Review of recent taxonomic changes to the emerald moths (Lepidoptera: Geometridae: Geometrinae)

Abstract

Background

The subfamily Geometrinae (Lepidoptera: Geometridae), commonly known as emerald moths, is an ecologically diverse group of moths with over 2,500 described species. Many taxonomic and phylogenetic studies of Geometrinae have been undertaken in the past decade, resulting in hundreds of new taxonomic changes since online publication of the most recent checklist in December 2007.

New information

This review synthesises the last 12 years of alpha-taxonomic research in Geometrinae. A comprehensive list of Geometrinae genus- and species-group descriptions, synonymies, combinations and other taxonomic changes, made since 2007, is provided. Since 2007, the known species richness of Geometrinae has increased from 2,529 to 2,642 species; an updated list of all these species is presented in a supplementary spreadsheet.

Introduction

The family Geometridae is an incredibly diverse lineage of moths that contains over 23,000 described species, making it the second-most speciose family in all of Lepidoptera (Scoble and Hausmann 2007, van Nieukerken et al. 2011). Although the subfamily Geometrinae only comprises roughly one-tenth of this species richness, it is one of the more recognisable geometrid subfamilies because of the green colouration found in most adults and some larvae. Geometrinae are consequently known as ‘emerald moths’ and have been the subjects of studies on phenotypic plasticity and polyphenism (Greene 1989, Canfield et al. 2008). Many authoritative taxonomic works on Geometrinae have been published over the years, but with many new species being described annually, it does not take long for an update to become necessary.

Parsons et al. (1999) published a two-volume catalogue of the geometrid moths of the world, which is currently the most recently printed work that contains a comprehensive checklist of the global emerald moth fauna. However, between 1999 and 2007, one of this checklist’s co-authors, Malcolm Scoble, worked with Axel Hausmann to update the checklist; these revisions are hosted on the Lepidoptera Barcode of Life website. At the time of the most recent update (December 2007), the online checklist contained 269 genera and 2,529 species of Geometrinae. Since then, hundreds of taxonomic changes have been made in this subfamily, including over 100 new species. In this review, we catalogue the last 12 years of emerald moth taxonomy and update the list of geometrines provided by Scoble and Hausmann (2007) to include all newly-described emerald moth species.

Materials and methods

Updates and changes to the online checklist of Geometrinae are presented in alphabetical order by genus, following the format used by Scoble and Hausmann (2007). Since their checklist was last updated in December 2007, the taxonomic literature published between January 2008 and December 2019, inclusive, was consulted. It was also found that some taxonomic works (e.g. Beljaev 2007) were published prior to December 2007, but were not incorporated into the update; these are consequently included in this review.

Within each genus section, valid species names are listed in alphabetical order, with junior synonyms placed on an indented line following the corresponding senior synonym. Only genus- and species-group taxonomic changes in Geometrinae are discussed here; a review of recent family-group taxonomic changes can be found in Ban et al. (2018) and Murillo-Ramos et al. (2019), both of whom have also proposed new taxonomic changes to the geometrine tribes and subtribes, based on molecular phylogenetic data. Scoble and Hausmann (2007) did not include subspecies in their checklist, but post-2007 taxonomic changes to subspecies are discussed in this review.

The type of taxonomic change is indicated in parentheses. In this catalogue, the word 'new' and the abbreviation 'nov.' (novus, -a, -um) both denote that a taxonomic change was recent enough to not appear in the checklist of Scoble and Hausmann (2007). In this review, no taxonomic changes are proposed for the first time; this is further emphasised by the use of quotation marks surrounding each record of taxonomic change in the Results. Citations for recent taxonomic changes are provided in corresponding Remarks sections for each genus. The vast majority of taxonomic changes discussed here were proposed based solely on morphological evidence, such as variation in colour patterns, wing venation and genitalic characters of the adults. If molecular evidence were used to justify a taxonomic change, this is noted in the corresponding Remarks section.

If the status of a subspecies has been changed or a new synonymy has been proposed, the name of the associated valid species name is provided for context. Otherwise, species that have not undergone any taxonomic changes since the publication of Scoble and Hausmann (2007) are not included in the main text; a full list of all current Geometrinae species names is provided in the supplementary material. Similarly, synonyms that are not directly associated with a recent taxonomic change are excluded from the text.

Multiple Latin abbreviations for standard taxonomic terms are used throughout the text. Since the abbreviations themselves are not standardised across all taxonomic literature and do not appear at all in Scoble and Hausmann (2007), the notation used by Kitching et al. (2018) for a recent checklist of bombycoid moths was applied here in both the main text and supplementary material. These abbreviations and their definitions, are as follows:

“comb. nov.” – new combination

“comb. rev.” – revived combination

“gen. nov.” – new genus

“nom. nov.” – new replacement name

“nom. nud.” – nomen nudum (without description, thus unavailable)

“sp. nov.” – new species

“ssp. nov.” – new subspecies

“stat. nov.” – new status

“stat. rev.” – revived status

“syn. nov.” – new synonym

“syn. rev.” – revived synonym

Data resources

The list of taxonomic changes made in Geometrinae since publication of Scoble and Hausmann (2007) and the updated list of emerald moth species of the world are provided as tables (Excel format) in Suppl. material 1.

Checklists

List of Geometrinae genera and species associated with recent taxonomic changes

Acidaliastis

Hampson, 1896

299EF5A0-BD51-546F-99E0-802C91EA1ADC

• Acidaliastis porphyretica Prout, 1925

• Acidaliastis subbrunnescens Prout, 1916

Notes

The AfroMoths database (De Prins and De Prins 2019) states, without a citation, that Acidaliastis porphyretica was transferred to the genus Acidromodes Hausmann, 1996 and that Acidaliastis subbrunnescens was transferred to Hemidromodes Prout, 1916. After searching the literature, these names were found on other online species lists, but there did not appear to be any formal publications that proposed these new combinations. Thus, Acidaliastis porphyretica and Acidaliastis subbrunnescens are currently considered the valid names for these species.

Agathia

Guenée, [1858]

77C41F2B-6DDF-5CB0-98B4-E155DB33EA4B

• Agathia microlaetata Goyal, Kirti & Saxena, 2018 ("sp. nov.")

Notes

The name and locality of Agathia microlaetata appeared in Kirti et al. (2012), but this new species was not formally described until it appeared in Goyal et al. (2018).

Albinospila

Holloway, 1996

B10F57EE-5614-51E3-916B-0750ACF3A7C5

• Albinospila juancarlosi Tautel & Barrion-Dupo, 2017 (“sp. nov.”)

• Albinospila variifrons (Prout, 1917) (“comb. nov.”)

Notes

One new species was described (Tautel and Barrion-Dupo 2017). Albinospila variifrons was transferred from Comostola Meyrick, 1888 by Tautel and Barrion-Dupo (2017).

Aoshakuna

Matsumura, 1925 (“stat. rev.”)

3830F8B2-D432-5631-B602-75E0E57D3A42

• Nipponogelasma Inoue, 1946 (“syn. nov.”)

• Aoshakuna lucia (Thierry-Mieg, 1916) (“comb. nov.”)

• Aoshakuna sachalinensis Matsumura, 1925 (“syn. nov.”)

• Aoshakuna lucia ussurica Beljaev, 2007 (“ssp. nov.”)

Notes

One new subspecies was described (Beljaev 2007).

Aoshakuna was previously a junior synonym of Chlorissa Stephens, but was re-instated by Beljaev (2007). In the same revision, Beljaev (2007) subsequently designated Nipponogelasma a junior synonym of Aoshakuna. As a result of this synonymy, Nipponogelasma lucia was transferred to Aoshakuna, creating the new combination A. lucia. Beljaev (2007) then synonymised this species with A. sachalinensis, the type species of Aoshakuna.

Assachlora

Viidalepp & Lindt, 2012 (“gen. nov.”)

C54EDD5E-CB5A-5F51-9575-D5E3B0DD1C8C

• Assachlora assa (Druce, 1892) (“comb. nov.”)

• Assachlora julietae Viidalepp & Lindt, 2012 (“sp. nov.”)

• Assachlora mitigata (Prout, 1912) (“comb. nov.”)

Notes

One new species was described in this new genus (Viidalepp and Lindt 2012). Assachlora assa and A. mitigata were transferred from Phrudocentra Warren, 1895 by Viidalepp and Lindt (2012). Assachlora currently contains three species, with A. assa designated as the type species.

Bathycolpodes

Prout, 1912

F9BDDDDF-7B4F-5050-9DF7-DE51779BE45B

• Bathycolpodes parexplanata Karisch & Hoppe, 2010 (“sp. nov.”)

• Bathycolpodes roehrichti Karisch, 2010 (“sp. nov.”)

• Bathycolpodes scheeli Karisch & Hoppe, 2010 (“sp. nov.”)

• Bathycolpodes subferrata Prout, 1930 (“stat. nov.”)

• Bathycolpodes subfuscata (Warren, 1902)

Notes

Three new species were described (Karisch 2010). Although Henri Hoppe is credited with co-authorship of the new species Bathycolpodes parexplanata and B. scheeli in Karisch (2010), he is not credited as an author of the publication.

The subspecies Bathycolpodes subfuscata subferrata was elevated to the species B. subferrata by Karisch (2010).

Bustilloxia

Expósito, 1979

893DA29D-F9BF-541D-9A6C-DDC7BE3BDE2C

• Bustilloxia saturata (Bang-Haas, 1996)

• Bustilloxia saturata iberica Hausmann, 1995 (“syn. nov.”, followed by “stat. rev.”)

Notes

Leraut (2009) changed the status of Bustilloxia saturata iberica from a subspecies to a junior synonym of B. saturata. Müller et al. (2019) later revived B. s. iberica as a valid subspecies.

Chlorissa

Stephens, 1831

9CF0591E-8492-50C5-91C5-D863DBBD43D9

• Chlorissa archipelago Tautel, 2016 (“sp. nov.”)

• Chlorissa obliterata (Walker, 1863) (“syn. nov.”, followed by “stat. rev.”)

• Chlorissa viridata (Linnaeaus, 1758)

Notes

One new species was described (Tautel 2016). Leraut (2009) synonymised Chlorissa obliterata with C. viridata and Müller et al. (2019) subsequently revived its status as a valid species.

Chloristola

Holloway, 1996

31A7DDCD-6C4E-5772-A26B-8CA41D66DAAF

• Chloristola murzini Tautel, 2016 (“sp. nov.”)

Notes

One new species was described (Tautel 2016).

Chlorochromodes

Warren, 1896

AA16DB10-4163-5DC0-BC5D-6E4F26B9FAAA

• Comostolodes Warren, 1896 (“syn. nov.”)

• Chlorochromodes albicatena (Warren, 1896) (“comb. nov.”)

• Chlorochromodes chlorochromodes (Prout, 1916) (“comb. nov.”)

• Chlorochromodes dialitha (West, 1930) (“comb. nov.”)

• Chlorochromodes rhodocraspeda Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Chlorochromodes tenera (Warren, 1896) (“comb. nov.”)

• Chlorochromodes tumona Tautel, 2016 (“sp. nov.”)

Notes

Two new species were described (Han et al. 2012, Tautel 2016). Comostolodes was designated a junior synonym of Chlorochromodes by Han et al. (2012), who consequently formed new combinations for four species formerly in Comostolodes.

Chloroglyphica

Warren, 1894

EB5AEAF1-C436-5EA5-8172-D2A3E98F5277

• Chloroglyphica glaucochrista (Prout, 1916)

• Chloroglyphica glaucochrista grearia (Oberthür, 1916) (“syn. nov.”)

Notes

The status of Chloroglyphica glaucochrista grearia was changed from subspecies to junior synonym of C. glaucochrista by Han and Xue (2011a).

Chlororithra

Butler, 1889

1793074D-B595-5AF3-9745-E66731F73377

• Chlororithra fea Butler, 1889

• Chlororithra missioniaria Oberthür, 1916 (“stat. nov.”)

Notes

Chlororithra missioniaria was originally described as a variation of C. fea by Oberthür (1916). Parsons et al. (1999) instead treated C. missioniaria as a junior synonym of C. fea, so the name was absent from the checklist of Scoble and Hausmann (2007); however, prior to the publication of the checklist, C. missioniaria was designated a distinct species of Chlororithra by Han et al. (2006).

Comibaena

Hübner, [1823]

DFDDA539-4486-55E6-92B5-BE2B4274B0CA

• Comibaena auromaculata Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Comibaena bellula Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Comibaena birectilinea Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Comibaena decora Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Comibaena levequei Leraut, 2009 (“sp. nov.”)

• Comibaena nigromacularia (Leech, 1897)

• Comibaena delicatior (Warren, 1897) (“syn. nov.”)

• Comibaena parornataria Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Comibaena pictipennis Butler, 1880

• Comibaena pictipennis superornataria (Oberthür, 1916) (“syn. nov.”)

• Comibaena sheni Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Comibaena tibetensis Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Comibaena theodori Hausmann & Parisi, 2014 (“sp. nov.”)

Notes

Nine new species were described (Han et al. 2012, Hausmann et al. 2014).

Müller et al. (2019) noted that Comibaena levequei may be identical to C. pseudoneriaria Wehrli, 1926, but tentatively accepted it as a distinct species. Comibaena delicatior was synonymised with C. nigromacularia by Han and Xue (2011a). Comibaena pictipennis superornataria had its status changed from subspecies to junior synonym of C. pictipennis by Han et al. (2012).

Comostola

Meyrick, 1888

8D8DC79F-0D7A-525B-9DE7-FF3ED61ABDBA

• Comostola christinaria (Oberthür, 1916) (“comb. nov.”)

• Comostola desdemona Tautel, 2015 (“sp. nov.”)

• Comostola romblonensis Tautel & Barrion-Dupo, 2017 (“sp. nov.”)

• Comostola stueningi Tautel & Barrion-Dupo, 2017 (“sp. nov.”)

Notes

Three new species were described (Tautel 2015, Tautel and Barrion-Dupo 2017). Comostola christinaria was transferred from Hemistola Warren, 1893 by Han and Xue (2009).

Crypsiphona

Meyrick, 1888

E9D7EDCA-D804-52FB-865E-F31C462128CB

• Crypsiphona tasmanica Õunap & Viidalepp, 2009 (“sp. nov.”)

Notes

One new species was described (Õunap and Viidalepp 2009).

Dindica

Moore, 1888

2D7F92BF-5C96-56F6-B57E-3B29AB0D28D5

• Dindica purpurata Bastelberger, 1911

• Dindica wytsmani Prout, 1927 (“stat. rev.”)

Notes

Dindica purpurata wytsmani was elevated from subspecies to species by Pitkin et al. (2007).

Dindicodes

Prout, 1912

8C8C8446-529E-5089-AC96-6874A2DBEB7F

• Dindicodes albodavidaria (Xue, 1992) (“comb. nov.”)

• Dindicodes apicalis (Moore, 1888)(“comb. rev.”)

• Dindicodes apicalis hunana (Xue, 1992) (“comb. nov.”)

• Dindicodes costiflavens (Wehrli, 1933) (“comb. nov.”)

• Dindicodes davidaria (Poujade, 1895) (“comb. rev.”)

• Dindicodes ectoxantha (Wehrli, 1933) (“comb. nov.”)

• Dindicodes euclidiaria (Oberthür, 1913) (“comb. rev.”)

• Dindicodes harutai (Yazaki, 1992) (“comb. nov.”)

• Dindicodes harutai infuscatus (Yazaki, 1992) (“comb. nov.”)

• Dindicodes leopardinata (Moore, 1868) (“comb. rev.”)

• Dindicodes moelleri (Warren, 1893) (“comb. rev.”)

Notes

The 11 species and subspecies listed here were formally transferred from the genus Pachyodes Guenée, [1858] to Dindicodes by Pitkin et al. (2007).

Dioscore

Warren, 1907

D51FF1E8-38D5-590A-8089-9AD4A147BA09

• Dioscore kirke Lindt, Lennuk & Viidalepp, 2017 (“sp. nov.”)

• Dioscore vilu Lindt, Lennuk & Viidalepp, 2017 (“sp. nov.”)

Notes

Two new species were described (Lindt et al. 2017a).

Dysphania

Hübner, [1819]

155808DB-13A3-59B9-81E5-78B59BE26E6F

• Dysphania discalis aureolina Inoue, 2007 (“ssp. nov.”)

Notes

One new species was described (Inoue 2007).

Epichrysodes

Han & Stüning, 2007 (“gen. nov.”)

CBC0752C-7CB1-574E-B934-20BAD011D9F9

• Epichrysodes tienmuensis Han & Stüning, 2007 (“sp. nov.”)

Notes

Epichrysodes is currently a monotypic genus containing only the type species, E. tienmuensis; both the genus and the species were described by Han et al. (2007).

Epipristis

Meyrick, 1888

2E229E94-55BF-570E-9008-E4B9E7876E9F

• Epipristis pullusa Han & Xue, 2009 (“sp. nov.”)

• Epipristis roseus Expósito & Han, 2009 (“sp. nov.”)

Notes

Two new species were described (Han et al. 2009a).

Episothalma

Swinhoe, 1893

D3F8A63B-077B-51A5-9956-985002CFDF6D

• Episothalma cuspidata Xue & Wang, 2009 (“sp. nov.”)

• Episothalma irrobustaria Xue & Wang, 2009 (“sp. nov.”)

Notes

Two new species were described (Xue et al. 2009).

Eucyclodes

Warren, 1894

783E4343-F800-588F-B51C-A5704DFCD722

• Eucyclodes aphrodite (Prout, 1933) (“stat. nov.”)

• Eucyclodes gavissima (Walker, 1861)

• Eucyclodes hiyasata Tautel, 2016 (“sp. nov.”)

• Eucyclodes insolita Han & Zhang, 2019 (“sp. nov.”)

• Eucyclodes omeica (Chu, 1981) (“comb. nov.”)

Notes

Two new species were described (Tautel 2016, Zhang et al. 2019). Han and Xue (2011a) elevated Eucyclodes gavissima aphrodite (Prout, 1933) from subspecies to species and transferred Chloromachia omeica Chu, 1981 to Eucyclodes. Chloromachia was already considered a junior synonym of Eucyclodes (Parsons et al. 1999), but Han and Xue (2011a) were the first to formally publish the new combination E. omeica.

Geometra

Linnaeus, 1758

973CBEA8-F5B6-5FA6-8E95-910C359EAD44

• Geometra neovalida Han, Galsworthy & Xue, 2009 (“sp. nov.”)

Notes

One new species was described (Han et al. 2009b).

Gnophosema

Prout, 1912

5C4211BE-ABB7-55FC-9F10-A36074437F07

• Gnophosema isometra (Warren, 1888)

• Gnophosema leucites Wiltshire, 1980 (“stat. nov.”)

Notes

Gnophosema isometra leucites Wiltshire, 1980 was elevated from subspecies to species by Hausmann (2009).

Haruchlora

Viidalepp & Lindt, 2014 (“gen. nov.”)

3C5E68CB-85C5-519A-B31E-C21D13C33821

• Haruchlora maesi Viidalepp & Lindt, 2014 (“sp. nov.”)

Notes

Haruchlora is currently a monotypic genus containing only the type species, H. maesi; both the genus and the species were described by Viidalepp and Lindt (2014).

Hemistola

Warren, 1893

F2412824-074D-5751-B1BB-9345D1FF44E6

• Hemistola arcilinea Han & Xue, 2009 (“sp. nov.”)

• Hemistola asymmetra Han & Xue, 2009 (“sp. nov.”)

• Hemistola flavifimbria Han & Xue, 2009 (“sp. nov.”)

• Hemistola flavitincta Warren, 1897 (“comb. rev.”)

• Hemistola fui Chang & Wu, 2013 (“sp. nov.”)

• Hemistola glauca Han & Xue, 2009 (“sp. nov.”)

• Hemistola hanae Wu, 2019 (“sp. nov.”)

• Hemistola liliana (Swinhoe, 1892) (“comb. rev.”)

• Hemistola orbiculosoides Han & Xue, 2009 (“sp. nov.”)

• Hemistola piceacola Chang & Wu, 2013 (“sp. nov.”)

• Hemistola stueningi Han & Xue, 2009 (“sp. nov.”)

• Hemistola taiwanensis Chang & Wu, 2013 (“sp. nov.”)

• Hemistola viridimargo Han & Xue, 2009 (“sp. nov.”)

Notes

Eleven new species were described (Han and Xue 2009, Chang and Wu 2013, Wu 2019). The species Hemistola flavitincta and Hemistola liliana were transferred to Herochroma Swinhoe, 1893 by Parsons et al. (1999). Pitkin et al. (2007) implied this was an editorial error and, citing a complete absence of Herochroma diagnostic characters, transferred both species back to Hemistola.

Hemithea

Duponchel, 1829

2A8D74AB-0726-52B5-BA88-A1508DF3CC5E

• Hemithea aestivaria (Hübner, 1789)

• Hemithea aestivaria alboundulata (Hedemann, 1879) (“stat. nov.”, followed by “syn. rev.”)

Notes

Hemithea alboundulata was a junior synonym of H. aestivaria until Leraut (2009) elevated it to subspecies. Müller et al. (2019) found the justification for this taxonomic change to be too vague and, consequently, revived its status as a synonym of H. aestivaria.

Herochroma

Swinhoe, 1893

D9460764-45AA-57E9-98D7-1ACEBC8FA45A

• Herochroma costata Kirti, Goyal & Kaur, 2012 (nom. nud.)

• Herochroma subspoliata (Prout, 1916)

• Herochroma xuthopletes (Prout, 1934) (“stat. rev.”)

Notes

The name and locality of Herochroma costata were published in Kirti et al. (2012), but its description and diagnosis can only be found in the first author’s unpublished thesis. This species name is thus considered a nomen nudum.

Herochroma subspoliata xuthopletes (Prout, 1934) was elevated from subspecies to species by Pitkin et al. (2007).

Hypobapta

Prout, 1912

68E712F1-8BA7-5CB4-BFE5-D7173EF3E278

• Hypobapta tachyhalotaria Hausmann & Sommerer, 2009 (“sp. nov.”)

Notes

One new species was described (Hausmann et al. 2009).

Jodis

Hübner, [1823]

AF1E1F69-36A4-587F-9F53-2298F123D086

• Jodis altitudinis Tautel, 2016 (“sp. nov.”)

• Jodis argentea Tautel, 2016 (“sp. nov.”)

• Jodis berde Tautel, 2016 (“sp. nov.”)

• Jodis mystica Tautel, 2016 (“sp. nov.”)

• Jodis omeiensis (Chu, 1981) (“comb. nov.”)

• Jodis orientalis Wehrli, 1923 (“stat. nov.”)

• Jodis angulata Inoue, 1961 (“syn. nov.”)

• Jodis putata (Linnaeus, 1758)

• Jodis sibuyana Tautel, 2016 (“sp. nov.”)

• Jodis tomopunctata Tautel, 2016 (“sp. nov.”)

Notes

Six new species were described (Tautel 2016).

Han and Xue (2011a) created the new combination Jodis omeiensis, stating that this species was transferred from the genus Gelasma Warren, 1893; however, Gelasma had been designated a junior synonym of Maxates Moore, [1887] by Holloway (1996). Despite this synonymy, the combination Maxates omeiensis (Chu, 1981) does not appear to have ever been published between 1996 and 2011.

Beljaev (2007) elevated Jodis putata orientalis Wehrli, 1923 from subspecies to species and subsequently synonymised it with J. angulata.

Kuchleria

Hausmann, 1995

5E3BE2C5-B3A0-5AE1-ADB2-57CDC3144125

• Kuchleria menadiara Thierry-Mieg, 1893

• Kuchleria insignata Hausmann, 1995 (“syn. nov.”, followed by “stat. rev.”)

• Kuchleria garciapitai Expósito, 2006 (“syn. nov.”)

Notes

Kuchleria garciapitai was designated a junior synonym of K. insignata by Leraut (2009). In the same publication, Leraut (2009) claimed that K. insignata was a “synonym or subspecies” of Kuchleria menadiara Thierry-Mieg, 1893. Müller et al. (2019) treated this claim as a formal synonymy of K. insignata and K. menadiara and, subsequently, provided molecular and morphological evidence to justify elevating it back to species. Kuchleria garciapitai remains a junior synonym of K. insignata.

Lindachlora

Viidalepp & Lindt, 2012 (“gen. nov.”)

161A4BBD-6804-51B6-93BC-00BFEDBA4B32

• Lindachlora flaccida (Warren, 1909) (“comb. nov.”)

• Lindachlora tanystys (Prout, 1931) (“comb. nov.”)

Notes

The genus Lindachlora currently contains two species, both of which were transferred from Phrudocentra Warren, 1895 by Viidalepp and Lindt (2012), with L. flaccida designated as the type species.

Linguisaccus

Han, Galsworthy & Xue, 2012 (“gen. nov.”)

15BDF026-E5B7-524B-AEEB-91C24C41144E

• Linguisaccus minor Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Linguisaccus subhyalina (Warren, 1899) (“comb. nov.”)

Notes

Comostolodes subhyalina Warren, 1899 was transferred to Comibaena by Han and Xue (2011a) and then designated as the type species of the new genus Linguisaccus by Han et al. (2012).

Lissocentra

Viidalepp & Lindt, 2012 (“gen. nov.”)

077F3EC9-8A2D-5FEE-9898-F4102DA55192

• Lissocentra hydatodes (Warren, 1906) (“comb. nov.”)

• Lissocentra vitiosaria (Dognin, 1912) (“comb. nov.”)

Notes

The recently described genus Lissocentra currently contains two species, both of which were transferred from Phrudocentra by Viidalepp and Lindt (2012), with L. hydatodes designated as the type species.

Lissochlora

Warren, 1900

F1DB66A7-E0A0-51F3-83B0-69F72E5CAD7D

• Lissochlora hinojosae Lindt & Viidalepp, 2014 (“sp. nov.”)

• Lissochlora janamariae Lindt & Viidalepp, 2014 (“sp. nov.”)

• Lissochlora klausi Viidalepp & Lindt, 2019 (“sp. nov.”)

• Lissochlora niveiceps (Prout, 1912) (“comb. nov.”)

• Lissochlora senescens (Prout, 1917) (“comb. nov.”)

Notes

Three new species were described (Lindt and Viidalepp 2014, Viidalepp and Lindt 2019a). One of those species, Lissochlora hinojosae, was described in Lindt et al. (2014) with the specific epithet spelled ‘hinoyosae’, the first time it appears in both the English and Spanish versions of the abstract. However, it is spelled ‘hinojosae’ the first time it appears in the main text. The etymological remarks provided in Lindt et al. (2014) confirm that ‘hinojosae’ is the intended spelling (cf. § 24.2; 32.2.1.; 32.5 Code ICZN).

Lissochlora niveiceps and L. senescens were transferred from Phrudocentra by Viidalepp and Lindt (2012).

Lophophelma

Prout, 1912

0EADF3D8-3F8D-55FF-879D-5BDD6E10B7C9

• Lophophelma albapex (Inoue, 1988) (“comb. nov.”)

• Lophophelma costistrigaria (Moore, 1868) (“comb. rev.”)

• Lophophelma iterans (Prout, 1926) (“comb. nov.”)

• Lophophelma pingbiana (Chu, 1981) (“comb. nov.”)

• Lophophelma taiwana (Wileman, 1912) (“comb. rev.”)

• Lophophelma tanatoraja Sommerer, Stüning & Tautel, 2015 (“sp. nov.”)

• Lophophelma varicoloraria (Moore, 1868) (“comb. rev.”)

Notes

One new species was described (Sommerer et al. 2015). Pitkin et al. (2007) transferred six species to Lophophelma: five from the genus Pachyodes (Lophophelma albapex, L. costistrigaria, L. iterans, L. taiwana, L. varicoloraria) and one from the genus Terpna Herrich-Schäffer, 1854 (Lophophelma pingbiana).

Loxochila

Butler, 1881 (“stat. rev.”)

FF7B47F1-DFD7-5903-A101-67B790385CF7

• Loxochila burmensis (Han, Galsworthy & Xue, 2009) (“sp. nov.”)

• Loxochila fragilis (Oberthür, 1916) (“comb. nov.”)

• Loxochila kina (Swinhoe, 1893) (“comb. nov.”)

• Loxochila sinoisaria (Oberthür, 1916) (“comb. nov.”)

• Loxochila smaragdus (Butler, 1880) (“comb. rev.”)

• Loxochila tibeta (Chu, 1982) (“comb. nov.”)

Notes

One new species was described (Han et al. 2009b). Loxochila was treated as a junior synonym of Geometra in Parsons et al. (1999) and Scoble and Hausmann (2007). Han et al. (2009b)retained this classification in their revision of Geometra, which divided the genus into two species groups. The type species of Loxochila, at the time named Geometra smaragdus (Butler), served as the eponymous taxon of the smaragdus species group. The molecular phylogeny of Ban et al. (2018) demonstrated that the smaragdus group is a strongly-supported clade that also contains one species from a different genus (Tanaorhinus kina Swinhoe). Ban et al. (2018) consequently re-instated the generic status of Loxochila and transferred the species in the smaragdus group, including T. kina, to this genus.

Maxates

Moore, [1887]

91CBABDF-028B-5C2C-B99F-96F87A694805

• Maxates acyra (Prout, 1935) (“comb. nov.”)

• Maxates dissimulata (Walker, 1861)

• Maxates semiprotrusa (Inoue, 1989) (“syn. nov.”)

• Maxates elegante Tautel, 2015 (“sp. nov.”)

• Maxates persona Tautel, 2016 (“sp. nov.”)

• Maxates szechwanensis (Chu, 1981) (“comb. nov.”)

Notes

Two new species were described (Tautel 2015, Tautel 2016). Maxates acyra was transferred from Hemistola by Han and Xue (2009) and M. szechwanensis was transferred from Jodis by Han and Xue (2011a). Han and Xue (2011a) also synonymised M. semiprotrusa with M. dissimulata.

Metaterpna

Yazaki, 1992

26B7D94E-D38B-5B4D-8D16-CD7270FED1BF

• Metaterpna batangensis Han & Stüning, 2016 (“sp. nov.”)

Notes

One new species was described (Jiang et al. 2016).

Microloxia

Warren, 1893

0BA906A5-C421-5608-AE53-B576A58BABD4

• Microloxia aistleitneri Hausmann, 2009 (“sp. nov.”)

• Microloxia chlorissoides (Prout, 1912) (“comb. nov.”)

• Microloxia herbaria (Hübner, 1813)

• Microloxia herbaria virideciliata (Bubacek, 1926) (“syn. nov.”, followed by “stat. rev.”)

Notes

One new species was described (Hausmann 2009).

After synonymising Aoshakuna and Nipponogelasma, Beljaev (2007) transferred Nipponogelasma chlorissoides (Prout, 1913) to Microloxia.

Leraut (2009) synonymised Microloxia herbaria virideciliata with M. h. herbaria (Hübner, 1813). Müller et al. (2019) cited molecular and morphological evidence to justify elevating M. h. virideciliata back to subspecies.

Nemoria

Hübner, 1818

17BD7106-FE9B-5A21-8D22-9C864392EF35

• Nemoria “Nemoria” erina (Dognin, 1896)

• Nemoria “Nemoria” nigrisquama (Dognin, 1904)

• Nemoria yellowrosea Koçak & Kemal, 2008 (“nom. nov.”)

• Nemoria albilineata Cassino, 1927

Notes

The molecular phylogeny of Murillo-Ramos et al. (2019) indicates that the current concept of Nemoria Hübner is polyphyletic and that N. erina (Dognin) and N. nigrisquama (Dognin) do not belong in Nemoria, though there is insufficient evidence to describe new genera or create new combinations for these two species. Brehm et al. (2019) consequently suggested that their generic names are listed in quotation marks, pending further taxonomic study.

In her revision of Neotropical Nemoria, Pitkin (1993) transferred Lissochlora albilineata Warren, 1909 to the genus Nemoria. This new combination, Nemoria albilineata (Warren, 1909) consequently became a senior homonym of the Texan species Nemoria albilineata Cassino, 1927. This homonymy went unnoticed for over a decade, until Koçak and Kemal (2008) designated a replacement name for the junior homonym.

Neochloroglyphica

Han & Skou, 2019 (“gen. nov.”)

5406FEB8-A06F-5FAC-B5AB-8D71C8F4B9A6

• Neochloroglyphica perbella Han & Skou, 2019 (“sp. nov.”)

Notes

Neochloroglyphica is currently a monotypic genus containing only the type species, N. perbella; both the genus and the species were described by Han et al. (2019).

Neohipparchus

Inoue, 1944

B7A6CCD0-ADFE-5048-9EDE-28725116C25B

• Neohipparchus maculata (Warren, 1897)

• Chloroglyphica orhanti Herbulot, 1994 (“syn. nov.”)

Notes

Chloroglyphica orhanti was synonymised with Neohipparchus maculata by Han and Xue (2011a).

Neromia

Staudinger, 1898

4BFE5813-C386-5353-B8D6-628A27A548D3

• Neromia integrata Hausmann, 2009 (“sp. nov.”)

Notes

One new species was described (Hausmann and Hebert 2009).

Oenospila

Swinhoe, 1892

E6A920AC-D4DD-556A-915A-D262FC9476C2

• Oenospila sacculstrix Kirti, Goyal & Kaur, 2012 (nom. nud.)

Notes

The name and locality of this species were formally published in Kirti et al. (2012), but its description and diagnosis can only be found in the first author’s unpublished thesis. This species name is thus considered a nomen nudum.

Oospila

Warren, 1897

2A00E10E-950B-5BCB-AE2B-76C03B6F75D7

• Oospila absaloni Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila agnetaforslundae Lindt, Hausmann & Viidalepp, 2018 (“nom. nov.”)

• Oospila bifida Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila brehmi Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila bulava Lindt & Viidalepp, 2015 (“sp. nov.”)

• Oospila cristae Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila ehakernae Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila falcata Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila imula Dognin, 1911 (“stat. nov.”)

• Oospila loreenae Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila moseri Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila pallidaria boliviensis Lindt, Hausmann & Viidalepp, 2018 (“ssp. nov.”)

• Oospila pipa Lindt, Hausmann & Viidalepp, 2018 (“sp. nov.”)

• Oospila poirieri Lévêque & Viidalepp, 2015 (“sp. nov.”)

• Oospila similiplaga Warren, 1900 (“stat. nov.”)

• Oospila similiplaga bolarpata Lindt, Hausmann & Viidalepp, 2018 (“ssp. nov.”)

Notes

Eleven species and two subspecies were described (Lévêque and Viidalepp 2015, Lindt and Viidalepp 2015, Lindt et al. 2018). Lindt et al. (2018) designated Oospila agnetaforslundae as a replacement name for Oospila marginata (Schaus, 1912), which had previously been erroneously synonymised with Oospila permagna (Warren, 1909) by Cook and Scoble (1995). The replacement name was necessary because O. marginata (Schaus, 1912) is a junior secondary homonym of O. marginata Warren, 1897. Lindt et al. (2018) also raised O. imula from synonymy with O. miccularia Guenée, [1858] and raised O. similiplaga from synonymy with O. arpata (Schaus, 1897).

Ornithospila

Warren, 1894

D7F00D60-3B82-5822-B539-45D4CE04A831

• Ornithospila explorator Tautel, 2015 (“sp. nov.”)

Notes

One new species was described (Tautel 2015).

Orothalassodes

Holloway, 1996

FC69FB22-34AE-53F4-B1A6-6EF1EDBE4FA4

• Orothalassodes leucospilota (Moore, [1887])

• Thalassodes albomaculata Hampson, 1895

• Orothalassodes albomaculata Kirti, Goyal & Kaur, 2012

Notes

Kirti et al. (2012) published the name Orothalassodes albomaculata as a new combination for Thalassodes albomaculata. However, T. albomaculata had already been synonymised with Thalassodes leucospilota Moore by Hampson (1896), which was then transferred to Orothalassodes by Holloway (1996). Thus, the current valid name for this species is still Orothalassodes leucospilota.

Pachyodes

Guenée, [1858]

02FB8F9B-6AA6-55CC-91E8-16663A5DED85

• Pachyodes jianfengensis Han & Xue, 2008 (“sp. nov.”)

• Pachyodes novata Han & Xue, 2008 (“sp. nov.”)

Notes

Two new species were described (Han and Xue 2008).

Paramaxates

Warren, 1894

FB500918-2293-5552-9223-0C94EA9CD198

• Paramaxates fournieri Tautel, 2016 (“sp. nov.”)

• Paramaxates vagata (Walker, 1861)

• Paramaxates hainana Chu, 1981 (“syn. nov.”)

Notes

One new species was described (Tautel 2016). Paramaxates hainana was synonymised with P. vagata by Han and Xue (2011a).

Paromphacodes

Warren, 1897

FF04FAD8-1439-5E54-8D70-DF049656AE5A

• Paromphacodes alpha Lindt, Tasane, Õunap & Viidalepp, 2017 (“sp. nov.”)

• Paromphacodes alticola Lindt, Tasane, Õunap & Viidalepp, 2017 (“sp. nov.”)

• Paromphacodes onae Lindt, Tasane, Õunap & Viidalepp, 2017 (“sp. nov.”)

• Paromphacodes spina Lindt, Tasane, Õunap & Viidalepp, 2017 (“sp. nov.”)

• Paromphacodes summita Lindt, Tasane, Õunap & Viidalepp, 2017 (“sp. nov.”)

Notes

Five new species were described (Lindt et al. 2017b).

Pelagodes

Holloway, 1996

67CE21E4-8201-5BCA-97FC-F20482D03AC4

• Pelagodes bellula Han & Xue, 2011 (“sp. nov.”)

• Pelagodes cancriformis Viidalepp, Han & Lindt, 2012 (“sp. nov.”)

• Pelagodes paraveraria Han & Xue, 2011 (“sp. nov.”)

• Pelagodes simplvalvae Han & Xue, 2011 (“sp. nov.”)

• Pelagodes sinuspinae Han & Xue, 2011 (“sp. nov.”)

Notes

Five new species were described (Han and Xue 2011b, Viidalepp et al. 2012).

Prasinocyma

Warren, 1897

337CD292-D817-5E68-9287-8B40CC377A16

• Prasinocyma amharensis Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma angolica pseudopedicata Hausmann, Sciarretta & Parisi, 2016 (“ssp. nov.”)

• Prasinocyma angolica yemenicola Hausmann & Wildfeuer, 2017 (“ssp. nov.”)

• Prasinocyma angulifera Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma aquamarina Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma batesi distans Hausmann, Sciarretta & Parisi, 2016 (“ssp. nov.”)

• Prasinocyma baumgaertneri Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma beryllaria Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma bongaensis Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma camerunalta (Herbulot, 1986) (“comb. nov.”)

• Prasinocyma discipuncta Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma fallax Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma fusca Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma gemmifera Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma getachewi Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma immaculata (Thunberg, 1784)

• Prasinocyma unipuncta Warren, 1897 (“syn. nov.”)

• Prasinocyma immaculata thiaucourti Herbulot, 1993 (“stat. nov.”)

• Prasinocyma leveneorum Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma lutulenta Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma magica Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma monikae Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma nereis Townsend, 1952 (“comb. rev.”)

• Prasinocyma pedicata aethiopica Hausmann, Sciarretta & Parisi, 2016 (“ssp. nov.”)

• Prasinocyma robusta Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma saba Hausmann & Wildfeuer, 2017 (“sp. nov.”)

• Prasinocyma septentrionalis Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma shoa yabellensis Hausmann, Sciarretta & Parisi, 2016 (“ssp. nov.”)

• Prasinocyma stefani Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma trematerrai Hausmann, Sciarretta & Parisi, 2016 (“sp. nov.”)

• Prasinocyma trematerraisimienensis Hausmann, Sciarretta & Parisi, 2016 (“ssp. nov.”)

Notes

Twenty new species and six new subspecies were described (Hausmann et al. 2016, Hausmann and Wildfeuer 2017). Hausmann et al. (2016) transferred Thalassodes camerunalta Herbulot, 1986 and Eretmopus nereis (Townsend, 1952) to the genus Prasinocyma. They also synonymised P. unipuncta with P. immaculata and changed the status of P. thiaucourti Herbulot, 1993 from a species to a subspecies of P. immaculata.

Protuliocnemis

Holloway, 1996

CFDFDB2F-508D-5D3D-B548-0E1EFFC0706F

• Protuliocnemis candida Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Protuliocnemis dissimilis Han, Galsworthy & Xue, 2012 (“sp. nov.”)

• Protuliocnemis falcipennis (Yazaki, 1991) (“comb. nov.”)

Notes

Two new species were described (Han et al. 2012). Protuliocnemis falcipennis was transferred from the genus Comibaena by Han and Xue (2011a).

Pseudepisothalma

Han, 2009 (“gen. nov.”)

4DC4C94E-94BF-5F8E-8D1D-B9E8F100263C

• Pseudepisothalma ocellata (Swinhoe, 1893) (“comb. nov.”)

Notes

Pseudepisothalma is currently a monotypic genus containing only the type species, P. ocellata, which was transferred from the genus Episothalma. The new genus description and new combination were presented in Xue et al. (2009), though only the third author (Han) is credited with authorship.

Psilotagma

Warren, 1894

119C2CF0-1DE0-559B-9323-23DAB825698B

• Psilotagma pictaria (Moore, 1888) (“comb. nov.”)

Notes

Psilotagma pictaria was transferred from the genus Pachyodes by Pitkin et al. (2007).

Pyrochlora

Warren, 1895

349190BC-3B3C-5E9F-9124-1B73A2BB301D

• Pyrochlora kuklase Viidalepp, 2009 (“sp. nov.”)

• Pyrochlora motilonia Viidalepp, 2009 (“sp. nov.”)

• Pyrochlora vogli Viidalepp, 2009 (“sp. nov.”)

Notes

Three new species were described (Viidalepp 2009).

Rhanidopsis

West, 1930

EBEC8DAF-08AF-5631-B81A-6D69D42A6000

• Rhanidopsis kogeri Viidalepp & Lindt, 2010 (“sp. nov.”)

Notes

One new species was described (Viidalepp and Lindt 2010).

Rhuma

Walker, 1860

CC00412B-937B-59EE-A03D-60417CD5D7BE

• Sterictopsis Warren, 1898 (“syn. nov.”)

• Oxyphanes Turner, 1936 (“syn. nov.”)

• Rhuma argyraspis (Lower, 1893) (“comb. nov.”)

• Rhuma divergens (Goldfinch, 1929) (“comb. nov.”)

• Rhuma thiobapta (Turner, 1936)(“comb. nov.”)

Notes

Pitkin et al. (2007) designated Sterictopsis and Oxyphanes as junior synonyms of Rhuma and consequently transferred S. argyraspis (Lower, 1893), S. divergens Goldfinch, 1929 and O. thiobapta Turner, 1936 to this genus.

Tachyphyle

Butler, 1881

D2C21DFB-4AB7-5212-962E-A5F4C0B1F727

• Tachyphyle nielseni Viidalepp & Lindt, 2017 (“sp. nov.”)

• Tachyphyle selini Viidalepp & Lindt, 2017 (“sp. nov.”)

Notes

Two new species were described (Viidalepp and Lindt 2017).

Tanaorhinus

Butler, 1879

C61601D3-1139-529B-98A9-7008914E9C2C

• Tanaorhinus baruensis Orhant, 2014 (“sp. nov.”)

• Tanaorhinus guitinguensis Tautel, 2014 (“sp. nov.”)

• Tanaorhinus sultan Tautel, 2014 (“sp. nov.”)

Notes

Three new species were described (Orhant 2014, Tautel 2014).

Telotheta

Warren, 1900

BC719B1C-FD28-5550-AE1E-1070F2AF50DD

• Telotheta fresei Lindt & Viidalepp, 2014 (“sp. nov.”)

• Telotheta unoi Lindt & Viidalepp, 2014 (“sp. nov.”)

Notes

Two new species were described (Lindt and Viidalepp 2014).

Thalera

Hübner, [1823]

1DD62E31-8E9E-5D9C-902F-54B709D18EC9

• Hethemia Ferguson, 1969 (“syn. nov.”)

• Thalera pistasciaria (Guenée, 1858) (“comb. nov.”)

Notes

Hethemia sensu Ferguson was a monotypic genus, containing only the type species H. pistasciaria. Ban et al. (2018) provided morphological and molecular evidence to justify the designation of Hethemia as a junior synonym of Thalera, creating the new combination T. pistasciaria.

Thetidia

Boisduval, 1840

21C772E9-3A1A-570A-BDFB-50480AB83FEB

• Thetidia chlorophyllaria (Hedemann, 1879)

• Thetidia pekingensis (Chu, 1981) (“comb. nov.”, followed by “syn. nov.”)

Notes

The name Thetidia pekingensis (Chu, 1981) was first published in Han and Xue (2011a); it was not designated a new combination, but since Euchloris Hübner, [1823] was already known to be a synonym of Thetidia (Parsons et al. 1999), Han and Xue (2011a) were presumably transferring Euchloris pekingensis Chu, 1981 to Thetidia. This was confirmed by Han et al. (2012), who subsequently synonymised T. pekingensis with T. chlorophyllaria.

Timandromorpha

Inoue, 1944

97240A09-A473-5F9E-9F46-293795562BD8

• Timandromorpha inouei Stüning & Yazaki, 2008 (“sp. nov.”)

• Timandromorpha pinratanai Stüning & Yazaki, 2008 (“sp. nov.”)

• Timandromorpha wangi Stüning & Yazaki, 2008 (“sp. nov.”)

• Timandromorpha xuedayongi Orhant, 2013 (“sp. nov.”)

Notes

Four new species were described (Stüning and Yazaki 2008, Orhant 2013).

Vallichlora

Viidalepp & Lindt, 2019 (“gen. nov.”)

B63E05DB-3604-557D-BAB1-C480C1CF4F08

• Vallichlora rara Viidalepp & Lindt, 2019 (“sp. nov.”)

• Vallichlora selva Viidalepp & Lindt, 2019 (“sp. nov.”)

Notes

Two new species were described (Viidalepp and Lindt 2019b).

Xenozancla

Warren, 1893

0AF7B21C-0814-5F7F-9644-4BD5258B2B2B

• Yinchie Yang, 1978 (“syn. nov.”)

• Xenozancla versicolor Warren, 1893

• Yinchie zaohui Yang, 1978 (“syn. nov.”)

Notes

Han et al. (2008) synonymised Yinchie zaohui with Xenozancla versicolor. Since Y. zaohui was the type species of its genus, Yinchie was consequently designated a junior synonym of Xenozancla.

Other species affiliated with Geometrinae

Pseudobiston pinratanai

Inoue, 1994

71E2CC97-856D-5829-B86F-147CC8FD6D34

Notes

Pseudobiston pinratanai Inoue, 1994, was classified as a geometrine in Scoble and Hausmann (2007) but was recently transferred to the new family Pseudobistonidae by Rajaei et al. (2015).

Cerura melanoglypta

(Lower, 1905)

0CF84C6D-3590-5713-9D32-CAF2CC432888

Notes

One species in the Scoble and Hausmann (2007) checklist, Cerura melanoglypta (Lower, 1905), is classified as a geometrine (Ollerenshaw 2012), but has never formally been transferred from the notodontid genus Cerura Schrank, 1802. We agree that this species should eventually be assigned to a genus in Geometrinae, but it is technically not in Geometrinae at this time.

Discussion

In summation, nine new genera, 128 new species and ten new subspecies of emerald moths have been described since the publication of Scoble and Hausmann (2007), along with over 80 new genus- and species-group changes within subfamily Geometrinae. Since 2007, the known species richness of Geometrinae has increased by ~4.5%, from 2,529 species (Scoble and Hausmann 2007) to 2,643 species.

Supplementary Material

Supplementary material 1

List of the Geometrinae species of the world

David Plotkin

Data type

Taxonomical checklist

Brief description

This table provides a list of all new species descriptions, combinations and other taxonomic changes in the subfamily Geometrinae (Lepidoptera: Geometridae) since 2007 (Sheet 1: "Taxonomic changes since 2007"). This table also provides a list of the 2,643 current valid species names in the subfamily Geometrinae (Lepidoptera: Geometridae), with their authorship and year of description (Sheet 2: "All current Geometrinae species").

File: oo_400377.xls