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[Preprint]. 2022 Jan 11:2020.09.16.20190694. Originally published 2020 Sep 18. [Version 2] doi: 10.1101/2020.09.16.20190694

KIM-1/TIM-1 is a Receptor for SARS-CoV-2 in Lung and Kidney

Yutaro Mori, Corby Fink, Takaharu Ichimura, Keisuke Sako, Makiko Mori, Nathan N Lee, Philipp Aschauer, Krishna M Padmanabha Das, SoonGweon Hong, Minsun Song, Robert F Padera, Astrid Weins, Luke P Lee, Mahmoud L Nasr, Gregory A Dekaban, Jimmy D Dikeakos, Joseph V Bonventre
PMCID: PMC7523142  PMID: 32995803

Abstract

SARS-CoV-2 precipitates respiratory distress by infection of airway epithelial cells and is often accompanied by acute kidney injury. We report that Kidney Injury Molecule-1/T cell immunoglobulin mucin domain 1 (KIM-1/TIM-1) is expressed in lung and kidney epithelial cells in COVID-19 patients and is a receptor for SARS-CoV-2. Human and mouse lung and kidney epithelial cells express KIM-1 and endocytose nanoparticles displaying the SARS-CoV-2 spike protein (virosomes). Uptake was inhibited by anti-KIM-1 antibodies and TW-37, a newly discovered inhibitor of KIM-1-mediated endocytosis. Enhanced KIM-1 expression by human kidney tubuloids increased uptake of virosomes. KIM-1 binds to the SARS-CoV-2 Spike protein in vitro . KIM-1 expressing cells, not expressing angiotensin-converting enzyme 2 (ACE2), are permissive to SARS-CoV-2 infection. Thus, KIM-1 is an alternative receptor to ACE2 for SARS-CoV-2. KIM-1 targeted therapeutics may prevent and/or treat COVID-19.

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