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[Preprint]. 2021 Jan 3:2020.12.31.424987. [Version 1] doi: 10.1101/2020.12.31.424987

Paired heavy and light chain signatures contribute to potent SARS-CoV-2 neutralization in public antibody responses

Bailey B Banach, Gabriele Cerutti, Ahmed S Fahad, Chen-Hsiang Shen, Matheus Oliveira de Souza, Phinikoula S Katsamba, Yaroslav Tsybovsky, Pengfei Wang, Manoj S Nair, Yaoxing Huang, Irene M Francino Urdániz, Paul J Steiner, Matias Gutiérrez-González, Lihong Liu, Sheila N López Acevedo, Alexandra Nazzari, Jacy R Wolfe, Yang Luo, Adam S Olia, I-Ting Teng, Jian Yu, Tongqing Zhou, Eswar R Reddem, Jude Bimela, Xiaoli Pan, Bharat Madan, Amy D Laflin, Rajani Nimrania, Kwon-Tung Yuen, Timothy A Whitehead, David D Ho, Peter D Kwong, Lawrence Shapiro, Brandon J DeKosky
PMCID: PMC7805438  PMID: 33442681

Summary

Understanding protective mechanisms of antibody recognition can inform vaccine and therapeutic strategies against SARS-CoV-2. We discovered a new antibody, 910-30, that targets the SARS-CoV-2 ACE2 receptor binding site as a member of a public antibody response encoded by IGHV3-53/IGHV3-66 genes. We performed sequence and structural analyses to explore how antibody features correlate with SARS-CoV-2 neutralization. Cryo-EM structures of 910-30 bound to the SARS-CoV-2 spike trimer revealed its binding interactions and ability to disassemble spike. Despite heavy chain sequence similarity, biophysical analyses of IGHV3-53/3-66 antibodies highlighted the importance of native heavy:light pairings for ACE2 binding competition and for SARS-CoV-2 neutralization. We defined paired heavy:light sequence signatures and determined antibody precursor prevalence to be ~1 in 44,000 human B cells, consistent with public antibody identification in several convalescent COVID-19 patients. These data reveal key structural and functional neutralization features in the IGHV3-53/3-66 public antibody class to accelerate antibody-based medical interventions against SARS-CoV-2.

Highlights

  • A molecular study of IGHV3-53/3-66 public antibody responses reveals critical heavy and light chain features for potent neutralization

  • Cryo-EM analyses detail the structure of a novel public antibody class member, antibody 910-30, in complex with SARS-CoV-2 spike trimer

  • Cryo-EM data reveal that 910-30 can both bind assembled trimer and can disassemble the SARS-CoV-2 spike

  • Sequence-structure-function signatures defined for IGHV3-53/3-66 class antibodies including both heavy and light chains

  • IGHV3-53/3-66 class precursors have a prevalence of 1:44,000 B cells in healthy human antibody repertoires

Full Text Availability

The license terms selected by the author(s) for this preprint version do not permit archiving in PMC. The full text is available from the preprint server.


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