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. 2021 Mar 22;9:e64212. doi: 10.3897/BDJ.9.e64212

The family Stratiomyidae in Egypt and Saudi Arabia (Diptera: Stratiomyoidea)

Magdi El-Hawagry 1,✉, Hathal Mohammed Al Dhafer 2, Mahmoud Abdel-Dayem 2, Martin Hauser 3
PMCID: PMC8007553  PMID: 33790686

Abstract

Background

This study systematically catalogues all known taxa of the family Stratiomyidae in Egypt and Saudi Arabia. It is one in a series of planned studies aiming to catalogue the whole order in both countries.

New information

Twenty species, belonging to seven genera and three subfamilies (Pachygastrinae, Stratiomyinae and Nemotelinae), are treated. One of these genera, Oplodontha and two species, Oplodontha pulchriceps Loew and Oxycera turcica Üstüner & Hasbenli, are recorded herein for the first time from Saudi Arabia. A lectotype for Nemotelus matrouhensis Mohammad et al., 2009 is designated. An updated classification, synonymies, type localities, world and local distributions, dates of collection and some coloured photographs are provided.

Keywords: soldier-flies, local distribution, dates of collection, new records

Introduction

The Stratiomyidae is a lower brachycerous Dipteran family, including more than 2650 species classified in 375 genera and 12 subfamilies worldwide (Woodley 2001). Flies of the family Stratiomyidae, commonly known as soldier-flies, are small to large, ranging from 2 to 28 mm in length, highly varied in shape and colouration, with wings hyaline to variously patterned or infuscated and some species likely mimic wasps and bees in their flight and body colour (Hauser 2008, Hauser et al. 2017b). These flies can be easily distinguished as adults by the following characters: radial veins grouped together anteriorly, ending before tip of the wing; costal vein (C) usually ending well before wing apex and discal cell (d) short, usually forming distinct short, often squarish cell at middle of the wing (Marshall et al. 2017).

Adult soldier-flies can be found sitting on foliage in sunny locations in damp forest habitats, near water or boggy areas and some flies can be found frequenting flowers, particularly those of the subfamily Stratiomyinae and some of the subfamily Clitellariinae (James 1981, Khaghaninia et al. 2015, Hauser et al. 2017b).

Immature stages of soldier-flies can be found in a variety of habitats. Larvae of the subfamilies Beridinae, Clitellariinae and Sarginae, which have not been represented in Egypt or Saudi Arabia, are usually associated with decaying organic matter, such as leaf litter and rotting fruits. However, larvae of the subfamilies Nemotelinae and Stratiomyinae are aquatic, occurring in slow moving or standing water in rivers, ponds, seepage areas and other transient water sources, with those of the genus Nemotelus Geoffroy, 1762 being remarkably tolerant to high levels of salinity. Larvae of the subfamily Pachygastrinae can be found under the bark of fallen trees (Rozkošný 1983, Hauser 2008, Hauser et al. 2017b).

Of the 12 subfamilies, seven are recorded in the Palaearctic Region with about 430 species (Woodley 2001). Of these, only three subfamilies (Pachygastrinae, Stratiomyinae and Nemotelinae) have been represented in Saudi Arabia and/or Egypt by seven genera comprising 20 species. One of these genera, Oplodontha and two species, Oxycera turcica Üstüner & Hasbenli and Oplodontha pulchriceps Loew, are recorded herein for the first time from Saudi Arabia. This is not surprising as no previous faunistic or systematic studies on the Stratiomyidae have been carried out in Saudi Arabia and only two species have been recorded from Saudi Arabia amidst two comprehensive checklists of the Saudi Arabian Diptera: El-Hawagry et al. (2017) [Aspidacantha atra Kertesz] and Abu-Zoherah et al. (1993) [Stratiomys deserticolor Lindner]. We think the overall number of Saudi Arabian taxa treated in the present study is still low and does not represent the real fauna of Stratiomyidae in this country and an extensive faunistic and systematic study on the Saudi Arabian Stratiomyidae is required. On the other hand, some previous studies have been carried out in Egypt to list the Stratiomyidae species (Steyskal and El-Bialy 1967) or to study the fauna and/or taxonomy of the family (Lindner 1930, Badrawy 2006, Mohammad et al. 2009), in addition to some miscellaneous studies which described some new species from Egypt (Olivier 1811, Becker 1902, Wiedemann 1830, Lindner 1925, Lindner 1930, Lindner 1937, Lindner 1974).

Egypt and Saudi Arabia are two neighbouring countries in the Middle East, separated by the Gulf of Aqaba and the Red Sea (Fig. 1). Egypt is a transcontinental country as its majority is located in the north-eastern corner of Africa, while its north-eastern extremity, Sinai Peninsula, is located in the south-western corner of Asia. On the other hand, Saudi Arabia is wholly located in the south-western corner of Asia. Both Egypt and Saudi Arabia are biogeographically located at the junction of the Palaearctic and the Afrotropical Regions. The faunal affiliation of the two countries is mainly Palaearctic. Exceptions are Gebel Elba, the south-eastern triangle of Egypt and the southwestern part of Saudi Arabia, south to the Tropic of Cancer, which are considered as having an Afrotropical faunal affiliation (Wallace 1876, Hölzel 1998, El-Hawagry and Gilbert 2014, El-Hawagry et al. 2017).

Figure 1.

Figure 1.

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A satellite map of Egypt and Saudi Arabia.

This study is one in a series of taxonomic studies on different Egyptian and Saudi Arabian dipteran taxa aiming to catalogue the whole order Diptera in the two countries (EL-Hawagry 2015, Al Dhafer and El-Hawagry 2016, El-Hawagry 2017, EL-Hawagry 2018, El-Hawagry et al. 2018, El-Hawagry and Al Dhafer 2019, El-Hawagry et al. 2019, El-Hawagry and Gilbert 2019, El-Hawagry et al. 2020b, El-Hawagry et al. 2020a, El-Hawagry et al. 2021, El-Hawagry and El-Azab 2019).

Materials and methods

Data sources. Data of the present study are obtained from three main sources: 1. Specimens preserved in the Egyptian and Saudi Arabian insect collections and museums, namely: Efflatoun Bey’s collection, Cairo University, Egypt (EFC); Collection of Entomological Society of Egypt (ESEC) and King Saud University Museum of Arthropods, Saudi Arabia (KSMA). 2. Previous studies on the Stratiomyidae in Egypt and Saudi Arabia. 3. Specimens collected by the authors and their co-workers, especially from Saudi Arabia using both Malaise traps and aerial nets. A great deal of faunistic and taxonomic information, including type species, type localities, Old World synonymies, world and local distributions and collection dates were obtained from relevant literature as well. These sources are listed in the following subsections.

Study area. Egypt and Saudi Arabia, the study area, are two neighbouring countries in the Middle East, separated by the Red Sea and the Gulf of Aqaba. They constitute a part of the Great Desert Belt, mainly with an arid desert climate characterised by hot summer and a mild winter (Alsharhan et al. 2001, Nasser et al. 2019).

Ecologists divide Egypt into eight ecological zones: the Coastal Strip, Lower Nile Valley & Delta, Upper Nile Valley, Fayoum, Eastern Desert, Western Desert, Sinai and Gebel Elba (Fig. 1, El-Hawagry and Gilbert 2014). These ecological zones are adopted in the present study in the sections of localities and dates of collection. However, Saudi Arabia is not divided into ecological zones, so ecologists usually adopt the administrative divisions instead. These divisions (also called emirates, regions or provinces) are adopted in the present study, namely: Makkah, Riyadh, Eastern Province, Asir, Jazan, Al-Madinah, Al-Qaseem, Tabuk, Hail, Najran, Al-Jawf, Al-Baha and Northern Frontier (El-Hawagry and Al Dhafer 2019).

Classification. The classification and arrangement of taxa in the present study basically follows that used in Woodley (2001) and subsequent updates by Hauser et al. (2017a).

World distribution. Sources of world distribution of each stratiomyid species are given between square brackets at the end of the list of countries.

Local distribution and dates of collection. Localities within each Egyptian ecological zone or Saudi Arabian administrative region are arranged alphabetically and written after a colon following each zone or region followed by the dates of collection between parentheses; for example, "Coastal Strip: Abu-Kir, Cleopatra, Dekhela (April to September)" and “Tabuk: Tabuk City (May)”. Sources for this distribution are given between square brackets at the end of the section. Coordinates of all Egyptian and Saudi Arabian localities of the family Stratiomyidae are listed (Table 1). Distribution maps of species were made using SimpleMappr (Shorthouse 2010).

Table 1.

A gazetteer of Egyptian and Saudi Arabian localities of the family Stratiomyidae.

Country Locality Governorate Ecological zone or Region Latitude (N) Longitude (E)
Egypt Abu-Kir Alexandria Coastal Strip 31.22429 33.8256
Abu-Rawash Giza Lower Nile Valley & Delta 30.0438 42.4101
Abu-Zaabal Al-Qalyubia Lower Nile Valley & Delta 30.24098 42.4089
Alexandria Alexandria Coastal Strip 31.2129 42.4089
Behaira Behaira Lower Nile Valley & Delta 30.62189 42.4102
Benha Al-Qalyubia Lower Nile Valley & Delta 30.46572 42.4089
Beni Sweif Beni Sweif Lower Nile Valley & Delta 29.07788 42.4089
Birket Qaroun Fayoum Fayoum 29.40879 42.4124
Cleopatra Alexandria Coastal Strip 31.22022 42.4101
Dakhla Oasis New Valley Western Desert 25.5000 41.3115
Damietta Damietta Lower Nile Valley & Delta 31.34595 41.3115
Dekhela Alexandria Coastal Strip 31.12098 41.3101
Ein Moussa South Sinai Sinai 29.8667 41.3101
El-Alag Al-Qalyubia Lower Nile Valley & Delta 30.18009 42.8348
El-Arish North Sinai Sinai 31.1244 39.8749
El-Baragil Giza Lower Nile Valley & Delta 30.07673 41.4107
El-Ferdan Ismailia Eastern Desert 30.65760 36.6297
El-Gebel El-Asfar Al-Qalyubia Lower Nile Valley & Delta 29.1689 34.1412
El-Kantara Ismailia Eastern Desert 30.79392 34.3836
El-Marg Al-Qalyubia Lower Nile Valley & Delta 31.0667 30.5833
El-Siala Alexandria Coastal Strip 31.20849 30.8180
El-Tour South Sinai Sinai 28.24024 29.1667
Ezbet El-Nakhl Al-Qalyubia Lower Nile Valley & Delta 31.1111 32.6500
Fayed Ismailia Eastern Desert 30.32382 29.8805
Fayoum Fayoum Fayoum 29.32061 32.3008
Gebel Elba Red Sea Gebel Elba 22.2008 30.2167
Gezeira Cairo Lower Nile Valley & Delta 30.04596 34.3836
Girza Fayoum Fayoum 29.49968 34.1412
Giza Giza Lower Nile Valley & Delta 30.01350 31.3333
Giza-Fayoum Road Giza Western Desert 29.85176 31.18121
Helwan Cairo Lower Nile Valley & Delta 29.8500 31.22435
Hurghada Red Sea Eastern Desert 27.2337 30.5833
Ismailia Ismailia Eastern Desert 30.59428 30.8180
Kerdassa Giza Lower Nile Valley & Delta 30.0297 30.48755
Kharga Oasis New Valley Western Desert 25.2500 31.6317
Khosous Cairo Lower Nile Valley & Delta 30.15957 31.1117
Kosseimah North Sinai Sinai 30.90307 30.9898
Maadi Cairo Lower Nile Valley & Delta 29.95772 31.3125
Mariout Alexandria Coastal Strip 31.0172 31.3531
Max Alexandria Coastal Strip 31.1636 30.2167
Mersa Matrouh Matrouh Coastal Strip 29.5696 30.6468
Moharram Bey Alexandria Coastal Strip 31.17796 31.10713
Moweileh South Sinai Sinai 30.3924 30.5833
Nazla Fayoum Fayoum 29.29792 30.5833
Nubar Bey Alexandria Coastal Strip 31.18079 31.2505
Nuzha (Alex.) Alexandria Coastal Strip 31.2001 31.3333
Quisna Menofiya Lower Nile Valley & Delta 30.53514 31.3333
Ramleh Alexandria Coastal Strip 31.2279 30.8180
Sandoub Dakahlyia Lower Nile Valley & Delta 31.00782 30.2167
Sherbin Dakahlyia Lower Nile Valley & Delta 31.19461 31.3333
Shubra Al-Qalyoubia Lower Nile Valley & Delta 30.1012 30.5833
Siwa Oasis Matrouh Western Desert 29.20427 26.4194
Suez Suez Eastern Desert 29.95278 32.3008
Tanta Gharbia Lower Nile Valley & Delta 30.75725 32.3008
Wadi El-Lega South Sinai Sinai 28.5469 32.3008
Wadi El-Natroun Behaira Western Desert 30.3814 29.8157
Wadi Hebran South Sinai Sinai 28.40225 31.3521
Wadi Hoff Cairo Eastern Desert 29.8821 32.3008
Wadi Watir South Sinai Sinai 29.02147 31.3110
Zaranik North Sinai Sinai 31.10345 30.3441
Saudi Arabia Al-Mekhwa Al-Mekhwa Al-Baha 19.798133 30.3441
Dawmat Al-Jandal Dawmat Al-Gandal Al-Jwaf 29.809552 30.8180
Hassan Ameen farm Tabouk Tabouk 28.36661 26.4194
Jabal Shada al-A’la Nature Reserve Al-Mekhwa Al-Baha 19.8429 29.7600
Jazan Jazan Jazan 16.9595 29.7600
Raydah Nature Reserve Abha Asir 18.20525 30.8180
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Abbreviations used:

  • AF, Afrotropical

  • CSCA, California State Collection of Arthropods, Sacramento, California, USA

  • EFC, Efflatoun Bey’s collection, Department of Entomology, Faculty of Science, Cairo University, Egypt

  • ESEC, Collection of Entomological Society of Egypt, Cairo, Egypt

  • KSMA, King Saud University Museum of Arthropods, Riyadh, Saudi Arabia

  • MCCB, Museum of Community College, Al-Baha University, Saudi Arabia

  • OR, Oriental

  • PA, Palearctic

Checklists

The Catalogue

Stratiomyidae

931D5370-5409-5F83-A6D0-7DB38E012480

Stratiomyiinae

33B098FD-8763-51CC-A118-4E3315DFF0E8

Aspidacantha

Kertsz, 1916

205E6D2F-CE54-5634-BC1A-FC3E58007A8F

https://www.gbif.org/species/1578168

  • Aspidacantha Kertész, 1916: 154. Type species: Aspidacantha atra Kertész, by original designation.

Aspidacantha atra

Kertsz, 1916

081AA6FB-0EBC-56F1-904D-113D9976E520

https://www.gbif.org/species/1578171

  • Aspidacantha atra Kertész, 1916: 155. Type locality: Eritrea (Assab) [Assab belongs now to Eritrea not to Ethiopia as written in world catalogues].

Materials

  1. Type status: Other material. Occurrence: recordedBy: H. Pohl; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Egypt; locality: Hurghada; decimalLatitude: 27.23370; decimalLongitude: 33.8256; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Light trap; eventDate: 9/11-22/1994; Record Level: institutionCode: CSCA

  2. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Saudi Arabia; locality: Raydah; decimalLatitude: 18.20525; decimalLongitude: 42.4101; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Sweeping; eventDate: 06-07-2014; Record Level: institutionCode: KSMA

  3. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Saudi Arabia; locality: Raydah; decimalLatitude: 18.201583; decimalLongitude: 42.4089; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Light trap; eventDate: 02-21-2014; Record Level: institutionCode: KSMA

  4. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Saudi Arabia; locality: Raydah; decimalLatitude: 18.201583; decimalLongitude: 42.4089; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Light trap; eventDate: 07-31-2015; Record Level: institutionCode: KSMA

  5. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Saudi Arabia; locality: Raydah; decimalLatitude: 18.20525; decimalLongitude: 42.4102; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Sweeping; eventDate: 09-05-2015; Record Level: institutionCode: KSMA

  6. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Saudi Arabia; locality: Raydah; decimalLatitude: 18.201583; decimalLongitude: 42.4089; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Light trap; eventDate: 09-05-2015; Record Level: institutionCode: KSMA

  7. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Saudi Arabia; locality: Raydah; decimalLatitude: 18.201583; decimalLongitude: 42.4089; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 09-05-2015; Record Level: institutionCode: KSMA

  8. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Saudi Arabia; locality: Raydah; decimalLatitude: 18.204417; decimalLongitude: 42.4124; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 02-20-2014; Record Level: institutionCode: KSMA

  9. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578171; scientificName: Aspidacantha atra; Location: country: Saudi Arabia; locality: Raydah; decimalLatitude: 18.20525; decimalLongitude: 42.4101; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 06-07-2014; Record Level: institutionCode: KSMA

Distribution

AF: Democratic Republic of Congo, Eritrea, Ethiopia, Saudi Arabia [as “south western part”], Tanzania, Uganda, United Arab Emirates, Zimbabwe. PA: Egypt, Israel, Russia, Turkmenistan. [Sources: Lindner (1936), Woodley (2001), Hauser (2008) and El-Hawagry et al. (2017)]

Local distribution and dates of collection (Fig. 2): EGYPT: Eastern Desert: Hurghada (September). Western Desert: Siwa Oasis (August). [Sources: Lindner (1930) and museum material]. SAUDI ARABIA: Asir: Raydah Nature Reserve (February to December). [Sources: El-Hawagry et al. (2017) and collected material]

Figure 2.

Figure 2.

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Distribution map of Aspidacantha atra Kertész and Sternobrithes sp.

Sternobrithes

Loew, 1857

6900F789-4AEF-5661-9FA8-AA1793F98088

https://www.gbif.org/species/1576881

  • Sternobrithes Loew, 1857: 264. Type species Sternobrithes tumidus Loew, by monotypy.

Sternobrithes sp.

A5F755C2-AE97-546D-A32C-99D07F4AA224

https://www.gbif.org/species/1576881

Materials

  1. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1576881; scientificName: Sternobrithes sp.; Location: country: Saudi Arabia; locality: Shada; decimalLatitude: 19.8429; decimalLongitude: 41.3115; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 11-15-2015; Record Level: institutionCode: KSMA

  2. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1576881; scientificName: Sternobrithes sp.; Location: country: Saudi Arabia; locality: Shada; decimalLatitude: 19.8429; decimalLongitude: 41.3115; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 05-05-2015; Record Level: institutionCode: KSMA

  3. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1576881; scientificName: Sternobrithes sp.; Location: country: Saudi Arabia; locality: Shada; decimalLatitude: 19.8388; decimalLongitude: 41.3101; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 11-15-2015; Record Level: institutionCode: KSMA

  4. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1576881; scientificName: Sternobrithes sp.; Location: country: Saudi Arabia; locality: Shada; decimalLatitude: 19.8388; decimalLongitude: 41.3101; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Light trap; eventDate: 09-02-2015; Record Level: institutionCode: KSMA

Distribution

The genus Sternobrithes is widely distributed all over the African continent (Botswana, Burundi, Cameroon, Cape Verde Islands, Democratic Republic of the Congo, Ethiopia, Ghana, Guinea, Ivory Coast, Kenya, Liberia, Malawi, Mozambique, Namibia, Nigeria, Senegal, Sierra Leone, Somalia, South Africa, Tanzania, Togo, Uganda and Zimbabwe. [Sources: Woodley (2001)]

Distribution of Sternobrithes sp.: AF: Kenya, Saudi Arabia [as “south western part”], Yemen.

Local distribution and dates of collection (Fig. 2): SAUDI ARABIA: Al-Baha: Jabal Shada al-A’la Nature Reserve (May to November). Asir: Raydah Nature Reserve (June). [Sources: El-Hawagry et al. (2017) and collected material]

Notes

El-Hawagry et al. (2017) reported this species as Sternobrithes sp. and further research indicated that this is an undescribed species (Fig. 3). As we do not have a male specimen available, we are not yet describing this species here. According the generic key in Hauser et al. (2017b), the difference between Argyrobrithes and Sternobrithes is the colour of the last antennal segment, but further studies indicate that Argyrobrithes is characterised by the dichoptic male eyes and a very long last antennal segment. The two genera are widely distributed through Africa and the Oriental Region and a more thorough revision is needed to clarify the limits of the genera and which species should be included in each genus. Material of this undescribed species are known from Yemen and Kenya.

Figure 3.

Figure 3.

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Sternobrithes sp.: A. female habitus, lateral; B. same, dorsal.

Stratiomyinae

E60C20CF-65A2-5F7A-ACC7-14B58E3F4B2B

Oxycerini

6E494BD5-9091-5FF6-BD63-9B9851B9821F

Oxycera

Meigen, 1803

4E5455B3-8E70-55AA-B7C3-DA5E5E3CD962

https://www.gbif.org/species/1577931

  • Oxycera Meigen, 1803: 265. Type species: Musca hypoleon Linnaeus [= Musca trilineata Linnaeus], by designation of Latreille (1810).

Oxycera orientalis

(Lindner, 1974)

68A7407C-36A4-5310-B2AE-27ACC7656E2C

https://www.gbif.org/species/5078177

  • Heraclina orientalis Lindner, 1974: 103. Type locality: Egypt (Sinai Mountains: Wadi Watir) [Wadi Watir is located in Sinai, Egypt, not in Israel as given in world catalogues].

Materials

  1. Type status: Other material. Occurrence: sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/5078177; scientificName: Oxycera orientalis; Location: country: Egypt; locality: Wadi Watir; decimalLatitude: 29.02147; decimalLongitude: 34.6731; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 09-06-1941; Record Level: institutionCode: EFC

Distribution

PA: Egypt. [Sources: Furth (1983), Woodley (2001) and Badrawy (2006)]

Local distribution and dates of collection (Fig. 4): EGYPT: Sinai: Wadi El-Lega, Wadi Watir (Ain Fortaga) (September to November). [Sources: original description (Lindner 1974) and museum material]

Figure 4.

Figure 4.

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Distribution map of Oxycera orientalis (Lindner) and Oxycera turcica Üstüner & Hasbenli.

Oxycera turcica

Üstüner & Hasbenli, 2005

3D828CE0-4966-5794-9AB6-159D8DBE9D5A

https://www.gbif.org/species/5078139

  • Oxycera turcica Üstüner & Hasbenli, 2005: 163. Type locality: Turkey (Sivas: Sarkisla, Karacaören Village).

Materials

  1. Type status: Other material. Occurrence: recordedBy: El-Hawagry; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/5078139; scientificName: Oxycera turcica; Location: country: Saudi Arabia; locality: Al-Mekhwa; decimalLatitude: 19.798133; decimalLongitude: 41.4107; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 03-15-2012; Record Level: institutionCode: MCCB

Distribution

PA: Saudi Arabia, Turkey. [Sources: Woodley (2011) and collected material]

Local distribution and dates of collection (Fig. 4): SAUDI ARABIA: Al-Baha: Al-Mekhwa (March). [Source: collected material]

Notes

This species is recorded herein for the first time from Saudi Arabia.

Stratiomyini

EC102921-A038-53EC-80F2-0CF9A9A17449

Odontomyia

Meigen, 1803

E15F2AE8-41FF-5252-9A3E-00AA2DA9CAB8

https://www.gbif.org/species/1580809

  • Odontomyia Meigen, 1803: 265. Type species: Musca hydroleon Linnaeus, by designation of Westwood (1840).

Odontomyia angulata

(Panzer, 1798)

DC32212A-8253-58EF-B199-38CBA0008E8A

https://www.gbif.org/species/1581104

  • Stratiomys angulata Panzer, 1798: 19. Type locality: Germany (Nürnberg).

  • Stratiomys vulpina Panzer, 1798: 24. Type locality: Germany (Nürnberg).

  • Stratiomys hydropota Meigen, 1822: 147. Type locality: Europe.

  • Odontomyia latifaciata Macquart, 1834: 248. Type locality: France.

  • Stratiomys brevicornis Loew, 1840: 25. Type locality: Poland (Poznañ).

  • Stratiomys brevicornis Loew, 1840: 557. Type locality: Poland (Poznañ). Preoccupied, primary homonym of Stratiomys brevicornis Loew, 1840.

  • Stratiomys ruficornis Zetterstedt, 1842: 139. Type locality: Sweden (Gotland: Ejsta). Preoccupied, primary homonym of Stratiomyia ruficornis Macquart, 1838.

  • Odontomyia hydrophila Loew, 1846: 486. Type locality: Turkey (Makri) and Italy (Sicily: Siracusa).

  • Eulalia latifasciata Kertész, 1908: 74. Unjustified emendation of Odontomyia latifaciata Macquart.

Distribution

PA: Afghanistan, Albania, Algeria, Austria, Belgium, Bulgaria, China, Czech Republic, Denmark, Egypt, England, Estonia, Finland, France, Germany, Greece, Hungary, Iran, Israel, Italy including Sardinia, Kazakhstan, Morocco, Netherlands, Poland, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Turkey, UAE, former Yugoslavia. [Sources: Woodley (2001), Mason et al. (2009), Hauser (2014) and Yimlahi et al. (2017)]

Local distribution and dates of collection: Unknown.

Notes

This species was listed by Steyskal and El-Bialy (1967), Woodley (2001) and Badrawy (2006) as recorded from Egypt, but no specimens have been collected or preserved in Egyptian museums to confirm these records.

Odontomyia megacephala

Olivier, 1811

2D7491E4-4962-5031-9DA8-1A82AB6A75FD

https://www.gbif.org/species/7863197

  • Odontomyia megacephala Olivier, 1811: 432. Type locality: Egypt (borders of the Nile [as “bords du Nil”]).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Moweileh; decimalLatitude: 30.3924; decimalLongitude: 34.1412; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Sept.1924; Record Level: institutionCode: ESEC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Kosseima; decimalLatitude: 30.90307; decimalLongitude: 34.3836; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 09-10-1924; Record Level: institutionCode: ESEC

  3. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Kharga Oasis; decimalLatitude: 25.25; decimalLongitude: 30.5833; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Oct.1924; Record Level: institutionCode: ESEC

  4. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Fayoum; decimalLatitude: 29.32061; decimalLongitude: 30.818; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Sept.1924; Record Level: institutionCode: ESEC

  5. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Alexandria; decimalLatitude: 31.2129; decimalLongitude: 29.9726; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 07-04-1920; Record Level: institutionCode: EFC

  6. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Nuzha; decimalLatitude: 31.2001; decimalLongitude: 29.9436; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-07-1921; Record Level: institutionCode: EFC

  7. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Nuzha; decimalLatitude: 31.2001; decimalLongitude: 29.9436; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 07-18-1921; Record Level: institutionCode: EFC

  8. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Cleopatra; decimalLatitude: 31.22021; decimalLongitude: 29.9348; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 07-10-1921; Record Level: institutionCode: EFC

  9. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Cleopatra; decimalLatitude: 31.22021; decimalLongitude: 29.9348; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 07-22-1921; Record Level: institutionCode: EFC

  10. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Cleopatra; decimalLatitude: 31.22021; decimalLongitude: 29.9348; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 08-01-1921; Record Level: institutionCode: EFC

  11. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Fayed; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 09-24-1925; Record Level: institutionCode: EFC

  12. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Fayed; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 10-20-1925; Record Level: institutionCode: EFC

  13. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/7863197; scientificName: Odontomyia megacephala; Location: country: Egypt; locality: Wadi El-Natroun; decimalLatitude: 30.3814; decimalLongitude: 30.3441; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 08-06-1929; Record Level: institutionCode: EFC

Distribution

PA: Egypt. [Sources: Lindner (1936), Woodley (2001) and Badrawy (2006)]

Local distribution and dates of collection (Fig. 5): EGYPT: Coastal Strip: Alexandria, Cleopatra, Mariout, Max, Nuzha (May and August). Eastern Desert: Fayed (September). Fayoum: Fayoum City (September). Lower Nile Valley: El-Baragil (April). Sinai: Kosseimah, Moweileh (September). Western Desert: Kharga Oasis, Wadi El-Natroun (August to October). [Sources: original description (Olivier 1811), Badrawy (2006) and museum material]

Figure 5.

Figure 5.

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Distribution map of Odontomyia megacephala Olivier and Odontomyia xanthopus Bezzi.

Odontomyia xanthopus

Bezzi, 1906

2FC4B24A-43C0-510A-953E-49D8BC43F870

https://www.gbif.org/species/1581000

  • Odontomyia xanthopus Bezzi, 1906: 225. Type locality: Eritrea (Adi Ugri) [Adi Ugri belongs now to Eritrea not to Ethiopia as written in world catalogues].

Materials

  1. Type status: Other material. Occurrence: recordedBy: M.Tewfik; sex: 1 male, 3 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1581000; scientificName: Odontomyia xanthopus; Location: country: Egypt; locality: Gebel Ela; decimalLatitude: 22.2008; decimalLongitude: 36.3331; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Apr 1928; Record Level: institutionCode: EFC

Distribution

AF: Eritrea, Malawi, Oman, Yemen (this is the first time this species has been recorded from Oman and Yemen). PA: Egypt, Israel. [Sources: original description (Bezzi 1906), Woodley (2001) and collected material from Yemen and Oman]

Local distribution and dates of collection (Fig. 5): EGYPT: Gebel Elba: ? (April). Sinai: El-Tour, Wadi Hebran (May to July). [Source: Lindner (1930), Lindner (1974) and museum material]

Oplodontha

Rondani, 1863

8AAE7DD9-5859-5413-A56C-050816A7C31E

https://www.gbif.org/species/1577328

  • Oplodontha Rondani, 1863: 78. Type species: Stratiomys viridula Fabricius, by original designation.

Oplodontha minuta

Fabricius, 1794

7F02B74F-4E16-588A-8469-8893168BA54B

https://www.gbif.org/species/1577386

  • Stratiomys minuta Fabricius, 1794: 268. Type locality: India (Tranquebar).

  • Nemotelus pusillus Fabricius, 1794: 271. Type locality: India (Tranquebar).

  • Musca minutior Turton, 1801: 631. New name for Stratiomys minuta Fabricius, 1794.

  • Musca minor Turton, 1801: 655. New name for Nemotelus pusillus Fabricius, 1794.

  • Oxycera indica Brunetti, 1907: 119. Type locality: India (Uttar Pradesh: Bareilly).

  • Odontomyia incompleta Brunetti, 1907: 128. Nomen nudum.

  • Odontomyia ochracea Brunetti, 1907: 129. Type locality: India (Calcutta).

  • Odontomyia submutica Brunetti, 1907: 130. Type locality: India (Siliguri, Calcutta and Tollygunge).

  • Eulalia oasina Lindner, 1925: 150. Type locality: Egypt (Khârga Oasis and Dakhla Oasis).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577386; scientificName: Oplodontha minuta; Location: country: Egypt; locality: Dakhla Oasis; decimalLatitude: 25.5; decimalLongitude: 29.1667; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-13-2018; Record Level: institutionCode: ESEC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577386; scientificName: Oplodontha minuta; Location: country: Egypt; locality: Ein Moussa; decimalLatitude: 29.8667; decimalLongitude: 32.65; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 09-20-1924; Record Level: institutionCode: ESEC

  3. Type status: Other material. Occurrence: recordedBy: Storey; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577386; scientificName: as Eulalia oasina; Location: country: Egypt; locality: Kharga Oasis; decimalLatitude: 25.25; decimalLongitude: 30.5833; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-10-1918; Record Level: institutionCode: EFC

  4. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577386; scientificName: as Eulalia oasina; Location: country: Egypt; locality: Kharga Oasis; decimalLatitude: 25.25; decimalLongitude: 30.5833; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 06-12-1918; Record Level: institutionCode: EFC

Distribution

AF: Socotra Island, United Arab Emirates, Yemen. OR: India, Sri Lanka. PA: Afghanistan, Egypt, Israel. [Sources: original description of O. oasina (Lindner 1925), Woodley (2001) and Tkoč and Rozkošný (2014)]

Local distribution and dates of collection (Fig. 6): EGYPT: Eastern Desert: Ein Moussa (September). Western Desert: Dakhla Oasis, Kharga Oasis (May and June). [Sources: Lindner (1925), Lindner (1930) and museum material]

Figure 6.

Figure 6.

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Distribution map of Oplodontha minuta Fabricius and Oplodontha pulchriceps Loew.

Oplodontha pulchriceps

Loew, 1858

3A60294B-A809-53E8-9EB9-788A7EF5F301

https://www.gbif.org/species/1577339

  • Odontomyia pulchriceps Loew, 1858: 335. Type locality: South Africa (Cape of Good Hope).

  • Odontomyia pulchriceps Loew, 1860: 80. Type locality: South Africa (Cape of Good Hope). Preoccupied, primary homonym of Odontomyia pulchriceps Loew, 1858.

  • Hoplodonta madagascariensis Lindner, 1936: 42. Type locality: Madagascar (Bekily).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577339; scientificName: Oplodontha pulchriceps; Location: country: Saudi Arabia; locality: Jazan; decimalLatitude: 16.9595; decimalLongitude: 42.8348; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 05-11-2018; Record Level: institutionCode: KSMA

  2. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577339; scientificName: Oplodontha pulchriceps; Location: country: Saudi Arabia; locality: Dawmat Al-Jandal; decimalLatitude: 29.809552; decimalLongitude: 39.8749; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Malaise trap; eventDate: 05-26-2018; Record Level: institutionCode: KSMA

  3. Type status: Other material. Occurrence: recordedBy: Aldhafer H.M. et al.; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577339; scientificName: Oplodontha pulchriceps; Location: country: Saudi Arabia; locality: Hassan Ameen farm; decimalLatitude: 28.36661; decimalLongitude: 36.6297; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Sweeping; eventDate: 05-27-2018; Record Level: institutionCode: KSMA

Distribution

AF: Democratic Republic of Congo, Lesotho, Madagascar, Mozambique, Saudi Arabia [as “south western part"], South Africa, United Arab Emirates. PA: Israel, Saudi Arabia [Jawf and Tabouk]. [Sources: Woodley (2001), Hauser (2008) and collected material]

Local distribution and dates of collection (Fig. 6): SAUDI ARABIA: Al-Jawf: Dawmat Al-Jandal (May). Jazan: Jazan (May). Tabouk: Hassan Ameen farm (May). [Source: collected material]

Notes

This species (Fig. 7) and the genus Oplodontha are recorded herein for the first time from Saudi Arabia. It looks similar to O. minuta Fabricius; however, this case requires more clarification in a future study.

Figure 7.

Figure 7.

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Oplodontha pulchriceps Loew: A. female habitus, lateral; B. same, dorsal; C. male habitus, lateral.

Stratiomys

Geoffroy, 1762

8AB342A4-CA7D-5E19-B448-513C3B81C41C

https://www.gbif.org/species/1501902

  • Stratiomys Geoffroy, 1762: 449, 475. Type species: Musca chamaeleon Linnaeus, by designation of I.C.Z.N. (1957).

Stratiomys cenisia

Meigen, 1822

FAACD0CB-FF58-5ABC-A724-35427CE6C111

https://www.gbif.org/species/1577708

  • Stratiomys cenisia Meigen, 1822: 136. Type locality: France (Mont Cenis).

  • Stratiomys flaviventris Loew, 1846: 464. Type locality: Italy (Sicily: Siracusa).

  • Stratiomyia ahngeri Pleske, 1901: 364. Type locality: “Transcaspian Region” [probably = Turkmenistan].

  • Stratiomyia cypria Pleske, 1902: 413. Type locality: Cyprus (Lárnax).

  • Stratiomyia kervillei Villeneuve, 1911: 4. Type locality: Syria (near “lac de Homs”).

  • Stratiomys hispanica ssp. planes James, 1941: 18. Type locality: Iran (Curum, 100 km from Bouchir).

Distribution

PA: Algeria, Armenia, Austria, Bulgaria, Cyprus, Czech Republic, Egypt, France, Germany, Hungary, Iran, Israel, Italy, Kazakhstan, Morocco, Poland, Romania, Russia, Slovakia, Spain, Syria, Tunisia, Turkey, Turkmenistan, Ukraine, former Yugoslavia. [Sources: Woodley (2001) and Yimlahi et al. (2017)]

Local distribution and dates of collection: Unknown.

Notes

This species was listed by Woodley (2001), Badrawy (2006) and Yimlahi et al. (2017) as recorded from Egypt, but no specimens have been collected or preserved in Egyptian museums to verify these records.

Stratiomys deserticolor

Lindner, 1930

F948E65B-43AB-5508-B4DB-AF868AB059B2

https://www.gbif.org/species/4293811

  • Stratiomyia segnis form deserticolor Lindner, 1930: 27. Type locality: Egypt (Siwa Oasis).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/4293811; scientificName: Stratiomys deserticolor; Location: country: Egypt; locality: Kharga Oasis; decimalLatitude: 25.2500; decimalLongitude: 30.5833; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 08-12-1926; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/4293811; scientificName: Stratiomys deserticolor; Location: country: Egypt; locality: Mariout; decimalLatitude: 31.0172; decimalLongitude: 29.76; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Jul 1934; Record Level: institutionCode: EFC

Distribution

PA: Egypt, Saudi Arabia. [Sources: original description (Lindner 1930), Abu-Zoherah et al. (1993) and Woodley (2001)]

Local distribution and dates of collection (Fig. 8): EGYPT: Coastal Strip: Mariout (July). Western Desert: Kharga Oasis, Siwa Oasis (February to August) [Sources: Original description (Lindner 1930) and museum material]. SAUDI ARABIA: localities and dates unknown. [Sources: Abu-Zoherah et al. (1993)]

Figure 8.

Figure 8.

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Distribution map of Stratiomys deserticolor Lindner and Stratiomys longicornis (Scopoli).

Stratiomys longicornis

(Scopoli, 1763)

4129030C-6E96-52DF-A33B-DD25BBBD46C9

https://www.gbif.org/species/1577660

  • Hirtea longicornis Scopoli, 1763: 367. Type locality: “Carniola” [= Slovenija?].

  • Musca tenebricus Harris, 1778: 45. Type locality: England.

  • Stratiomys strigata Fabricius, 1781: 417. Type locality: Italy.

  • Stratiomys tomentosa Schrank, 1803: 94. Type locality: Germany (Ingolstadt).

  • Stratiomys villosa Meigen, 1804: 124. Type locality: Europe.

  • Stratiomys nubeculosa Meigen, 1804: 125. Type locality: Europe.

  • Stratiomys thoracica Fabricius, 1805: 79. Type locality: France.

  • Stratiomys hirtuosa Meigen, 1830: 347. Type locality: Europe.

  • Stratiomys anubis Wiedemann, 1830: 60. Type locality: Egypt.

  • Stratiomyia flavifrons Macquart, 1838: 179. Type locality: “Mesopotamie” [= Iraq].

  • Stratiomys strigata var. pallida Loew, 1840: 25. Type locality: Poland (Poznañ).

  • Stratiomys strigata var. pallida Loew, 1840: 557. Type locality: Poland (Poznañ). Preoccupied, primary homonym of Stratiomys strigata var. pallida Loew, 1840.

  • Stratiomys lambessiana Bigot, 1879: 212. Type locality: Algeria (Lambessa).

  • Stratiomys flavolimbata Costa, 1893: 21. Type locality: Tunisia.

  • Stratiomyia pleskei Wagner, 1903: 108. Type locality: Uzbekistan (Fergana).

  • Stratiomyia segnis Becker, 1906: 8. Type locality: Tunisia (vicinity of Tunis).

  • Hirtea efflatouni Lindner, 1925: 148. Type locality: Egypt (Giza).

  • Stratiomyia (Hirtea) surcoufi Séguy, 1930: 63. Type locality: Algeria (Touggourt).

  • Hirtea surcoufi Séguy, 1932: 125. Type locality: Algeria (Touggourt). Preoccupied, secondary homonym of Stratiomyia surcoufi Séguy, 1930.

  • Stratiomyia longicornis ssp. palaestinensis Lindner, 1937: 64. Type locality: Israel (Upper-Galilee, Kfar-Giladi).

  • Stratiomyia (Hirtea) longicornis ssp. flavoscutellata Lindner, 1940: 24. Type locality: China (Shanxi Province: “Ta-tong-fou”). Preoccupied, primary homonym of Stratiomyia flavoscutellata Wulp, 1885.

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Siala; decimalLatitude: 31.20849; decimalLongitude: 29.8805; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Nov 1913; Record Level: institutionCode: ESEC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Fayed; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Oct 1924; Record Level: institutionCode: ESEC

  3. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Marg; decimalLatitude: 31.0667; decimalLongitude: 30.2167; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 03-24-1918; Record Level: institutionCode: ESEC

  4. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Kosseima; decimalLatitude: 30.90307; decimalLongitude: 34.3836; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Aug 1924; Record Level: institutionCode: ESEC

  5. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Moweileh; decimalLatitude: 30.3924; decimalLongitude: 34.1412; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 08-21-1924; Record Level: institutionCode: ESEC

  6. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 11-25-1913; Record Level: institutionCode: ESEC

  7. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Benha; decimalLatitude: 30.46572; decimalLongitude: 31.18121; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-09-1918; Record Level: institutionCode: ESEC

  8. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Gezeira; decimalLatitude: 30.04596; decimalLongitude: 31.22435; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 09-21-1917; Record Level: institutionCode: ESEC

  9. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1577660; scientificName: Stratiomys longicornis; Location: country: Egypt; locality: Kharga Oasis; decimalLatitude: 25.25; decimalLongitude: 30.5833; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 01-03-1914; Record Level: institutionCode: ESEC

Distribution

PA: Afghanistan, Albania, Algeria, Armenia, Austria, Azerbaijan, Belgium, Bulgaria, China, Cyprus, Czech Republic, Denmark, Egypt, England, France, Germany, Greece, Hungary, Iran, Israel, Italy including Sardinia, Korea, Lithuania, Malta, Mongolia, Morocco, Netherlands, Poland, Portugal, Romania, Russia, Scotland, Slovakia, Slovenia, Spain, Sweden, Switzerland, Tunisia, Turkey, former Yugoslavia. [Sources: Woodley (2001), Mason et al. (2009) and Yimlahi et al. (2017)]

Local distribution and dates of collection (Fig. 8): EGYPT: Coastal Strip: El-Siala, Cleopatra, Dekhela, Nuzha (April to November). Eastern Desert: Fayed (October). Fayoum: Fayoum City, Nazla, Sanhur (April, May and September). Lower Nile Valley and Delta: Abu-Rawash, Benha, Faraskour, Gezeira, Giza, Helwan, Kerdassa, Marg, Sandoub, Shubra (March to November). Sinai: El-Arish, Kosseimah, Moweileh (April and August). Western Desert: Kharga Oasis, Siwa Oasis (January and May). [Sources: Lindner (1925) and museum material]

Stratiomys singularior

(Harris, 1776)

331107BE-392C-566A-A150-01AE52A595BC

https://www.gbif.org/species/4293822

  • Musca singularius Harris, 1778: 45. Type locality: England.

  • Stratiomys furcata Fabricius, 1794: 264. Type locality: Germany.

  • Stratiomys panthaleon Fallén, 1817: 7. Type locality: Sweden.

  • Stratiomys riparia Meigen, 1822: 138. Type locality: Europe.

  • Stratiomys paludosa Siebke, 1863: 149. Type locality: Norway (Dovre Mountains: Jerkin). Preoccupied, primary homonym of Stratiomys paludosa Schummel in Gravenhorst, 1837.

Materials

  1. Type status: Other material. Occurrence: sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/4293822; scientificName: Stratiomys singularior; Location: country: Egypt; locality: Fayoum; decimalLatitude: 29.32061; decimalLongitude: 30.818; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 03-01-1947; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: sex: 2 males; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/4293822; scientificName: Stratiomys singularior; Location: country: Egypt; locality: Girza; decimalLatitude: 29.49968; decimalLongitude: 31.0738; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-09-1950; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/4293822; scientificName: Stratiomys singularior; Location: country: Egypt; locality: Abu-Zaabal; decimalLatitude: 30.24098; decimalLongitude: 31.35211; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 03-19-1950; Record Level: institutionCode: EFC

Distribution

PA: Armenia, Austria, Belgium, Bulgaria, China, Czech Republic, Denmark, Egypt, England, Estonia, Finland, France, Germany, Hungary, Iran, Ireland, Italy, Kazakhstan, Lithuania, Mongolia, Netherlands, Norway, Poland, Romania, Russia, Slovakia, Spain, Sweden, Switzerland, Ukraine, former Yugoslavia. [Sources: Woodley (2001) and Badrawy (2006)]

Local distribution and dates of collection (Fig. 9): EGYPT: Fayoum: Fayoum City, Girza (March and April). Lower Nile Valley & Delta: Abu-Zaabal (March). [Source: Badrawy (2006) and museum material]

Figure 9.

Figure 9.

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Distribution map of Stratiomys singularior (Harris) and Nemotelus anchora Loew.

Nemotelinae

2ABD0C30-4A87-5082-B24B-69C3CF321C1A

Nemotelus

Geoffroy, 1762

49710B4B-7BC2-5B6B-A8E6-E4102126E652

https://www.gbif.org/species/1501964

  • Nemotelus Geoffroy, 1762: 450, 542. Type species: Musca pantherina Linnaeus, by designation of I.C.Z.N. (1957).

Nemotelus

Geoffroy, 1762

059ED454-0BEF-53E2-A1DF-D786D1F51441

Nemotelus (Nemotelus) anchora

Loew, 1846

4D1316EF-9C6F-5EC9-BE6E-C597F3125FD4

https://www.gbif.org/species/1578822

  • Nemotelus anchora Loew, 1846: 429. Type locality: Italy (Sicily: Siracusa).

  • Nemotelus siculus Jaennicke, 1866: 224. Type locality: Italy (Sicily).

  • Nemotelus persicus Pleske in Lindner, 1937: 137. Type locality: Iran (Irak-Adzhemi: Buyun Village).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 32 males, 5 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578822; scientificName: Nemotelus anchora; Location: country: Egypt; locality: Fayoum; decimalLatitude: 29.32061; decimalLongitude: 30.818; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 03-01-1947; Record Level: institutionCode: EFC

Distribution

PA: Algeria, Egypt, Iran, Israel, Italy (including Sicily and Sardinia), Malta, Russia, Tunisia. [Sources: Woodley (2001), Mason et al. (2009) and Mohammad et al. (2009)]

Local distribution and dates of collection (Fig. 9): EGYPT: Fayoum: ? (March). [Source: Mohammad et al. (2009)]

Nemotelus (Nemotelus) candidus

Becker, 1906

D2323DD6-1BD1-51F5-B458-C7959CF4432A

https://www.gbif.org/species/1578675

  • Nemotelus candidus Becker, 1906: 4. Type locality: Algeria (Biskra: Hammam-Salahin).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 4 males, 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578675; scientificName: Nemotelus candidus; Location: country: Egypt; locality: Wadi El-Natroun; decimalLatitude: 30.3814; decimalLongitude: 30.3441; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 08-06-1929; Record Level: institutionCode: EFC

Distribution

PA: Algeria, Egypt. [Sources: Woodley (2001) and Mohammad et al. (2009)]

Local distribution and dates of collection (Fig. 10): EGYPT: Coastal Strip: Dekhela (May and September). Western Desert: Wadi El-Natroun (August). [Source: Mohammad et al. (2009)]

Figure 10.

Figure 10.

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Distribution map of Nemotelus candidus Becker and Nemotelus dentatus Becker.

Nemotelus (Nemotelus) dentatus

Becker, 1902

CA086D1F-AD03-5D5E-8BB4-6CD0A066026E

https://www.gbif.org/species/1578829

  • Nemotelus dentatus Becker, 1902: 7. Type localities: Egypt (Birket Qaroun, Damietta and Alexandria).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Maadi; decimalLatitude: 29.95772; decimalLongitude: 31.2505; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 06-05-1916; Record Level: institutionCode: ESEC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 02-11-1924; Record Level: institutionCode: ESEC

  3. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 02-12-1924; Record Level: institutionCode: ESEC

  4. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 males, 2 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Fayoum; decimalLatitude: 29.32061; decimalLongitude: 30.818; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 03-01-1947; Record Level: institutionCode: EFC

  5. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Marg; decimalLatitude: 31.0667; decimalLongitude: 30.2167; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-01-1923; Record Level: institutionCode: EFC

  6. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-02-1939; Record Level: institutionCode: EFC

  7. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 males; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Kharga Oasis; decimalLatitude: 25.2500; decimalLongitude: 30.5833; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 03-13-1932; Record Level: institutionCode: EFC

  8. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 males; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Mirsa Matrouh; decimalLatitude: 29.5696; decimalLongitude: 26.4194; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-08-1935; Record Level: institutionCode: EFC

  9. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 2 males, 2 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Ismaila; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-07-1924; Record Level: institutionCode: EFC

  10. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Ismaila; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 10-14-1926; Record Level: institutionCode: EFC

  11. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 males; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Ismaila; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 10-14-1926; Record Level: institutionCode: EFC

  12. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Dekhela; decimalLatitude: 31.12098; decimalLongitude: 29.8157; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: May-April 1923; Record Level: institutionCode: EFC

  13. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578829; scientificName: Nemotelus dentatus; Location: country: Egypt; locality: Abu-Zaabal; decimalLatitude: 30.24098; decimalLongitude: 31.3521; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 03-19-1950; Record Level: institutionCode: EFC

Distribution

PA: Egypt. [Source: Woodley (2001)]

Local distribution and dates of collection (Fig. 10): EGYPT: Coastal Strip: Alexandria, Dekhela, Mariout, Mersa Matrouh (March to May). Eastern Desert: Ismailia, El-Kantara (April, May & October). Fayoum: Birket Qaroun, El-Athar [(April & May). Lower Nile Valley and Delta: Abu-Zaabal, Damietta, El-Alag to Marg, El-Gabal El-Asfar, Ezbet El-Nakhl, Helwan, Maadi (February to April & November). Western Desert: Kharga Oasis, Wadi El-Natroun (March). [Sources: original description Becker (1902), Lindner (1925), Mohammad et al. (2009) and museum material]

Nemotelus (Nemotelus) marinus

Becker, 1902

E9E34996-28CE-5C91-9FA0-B7D217AF6AF6

https://www.gbif.org/species/1578792

  • Nemotelus marinus Becker, 1902: 9. Type locality: Egypt (Suez, at the seashore).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578792; scientificName: Nemotelus marinus; Location: country: Egypt; locality: Fayed; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 09-24-1925; Record Level: institutionCode: EFC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578792; scientificName: Nemotelus marinus; Location: country: Egypt; locality: Wadi Hoff; decimalLatitude: 29.8821; decimalLongitude: 31.311; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 08-09-1927; Record Level: institutionCode: EFC

  3. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578792; scientificName: Nemotelus marinus; Location: country: Egypt; locality: Wadi El-Natroun; decimalLatitude: 30.3814; decimalLongitude: 30.3441; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 08-06-1929; Record Level: institutionCode: EFC

Distribution

PA: Egypt. [Source: original description Becker (1902) and Woodley (2001)]

Local distribution and dates of collection (Fig. 11): EGYPT: Eastern Desert: El-Ferdan, Fayed, Ismailia, Suez, Wadi Hoff (April to October). Western Desert: Wadi El Natroun (August). [Sources: original description (Becker 1902), Mohammad et al. (2009) and museum material]

Figure 11.

Figure 11.

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Distribution map of Nemotelus marinus Becker and Nemotelus matrouhensis Mohammad et al.

Nemotelus (Nemotelus) matrouhensis

Mohammad et al., 2009

2097C6B9-034B-5F2F-B299-255920000E98

  • Nemotelus matrouhensis Mohammad, Fadl, Gadalla & Badrawy, 2009: 103. Type locality: Egypt (Mersa Matrouh).

Materials

  1. Type status: Lectotype. Occurrence: recordedBy: H.C.E & M.T.; sex: male; lifeStage: Adult; Taxon: scientificName: Nemotelus matrouhensis; Location: country: Egypt; locality: Mersa Matrouh; decimalLatitude: 29.5696; decimalLongitude: 26.4194; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: July & Aug 1931; Record Level: institutionCode: EFC

Distribution

PA: Egypt. [Source: Woodley (2001)]

Local distribution and dates of collection (Fig. 11): EGYPT: Coastal Strip: Mersa Matrouh (July and August). [Source: original description (Mohammad et al. (2009)]

Notes

Mohammad et al. (2009) published this species and inaccurately cited 13 males and 8 females as "holotype". They did not explicitly select a single specimen as holotype, so all cited specimens are considered as syntypes and a lectotype should be designated. As the illustration of the male genitalia was made from an intact specimen and did not show all the important structures, we are providing an illustration of dissected genitalia (Fig. 12) and a habitus photograph of the specimen (Fig. 13). When one of us (MH) requested a specimen, we were told that this would be a paratype, but because the specimen had no identification or type label at all, a red label was generated and attached to this specimen. This label reads: Paratype ♂/ Nemotelus matrouhensis/ det.: Haitham Badrawy 2009. However, now it is clear that none of the specimens had identification labels and also that this was not a paratype, but a syntype. The male genitalia resemble N. crenatus Egger, 1859 and N. obscuripes Loew, 1871. Lectotype designation: a male specimen in EFC is herein designated as lectotype and is labelled: Mirsa Matruh, July & Aug 1931 (leg. H.C.E. & M.T.), with a red lectotype label. Paralectotypes: 12 males and 8 females, same data.

Figure 12.

Figure 12.

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Nemotelus matrouhensis Mohammad et al.: A. Male terminalia, dorsal view; B. Phallic complex, dorsal view; C. same, lateral view.

Figure 13.

Figure 13.

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Nemotelus matrouhensis Mohammad et al.: A. Male habitus, dorsal view; B. Head, frontal view; C. Head and thorax, frontolateral view; D. Male habitus, lateral view.

Nemotelus (Nemotelus) niloticus

Olivier, 1811

E355FEE9-6CA4-509F-B3B7-A142D0634414

https://www.gbif.org/species/1578794

  • Nemotelus niloticus Olivier, 1811: 183. Type locality: Egypt.

  • Nemotelus fasciatus Olivier, 1811: 183. Type locality: Egypt (“bords du Nil & des canaux qui en dérivent”). Preoccupied by Nemotelus fasciatus Geoffroy in Fourcroy, 1785.

  • Nemotelus albifacies Becker, 1902: 9. Type locality: Egypt (Alexandria).

  • Nemotelus oasis Becker, 1906: 6. Type locality: Algeria (Biskra).

  • Nemotelus theodori Lindner, 1974: 95. Type locality: Israel (Arava Valley: Hazeva).

  • Nemotelus duofasciatus Woodley 2001. Replacement name for N. fasciatus Olivier, 1811.

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Fayoum City; decimalLatitude: 29.32061; decimalLongitude: 30.818; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-12-1918; Record Level: institutionCode: ESEC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Beheira; decimalLatitude: 30.62189; decimalLongitude: 30.48755; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Apr 1924; Record Level: institutionCode: ESEC

  3. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Damietta; decimalLatitude: 31.34595; decimalLongitude: 31.6317; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-11-1918; Record Level: institutionCode: ESEC

  4. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Quisna; decimalLatitude: 30.53514; decimalLongitude: 31.1117; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Mar 1924; Record Level: institutionCode: ESEC

  5. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Mariout; decimalLatitude: 31.0172; decimalLongitude: 29.76; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 07-09-1927; Record Level: institutionCode: EFC

  6. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 4 males, 11 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Mariout (Mallaha); decimalLatitude: 31.0172; decimalLongitude: 29.76; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 06-16-1929; Record Level: institutionCode: EFC

  7. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 males; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Fayoum City; decimalLatitude: 29.32061; decimalLongitude: 30.818; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-23-1943; Record Level: institutionCode: EFC

  8. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 2 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Fayoum City; decimalLatitude: 29.32061; decimalLongitude: 30.818; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-20-1945; Record Level: institutionCode: EFC

  9. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Giza-Fayoum Road; decimalLatitude: 29.5564; decimalLongitude: 30.8869; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-14-1947; Record Level: institutionCode: EFC

  10. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-08-1934; Record Level: institutionCode: EFC

  11. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-14-1934; Record Level: institutionCode: EFC

  12. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-24-1934; Record Level: institutionCode: EFC

  13. Type status: Other material. Occurrence: recordedBy: Farag; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: WadiHoff; decimalLatitude: 29.8821; decimalLongitude: 31.311; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-04-1930; Record Level: institutionCode: EFC

  14. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Ramleh; decimalLatitude: 31.2279; decimalLongitude: 29.976; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 09-17-1921; Record Level: institutionCode: EFC

  15. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Dekhela; decimalLatitude: 31.12098; decimalLongitude: 29.8156; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 06-24-1926; Record Level: institutionCode: EFC

  16. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Dekhela; decimalLatitude: 31.12098; decimalLongitude: 29.8156; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 06-24-1926; Record Level: institutionCode: EFC

  17. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Dekhela; decimalLatitude: 31.12098; decimalLongitude: 29.8156; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 06-18-1929; Record Level: institutionCode: EFC

  18. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578794; scientificName: Nemotelus niloticus; Location: country: Egypt; locality: Ismaila; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 10-14-1926; Record Level: institutionCode: EFC

Distribution

AF: United Arab Emirates. PA: Algeria, Egypt, Israel, Italy (Sardinia), Tunisia. [Sources: Woodley (2001), Mason et al. (2009) and Hauser (2008)]

Local distribution and dates of collection (Fig. 14): EGYPT: Coastal Strip: Alexandria, Dekhela, Mariout (Mallaha), Ramleh (May to September). Eastern Desert: Fayed, Ismailia, Wadi Hoff (April, May and October). Fayoum: Fayoum City, Giza-Fayoum Road (April and May). Lower Nile Valley and Delta: Beheira, Damietta, Helwan, Quisna, Sherbin (March to September). Sinai: Zaranik Protectorate (April). [Sources: original description of N. albifacies (Becker 1902, Mohammad et al. (2009) and museum material]

Figure 14.

Figure 14.

Open in a new tab

Distribution map of Nemotelus niloticus Olivier and Nemotelus notatus Zetterstedt.

Nemotelus (Nemotelus) notatus

Zetterstedt, 1842

B981BA33-3E05-57CF-B384-522534F45E6C

https://www.gbif.org/species/1578742

  • Nemotelus notatus Zetterstedt, 1842: 148. Type locality: Denmark (Copenhagen).

  • Nemotelus brachystomus Loew, 1846: 443. Type locality: Croatia (as “Dalmatien”).

  • Nemotelus leucorhynchus A. Costa, 1884: 61. Type locality: Italy (Sardinia: Stagno di Caliari).

  • Nemotelus nigroaeneus Verhoeff, 1891: 3. Type locality: Germany (Insel Norderney).

  • Nemotelus punctiventris Becker, 1902: 8. Type locality: Egypt (Alexandria).

  • Nemotelus brachystomus form aegyptiacus Lindner, 1925: 146. Type locality: Egypt (Alexandria: Nuzha, Cleopatra, Moharrem Bey, and Aboukir).

  • Nemotelus nigroaeneus ab. portalis Szilády, 1932: 33. Type locality: Germany (Borkum).

  • Nemotelus balearicus Lindner, 1937: 121. Type locality: Spain (Balearic Islands).

  • Nemotelus zernyi Lindner, 1937: 147. Type locality: Spain (Andalucía: Algeciras).

Materials

  1. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Tanta; decimalLatitude: 30.75725; decimalLongitude: 30.9898; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-11-1918; Record Level: institutionCode: ESEC

  2. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Khusus; decimalLatitude: 30.15957; decimalLongitude: 31.3125; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Apr 1918; Record Level: institutionCode: ESEC

  3. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Alag; decimalLatitude: 30.18009; decimalLongitude: 31.3521; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Nov 1924; Record Level: institutionCode: ESEC

  4. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Marg; decimalLatitude: 31.0667; decimalLongitude: 30.2167; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Mar 1918; Record Level: institutionCode: ESEC

  5. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: El-Mahareeq (Kharga); decimalLatitude: 25.618206; decimalLongitude: 30.6468; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Feb 1914; Record Level: institutionCode: ESEC

  6. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Beni Sweif; decimalLatitude: 29.07788; decimalLongitude: 31.10713; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 02-14-1914; Record Level: institutionCode: ESEC

  7. Type status: Other material. Occurrence: recordedBy: Efflatoun; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Kharga Oasis; decimalLatitude: 25.25; decimalLongitude: 30.5833; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: Feb 1914; Record Level: institutionCode: ESEC

  8. Type status: Other material. Occurrence: recordedBy: Priesner; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Kharga Oasis; decimalLatitude: 25.25; decimalLongitude: 30.5833; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 03-10-1924; Record Level: institutionCode: ESEC

  9. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Cleopatra; decimalLatitude: 31.22021; decimalLongitude: 29.9348; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 07-12-1921; Record Level: institutionCode: EFC

  10. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 1 male; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Dekhela; decimalLatitude: 31.12098; decimalLongitude: 29.8156; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 06-24-1926; Record Level: institutionCode: EFC

  11. Type status: Other material. Occurrence: recordedBy: H.C.E & M.T.; sex: 3 males, 5 females; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Dekhela; decimalLatitude: 31.12098; decimalLongitude: 29.8156; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 05-24-1925; Record Level: institutionCode: EFC

  12. Type status: Other material. Occurrence: recordedBy: Efflatoun; sex: 1 male, 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Dekhela; decimalLatitude: 31.12098; decimalLongitude: 29.8156; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 07-09-1927; Record Level: institutionCode: EFC

  13. Type status: Other material. Occurrence: sex: 1 female; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Fayed; decimalLatitude: 30.32382; decimalLongitude: 32.3008; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 07-24-1924; Record Level: institutionCode: EFC

  14. Type status: Other material. Occurrence: recordedBy: Farag; sex: ?; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Helwan; decimalLatitude: 29.8500; decimalLongitude: 31.3333; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 04-08-1934; Record Level: institutionCode: EFC

  15. Type status: Other material. Occurrence: recordedBy: Tewfik; sex: ?; lifeStage: Adult; Taxon: taxonID: https://www.gbif.org/species/1578742; scientificName: Nemotelus notatus; Location: country: Egypt; locality: Wadi Hoff; decimalLatitude: 29.8821; decimalLongitude: 31.311; Identification: identifiedBy: M. El-Hawagry & M. Hauser; dateIdentified: 2020-2021; Event: samplingProtocol: Not given; eventDate: 06-09-1927; Record Level: institutionCode: EFC

Distribution

PA: Albania, Austria, Azerbaijan, Belgium, Bulgaria, Croatia, Cyprus, Denmark, Egypt, England, Finland, France, Germany, Greece, Hungary, Iran, Ireland, Israel, Italy (including Sardinia), Netherlands, Norway, Poland, Romania, Russia, Spain, Sweden, Turkey, former Yugoslavia. [Sources: Woodley (2001), Üstüner and Hasbenli (2005), Mason et al. (2009) and Khaghaninia et al. (2015)]

Local distribution and dates of collection (Fig. 14): EGYPT: Coastal Strip: Abu-Kir, Cleopatra, Dekhela, Mariout, Moharram Bey, Nubar Bey, Nuzha, Ramleh (April to September). Eastern Desert: Ismailia, Wadi Hoff (May to July). Lower Nile Valley and Delta: Beni Sweif, Damietta, El-Alag, Helwan, El-Marg, Khusous, Tanta (February to May and November). Sinai: Zaranik Protectorate (April). Western Desert: Kharga Oasis (February and March). [Sources: original description of N. b. aegyptiacus (Lindner 1925), Mohammad et al. (2009) and museum material]

Supplementary Material

XML Treatment for Stratiomyidae
BDJ.9.e64212-treatment1.xml (1.5KB, xml)
XML Treatment for Stratiomyiinae
BDJ.9.e64212-treatment2.xml (1.5KB, xml)
XML Treatment for Aspidacantha
BDJ.9.e64212-treatment3.xml (2.3KB, xml)
XML Treatment for Aspidacantha atra
BDJ.9.e64212-treatment4.xml (31.2KB, xml)
XML Treatment for Sternobrithes
BDJ.9.e64212-treatment5.xml (2.3KB, xml)
XML Treatment for Sternobrithes sp.
BDJ.9.e64212-treatment6.xml (19.3KB, xml)
XML Treatment for Stratiomyinae
BDJ.9.e64212-treatment7.xml (1.5KB, xml)
XML Treatment for Oxycerini
BDJ.9.e64212-treatment8.xml (1.5KB, xml)
XML Treatment for Oxycera
BDJ.9.e64212-treatment9.xml (3.2KB, xml)
XML Treatment for Oxycera orientalis
BDJ.9.e64212-treatment10.xml (9.7KB, xml)
XML Treatment for Oxycera turcica
BDJ.9.e64212-treatment11.xml (7.5KB, xml)
XML Treatment for Stratiomyini
BDJ.9.e64212-treatment12.xml (1.5KB, xml)
XML Treatment for Odontomyia
BDJ.9.e64212-treatment13.xml (3.3KB, xml)
XML Treatment for Odontomyia angulata
BDJ.9.e64212-treatment14.xml (12.2KB, xml)
XML Treatment for Odontomyia megacephala
BDJ.9.e64212-treatment15.xml (41.3KB, xml)
XML Treatment for Odontomyia xanthopus
BDJ.9.e64212-treatment16.xml (9.4KB, xml)
XML Treatment for Oplodontha
BDJ.9.e64212-treatment17.xml (2.3KB, xml)
XML Treatment for Oplodontha minuta
BDJ.9.e64212-treatment18.xml (20.7KB, xml)
XML Treatment for Oplodontha pulchriceps
BDJ.9.e64212-treatment19.xml (16.4KB, xml)
XML Treatment for Stratiomys
BDJ.9.e64212-treatment20.xml (2.3KB, xml)
XML Treatment for Stratiomys cenisia
BDJ.9.e64212-treatment21.xml (7.7KB, xml)
XML Treatment for Stratiomys deserticolor
BDJ.9.e64212-treatment22.xml (11.6KB, xml)
XML Treatment for Stratiomys longicornis
BDJ.9.e64212-treatment23.xml (39.4KB, xml)
XML Treatment for Stratiomys singularior
BDJ.9.e64212-treatment24.xml (15.2KB, xml)
XML Treatment for Nemotelinae
BDJ.9.e64212-treatment25.xml (1.5KB, xml)
XML Treatment for Nemotelus
BDJ.9.e64212-treatment26.xml (2.3KB, xml)
XML Treatment for Nemotelus
BDJ.9.e64212-treatment27.xml (1.6KB, xml)
XML Treatment for Nemotelus (Nemotelus) anchora
BDJ.9.e64212-treatment28.xml (10.4KB, xml)
XML Treatment for Nemotelus (Nemotelus) candidus
BDJ.9.e64212-treatment29.xml (8.7KB, xml)
XML Treatment for Nemotelus (Nemotelus) dentatus
BDJ.9.e64212-treatment30.xml (41.6KB, xml)
XML Treatment for Nemotelus (Nemotelus) marinus
BDJ.9.e64212-treatment31.xml (14.5KB, xml)
XML Treatment for Nemotelus (Nemotelus) matrouhensis
BDJ.9.e64212-treatment32.xml (12.5KB, xml)
XML Treatment for Nemotelus (Nemotelus) niloticus
BDJ.9.e64212-treatment33.xml (58.1KB, xml)
XML Treatment for Nemotelus (Nemotelus) notatus
BDJ.9.e64212-treatment34.xml (52.2KB, xml)

Acknowledgements

The authors extend their appreciation to the Deanship of Scientific Research at King Saud University for funding this work through research group No [RG-1437-009].

We are grateful to Dr Neal Evenhuis, Bishop Museum, Honolulu, Hawaii, USA and Dr Mohammed Gamal El-Den Nasser, Entomology Department, Faculty of Science, Ain Shams University, Cairo, Egypt for providing critical assistance throughout this study. We thank Haitham Badrawy for the loan of a Nemotelus matrouhensis specimen. We are also grateful to the research team in KSMA for collecting, mounting and photographing specimens.

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Associated Data

This section collects any data citations, data availability statements, or supplementary materials included in this article.

Supplementary Materials

XML Treatment for Stratiomyidae
BDJ.9.e64212-treatment1.xml (1.5KB, xml)
XML Treatment for Stratiomyiinae
BDJ.9.e64212-treatment2.xml (1.5KB, xml)
XML Treatment for Aspidacantha
BDJ.9.e64212-treatment3.xml (2.3KB, xml)
XML Treatment for Aspidacantha atra
BDJ.9.e64212-treatment4.xml (31.2KB, xml)
XML Treatment for Sternobrithes
BDJ.9.e64212-treatment5.xml (2.3KB, xml)
XML Treatment for Sternobrithes sp.
BDJ.9.e64212-treatment6.xml (19.3KB, xml)
XML Treatment for Stratiomyinae
BDJ.9.e64212-treatment7.xml (1.5KB, xml)
XML Treatment for Oxycerini
BDJ.9.e64212-treatment8.xml (1.5KB, xml)
XML Treatment for Oxycera
BDJ.9.e64212-treatment9.xml (3.2KB, xml)
XML Treatment for Oxycera orientalis
BDJ.9.e64212-treatment10.xml (9.7KB, xml)
XML Treatment for Oxycera turcica
BDJ.9.e64212-treatment11.xml (7.5KB, xml)
XML Treatment for Stratiomyini
BDJ.9.e64212-treatment12.xml (1.5KB, xml)
XML Treatment for Odontomyia
BDJ.9.e64212-treatment13.xml (3.3KB, xml)
XML Treatment for Odontomyia angulata
BDJ.9.e64212-treatment14.xml (12.2KB, xml)
XML Treatment for Odontomyia megacephala
BDJ.9.e64212-treatment15.xml (41.3KB, xml)
XML Treatment for Odontomyia xanthopus
BDJ.9.e64212-treatment16.xml (9.4KB, xml)
XML Treatment for Oplodontha
BDJ.9.e64212-treatment17.xml (2.3KB, xml)
XML Treatment for Oplodontha minuta
BDJ.9.e64212-treatment18.xml (20.7KB, xml)
XML Treatment for Oplodontha pulchriceps
BDJ.9.e64212-treatment19.xml (16.4KB, xml)
XML Treatment for Stratiomys
BDJ.9.e64212-treatment20.xml (2.3KB, xml)
XML Treatment for Stratiomys cenisia
BDJ.9.e64212-treatment21.xml (7.7KB, xml)
XML Treatment for Stratiomys deserticolor
BDJ.9.e64212-treatment22.xml (11.6KB, xml)
XML Treatment for Stratiomys longicornis
BDJ.9.e64212-treatment23.xml (39.4KB, xml)
XML Treatment for Stratiomys singularior
BDJ.9.e64212-treatment24.xml (15.2KB, xml)
XML Treatment for Nemotelinae
BDJ.9.e64212-treatment25.xml (1.5KB, xml)
XML Treatment for Nemotelus
BDJ.9.e64212-treatment26.xml (2.3KB, xml)
XML Treatment for Nemotelus
BDJ.9.e64212-treatment27.xml (1.6KB, xml)
XML Treatment for Nemotelus (Nemotelus) anchora
BDJ.9.e64212-treatment28.xml (10.4KB, xml)
XML Treatment for Nemotelus (Nemotelus) candidus
BDJ.9.e64212-treatment29.xml (8.7KB, xml)
XML Treatment for Nemotelus (Nemotelus) dentatus
BDJ.9.e64212-treatment30.xml (41.6KB, xml)
XML Treatment for Nemotelus (Nemotelus) marinus
BDJ.9.e64212-treatment31.xml (14.5KB, xml)
XML Treatment for Nemotelus (Nemotelus) matrouhensis
BDJ.9.e64212-treatment32.xml (12.5KB, xml)
XML Treatment for Nemotelus (Nemotelus) niloticus
BDJ.9.e64212-treatment33.xml (58.1KB, xml)
XML Treatment for Nemotelus (Nemotelus) notatus
BDJ.9.e64212-treatment34.xml (52.2KB, xml)

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