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The following funding information was missed from the final version:
This work was supported by the National Institutes of Health (NIH) Grant R01AR070222 to DC. This work was also partially supported by grants from the National Nature Science Foundation of China (NSFC) (81620108018, 81930070), and Tianjin key research and development plan, key projects for science and technology support (19YFZCSY00660) to SF. This work was also partially supported by NSFC grants (81874011, 81 572 104 and 81301531) to TW. This work was also supported by the NSFC grant (81672227) and the Frontier Science of CAS grant (QYZDB-SSW-JSC030) to HP.