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Journal of Integrative Bioinformatics logoLink to Journal of Integrative Bioinformatics
. 2019 Jun 20;16(2):20190021. doi: 10.1515/jib-2019-0021

The Systems Biology Markup Language (SBML): Language Specification for Level 3 Version 2 Core Release 2

Michael Hucka 1, Frank T Bergmann 1, Claudine Chaouiya 2, Andreas Dräger 3,4,5, Stefan Hoops 6, Sarah M Keating 7, Matthias König 8, Nicolas Le Novère 9, Chris J Myers 10, Brett G Olivier 11, Sven Sahle 12, James C Schaff 13, Rahuman Sheriff 7, Lucian P Smith 14,, Dagmar Waltemath 15, Darren J Wilkinson 16, Fengkai Zhang 17
PMCID: PMC6798823  PMID: 31219795

Abstract

Computational models can help researchers to interpret data, understand biological functions, and make quantitative predictions. The Systems Biology Markup Language (SBML) is a file format for representing computational models in a declarative form that different software systems can exchange. SBML is oriented towards describing biological processes of the sort common in research on a number of topics, including metabolic pathways, cell signaling pathways, and many others. By supporting SBML as an input/output format, different tools can all operate on an identical representation of a model, removing opportunities for translation errors and assuring a common starting point for analyses and simulations. This document provides the specification for Release 2 of Version 2 of SBML Level 3 Core. The specification defines the data structures prescribed by SBML as well as their encoding in XML, the eXtensible Markup Language. Release 2 corrects some errors and clarifies some ambiguities discovered in Release 1. This specification also defines validation rules that determine the validity of an SBML document, and provides many examples of models in SBML form. Other materials and software are available from the SBML project website at http://sbml.org/.

Keywords: Systems Biology Markup Language, Standards, Visualization, Representation


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