1Department of Biological Sciences, University of Idaho, Moscow, USA
2Faculty of Life and Environmental Sciences, University of Tsukuba, Ibaraki, Japan
3Integrative Research Center, Field Museum of Natural History, Chicago, USA
4Muroto Geopark Promotion Committee, Muroto Global Geopark Center, Kochi, Japan
5Center for Northeast Asian Studies, Tohoku University, Miyagi, Japan
6Graduate school of Life Sciences, Tohoku University, Miyagi, Japan
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