Abstract
A detailed catalogue of the type material of the Scarabaeoidea housed in the Research Institute of Evolutionary Biology, Tokyo, Japan is listed. The catalogue includes the data of the type material of four families and 111 species.
Keywords: Allotype, Geotrupidae , holotype, Lucanidae , neotype, paratype, Scarabaeidae , Trogidae
Introduction
In this report we present a catalogue of the type material of Scarabaeoidea housed in the Research Institute of Evolutionary Biology, Tokyo, Japan (RIEB). The beginning of the RIEB collection dates back to 1950, when the first natural objects were deposited at the Research Institute of Thremmatology of the Tokyo University of Agriculture (RIT) established that year. RIT was transformed into an incorporated foundation in 1974 and renamed Research Institute of Evolutionary Biology (Yamamoto 2000). The Lepidoptera collection in the RIEB was listed in detail by Aoki et al. (2005, 2008) and Yamaguchi et al. (2006, 2008, 2009, 2011, 2017), but the Coleoptera collection has never been catalogued. This creates a problem because well-organized museum collections and published type specimen catalogues are very important to facilitate taxonomic and systematic investigations of animals.
The majority of the types of Scarabaeidae deposited in the RIEB originate from the collection of Mr Yoshikazu Miyake (1926–2003), a well-known Japanese amateur specialist on Scarabaeidae, who was also the author of those taxa. A part of types of Miyake’s species is also housed in the National Museum of Nature and Science, Tokyo (NMNS) and the Nagaoka Municipal Science Museum, Niigata (NMSM). Unfortunately, some of his types are missing.
In addition, other researchers that contributed to the Scarabaeoidea type collection in RIEB are: Dirk Ahrens, Aleš Bezděk, Toshiyuki Ichikawa, Takeshi Itoh (Takeshi Matsumoto), Kimio Masumoto, Sizumu Nomura, Teruo Ochi, and Kaoru Wada.
Materials and methods
The scientific names of higher taxa in this catalogue follow Scholtz and Grebennikov (2016). The structure of each entry is as follows:
– original combination of the taxon name;
– original combination and spelling of the taxon name, followed by the author, year of description, and pagination;
– type material, number of specimens (including sex, if known), and exact label data. A double slash ‘//’ indicates separate labels and single slash ‘/’ indicates lines within each label. The words in Japanese or Chinese were transcribed into Roman alphabet. Paratypes with discrepancies between collection data on the label and data quoted in the original description are indicated by ‘ [[]]’;
– remarks on types condition (given only for holotypes and neotypes);
– current taxonomic status;
– remarks, if any.
The majority of holotypes and neotypes were photographed with a Nikon D7200 camera using a Nikon AF-S DX Micro NIKKOR 40 mm f/2.8 G lens, and some types were photographed with a KEYENCE VHX-1000 Digital Microscope.
Catalogue
Family Geotrupidae
Subfamily Geotrupinae
Genus Sinogeotrupes
Sinogeotrupes taiwanus
Miyake & Yamaya
966AF9F9-8E1F-5A78-BF35-005255BDABBB
Sinogeotrupes taiwanus Miyake & Yamaya, 1995: 32−34.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
3 exs.: 1 ex. ‘Wusha / –VI–1941 / R. Boloudo // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘Wu sha / Formosa / 21. Viii. 1941 / Col. Bumua Tin // Geotrupes / subshiatelluo FAIRMAIR / DET. Y. MIYAKE // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘台湾= taiwan 台中= taichû / 関刀山= guandaoshan / 1993. V. 31 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’.
Current status.
Phelotrupes (Sinogeotrupes) taiwanus (Miyake and Yamaya, 1995), see Nikolajev et al. (2016).
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 3 exs. ‘Tattaka / Formosa / 10. Vi. 1965 / T. Shirozu // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘(FORMOSA) / Mt Rata / Vi–Vii–1972 / L. F, Hang // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ex. ‘(FORMOSA) / Mt Rata / –Vii–1972 / L. F, Hang // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 2 exs. ‘ [Taiwan] / 台中= taichû 関刀山= guandaoshan / 3–VI–1993 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 1 ♂ ‘ [Taiwan] / 台中= taichû 関刀山= guandaoshan / 3–VI–1993 // Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’. 3 exs. ‘Paratype / Sinogeotrupes taiwanus / Y. MIYAKE et YAMAYA, 1995’.
Family Lucanidae
Subfamily Lucaninae
Genus Aegus
Aegus laevicollis nakanei
Ichikawa & Imanishi
21CE787E-F0F2-55A9-AC6C-39D33669E543
Aegus laevicollis nakanei Ichikawa & Imanishi, 1976: 9.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Kusui):
Paratype.
1 ex.: [[1 ♀ ‘Mt. Yonahadake, / Kunigami-son, / Is. Okinawa-Hontō, / 28. VI. 1972 (the date given in the original description is 16–17. VI. 1970), / HEIKICHI IRIE leg., // KUSUI // Paratype’]].
Current status.
Aegus nakanei nakanei Ichikawa and Imanishi, 1976, see Huang and Chen (2017).
Family Scarabaeidae
Subfamily Aphodiinae
Genus Aphodius
Aphodius (Agoliinus) tanakai
Masumoto
CF4CEE41-C8E6-5494-8E87-B13695D5B206
Aphodius (Agoliinus) tanakai Masumoto, 1981: 73−74.
Note.
The following paratype is deposited in RIEB (ex coll. K. Masumoto):
Paratype.
1 ex.: ‘Nikko / Date: 23 V 1981 / K. MASUMOTO leg. // Paratype / Aphodius (Agr) / tanakai MASUMOTO’.
Current status.
Agoliinus tanakai (Masumoto, 1981), see Dellacasa et al. (2016).
Aphodius (Phaeaphodius) himalorectus
Ahrens & Stebnicka
3D25AD27-F087-541C-BBF8-7D063B0A2768
Aphodius (Phaeaphodius) himalorectus Ahrens & Stebnicka, 1997: 12−13.
Note.
The following paratypes are deposited in RIEB (ex coll. D. Ahrens):
Paratypes.
2 exs.: ‘Phalante 20. 3. / unt. Nebelwaldstufe / 2100–2300 m // PARATYPUS / Aphodius (Phaeaphodius) himalorectus sp. n. / det. D. AHRENS 1994’.
Current status.
Phaeaphodius himalorectus (Ahrens and Stebnicka, 1997), see Dellacasa et al. (2016).
Aphodius (Sinodiapterna) zeni
Ochi
D196A45C-3FF4-5BDC-93F4-9C9ACFD97AE1
Aphodius (Sinodiapterna) zeni Ochi, 1991: 53−55.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: ‘SUNGKAN / C. TAIWAN / 29–V. 1987 / K. LAH // PARATYPE / Aphodius (Sinodiapterna) / zeni / OCHI 1991’.
Current status.
Sinodiapterna zeni (Ochi, 1991), see Dellacasa et al. (2016).
Genus Rhyparus
Rhyparus kitanoi
Miyake
77DC2BCD-51B6-5F0F-B2B5-CFCED9337CD1
Figure 1.
Habitus of holotype specimens. ARhyparus kitanoi (Miyake) BAdoretus (Lepadoretus) ubonensis (Miyake, Yamaguchi et Aoki) CAnomala flavoguttata Miyake DAnomala ohmomoi Miyake, Yamaguchi et Aoki EAnomala thailandiana Miyake, Yamaguchi et Akiyama FCallistethus isidai Miyake GMalaia castanoptera Miyake HMalaia macassara Miyake.
Rhyparus kitanoi Miyake, 1982: 65−67.
Note.
The holotype and following specimen are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘C, Sata /〔KYUSHU〕/ Japan / 25, VI 1971 / T. Kitano leg. // PARATYPE // Holotype: / Rhyparus / kitanoi / Y. MIYAKE, 1992’. (Fig. 1A)
Type condition.
The right and left protarsus and the right antenna of the holotype are missing.
Current status.
Rhyparus kitanoi kitanoi Miyake, 1982, see Ochi (2001).
Remark.
In addition to the holotype mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ex.: ‘Sesonoura / Is. Shimokoshiki / Kagoshima-pref. / 18. VI. 1982 / S. Imasaka leg. // Paratype: Rhyparus / kitanoi / Y. MIYAKE, 1992’.
Subfamily Scarabaeinae
Genus Caccobius
Caccobius bawangensis
Ochi, Kon & Kikuta
F7535001-A039-5B7D-B745-B4A47296800D
Caccobius bawangensis Ochi, Kon & Kikuta, 1997: 262−264.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: ‘Mt. Bawang / Sarawak S. / VIII. 1991 / M. Kon Coll. // PARATYPE / Caccobius / bawangensis OCHI, KON and KIKUTA’.
Current status.
Valid species.
Genus Copris
Copris (Microcopris) mindorensis
Ochi
6DF1E607-BEF2-52B9-94CA-57E10473BA39
Copris (Microcopris) mindorensis Ochi, 1992: 13.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: ‘MT. HALKON / IS. MINDRO / 28–XII. 1988 / D. MOHAGAN // PARATYPE / Copris / (Microcopris) / mindorensis / OCHI, 1992.
Current status.
Valid species.
Genus Onthophagus
Onthophagus setchan
Masumoto
F4B7452C-5C3F-551A-93E0-4CA97A3BADF9
Onthophagus setchan Masumoto, 1984: 80−81.
Note.
The following paratype is deposited in RIEB (ex coll. K. Masumoto):
Paratype.
1 ex.: ‘FENGCHIFU / FORMOSA / 7–VII. 74. / T. OCHI // PARATYPE / Onthophagus / setchan / MASUMOTO (1984)’.
Current status.
Valid species.
Onthophagus (Parascatonomus) acuticollissakishimanus
Nomura
71EEA96C-0871-54B2-8640-A8C6227EBA91
Onthophagus (Parascatonomus) acuticollis sakishimanus Nomura, 1976a: 29.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
6 exs.: 3 ♂ ‘Kanpira / Iriomote Is. / Yaeyama Iss. / 12. VII. 1975 / R. YANO leg. // ♂ // Onthophagus (P.) / acuticollis / sakishimanus / NOMURA (1976)’. 3 ♀ ‘Kanpira / Iriomote Is. / Yaeyama Iss. / 12. VII. 1975 / R. YANO leg. // ♀ // Onthophagus (P.) / acuticollis / sakishimanus / NOMURA (1976)’.
Current status.
Parascatonomus (Necramator) acuticollis sakishimanus (Nomura, 1976), see Ochi and Kon (2017).
Onthophagus (Parascatonomus) murasakianus
Nomura
B86813CD-6881-58EB-A9F2-4A6A0912486F
Onthophagus (Parascatonomus) murasakianus Nomura, 1976a: 30.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratype.
1 ex.: ‘Kanpira, / Iriomote Is. / Yaeyama Iss. / 12. VII 1975 / R. YANO leg. // PARATYPE / Onthophagus (P.) / murasakianus / NOMURA (1976)’.
Current status.
Parascatonomus (Necramator) murasakianus (Nomura, 1976), see Ochi and Kon (2017).
Onthophagus (Parascatonomus) takedai
Ochi & Araya
7562CF62-422A-5F5B-BD6B-0FC6CDFCC2EA
Onthophagus (Parascatonomus) takedai Ochi & Araya, 1992: 99−100.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: ‘QUESON. N. P / IS. LUZON / 10–VIII. 1981 / K. SUGINO // PARATYPE / Onthophagus / (Parascatonomus) / takedai / OCHI et ARAYA, 1992’.
Current status.
Parascatonomus (Granulidorsum) takedai (Ochi et Araya, 1992), see Ochi and Kon (2017).
Onthophagus (Proagoderus) watanabei
Ochi & Kon
E282C728-2248-58F9-9300-AB84DB9B23F1
Onthophagus (Proagoderus) watanabei Ochi & Kon, 2002: 306−311.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: ‘SEPIROK / N. BORNEO / 4–VII. 1985 / M. KON // PARATYPE / Onthophagus / (Proagoderus) / watanabei / OCHI and KON. 2002’.
Current status.
Proagoderus watanabei (Ochi et Kon, 2002), see Philips (2016).
Onthophagus (Strandius) hiurai
Ochi
0955D712-4CC3-5698-B6B9-49F9285CFC80
Onthophagus (Strandius) hiurai Ochi, 1984: 63−65.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: ‘PILU / Taiwan / 21, VIII, 1979 / K. Sugino leg. // PARATYPE / Onthophagus (St) / hiurai / OCHI (1984)’.
Current status.
Onthophagus (Phanaeomorphus) potanini hiurai Ochi, 1984, see Ziani and Bezděk (2016).
Subfamily Rutelinae
Genus Adoretus
Adoretus (Lepadoretus) ubonensis
Miyake, Yamaguchi & Aoki
6E95AC70-A412-52F8-876E-2E0E31040CDF
Adoretus (Lepadoretus) ubonensis Miyake, Yamaguchi, Aoki & Akiyama, 2002: 62−63.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Shirinthorn (light) / Ubon Ratchatani / E–THAILAND / 27. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // No. 1 / ♂ // Holotype: / Adoretus / ubonensis // Holotype / Adoretus (Lepadoretus) ubonensis / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig. 1B)
Type condition.
The left mesotarsus and the left metaleg of the holotype are missing.
Current status.
Junior subjective synonym of Adoretus compressus (Weber, 1801), see Kobayashi (2018).
Genus Anomala
Anomala flavoguttata
Miyake
F5A76A0A-A44E-57BA-BB1C-1DEA9E3417AA
Anomala flavoguttata Miyake, 2000: 108−109.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Sapa=Chapa / N. Vietnam / 20. V. 1994 / N. KATSURA // Holotype: Anomala / flavoguttata / Y. MIYAKE, 2000’. (Fig. 1C)
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Valid species.
Anomala ohmomoi
Miyake, Yamaguchi & Aoki
41E2EE96-E344-5136-98EE-92FE6B007ABC
Anomala ohmomoi Miyake, Yamaguchi, Aoki & Akiyama, 2002: 58−60.
Note.
The holotype and the following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Ban Lao Kram / Mukdahan (light) / E–THAILAND沢= sawa / 8. Mar. 1999 / S. Yamaguchi and / T. Aoki leg. // Holo Type: / Anomala / ohmomoi // Holotype / Anomala ohmomoi / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig. 1D)
Paratypes.
4 exs.: ‘Ban Lao Kram / Mukdahan (light) / E–THAILAND沢= sawa / 8. Mar. 1999 / S. Yamaguchi and / T. Aoki leg. // Paratype / Anomala ohmomoi / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’.
Current status.
Valid species.
Anomala thailandiana
Miyake, Yamaguchi & Akiyama
86A394A0-0AAB-5621-B49F-B06FAEC8625E
Anomala thailandiana Miyake, Yamaguchi, Aoki & Akiyama, 2002: 60−61.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Don Far Huan (light) / Ubon Ratchathani / E–THAILAND / 17. Sep. 1999 / S. Yamaguchi and / M. Akiyama leg. [white label, front] // Apigoni / 5 [white label, back] // Holotype / Anomala thailandiana / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig. 1E)
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Valid species.
Genus Callistethus
Callistethus isidai
Miyake
88B1C0B4-7A2D-5135-BD6E-3B847A7ADC06
Callistethus isidai Miyake, 1987: 5−6.
Note.
The holotype and the following paratype are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘ [Taiwan] / Lishan to Pilu / 31–VII–1972 / Y. Miyake leg. // Holotype: / Callistethus isidai / Y. MYAKE, 1987’. (Fig. 1F)
Paratype.
1 ex.: ‘ [Taiwan] / Lishan to Pilu / 31–VII–1972 / Y. Miyake leg. // Paratype / Callistethus isidia / Y. MIYAKE, 1987’.
Type condition.
The right and left metatarsi of the holotype are missing.
Current status.
Callistethus plagiicollis isidai Miyake, 1987, see Zorn and Bezděk (2016).
Remark.
The habitus photograph in the original description does not agree with the holotype. This is also the case with some other species described by Miyake. Apparently, he did not intend to provide the photographs of the holotypes. Also, as noted below, we were unable to trace some specimens illustrated in the original descriptions.
Genus Kibakoganea
Kibakoganea ohtanii
Miyake & Muramoto
6EF2AFCD-F5E1-5217-B09E-B7C8AEC0FB2E
Kibakoganea ohtanii Miyake & Muramoto, 1992: 22−24.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
2 exs.: [[1 ♂, 1 ♀ ‘Tam Dao / N. VIETNAM / 28–30, IV, 1991 (the date given in the original description is 28–30. III. 1991) / M. Fujioka and / R. Muramoto – leg. // PARATYPE / Kibakoganea / ohtanii / Y. MIYAKE et / MURAMOTO, 1992.’]].
Current status.
Junior subjective synonym of Kibakoganea sexmaculata (Kraatz, 1900), see Miyake (2003).
Genus Malaia
Malaia castanoptera
Miyake
8DD2B02D-0D3F-50C6-AE1B-E6F73F1FEF26
Malaia castanoptera Miyake, 2000: 112.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Holotype: Malaia / castanoptera / Y. MIYAKE, 2000’. (Fig. 1G)
Paratypes.
7 exs.: 1 ♂, 3 ♀ ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’. 2 exs. ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’. 1 ex. ‘malino / 2603.11.21 // Paratype: Malaia / castanoptera / Y. MIYAKE, 2000’.
Current status.
Valid species.
Remark.
The habitus photo in the original description belongs to the female paratype with the label ‘Malino, Celebes / Indonesia / 21 Nov 1944 // malino / 2603.11.21’.
Malaia cyanea
Miyake
367CEDFC-08CF-55EA-8F71-404677E01B2A
Malaia cyanea Miyake, 1996: 38−39.
Note.
The following paratypes are deposited is RIEB (ex coll. Y. Miyake):
Paratypes.
2 exs.: 1 ♂ ‘32 km from / Palopo / Surawesi / 10. VI. 1982 // Paratype: / Malaia cyanea / Y. MIYAKE, 1996’. 1 ♀ ‘Punchak–Lombanan / Mamasa, / S. SULAWESI / 1991. AUG. 10. / N. Kashiwai–leg. // Paratype: / Malaia cyanea / Y. MIYAKE, 1996’.
Current status.
Valid species.
Malaia macassara
Miyake
267BC972-6F84-5456-B861-E914FFBDC5CA
Malaia macassara Miyake, 2000: 109−110.
Note.
The holotype and the following paratype are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Lampbatter / Sulawesi / 23–25. VIII. 1999 / M. Takai // Holotype: Malaia / macassara / Y. MIYAKE, 2000’. (Fig. 1H)
Paratype.
1ex.: 1 ♂ ‘Lampbatter / Sulawesi / 23–25. XII. 1999 / M. Takai leg. // Paratype: Malaia / macassara / Y. MIYAKE, 2000’.
Type condition.
The left protarsus and metatarsus of the holotype are missing.
Current status.
Valid species.
Malaia rufofemorata
Miyake
B699423B-6F15-5832-B723-4FB660923D7E
Malaia rufofemorata Miyake, 1996: 40−41.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
4 exs.: 1 ♂, 1 ♀ ‘Puncak, Palopo / VI–VII, 1989 / Central SULAWESI // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’. 2 ♂ ‘Palu Palolo / VIII. 1990 / Central SULAWESI // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’.
Current status.
Valid species.
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 2 ♂, 1 ♀ ‘Palolo, Palu / 01–10/XI. 1990 / native coll. / CELEBES // Paratype: Malaia / rufofemorata / Y. MIYAKE, 1996’.
Malaia sulawesi
Miyake
37ADCBA8-9015-51C9-857F-6E8E77A08B24
Malaia sulawesi Miyake, 1996: 41−42.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
58 exs.: 1 ♂ ‘Mt. Pedamaran / Tana Toraja / 1985. III–17 / South SULAWESI / Soma, K. leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. 1 ex. ‘Puncak Palopo / 1989. III. 31 / Centr. CELEBES // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. 3 exs. ‘Palu Pulu / C. Sulawesi / II. 1989 // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’. [[44 exs. ‘Mt. Pedamaran / S. Sulawesi (real data is C. Sulawesi) / 20. X. 1983 (real date is 20. V. 1983) / K. Soma leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’]]. [[9 exs. ‘Mt. Pedamaran / Tana Toraja / 1983. X–20 (real date is 1983. V–20) / South SULAWESI (real data is Central SULAWESI) / Soma, K. leg. // Paratype: / Malaia / sulawesi / Y. MIYAKE, 1996’]].
Current status.
Valid species.
Malaia taoi
Miyake
EDCF8BFC-B447-5C78-880E-BBF9296C0B14
Malaia taoi Miyake, 1996: 39−40.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
4 exs.: 1 ♂ ‘37 km from / Palopo C.- / Sulawesi / 9. IV. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♀ ‘32 km from / Palopo / Surawesi / 10. VI. 1982 // 37 km from / Palopo C.- / Sulawesi / 10. VI. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♂ ‘37 km from / Palopo C.- / Sulawesi / 11. IV. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’. 1 ♀ ‘32 km from / Palopo / Surawesi / 13. VI. 1982 // 37 km from / Palopo C.- / Sulawesi / 13. VI. 1982 // Paratype: / Malaia / taoi / Y. MIYAKE, 1996’.
Current status.
Valid species.
Malaia tondanoensis
Miyake
768FB042-6157-5BEC-A61B-50BC5A907CCF
Malaia tondanoensis Miyake, 1996: 42−43.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
4exs.: ‘Tondano / N. Sulawesi / 10–IV–1989 / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’.
[[1 ex. ‘Tangkoko N. / N. Sulawesi / 10–III–1989 (the date given in the original description is 10–IV–1989) / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’]]. [[2 exs. ‘Tondano / N. Sulawesi / 9–IV–1989 (the date given in the original description is 10–IV–1989) / Y. Miyake leg. // Paratype: / Malaia / tondanoensis / Y. MIYAKE, 1996’]].
Current status.
Valid species.
Malaia toraja
Miyake
788C21F5-7F76-51EC-96D9-E8C359BBC754
Figure 2.
Habitus of holotype and neotype specimens. AMalaia toraja Miyake BMimela kitanoi Miyake CMimela marginipennis Miyake DSpinanomala moritai Miyake EPeltonotus morio Burmeister FPeltonotus morio sawaii Miyake GApogonia ohmomoi Miyake HApogonia ovata Miyake et Yamaguchi.
Malaia toraja Miyake, 2000: 110−111.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. / South SULAWESI // Holotype: Malaia / toraja / Y. MIYAKE, 2000’. (Fig. 2A)
Paratypes.
27 exs.: 1 ♂ ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 9 exs. ‘Mt. Pedamaran / Tana Traja / 20–X. 1983 / K. SOHMA lgd. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Mt. Pedamaran / S. Sulawesi / 20. X. 1983 / K. Soma leg. // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.8 exs. ‘Puncak, Palopo / VI–VII, 1989 / Central SULAWESI // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Central SULAWESI / Puncak, Palopo / VI–VII, 1989 // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ♂ ‘Puncak Palopo / 1989. / Centr. CELEBES // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 4 exs. ‘Puncak Palopo / 1989. / Centr. CELEBES // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.
Current status.
Valid species.
Remark.
The habitus photo in the original description belongs to the female paratype with the label ‘Mt. Pedamaran / Tana Toraja / S. Sulawesi / 7, VIII, 1988 / K. Soma leg.’.
In addition to the paratypes listed above, the following specimens labeled as the paratypes are not designated in the original description: 1 ♀ ‘Mt. Pedamaran / Tana Toraja / S. Sulawesi / 7, VIII, 1988 / K. Soma leg. // SI [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ex. ‘(Pulu Pulu) / C. Sulawesi / INDONESIA / Dec. 1988 // [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 2 exs. ‘Puncak, Palopo / Centr. CELEBES / 1989. I / INDONESIA // [yellow label] // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 1 ex. ‘Puncak Palopo / 1989. III. 31 / Centr. CELEBES // Paratype: Malaia / toraja / Y. MIYAKE, 2000’. 3 exs. ‘Palu, Palolo / VII 1989 / Central SULAWESI // Paratype: Malaia / toraja / Y. MIYAKE, 2000’.
Genus Mimela
Mimela kitanoi
Miyake
BD497983-A9A9-533E-A178-6531EAFC9751
Mimela kitanoi Miyake, 1987: 7−8.
Note.
The holotype is deposited in RIEB (ex coll. K. Wada):
Holotype
(♀). ‘Larashan / Hsinchu Hs. / VI. 1975, / Fang leg // Holotype: / Mimela kitanoi / Y. MIYAKE, 1987 // Kaoru WADA / Collection 2001’. (Fig. 2B)
Current status.
Junior subjective synonym of Mimela flavocincta Lin, 1966, see Zorn and Bezděk (2016).
Mimela marginipennis
Miyake
16DCDA55-40C3-593D-96D1-6DF7CA6F3756
Mimela marginipennis Miyake, 1994: 151.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘[Borneo] / Crocker Range / 1800−1400m alt. / nr. Keningau C. / X. 1988. M. ITOI leg. // Holotype: / Mimela / marginipennis / Y. MIYAKE, 1994’. (Fig. 2C)
Type condition.
The aedeagus of the holotype is pinned separately, and the left mesotarsus is missing.
Current status.
Junior subjective synonym of Mimela margarita Arrow, 1910, see Wada (2001).
Mimela sawaii
Miyake
BD07E97C-6368-5B68-9294-1E00504CA31F
Mimela sawaii Miyake, 1994: 149−150.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: 1 ♂ ‘Crocker Range / (1000–1400 m) / Nr. Keningau C. / N. Borneo, – / 15–V–1989 // [blue label] // Paratype: / Mimela / sawaii / Y. MIYAKE, 1994’.
Current status.
Junior subjective synonym of Mimela pallidicauda Arrow, 1910, see Wada (2001).
Mimela suspecta
Miyake
4DBBC7A8-EAF4-5A58-9AB7-D6790A0E5BD7
Mimela suspecta Miyake, 1994: 150.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratype.
2ex.: 1 ♂ ‘Mt. Trus Madi / (SW Slopa 1200m alt.) / Sabah, MALAYSIA / 14–IV–1993 / Minoru SAWAI leg. // Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’. [[1 ex. ‘Mt. Trus Madi / (SW Slopa 1200m alt.) / Sabah, MALAYSIA / 18–IV–1993 (the date given in the original description is 4–IV–1993) / Minoru SAWAI leg. // [yellow label] // Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’]].
Current status.
Valid species.
Remark.
In addition to the paratypes mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ♂ ‘Trus Madi Mts. / near Keningau / Saba MALAYSIA / 1993. iv. 17–30. // F //Paratype: / Mimela / suspecta / Y. MIYAKE, 1994’.
Genus Popillia
Popillia iwasei
Miyake
96642894-B568-5A84-BDC0-4193ADE24E3C
Popillia iwasei Miyake, 1996: 43−44.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
6 exs.: 2 ♀ ‘Puncak, Palopo / Centr. CELEBES / 1989. I / INDONESIA // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. 2 ♀ ‘Puncak Palopo / C. Sulawesi / 1. 1989 // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. 1 ♀ ‘Puncak Palopo / C. Sulawesi / 1989. V / INDONESIA // SI [yellow label] // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’. [[1 ♂ ‘Palolo (the locality given in the original description is Palopo) / II. 1991 (the date given in the original description and Telnov and Zorn (2019) are I. 1989) // Paratype: / Popillia / iwasei / Y. MIYAKE, 1996’]].
Current status.
Valid species.
Genus Spilopopillia
Spilopopillia rugosa
Miyake
5E5C8BEF-A9FA-55CA-8CDE-92960A75C283
Spilopopillia rugosa Miyake, 1989a: 180−181.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
18 exs.: [[1 ex. ‘ [Malaya] / Fraser’s Hill / 29–III–1977 (the date given in the original description is 28. III. 1977) / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]]. [[1 ex. ‘ [Malaya] / Freaser’s Hill (the locality given in the original description is Fraser’s Hill) / 30–III–1977 (the date given in the original description is 28. III. 1977) / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]]. 6 exs. ‘ [Malaya] / Tanah Rata / 30–III–1974 / Y, Miyake leg // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 9 exs. ‘TANARATA / MALAYA / 30 III 1974 / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. [[1 ex. ‘Gentig / Hiland (the locality given in the original description is Genting Highland) MALAYA / 5. IV. 1974 (the date given in the original description is 5. IV. 1975) / Y Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’]].
Current status.
Valid species.
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 8 exs. ‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 5 exs. ‘TANARATA / MALAYA / 28, 29, 30 III 1974 / Y. Miyake leg. // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 1 ex. ‘84. IV // Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’. 2 exs. ‘Paratype: / Spilopopillia / rugosa / Y. MIYAKE, 1989’.
Genus Spinanomala
Spinanomala moritai
Miyake
7D0865EF-5AB8-5182-A8D6-36332D5BEA2A
Spinanomala moritai Miyake, 1996: 38.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Tanah Rata / MALAYA / IV. 1984 / M. Hata // Holotype: / Spinanomala / moritai / Y. MIYAKE, 1996’. (Fig. 2D)
Paratypes.
11 exs.: 1 ex. ‘Tanah Rata / MALAYA / I–II. 1973 / MORITA, Kozo // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’. 2 ♂, 3 ♀ ‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’. 3 ♂, 2 ♀ ‘Tanah Rata / MALAYA / IV. 1984 / M. Hata // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’.
Type condition.
The aedeagus of the holotype is pinned separately and the right metaleg is missing.
Current status.
Valid species.
Remark.
The collecting data of the specimen labelled as the holotype do not match the original description. There are two male paratypes (‘Tanah Rata / MALAYA / 1. III. 1977 / Y. Miyake // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’) that match the holotype data.
In addition to the paratypes mentioned above, the following specimen labeled as the paratype is not designated in the original description: 1 ex. ‘フレーザーヒル= furêzâhiru / MALAYA / 4 – 6. IV. 1977 / M. Hata // Paratype: Spinanomala / moritai / Y. MIYAKE, 1996’.
Subfamily Dynastinae
Genus Papuana
Papuana timorensis
Miyake & Yamaya
82876068-C811-5498-A2F0-01E96156A7AF
Papuana timorensis Miyake & Yamaya, 1999: 103−104.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: 1 ♂ ‘Timor Is. / Indonesia / V. 1993 / Native Coll. // Timor / Is. [aedeagus mount] // Paratype: / Papuana / timorensis YAMAYA / Y. MIYAKE, 1999’.
Current status.
Valid species.
Genus Peltonotus
Peltonotus morio
Burmeister
AB63F9A4-2EC0-516F-B5D8-0A53585422F7
Peltonotus morio Burmeister, 1847: 75.
Note.
The neotype is deposited in RIEB (ex coll. K. Wada):
Neotype
(♂). ‘NEPAL・HIMALAYA / 10. 5. 1993 Chhachok Madi– / Khola/ Gandaki (Kaski– / Destr.) 1300m lg: D. Ahrens //Peltonotus / morio Burn. / det: D. Ahrens 1997 // Neotype / Peltonotus / morio Burmeister, 1847 / Jameson and Wada, 2004’. (Fig. 2E)
Type condition.
The aedeagus of the neotype is pinned separately.
Current status.
Valid species.
Remark.
The neotype was designated by Jameson and Wada (2004).
Peltonotus morio sawaii
Miyake
8A162BF8-C97D-56C7-A6DB-BAC71CAA3502
Peltonotus morio sawaii Miyake, 2000: 112−113.
Note.
The holotype and the following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Doi Suthep / Chieng Mai / N–THAILAND/ 22. V. 1983 / leg. M. Sawai // 12 // P. morio / Burmi. [white label, front] // Cameron / Highland / Pahang [white label, back] // Holotype: Peltonotus / moriosawaii / Y. MIYAKE, 2000’. (Fig. 2F)
Paratypes.
4 exs.: 2 ♂, 1 ♀ ‘Doi Suthep / Chieng Mai / N–THAILAND / 22. V. 1983 / leg. M. Sawai // Paratype: Peltonotus / moriosawaii / Y. MIYAKE, 2000 // P. morio /Burmeister’. 1 ♀ ‘Doi Suthep / Chieng Mai / N–THAILAND / 22. V. 1983 / leg. M. Sawai // Paratype: Peltonotus / moriosawaii / Y. MIYAKE, 2000’.
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Junior subjective synonym of Peltonotus morio Burmeister, 1847, see Krell and Bezděk (2016).
Peltonotus podocrassus
Jameson & Wada
904A01CA-2B9F-5120-B322-450EBD040D07
Peltonotus podocrassus Jameson & Wada, 2004: 34−37.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
7 exs.: 2 exs. ‘ [Malaya] / Tanah Rata / 30–III–1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘39 miles from / Tapah, Malaya / 30–III–1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘TANARATA / MALAYA / 30. III. 1976 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ♂ ‘V. R〔C. H.〕/ Malaya / 21. III. 1978 // 46 // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ex. [Malaya] / Tanah Rata / 1–IV–1984 / Y. Miyake leg. // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’. 1 ex. ‘TANARATA / MALAYA / 1. IV. 1984 / Y. Miyake // Paratype: Peltonotus peninsularis / Y. MIYAKE, 2000 // Paratype / Peltonotus podocrassus / Jameson and Wada, 2004’.
Current status.
Valid species.
Peltonotus sakaii
Miyake & Yamaya
AC24DDB8-DCB4-55C4-8285-056BEC4895A4
Peltonotus sakaii Miyake & Yamaya, 1994: 39−40, 42.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: ‘Mt. Serapi / SARAWAKU / 1991/III. // 20 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’.
Current status.
Junior subjective synonym of Peltonotus similis Arrow, 1931, see Krajcik (2012).
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ♂ ‘Mt. Serapi / Kuching, Sarawak / IX. 1990 / BORNEO // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 1 ex. ‘Lembah Anai / W. Sumatra / VI. 1990 / INDONESIA // [yellow label] // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 1 ♂ ‘Kimanis Road / Near Keningau / N. Borneo / 3–V–1994 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’. 2 exs. ‘Kimanis Road / Near Keningau / N. Borneo / 3–V–1994 // Paratype: / Peltonotus sakaii / Y. MIYAKE et YAMAYA, 1994 // Peltonotus / similis Arrow, 1931 / det. Kaoru Wada, 2004’.
Peltonotus sulawesiensis
Wada
119EB035-A2BA-579E-A9CF-52B909ADDDC9
Peltonotus sulawesiensis Wada, 1990: 1−2.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
4 exs.: ‘Tondano N. Celebes / INDONESIA / V 1988 / leg. N. Nisikawa // PARATYPE / Peltonotus sulawesiensis / Kaoru WADA, 1990’.
Current status.
Junior subjective synonym of Neohyphus celebesus Heller, 1896, see Kuijten (1994).
Subfamily Melolonthinae
Genus Amiserica
Amiserica rufidula
Nomura
A33D6B93-3142-5882-A021-B0E4A64F3B43
Amiserica rufidula Nomura, 1974: 84.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
2 exs.: 1 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // PARATYPE / Amiserica / rufidula / NOMURA (1974)’. 1 ♂ ‘Li-shan / Taiwan / 1. VIII. 1974 / Y. Miyake // ♂ // PARATYPE / Amiserica / rufidula / NOMURA (1974)’.
Current status.
Valid species.
Genus Apogonia
Apogonia fujiokai
Miyake & Yamaya
E96EFD87-9EE6-5DA3-BFF4-D582BBE73B5B
Apogonia fujiokai Miyake & Yamaya, 1997: 6−7.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: ‘Flores Is. / Indonesia / May. 1995 // Y // Paratype: / Apogonia fujiokai / Y. MIYAKE et YAMAYA, 1997’.
Current status.
Valid species.
Apogonia fusciventris
Miyake & Yamaya
C146E0CC-97AF-540E-852D-EEFA391B54EC
Apogonia fusciventris Miyake & Yamaya, 1997: 5−6.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: ‘Vangvieng / P. Vietiane / LAO. P. D. R. / 25–26. MAY. 1994. / K. Miura–leg. // Y // Paratype: Apogonia / fuscoventris / Y. MIYAKE et YAMAYA, 1997’.
Current status.
Junior subjective synonym of of Apogonia striatipennis Frey, 1971, see Kobayashi (2010).
Remark.
The species name on the paratype label is ‘fuscoventris’, which is invalid. Probably, the name was confused with fusciventris by Y. Miyake.
Apogonia hongkongica
Miyake
05D72C6E-19CC-5739-B9EA-0DFAAD654538
Apogonia hongkongica Miyake, 1989b: 38−39.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
11 exs.: 1 ♂ ‘Hong Kong / China / C. K. Yu / 1974. V. [white label, front] // Native Collector / legt. [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♂ ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / Native / Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♂ ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Native Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘♀ // Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / Native / Collector // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘♀ // Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Native / Collector / legt. // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘Hong Kong / 1974 / From C. K. Yu ♀ // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ex. ‘Hong Kong / China / V. 1974 / C. K. Yu [white label, front] // Legt. / native / Collector [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 2 exs. ‘Hang Kong / China / V. 1974 / C. K. Yu Col [white label, front] // Native / Collector / legt. // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’. 2 exs. ‘Hong Kong / V. 1974 / From C. K. Yu // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’.
Current status.
Valid species.
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1ex. ‘Hong Kong / 1975 / From. C. K-Yu // Paratype: Apogonia / hongkongica / Y. MIYAKE, 1989’. 1 ♀ ‘Hong Kong / ♀ 1975 [white label, front] // from / C. K. Yu [white label, back] // Paratype: / Apogonia / hongkongica / Y. MIYAKE, 1989’.
Apogonia inconstans
Kobayashi & Bezděk
9D598169-F5EA-5D40-BFB9-2D9F962506CA
Apogonia inconstans Kobayashi & Bezděk, 2011: 64−65.
Note.
The following paratypes are deposited in RIEB (ex coll. A. Bezděk):
Paratypes.
4 exs.: 1 ♂ ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 5/18 夜= yoru [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 5/18 夜= yoru [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // 夜= yoru 5/18 [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’. 1 ex. ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N-Thailand / Col. S. Ohmomo [white label, front] // night [white label, back] // PARATYPE / Apogonia / inconstans / Kobayashi et Bezděk, 2011’.
Current status.
Valid species.
Apogonia ohmomoi
Miyake
61714DF5-6C92-52D4-BA22-3816312E7F35
Apogonia ohmomoi Miyake, 1989: 177−178.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘1985. 5. 18−19. / Wieng Ko Sai Nat. / Park, Phrae Prov., / N−Thailand / Col. S. Ohmomo [white label, front] // 夜= yoru 5/18 [white label, back] // 14 // Holotype: / Apogonia / scabra / Y. MIYAKE // Apogonia ohmomoi / Miyake, 1989 / HOLOTYPUS, ♂ / A. Bezděk det. 2011’. (Fig. 2G)
Type condition.
The aedeagus of the holotype is pinned separately, and the right antenna, right and left protarsus, right mesoclaw, left mesotarsus, right metatarsus and mesonotum are missing.
Current status.
Valid species.
Remark.
The label ‘Apogonia scabra’ was apparently accidentally put by Miyake.
Apogonia ovata
Miyake & Yamaguchi
3A00381F-AB5C-57B8-AF75-93F23E810DEA
Apogonia ovata Miyake & Yamaguchi, 1998: 25−27.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Kaeng Tana / Khong Chiam / Ubonratchathani / [NE-THAILAND] / 29. Aug. 1997 / S. Yamaguchi leg. // Holotype / Apogonia ovata / Y. MIYAKE et YAMAGUCHI 1998’. (Fig. 2H)
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Senior synonym of Apogonia miyakei Bezděk, 2004, see Bezděk (2004).
Apogonia terminalis
Miyake, Yamaguchi & Akiyama
788E5484-1931-57BD-9F86-4438A8A1B71B
Figure 3.
Habitus of holotype specimens. AApogonia terminalis Miyake, Yamagushi et Akiyama BApogonia unidentata Miyake, Yamaguchi et Akiyama CDichelomorpha sublineata Miyake DHolomelia gigantea Miyake EHolotrichia loochooana umebayashii Miyake FHolotrichia yamayai Miyake et Yamaguchi GHoplia choui Miyake HHoplia nakanei Miyake.
Apogonia terminalis Miyake, Yamaguchi & Akiyama, 2002: 56−58.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Don Far Huan (light) / Ubon Ratchathani / E-THAILAND / 17. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Holotype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig. 3A)
Paratypes.
4exs.: 1 ♂ ‘Ban Lao Kram / Mukdahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Paratype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 3 ♀ ‘Don Far Huan (light) / Ubon Ratchathani / E-TAILAND / 17. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // ♀ // Paratype / Apogonia terminalis / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’.
Type condition.
The aedeagus of the holotype is pinned separately, and the left proclaw, right and left mesotarsus, right metaclaw and left metatarsus are missing.
Current status.
Valid species.
Apogonia unidentata
Miyake, Yamaguchi & Akiyama
A81AAD25-5775-590C-89DB-534062211D3B
Apogonia unidentata Miyake, Yamaguchi & Akiyama, 2002: 55−56.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Miyake):
Holotype
(♂). ‘Ban Lao Kram / Mukdahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Holotype / Apogonia / unidentata // Holotype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. (Fig. 3B)
Paratypes.
6 exs.: 4exs. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 1 ex. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 5. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. // 63 // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’. 1 ex. ‘Ban Lao Kram / Mukudahan (light) / NE-THAILAND / 14. Sep. 1999 / S. Yamaguchi & / M. Akiyama leg. / シミレア= shimirea // Paratype / Apogonia unidentata / Y. MIYAKE, YAMAGUCHI / et AKIYAMA 2002’.
Type condition.
The aedeagus of the holotype is pinned separately, and the right protarsus and right and left metatarsi are missing.
Current status.
Valid species.
Genus Dichelomorpha
Dichelomorpha sublineata
Miyake
A53A781B-F26E-59A5-B014-70E43FD6E0FD
Dichelomorpha sublineata Miyake, 1994: 140−141.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Is. Tioman, east of / Malay Pennisular March. 1993, nat, col // 1 [aedeagus mount] // Holotype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. (Fig. 3C)
Paratypes.
34 exs.: 10 ♂ ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 14 exs. ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 1 ♂ ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // 11 // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. 1 ex. ‘Is. Tioman, east of / Malay Pennisular / March. 1993. nat, col // 4 // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’. [[8 exs. ‘Tioman Is. / E. of Malaya / III–1933 (the date given in the original description is III–1993), W. // Paratype: / Dichelomorpha / sublineata / Y. MIYAKE, 1994’]].
Type condition.
The aedeagus of the holotype is pinned separately. The right mesotarsus is missing.
Current status.
Valid species.
Remark.
Miyake (1994) did not provide a photograph of the holotype. Apparently he provided a photograph of a paratype, but we could not trace that specimen.
Genus Holomelia
Holomelia gigantea
Miyake
57CD590E-E82D-5FDD-95EA-5949C1FA3046
Holomelia gigantea Miyake, 1996: 35−36.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake).
Holotype
(♂). ‘19 miles from / Tapah, Malaya / 1–II–1974 / Y. Miyake leg [white label, front] // Wu TET KONG / legt. [white label, back] // 29・5 // 家= ie [aedeagus mount] // Holotype: / Holomelia / gigantea / Y. MIYAKE, 1996’. (Fig. 3D)
Paratypes.
2 exs.: 1 ♂ ‘Cameron H. / MALAYA / Jan–Feb 1974 / Y. Miyake leg. // 30.5 // Paratype: / Holomelia gigantea / Y. MIYAKE, 1996 // 19 miles from / Tapah, Malaya / 1–II–1974 [white label, front] // Wu Tet Kong legt. [white label, back]’. 1 ex. ‘19 miles from / Tapah, Malaya / 1–II–1974 [white label, front] // Wu Tet Kong legt. [white label, back] // Paratype: / Holomelia gigantea / Y. MIYAKE, 1996’.
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Valid species.
Remark.
Miyake (1996) did not provide a photograph of the holotype. Apparently he provided a photograph of a paratype, but we could not trace that specimen.
Genus Holotrichia
Holotrichia loochooana umebayashii
Miyake
5A25A8A7-0A63-570C-A082-7BBD644E57A0
Holotrichia loochooana umebayashii Miyake, 1986a: 1−2.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘〔Loo-Choo〕/ Okinoerabuzima / 21. IV. 1957. / K. Umebayashi // 7 [white label, mounted on a part of tarsus] // HOLOTYPE / Holotrichia / loochooana / umebayashii / Y. MIYAKE, 1986’. (Fig. 3E)
Type condition.
The aedeagus of the holotype is pinned separately. The left protarsus and the right and left metatarsus are missing.
Current status.
Junior subjective synonym of Nigrotrichia loochooana okinawana (Nomura, 1964), see Matsumoto (2016).
Remark.
The collecting date in the original description is 1.V.1957, which disagrees with the label data.
Holotrichia yamayai
Miyake & Yamaguchi
78F48E7C-5440-547E-928D-E7C3EFA269F1
Holotrichia yamayai Miyake & Yamaguchi, 1998: 27−28.
Note.
The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘North Thailand / Chiangmai-Saraphee / 15th, June, 1996 / Native collector // Holotype / Holotricchia yamayai / Y. MIYAKE et YAMAGUCHI 1998’. (Fig. 3F)
Paratype.
1 ex.: 1 ♀ ‘Kaeng Tana / Khong Chiam / Ubonratohathani / [NE-THAILAND] / 27. Aug. 1997 / S. Yamaguchi leg. // Paratype / Holotrichia yamayai / Y. MIYAKE et YAMAGUCHI 1998’.
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Valid species.
Genus Hoplia
Hoplia choui
Miyake
3894136C-383B-503D-8CEE-E5FC9E0DBCFB
Hoplia choui Miyake, 1986b: 203−204.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Sozan / Formosa / 26. V. 1940 / Coll. M. Uno // HOLOTYPE / Hoplia / Choui Y. MIYAKE / 1986’. (Fig. 3G)
Type condition.
Abdomen only.
Current status.
Valid species.
Remark.
Most part of the holotype specimen was destroyed by Anthrenus verbasci.
Hoplia nakanei
Miyake
91A3288A-ADAE-5303-B357-F3A40BF0FAAA
Hoplia nakanei Miyake, 1986b: 207.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘ (Taiwan) / Jibbon / 2. iv. 1970 / Y. Miyake // 1 // HOLOTYPE / Hoplia / nakanei Y. MIYAKE’. (Fig. 3H)
Paratypes.
29 exs.: 4 exs. ‘ [Taiwan] / 六亀= liugui．石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 5 exs. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / 六亀= liugui．石山= shi shan / 9–VI–1976 / 陳文竜= chen wenzhao leg. // Paratype / Hoplia nakanei / Y. Miyake, 1986’. 4 exs. ‘ [Taiwan] / Baibara / –V–1963 / T. Shirozu leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / Baibara / –V–1963 / Shirozu leg. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘ [Taiwan] / Baibara / –V–1963 / T. Shirozu // Paratype: Hoplia / nakanei Y. Miyake, 1986’. 1 ♂ ‘六亀= liugui．石山= shi shan / alt. 2000 m / 9. VI. 1976 // Paratype: / Hoplia / nakanei / Y. MIYAKE, 1986’. 1 ex. ‘FENCHIFO / FORMOSA / 24. V. 1975 / K. MATSUDA // Paratype: / Hoplia nakanei / Y. MIYAKE, 1986’. 1 ex. ‘Swakang / Taiwan / 15. VI. 1970 / Fukuda // Paratype: / Hoplia nakanei / 1986. Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 1. IV. 1970 / H. NOMURA // PARATYPE / Hoplia / nakanei / Y. MIYAKE, 1986’. [[4 exs. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake’]]. [[2 exs. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ♂ ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ex. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / Y. Miyake, 1986’]]. [[1 ex. ‘ [Taiwan] / Baibara / V. 1965 (the date given in the original description is V. 1963) / Col. T. Shirozu // Paratype / Hoplia nakanei / 1986. Y. Miyake’]].
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Valid species.
Remark.
The label data the holotype do not match the original description. There is one male paratype (‘六亀= liugui．石山= shi shan / alt. 2000 m / 9. VI. 1976 // Paratype: / Hoplia / nakanei / Y. MIYAKE, 1986’) that agrees with the data of the holotype. Probably the type labels were mixed up and this specimen should be considered the holotype.
In addition to the paratypes listed above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘ [Taiwan] / Liukuei / –V–1963 / T. Shirozu col. // Paratype / Hoplia / nakanei / Y. Miyake, 1986’. 1 ex. ‘APR II. 1971 / 榮華= ronghua / B-S. CHANG // PARATYPE / Hoplia / nakanei / Y. MIYAKE, 1986’. 1 ex. ‘Paratype / Hoplia nakanei / Y. MIYAKE, 1986’.
Hoplia nengkaoshana
Miyake
C1DE97FD-1C02-5F60-8CAE-CE2832FF5B81
Hoplia nengkaoshana Miyake, 1986b: 211.
Note.
The allotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Allotype (♀). ‘Allotype // mt. Noko (2650 m) / V. 20–VI. 2, 1966 // 1 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’.
Paratypes.
3ex.: 1 ♂ ‘mt. Nōhō / C-Formosa / V–20 ～ Vi–21 1966 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’. 1 ♂ ‘MT. HÔFAN / FORMOSA / 5–V. 73 / T. OCHI // 1 // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986 // Hoplia / inornata / H. Kobayashi, 1990 / Det. H. Kobayashi, 2015’. 1 ex. ‘JUL. 16. 1966 / 合欢山= hehuan shan / B. S. Chang // PARATYPE / nengkaoshana / Y. MIYAKE, 1986 // Hoplia / inornata / H. Kobayashi, 1990 / Det. H. Kobayashi, 2015’.
Current status.
Valid species.
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘台中= tai chung 和平郷= heping xiang 早等村= zaodeng cun / 横貫公路= hengguan gonglu. 梨山站= lishan zhan / 海抜= kaibatsu 1679–2196 m. / 13–16. V. 1958. 陶家駒= tao jiayu // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’. 1 ex. ‘AUG 13. 1970 / 新坡= xinpo / B-S. CHANG // PARATYPE / Hoplia / nengkaoshana / Y. MIYAKE, 1986’.
Hoplia simillima
Miyake
E2A3BA86-D46A-575B-834E-1B0CF8CB43B4
Figure 4.
Habitus of holotype specimens. AHoplia simillima Miyake BHoplia taiwana Miyake CMaladera hiranoi Miyake DMaladera kusuii Miyake ENematophylla sugiharai Miyake FPseudohoplia shibatai makiharai Miyake GPseudohoplia shibatai matsudai Miyake HParatrichius kyushuensis Miyake.
Hoplia simillima Miyake, 1986b: 206−207.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1979 / Y. Miyake leg. // HOLOTYPE / Hoplia / simillima / Y. MIYAKE, 1986’. (Fig. 4A)
Paratypes.
5 exs.: 3 exs. ‘(FORMOSA) / Baibara / –V–1969 // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 2 exs. ‘(FORMOSA) / Baibara / –V–1969 / T. Shirozu // Paratype / Hoplia / simillima / 1989, Y. Miyake’.
Current status.
Valid species.
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1979 / Y. Miyake leg. // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 1 ♂ ‘石= ishi // [Taiwan] / 六亀= liugui 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // Paratype / Hoplia / simillima / 1989, Y. Miyake’. 1 ex. ‘(FORMOSA) / Mt. Ali / 14 V 1968 / Y. HAYASHI // Paratype’.
Hoplia taiwana
Miyake
8EF55CF4-A983-5C6F-BBBD-8F7CFDAF720C
Hoplia taiwana Miyake, 1986b: 204.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘HOLOTYPE / Hoplia / Taiwana Y. MIYAKE / 1986’. (Fig. 4B)
Paratypes.
9 exs.: 1 ex. ‘NANSHANCHI / TAIWAN / 3. IV. 1970 / H. NOMURA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 4. IV. 1970 / H. NOMURA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 5. IV. 1970 / Y. KIYOYAMA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘SUNGKANG / FORMOSA / 29. VI. 1971 / Y. MAEDA // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 31. III. 1981 / Y. YAMAMOTO // Paratype / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘NANSHANCHI / TAIWAN / 1. IV. 1981 / F. KIMURA // Paratype / Hoplia / taiwana Y. MIYAKE / (1968)’. 1 ex. ‘NANSHANCHI / TAIWAN / 2. IV. 1981 / F. KIMURA // Paratype / Hoplia / taiwana Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 2. IV. 1981 / F. KIMURA // Paratype / Hoplia taiwana / 1986 Y. MIYAKE’. 1 ex. ‘NANSHANCHI / TAIWAN / 3. IV. 1981 / Y. YAMAMOTO // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’.
Current status.
Valid species.
Remark.
The locality label data of the holotype are missing but the specimen agrees with the original description.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes were not designated in the original description: 3 exs. ‘ [Taiwan] / Liukuei 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘ [Taiwan] / 六亀= liugui 石山= shi shan / 9–VI–1976 / Y. Miyake leg. // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 2 exs. ‘Mt. YUSHAN / TAIWAN / 8. VI. 1980 / M. YAMAMOTO // Paratype / Hoplia / taiwana Y. MIYAKE’. 4 exs. ‘TAIWAN / 陽明山= mt. yangmingshan / 20. IV. 1956 / COL. K. S. LIN // Paratype / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘LUSHAN / FORMOSA / 13–IV. 74 / S. TAKEDA // PARATYPE / Hoplia / taiwana / Y. MIYAKE, 1986’. 1 ex. ‘榮華= ronghua / APR. 7. 1971 / BS, CHANG // PARATYPE / Hoplia taiwana / Y. MIYAKE, 1986’. 1 ex. ‘ [Taiwana] / Baibara / –V–1965 / T. Shirozu leg. // Paratype: Hoplia / taiwana Y. MIYAKE. 1986’.
Genus Maladera
Maladera hiranoi
Miyake
E833BA3A-DD9C-5399-B697-9ED67E6D5A63
Maladera hiranoi Miyake, 1986b: 3−5.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘三京= mikyô / 21/VII–82 // HOLOTYPE / Maladera / hiranoi MIYAKE / (1986) // Maladera / impressthorax / hiranoi Y. MIYAKE / DET. Y. MIYAKE, 199 // ムナクボビロウドコガネ= munakubobirôdokogane / Maladera (japonica–G) / impressithorax / Nomura, 1973 / Det. H. Hirasawa 2012’. (Fig. 4C)
Current status.
Junior subjective synonym of Maladera okinoerabuana Kobayashi, 1978, see Ahrens and Bezděk (2016).
Remark.
The date on the collecting data label of the holotype is 21.VII.1982, but the collecting date quoted in the original description is 22.VII.1982. Probably, Miyake made a typographical error when writing the description.
Maladera kobayashii
Nomura
4BCAA1B3-A149-563D-A100-6D1F62049CE7
Maladera kobayashii Nomura, 1974: 106−107.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
2 exs.: 1 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // PARATYPE / Maladera (s. str) / kobayashii / NOMURA (1974)’. 1 ♀ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♀ // PARATYPE / Maladera / kobayashii / NOMURA (1974)’.
Current status.
Valid species.
Maladera kusuii
Miyake
0D97437C-AEE7-5D0A-9C7B-AD4C918AD396
Maladera kusuii Miyake, 1986b: 2−3.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘三宅= miyake / コガネムシ上科標準図鑑= koganemushijyôka-hyôjyunzukan / ビロウドコガネ族= birôdokoganezoku / 図鑑使用標本= zukanshiyôhyôhon / 執筆担当= shippitsutantô：平沢 伴明= hirasawa tomoaki / 学研教育出版= gakkenkyôikushuppan (2012) // GAKIYA / Iheya-jima / Aug. 11 1977 / leg. Y Kusui // イヘヤビロウドコガネ= iheyabirôdokogane / Maladera (japonica–G) / kusuii / Y. Miyake, 1986 / Det. H. Hirasawa 2012 // Holotype: / Maladera kusuii / Y. MIYAKE, 1986’.
Type condition.
The aedeagus of the holotype is pinned separately. The right and left protarsi, the left mesotarsus, and the right and left metatarsi are missing, but some of the missing parts are pinned separately.
Current status.
Valid species.
Maladera lishana
Miyake
4C47BEBD-4B6C-5DCF-9699-F2B9AC54951F
Maladera lishana Miyake, 1989b: 37−38.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: 1 ♀ ‘Lishan / Taiwan / 28. VII. 1973 / Y. Miyake // Paratype: / Maladera / lishana / Y. MIYAKE, 1989’.
Current status.
Valid species.
Maladera nanshanchiana
Nomura
E829D807-21AE-5780-9CA9-5F64D50D126A
Maladera nanshanchiana Nomura, 1974: 111−112.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
2 exs.: 1 ex. ‘Puli 24 // PARATYPE / Maladera / nanshanchiana / NOMURA (1974)’. 1 ♀ ‘Puli 24 // ♀ // PARATYPE / Maladera / nanshanchiana / NOMURA (1974) // ナンザンケイビロウドコガネ= nanzenkeibirodokogane / Maladera (Cephaloserica) / nanshanchiana / Nomura, 1974 / Det. H. Hirasawa 2012’.
Current status.
Valid species.
Maladera secreta horaiana
Nomura
7D4441E9-A5DD-5463-944A-DBA57E2EB4FE
Maladera secreta horaiana Nomura, 1974: 112.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratype.
1 ex.: 1 ♂ ‘Lishan / C. Taiwan / 29. vii. 1973 / Y. Miyake // 三宅= miyake // PARATYPE / Maladera / secreta / horaiana / NOMURA (1974)’.
Current status.
Valid species.
Genus Melanomaladera
Melanomaladera yunnana
Miyake & Yamaya
D8508FD0-D29C-59F2-84A4-314045E0FE8D
Melanomaladera yunnana Miyake & Yamaya, 2001: 38−39.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
2 exs.: 1 ♂ ‘China, N W Yunnan / Degen City env. / 3300 m alt. / 29. Jun. 1998 / A. Gorodinsky leg. // 13 // Paratype: / Melanoserica / yunnana’. 1 ♀ ‘China, N W Yunnan / Degen City env. / 3300 m alt. / 29. Jun. 1998 / A. Gorodinsky leg. // 13 // Paratype: / Melanoserica / yunnana’.
Current status.
Archeohomaloplia yunnana (Miyake et Yamaya, 2001), see Ahrens and Bezděk (2016).
Remark.
The genus name on the paratype labels is ‘Melanoserica’, which is invalid. Probably, it was confused with ‘Melanomaladera’ by Y. Miyake.
Genus Microserica
Microserica (Parvulomaladera) annapurnae
Ahrens
449910E6-6CA0-5765-AD9B-9C96964C31AA
Microserica ( Parvulomaladera) annapurnae Ahrens, 1995: 46−48.
Note.
The following paratype is deposited in RIEB (ex coll. D. Ahrens):
Paratype.
1 ex.: ‘NEPAL–HIMALAYA / Annapurna–Mts. / leg. Ahrens 1993 // Pisang bis / Manang, 30. 5. / 3000–3300 m // PARATYPEUS / Parvulomaladera / annapumae sp. n. / det. D. AHRENS 1994’.
Current status.
Oxyserica pygidialis annapurnae (Ahrens, 1995), see Ahrens and Bezděk (2016).
Microserica nitidipyga
Nomura
107FF582-A99F-5A8C-BD61-87C3448AB785
Microserica nitidipyga Nomura, 1974: 99.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratype.
1 ex.: ‘Lu-SHAN / FORMOSA / 27. IV. 1973 / K. MATSUDA // 三宅= miyake // Paratype / Microserica / nitidipyga / NOMURA (1974) // Microserica / nitidipyga NOM. / DET. Y. MIYAKE, 1994’.
Current status.
Valid species.
Genus Nematophylla
Nematophylla sugiharai
Miyake
D799782B-4E7E-58C4-A6B9-710CC7DB1C61
Nematophylla sugiharai Miyake, 2000: 106−108.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Kiau Gap / Mt. Kinabalu / Sabah, Borneo / 1. IV. 2000 / H. SUGIHARAI leg. // Holotype: Nematophylla / sugiharai / Y. MIYAKE 2000’. (Fig. 4E)
Current status.
Junior subjective synonym of Nematophylla carinicollis Arrow, 1938, see Krajcik (2012).
Genus Nipponoserica
Nipponoserica pubiventris
Nomura
B6640F5D-26A5-5BA2-BACB-DF131D0F9C59
Nipponoserica pubiventris Nomura, 1976b: 189−190.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratype.
1 ex.: ‘Takashinohama, / Osaka Pref. / 3. VI. 1974 / Coll. Mitsuo Goto // at light // PARATYPE / Nipponoserica / pubiventris / NOMURA (1976)’.
Current status.
Valid species.
Genus Paramaladera
Paramaladera makiharai
Nomura
0A6553B9-51DD-5B8D-99CC-21368211657C
Paramaladera makiharai Nomura, 1974: 101−102.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
4 exs.: 4 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // Paratype / Paramaladera / makiharai / NOMURA (1974)’.
Current status.
Valid species.
Genus Pseudohoplia
Pseudohoplia shibatai
Miyake
E6E6C84E-F6DE-556A-AF6B-0096F0C3EFD7
Pseudohoplia shibatai Miyake, 1986b: 201.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
2 exs.: 1 ♂ ‘（FORMOSA）/ Mt. Ali / 15 V 1968 / Y. HAYASHI // Paratype // Hoplia / shibataishibatai / Y. Miyake, 1986 / Det. H. Kobayashi, 2015’. 1 ex. ‘（FORMOSA）/ Mt. Ali / 15 V 1968 / Y. HAYASHI // Paratype // Hoplia / shibataishibatai / Y. Miyake, 1986 / Det. H. Kobayashi, 2015’.
Current status.
Hoplia shibatai shibatai (Miyake, 1986), see Kobayashi (2017).
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘Mt. Arishan / Formosa / 1. V. 1973 / K. MATSUDA // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986 // 贈台湾農試= zou-taiwan-noushi’. 1 ♂ ‘Houanchi- / Sundchuankang / Nantou Hsien / Taiwan / 25. VI. 1976 / H. Makihara leg. // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’. 1 ex. ‘Mt. Arishan / Formosa / 1. V. 1973 / K. MATSUDA // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’. 1 ex. ‘MT. ARI / FORMOSA / 1. V. 1973 / K. MATSUDA // Paratype // Paratype / Pseudohoplia / shibatai / Y. Miyake, 1986’.
Pseudohoplia shibatai makiharai
Miyake
0A49E652-A2BD-500A-AE0D-D633F9C22127
Pseudohoplia shibatai makiharai Miyake, 1986b: 202.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Mt. Lalashan / Taoyuan Hsien / Taiwan / 21–24. V. 1980 / H. Makihara // HOLOTYPE / Pseudohoplia / shibataimakiharai / Y. MIYAKE, 1886’. (Fig. 4F)
Type condition.
The aedeagus of the holotype is pinned separately. The right antenna is missing.
Current status.
Hoplia shibatai makiharai (Miyake, 1986), see Kobayashi (2017).
Pseudohoplia shibatai matsudai
Miyake
E4D86FE5-5862-5010-9784-DE4376BD3B2A
Pseudohoplia shibatai matsudai Miyake, 1986b: 202.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Mt. Nōko / C-Formosa / v–20～vi–21’ 66 // HOLOTYPE / Pseudohoplia / shibataimatsudai / Y. MIYAKE, 1986’. (Fig. 4G)
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Junior subjective synonym of Hoplia shibatai shibatai (Miyake, 1986), see Kobayashi (2017).
Genus Pseudomaladera
Pseudomaladera nitidifrons
Nomura
9DEE74B8-2B9B-59B4-A30C-2F99F861E129
Pseudomaladera nitidifrons Nomura, 1974: 96.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
2 exs.: 1 ♂ ‘FUNCHIHO / FORMOSA / 1. VIII. 1973 / H. OHASHI // ♂ // 三宅= miyake // Paratype / Pseudomaladera / nitidifrons / NOMURA, 1974’. 1 ♂ ‘MEIFENG / FORMOSA / 1. VIII. 1973 / T. OCHI // ♂ // Paratype / Pseudomaladera / nitidifrons / NOMURA, 1974’.
Current status.
Valid species.
Genus Sinoserica
Sinoserica maculipennis
Miyake & Yamaya
C96AC3DB-B6FC-5375-ABC4-68BFAC52E78D
Sinoserica maculipennis Miyake & Yamaya, 2001: 40−41.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
2 exs.: ‘China, Yunnan / Boshan C. env. / 2100 m alt. / 10. VII. 1998 / A. Gorodinski // 12 // Paratype / Sinoserica / maculipennis’.
Current status.
Trioserica maculipennis (Miyake et Yamaya, 2001), see Ahrens and Bezděk (2016).
Genus Serica
Serica fusifemorata
Nomura
405A8EA4-C8DC-56B6-8F16-5FFB567CD607
Serica fusifemorata Nomura, 1974: 91.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
11 exs.: 1 ♀ ‘Lishan / C. Taiwan / 29. vii. 1973 / Y. Miyake // ♀ // 三宅= miyake // Paratype / Serica / fusifemorata / NOMURA (1974)’. 2 ♀ ‘Li-shan / C. Taiwan / 26. VII. 1974 / Y. Miyake // ♀ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 2 ♂ ‘Li-shan / C. Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 2 ♀ ‘Li-shan / C. Taiwan / 29. VII. 1974 / Y. Miyake // ♀ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 1 ♂ ‘Li-shan / C. Taiwan / 1. VIII. 1974 / Y. Miyake // ♂ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 2 ♀ ‘Li-shan / C. Taiwan / 1. VIII. 1974 / Y. Miyake // ♀ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’. 1 ♀ ‘27 // ♀ // Paratype / Serica / fusifemorata / S. NOMURA (1974)’.
Current status.
Valid species.
Serica gansuensis
Miyake & Yamaya
26F569ED-6B83-5728-9BFE-D6040B09D546
Serica gansuensis Miyake & Yamaya, 2001: 35−36.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: ‘China, Gangu / Mts. Minshan / nr. Wudn. 2100 / 1. VI. 1997 / A. Gorodinski // 4 // Paratype: / Serica / gansuensis’.
Current status.
Valid species.
Genus Sericania
Sericania miyakei
Nomura
77BAE2CB-4596-547A-830A-CEF2400A4671
Sericania miyakei Nomura, 1960: 60.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratype.
1 ex.: ‘Paratype // TASHIRO / CHIKUGO / 4. V. 1952 / Y. MIYAKE // Sericania / miyakei / NOMURA (1960)’.
Current status.
Valid species.
Genus Sophrops
Sophrops takatoshii
Itoh
ABB7658D-82E2-5FDA-AFAD-69E263597833
Sophrops takatoshii Itoh, 1990: 9−10.
Note.
The following paratypes are deposited in RIEB (ex coll. T. Itoh):
Paratypes.
3 exs.: ‘1989–4–17 / 平良植物園= hirara-syokubutsuen /（宮古島）= miyako-jima / leg. T. Ueno // Paratype / Sophrops / takatoshii / T. Itoh’.
Current status.
Valid species.
Genus Taiwanoserica
Taiwanoserica elongata
Nomura
AAC3C5FA-BDAA-5AC2-AA48-50802597F146
Taiwanoserica elongata Nomura, 1974: 86.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratypes.
1 ex.: 1 ♂ ‘MT. ARI / FORMOSA / 30. VII. 1973 / H. OHASHI // ♂ // 三宅= miyake // Paratype / Taiwanoserica / elongata / NOMURA (1974)’.
Current status.
Serica (Taiwanoserica) elongata (Nomura, 1974), see Ahrens and Bezděk (2016).
Taiwanoserica lishana
Nomura
41FA8D7B-9F74-51D4-BA84-DACE70F2825A
Taiwanoserica lishana Nomura, 1974: 87.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
22 exs.: 2 ♂ ‘Lishan / C. Taiwan / 30. vii. 1970 / Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 6 ♂ ‘Lishan / C-Taiwan / 29. vii. 1973 / Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 4 ♀ ‘Lishan / C-Taiwan / 29. vii. 1973 / Y. Miyake // ♀ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 4 ♂ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 1 ♀ ‘Li-shan / Taiwan / 29. VII. 1974 / Y. Miyake // ♀ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 4 ♂ ‘Li-shan / Taiwan / 1. VII. 1974 // Y. Miyake // ♂ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’. 1 ♀ ‘Li-shan / Taiwan / 1. VII. 1974 // Y. Miyake // ♀ // PARATYPE / Taiwanoserica / lishana / S. Nomura (1974)’.
Current status.
Serica (Taiwanoserica) lishana (Nomura, 1974), see Ahrens and Bezděk (2016).
Genus Trichomaladera
Trichomaladera elongata
Nomura
072CD2CC-FE5D-50B0-9BE8-9739964B7400
Trichomaladera elongata Nomura, 1974: 93−94.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratype.
1 ex.: 1 ♂ ‘Paratype / Trichomaladera / elongata / NOMURA (1974)’.
Current status.
Valid species.
Remark.
The collecting data label of the paratype is missing.
Genus Trichoserica
Trichoserica elongata nitididorsis
Nomura
0D69CA90-4BAE-5465-B80F-DBB1798A5E28
Trichoserica elongata nitididorsis Nomura, 1971: 70−71.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratype.
1 ex.: ‘Mt. Hiko / [KYUSHU] / 27. 7. 1960 / TAKEISHI // Paratype / Trichoserica / elongata / nitididorsis / Nomura’.
Current status.
Serica nitididorsis (Nomura, 1971), see Ahrens and Bezděk (2016).
Trichoserica incurvata
Nomura
937C9036-527A-55E4-8EE3-B8B533B6C78C
Trichoserica incurvata Nomura, 1971: 67−68.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
8 exs.: 1 ex. ‘Aomori-pref / 17–vii–1956 / S. Kimotog // Paratype / Trichoserica / incurvata / Nomura (1971) // Aomori / 17. VII. 1956 / S. Kimoto’. 1 ♂ ‘17. VII. 1956 / S. Kimoto // Paratype / Trichoserica / incurvata / Nomura (1971)’. 2 exs. ‘Aomori-pref / 19–vii–1956 / S. Kimotog // Paratype / Trichoserica / incurvata / Nomura (1971) // Kimoto S. / 19. VII. 1956’. 1 ex. ‘Aomori-pref / 19–vii–1956 / S. Kimotog // Paratype / Trichoserica / incurvata / Nomura (1971) // 19. VII. 1956 / S. Kimoto’. 2 exs. ‘Sensuidani 1200 m / Okutama Tokyo / 21–VII–1968 / Coll. H. Yuasa // Paratype / Trichoserica / incurvata / Nomura (1971)’. 1 ex. ‘Okutama / 21. VII. 1968 / YUASA // Paratype / Trichoserica / incurvata / Nomura (1971)’.
Current status.
Serica incurvata (Nomura, 1971), see Ahrens and Bezděk (2016).
Trichoserica nitidiceps
Nomura
CCC606C9-2E36-55D4-AEA2-F3DD74E683E9
Trichoserica nitidiceps Nomura, 1971: 69−70.
Note.
The following paratype is deposited in RIEB (ex coll. S. Nomura):
Paratype.
1 ex.: 1 ♂ ‘(Kyushu) / Mt. Sobo / 23. V. 1952 / Y. Miyake //Paratype / Trichoserica / nitidiceps / NOMURA’.
Current status.
Junior subjective synonym of Serica inexspectata (Kontkanen, 1956), see Ahrens and Bezděk (2016).
Remark.
In addition to the paratype mentioned above, the following specimen labeled as paratype is not designated in the original description: 1 ex. ‘Mt. Kuju / Bungo / 28. vii. 1961 / Y. Miyake // Paratype / Trichoserica / nitidiceps / NOMURA (1971)’.
Trichoserica opacifrons
Nomura
4489A415-A7B2-5974-9793-05208846DC21
Trichoserica opacifrons Nomura, 1971: 68−69.
Note.
The following paratypes are deposited in RIEB (ex coll. S. Nomura):
Paratypes.
3 exs.: 1 ♂ ‘Mt. Hiko / Buzen / 1. VIII. 1953 / Y. Miyake // Paratype / Trichoserica / opacifrons / NOMURA’. 1 ex. ‘mt. Kujū / Hokkeein / 27. VII. 1961 / Y. Miyake Coll // Paratype / Trichoserica / opacifrons / NOMURA’. 1 ♂ ‘ [Kyushu] / Yakushima / 21. Vi. 1954 / Col. K. Matsuda // Paratype / Trichoserica / opacifrons / NOMURA // Ophthalmoserica/ matsudai / sp.n. / DET. Y. MIYAKE’.
Current status.
Senior objective synonym of Serica planifrons Nomura, 1972, see Ahrens and Bezděk (2016). Nomura (1972) proposed Serica planifrons as a replacement name for Serica opacifrons (Nomura, 1971), a junior secondary homonym of Serica opacifrons Fairmaire, 1891.
Subfamily Cetoniinae
Genus Anomalocera
Anomalocera paotao
Masumoto & Sakai
357351E6-02D3-5EC6-80C1-0E7BBE2F3540
Anomalocera paotao Masumoto & Sakai, 1987a: 51−54.
Note.
The following paratypes are deposited in RIEB (ex coll. Masumoto):
Paratypes.
2 exs.: 1 ex. ‘Lishan / Taiwan / 28. VII. 1974 / Y. Miyake // Paratype / Anomalocera / paotao MAS. et SAK.’. 1 ex. ‘Lishan / C, -Taiwan / 29 vii. 1973 / Y. Miyake // Paratype / Anomalocera / paotao MAS. et Sak.’.
Current status.
Diphyllomorpha paotao (Masumoto et Sakai, 1987), see Bezděk (2016).
Genus Rhomborhina
Rhomborhina kurosawai
Masumoto & Sakai
1AEF134E-A348-500D-8AA3-6B909D4DBB88
Rhomborhina kurosawai Masumoto & Sakai, 1987b: 104−105, 107.
Note.
The following paratypes are deposited in RIEB (ex coll. Masumoto):
Paratypes.
3 exs.: 2 ♂ ‘FORMOSA / tattaka / 9, VIII 1971 / H. OHARA // Paratype / Rhomborrhina / kurosawai / MASUMOTO et K. SAKAI’. [[1 ♂ ‘Tsuifeng / C. Tsiwan / 26. vii. 1970 (the date given in the original description is 27.vii.1970) // Y. Miyake // Paratype / Rhomborrhina / kurosawai / MASUMOTO et K. SAKAI’]].
Current status.
Valid species.
Genus Thaumastopeus
Thaumastopeus chicheryi tenuigenius
Miyake & Yamaya
2DC8B93A-3ED6-5DC0-ACF5-A8D49E5303A0
Thaumastopeus chicheryi tenuigenius Miyake & Yamaya, 1995: 38.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: 1 ♂ ‘N. Keningau / N. Borneo / 30. V. 1994 // Paratype: Thaumastopeus / chicheryitenuigenius / Y. MIYAKE et YAMAYA, 1995’.
Current status.
Valid species.
Genus Paratrichius
Paratrichius hatai
Miyake
3BDD0267-EC48-5C35-B9A4-B703F440D8A9
Paratrichius hatai Miyake, 1989: 41−42.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
4 exs.: [[‘Tanah Rata / MALAYA / 7. IV. 1972 (the date given in the original description is 7.IV.1978) / M. Hata // Paratype: / Paratrichius / hatai / Y. MIYAKE, 1989’]].
Current status.
Valid species.
Paratrichius kyushuensis
Miyake
E5E0CDE7-4F2C-5542-BDAF-A447CA3897D0
Paratrichius kyushuensis Miyake, 1990: 30−32.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘MT. SOBO / BUNGO / 28. vii. 1954 / Y. MIYAKE // Holotype: / Paratrichius / kyushuensis / Y. MIYAKE, 1990’. (Fig. 4H)
Paratypes.
5 exs.: 1 ex. ‘(Kyushu) / Mt. Sobo / 24. VII. 1950 / Y. Miyake // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ♂ ‘(Kyushu) / Mt. Sobo / 24. VII. 1952 / Y. Miyake // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ex. ‘(Kyushu) / Mt. Ichi Fu Sa / 28 VII 1952 / Col. S. Kimoto // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’. 1 ex. 1 ♂ ‘(Kyushu) / Mt. Sobo / 24. VII. 1953 / K. MATSUDA // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’.
Type condition.
The left metatarsus of the holotype is missing.
Current status.
Valid species.
Remark.
The date on the label of the holotype is 28.VII.1954 while that in the original description is 28.VII.1952.
In addition to the paratypes mentioned above, the following specimen labeled as paratype is not designated in the original description.: 1 ex. ‘⑦ // Paratype: Paratrichius / kyushuensis / Y. MIYAKE, 1990’.
Genus Tibiotrichius
Tibiotrichius vietnamensis
Miyake
EBC2D9E6-06B4-5EDD-BD77-EFDD01573701
Figure 5.
Habitus of holotype specimens. ATibiotrichius vietnamensis Miyake BCharitovalgus banzai Sawada CDasyvalgus annamensis Miyake DDasyvalgus castaneodorsalis Miyake EDasyvalgus decamaculatus Miyake FDasyvalgus flavicauda Miyake GDasyvalgus macacus Miyake HDasyvalgus multicus Miyake.
Tibiotrichius vietnamensis Miyake, 1996: 44.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘M. ITO leg / Sapa=Chapa / N. Vietnam / 23–V–1995 // サパ= sapa [aedeagus mount] // Holotype: Tibiotrichius / vietnamensis Y. MIYAKE, 1996’. (Fig. 5A)
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Valid species.
Subfamily Valginae
Genus Charitovalgus
Charitovalgus banzai
Sawada
C1A03233-EFFC-5E3F-96CD-37A074F67167
Charitovalgus banzai Sawada, 1941: 8−9.
Note.
The neotype is deposited in RIEB (ex coll. Y. Miyake):
Neotype
(♂). ‘Hourng Shoei ken / near Liukuei, / Kao Hsung Hsien / VII. 1972 W. Chen leg // Holotype: Neotype / Charitovalgus / banzai SAWADA / Y. MIYAKE, 1993’. (Fig. 5B)
Type condition.
The right and left metatarsi of the neotype are missing.
Current status.
Valid species.
Remark.
The neotype was designated by Miyake (1993).
Genus Dasyvalgus
Dasyvalgus annamensis
Miyake
F621BF50-71E8-5912-AB11-752C31A172A3
Dasyvalgus annamensis Miyake, 1993a: 21−22.
Note.
The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘4. 4 / 203 // Iluon Ma Thuot / Annam M. ITO // Holotype: Dasyvalgus / Annamensis / Y. MIYAKE, 1993’. (Fig. 5C)
Paratype.
1 ex.: 1 ♂ ‘Iluon Ma Thuot / Annam M. ITO // Paratype: Dasyvalgus / annamensis / Y. MIYAKE, 1993’.
Current status.
Valid species.
Remark.
The locality on the collecting data label of the holotype is ‘Iluon Ma Thuot’, but the locality given in the original description is ‘Ban Me Thuot’. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake made a typographical error when writing the description.
Dasyvalgus castaneodorsalis
Miyake
A920D72A-FEBB-5DF3-B0B4-8F68D6F9C77F
Dasyvalgus castaneodorsalis Miyake, 1993a: 24−25.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘〔Thailand〕/ Pupin–Doi Phui / 19–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / casraneodorsalis / Y. MIYAKE, 1993’. (Fig. 5D)
Paratypes.
6 exs.: 2 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 19–IV–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–IV–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 2 ♂ ‘〔Thailand〕/ Pupin / 21–IV–1985 / M. TAO // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Doi Suthep / 10–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / castaneodorsalis / Y. MIYAKE, 1993’.
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Valid species.
Dasyvalgus decamaculatus
Miyake
009FA200-B1A7-5663-B4A4-13E3612B57D2
Dasyvalgus decamaculatus Miyake, 1993a: 22−23.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Iluon Ma Thuot / Annam M. ITO // Holotype: / Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’. (Fig. 5E)
Paratypes.
2 exs.: 1 ex. ‘Near Chang Mai / N. Thailand / 7–V–1994 / K. KUME leg. // Paraype: Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’. 1 ex. ‘Paratype: Dasyvalgus / decamaculatus / Y. MIYAKE, 1993’.
Type condition.
The right and left metatarsus are missing.
Current status.
Valid species.
Remark.
The locality on the collecting data label of the holotype is ‘Iluon Ma Thuot’, but the locality given in the original description is ‘Ban Me Thuot’. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.
Dasyvalgus flavicauda
Miyake, Yamaguchi et Aoki
71C09282-C078-521A-B055-8FCAA7DAD4C3
Dasyvalgus flavicauda Miyake, Yamaguchi, Aoki et Akiyama, 2002: 65−66.
Note.
The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Ban Lao Kram / Mukdahan / E-THAILAND / 17. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // Holotype: / Dasyvalgus / flavicauda // Holotype / Dasyvalgus flavicauda / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig. 5F)
Paratype.
1 ex.: 1 ♂ ‘Phu Huai Sing / Mukdahan / E–THAILAND / 19. Feb. 1999 / S. Yamaguchi and / T. Aoki leg. // Paratype / Dasyvalgus flavicauda / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’.
Current status.
Valid species.
Dasyvalgus macacus
Miyake
3E894F15-AE19-5519-A811-43E6A8ABA179
Dasyvalgus macacus Miyake, 1993a: 24.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘〔Thailand〕Pupin–Doi Phui / 19–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / macacus / Y. MIYSKE, 1993’. (Fig. 5G)
Paratypes.
4 exs.: 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 1–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 5–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 12–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi Phui / 21–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / macacus / Y. MIYAKE, 1993’.
Current status.
Valid species.
Remark.
The habitus photograph in the original description does not refer to the holotype.
Dasyvalgus makiharai
Miyake
B2A5823C-EA37-57E7-97DA-D57420920DCA
Dasyvalgus makiharai Miyake, 1985: 11−12.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: ‘MEIFENG / FORMOSA / 3–VII. 74 / T, OCHI // Paratype: / Dasyvalgus / makiharai Y. MIYAKE / (1985)’.
Current status.
Valid species.
Remark.
In addition to the paratype mentioned above, the following specimens labeled as paratype are not designated in the original description: 1 ex. ‘松崗= songgang / 27～29. VII 1968 / K. YAMAMOTO // Paratype: Dasyvalgus / makiharai / Y. MIYAKE, 1985’. 1 ♂ ‘SUNG KANG / FORMOSA / 30–VI, 74 / T. OCHI // Paratype: / Dasyvalgus / makiharai Y. MIYAKE / (1985)’.
Dasyvalgus minahasanus
Miyake
A4E3A16F-21E7-594C-B01D-F5F9183FB20D
Dasyvalgus minahasanus Miyake, 1989b: 42−43.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
78 exs.: 75 exs. ‘Tondano / N. Sulawesi / 10–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’. 3 exs. ‘Tondano / N. Sulawesi / 9–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’.
Current status.
Valid species.
Remark.
In addition to the paratypes mentioned above, the following specimen labeled as paratypes is not designated in the original description: 1 ex. ‘Tondano / N. Sulawesi / 11–IV–1989 / Y. Miyake leg. // Paratype: / Dasyvalgus / minahasanus / Y. MIYAKE, 1989’.
Dasyvalgus multicus
Miyake
415E32BB-7257-567E-BC1A-24AB1AF83AD6
Dasyvalgus multicus Miyake, 1993a: 25−26.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘ [Thailand] / プピン-プイ= pupin-pui / 5–V–1985 / M. Tao leg. // Holotype: Dasyvalgus / multicus / Y. MIYAKE, 1993’. (Fig. 5H)
Type condition.
The aedeagus of the holotype is pinned separately. The right and left protarsus are missing.
Current status.
Valid species.
Dasyvalgus nudis
Miyake
E38711E1-4687-505C-8356-B3418077BE07
Figure 6.
Habitus of holotype specimens. ADasyvalgus nudis Miyake BDasyvalgus rubrothoracicus Miyake CHybovalgus matsudai Miyake DNeovalgus formosanus Miyake EOreoderus quadrimaculatus Miyake FTarsovalgus hatai Miyake.
Dasyvalgus nudis Miyake, 1994: 153.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Cao Bang / N. Vietnam / 9–11. VI, 1993 / M. ITO leg // Holotype: Dasyvalgus / nudis / Y. MIYAKE, 1994’. (Fig. 6A)
Paratypes.
9 exs.: 3 ♂ ‘Cao Bang / N. Vietnam / 8–10 VI, 1993 / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’. 3 ♂ ‘Cao Bang / N. Vietnam / 10. VI, 1993 / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’. [[2 ♂ ‘Cao Bang / N. Vietnam / 9–11. VI, 1993 (real date is 8–10. VI. 1993) / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’]]. [[1 ex. ‘Cao Bang / N. Vietnam / 11. VI, 1993 (real date is 8–10. VI. 1993) / M. ITO leg. // Paratype: / Dasyvalgus / nudis / Y. MIYAKE, 1994’]].
Type condition.
The aedeagus of the holotype is pinned separately. The right antenna is missing.
Current status.
Valid species.
Remark.
The date on the collecting data label of the holotype is 9–11,VI,1993, but the collecting date quoted in the original description is 8–10,VI,1993. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.
Dasyvalgus rubrothoracicus
Miyake
A362D698-39B2-5BBA-9145-1F0E32E71F1E
Dasyvalgus rubrothoracicus Miyake, 1993a: 23−24.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘〔Thailand〕/ Pupin–Doi Phui / 21–IV–1985 / M. TAO leg. // Holotype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. (Fig. 6B)
Paratypes.
11 exs.: 1 ex. ‘Doi Sutehp / Thailand / 28–IV–1985 / M Tao leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 1–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 1–V–1985 / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 5–V–1985 / M. TAO leg. // TDvA // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. 1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 11–V–1985 / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 17–V–1985 (the date given in the original description is 17–IV–1985) / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–V–1985 (the date given in the original description is 21–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 21–V–1985 (the date given in the original description is 21–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[1 ♂‘〔Thailand〕/ Pupin-Doi-Phui / 23–V–1985 (the date given in the original description is 23–IV–1985) / M. TAO leg. // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]]. [[2 ♂ ‘〔Thailand〕/ Pupin-Doi-Phui / 28–V–1985 (the date given in the original description is 28–IV–1985) / M. TAO leg. // TDvA // Paratype: Dasyvalgus / rubrothoracicus / Y. MIYAKE, 1993’]].
Type condition.
The aedeagus of the holotype is pinned separately.
Current status.
Valid species.
Remark.
The date on the collecting data label of the holotype is 21–IV–1985, but the collecting date quoted in the original description is 5–V–1985. However, the habitus of the holotype and the figure in the original description match. Probably, Miyake caused a typographical error when writing the description.
Dasyvalgus similis
Miyake
5167838A-9A15-596E-A172-79D561BA425F
Dasyvalgus similis Miyake, 1985: 8−9.
Note.
The following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Paratypes.
3 exs.: 1 ♂ ‘蓬来= hôrai / 20, 21 VIII 1968 / H. MAKIHARA // Paratype’. 1 ♂ ‘FUNCHIIFO / FORMOSA / 1. VI. 1970 / Y. KIYOYAMA // Paratype // Paratype / Dasyvalgus / similis, Y. MIYAKE / 1985’. 1 ♂ ‘FUNCHIIFO / FORMOSA / 1. VI. 1970 / Y. KIYOYAMA // Paratype’.
Current status.
Senior objective synonym of Dasyvalgus taiwanus Miyake, 1991. Miyake in Miyake et al. (1991) proposed Dasyvalgus taiwanus as a replacement name for Dasyvalgus similis Miyake, 1985, a junior primary homonym of Dasyvalgus similis Moser, 1908.
Remark.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 1 ex. ‘LIUKUEI / FORMOSA / 3. X. 1970 / Y. KIYOYAMA // 16 // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘NANSHANCHI / FORMOSA / 27. VI. 1971 / Y. MAEDA // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / Chen. W. // Paratype’. 1 ♂ ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / Chen. W. // Paratype’. 1 ex. ‘紅水坑= hongshuikeng / Liukuei / 11. V. 1978 / W. Chen // Paratype // Paratype / Dasyvalgus / similis Y. MIYAKE, 1985’. 1 ex. ‘Paratype’.
Dasyvalgus wadai
Miyake
D0B3AFA2-D024-57A6-A144-F4C74A23B05D
Dasyvalgus wadai Miyake, 1985: 9−10.
Note.
The following paratype is deposited in RIEB (ex coll. Y. Miyake):
Paratype.
1 ex.: 1 ♂ ‘Paratype / Dasyvalgus / wadai Y. MIYAKE, 1985’.
Current status.
Valid species.
Remark.
This paratype has no collecting data label.
Genus Hybovalgus
Hybovalgus matsudai
Miyake
4EA313E3-E144-5B78-9B36-AA5A92447B6E
Hybovalgus matsudai Miyake, 1985: 6.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Paratype // Holotype / Hybovalgus / matsudai Y. MIYAKE / (1985)’.
Type condition.
The right protarsus is missing.
Current status.
Valid species.
Remark.
There is no data label attached to the holotype, but the habitus of the holotype matches the figure in the original description. If confidence is placed in the pinned holotype label and the habitus, the above holotype specimen is the true holotype.
Genus Neovalgus
Neovalgus formosanus
Miyake
1B144BD3-2306-5C0A-837F-1049679B8679
Neovalgus formosanus Miyake, 1985: 3.
Note.
The holotype and following paratypes are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘六亀= liugui．石山= shi shan / 9. VI. 1976 / 2000m // Holotype: / Neovalgus / formosanus / Y, MIYAKE, 1985’. (Fig. 6D)
Paratypes.
2 exs.: 1 ex. ‘ [Taiwan] / Baibara, mt. Puli / –V–1965 // Paratype: Neovalgus / formosanus Y. MIYAKE, 1985’. 1 ex. ‘ [Taiwan] / Liukuei / 16–V–1978 // PARATYPE / Neovalgus / formosanus / Y. MIYAKE, 1985’.
Current status.
Valid species.
Remark.
The date on the collecting label of the holotype is 9.VI.1976, but the collecting date quoted in the original description is 16.V.1978. However, the habitus of the holotype matches the figure in the original description. Probably, Miyake caused a typographical error when writing the description.
In addition to the paratypes mentioned above, the following specimens labeled as paratypes are not designated in the original description: 3 exs. ‘ [Taiwan] / Baibara, mt. Puli / –V–1965 // PARATYPE / Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘Formosa / Wushe / V. 1965 / T. Shiroz / Paratype: Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘六 亀= liugui, 石山= shi shan / 2000 m / 9–VI–1976 / Paratype / Neovalgus / formosanus / Y. MIYAKE, 1985’. 1 ex. ‘Paratype: / Neovalgus / formosanus / Y. MIYAKE, 1985’.
Genus Oreoderus
Oreoderus quadrimaculatus
Miyake, Yamaguchi et Aoki
C82B01EE-7E4C-5CE6-80B0-417B8A698CE0
Oreoderus quadrimaculatus Miyake, Yamaguchi, Aoki et Akiyama, 2002: 64−65.
Note.
The holotype is deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘Phu Huai Sing / Mukdahan / E-THAILAND / 19. Feb. 1999 / S. Yamahuchi and T. Aoki leg. // Holotype: / Oreoderus / quadrumaculatus // Or. 4ml / Coll. // Holotype / Oreoderus quadrimaculatus / Y. MIYAKE, YAMAGUCHI / et AOKI 2002’. (Fig. 6E)
Type condition.
The aedeagus of the holotype is pinned separately. The right protarsus is missing.
Current status.
Valid species.
Genus Tarsovalgus
Tarsovalgus hatai
Miyake
67027003-E0F6-5F5B-875B-72327D176BFF
Tarsovalgus hatai Miyake, 1993b: 32−33.
Note.
The holotype and following paratype are deposited in RIEB (ex coll. Y. Miyake):
Holotype
(♂). ‘ [Malaya] / – –1992 // Holotype: Tarsovalgus / hatai Y. MIYAKE, 1993’. (Fig. 6F)
Paratype.
1 ex.: ‘ [Malaya] / – –1992 // Paratype: / Tarsovalgus / hatai / Y. MIYAKE, 1993’.
Current status.
Valid species.
Family Trogidae
Genus Trox
Trox kyotensis
Ochi & Kawahara
5D0F00DB-1B88-5D3F-812D-B59FE8084411
Trox kyotensis Ochi & Kawahara, 2000: 53−56.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: ‘YAMAZAKI- / CHO, KYOTO. / 5–IV, 1999 / M. KAWAHARA // PARATYPE / Trox / kyotensis / OCHI and KAWAHARA / 2000’.
Current status.
Trox (Niditrox) kyotensis Ochi et Kawahara, 2000, see Kalz (2019).
Trox sabulosus fujiokai
Ochi
F2C2474D-E5CB-5695-B111-A24B8FE255AB
Trox sabulosus fujiokai Ochi, 2000: 43−44.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: ‘Yukyuzan-Park, Nagaoka / Niigata Pref., Japan / 28. V. 1997 / Y and M. Kawahara leg // PARATYPE / Trox / Sabulosus / Subsp. fujiokai / OCHI, 2000’.
Current status.
Valid species.
Trox setifer horiguchii
Ochi & Kawahara
CB4CF742-61BA-501D-90E6-D88EB3DCFBCF
Trox setifer horiguchii Ochi & Kawahara, 2002: 54−55.
Note.
The following paratype is deposited in RIEB (ex coll. T. Ochi):
Paratype.
1 ex.: 1♂ ‘MEBORO-DAM / KAMIAGATA. T. / IS. TSUSHIMA / NAGASAKI. J. P / 25–V. 2002 / Y and M. KAWAHARA // 対県= tsu-ken上県町= kamiagata-chô / 目保呂ダム= meborodamu / 25. V. 2002 / Y and M. Kawahara // ♂ // PARATYPE / Trox / setifer / subsp. horiguchii / OCHI and KAWAHARA / 2002’.
Current status.
Trox horiguchii Ochi and Kawahara, 2002, see Pittino (2006).
Acknowledgments
We are grateful to Toshiaki Aoki (RIEB), Shûhei Yamagushi (RIEB), Reiichiro Katsuyama (RIEB), Shûhei Nomura (NMNS), and Kônosuke Hoshino (NMSM) for their help in the type material survey. Our thanks go to Hao Xu (Mianyang Normal University) for providing the information about Miyake’s type material. We express our sincere gratitude to Carsten Zorn (Gnoien) for reviewing the manuscript.
Citation
Kaneko N, Wada K (2020) Catalogue of the type material of Scarabaeoidea (Coleoptera) deposited in the Research Institute of Evolutionary Biology, Tokyo, Japan. ZooKeys 958: 35–89. https://doi.org/10.3897/zookeys.958.52799
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