: 68. 1848. Type: Eastern Cape, Bathurst, Burchell 3924, (holotype: K).
Distribution.
South Africa: S. Cape Prov.
Plectranthus macilentus
Hedge, Fl. Madag. 175: 170. 1998
A7E828C4CB2B5A1EB6422FFD06EAA741
Plectranthus macilentus Hedge, Fl. Madag. 175: 170. 1998. Type: Madagascar, collines et plateaux de l’Ankarana du Nord, (Province de Diégo-Suarez), W de la grande route Mt Ambatopiraka, Antsiranana Humbert & Cours 32655 (holotype: P).
Distribution.
Madagascar.
Plectranthus malvinus
van Jaarsv. & T.J.Edwards, Bothalia 27: 1. 1997
74123B51B33351BFA0E4CD49D1CC22D7
Plectranthus malvinus van Jaarsv. & T.J.Edwards, Bothalia 27: 1. 1997. Type: Eastern Cape, 3129 (Port St Johns), Mount Sullivan, (-DA), E. van Jaarsveld & Bingham 10522 (holotype: NBG).
Distribution.
South Africa: E. Cape Prov.
Plectranthus mandalensis
Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 405. 1900
74F4AA4B99BC56C5A773408614F102A0
Plectranthus mandalensis Baker in D.Oliver & auct. suc. (eds.), Fl. Trop. Afr. 5: 405. 1900. Type: Malawi, Mandala, cultivated at Kew, Feb. 1893, Moir s.n. (holotype: K).
Distribution.
Malawi (Mt. Mulanje), Mozambique (Mt. Namuli).
Plectranthus membranaceus
(Scott-Elliot) Hedge, Fl. Madag. 175: 145. 1998
2434E96D519056F7AA1FB8FB7F75C5DD
Plectranthus canescens var. membranaceus Scott-Elliot, J. Linn. Soc., Bot. 29: 49. 1891. Type: Madagascar, between Angalampena and the sea, Scott-Elliot 3072 (holotype: K; isotype: P).
Distribution.
Madagascar.
Plectranthus mocquerysii
Briq., Annuaire Conserv. Jard. Bot. Genève 2: 233. 1898
B639645114B659CDB66AEA6D9DC5FD38
Plectranthus mocquerysii Briq., Annuaire Conserv. Jard. Bot. Genève 2: 233. 1898. Type: Madagascar, Maroa, forest inland from bay d’ Antongil, Mocquerys 239 (holotype: G).
Distribution.
Madagascar.
Plectranthus mzimvubuensis
van Jaarsv., Bothalia 34: 30. 2004
6344B3F5B6CE5FA6925EBCEB62E4D91B
Plectranthus mzimvubuensis van Jaarsv., Bothalia 34: 30. 2004. Type: South Africa, Eastern Cape, near Ludonga village, Mzimvubu River, Ecca group shale cliffs, Van Jaarsveld, Xaba, Harower & Zwide 92 (holotype: PRE).
Distribution.
South Africa: SE. Cape Prov.
Plectranthus oblanceolatus
Hedge, Fl. Madag. 175: 161. 1998
C66448B8D32E5D4EB41F776C94A3C529
Plectranthus oblanceolatus Hedge, Fl. Madag. 175: 161. 1998. Type: Madagascar, partie occidentale du Massif du Marojejy, de la vallée de l’Ambatoharanana au bassin supérieur de l’Antsahaberoka, Humbert & Saboureau 31614 (holotype: P; isotype: P).
Distribution.
Madagascar.
Plectranthus oertendahlii
T.C.E.Fr., Acta Horti Gothob. 1: 253. 1924
77F5939DC9125CAC8D27BEA28824A30C
Plectranthus oertendahlii T.C.E.Fr., Acta Horti Gothob. 1: 253. 1924. Type: cultivated from material collected near Port Shepstone, Uppsala, (holotype, UPS).
Distribution.
South Africa: S. KwaZulu-Natal.
Plectranthus ombrophilus
Hedge, Fl. Madag. 175: 166. 1998
76F7CAC2B4C05747B4AEFDBD904F6692
Plectranthus ombrophilus Hedge, Fl. Madag. 175: 166. 1998. Type: Madagascar, haute vallée de la Manampanihy, col de Saindro, Humbert 14005 (holotype: P; isotypes: B, E, G, MO, PRE).
Distribution.
Madagascar.
Plectranthus oribiensis
Codd, Fl. Pl. Africa 46: t. 1809. 1980
186933722A5957FE85687577C346B07D
Plectranthus oribiensis Codd, Fl. Pl. Africa 46: t. 1809. 1980. Type: South Africa, Port Shepstone, Oribi Gorge, van Jaarsveld 2198, cultivated in Pretoria sub Garden 24530 (holotype: PRE).
Distribution.
South Africa: KwaZulu-Natal.
Plectranthus papilionaceus
Ranir. & Phillipson, Candollea 62: 158. 2007
2E02B06139B3526FA1CD82685DE4D729
Plectranthus papilionaceus Ranir. & Phillipson, Candollea 62: 158. 2007. Type: Madagascar, Province de Diego-Suarez/Antsiranana: sous-préfecture de Vohémar, commune rurale de Daraina, forêt de Binara, P.Ranirison & L.Nusbaumer PR 993 (holotype: G; isotypes: K, MO, P, TEF).
Distribution.
Madagascar.
Plectranthus pichompae
Hedge, Fl. Madag. 175: 163. 1998
0138C0FC95EE5CFF918686E0762C0811
Plectranthus pichompae Hedge, Fl. Madag. 175: 163. 1998. Type: Madagascar, partie occidentale du massif du Marojejy, de la vallée de l’Ambatoharanana au bassin supérieur de l’Antsahaberoka, Humbert & Saboureau 31546 (holotype: P; isotype P).
Distribution.
Madagascar.
Plectranthus praetermissus
Codd, Fl. Pl. Africa 45: t. 1791 1979
16D6C004841C5B7A9831658786522F6A
Plectranthus praetermissus Codd, Fl. Pl. Africa 45: t. 1791. 1979. Type: South Africa, Port St Johns, Stutterrhiem sub PRE 57330 (holotype: PRE).
Distribution.
South Africa: SE. Cape Prov.
Plectranthus purpuratus
Harv., Thes. Cap. 1: 53. 1860
ECBB4BDCC61D5C55826DAC75D6750686
Plectranthus purpuratus Harv., Thes. Cap. 1: 53. 1860. Type: cultivated, ex Hort., Kew, from seed sent from Port Natal [KwaZulu-Natal: Durban], R. Vause s.n. (holotype: K).
Distribution.
South Africa.
Plectranthus purpuratus subsp. purpuratus
55E492A4B2BD5088A5E08263C10CDD35
Distribution.
South Africa: KwaZulu-Natal.
Plectranthus purpuratus subsp. montanus
van Jaarsv. & T.J.Edwards, Bothalia 27: 4. 1997
9C9542D8B5AC5F62AB4BB59CA4930494
Plectranthus purpuratus subsp. montanus van Jaarsv. & T.J.Edwards, Bothalia 27: 4. 1997. Type: Swaziland, 2631 (Mbabane): Mbabane, Compton 32207 (holotype: NBG).
Distribution.
South Africa: S. Mpumalanga to Swaziland.
Plectranthus purpuratus subsp. tongaensis
van Jaarsv. & T.J.Edwards, Bothalia 27: 2. 1997
FEA1C7B83B7D58F499E65EA5D6C8EAAD
Plectranthus purpuratus subsp. tongaensis van Jaarsv. & T.J.Edwards, Bothalia 27: 2. 1997. Type: South Africa, KwaZulu-Natal, 2732 (Ubombo): Kosi Bay, (-BB), Van Jaarsveld 12206 (holotype: NBG).
Distribution.
South Africa: KwaZulu-Natal.
Plectranthus reflexus
van Jaarsv. & T.J.Edwards, Fl. Pl. Africa 51: t. 2034 1991
191E3FBD51E752F5800927BB1FAD64D2
Plectranthus reflexus van Jaarsv. & T.J.Edwards, Fl. Pl. Africa 51: t. 2034 1991. Type: South Africa, Port St Johns, western embankment of Bulowe River, van Jaarsveld, Campher & Dekker 10156 (holotype, NBG).
Distribution.
South Africa: SE. Cape Prov.
Plectranthus rosulatus
Hedge, Adansonia, sér. 3, 27: 321. 2005
EE80C631A24856D4A3EE7DAFDBC1DBF2
Plectranthus rosulatus Hedge, Adansonia, sér. 3, 27: 321. 2005. Type: Cultivated plants under glass, Royal Botanic Garden Edinburgh; specimen from Munich Botanic Garden, originally grown from seed collected in NW Madagascar, c. 50 km west of Sambava, in moss cushions in rain forest, by Mangelsdorff HC22 (holotype: E).
Distribution.
NW. Madagascar.
Plectranthus rubropunctatus
Codd, Bothalia 11: 420. 1975
D4522FE29468538A90B8D8E583EB4AA0
Plectranthus rubropunctatus Codd, Bothalia 11: 420. 1975. Type: South Africa, Barberton, Nelshootge Forest Station, Strey 4081 (holotype: PRE).
Distribution.
South Africa: Limpopo to Swaziland.
Plectranthus rubroviolaceus
Hedge, Fl. Madag. 175: 151. 1998
2267087C577854DE8BB0360125854A0A
Plectranthus rubroviolaceus Hedge, Fl. Madag. 175: 151. 1998. Type: Madagascar, contreforts occidentaux du massif du Marojejy, près du col de Boanyanala, limite des bassins de la Lokoho et de l’Androranga, Humbert 23127 (holotype: P; isotypes: BR, G, MO, NY, S, TAN).
Distribution.
Madagascar.
Plectranthus saccatus
Benth. in E.H.F.Meyer, Comm. Pl. Afr. Austr.: 227. 1838
0D35B95CD4D75C12964435023D50AFF7
Plectranthus saccatus Benth. in E.H.F.Meyer, Comm. Pl. Afr. Austr.: 227. 1838. Type: South Africa, Cape, Omsamwubo [Umzimvubu river, near Port St Johns], Drège 4771 (lectotype: K (K000430872), designated by Codd 1975; isolectotypes: G, P, S).
Distribution.
South Africa: SE. Cape Prov. to KwaZulu-Natal.
Plectranthus saccatus subsp. saccatus
E51B98050C205F1D903EB43C3DDECF30
Distribution.
South Africa: SE. Cape Prov. to KwaZulu-Natal.
Plectranthus saccatus var. saccatus
A811943D252A543AAD87D015D0B9903E
Distribution.
SE. Cape Prov. to KwaZulu-Natal.
Plectranthus saccatus var. longitubus
AE8B205C16CA531E8C92F7A19FBBAFCB
Plectranthus saccatus var. longitubus Codd, Bothalia 11: 428. 1975. Type: South Africa, Natal, Ingwavuma District, 2732 (Ubombo), Gwalaweni forest (-AC), Edwards 2930 (holotype: PRE).
Distribution.
South Africa: KwaZulu-Natal.
Plectranthus saccatus subsp. pondoensis
van Jaarsv. & T.J.Edwards, Bothalia 27: 4. 1997
9B6D5874C470538FBD489B26215517E4
Plectranthus saccatus subsp. pondoensis van Jaarsv. & T.J.Edwards, Bothalia 27: 4. 1997. Type: South Africa KwaZulu-Natal, 3030 (Port Shepstone): Oribi Gorge, (-CA), E. van Jaarsveld 2201 (holotype: PRE).
Distribution.
South Africa: SE. Cape Prov. to S. KwaZulu-Natal.
Plectranthus scaposus
Hedge, Fl. Madag. 175: 157. 1998
F8E34777919555C4919489ECD792053B
Plectranthus scaposus Hedge, Fl. Madag. 175: 157. 1998. Type: Madagascar, Antsiranana, rés. nat. du Marojejy, pentes au nord d’Ambatosoratra, Miller 4232 (holotype: P; isotype: MO).
Distribution.
Madagascar.
Plectranthus secundiflorus
(Baker) Hedge, Fl. Madag. 175: 180. 1998
B2175DB4E9CC5477AC41BA30054327C9
Orthosiphon secundiflorus Baker, J. Linn. Soc., Bot. 21: 433. 1885. Type: Madagascar, central, Baron 1226 (holotype: K).
Distribution.
Madagascar.
Plectranthus strangulatus
A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 246. 2009
1601029D97B4558CBAE1FF79455F4869
Plectranthus strangulatus A.J.Paton, Fl. Trop. E. Afr., Lamiac.: 246. 2009. Type: Tanzania, Morogoro District: Uluguru Mts, Kitundu, E.M. Bruce 957 (holotype: K (K000431930); isotypes: BR, K (K000431929)).
Distribution.
Tanzania (Uluguru Mts.).
Plectranthus strigosus
Benth. ex E.Mey., Comm. Pl. Afr. Austr.: 229. 1838
409819377B235A12B956B271A2D3EDC4
Plectranthus strigosus Benth. ex E.Mey., Comm. Pl. Afr. Austr.: 229. 1838. Type: South Africa, Cape, Olifantshoek Forest (Alexandria), 1836, Eklon s.n. (lectotype: K (K000430853), designated by Codd 1975).
Plectranthus parviflorus Gürke in C.E.O.Kuntze, Revis. Gen. Pl. 3(2): 261. 1898., nom. illeg., non P. parviflorus Willd.
Plectranthus kuntzeanus Domin, Biblioth. Bot. 89: 564. 1928. Type: South Africa, East London, Kuntze s.n. (lectotype: NY, designated by Codd 1975).
Distribution.
South Africa: S. Cape Prov.
Plectranthus stylesii
T.J.Edwards, Bothalia 35: 149. 2005
316BE5369FBE54E7AC4BA86B45EDE606
Plectranthus stylesii T.J.Edwards, Bothalia 35: 149. 2005. Type: South Africa, Port St Johns, Msikaba River Gorge, c. 4km from the mouth, Edwards 3259 (holotype: NU; isotypes: E, GRA, K, PRE).
Distribution.
South Africa: Eastern Cape Prov.
Plectranthus swynnertonii
S.Moore, J. Linn. Soc., Bot. 40: 176. 1911
097220843D3058A697BE79DEE617D46D
Plectranthus swynnertonii S.Moore, J. Linn. Soc., Bot. 40: 176. 1911. Type: Zimbabwe, Chirinda forest, Swynnerton 337 (holotype: K).
Distribution.
S. Trop. Africa to South Africa, Mpumalanga.
Plectranthus trilobus
Hedge, Fl. Madag. 175: 205. 1998
B386CECC7A385A50BDCE2272A6C31296
Plectranthus trilobus Hedge, Fl. Madag. 175: 205. 1998. Type: Madagascar, dans le haut bassin de l’Andranomaizina, affluent de l’Imonty, au-dessous du col d’Etsilotsilo, Capuron SF 22442 (holotype: P).
Distribution.
Madagascar.
Plectranthus verticillatus
(L.f.) Druce, Rep. Bot. Soc. Exch. Club Brit. Isles 4: 640. 1916 (publ. 1917)
B5D1AB15F38E5ED8B0ADCDE77017F840
Ocimum verticillatum L.f., suppl. Pl.: 276. 1782.
Plectranthus thunbergii Benth., Labiat. Gen. Spec.: 37. 1832., nom. illeg. Type: South Africa, Cape, Thunbergs.n. (holotype: LINN 749.4).
Ocimum racemosum Thunb., Prodr. Pl. Cap.: 96. 1800. Type: South Africa, Cape, Houteniquas, Thunberg s.n. (holotype: UPS; isotype: SBT).
Plectranthus nummularius Briq., Bull. Herb. Boissier, sér. 2, 3: 1072. 1903. Type: South Africa, KwaZulu-Natal, Camperdown, n.d., Rehmann 7702 (holotype: Z).
Distribution.
S. Mozambique to South Africa.
Plectranthus vestitus
Benth., Labiat. Gen. Spec.: 32. 1832
A5FF108F6640526F8B0F01CB527F7066
Plectranthus vestitus Benth., Labiat. Gen. Spec.: 32. 1832. Type: Madagascar, Prov. de Bezanozano, Bojer s.n. (holotype: K).
Distribution.
Madagascar.
Plectranthus vinaceus
Hedge, Fl. Madag. 175: 148. 1998
39E59707D1F45D10826E398496924F0B
Plectranthus vinaceus Hedge, Fl. Madag. 175: 148. 1998. Type: Madagascar, sommet oriental du massif du Marojejy, ál’ouest de la haute Manantenina, affluent de la Lokoho, Humbert 22737 (holotype: P).
Distribution.
Madagascar.
Plectranthus zuluensis
T.Cooke, Bull. Misc. Inform. Kew 1909: 379. 1909
E6F01BD1B72058148AA969D3EF039F77
Plectranthus zuluensis T.Cooke, Bull. Misc. Inform. Kew 1909: 379. 1909. Type: South Africa, Natal, Gerrard 1675 (holotype: K).
Distribution.
South Africa: SE. Cape Prov. to Swaziland
Unplaced names
Anisochilus siamensis Ridl., J. Straits Branch Roy. Asiat. Soc. 59: 159. 1911. Type: Thailand, Bangsaphan Distr., Prachuap Khirikhan Province, Keith s.n. (not seen- see also Suddee & Paton (2009)).
Coleus anamallayensis Bedd., Madras J. Lit. Sci., ser. 2, 22: 73. 1861. Type: not seen, not cited in protologue.
Coleus aulihanensis Schweinf. & Volkens, Liste Pl. Somalis: 211. 1897. Type: Ethiopia, Aulihan, Dec. 1895 to Jan 1996, Schweinfurth & Volkens s.n. (not seen). Protologue suggests similarity to C. igniarius.
Coleus kisanfuensis De Wild., Contr. Fl. Katanga: 174. 1921. Types: Democratic Republic of Congo, Elizabethville, Hock s.n. (syntype: not seen); Kiasanfu, Hock s.n. (syntype: not seen).
Coleus luengerensis Gürke, Bot. Jahrb. Syst. 38: 167. 1906. Type: Tanzania, Usambara, Tanga, x. 1895, Heinsen 121 (syntype: B, destroyed); Sept. 1902, Engler 890 (syntype: B, destroyed).
Coleus mechowianus Briq., Bot. Jahrb. Syst. 19: 183. 1894. Type: Angola, Pungo Andongo, Mechow 123 (B, holotype destroyed). Protologue suggests similarity to C. tenuicaulis.
Coleus nervosus Briq., Bot. Jahrb. Syst. 19: 185. 1894. Type: DRC; South Central Kasongo; near the Lomami River, Pogge 1034 (B, holotype, destroyed).
Coleus parensis Gürke, Bot. Jahrb. Syst. 41: 317. 1908. Types: Tanzania, Pare District: foot of Pare Mts, Sengina–Simba, Engler 1620 (syntype: B, destroyed) & Uluguru Mts, Meyer 188 (syntype: B, destroyed).
Coleus petrophilus Gürke, Bot. Jahrb. Syst. 41: 319. 1908. Type: Tanzania, Lushoto District: W Usambara Mts, Mbalu, Engler 1488 (holotype: B, destroyed)
Coleus petersianus Vatke, Linnaea 40: 180. 1876. Type: Mozambique, Rios de Sena, Peters s.n. (holotype: B, destroyed). The protologue suggests that this is either C. bojeri or C. welwitschii.
Coleus poggeanus Briq., Bot. Jahrb. Syst. 19: 182. 1894. Type: DRC, Mussumba des Muata Jamwo, Jan. 1876, Pogge 364 (type: B, destroyed).
Coleus preussii Gürke, Bot. Jahrb. Syst. 19: 219. 1894. Type: Cameroon, Buea, Preuss 691 (B, destroyed, isotype); Dusen 309 (B, destroyed).
Coleus saxicola Gürke, Bot. Jahrb. Syst. 41: 316. 1908. Type: Tanzania, Lushoto District: W Usambara Mts, Kwai, Eick 27 (syntype: B, destroyed) & Engler 2250 (syntype: B, destroyed).
Coleus subscandens Gürke, Bot. Jahrb. Syst. 41: 319. 1908. Type: Tanzania, Lushoto District: W Usambara Mts, Mlalo, Engler 1394 (holotype: B, destroyed).
Coleus ulugurensis Gürke, Bot. Jahrb. Syst. 28: 470. 1900. Type: Tanzania, Morogoro District: NE Uluguru Mts, Kitope, Stuhlmann 9060 (holotype: B, destroyed).
Coleus viridis Briq., Bot. Jahrb. Syst. 19: 181. 1894. Type: DRC, Mussumba, Pogge 365, (B, holotype, destroyed).
Coleus wugensis Gürke, Bot. Jahrb. Syst. 41: 318. 1908. Type: Tanzania, Lushoto District: W Usambara Mts, Wuga area, Engler 1165 (holotype: B, destroyed)
Plectranthus crenatus Gürke, Bot. Jahrb. Syst. 19: 208. 1894. Type: Tanzania, Lushoto District: Usambara Mts, Maschewa, Holst 8730 (holotype: B, destroyed).
Plectranthus eminii Gürke, Bot. Jahrb. Syst. 19: 207. 1894. Type: Tanzania, Bukoba, Stuhlmann 3882 (holotype: B, destroyed)
Plectranthus holstii Gürke, Bot. Jahrb. Syst. 19: 204. 1894. Type: Tanzania, Lushoto District: W Usambara Mts, Mlalo, Holst 248 (holotype: B, destroyed). Perhaps close to E. orbiculare (Gürke) Mwany. & A.J.Paton.
Plectranthus lycopifolius A.Rich., Tent. Fl. Abyss. 2: 181. 1850. Type Ethiopia, Tigre, near Adoa, Quartin-Dillon (holotype: P, not seen, no Richard specimen in P which matches protologue.).
Plectranthus mbaluensis Gürke, Bot. Jahrb. Syst. 36: 136. 1905. Type: Tanzania, W Usambara Mts, Mbalu, Engler 1479 (holotype: B, destroyed). Note. Similar to E. kamerunense.
Plectranthus lindblomii T.C.E.Fr., Notizbl. Bot. Gart. Berlin-Dahlem 11: 28. 1930. Type: Kenya, Mt Elgon, Kitosh, Lindblom s.n. (holotype: B, destroyed).
Plectranthus marquesii Gürke, Bol. Soc. Brot. 16: 70. 1896. Type: Angola, R. Cuillo, Marques 197 (holotype: B, destroyed).
Plectranthus mildbraedii Perkins in G.W.J.Mildbraed (ed.), Wiss. Erg. Deut. Zentr.-Afr. Exped., Bot. 2: 549. 1913. Type: Uganda, Virunga Mts (Vulkan Mts), Karissimbi, Mildbraed 1618 (holotype: B, destroyed). Note. Protologue suggests relationship to E. longipes or E. kamerunense.
Plectranthus microphyllus Gürke, Bot. Jahrb. Syst. 28: 467. 1900. Type: Tanzania, Morogoro District: E Uluguru Mts, Stuhlmann 8748 (holotype: B, destroyed).
Plectranthus miserabilis Briq., Bot Jarbh. Syst 19: 180. 1895. Type: Democratic Republic of Congo, South Central Lunda; Mukenge, in Bashilange district, Pogge 1022 (type: not seen).
Plectranthus monticola Gürke, Bot. Jahrb. Syst. 28: 468. 1900. Type: Tanzania, Morogoro District: Uluguru Mts, Lukwangule, Stuhlmann 9166 (holotype: B, destroyed). Schlieben 2854 (BM, LISC) has been determined as C. monticola. These specimens are C. trullatus. However the description is a better match for the closely related C. triangularis which also occurs in the Uluguru Mts.
Plectranthus nodosus Bellardi ex Colla, Herb. Pedem. 4: 475. 1835. Listed as dubious species, no type details.
Plectranthus pallidus Biroli ex Colla, Herb. Pedem. 4: 475. 1835. Listed as dubious species, no type details.
Plectranthus pendulus Gürke, Bot. Jahrb. Syst. 41: 314. 1908. Type: Tanzania, Lushoto District: W Usambara Mts, Engler 1180 (holotype: B, destroyed).
Plectranthus rupicola Gürke, Bot. Jahrb. Syst. 41: 313. 1908. Type: Tanzania, Lushoto District: W Usambara Mts, Mbalu, Engler 1457 (holotype: B, destroyed).
Plectranthus pratensis Gürke, Bot. Jahrb. Syst. 19: 201. 1894. Type: Tanzania, Kilimanjaro, Marangu, Volkens 634 & W Usambara Mts, Mlalo, Holst 459 (syntype: B, destroyed).
Plectranthus sakarensis Gürke, Bot. Jahrb. Syst. 41: 313. 1908. Type: Tanzania, Lushoto District: W Usambara Mts, Sakare, Engler 989a (holotype: B, destroyed)
Plectranthus schweinfurthii Sprenger, Wiener Ill. Gart.-Zeitung 19: 1. 1894, non C. schweinfurthii Vatke Type: Arabian Peninsula (type: not seen).
Plectranthus saxatilis Gürke, Bot. Jahrb. Syst. 36: 133. 1905. Types: Tanzania, Lushoto District: W Usambara Mts, Mlalo, Holst 345 & 611 & Engler 1485 (syntypes: B, destroyed).
Plectranthus stuhlmannii Gürke, Bot. Jahrb. Syst. 28: 469. 1900. Type: Tanzania, Morogoro District: S Uluguru Mts, Dundumi stream, Stuhlmann 9291 (holotype: B, destroyed) Protologue suggests similarity to E. laxiflorum.
Plectranthus usambarensis Gürke, Bot. Jahrb. Syst. 19: 207. 1894. Type: Tanzania, Lushoto District: Usambara Mts, Mlalo, Holst 487 (holotype: B, destroyed).
Saccostoma urticifolia Wall. ex Voigt Hortus Suburbanus Calcuttensis 449. 1845. Type: not seen.
Solenostemon bullatus Briq., Bot. Jahrb. Syst. 19: 180. 1894.
Coleus bullulatus Briq., Bull. Soc. Roy. Bot. Belgique 37: 69. 1899. nom. illeg. Type: DRC, Lulua River, 9.30 S, May 1876, Pogge 356. (holotype: not seen)
Excluded names
Coleus wulfenioides Diels: Orthosiphon wulfenioides (Diels) Hand.-Mazz.
Plectranthus adenanthus Diels: Isodon adenanthus (Diels) Kudô
Plectranthus adenolomus Hand.-Mazz.: Isodon adenolomus (Hand.-Mazz.) H.Hara
Plectranthus albidus Baker: Capitanopsis albida (Baker) Hedge
Plectranthus allenii C.H.Wright: Orthosiphon allenii (C.H.Wright) Codd
Plectranthus amethystoides Benth.: Isodon amethystoides (Benth.) H.Hara
Plectranthus angustifolius Dunn: Isodon angustifolius (Dunn) Kudô
Plectranthus assamicus Mukerjee: Isodon assamicus (Mukerjee) H.Hara
Plectranthus axillariflorus Honda: Isodon shikokianus var. intermedius (Kudô) Murata
Plectranthus barrelieri (Roth) Spreng.: Ocimum basilicum L.
Plectranthus benthamianus Miq.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus bifidocalyx Dunn: Isodon macrocalyx (Dunn) Kudô
Plectranthus bolusii T.Cooke: Orthosiphon thymiflorus (Roth) Sleesen
Plectranthus brandisii Prain: Isodon walkeri (Arn.) H.Hara
Plectranthus brevifolius Hand.-Mazz.: Isodon brevifolius (Hand.-Mazz.) H.W.Li
Plectranthus buckia Buch.-Ham. ex D.Don: Isodon ternifolius (D.Don) Kudô
Plectranthus buergeri Miq.: Isodon japonicus var. japonicus
Plectranthus bullatus Benth.: Isodon nilgherricus (Benth.) H.Hara
Plectranthus bullatus Robyns & Lebrun: Isodon ramosissimus (Hook.f.) Codd
Plectranthus bulleyanus Diels: Isodon bulleyanus (Diels) Kudô
Plectranthus calcicola Hand.-Mazz.: Isodon calcicola (Hand.-Mazz.) H.Hara
Plectranthus calcicola var. subcalvus Hand.-Mazz.: Isodon calcicola var. subcalvus (Hand.-Mazz.) H.W.Li
Plectranthus capillipes Benth.: Isodon capillipes (Benth.) H.Hara
Plectranthus cardiophyllus Hemsl.: Heterolamium debile var. cardiophyllum (Hemsl.) C.Y.Wu
Plectranthus cavaleriei H.Lév.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus chamaedrys H.Lév., = Triumfetta annuaL. (Malvaceae)
Plectranthus chenmui Y.Z.Sun: Isodon phyllopodus (Diels) Kudô
Plectranthus chienii Y.Z.Sun: Isodon hispidus (Benth.) Murata
Plectranthus coetsa Buch.-Ham. ex D.Don: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus colaniae Doan: Isodon colaniae H.Hara ex Suddee & A.J.Paton
Plectranthus coloratus D.Don: Platostoma coloratum (D.Don) A.J.Paton
Plectranthus coreanus Vaniot: Isodon inflexus (Thunb.) Kudô
Plectranthus crenulatus Hook.f.: Paralamium griffithii (Hook.f.) Suddee & A.J.Paton
Plectranthus daitonensis Hayata: Isodon amethystoides (Benth.) H.Hara
Plectranthus daoi Phuong: Paralamium griffithii (Hook.f.) Suddee & A.J.Paton
Plectranthus dawoensis Hand.-Mazz.: Isodon dawoensis (Hand.-Mazz.) H.Hara
Plectranthus dhankutanus (Murata) Bennet: Isodon dhankutanus Murata
Plectranthus dichromophyllus Diels: Isodon rubescens (Hemsl.) H.Hara
Plectranthus diffusus Merr.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus discolor Dunn: Isodon parvifolius (Batalin) H.Hara
Plectranthus drogotschiensis Hand.-Mazz.: Isodon barbeyanus (H.Lév.) H.W.Li
Plectranthus drosocarpus Hand.-Mazz.: Isodon macrocalyx (Dunn) Kudô
Plectranthus dubius Vahl ex Benth.: Isodon amethystoides (Benth.) H.Hara
Plectranthus effusus (Maxim.) Honda: Isodon effusus (Maxim.) H.Hara
Plectranthus enanderianus Hand.-Mazz.: Isodon enanderianus (Hand.-Mazz.) H.W.Li
Plectranthus eriocalyx Dunn: Isodon eriocalyx (Dunn) Kudô
Plectranthus esquirolii H.Lév.: Isodon lophanthoides var. lophanthoides
Plectranthus etiolatus Chiov.: Platostoma rotundifolium (Briq.) A.J.Paton
Plectranthus excisoides Y.Z.Sun ex C.H.Hu: Isodon excisoides (Y.Z.Sun ex C.H.Hu) H.Hara
Plectranthus excisus Maxim.: Isodon excisus (Maxim.) Kudô
Plectranthus excisus var. shikokianus Makino: Isodon shikokianus (Makino) H.Hara
Plectranthus fangii Y.Z.Sun: Isodon lophanthoides var. lophanthoides
Plectranthus flavidus Hand.-Mazz.: Isodon flavidus (Hand.-Mazz.) H.Hara
Plectranthus forrestii Diels: Isodon forrestii (Diels) Kudô
Plectranthus garrettii Craib: Platostoma calcaratum var. garrettii (Craib) Suddee
Plectranthus gerardianus Benth.: Isodon lophanthoides (Buch.-Ham. ex D.Don) H.Hara
Plectranthus gesneroides J.Sinclair: Isodon gesneroides (J.Sinclair) H.Hara
Plectranthus glaucocalyx Maxim.: Isodon japonicus var. glaucocalyx (Maxim.) H.W.Li
Plectranthus graciliflorus Benth.: Isodon lophanthoides var. graciliflorus (Benth.) H.Hara
Plectranthus grandifolius Hand.-Mazz.: Isodon grandifolius (Hand.-Mazz.) H.Hara
Plectranthus griffithii Hook.f.: Paralamium griffithii (Hook.f.) Suddee & A.J.Paton
Plectranthus grosseserratus Dunn: Isodon grosseserratus (Dunn) Kudô
Plectranthus hanceiformis H.Lév.: Teucrium bidentatum Hemsl.
Plectranthus henryi Hemsl.: Isodon henryi (Hemsl.) Kudô
Plectranthus hians Benth.: Isodon nigrescens (Benth.) H.Hara
Plectranthus hirtellus Hand.-Mazz.: Isodon hirtellus (Hand.-Mazz.) H.Hara
Plectranthus hispidus Benth.: Isodon hispidus (Benth.) Murata
Plectranthus hispidus var. glabrior Benth.: Isodon lophanthoides var. lophanthoides
Plectranthus hoslundioides Baker: Isodon ramosissimus (Hook.f.) Codd
Plectranthus hosseusii Muschl.: Isodon ternifolius (D.Don) Kudô
Plectranthus inconspicuus Miq.: Isodon inflexus (Thunb.) Kudô
Plectranthus indicus (Roth) Spreng.: Ocimum filamentosum Forssk.
Plectranthus inflexus (Thunb.) Vahl ex Benth.: Isodon inflexus (Thunb.) Kudô
Plectranthus inflexus var. macrophyllus Maxim.: Isodon inflexus (Thunb.) Kudô
Plectranthus inflexus f. macrophyllus (Maxim.) Kitag.: Isodon inflexus (Thunb.) Kudô
Plectranthus inflexus f. subglabrus Makino: Isodon inflexus (Thunb.) Kudô
Plectranthus inflexus var. umbrosus Maxim.: Isodon umbrosus (Maxim.) H.Hara
Plectranthus inflexus var. verticillatus Makino: Isodon inflexus (Thunb.) Kudô
Plectranthus intermedius Zoll. & Moritzi: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus intermedius Benth. nom. illeg.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus irroratus Forrest ex Diels: Isodon irroratus (Forrest ex Diels) Kudô
Plectranthus japonicus (Burm.f.) Koidz.: Isodon japonicus (Burm.f.) H.Hara
Plectranthus japonicus var. macraei (Benth.) S.R.Sriniv.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus javanicus (Blume) Benth.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus kameba (Okuyama ex Ohwi) Ohwi: Isodon umbrosus (Maxim.) H.Hara
Plectranthus kurzii Prain: Isodon kurzii (Prain) H.Hara
Plectranthus lanceus Nakai: Isodon shikokianus var. intermedius (Kudô) Murata
Plectranthus lasiocarpus Hayata: Isodon serra (Maxim.) Kudô
Plectranthus leptobotrys Diels: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus leucanthus Diels: Isodon phyllopodus (Diels) Kudô
Plectranthus leucophyllus Dunn: Isodon leucophyllus (Dunn) Kudô
Plectranthus longicornis F.Muell.: Platostoma longicorne (F.Muell.) A.J.Paton
Plectranthus longitubus Miq.: Isodon longitubus (Miq.) Kudô
Plectranthus longitubus var. albiflorus Makino: Isodon longitubus (Miq.) Kudô
Plectranthus longitubus var. contractus Maxim.: Isodon longitubus (Miq.) Kudô
Plectranthus longitubus var. effusus Maxim.: Isodon effusus (Maxim.) H.Hara
Plectranthus lophanthoides (Buch.-Ham. ex D.Don) Grierson & D.G.Long: Isodon lophanthoides (Buch.-Ham. ex D.Don) H.Hara
Plectranthus loxothyrsus Hand.-Mazz.: Isodon loxothyrsus (Hand.-Mazz.) H.Hara
Plectranthus macraei Benth.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus macranthus Hook.f.: Siphocranion macranthum (Hook.f.) C.Y.Wu
Plectranthus macrocalyx Dunn: Isodon macrocalyx (Dunn) Kudô
Plectranthus maddenii Benth. ex Hook.f.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus mairei H.Lév.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus marmoritis Hance: Orthosiphon thymiflorus (Roth) Sleesen
Plectranthus maximowiczii Miq.: Isodon japonicus var. glaucocalyx (Maxim.) H.W.Li
Plectranthus meeboldii W.W.Sm.: Isodon meeboldii (W.W.Sm.) Suddee
Plectranthus megathyrsus Diels: Isodon megathyrsus (Diels) H.Hara
Plectranthus melissoides Benth.: Isodon melissoides (Benth.) H.Hara
Plectranthus menthoides Moritzi: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus menthoides Willd.: Ocimum filamentosum Forssk.
Plectranthus menthoides Benth.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus micranthus Spreng.: Basilicum polystachyon (L.) Moench
Plectranthus mollissimus Wall. ex Benth.: Orthosiphon rubicundus var. rubicundus
Plectranthus monachorum (L.) Spreng.: Ocimum tenuiflorum L.
Plectranthus moschatus R.Br.,: Basilicum polystachyon (L.) Moench
Plectranthus moslifolius H.Lév.: Isodon nervosus (Hemsl.) Kudô
Plectranthus muliensis W.W.Sm.: Isodon muliensis (W.W.Sm.) Kudô
Plectranthus multiflorus Hochst. ex Benth.: Tetradenia multiflora (Benth.) Phillipson
Plectranthus namikawanus (Murata) Bennet: Isodon namikawanus Murata
Plectranthus nankinensis Spreng., Syst. Veg. 2: 691. 1825). based on Dentidia nankinensis Lour, Flora Cochinensis ed. 2: 448. 1793. Type: China, Nanjing, Loureiro s.n. (holotype: P) = Perilla ocymoides L. var. crispa Benth.
Plectranthus neglectus Dinter: Aeollanthus neglectus (Dinter) Launert
Plectranthus nepetifolius Benth.: Isodon wightii (Benth.) H.Hara
Plectranthus nervosus Hemsl.: Isodon nervosus (Hemsl.) Kudô
Plectranthus nigrescens Benth.: Isodon nigrescens (Benth.) H.Hara
Plectranthus nigrescens var. pilosus L.H.Cramer: Isodon nigrescens (Benth.) H.Hara
Plectranthus nilgherricus Benth.: Isodon nilgherricus (Benth.) H.Hara
Plectranthus nudipes Hemsl.: Siphocranion nudipes (Hemsl.) Kudô
Plectranthus oblongifolius Wall. ex Benth.: Isodon lophanthoides var. graciliflorus (Benth.) H.Hara
Plectranthus oreophilus Diels: Isodon oreophilus (Diels) A.J.Paton & Ryding
Plectranthus oreophilus var. elongatus Hand.-Mazz.: Isodon oreophilus (Diels) A.J.Paton & Ryding
Plectranthus oresbius W.W.Sm.: Isodon oresbius (W.W.Sm.) Kudô
Plectranthus ovatifolius Oliv.: Ocimum lamiifolium Hochst. ex Benth.
Plectranthus pachythyrsus Hand.-Mazz.: Isodon leucophyllus (Dunn) Kudô
Plectranthus palisotii Benth.: Plectranthus monostachyus (P.Beauv.) B.J.Pollard
Plectranthus paniculatus Baker: Isodon ramosissimus (Hook.f.) Codd
Plectranthus pantadenius Hand.-Mazz.: Isodon pantadenius (Hand.-Mazz.) H.W.Li
Plectranthus parviflorus R.Br.: Basilicum polystachyon (L.) Moench
Plectranthus parvifolius (Batalin) C.Pei: Isodon parvifolius (Batalin) H.Hara
Plectranthus parvifolius Talbot, J. Bombay Nat. Hist. Soc. 11: 238 plate x. 1897.
S. India. Type: Illustration cited above: Isodon
Plectranthus patchoulii C.B.Clarke ex Hook.f.: Microtoena patchoulii (C.B.Clarke ex Hook.f.) C.Y.Wu & S.J.Hsuan
Plectranthus pekinensis Maxim.: Isodon amethystoides (Benth.) H.Hara
Plectranthus pharicus Prain: Isodon pharicus (Prain) Murata
Plectranthus phulchokiensis (Murata) Bennet: Isodon phulchokiensis (Murata) H.Hara
Plectranthus phyllopodus Diels: Isodon phyllopodus (Diels) Kudô
Plectranthus phyllostachys Diels: Isodon phyllostachys (Diels) Kudô
Plectranthus pleiophyllus Diels: Isodon pleiophyllus (Diels) Kudô
Plectranthus polystachys Y.Z.Sun ex C.H.Hu: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus prainianus (H.Lév.) Dunn: Siphocranion macranthum (Hook.f.) C.Y.Wu
Plectranthus provicarii H.Lév.: Isodon bulleyanus (Diels) Kudô
Plectranthus pulneyensis Hook.f.: Isodon wightii (Benth.) H.Hara
Plectranthus racemosus Hemsl.: Isodon racemosus (Hemsl.) Murata
Plectranthus ramosissimus Hook.f.: Isodon ramosissimus (Hook.f.) Codd
Plectranthus rariflorus Hochst.: Endostemon obtusifolius (E.Mey.) N.E.Br.
Plectranthus repens Wall. ex Benth.: Isodon repens (Wall. ex Benth.) Murata
Plectranthus ricinispermus Pamp.: Isodon rubescens (Hemsl.) H.Hara
Plectranthus riparius Hochst.: Tetradenia riparia (Hochst.) Codd
Plectranthus rivularis Wight ex Hook.f.: Isodon rivularis (Wight ex Hook.f.) H.Hara
Plectranthus rosthornii Diels: Isodon rosthornii (Diels) Kudô
Plectranthus rubescens Hemsl.: Isodon rubescens (Hemsl.) H.Hara
Plectranthus rubicundus D.Don: Orthosiphon rubicundus (D.Don) Benth.
Plectranthus rufescens Benth.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus rugosiformis Hand.-Mazz.: Isodon rugosiformis (Hand.-Mazz.) H.Hara
Plectranthus rugosus Wall. ex Benth.: Isodon rugosus (Wall. ex Benth.) Codd
Plectranthus rugosus var. tomentosus Benth.: Isodon coetsa (Buch.-Ham. ex D.Don) Kudô
Plectranthus rupicola Dinter ex Gooss.: Aeollanthus buchnerianus Briq.
Plectranthus salicarius Hand.-Mazz.: Isodon nervosus (Hemsl.) Kudô
Plectranthus salvioides (B.Heyne ex Roth) Benth.: Perilla frutescens L.
Plectranthus scabrellus Benth.: Isodon nigrescens (Benth.) H.Hara
Plectranthus schimperi Vatke: Isodon schimperi (Vatke) J.K.Morton
Plectranthus schimperi Chiov., nom. illeg.: Isodon schimperi (Vatke) J.K. Morton
Plectranthus scrophularioides Wall. ex Benth.: Isodon scrophularioides (Wall. ex Benth.) Murata
Plectranthus sculponiatus Vaniot: Isodon sculponiatus (Vaniot) Kudô
Plectranthus serra Maxim.: Isodon serra (Maxim.) Kudô
Plectranthus setschwanensis Hand.-Mazz.: Isodon setschwanensis (Hand.-Mazz.) H.Hara
Plectranthus shikokianus (Makino) Makino: Isodon shikokianus (Makino) H.Hara
Plectranthus shikokianus var. intermedius (Kudô) Ohwi: Isodon shikokianus var. intermedius (Kudô) Murata
Plectranthus shikokianus var. occidentalis (Murata) Ohwi: Isodon shikokianus var. occidentalis Murata
Plectranthus sinensis Miq.: Isodon amethystoides (Benth.) H.Hara
Plectranthus smithianus Hand.-Mazz.: Isodon smithianus (Hand.-Mazz.) H.Hara
Plectranthus stocksii Hook.f.: Isodon lophanthoides var. lophanthoides
Plectranthus stracheyi Benth. ex Hook.f.: Isodon walkeri (Arn.) H.Hara
Plectranthus striatus Hook.f. & Thomson: Isodon lophanthoides var. lophanthoides
Plectranthus striatus Wight ex Hook.f.: Isodon wightii (Benth.) H.Hara
Plectranthus striatus Benth.: Isodon lophanthoides var. graciliflorus (Benth.) H.Hara
Plectranthus striatus var. gerardianus Hand.-Mazz.: Isodon lophanthoides (Buch.-Ham. ex D.Don) H.Hara
Plectranthus striatus var. graciliflorus (Benth.) Hand.-Mazz.: Isodon lophanthoides var. graciliflorus (Benth.) H.Hara
Plectranthus striatus var. kerrii Doan: Isodon lophanthoides var. lophanthoides
Plectranthus subacaulis Baker: Aeollanthus subacaulis (Baker) Hua & Briq.
Plectranthus succulentus R.A.Dyer & E.A.Bruce: Thorncroftia succulenta (R.A.Dyer & E.A.Bruce) Codd
Plectranthus tatei Hemsl.: Isodon lophanthoides var. graciliflorus (Benth.) H.Hara
Plectranthus tenuifolius W.W.Sm.: Isodon tenuifolius (W.W.Sm.) Kudô
Plectranthus ternifolius D.Don: Isodon ternifolius (D.Don) Kudô
Plectranthus teysmannii Miq.: Isodon teysmannii (Miq.) H.W.Li
Plectranthus thiothyrsus Hand.-Mazz.: Isodon leucophyllus (Dunn) Kudô
Plectranthus thorncroftii S.Moore: Thorncroftia thorncroftii (S.Moore) Codd
Plectranthus thymiflorus (Roth) Spreng.: Orthosiphon thymiflorus (Roth) Sleesen
Plectranthus trichocarpus Maxim.: Isodon trichocarpus (Maxim.) Kudô
Plectranthus tristis (Roth) Spreng.: Orthosiphon thymiflorus (Roth) Sleesen
Plectranthus tuberosus Roxb. ex Benth.: Orthosiphon rubicundus var. rubicundus
Plectranthus umbrosus (Maxim.) Makino: Isodon umbrosus (Maxim.) H.Hara
Plectranthus veronicifolius Hance: Isodon walkeri (Arn.) H.Hara
Plectranthus virgatus D.Don: Orthosiphon rubicundus var. rubicundus
Plectranthus viscosus (Roth) Spreng.: Endostemon viscosus (Roth) M.R.Ashby
Plectranthus volkensianus Muschl.: Isodon lophanthoides var. lophanthoides
Plectranthus volkmannae Dinter: Aeollanthus buchnerianus Briq.
Plectranthus walkeri Arn.: Isodon walkeri (Arn.) H.Hara
Plectranthus wardii C.Marquand & Airy Shaw: Isodon wardii (C.Marquand & Airy Shaw) H.Hara
Plectranthus websteri Hemsl.: Isodon websteri (Hemsl.) Kudô
Plectranthus whytei Baker: Isodon ramosissimus (Hook.f.) Codd
Plectranthus wightii Benth.: Isodon wightii (Benth.) H.Hara
Plectranthus wightii J.Graham: Isodon lophanthoides var. lophanthoides
Plectranthus wikstroemioides Hand.-Mazz.: Isodon wikstroemioides (Hand.-Mazz.) H.Hara
Plectranthus wui Y.Z.Sun: Isodon adenanthus (Diels) Kudô
Plectranthus yuennanensis Hand.-Mazz.: Isodon yuennanensis (Hand.-Mazz.) H.Hara
Plectranthus zenkeri Gürke: Isodon ramosissimus (Hook. F.) Codd
Plectranthus zollingeri Briq.: Isodon teysmannii (Miq.) H.W.Li
Acknowledgements
Thanks are due to the Norwegian Ministry of Foreign Affairs, SADC Biodiversity Support Programme MWI-04/333 for funding Montfort Mwanyambo during his PhD studies. Alan Paton received support for visits to P and C from the SYNTHESYS project http://www.synthesys.info/ which is financed by European Community Research Infrastructure Action under the FP7 “Capacities” Program. The following are thanks for providing photographs: Jean Bosser, Neil Crouch, Vincent Droissart, David Goyder, Witawat Keiwbang, Thamarat Phutthai, Naiyana Tetsana, Ernst van Jaarsveld, Bart Wursten. Staff of AAU, B, BK, BKF, BM, BOL; BR, C, CALI, COI, E, EA, ETH, FT, G, HBG, K, L, LE, LMA, LISC, LMU, MAL, MEL, MH, MO, NHT, P, PRE, RNG, S, SAM, STU, TCD, UPS, W, WAG are thanked for proividing access to herbarium specimens.
Citation
Paton AJ, Mwanyambo M, Govaerts RHA, Smitha K, Suddee S, Phillipson PB, Wilson TC, Forster PI, Culham A (2019) Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera. PhytoKeys 129: 1–158. https://doi.org/10.3897/phytokeys.129.34988
Funding Statement
Norwegian Ministry of Foreign Affairs, SADC Biodiversity Support Programme
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