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. 2021 Jan 13;9:e60287. doi: 10.3897/BDJ.9.e60287

The family Conopidae (Diptera) in Egypt and Saudi Arabia

Magdi El-Hawagry 1,, Ahmed Mostafa Soliman 2,3,, Hathal Mohammed Al Dhafer 2,
PMCID: PMC7817652  PMID: 33519263

Abstract

Background

The present study is one in a series of planned studies aiming to catalogue the whole order Diptera in both Egypt and Saudi Arabia.

New information

All known Egyptian and Saudi Arabian conopid taxa are systematically catalogued in the present study. Three species are recorded herein for the first time from Saudi Arabia: Conops (Asiconops) elegans Meigen, 1804 and Thecophora atra (Fabricius, 1775) (Al-Baha region, south-western of Saudi Arabia), and Conops (Conops) quadrifasciatus De Geer, 1776 (Tabuk region, north-western of Saudi Arabia). Physocephala variegata (Meigen, 1924) is also recorded for the first time from Gebel Elba, the south-eastern triangle of Egypt. Considering that Gebel Elba in Egypt and Al-Baha in Saudi Arabia are affiliated to the Afrotropical Region, this is the first time Physocephala variegata and Thecophora atra have been recorded from the Afrotropical Region. An updated taxonomy, world and local distributions, dates of collection and some coloured photographs are provided.

Keywords: thick-headed flies, local distribution, dates of collection, new records

Introduction

The Conopidae, commonly known as thick-headed flies, are an interesting brachycerous family of usually large-sized wasp-like flies. Many conopids are most frequently found at flowers feeding on nectar using their long proboscis and considered as important pollinators. The majority of conopid larvae are obligatory endoparasitoids of adult Hymenoptera, such as bees and wasps. Adult conopid females aggressively intercept their hymenopterous hosts in the field to deposit their eggs (Freeman (1966), Gibson and Skevington (2013), Pape et al. (2011), Stuke (2017), Azmy et al. (2016)). The family Conopidae is represented by 808 valid species assigned to 57 genera worldwide (Stuke 2017). Conopids are found in all parts of the world, except Antarctica and the Pacific Islands (Gibson et al. 2013).

Egypt and Saudi Arabia are two neighbouring Middle Eastern countries, separated by the Gulf of Aqaba and the Red Sea (Fig. 1). Both countries are located at the junction of the Palaearctic and the Afrotropical Regions and the faunal affiliation of them is mainly Palaearctic, except Gebel Elba, the south-eastern triangle of Egypt and the south-western part of Saudi Arabia, south to the Tropic of Cancer, which are considered as Afrotropical (Hölzel 1998, Wallace 1876, El-Hawagry and Gilbert 2014, El-Hawagry et al. 2020).

Figure 1.

Figure 1.

A satellite map of Egypt and Saudi Arabia.

Some previous studies have been carried out in Egypt on the taxonomy and faunistics of the family Conopidae, including Kröber (1924), Kröber (1925), Kröber (1927), Kröber (1929), Steyskal and El-Bialy (1967), Mohammed and Negm (1991), Azmy (2016) and Azmy et al. (2016), in addition to some miscellaneous studies which described some new species from Egypt as Kröber (1915a), Wiedemann (1830), Macquart (1835) and Rondani (1850). On the other hand, no previous faunistic or taxonomic studies on the family Conopidae have been carried out before in Saudi Arabia and only a few species have been described or recorded there amid some miscellaneous studies, such as Macquart (1851), Kröber (1915a), Zimina (1963), Walker and Pittaway (1987), Abu-Zoherah et al. (1993), Stuke and Clements (2008), El-Hawagry et al. (2016) and El-Hawagry et al. (2017).

The abundance and diversity of conopid species in both Egypt and Saudi Arabia seem to be low (17 species in Egypt and 12 species in Saudi Arabia) comparable with the total number of species known from the Palaearctic Region (173 species) and from the Afrotropical Region (187 species) (Stuke 2017). However, this should be taken with caution, since the family Conopidae seems to lack sampling efforts, especially in Saudi Arabia. This is most likely due to the fact that documentation of biological diversity in Saudi Arabia began in the second half of the 1960s and no systematic studies on the Conopidae have been previously conducted in this country. We think that the faunistic data of these flies in Egypt and Saudi Arabia is still scanty and more efforts would be highly required.

The taxonomy of Conopidae in both Egypt and Saudi Arabia is updated in the present study. Twenty species assigned to five genera and three subfamilies (Conopinae, Myopinae and Zodioninae) are treated (Table 1). Three species, Conops (Asiconops) elegans Meigen, 1804, Conops (Conops) quadrifasciatus De Geer, 1776 and Thecophora atra (Fabricius, 1775), are recorded for the first time from Saudi Arabia.

Table 1.

Conopid species treated in the present study as recorded from Egypt and Saudi Arabia (* = recorded, - = not recorded).

Species Egypt Saudi Arabia
Subfamily CONOPINAE
Tribe CONOPINI
Conops (Asiconops) elegans Meigen, 1804 * * (new record)
Conops (Asiconops) flavifrons Meigen, 1804 * -
Conops (Asiconops) nubeculipennis Bezzi, 1901 * *
Conops (Conops) flavicauda (Bigot, 1880) * -
Conops (Conops) quadrifasciatus De Geer, 1776 - * (new record)
Conops (Conops) rufiventris Macquart, 1849 * -
Conops (Conops) tomentosus Kröber, 1916 - *
Tribe PHYSOCEPHALINI
Physocephala antiqua (Wiedemann, 1830) * *
Physocephala chrysorrhoea (Meigen, 1824) * *
Physocephala pusilla (Meigen, 1804) * *
Physocephala variegata (Meigen, 1824) * *
Physocephala vittata (Fabricius, 1794) * -
Subfamily MYOPINAE
Tribe MYOPINI
Myopa dorsalis Fabricius, 1794 * -
Myopa picta Panzer, 1798 * -
Myopa stigma Meigen, 1824 - *
Myopa testacea (Linnaeus, 1767) * -
Tribe THECOPHORINI
Thecophora atra (Fabricius, 1775) * * (new record)
Thecophora fulvipes (Robineau-Desvoidy, 1830) * *
Subfamily ZODIONINAE
Zodion cinereum (Fabricius, 1794) * *
Zodion erythrurum Rondani, 1865 * -

Physocephala variegata (Meigen) is recorded for the first time from Gebel Elba, the south-eastern triangle of Egypt and Thecophora atra (Fabricius) is recorded for the first time from the south-western part of Saudi Arabia. Considering that Gebel Elba in Egypt and the south-western part in Saudi Arabia including Al-Baha Region are affiliated to the Afrotropical Region, this is the first time that these two species from the Afrotropical Region have been recorded.

Walker (1871) described Conops auratus from “Hor Tamanib” which belonged to Egypt at the time of description; however, it lies now in Sudan. Steyskal and El-Bialy (1967) inaccurately listed this species as an Egyptian species, based mainly on the misleading type locality. Thus, this species is excluded from the Egyptian list in the present study.

This study is one in a series of taxonomic studies on different Egyptian and Saudi Arabian dipteran taxa aiming to catalogue the whole order Diptera in the two countries.

Materials and methods

Data sources. Data of the present study are obtained from three main sources: 1. Specimens preserved in the Egyptian and Saudi Arabian insect collections and museums, namely: Efflatoun Bey’s collection, Cairo University, Egypt (EFC); Collection of Entomological Society of Egypt (ESEC); King Saud University Museum of Arthropods, Saudi Arabia (KSMA) and Personal collection M. El-Hawagry (MSHC). 2. Material collected by the authors and their co-workers. 3. Previous studies on the Egyptian and Saudi Arabian Conopidae. The majority of specimens preserved in EFC and ESEC were determined by Dr. Otto Kröber. However, all identifications have been rechecked by the first author using Kröber (1927), Stuke and Clements (2008), Azmy (2016) and Azmy et al. (2016). Photos of Thecophora atra have been checked by Dr. Jens-Hermann Stuke (personal communication).

Study area. Ecologists divide Egypt into eight ecological zones: the Coastal Strip (= Mediterranean Coastal Strip), Lower Nile Valley and Delta, Upper Nile Valley, Fayoum, Eastern Desert, Western Desert, Sinai and Gebel Elba (El-Hawagry and Gilbert 2014). These ecological zones are used in the present study in the sections of local distribution. However, Saudi Arabia is not divided into ecological zones by ecologists, so the administrative divisions (also called regions, provinces or emirates) are used instead, namely: Makkah, Riyadh, Eastern Province, Asir, Jazan, Al-Madinah, Al-Qaseem, Tabuk, Hail, Najran, Al-Jawf, Al-Baha and Northern Frontier (Fig. 1) (El-Hawagry et al. 2020).

Classification. The classification and nomenclature in the present study basically follows that used in Stuke (2017), in addition to the tribal arrangement suggested by Gibson and Skevington (2013). Full lists of synonyms for genera and species are not given in the present study as they are already listed in world and regional catalogues; however, we listed only the synonyms of species that have been mentioned in previous Egyptian and Saudi Arabian literature.

World distribution. Stuke (2017) has collected all sources of world distribution of species and it is used here as our basic source. The additional sources, if any, are given between parentheses after each record.

Local distribution and dates of collection. Localities within each Egyptian ecological zone or each Saudi Arabian administrative region are arranged alphabetically and written after a colon following each zone or region, followed by the dates of collection between parentheses. Sources for the local distribution of each species in each country are given between square brackets at the end of the section. Coordinates of Egyptian and Saudi Arabian localities of the family Conopidae are listed (Table 2). Local distribution maps of species based on all available records are given, using SimpleMappr (Shorthouse 2010).

Table 2.

A gazetteer of some Egyptian and Saudi Arabian localities of the family Conopidae.

Country Locality Governorate Ecological zone or Region Latitude (N) / Longitude (E)
Egypt Abu-Kir Alexandria Coastal Strip 31.22429, 29.94664
Abu-Rawash Giza Lower Nile Valley and Delta 30.0438, 31.0929
Abu-Sueir Ismailia Eastern Desert 30.5766, 32.1076
Amria Alexandria Coastal Strip 31.0037, 29.7983
Ashmoun Menofiya Lower Nile Valley and Delta 30.29414, 30.96941
Aswan, south Aswan Upper Nile Valley 22.6651, 31.7943
Balteem Kafr Al-Sheikh Coastal Strip 31.53811, 31.19606
Barrage Qalyoubia Lower Nile Valley and Delta 30.1835, 31.1107
Behig Alexandria Coastal Strip 30.931, 29.5988
Burg Alexandria Coastal Strip 30.9081, 29.5464
Burgash Giza Lower Nile Valley and Delta 30.1636, 31.0379
Cairo Cairo Lower Nile Valley and Delta 30.0861, 31.2856
Cairo-Suez Road Cairo Eastern Desert 30.0849, 32.0542
Dahshour Giza Lower Nile Valley and Delta 29.7539, 31.2431
Dakhla New Valley Western Desert 25.5, 29.1667
Dekhela Alexandria Coastal Strip 31.12098, 29.81563
Edfina Al-Behaira Lower Nile Valley and Delta 31.2922, 30.51599
Ein Gedeirat North Sinai Sinai 30.65, 34.4333
Ein Moussa South Sinai Sinai 29.8667, 32.65
El-Arish North Sinai Sinai 31.1442, 33.8056
El-Gebel El-Asfar Qalyoubia Lower Nile Valley and Delta 30.201416, 31.356162
El-Katta Giza Lower Nile Valley and Delta 30.2191, 30.9678
El-Magadlah Giza Lower Nile Valley and Delta 30.03986, 31.1046
El-Mallah East Cairo Eastern Desert 30.8167, 32.1
El-Wasfia Ismailia Eastern Desert 30.58076, 32.17235
Esna Qena Upper Nile Valley 25.2919, 32.5532
Ezbet Nakhl Qalyoubia Lower Nile Valley and Delta 31.1111, 32.1625
Fayed Ismailia Eastern Desert 30.32382, 32.30079
Fayoum Fayoum Fayoum 29.32061, 30.818
Gebel Elba Red Sea Gebel Elba 22.2008, 36.3331
Geneifa Suez Eastern Desert 30.1516, 32.429
Gezira Cairo Lower Nile Valley and Delta 30.04596, 31.22435
Girza Fayoum Fayoum 29.49968, 31.0738
Giza Giza Lower Nile Valley and Delta 30.0135, 31.21127
Halaib Red Sea Eastern Desert 22.2203, 36.6427
Hawamdia Giza Lower Nile Valley and Delta 29.9092, 31.2615
Helwan Cairo Lower Nile Valley and Delta 29.85, 31.3333
Ismailia Ismailia Eastern Desert 30.59428, 32.26026
Kafr Ghatati Giza Lower Nile Valley and Delta 30.00415, 31.12157
Kafr Hakim Giza Lower Nile Valley and Delta 30.0808, 31.1164
Kerdassa Giza Lower Nile Valley and Delta 30.0297, 31.1061
Kom Ombo Aswan Upper Nile Valley 24.4761, 32.9483
Kom Osheem Fayoum Fayoum 29.5564, 30.8869
Kubba Cairo Lower Nile Valley and Delta 30.0876, 31.2854
Maadi Cairo Lower Nile Valley and Delta 29.95772, 31.25054
Madinet El-Sadat Menofiya Western Desert 30.35108, 30.51109
Madinet Badr Cairo Lower Nile Valley and Delta 30.153, 31.7103
Mansouriah Giza Lower Nile Valley and Delta 30.1236, 31.0725
Mariout Alexandria Coastal Strip 31.0172, 29.76
Mazghouna Giza Lower Nile Valley and Delta 29.7457, 31.2633
Mersa Matrouh Matrouh Coastal Strip 29.5696, 26.4194
Orman Giza Lower Nile Valley and Delta 30.02912, 31.21271
Pyramids Giza Lower Nile Valley and Delta 29.9816, 31.1337
Rafah North Sinai Sinai 31.2667, 34.2333
Salloum Matrouh Coastal Strip 31.5888, 25.1026
Saqqara Giza Lower Nile Valley and Delta 29.84955, 31.21641
Sennouris Fayoum Fayoum 29.40662, 30.86602
Sentris Menofiya Lower Nile Valley and Delta 30.31122, 31.0577
Siwa Oasis Matrouh Western Desert 29.20427, 25.51928
Suez Suez Eastern Desert 29.95278, 32.56582
Tamiya Fayoum Fayoum 29.475479, 30.95228
Turah Cairo Lower Nile Valley and Delta 29.9467, 31.2728
W. Asal Red Sea Eastern Desert 25.9497, 34.383
W. Dar El-Maskhara Cairo Eastern Desert 29.7833, 31.4167
W. Digla Cairo Eastern Desert 29.9578, 31.3348
W. El-Arbaein South Sinai Sinai 28.5469, 33.953
W. El-Daiqa North Sinai Sinai 30.8278, 34.1401
W. El-Lega South Sinai Sinai 28.5469, 33.953
W. El-Natroun Al-Behaira Western Desert 30.3814, 30.3441
W. El-Rahba South Sinai Sinai 28.82, 33.6419
W. Firan South Sinai Sinai 28.7092, 33.3222
W. Garawi Cairo Eastern Desert 29.7833, 31.3167
W. Gedeirat North Sinai Sinai 30.6511, 34.4321
W. Gharagid Cairo Eastern Desert 28.955, 31.4822
W. Hoff Cairo Eastern Desert 29.8821, 31.311
W. Ibib Red Sea Gebel Elba 22.83333, 35.76667
W. Isla South Sinai Sinai 28.201, 34.244
W. Morrah Cairo Eastern Desert 22.35, 33.75
W. Rishrash Giza Eastern Desert 29.4642, 31.3672
W. Um Elek Cairo Eastern Desert 29.8833, 31.5167
W. Zohleiga Cairo Eastern Desert 26.1333, 33.75
Saudi Arabia Abu-Arish Jazan Jazan 16.9821, 42.8389
Al-Aqiq Al-Aqiq Al-Baha 20.2866, 41.6312
Al-Mekhwa Al-Mekhwa Al-Baha 19.8429, 41.3115
Ghabet Shahba Al-Baha Al-Baha 20.02723, 41.28565
Jabal Shada al-A’la Nature Reserve Al-Mekhwa Al-Baha 19.8388, 41.3101
Jazan Jazan Jazan 16.90245, 42.5705
Jeddah Jeddah Makka Al-Mukarramah 21.5922, 39.2631
Marabah Marabah Asir 18.0426, 42.3739
Najran Najran Najran 17.4833, 44.1166
Raydah Nature Reserve Abha Asir 18.20525, 42.41011
Tabouk Tabouk Tabouk 28.36661, 36.629747
Unayzah Unayzah Al-Qassim 26.08548, 43.9768

Abbreviations used:

  • AF, Afrotropical.

  • EFC, Efflatoun Bey’s collection, Department of Entomology, Faculty of Science, Cairo University, Egypt.

  • ESEC, Collection of Entomological Society of Egypt, Cairo, Egypt.

  • KSMA, King Saud University Museum of Arthropods, Riyadh, Saudi Arabia.

  • MSHC, Personal collection M. El-Hawagry.

  • OR, Oriental.

  • PA, Palearctic.

Checklists

The catalogue

CONOPIDAE

2886EBB0-F4EB-58D7-8BF6-EAD3A69B6FFE

CONOPINAE

A9C02511-9D07-57D5-BD4F-EA81E057285E

CONOPINI

07DBF252-FE1F-5C41-96FD-2D0273F8B1BC

Conops

Linnaeus

BC310CE2-5316-5909-895B-815792C7DEDB

https://www.gbif.org/species/1444664

  • Conops Conops Linnaeus, 1758: 604. Type species: Conops flavipes Linnaeus, by subsequent designation of Curtis (1831).

Asiconops

Chen

5168C48E-6A17-5C60-AC1A-7862169A67D9

  • Asiconops Asiconops Chen, 1939: 171. Type species: Conops aureomaculatus Kröber, by original designation.

Conops (Asiconops) elegans

Meigen, 1804

EA1D392D-75C8-5867-AD62-3DD8AD852EAA

https://www.gbif.org/species/5076777

  • Conops elegans Meigen, 1804: 275. Type locality: France (Marseille).

  • Conops elegans var. fuscipennis Macquart 1849: 472. Type locality: Algeria.

  • Conops algirus Macquart 1849: 473. Type locality: Algeria (environs du Cercle de Lacalle).

  • Conopaejus aegyptiacus Rondani, 1850: 167. Type locality: Egypt.

  • Conops fuscanipennis Bigot, 1887: 32. Type locality: Italy (Sicily).

  • Conops elegans var. minutus Kröber, 1916: 48. Type locality: Not given (Cyprus, Limassol in Kröber 1939).

Materials
  1. Type status: Other material. Occurrence: recordedBy: Aly; sex: 2 males, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Fayid; decimalLatitude: 30.323823; decimalLongitude: 32.30079; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 03-16-1959; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: W. Garawi; decimalLatitude: 29.7833; decimalLongitude: 31.3167; Identification: identifiedBy: M.El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-06-1927; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: Shafik & Str.; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Girza; decimalLatitude: 29.499684; decimalLongitude: 31.07380; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-16-1952; Record Level: institutionCode: EFC

  4. Type status: Other material. Occurrence: recordedBy: Str. & Sh.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Tameyah; decimalLatitude: 29.475479; decimalLongitude: 30.95228; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 07-25-1948; Record Level: institutionCode: EFC

  5. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.0929; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-17-1925; Record Level: institutionCode: EFC

  6. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 2 males; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.0929; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 09-16-1925; Record Level: institutionCode: EFC

  7. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.0929; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 10-13-1926; Record Level: institutionCode: EFC

  8. Type status: Other material. Occurrence: recordedBy: Mistikawi; sex: 2 males, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Edfina; decimalLatitude: 31.29252; decimalLongitude: 30.51322; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 06-08-1914; Record Level: institutionCode: EFC

  9. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Ezbet-Naghl; decimalLatitude: 31.1111; decimalLongitude: 32.1625; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 04-13-1922; Record Level: institutionCode: EFC

  10. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Ezbet-Naghl; decimalLatitude: 32.1111; decimalLongitude: 32.1625; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 06-06-1926; Record Level: institutionCode: EFC

  11. Type status: Other material. Occurrence: recordedBy: Alfieri; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Ghezirah; decimalLatitude: 30.012106; decimalLongitude: 31.21200; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-20-1920; Record Level: institutionCode: EFC

  12. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 04-21-1934; Record Level: institutionCode: EFC

  13. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 04-28-1934; Record Level: institutionCode: EFC

  14. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-26-1934; Record Level: institutionCode: EFC

  15. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kafr Hakim; decimalLatitude: 30.0808; decimalLongitude: 31.1164; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 10-17-1925; Record Level: institutionCode: EFC

  16. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kafr Hakim; decimalLatitude: 30.0808; decimalLongitude: 31.1164; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 06-08-1927; Record Level: institutionCode: EFC

  17. Type status: Other material. Occurrence: recordedBy: L.H.G; sex: 2 males; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Maadi; decimalLatitude: 29.957721; decimalLongitude: 31.25054; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 06-20-12; Record Level: institutionCode: EFC

  18. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kerdassa; decimalLatitude: 30.0297; decimalLongitude: 31.1061; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 06-10-1925; Record Level: institutionCode: EFC

  19. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kerdassa; decimalLatitude: 30.0297; decimalLongitude: 31.1061; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 10-09-1926; Record Level: institutionCode: EFC

  20. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kerdassa; decimalLatitude: 30.0297; decimalLongitude: 31.1061; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 10-30-1926; Record Level: institutionCode: EFC

  21. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kerdassa; decimalLatitude: 30.0297; decimalLongitude: 31.1061; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-07-1927; Record Level: institutionCode: EFC

  22. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Mansouriah; decimalLatitude: 30.1236; decimalLongitude: 31.0725; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 08-22-1925; Record Level: institutionCode: EFC

  23. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Mansouriah; decimalLatitude: 30.1236; decimalLongitude: 31.0725; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-12-1926; Record Level: institutionCode: EFC

  24. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Mansouriah; decimalLatitude: 30.1236; decimalLongitude: 31.0725; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-12-1926; Record Level: institutionCode: EFC

  25. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Mansouriah; decimalLatitude: 30.1236; decimalLongitude: 31.0725; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 04-26-1930; Record Level: institutionCode: EFC

  26. Type status: Other material. Occurrence: recordedBy: Shafik; sex: 2 females; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Rafah; decimalLatitude: 31.2667; decimalLongitude: 34.2333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-17-1951; Record Level: institutionCode: EFC

  27. Type status: Other material. Occurrence: sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Sakkara; decimalLatitude: 29.849552; decimalLongitude: 31.2164; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 06-08-1917; Record Level: institutionCode: EFC

  28. Type status: Other material. Occurrence: recordedBy: El-Hawagry; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076777; scientificName: Conops elegans; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; locality: Al-Mekhwa; decimalLatitude: 19.8429; decimalLongitude: 41.3115; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 03-17-1912; Record Level: institutionCode: MSHC

Distribution

AF: Democratic Republic of the Congo, Ethiopia, Gabon, Guinea, Guinea-Bissau, Malawi, Mozambique, Namibia, Nigeria, Saudi Arabia [as “South-western part”] (first record), South Africa, Tanzania, Yemen. PA: Algeria, Bulgaria, Cyprus, Egypt, France, Israel, Italy, Morocco, Romania, Spain, Syria, Tunisia, Turkey.

Local distribution and dates of collection (Fig. 2): EGYPT: Coastal Strip: Balteem, Burg (March to August); Eastern Desert: Abu-Sueir, El-Mallah (= Mallah East), Fayed, Suez, Wadi Garawi, Wadi Hoff, Wadi Morrah, Wadi Rishrash, Wadi Um-Elek, Wadi Zohleiga (March to August); Fayoum: Fayoum City, Kom Osheem, Tamiya (May to November); Lower Nile Valley and Delta: Abu-Rawash, Barrage, Edfina, El-Gebel El-Asfar, El-Magadlah, Ezbet El-Nakhl, Gezira, Giza, Helwan, Kafr Ghatati, Kafr Hakim, Kerdassa, Maadi, Mansouriah, Orman, Pyramids, Sentris (throughout the year); Sinai: Rafah (August); Western Desert: Saqqara, Madinet El-Sadat, Wadi El-Natroun (June and October) [Sources: Kröber (1927), Azmy (2016) and museum material in EFC and ESEC]. SAUDI ARABIA: Al-Baha: Al-Mekhwa (March) [Sources: collected material].

Figure 2.

Figure 2.

Local distribution map of Conops (Asiconops) elegans Meigen and Conops (Asiconops) nubeculipennis Bezzi in Egypt and Saudi Arabia.

Notes

This species (Fig. 12a) is recorded herein for the first time from Saudi Arabia.

Habitus.

Figure 12a.

Figure 12a.

Conops (Asiconops) elegans Meigen male, lateral

Figure 12b.

Figure 12b.

Thecophora atra (Fabricius) female, lateral

Conops (Asiconops) flavifrons

Meigen, 1804

D4A808AA-1F11-533D-B289-291E663950AC

https://www.gbif.org/species/5076765

  • Conops flavifrons Meigen, 1804: 281. Type locality: France (Lyon).

  • Physocephala pugioniformis Becker, 1913: 611. Type locality: Iran.

  • Conops minor Becker, 1922: 201. Type locality: Russia, Italy, Hungary & Greece.

  • Conops kroeberi var. immaculata Paramonov, 1927: 4. Type locality: Armenia.

  • Conops kroeberi Paramonov, 1927: 4. Type locality: Armenia.

Distribution

AF: Yemen. PA: Albania, Armenia, Austria, ?Belgium, Bulgaria, ?China, Croatia, Czech Republic, Egypt, France, ?Germany, Greece, Hungary, Iran, Italy, Poland, Russia, Serbia, Spain, Syria, Tunisia, Turkey.

Notes

This species was listed by Steyskal and El-Bialy (1967), Chvála and Smith (1988), Azmy (2016) as recorded from Egypt, but these records could not be verified and require confirmation as no specimens have been collected or preserved in Egyptian museums.

Conops (Asiconops) nubeculipennis

Bezzi, 1901

8C73F907-C774-50D8-964C-66C249D68A06

https://www.gbif.org/species/5076815

  • Conops nubeculipennis Bezzi, 1901: 21. Type locality: Eritrea.

  • Conops atrogonius Séguy 1930: 134. Type locality: Morocco.

Materials
  1. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076815; scientificName: Conops nubeculipennis; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; stateProvince: Al-Qassim; locality: Unayzah; decimalLatitude: 26.085478; decimalLongitude: 43.9768123; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Malaise trap; eventDate: 05-01-2018; Record Level: institutionCode: KSMA

  2. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076815; scientificName: Conops nubeculipennis; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; stateProvince: Asir; locality: Garf Raydah Nature Reserve; decimalLatitude: 18.194917; decimalLongitude: 42.396967; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Malaise trap; eventDate: 05-08-2015; Record Level: institutionCode: KSMA

Distribution

AF: Cameroon, Ethiopia, Guinea-Bissau, Mozambique, Nigeria, Saudi Arabia [as “South-western part”] (El-Hawagry et al. 2017), Yemen, Zambia. PA: Egypt, Israel, Morocco.

Local distribution and dates of collection (Fig. 2): EGYPT: Upper Nile Valley: South Aswan (as “Nubien”) [Source: Kröber (1915b)]. SAUDI ARABIA: Al-Qassim: Unayzah (May); Asir: Raydah Nature Reserve (May and June); South-western part of Saudi Arabia including Jazan and Najran (September and October) [Sources: Walker and Pittaway (1987), El-Hawagry et al. (2017)) and collected material].

Notes

This species was listed by Kröber (1915b), Steyskal and El-Bialy (1967), Azmy (2016), Stuke (2017) as recorded from Egypt, but no specimens have been collected or preserved in Egyptian museums.

Conops

Linnaeus

95513370-78CD-5EE7-B198-0A8838230483

Conops (Conops) flavicauda

(Bigot, 1880)

C37A3C82-52E9-5907-A837-5A5656702F6D

https://www.gbif.org/species/5076720

  • Sphixosoma flavicauda Bigot, 1880: 149. Type locality: Iran (Northern Iran and Caucasus).

  • Conops euzonatus Bigot, 1887: 33. Type locality: Caucasus Mountains.

  • Conops superbus Röder, 1889: 6. Type locality: Lebanon (Beirut) [as “Beyrut, Syria (Asia minor)”].

  • Conops cypris Janssens, 1955: 2. Type locality: Cyprus (Akrotiri Bay, Yermdoyia & Erimi).

Distribution

AF: Uganda. PA: Afghanistan, Armenia, Cyprus, Egypt, Greece, Iran, Israel, Lebanon, Syria, Turkey.

Local distribution and dates of collection (Fig. 3): EGYPT: Sinai: Wadi Isla (August) [Sources: Zalat et al. 2008].

Figure 3.

Figure 3.

Local distribution map of Conops (Conops) flavicauda (Bigot) and Conops (Conops) quadrifasciatus De Geer in Egypt and Saudi Arabia.

Conops (Conops) quadrifasciatus

De Geer, 1776

527F63CF-C9C2-5220-AFC6-5EBFE9727F0E

https://www.gbif.org/species/5076797

  • Conops De Geer, 1776: 261. Type locality: Presumably Sweden.

Materials
  1. Type status: Other material. Occurrence: recordedBy: El-Hawagry; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076797; scientificName: Conops quadrifasciatus; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; stateProvince: Tabouk; locality: Tabouk; decimalLatitude: 28.36661; decimalLongitude: 36.629747; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Malaise trap; eventDate: 10-27-2014; Record Level: institutionCode: MSHC

Distribution

PA: Andorra, Austria, Belarus, Belgium, Bosnia and Herzegovina, Bulgaria, Croatia, Czech Republic, Denmark, Finland, France, Germany, Great Britain, Hungary, Iran, Ireland, Italy, Kazakhstan, Latvia, Lithuania, Netherlands, Norway, Poland, Romania, Russia, Saudi Arabia (first record), Slovakia, Spain, Sweden, Switzerland, Turkey, Ukraine.

Local distribution and dates of collection (Fig. 3): SAUDI ARABIA: Tabouk (October) [Source: collected material].

Notes

This species is recorded herein for the first time from Saudi Arabia.

Conops (Conops) rufiventris

Macquart, 1849

0BBA3D38-99F8-55E7-8D52-4013A9F89E3B

https://www.gbif.org/species/5076705

  • Conops rufiventris Macquart, 1849: 474. Type locality: Algeria (Constantine).

Materials
  1. Type status: Other material. Occurrence: recordedBy: Willson; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/5076705; scientificName: Conops rufiventris; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Salloum; decimalLatitude: 31.588751; decimalLongitude: 25.102649; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-14-1917; Record Level: institutionCode: EFC

Distribution

Algeria, Egypt, Tunisia.

Local distribution and dates of collection (Fig. 4): EGYPT: Coastal Strip: Salloum (April and May) [Azmy (2016) and museum material in EFC].

Figure 4.

Figure 4.

Local distribution map of Conops (Conops) rufiventris Macquart and Conops (Conops) tomentosus Kröber in Egypt and Saudi Arabia.

Conops (Conops) tomentosus

Kröber, 1916

C35CEF87-DDD3-53A9-BB2B-012315BB75EC

https://www.gbif.org/species/5076849

  • Conops tomentosus Kröber, 1916: 45. Type locality: Turkmenistan.

Distribution

AF: Saudi Arabia [as “South-western part”]. PA: Iraq, Tajikistan, Turkey, Turkmenistan.

Local distribution and dates of collection (Fig. 4): SAUDI ARABIA: Jazan: Abu-Arish (March) [Source: Stuke et al. (2008)].

PHYSOCEPHALINI

0CF0A37A-3709-53B4-8806-EF4E4B18A3A7

Physocephala

Schiner

2ABD2B29-4CB6-56BF-B851-931FE8B51E63

https://www.gbif.org/species/1568868

  • Physocephala Physocephala Schiner, 1861: 138. Type species: Conops rufipes Fabricius, by original designation.

Physocephala antiqua

(Wiedemann, 1830)

F05F8C8C-6F7A-539B-8DE6-96B58ED73222

https://www.gbif.org/species/1569080

  • Conops antiqua Wiedemann, 1830: 239. Type locality: Egypt.

  • Conops arabica Macquart, 1851: 134. Type locality: Saudi Arabia (Jeddah).

  • Physocephala furax Becker, 1913: 612. Type locality: Iran.

  • Physocephala syriaca Kröber, 1915: 72. Type locality: Egypt (Cairo) & Palestinian West Bank (Jericho).

  • Physocephala obscurifacies Kröber, 1924: 67. Type locality: Egypt (Wadi Hoff).

Materials
  1. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Sueir; decimalLatitude: 30.5766; decimalLongitude: 32.107622; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-17-1925; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Dar El-Maskhara; decimalLatitude: 29.7833; decimalLongitude: 31.4167; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 04-11-1927; Record Level: collectionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: El-Mallah; decimalLatitude: 30.8167; decimalLongitude: 32.1000; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 04-29-1927; Record Level: institutionCode: EFC

  4. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: W. Garawi; decimalLatitude: 30.7833; decimalLongitude: 31.3167; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 05-15-1927; Record Level: institutionCode: EFC

  5. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Digla; decimalLatitude: 29.9578; decimalLongitude: 31.3348; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 09-03-1926; Record Level: institutionCode: EFC

  6. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Rishrash; decimalLatitude: 29.4642; decimalLongitude: 31.3672; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 06-17-1932; Record Level: institutionCode: EFC

  7. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.092932; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 08-15-1925; Record Level: institutionCode: EFC

  8. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 2 males; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.092932; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 07-18-1925; Record Level: institutionCode: EFC

  9. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.092932; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 06-06-1925; Record Level: institutionCode: EFC

  10. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 04-23-1935; Record Level: institutionCode: EFC

  11. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 2 males; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kerdassa; decimalLatitude: 30.0297; decimalLongitude: 31.1061; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 04-17-1926; Record Level: institutionCode: EFC

  12. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 2 males; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Mansouriah; decimalLatitude: 30.1236; decimalLongitude: 31.07246; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 09-02-1925; Record Level: institutionCode: EFC

  13. Type status: Other material. Occurrence: recordedBy: El-Hawagry; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; stateProvince: Al-Baha; locality: Al-Mekhwa; decimalLatitude: 19.8429; decimalLongitude: 41.3115; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Sweeping; eventDate: 03-17-2012; Record Level: institutionCode: MSHC

  14. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569080; scientificName: Physocephala antiqua; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; stateProvince: Jazan; locality: Jazan; decimalLatitude: 16.902448; decimalLongitude: 42.570516; Identification: identifiedBy: M. El-Hawagry; dateIdentified: May 2020; Event: samplingProtocol: Malaise trap; eventDate: 05-12-2019; Record Level: institutionCode: KSMA

Distribution

AF: Egypt [as “Gebel Elba”], Saudi Arabia [as “South-western part”], United Arab Emirates, Yemen. PA: Afghanistan, Algeria, China, Cyprus, Egypt, Greece, Iran, Iraq, Israel, Italy, Jordan, Kazakhstan, Kyrgyzstan, Mongolia, Palestinian Territories, Russia, Saudi Arabia (Jeddah), Syria, Tunisia, Turkey, Turkmenistan.

Local distribution and dates of collection (Fig. 5): EGYPT: Coastal Strip: Abu-Kir, Dekhela, Mariout (April to August); Eastern Desert: Abu Sueir, Cairo-Suez Road, El-Mallah, Geneifa, Ismailia, Wadi Asal (30 km S. Kosseir), Wadi Digla, Wadi El-Maskhara, Wadi Garawi, Wadi Gharagid, Wadi Hoff, Wadi Ibtadi, Wadi Rishrash (February to September); Fayoum: Kom Osheem (May, August and November); Gebel Elba: locality unknown (January); Lower Nile Valley and Delta: Abu-Rawash, Barrage, Burgash, Cairo, El-Katta, El-Magadlah, Ezbet El-Nakhl, Helwan, Kafr Hakim, Kerdassa, Maadi, Mansouriah, Pyramids, Turah (April to October); Sinai: Wadi Gedeirat (June and July); Upper Nile Valley: Kom Ombo (January); Western Desert: Dahshour, Dakhla Oasis (Balat, Ewina & Rashda), Saqqara, Siwa Oasis (March to December) [Sources: Kröber (1915a), Kröber (1924), Kröber (1927), Stuke and Clements (2008), Stuke (2014), Azmy (2016) and museum material in EFC and ESEC]. SAUDI ARABIA: Al-Baha: Al-Mekhwa (March); Asir: Marabah (April); Jazan: Abu-Arish Road (Al-Mahdag and Arada), Jazan City (March to May); Makka Al-Mukarramah: Jeddah (date unknown) [Sources: Kröber (1915a), Stuke and Clements (2008) and collected material].

Figure 5.

Figure 5.

Local distribution map of Physocephala antiqua (Wiedemann) in Egypt and Saudi Arabia.

Physocephala chrysorrhoea

(Meigen, 1824)

9F4DA306-0CC6-5601-8A31-9FE15AF9280D

https://www.gbif.org/species/1569093

  • Conops chrysorrhoea Meigen, 1824: 128. Type locality: Austria.

  • Conops pallasi Meigen, 1824: 128. Type locality: Russia.

  • Conops serpylleti Zeller, 1842: 837. Type locality: Germany.

  • Physocephala zarudnyi Becker, 1913: 614. Type locality: Iran.

  • Physocephala truncata var. pseudomaculigera Kröber, 1915: 72. Type locality: Egypt & Tunisia.

Materials
  1. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg; decimalLatitude: 30.9081; decimalLongitude: 29.5464; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-25-1927; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg; decimalLatitude: 30.9081; decimalLongitude: 29.5464; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 05-10-1927; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: Selim; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Ein Moussa; decimalLatitude: 29.8667; decimalLongitude: 32.6500; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-07-1928; Record Level: institutionCode: ESEC

  4. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: W. Dar El-Maskhara; decimalLatitude: 29.7833; decimalLongitude: 31.4167; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-21-1927; Record Level: institutionCode: EFC

  5. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: W. Garawi; decimalLatitude: 29.7833; decimalLongitude: 31.3167; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-25-1932; Record Level: institutionCode: EFC

  6. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 2 males; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: W. Morrah; decimalLatitude: 22.3500; decimalLongitude: 33.7500; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-01-1925; Record Level: institutionCode: EFC

  7. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Hoff; decimalLatitude: 29.8820558; decimalLongitude: 31.3109855; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-04-1927; Record Level: institutionCode: EFC

  8. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Um-Elek; decimalLatitude: 29.8833; decimalLongitude: 31.5167; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-12-1924; Record Level: institutionCode: EFC

  9. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Um-Elek; decimalLatitude: 29.8833; decimalLongitude: 31.5167; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-13-1930; Record Level: institutionCode: EFC

  10. Type status: Other material. Occurrence: recordedBy: Shafik & Str.EFC; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Girza; decimalLatitude: 29.499684; decimalLongitude: 31.073799; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-05-1950; Record Level: institutionCode: EFC

  11. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.092932; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 05-05-1926; Record Level: institutionCode: EFC

  12. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.092932; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 04-03-1926; Record Level: institutionCode: EFC

  13. Type status: Other material. Occurrence: recordedBy: Farag; sex: 3 females; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 04-13-1935; Record Level: institutionCode: EFC

  14. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male, 2 females; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 04-28-1934; Record Level: institutionCode: EFC

  15. Type status: Other material. Occurrence: recordedBy: Shafik; sex: 3 males, 3 females; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Ein Gedeirat; decimalLatitude: 30.6500; decimalLongitude: 34.4333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 5/24-29/1938; Record Level: institutionCode: EFC

  16. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 2 males, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569093; scientificName: Physocephala chrysorrhoea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: W. El-Arbaein (Sinai); decimalLatitude: 28.5469; decimalLongitude: 33.9530; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 4/19-27/1939; Record Level: institutionCode: EFC

Distribution

AF: Saudi Arabia [as “South-western part”]. PA: Algeria, Armenia, Austria, Belgium, Bulgaria, China, Croatia, Czech, Egypt, France, Germany, Greece, Hungary, Iran, Italy, Kyrgyzstan, Lithuania, Mongolia, Morocco, Netherlands, Poland, Romania, Russia, Serbia, Slovakia, Spain, Switzerland, Turkey, Turkmenistan, Ukraine.

Local distribution and dates of collection (Fig. 6): EGYPT: Coastal Strip: Burg El-Arab, Mariout (February to May); Eastern Desert: Bir El-Fahm, Ogret El-Sheikh, Wadi El-Mallah, Wadi El-Maskhara, Wadi Garawi, Wadi Hoff, Wadi Morrah, Wadi Um-Elek, Wadi Zohleiga (March and April); Fayoum: Girza, Sennouris (January to March); Gebel Elba: locality unknown (January); Lower Nile Valley and Delta: Abu-Rawash, Barrage, Ezbet El-Nakhl, Hawamdiya, Helwan, Kafr Hakim, Kubba, Maadi, Mansouriah, Mazghouna (February to August); Sinai: Ein Gedeirat, Ein Moussa, Wadi El-Arbaein, Wadi El-Rabba (February to May); Western Desert: Dahshour (April) [Sources: Azmy (2016) and museum material in EFC and ESEC]. SAUDI ARABIA: locality and date unknown [Sources: Stuke (2017)].

Figure 6.

Figure 6.

Local distribution map of Physocephala chrysorrhoea (Meigen) and Physocephala pusilla (Meigen) in Egypt and Saudi Arabia.

Physocephala pusilla

(Meigen, 1804)

6CEC5DCC-417E-50CD-A2FF-8D365781D445

https://www.gbif.org/species/1568897

  • Conops pusilla Meigen, 1824: 280. Type locality: Germany (Frankreich).

  • Conops lacera Meigen, 1824: 130. Type locality: Austria.

  • Conops pumila Macquart, 1835: 26. Type locality: France.

  • Conops persica Becker, 1913: 609. Type locality: Iran.

  • Conops punctithorax Becker, 1913: 611. Type locality: Iran.

Materials
  1. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568897; scientificName: Physocephala pusilla; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.092932; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 05-26-1926; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568897; scientificName: Physocephala pusilla; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Abu-Rawash; decimalLatitude: 30.043845; decimalLongitude: 31.092932; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 07-18-1925; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: Bdair; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568897; scientificName: Physocephala pusilla; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Ezbet-Naghl; decimalLatitude: 32.1111; decimalLongitude: 32.1625; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 05-06-1916; Record Level: institutionCode: EFC

  4. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568897; scientificName: Physocephala pusilla; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kafr Hakim; decimalLatitude: 30.0808; decimalLongitude: 31.1164; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 05-20-1930; Record Level: institutionCode: EFC

Distribution

PA: Afghanistan, China, Egypt, Europe (widespread), Kazakhstan, Mongolia, Morocco, Saudi Arabia, Syria, Tunisia, Turkmenistan.

Local distribution and dates of collection (Fig. 6): EGYPT: Eastern Desert: Cairo-Suez Road, El-Wasfia, Geneifa, Wadi Digla, Wadi Gharagid (April to September); Fayoum: Fayoum City, Kom Osheem (March to October); Lower Nile Valley and Delta: Abu-Rawash, Ashmoun, Ezbet El-Nakhl, Kafr Hakim, Maadi, Mansouriah, Pyramids (April to October); Sinai: Wadi El-Lega, Wadi El-Rabba, Wadi Firan, Wadi Isla, Wadi Khoshbi (April to August); Western Desert: Madinet El-Sadat, Siwa Oasis (July to October) [Sources: Zalat et al. (2008), Azmy (2016) and museum material in EFC and ESEC]. SAUDI ARABIA: locality and date unknown [Source: Abu-Zoherah et al. (1993) as Conops lacera].

Physocephala variegata

(Meigen, 1824)

B5A64EBE-FF12-52AE-A76D-CE6400E12594

https://www.gbif.org/species/1569101

  • Conops variegata Meigen, 1824: 132. Type locality: Austria & France.

Materials
  1. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Dar El-Maskhara; decimalLatitude: 29.7833; decimalLongitude: 31.4167; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 04-29-1925; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Digla; decimalLatitude: 29.9578; decimalLongitude: 31.3348; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 07-25-1926; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: Kassem; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Gharagid; decimalLatitude: 28.9550; decimalLongitude: 31.4822; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 08-24-1925; Record Level: institutionCode: EFC

  4. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Hoff; decimalLatitude: 29.8820558; decimalLongitude: 31.3109855; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 05-17-1928; Record Level: institutionCode: EFC

  5. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Wadi Hoff; decimalLatitude: 29.8820558; decimalLongitude: 31.3109855; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 06-06-1927; Record Level: institutionCode: EFC

  6. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Halaib; decimalLatitude: 22.2203; decimalLongitude: 36.6427; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 01-12-1929; Record Level: institutionCode: EFC

  7. Type status: Other material. Occurrence: sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: W. Ibib; decimalLatitude: 22.83333; decimalLongitude: 35.76667; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-11-1928; Record Level: institutionCode: EFC

  8. Type status: Other material. Occurrence: recordedBy: R.M.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kafr Katati; decimalLatitude: 30.004146; decimalLongitude: 31.121566; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 05-25-1927; Record Level: institutionCode: EFC

  9. Type status: Other material. Occurrence: recordedBy: Sh.M.; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569101; scientificName: Physocephala variegata; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Ein Gedeirat; decimalLatitude: 30.6500; decimalLongitude: 34.4333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 06-25-1934; Record Level: institutionCode: EFC

Distribution

AF: Egypt [as “Gebel Elba”]. PA: Austria, Bulgaria, Egypt, France, Greece, Hungary, Iran, Kazakhstan, Kyrgyzstan, Mongolia, Romania, Russia, Saudi Arabia, Serbia, Turkey, Turkmenistan, Ukraine.

Local distribution and dates of collection (Fig. 7): EGYPT: Coastal Strip: Mariout, Salloum (May to August); Eastern Desert: Abu Sueir, Geneifa, Wadi Digla, Wadi Dir El-Maskhara, Wadi Garrariyat, Wadi Gharagid, Wadi Hoff, Wadi Ibtadi, Wadi Rishrash, Wadi Um-Elek (April to August); Gebel Elba: Mersa Halaib, Wadi Edeib, Wadi Ibib (January to March); Lower Nile Valley and Delta: Abu-Rawash, Kafr Ghatati (May to July); Sinai: El-Arish, Wadi El-Daiqa, Wadi El-Rabba, Wadi Firan, Wadi Gedeirat (April to August) [Sources: Kröber (1927), Kröber (1929), Azmy (2016) and museum material in EFC and ESEC]. SAUDI ARABIA: locality and date unknown [Source: Zimina (1963), Stuke (2017)].

Figure 7.

Figure 7.

Local distribution map of Physocephala variegata (Meigen) and Physocephala vittata (Fabricius) in Egypt and Saudi Arabia.

Physocephala vittata

(Fabricius, 1794)

86C49695-1BE7-59CB-8642-60F1BC8B84A8

https://www.gbif.org/species/1569046

  • Conops vittata Fabricius, 1794: 392. Type locality: Germany (Kiel).

  • Conops dorsalis Wiedemann in Meigen, 1824: 133. Type locality: Austria.

  • Conops solaeformis Gimmerthal, 1842: 672. Type locality: Latvia.

  • Conops semiatrata Costa, 1844: 89. Type locality: Italy.

  • Conops fraternus Loew, 1847: 18. Type locality: Turkey, Greece, Croatia & Italy.

  • Conops truncata Loew, 1847: 21. Type locality: Italy (Sicily).

  • Physocephala detecta Becker, 1913: 615. Type locality: Iran.

  • Physocephala vittata var. abdominalis Kröber, 1915: 57. Type locality: Turkey, Syria & Cyprus.

  • Physocephala vittata var. semirufa Kröber, 1915: 58. Type locality: Israel.

  • Physocephala truncata var. maculigera Kröber, 1915: 71. Type locality: Egypt, Tunisia, Algeria & Syria.

  • Physocephala vittata var. immaculata Kröber, 1939: 365. Type locality: Cyprus.

Distribution

AF: Ethiopia, Kenya, Yemen. PA: Afghanistan, Algeria, Armenia, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Cyprus, Czech Republic, Egypt, France, Germany, Hungary, Iran, Israel, Italy, Lithuania, Malta, Mongolia, Morocco, Netherlands, Norway, Palestinian Territories, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Spain, Sweden, Switzerland, Syria, Tunisia, Turkey, Turkmenistan, Ukraine, Uzbekistan.

Local distribution and dates of collection (Fig. 7): EGYPT: Coastal Strip: El-Burg, Mariout (May); Sinai: locality and date unknown; Upper Nile Valley: locality and date unknown [Sources: Kröber (1915a), Kröber (1927), Azmy (2016)].

MYOPINAE

CBC63EAC-B7F2-5B01-A625-1D7784F019FA

MYOPINI

113B4D60-A103-5B85-8BE8-F621A2CB8191

Myopa

Fabricius

A1376120-EB6D-5604-AD52-58567D1CB98E

https://www.gbif.org/species/1568357

  • Myopa Fabricius, 1775: 798. Type species: Conops buccata Linnaeus, by subsequent designation of Curtis (1831).

Myopa dorsalis

Fabricius, 1794

4561A5BB-F557-5A95-A89B-4FC0F211ACC5

https://www.gbif.org/species/1568413

  • Myopa dorsalis Fabricius, 1794: 397. Type locality: Germany.

  • Myopa ferruginea Panzer, 1794: 24. Type locality: Germany.

  • Myopa grandis Meigen, 1804: 284. Type locality: Germany.

  • Myopa dorsalis var. nigrifacies Becker, 1922: 289. Type locality: Not given.

Materials
  1. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568413; scientificName: Myopa dorsalis; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Behig; decimalLatitude: 30.9310; decimalLongitude: 29.5988; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-28-1927; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568413; scientificName: Myopa dorsalis; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg El-Arab; decimalLatitude: 30.908084; decimalLongitude: 29.546389; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-10-1926; Record Level: institutionCode: EFC

Distribution

OR: India. PA: Albania, Algeria, Armenia, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Denmark, Egypt, Finland, France, Germany, Greece, Hungary, Iran, Italy, Kazakhstan, Luxembourg, Morocco, Netherlands, Norway, Poland, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Syria, Tunisia, Turkey, Turkmenistan, Ukraine.

Local distribution and dates of collection (Fig. 8): EGYPT: Coastal Strip: Behig, Burg El-Arab, Mariout (February and March); Lower Nile Valley and Delta: Madinet Badr (January); Sinai: Wadi El-Rabba (April) [Sources: Azmy et al. (2016) and museum material in EFC and ESEC].

Figure 8.

Figure 8.

Local distribution map of Myopa dorsalis Fabricius and Myopa picta Panzer in Egypt and Saudi Arabia.

Myopa picta

Panzer, 1798

2657AEC4-4949-5D2C-8006-B3693AFC63B2

https://www.gbif.org/species/1568425

  • Myopa picta Panzer, 1798: 22. Type locality: Austria.

  • Myopa varia Wiedemann, 1830: 242. Type locality: Egypt.

  • Myopa meridionalis Macquart, 1835: 34. Type locality: Egypt.

Materials
  1. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568425; scientificName: Myopa picta; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Behig; decimalLatitude: 30.9310; decimalLongitude: 29.5988; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-28-1927; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568425; scientificName: Myopa picta; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg; decimalLatitude: 30.908084; decimalLongitude: 29.546389; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-25-1932; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568425; scientificName: Myopa picta; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg El-Arab; decimalLatitude: 30.908084; decimalLongitude: 29.546389; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-10-1926; Record Level: institutionCode: EFC

  4. Type status: Other material. Occurrence: recordedBy: Willson; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568425; scientificName: Myopa picta; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Mersa Matrouh; decimalLatitude: 29.5696; decimalLongitude: 26.4194; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: March.1920; Record Level: institutionCode: EFC

Distribution

AF: Ethiopia. OR: India, Pakistan. PA: Afghanistan, Austria, Belarus, Belgium, Bosnia and Herzegovina, Bulgaria, China, Croatia, Czech Republic, Egypt, France, Germany, Greece, Iran, Italy, Kyrgyzstan, Morocco, Poland, Romania, Russia, Slovakia, Spain, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine.

Local distribution and dates of collection (Fig. 8): EGYPT: Coastal Strip: Amria, Behig, Burg El-Arab, Mariout, Mersa Matrouh (February to May); Sinai: Ein Gedeirat (June) [Sources: Kröber (1925), Azmy et al. (2016) and museum material in EFC and ESEC].

Myopa stigma

Meigen, 1824

33C3A52E-EDDF-5238-8854-C8C13C857252

https://www.gbif.org/species/1568382

  • Myopa stigma Meigen, 1824: 148. Type locality: Austria.

  • Myopa arabica Macquart 1851: 138. Type locality: Saudi Arabia (Jeddah).

Distribution

AF: Ethiopia. PA: Afghanistan, Algeria, Austria, Belarus, Bulgaria, Cyprus, Czech Republic, France, Germany, Greece, Hungary, Italy, Kazakhstan, Morocco, Poland, Portugal, Romania, Russia, Saudi Arabia (Jeddah), Slovakia, Spain, Switzerland, Tunisia, Turkey, Ukraine.

Local distribution and dates of collection (Fig. 9): SAUDI ARABIA: Makka Al-Mukarramah: Jeddah (date unknown) [Source: original description of M. arabica Macquart and Stuke and Clements 2008].

Figure 9.

Figure 9.

Local distribution map of Myopa stigma Meigen and Myopa testacea (Linnaeus) in Egypt and Saudi Arabia.

Myopa testacea

(Linnaeus, 1767)

5DBDE653-63AB-533E-8BF5-2E34BCA1A11D

https://www.gbif.org/species/1568398

  • Conops testacea Linnaeus, 1767: 1006. Type locality: "Europa australis Ascanius".

  • Myopa longirostris Robineau-Desvoidy, 1830: 243. Type locality: Not given.

  • Myopa pictipennis Robineau-Desvoidy, 1830: 243. Type locality: Not given.

  • Myopa umbripennis Robineau-Desvoidy, 1830: 243. Type locality: Not given.

  • Myopa testacea var. japonica Kröber, 1916: 89. Type locality: Japan.

Materials
  1. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568398; scientificName: Myopa testacea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg; decimalLatitude: 30.9310; decimalLongitude: 29.5988; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-25-1926; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568398; scientificName: Myopa testacea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg El-Arab; decimalLatitude: 30.908084; decimalLongitude: 29.546389; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-10-1926; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568398; scientificName: Myopa testacea; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Kubba; decimalLatitude: 30.0876; decimalLongitude: 31.2854; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-20-1921; Record Level: institutionCode: ESEC

Distribution

OR: India. PA: Afghanistan, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Denmark, Egypt, Finland, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Japan, Kazakhstan, Kyrgyzstan, Lithuania, Luxembourg, Malta, Mongolia, Netherlands, Norway, Poland, Portugal, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine.

Local distribution and dates of collection (Fig. 9): EGYPT: Coastal Strip: Amria, Burg El-Arab, Mariout (February to June); Lower Nile Valley and Delta: Kubba (February) [Sources: Azmy et al. (2016) and museum material in EFC and ESEC].

THECOPHORINI

F28F8D07-2E94-5BAC-8C4A-460D355BB105

Thecophora

Rondani

4725D923-F9C0-5829-9C2C-7A4F875AC92F

  • Thecophora Thecophora Rondani, 1845: 15. Type species: Myopa atra Fabricius, by monotypy.

Thecophora atra

(Fabricius, 1775)

C9C45211-4E80-5505-9B32-38BB065160CD

https://www.gbif.org/species/1569366

  • Myopa atra Fabricius, 1775: 799. Type locality: Denmark (Copenhagen).

  • Myopa annulata Fabricius, 1794: 399. Type locality: Italy.

  • Myopa maculata Meigen, 1804: 287. Type locality: France.

  • Myopa micans Meigen, 1804: 288. Type locality: France.

  • Myopa femorata Fabricius, 1805: 181. Type locality: Germany.

  • Occemya fulvifrons Robineau-Desvoidy, 1853: 134. Type locality: France.

  • Occemya meigeni Robineau-Desvoidy, 1853: 135. Type locality: France.

  • Occemya grisea Robineau-Desvoidy, 1853: 137. Type locality: France.

  • Occemya lamarckii Robineau-Desvoidy, 1853: 140. Type locality: France.

  • Occemya guerini Robineau-Desvoidy, 1853: 141. Type locality: France.

  • Occemya bigoti Robineau-Desvoidy, 1853: 142. Type locality: France.

  • Occemya brunipes Robineau-Desvoidy, 1853: 143. Type locality: France.

  • Occemya lucasi Robineau-Desvoidy, 1853: 144. Type locality: France.

Materials
  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569366; scientificName: Thecophora atra; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Ezbet El-Nakhl; decimalLatitude: 32.1111; decimalLongitude: 32.1625; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 04-25-1924; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569366; scientificName: Thecophora atra; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Hawamdia; decimalLatitude: 29.9092; decimalLongitude: 31.2615; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: August.1925; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569366; scientificName: Thecophora atra; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 12-11-1934; Record Level: institutionCode: EFC

  4. Type status: Other material. Occurrence: recordedBy: El-Hawagry; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569366; scientificName: Thecophora atra; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; locality: Al-Aqiq; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 4-20-2011; Record Level: institutionCode: MSHC

Distribution

AF: Saudi Arabia [as “South-western part”] (first record). OR: India, Sri Lanka. PA: Afghanistan, Albania, Algeria, Austria, Belgium, Bosnia and Herzegovina, Bulgaria, China, Croatia, Cyprus, Czech Republic, Denmark, Egypt, Finland, France, Germany, Great Britain, Greece, Hungary, Iran, Ireland, Italy, Japan, Kyrgyzstan, Lithuania, Luxembourg, Macedonia, Malta, Mongolia, Montenegro, Morocco, Netherlands, North Korea, Norway, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Spain, Sweden, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine.

Local distribution and dates of collection (Fig. 10): EGYPT: Coastal Strip: Mariout (April and May); Lower Nile Valley and Delta: Ezbet El-Nakhl, Hawamdia, Helwan, Kafr Hakim, Mazghouna (March to December) [Sources: Azmy et al. 2016 and museum material in EFC and ESEC]. SAUDI ARABIA: Al-Baha: Al-Aqiq (April) [Source: collected material].

Figure 10.

Figure 10.

Local distribution map of Thecophora atra (Fabricius) and Thecophora fulvipes (Robineau-Desvoidy) in Egypt and Saudi Arabia.

Notes

This species (Fig. 12b) is recorded herein for the first time from Saudi Arabia and this is the first record in the Afrotropical Region considering the south-western part of Saudi Arabia to be affiliated with the Afrotropical Region.

Thecophora fulvipes

(Robineau-Desvoidy, 1830)

C580F758-DF0E-52A7-A4B8-24E460A2EEA4

https://www.gbif.org/species/1569429

  • Myopa fulvipes Robineau-Desvoidy, 1830: 246. Type locality: France (Paris).

  • Myopa sundewalli Zetterstedt, 1844: 942. Type locality: Sweden.

Materials
  1. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1569429; scientificName: Thecophora fulvipes; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; stateProvince: Asir; locality: Garf Raydah Nature Reserve; decimalLatitude: 18.194917; decimalLongitude: 42.4072; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Malaise trap; eventDate: 06-07-2014; Record Level: institutionCode: KSMA

Distribution

AF: Saudi Arabia [as “South-western part”] (El-Hawagry et al. 2017El-Hawagry et al. 2017). PA: Austria, Belgium, Bulgaria, China, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britain, Hungary, Iran, Ireland, Italy, Latvia, Lithuania, Mongolia, Netherlands, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Spain, Sweden, Switzerland, Turkey, Ukraine.

Local distribution and dates of collection (Fig. 10): EGYPT: Fayoum: Kom Osheem (June); Lower Nile Valley and Delta: Helwan (December) [Sources: Azmy 2016]. SAUDI ARABIA: Asir: Raydah Nature Reserve (June) [Source: El-Hawagry et al. 2017 and collected material].

ZODIONINAE

31F6A9C8-4794-58AF-90F5-A2645E6B4DED

Zodion

Latreille

7C6AD3BD-43E6-5BB2-BC92-2A7FB81957C9

https://www.gbif.org/species/1568754

  • Zodion Zodion Latreille, 1797: 162. Type species: Myopa cinerea Fabricius, by Subsequent monotypy of Latreille (1802).

Zodion cinereum

(Fabricius, 1794)

B669E2C8-1C91-559F-A10D-8DB9A0F9AE59

https://www.gbif.org/species/1568846

  • Myopa cinerea Fabricius, 1794: 399. Type locality: Italy.

  • Myopa notata Meigen, 1804: 288. Type locality: France.

  • Myopa tibialis Fabricius, 1805: 182. Type locality: Europe.

  • Zodion conopsoides Latreille, 1809: 337. Type locality: France.

  • Zodion pedicillatum Robineau-Desvoidy, 1830: 252. Type locality: Not given.

  • Zodion fuliginosum Robineau-Desvoidy, 1853: 156. Type locality: France.

  • Zodion fulvipes Robineau-Desvoidy, 1853: 157. Type locality: France.

  • Zodion fulvicorne Robineau-Desvoidy, 1853: 158. Type locality: France.

  • Zodion cinereum var. rubescens Szilády, 1925: 223. Type locality: Hungary.

  • Zodion cinereum var. kerteszi Szilády, 1926: 589. Type locality: Hungary.

Materials
  1. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568846; scientificName: Zodion cinereum; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Behig; decimalLatitude: 30.9310; decimalLongitude: 29.5988; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-28-1927; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568846; scientificName: Zodion cinereum; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg; decimalLatitude: 30.908084; decimalLongitude: 29.546389; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 03-05-1930; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568846; scientificName: Zodion cinereum; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Burg; decimalLatitude: 30.908084; decimalLongitude: 29.546389; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-25-1932; Record Level: institutionCode: EFC

  4. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568846; scientificName: Zodion cinereum; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Mariout; decimalLatitude: 31.017186; decimalLongitude: 29.759966; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 02-15-1923; Record Level: institutionCode: EFC

  5. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568846; scientificName: Zodion cinereum; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Egypt; locality: Mariout: El-Burg; decimalLatitude: 30.908084; decimalLongitude: 29.546389; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 06-16-1929; Record Level: institutionCode: EFC

  6. Type status: Other material. Occurrence: recordedBy: El-Hawagry; sex: 1 male, 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568846; scientificName: Zodion cinereum; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; stateProvince: Al-Baha; locality: Ghabet Shahba; decimalLatitude: 20.02723; decimalLongitude: 41.28565; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Sweeping; eventDate: 05-06-2012; Record Level: institutionCode: MSHC

  7. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; sex: 1 female; lifeStage: adult; Taxon: taxonID: https://www.gbif.org/species/1568846; scientificName: Zodion cinereum; kingdom: Animalia; phylum: Arthropoda; class: Insecta; order: Diptera; family: Conopidae; Location: country: Saudi Arabia; stateProvince: Al-Baha; locality: Jabal Shada al-A’la Nature Reserve; decimalLatitude: 19.8388; decimalLongitude: 41.3101; Identification: identifiedBy: M. El-Hawagry; dateIdentified: June 2020; Event: samplingProtocol: Malaise trap; eventDate: 05-05-2015; Record Level: institutionCode: KSMA

Distribution

AF: Central African Republic, Saudi Arabia [as “South-western part”] (El-Hawagry et al. 2016, United Arabian Emirates. OR: India. PA: Afghanistan, Albania, Algeria, Armenia, Austria, Belgium, Bosnia and Herzegovina, Bulgaria, China, Croatia, Cyprus, Czech Republic, Denmark, Egypt, Estonia, Finland, France, Germany, Great Britain, Greece, Hungary, Iran, Ireland, Italy, Japan, Kazakhstan, Kyrgyzstan, Latvia, Lebanon, Lithuania, Luxembourg, Macedonia, Moldova, Mongolia, Montenegro, Morocco, Netherlands, Norway, Poland, Portugal, Romania, Russia, Serbia, Slovakia, Spain, Sweden, Switzerland, Tunisia, Turkey, Turkmenistan, Ukraine.

Local distribution and dates of collection (Fig. 11): EGYPT: Coastal Strip: Behig, Burg El-Arab, Mariout (February and March); Sinai: Wadi El-Lega, Wadi El-Rabba (April, August and September) [Sources: Azmy (2016) and museum material in EFC]. SAUDI ARABIA: Al-Baha: Ghabet Shahba (Al-Baha City) (May and June 2012), Jabal Shada al-A’la Nature Reserve (May) [Source: Stuke (2008), El-Hawagry et al. (2016) and collected material].

Figure 11.

Figure 11.

Local distribution map of Zodion cinereum (Fabricius) and Zodion erythrurum Rondani in Egypt and Saudi Arabia.

Zodion erythrurum

Rondani, 1865

A862FD02-D012-50A3-8142-283DF648131F

https://www.gbif.org/species/1568759

  • Zodion erythrurum Rondani, 1865: 146. Type locality: Italy (Etruria).

  • Zodion pulchrum Loew, 1868: 384. Type locality: Italy.

  • Zodion vittipes Strobl, 1906: 331. Type locality: Spain.

Distribution

PA: Algeria, Bulgaria, Egypt, France, Hungary, Israel, Italy, Kazakhstan, Kyrgyzstan, Morocco, Poland, Romania, Russia, Serbia, Spain, Tunisia, Turkey, Turkmenistan, Ukraine.

Local distribution and dates of collection (Fig. 11): EGYPT: Coastal Strip: Burg El-Arab, Mariout (February to April); Eastern Desert: Wadi Digla (July and August); Gebel Elba: locality and unknown (April and May); Upper Nile Valley: Esna (January) [Sources: Kröber (1927), Azmy (2016) and museum material in ESEC].

Supplementary Material

XML Treatment for CONOPIDAE
XML Treatment for CONOPINAE
XML Treatment for CONOPINI
XML Treatment for Conops
XML Treatment for Asiconops
XML Treatment for Conops (Asiconops) elegans
XML Treatment for Conops (Asiconops) flavifrons
XML Treatment for Conops (Asiconops) nubeculipennis
XML Treatment for Conops
XML Treatment for Conops (Conops) flavicauda
XML Treatment for Conops (Conops) quadrifasciatus
XML Treatment for Conops (Conops) rufiventris
XML Treatment for Conops (Conops) tomentosus
XML Treatment for PHYSOCEPHALINI
XML Treatment for Physocephala
XML Treatment for Physocephala antiqua
XML Treatment for Physocephala chrysorrhoea
XML Treatment for Physocephala pusilla
XML Treatment for Physocephala variegata
XML Treatment for Physocephala vittata
XML Treatment for MYOPINAE
XML Treatment for MYOPINI
XML Treatment for Myopa
XML Treatment for Myopa dorsalis
XML Treatment for Myopa picta
XML Treatment for Myopa stigma
XML Treatment for Myopa testacea
XML Treatment for THECOPHORINI
XML Treatment for Thecophora
XML Treatment for Thecophora atra
XML Treatment for Thecophora fulvipes
XML Treatment for ZODIONINAE
XML Treatment for Zodion
XML Treatment for Zodion cinereum
XML Treatment for Zodion erythrurum

Acknowledgements

The authors would like to extend their sincere appreciation to the Deanship of Scientific Research at King Saud University for funding this work [Research group No: RG-1437-009].

We are grateful to Dr. Jens-Hermann Stuke, Roter Weg 22, D-26789 Leer, Germany and Dr. David Clements, 7 Vista Rise, Llandaff, Cardiff, CF5 2SD, U.K. for providing critical assistance throughout this study and for supplying us with some relevant papers. We are also grateful to the research team in KSMA for collecting, mounting and photographing specimens.

Contributor Information

Magdi El-Hawagry, Email: elhawagry@gmail.com.

Ahmed Mostafa Soliman, Email: ammsoliman@gmail.com.

Hathal Mohammed Al Dhafer, Email: hdhafer@ksu.edu.sa.

References

  1. Abu-Zoherah R., Al-Taher K., Tilkian S. List of insects recorded from Saudi Arabia. Ministry of Agriculture and Water, National Agriculture and Water Research Centre. Kingdom of Saudi Arabia; Riyadh: 1993. 394 [Google Scholar]
  2. Azmy R. M. Taxonomic and Morphological Studies on Family Conopidae (Order: Diptera) in Egypt, A thesis presented in partial fulfillment for the award of the M.Sc. Degree (Entomology) Department of Entomology, Faculty of Science, Ain Shams University; Cairo: 2016. 180 [Google Scholar]
  3. Azmy R. M., Soliman A. M., Aly F. I., Gad Allah S. M., Mohammad S. K. A Taxonomic review of the subfamily Myopinae Macquart, 1834 (Conopidae, Diptera) from Egypt. African Journal of Biological Sciences. 2016;12(1):169–178. [Google Scholar]
  4. Chvála M., Smith K. G. V. Conopidae . In: Sóos A., Papp L., editors. Catalogue of Palearctic Diptera. Vol. 8. Elsevier, Amsterdam-Oxford-New York-Tokyo; 1988. 363 [Google Scholar]
  5. Curtis J. British Entomology; being illustrations and descriptions of the genera of insects found in in Great Britain and Ireland: containing coloured figures from nature of the most rare and beautiful species, and in many instances of the plants upon which they are found. Vol. 15. Curtis; London: 1831. [Google Scholar]
  6. El-Hawagry Magdi, Gilbert Francis. Zoogeographical affinities and faunal relationships of bee flies (Diptera: Bombyliidae) in Egypt. Zoology in the Middle East. 2014;60(1):50–56. doi: 10.1080/09397140.2014.892339. [DOI] [Google Scholar]
  7. El-Hawagry Magdi S., Sharaf Mostafa R., Al Dhafer Hathal M., Fadl Hassan H., Aldawood Abdulrahman S. Addenda to the insect fauna of Al-Baha Province, Kingdom of Saudi Arabia with zoogeographical notes. Journal of Natural History. 2016;50:1209–1236. doi: 10.1080/00222933.2015.1103913. [DOI] [Google Scholar]
  8. El-Hawagry M. S., Abdel-Dayem M. S., El-Sonbati S. A., HM Al. A preliminary account of the fly fauna in Garf Raydah Nature Reserve, Kingdom of Saudi Arabia, with new records and biogeographical remarks (Diptera: Insecta) Journal of Natural History. 2017;51:1499–1530. doi: 10.1080/00222933.2017.1347299. [DOI] [Google Scholar]
  9. El-Hawagry M. S., Abdel-Dayem M. S., Al Dhafer H. M. The family Oestridae in Egypt and Saudi Arabia (Diptera, Oestroidea. ZooKeys. 2020;947:113–142. doi: 10.3897/zookeys.947.52317. [DOI] [PMC free article] [PubMed] [Google Scholar]
  10. Freeman B. A. Notes on conopid flies, including insect host, plant and phoretic relationships (Diptera: Conopidae. Journal of the Kansas Entomological Society. 1966;39:123–131. [Google Scholar]
  11. Gibson Joel F., Skevington Jeffrey H. Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data. Zoological Journal of the Linnean Society. 2013;167(1):43–81. doi: 10.1111/j.1096-3642.2012.00873.x. [DOI] [Google Scholar]
  12. Gibson J. F., Skevington J. H., Kelso S. A phylogenetic analysis of relationships between genera of Conopidae (Diptera) based on molecular and morphological data. Cladistics. 2013;29(2):193–226. doi: 10.1111/j.1096-0031.2012.00422.x. [DOI] [PubMed] [Google Scholar]
  13. Hölzel H. Zoogeographical features of Neuroptera of the Arabian Peninsula. Acta Zoologica Fennica. 1998;209:120–140. [Google Scholar]
  14. Kröber O. Die palaearktischen Arten der Gattung Physocephala. Archiv fr Naturgeschichte, Abteilung A. 1915;80:43–77. [Google Scholar]
  15. Kröber O. Die palaearktischen Arten der Gattung Conops L. Archiv fr Naturgeschichte, Abteilung A. 1915;81:35–60. [Google Scholar]
  16. Kröber O. Aegyptische Dipteren aus den Familien der Conopidae, Omphralidae und Therevidae. Bulletin de la Socit Royale Entomologique d'Egypte. 1924;7:57–116. [Google Scholar]
  17. Kröber O. Neue Dipteren aus Aegypten. Bulletin de la Socit Royale Entomologique d'Egypte. 1925;9:232––243. [Google Scholar]
  18. Kröber O. Aegyptische Dipteren aus den Familien der Conopidae und Therevidae. Bulletin de la Socit Royale Entomologique d'Egypte. 1927;11:9–16. [Google Scholar]
  19. Kröber O. Neue Dipteren aus Aegypten aus den Familien Tabanidae, Therevidae, Omphralidae und Conopidae. Bulletin de la Socit Royale Entomologique d'Egypte. 1929;13:73–84. [Google Scholar]
  20. Kröber O. Beitrge zur Kenntnis der Conopiden. I, II. Die Conopiden der Aethiopischen Region; III. Die Conopiden Sud- und Mittelamerikas (incl. Antillen und Mexiko); IV. Die Conopiden der Australischen Region; V. Die Conopiden der Orientalischen fauna. Annals and Magazine of Natural History. 1939;4:362–395. [Google Scholar]
  21. Latreille P. A. Histoire naturelle, generale et particuliere, des crustaces et des insectes. Vol. 3. Dufart; Paris: 1802. 467 [Google Scholar]
  22. Macquart P. J.M. Histoire Naturelle des insectes. Diptères. Vol. 2. Roret; Paris: 1835. 710 [Google Scholar]
  23. Macquart P. J.M. Diptères exotiques nouveaux ou peu connus. Suite du 4.e Supplement. Mémoires de la Société des sciences, de l’agriculture et des arts de Lille, 1851;1850:134–249. [Google Scholar]
  24. Mohammed S. K., Negm F. A list of Conopidae of Egypt (Cyclorrhapha, Diptera) Egyptian Journal of Biological Pest Control. 1991;1:109–115. [Google Scholar]
  25. Pape T., Blagoderov V., Mostovski M. B. Order Diptera Linnaeus, 1758. In: Zhang Z-Q (Ed.), Animal biodiversity: An outline of higher–level classification and survey of taxonomic richness. Zootaxa. 2011;3148:222–229. doi: 10.11646/zootaxa.3148.1.42. [DOI] [PubMed] [Google Scholar]
  26. Rondani A. C. Osservazioni sopra alquante specie di esapodi ditteri del Museo Torinese. Nuovi Annali delle Scienze Naturali e Rendiconto dei Lavori dell’Accademia delle Scienze dell’Istituto di Bologna con Appendice Agraria. 1850;2:165–179. [Google Scholar]
  27. Shorthouse D. SimpleMappr, an online tool to produce publication-quality point maps. https://www.simplemappr.net. [2020-10-25T00:00:00+03:00];
  28. Steyskal G. C., El-Bialy S. A list of Egyptian Diptera with a bibliography and key to families. Ministry of Agriculture Technical Bulletin. 1967;3:24–25. [Google Scholar]
  29. Stuke J. H. Order Diptera, family Conopidae A. Arthropod fauna of the United Arab Emirates. In: van Harten A., editor. Arthropod Fauna of the United Arab Emirates. Vol. 1. Dar Al Ummah Publishers; Abu Dhabi: 2008. 754 [Google Scholar]
  30. Stuke J H, Clements D K. Revision of the Myopa testacea species-group in the Palaearctic region (Diptera: Conopidae) Zootaxa. 2008;1713:1–26. doi: 10.5281/zenodo.274119. [DOI] [Google Scholar]
  31. Stuke J. H., Hayat R., Özbek H. Records of notable Conopidae (Diptera) from Turkey. Faunistische Abhandlungen. 2008;26:109–117. [Google Scholar]
  32. Stuke J. H. A contribution to the knowledge of Physocephala antiqua (Wiedemann) (Diptera: Conopidae. Studia dipterologica. 2014;21(2):209–220. [Google Scholar]
  33. Stuke J. H. World catalogue of insects. Conopidae (Diptera) Vol. 15. e.J. Brill, leiden; Boston: 2017. xxxviii + 354 [Google Scholar]
  34. Walker D. H., Pittaway A. R. Insects of Eastern Arabia. Patron: H.E. Sheikh Nahayan bin Mubarak Al-Nahayan, Emirates Natural History Group; 1987. 175 [Google Scholar]
  35. Walker F. List of Diptera collected in Egypt and Arabia, by J. K. Lord, Esq., with descriptions of the species new to science. Entomologist. 1871;5:255–263. [Google Scholar]
  36. Wallace Alfred Russel. The geographical distribution of animals. Vol.1 & 2. Macmillan; London: 1876. 574 + 650 [Google Scholar]
  37. Wiedemann C. R.W. Aussereuropäische zweiflügelige Insekten. Als Fortsetzung des Meigenschen Werkes. Vol. 2. Schulz, Hamm; 1830. xii + 684 [Google Scholar]
  38. Zalat S. M., Gilbert F., Fadl H. H., El-Hawagry M. S., Abdel-Dayem M. S., Kamel S., Gilbert J. Biological explorations of Sinai: flora and fauna of Wadi Isla and Hebran, St Katherine Protectorate. Egypt Egyptian Journal of Natural History. 2008;5:6–15. [Google Scholar]
  39. Zimina L. V. The genus Myopa Fabr. (Diptera, Conopidae) in the USSR. Entomological Review. 1963;42:211–218. [Google Scholar]

Associated Data

This section collects any data citations, data availability statements, or supplementary materials included in this article.

Supplementary Materials

XML Treatment for CONOPIDAE
XML Treatment for CONOPINAE
XML Treatment for CONOPINI
XML Treatment for Conops
XML Treatment for Asiconops
XML Treatment for Conops (Asiconops) elegans
XML Treatment for Conops (Asiconops) flavifrons
XML Treatment for Conops (Asiconops) nubeculipennis
XML Treatment for Conops
XML Treatment for Conops (Conops) flavicauda
XML Treatment for Conops (Conops) quadrifasciatus
XML Treatment for Conops (Conops) rufiventris
XML Treatment for Conops (Conops) tomentosus
XML Treatment for PHYSOCEPHALINI
XML Treatment for Physocephala
XML Treatment for Physocephala antiqua
XML Treatment for Physocephala chrysorrhoea
XML Treatment for Physocephala pusilla
XML Treatment for Physocephala variegata
XML Treatment for Physocephala vittata
XML Treatment for MYOPINAE
XML Treatment for MYOPINI
XML Treatment for Myopa
XML Treatment for Myopa dorsalis
XML Treatment for Myopa picta
XML Treatment for Myopa stigma
XML Treatment for Myopa testacea
XML Treatment for THECOPHORINI
XML Treatment for Thecophora
XML Treatment for Thecophora atra
XML Treatment for Thecophora fulvipes
XML Treatment for ZODIONINAE
XML Treatment for Zodion
XML Treatment for Zodion cinereum
XML Treatment for Zodion erythrurum

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