1Department of Medicine, University of Cambridge, Cambridge, United Kingdom
2Present Address: Faculty of Biology, Medicine and Health, University of Manchester, Michael Smith Building, Oxford Road, Manchester, M13 9PT UK
3Wellcome Centre for Human Genetics, University of Oxford, Oxford, United Kingdom
4Department of Pathology, University of Cambridge, Cambridge, United Kingdom
5Cambridge Institute of Therapeutic Immunology and Infectious Disease, Jeffrey Cheah Biomedical Centre, University of Cambridge, Cambridge, United Kingdom
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