Review of genus-group names in the family Tenebrionidae (Insecta, Coleoptera)

Abstract

A review of genus-group names for darkling beetles in the family Tenebrionidae (Insecta: Coleoptera) is presented. A catalogue of 4122 nomenclaturally available genus-group names, representing 2307 valid genera (33 of which are extinct) and 761 valid subgenera, is given. For each name the author, date, page number, gender, type species, type fixation, current status, and first synonymy (when the name is a synonym) are provided. Genus-group names in this family are also recorded in a classification framework, along with data on the distribution of valid genera and subgenera within major biogeographical realms. A list of 535 unavailable genus-group names (e.g., incorrect subsequent spellings) is included. Notes on the date of publication of references cited herein are given, when known.

The following genera and subgenera are made available for the first time: Anemiadena Bouchard & Bousquet, subgen. nov. (in Cheirodes Gené, 1839), Armigena Bouchard & Bousquet, subgen. nov. (in Nesogena Mäklin, 1863), Debeauxiella Bouchard & Bousquet, subgen. nov. (in Hyperops Eschscholtz, 1831), Hyperopsis Bouchard & Bousquet, subgen. nov. (in Hyperops Eschscholtz, 1831), Linio Bouchard & Bousquet, subgen. nov. (in Nilio Latreille, 1802), Matthewsotys Bouchard & Bousquet, gen. nov., Neosolenopistoma Bouchard & Bousquet, subgen. nov. (in Eurynotus W. Kirby, 1819), Paragena Bouchard & Bousquet, subgen. nov. (in Nesogena Mäklin, 1863), Paulianaria Bouchard & Bousquet, gen. nov., Phyllechus Bouchard & Bousquet, gen. nov., Prorhytinota Bouchard & Bousquet, subgen. nov. (in Rhytinota Eschscholtz, 1831), Pseudorozonia Bouchard & Bousquet, subgen. nov. (in Rozonia Fairmaire, 1888), Pseudothinobatis Bouchard & Bousquet, gen. nov., Rhytinopsis Bouchard & Bousquet, subgen. nov. (in Thalpophilodes Strand, 1942), Rhytistena Bouchard & Bousquet, subgen. nov. (in Rhytinota Eschscholtz, 1831), Spinosdara Bouchard & Bousquet, subgen. nov. (in Osdara Walker, 1858), Spongesmia Bouchard & Bousquet, subgen. nov. (in Adesmia Fischer, 1822), and Zambesmia Bouchard & Bousquet, subgen. nov. (in Adesmia Fischer, 1822).

The names Adeps Gistel, 1857 and Adepsion Strand, 1917 syn. nov. [= Tetraphyllus Laporte & Brullé, 1831], Asyrmatus Canzoneri, 1959 syn. nov. [= Pystelops Gozis, 1910], Euzadenos Koch, 1956 syn. nov. [= Selenepistoma Dejean, 1834], Gondwanodilamus Kaszab, 1969 syn. nov. [= Conibius J.L. LeConte, 1851], Gyrinodes Fauvel, 1897 syn. nov. [= Nesotes Allard, 1876], Helopondrus Reitter, 1922 syn. nov. [= Horistelops Gozis, 1910], Hybonotus Dejean, 1834 syn. nov. [= Damatris Laporte, 1840], Iphthimera Reitter, 1916 syn. nov. [= Metriopus Solier, 1835], Lagriomima Pic, 1950 syn. nov. [= Neogria Borchmann, 1911], Orphelops Gozis, 1910 syn. nov. [= Nalassus Mulsant, 1854], Phymatium Billberg, 1820 syn. nov. [= Cryptochile Latreille, 1828], Prosoblapsia Skopin & Kaszab, 1978 syn. nov. [= Genoblaps Bauer, 1921], and Pseudopimelia Gebler, 1859 syn. nov. [= Lasiostola Dejean, 1834] are established as new synonyms (valid names in square brackets). Anachayus Bouchard & Bousquet, nom. nov. is proposed as a replacement name for Chatanayus Ardoin, 1957, Genateropa Bouchard & Bousquet, nom. nov. as a replacement name for Apterogena Ardoin, 1962, Hemipristula Bouchard & Bousquet, nom. nov. as a replacement name for Hemipristis Kolbe, 1903, Kochotella Bouchard & Bousquet, nom. nov. as a replacement name for Millotella Koch, 1962, Medvedevoblaps Bouchard & Bousquet, nom. nov. as a replacement name for Protoblaps G.S. Medvedev, 1998, and Subpterocoma Bouchard & Bousquet, nom. nov. is proposed as a replacement name for Pseudopimelia Motschulsky, 1860. Neoeutrapela Bousquet & Bouchard, 2013 is downgraded to a subgenus (stat. nov.) of Impressosora Pic, 1952. Anchomma J.L. LeConte, 1858 is placed in Stenosini: Dichillina (previously in Pimeliinae: Anepsiini); Entypodera Gerstaecker, 1871, Impressosora Pic, 1952 and Xanthalia Fairmaire, 1894 are placed in Lagriinae: Lagriini: Statirina (previously in Lagriinae: Lagriini: Lagriina); Loxostethus Triplehorn, 1962 is placed in Diaperinae: Diaperini: Diaperina (previously in Diaperinae: Diaperini: Adelinina); Periphanodes Gebien, 1943 is placed in Stenochiinae: Cnodalonini (previously in Tenebrioninae: Helopini); Zadenos Laporte, 1840 is downgraded to a subgenus (stat. nov.) of the older name Selenepistoma Dejean, 1834.

The type species [placed in square brackets] of the following available genus-group names are designated for the first time: Allostrongylium Kolbe, 1896 [Allostrongyliumsilvestre Kolbe, 1896], Auristira Borchmann, 1916 [Auristiraoctocostata Borchmann, 1916], Blapidocampsia Pic, 1919 [Campsiapallidipes Pic, 1918], Cerostena Solier, 1836 [Cerostenadeplanata Solier, 1836], Coracostira Fairmaire, 1899 [Coracostiraarmipes Fairmaire, 1899], Dischidus Kolbe, 1886 [Helopssinuatus Fabricius, 1801], Eccoptostoma Gebien, 1913 [Taraxidesruficrus Fairmaire, 1894], Ellaemus Pascoe, 1866 [Emcephalussubmaculatus Brême, 1842], Epeurycaulus Kolbe, 1902 [Epeurycaulusaldabricus Kolbe, 1902], Euschatia Solier, 1851 [Euschatiaproxima Solier, 1851], Heliocaes Bedel, 1906 [Blaps emarginata Fabricius, 1792], Hemipristis Kolbe, 1903 [Hemipristisukamia Kolbe, 1903], Iphthimera Reitter, 1916 [Stenocararuficornis Solier, 1835], Isopedus Stein, 1877 [Helopstenebrioides Germar, 1813], Malacova Fairmaire, 1898 [Malacovabicolor Fairmaire, 1898], Modicodisema Pic, 1917 [Disemasubopaca Pic, 1912], Peltadesmia Kuntzen, 1916 [Metriopusplatynotus Gerstaecker, 1854], Phymatium Billberg, 1820 [Pimeliamaculata Fabricius, 1781], Podoces Péringuey, 1886 [Podocesgranosula Péringuey, 1886], Pseuduroplatopsis Pic, 1913 [Borchmanniajavana Pic, 1913], Pteraulus Solier, 1848 [Pteraulussulcatipennis Solier, 1848], Sciaca Solier, 1835 [Hylithusdisctinctus Solier, 1835], Sterces Champion, 1891 [Stercesviolaceipennis Champion, 1891] and Teremenes Carter, 1914 [Tenebriolongipennis Hope, 1843].

Evidence suggests that some type species were misidentified. In these instances, information on the misidentification is provided and, in the following cases, the taxonomic species actually involved is fixed as the type species [placed in square brackets] following requirements in Article 70.3 of the International Code of Zoological Nomenclature: Accanthopus Dejean, 1821 [Tenebriovelikensis Piller & Mitterpacher, 1783], Becvaramarygmus Masumoto, 1999 [Dietysusnodicornis Gravely, 1915], Heterophaga Dejean, 1834 [Opatrumlaevigatum Fabricius, 1781], Laena Dejean, 1821, [Scaurusviennensis Sturm, 1807], Margus Dejean, 1834 [Colydiumcastaneum Herbst, 1797], Pachycera Eschscholtz, 1831 [Tenebriobuprestoides Fabricius, 1781], Saragus Erichson, 1842 [Celibecostata Solier, 1848], Stene Stephens, 1829 [Colydiumcastaneum Herbst, 1797], Stenosis Herbst, 1799 [Tageniaintermedia Solier, 1838] and Tentyriopsis Gebien, 1928 [Tentyriopsispertyi Gebien, 1940].

The following First Reviser actions are proposed to fix the precedence of names or nomenclatural acts (rejected name or act in square brackets): Stenosisciliaris Gebien, 1920 as the type species for Afronosis G.S. Medvedev, 1995 [Stenosisleontjevi G.S. Medvedev, 1995], Alienoplonyx Bremer, 2019 [Alienolonyx], Amblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 2018 [Amplypteraca], Caenocrypticoides Kaszab, 1969 [Caenocripticoides], Deriles Motschulsky, 1872 [Derilis], Eccoptostira Borchmann, 1936 [Ecoptostira], †Eodromus Haupt, 1950 [†Edromus], Eutelus Solier, 1843 [Lutelus], Euthriptera Reitter, 1893 [Enthriptera], Meglyphus Motschulsky, 1872 [Megliphus], Microtelopsis Koch, 1940 [Extetranosis Koch, 1940, Hypermicrotelopsis Koch, 1940], Neandrosus Pic, 1921 [Neoandrosus], Nodosogylium Pic, 1951 [Nodosogilium], Notiolesthus Motschulsky, 1872 [Notiolosthus], Pseudeucyrtus Pic, 1916 [Pseudocyrtus], Pseudotrichoplatyscelis Kaszab, 1960 [Pseudotrichoplatynoscelis and Pseudotrichoplatycelis], Rhydimorpha Koch, 1943 [Rhytimorpha], Rhophobas Motschulsky, 1872 [Rophobas], Rhyssochiton Gray, 1831 [Ryssocheton and Ryssochiton], Sphaerotidius Kaszab, 1941 [Spaerotidius], Stira Agassiz, 1846 (Mollusca) [Stira Agassiz, 1846 (Coleoptera)], Sulpiusoma Ferrer, 2006 [Sulpiosoma] and Taenobates Motschulsky, 1872 [Taeniobates].

Supporting evidence is provided for the conservation of usage of Cyphaleus Westwood, 1841 nomen protectum over Chrysobalus Boisduval, 1835 nomen oblitum.

Introduction

The last world catalogue of darkling beetles in the family Tenebrionidae Latreille, 1802 (Insecta: Coleoptera) was published by Gebien (1937a, 1938a, 1939, 1940, 1941, 1942a, 1943, 1948) more than 70 years ago. At the time, entire lineages were treated by coleopterists as separate families (e.g., Alleculidae Laporte, 1840, Lagriidae Latreille, 1825 (1820), Nilionidae Oken, 1843, Cossyphodidae Wasmann, 1899) and were therefore excluded from Gebien’s “Katalog der Tenebrioniden”. Since then, important studies on the relationships of taxa classified within Tenebrionidae and their close relatives, based on morphological (e.g., Doyen and Tschinkel 1982) and molecular data (e.g., Kergoat et al. 2014b; Kamiński et al. 2021b), have led to significant improvements in our understanding of the limits of the family and the relationships of major clades within the family. In addition, several taxa traditionally included in Tenebrionidae have now been transferred into closely related families (Table 1).

Table 1.

Genus-group names transferred from Tenebrionidae to other families. An example of the current placement of each genus is given in the “Source” column.

Genus	Current placement	Source
Acotulus Reitter, 1891	Zopheridae	Schuh (2020: 68)
†Adelidium Tillyard, 1918	Coleoptera incertae sedis	Nabozhenko and Bukejs (2021: 55)
Aegialites Mannerheim, 1853	Salpingidae	Lawrence et al. (2010b: 722)
Ageonoma Pascoe, 1866	Zopheridae	Foley and Ivie (2008b: 37)
Agnathus Germar, 1818	Pyrochroidae	Young and Pollock (2010: 715)
Anaplopus Blackburn, 1890	Pythidae	Pollock (2010b: 709)
Antarcticodomus Brookes, 1951	Salpingidae	Lawrence et al. (2010b: 722)
Apelta Montrouzier 1864	Corylophidae	Fauvel (1903: 289)
Aposyla Pascoe, 1862	Boridae	Lawrence and Pollock (1994: 37)
Archeocrypticus Kaszab, 1964	Archeocrypticidae	Gimmel et al. (2018: 275)
Arthopus Sharp, 1876	Ulodidae	Leschen et al. (2016: 468)
Bancous Pic, 1946	Erotylidae	Skelley and Alonso-Zarazaga (2003: 107)
Boros Herbst, 1797	Boridae	Bouchard et al. (2011: 444)
Brachyhelops Fairmaire, 1885	Chrysomelidae	Reid (2014: 248)
Brouniphylax Strand, 1943	Ulodidae	Leschen et al. (2016: 468)
Caanthus Champion, 1894	Zopheridae	Ślipiński and Lawrence (1997: 372)
Calophthalmus J. Thomson, 1860	Mycteridae	Pollock (2010a: 693, as Stilpnonotus)
Chalcodrya Redtenbacher, 1867	Chalcodryidae	Lawrence and Leschen (2010: 567)
Chanopterus Boheman, 1858	Promecheilidae	Lawrence et al. (2010a: 563)
Chitoniscus C.O. Waterhouse, 1875	Promecheilidae	Kulzer (1963: 602)
†Cistelites Heer, 1864	Coleoptera incertae sedis	Nabozhenko (2019: 8)
Cleteus Fairmaire, 1906	Zopheridae	Freude (1974: 258)
Coeloderes Mulsant and Rey, 1859	Zopheridae	Bedel (1887: 199)
Cotulades Pascoe, 1860	Zopheridae	Ślipiński and Lawrence (2010: 549)
Cycloderus Solier, 1851	Pyrochroidae	Young and Pollock (2010: 715)
Dacoderus J.L. LeConte, 1859	Salpingidae	Bouchard et al. (2011: 446)
Darwinella Enderlein, 1912	Promecheilidae	Lawrence et al. (2010a: 563)
Deridea Westwood, 1875	Meloidae	Bouchard et al. (2011: 437)
Diacalla Pascoe, 1863	Anthicidae	Chandler (2010: 730)
Diacallina Champion, 1916	Anthicidae	Chandler (2010: 730)
Dipsaconia Pascoe, 1860	Ulodidae	Leschen et al. (2016: 468)
Docalis Pascoe, 1860	Zopheridae	Ślipiński and Lawrence (2010: 549)
Egestriomima Champion, 1916	Anthicidae	Chandler (2010: 730)
Elascus Pascoe, 1860	Zopheridae	Ślipiński and Lawrence (1999: 33)
Endophloeus Dejean, 1834	Zopheridae	Schuh (2020: 71)
Enhypnon Carter, 1919	Zopheridae	Turco et al. (2013: 371)
Enneboeopsis Champion, 1894	Archeocrypticidae	Gimmel et al. (2018: 275)
Enneboeus Waterhouse, 1878	Archeocrypticidae	Gimmel et al. (2018: 276)
Eucistela Carter, 1922	Pyrochroidae	Bousquet et al. (2015: 131)
Eurypus Kirby, 1819	Mycteridae	Pollock (2010a: 693)
Exohadrus Broun, 1893	Ulodidae	Leschen et al. (2016: 468)
Falsoxanthalia Pic,1934	Tetratomidae	Nikitsky (2020: 44)
Ganyme Pascoe, 1869	Ulodidae	Leschen et al. (2016: 468)
†Helopides Roemer, 1876	Cupedidae incertae sedis	Kirejtshuk et al. (2016: 146)
Hymaea Pascoe, 1869	Phloeostichidae	Bouchard et al. (2011: 362)
Hydromedion Waterhouse, 1875	Promecheilidae	Lawrence et al. (2010a: 563)
Ictistygna Pascoe, 1866	Anthicidae	Chandler (2010: 730)
Ictistygnina Champion, 1916	Anthicidae	Chandler (2010: 730)
Ischyomius Chevrolat, 1878	Pythidae	Pollock (2010b: 709)
Latometus Erichson, 1842	Zopheridae	Ślipiński and Lawrence (2010: 556)
Loboglossa Solier, 1851	Mycteridae	Pollock (2010a: 693)
Malacodrya Sharp, 1886	Promecheilidae	Lawrence et al. (2020: 28)
Megazopherus Casey, 1907	Zopheridae	Bousquet et al. (2018: 19)
Melytra Pascoe, 1869	Promecheilidae	Lawrence et al. (2010a: 563)
†Menephiloides Fujiyama, 1973	Coleoptera incertae sedis	Nabozhenko and Bukejs (2021: 55)
Meralius Casey, 1907	Zopheridae	Bousquet et al. (2018: 19)
Meryx Latreille, 1802	Ulodidae	Leschen et al. (2016: 468)
†Mesothoris Tillyard, 1916	Cupedidae incertae sedis	Kirejtshuk et al. (2016: 148)
Micruloma Carter, 1919	Cerylonidae	Doyen et al. (1990: 238)
Mnionophilus Carter, 1919	Zopheridae	Turco et al. (2013: 371)
Mnionychus Carter, 1926	Zopheridae	Lawrence and Ślipiński (2013: 12)
Mylops Fairmaire, 1884	Promecheilidae	Lawrence et al. (2010a: 563, as Hydromedion)
Neboissianus Kaszab, 1981	Archeocrypticidae	Gimmel et al. (2018: 276)
Noserinus Casey, 1907	Zopheridae	Foley and Ivie (2008: 18)
Noserodes Casey, 1907	Zopheridae	Bousquet et al. (2018: 19)
Noserus J.L. LeConte, 1862	Zopheridae	Bousquet et al. (2018: 19)
Nosoderma Solier, 1841	Zopheridae	Bousquet et al. (2018: 19)
Notocerastes Carter, 1926	Ulodidae	Leschen et al. (2016: 468)
Notolea Carter, 1915	Promecheilidae	Lawrence et al. (2010a: 566)
Ocholissa Pascoe, 1863	Salpingidae	Lawrence et al. (2010b: 724)
Onysius Broun, 1886	Promecheilidae	Lawrence et al. (2020: 33)
Parahelops Waterhouse, 1875	Promecheilidae	Lawrence et al. (2010a: 563)
Paraphylax Broun, 1880	Ulodidae	Leschen et al. (2016: 468)
Parenneboeus Kaszab, 1981	Archeocrypticidae	Gimmel et al. (2018: 276)
Perimylops Müller 1884	Promecheilidae	Lawrence et al. (2010a: 563)
Phaennis Champion, 1894	Ulodidae	Leschen et al. (2016: 469)
Phaeogala Fairmaire, 1896	Mycteridae	Pollock (2010a: 693)
Phellopsis J.L. LeConte, 1862	Zopheridae	Bousquet et al. (2018: 19)
Philpottia Broun, 1915	Chalcodryidae	Lawrence and Leschen (2010: 567)
Phloeodes J.L. LeConte, 1862	Zopheridae	Bousquet et al. (2018: 19)
PhloeopsidiusGebien, 1925	Zopheridae	Ivie et al. (2016: 780)
Phycosecis Pascoe, 1875	Phycosecidae	Bouchard et al. (2011: 349)
Promecheilus Solier, 1851	Promecheilidae	Lawrence et al. (2010a: 563)
Pseudenneboeus Kaszab, 1981	Archeocrypticidae	Gimmel et al. (2018: 277)
Pseudonosoderma Heyden, 1885	Zopheridae	Foley and Ivie (2008a: 8)
Psilonycha Fåhraeus, 1870	Scraptiidae	Bousquet et al. (2015: 131)
Pteroderes Germain, 1894	Ulodidae	Leschen et al. (2016: 469)
Pycnidium Erichson, 1847	Leiodidae	A. Newton (pers. comm.)
Rhyssopera Pascoe, 1860	Ulodidae	Leschen et al. (2016: 468)
Rygmodus White, 1846	Hydrophilidae	Bouchard et al. (2011: 158)
Scoriaderma Fairmaire, 1894	Zopheridae	Foley and Ivie (2008b: 21)
Sesaspis Casey, 1907	Zopheridae	Bousquet et al. (2018: 19)
Sirrhas Champion, 1893	Promecheilidae	Lawrence et al. (2010a: 563)
Sivacrypticus Kaszab, 1964	Archeocrypticidae	Gimmel et al. (2018: 277)
Stilpnonotus Gray, 1832	Mycteridae	Pollock (2010a: 693)
Sympiezocera P.H. Lucas, 1851	Cerambycidae	Danilevsky and Lin (2020: 211)
Synercticus Newman, 1842	Boridae	Pollock (2010: 699)
Synopticus J. Thomson, 1858	Tenebrionoidea incertae sedis	Bremer (2013a: 72)
Syrphetodes Broun, 1875	Ulodidae	Leschen et al. (2016: 469)
Szekessya Kaszab, 1955	Salpingidae	Lawrence et al. (2010b: 727)
Tarphiomimus Wollaston, 1873	Zopheridae	Ślipiński and Lawrence (2010: 553)
Trachelolagria Pic, 1941	Cleridae	Merkl (2004: 285)
Trachyderas Philippi and Philippi, 1864	Ulodidae	Leschen et al. (2016: 469)
Trachyderastes Kaszab, 1982	Ulodidae	Leschen et al. (2016: 469)
Tretothorax Lea, 1911	Salpingidae	Lawrence et al. (2010b: 722)
Trictenotoma Gray, 1831	Trictenotomidae	Bouchard et al. (2011: 444)
Ulodes Erichson, 1942	Ulodidae	Leschen et al. (2016: 469)
Ulodica Pascoe, 1769	Ulodidae	Leschen et al. (2016: 469)
†Ulomites Tillyard, 1916	Coleoptera incertae sedis	Martin (2010: 939)
Uloporus Casey, 1889	Archeocrypticidae	Gimmel et al. (2018: 276)
Usechimorpha Blaisdell, 1929	Zopheridae	Ślipiński and Lawrence (2010: 552)
Usechus Motschulsky, 1845	Zopheridae	Bousquet et al. (2018: 19)
Verodes Casey, 1907	Zopheridae	Bousquet et al. (2018: 19)
Wattianus Kaszab, 1981	Archeocrypticidae	Gimmel et al. (2018: 278)
Zopherinus Casey, 1907	Zopheridae	Bousquet et al. (2018: 19)
Zopherodes Casey, 1907	Zopheridae	Bousquet et al. (2018: 19)
Zopherosis White, 1859	Zopheridae	Ślipiński and Lawrence (2010: 549)
Zopherus Laporte, 1840	Zopheridae	Bousquet et al. (2018: 19)

With more than 30 000 described species worldwide (RLA, unpubl. data) and many new species described each year, the family Tenebrionidae has been described as “hyperdiverse” (Kergoat et al. 2014a). This overwhelming diversity, combined with significant changes in classification over the last decades, has hindered the production of recent catalogues at a world scale. Although some catalogues that cover large geographic areas have been published (e.g., Matthews and Bouchard 2008; Bousquet et al. 2018; Iwan et al. 2020), the nomenclatural information contained therein often differs in scope and major regional gaps of knowledge (e.g., fauna in the Afrotropical and Neotropical biogeographic realms) still remain.

The main objective of this publication is to provide a nomenclatural review of all genus-group names in the family Tenebrionidae. A table that includes an up-to-date synthesis of the classification of available genus-group names, along with the distribution of each valid genus and subgenus within the world’s major biogeographical realms, is included. Following bibliographic research, dating of references cited in this work is provided (whenever data is available) in order to establish the priority of genus-group names and therefore promote their nomenclatural stability in the future.

Methods

Nomenclatural data

All nomenclaturally available genus-group names in the family Tenebrionidae are listed alphabetically. For each name the author, year of publication, page number, gender (in square brackets: M = masculine, F = feminine, N = neuter), type species, typification, current status, and first synonymy (when the name is a synonym) are provided. The type species and type species fixation given for unjustified emendations and replacement names are identical to those of the available names they replace (International Commission on Zoological Nomenclature (henceforth ICZN) 1999, Article 67.8) and are listed following “Type species [automatic]:”. We follow previous authors (e.g., Alonso-Zarazaga and Lyal 2009; Bousquet et al. 2015, 2018) in considering type-species designations in R. Lucas (1920) potentially valid when a single species is listed under a particular genus-group name. According to ICZN (1999, Article 67.2.5) “a nominal species is deemed not to be originally included if it was doubtfully or conditionally included, or was cited as a species inquirenda, or as a species incertae sedis.” For this catalogue, any nominal species associated with a new genus with a question mark “?” was deemed not to be originally included. In accordance with Article 11.9.3.2 (ICZN 1999) we used the correct spelling of the species name (i.e., in agreement with the gender of the generic name) for all species-group name combinations. The author of each new synonym or new placement is given in square brackets (e.g., “[RLA]”) in the alphabetical list of available genus-group names. We included all genus-group names known to us up to the date of publication of this article.

While the identification of the type species is generally assumed to be correct (ICZN 1999, Article 70.1), evidence suggests that the type species of a small number of genus-group names were misidentified. In these cases, we include information about the misidentification and select the species (either the nominal species previously cited as the type species or the taxonomic species actually involved) that will best serve stability and universality in our opinion (ICZN 1999, Article 70.3). If we select the taxonomic species actually involved as type species, an action that could not have been taken under previous editions of the International Code of Zoological Nomenclature and that must be accompanied with a reference to Article 73 (ICZN 1999), it is given in the format “fixed herein (ICZN 1999, Article 70.3) as Tenebriovelikensis Piller and Mitterpacher, 1783, misidentified as Tenebriocaraboides Linnaeus, 1758 in the original designation by monotypy in Dejean (1821).”

In some cases, we have noted that the original combination of the accepted name of the type species is a primary homonym of an older species name (e.g., Helopstenebrioides Germar, 1813, the type species of Isopedus Stein, 1877, is a junior primary homonym of Helopstenebrioides Palisot de Beauvois, 1812). We refrain proposing replacement names for two raisons: first, the status of some senior homonyms, which in most cases involved pre-1820 names, is uncertain; second, in some cases the two homonyms belong to taxa in use but the names apply to taxa not considered congeneric after 1899, and the Commission (ICZN 1999: Article 23.9.5) mandates that the author(s) must not automatically replace the junior homonym but instead should refer to the Commission for a ruling. We hope that experts on the taxonomy of the taxa involved will look at all evidence and propose solutions to remove the homonymies as needed.

Genus-group names encountered in the literature that are not nomenclaturally available are listed in Appendix 1. Reasons for assigning the status of “nomenclaturally unavailable” for these names include: name proposed as an unmodified vernacular word, not appropriately latinized (ICZN 1999, Article 11.3, Recommendation 11A); name not used as valid when proposed (ICZN 1999, Article 11.5); name first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym (ICZN 1999, Article 11.6); name published before 1931 without a description, a definition or an indication (Article 12.1); name proposed as a replacement name for an unavailable name (ICZN 1999, Article 12.2.3); name published after 1930 without a description, a definition or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1); name published after 1930 without a type species (ICZN 1999, Article 13.3); replacement name published after 1930, for a name without valid typification, without designating a type species (ICZN 1999, Article 13.3.1); new nominal genus or subgenus published in a combined description with a new species not marked by “gen. nov., spec. nov.” or an equivalent expression (ICZN 1999, Article 13.4); name published after 1999 and not explicitly indicated as intentionally new (ICZN 1999, Article 16.1); rejected alternative original spelling (ICZN 1999, Article 24.2.3); original spelling corrected in the same work (ICZN 1999, Article 32.5.1.1); incorrect subsequent spelling that is not in prevailing usage (ICZN 1999, Article 33.3).

The symbol for fossil (†) and the acronym for plate (pl.) were used when relevant. The following initials were necessary to distinguish different authors with an identical family name: F.M. Brown, K.W. Brown, W. Kirby, W.F. Kirby, J.E. LeConte, J.L. LeConte, P.H. Lucas, R. Lucas, W.J. MacLeay, W.S. MacLeay, G.S. Medvedev, L.N. Medvedev, G.-A. Olivier, E. Olivier, F. Soldati, L. Soldati, C.G. Thomson, J. Thomson, C.O. Waterhouse, F.H. Waterhouse, G.R. Waterhouse. Note that the honorary title “Tjan-Schansky” was added by the Emperor of All Russia Nikolay II to the surname of A.P. Semenov’s father (and all his family members) in 1906 (M. Nabozhenko, pers. comm.). We use the author name “Semenov” for works published up to the end of 1906 and “Semenov-Tjan-Shansky” for those published after 1906. We also follow Bessudnova (2012: 102) and use the author’s surname “Fischer” for G. Fischer’s works up to the end of 1832 and use “Fischer von Waldheim” from 1833, the year he became a member of the Russian nobility.

Original literature for species-group names (i.e., type species) was verified; however, those works are not included in the “References” section because they are not the principal focus of this article, to conserve space and because they are, for the most part, available in recent publications. When the author(s) of a scientific name is different from the author(s) of the publication in which the name was proposed, the authorship of the scientific name is given in the format “Gray in Griffith and Pidgeon, 1832” (ICZN, Article 50.1.1 and Recommendation 51E) and only the reference pertaining to the whole work is given in the “References” section. Titles of references using a non-Latin alphabet were translated in English.

The name Xylotinus, as used by Sturm (1826: 59) to establish Xylotinusflabellicornis Sturm, 1826, the type species of Rhipidandrus J.L. LeConte, 1862, is considered an incorrect subsequent spelling for Xyletinus Latreille, 1810 [Coleoptera: Anobiidae] and is therefore not included here as an available genus-group name in Tenebrionidae (see Schwarz and Barber 1914: 175).

Two new synonymies were proposed in the fascicle on Coleoptera of the ‘Nomenclator Zoologicus’ by Agassiz (1846a). These synonymies are credited here to “Erichson in Agassiz” since Agassiz, who was not a beetle expert, acknowledged the contribution of the Coleoptera specialist W.F. Erichson, on the fascicle's title page, in the production of his catalogue.

Classification

The classification presented in Table 2 is a synthesis starting from works on Tenebrionidae family-group names (e.g., Bouchard et al. 2005, 2011; Bouchard and Bousquet 2020a) with additional data published in recent regional catalogues (e.g., Matthews and Bouchard 2008; Bousquet et al. 2018; Iwan et al. 2020), on fossils (Nabozhenko 2019), as well as on the diversity, taxonomy and phylogenetics of various clades (e.g., Bremer and Lillig 2014; Aalbu et al. 2017, 2018; Kamiński et al. 2019a, b, c, 2021b; Matthews and Lawrence 2019; Lumen et al. 2020; Lillig and Bremer 2021; Gearner et al. 2021). Only nomenclaturally available names are included in Table 2 wherein the valid genus-group names are listed in alphabetical order under each valid family-group name and the synonyms (preceded by “=”) are given in chronological order. Valid subgenera are given following the acronym “SG”.

Table 2.

List of nomenclaturally available genus-group names in the family Tenebrionidae presented in a classification framework. The presence of each valid extant genus and subgenus within a major biogeographical realm is indicated with the symbol “X” (usage of italics for the same symbol “X” indicates that a genus or subgenus is represented only by adventive species in a particular realm). * = status undetermined.

Genus-group names		Nearctic	Neotropic	Palaearctic	Afrotropic	Indo-Malay	Australasia	Oceania
Pimeliinae Latreille, 1802
Adelostomini Solier, 1834
Acanthioides Fairmaire, 1894					X
Acestophanus Koch, 1950					X
= Acestus Haag-Rutenberg, 1875
Adelostoma Duponchel, 1827				X	X
SGAdelostoma Duponchel, 1827				X	X
= Polyscopus Waltl, 1835
SGOmandelostoma Purchart, 2017					X
SGPsaryphulum Koch, 1952					X
SGZarudnionymus Semenov & Bogatchev, 1947				X	X
= Falsaspila Koch, 1952
= Adelostomoides Carl, 1991
Argasidus Péringuey, 1899					X
Arthrochora Gebien, 1938					X
Aspilomorpha Koch, 1952					X
Basilewskyum Koch, 1952					X
Brachymoschium Fairmaire, 1896					X
Carinosella Purchart, 2010					X
Cimicia Fairmaire, 1891					X
Cimicichora Koch, 1952					X
Cimiciopsis Koch, 1952					X
Eurychora Thunberg, 1789					X
Eurychorula Koch, 1952					X
Eutichus Haag-Rutenberg, 1875					X
Geophanus Haag-Rutenberg, 1875					X
Herpsis Haag-Rutenberg, 1875					X
Lepidochora Koch, 1952					X
Lycanthropa J. Thomson, 1860					X
= Zygas Pascoe, 1866
Machlopsis Pomel, 1871				X	X
SGHidrosella Koch, 1952				X	X
SGMachlopsis Pomel, 1871				X	X
= Hidrosis Haag-Rutenberg, 1875
Phytolostoma Koch, 1952					X
Platyphanus Koch, 1952					X
Platysemodes Strand, 1935					X
= Platysemus Haag-Rutenberg, 1875
Pogonobasis Solier, 1837				X	X
= Peristeptus Haag-Rutenberg, 1875
Pogonocanta Koch, 1952					X
Prunaspila Koch, 1950					X
= Aspila Fåhraeus, 1870
Psaryphis Erichson, 1843					X
Serrichora Koch, 1952					X
Smiliophanus Koch, 1950					X
= Smiliotus Haag-Rutenberg, 1875
Steptochora Koch, 1952					X
Stips Koch, 1950					X
= Steira Westwood, 1837
= Stira Agassiz, 1846
Stipsostoma Koch, 1952					X
Symphochora Koch, 1952					X
Adesmiini Lacordaire, 1859
Adesmia Fischer von Waldheim, 1822				X	X	X
SGAdesmia Fischer von Waldheim, 1822				X	X
= Sarachus Gistel, 1848
SGAdesmina Reitter, 1916				X	X	X
SGMacradesmia Kaszab, 1959				X	X
= Macradesmia Löbl & Merkl, 2020
SGMacropoda Solier, 1835					X
SGMacropodesmia Löbl & Merkl, 2020					X
SGOteroscelis Solier, 1835				X	X
= Heteroscelis Agassiz, 1846
SGOteroscelopsis Löbl & Merkl, 2020				X
SGPhysosterna Dejean, 1834					X
SGSomaladesmia Koch, 1944					X
SGSpongesmia Bouchard & Bousquet, subgen. nov.					X
SGZambesmia Bouchard & Bousquet, subgen. nov.					X
Alogenius Gebien, 1910					X
SGAequigula Penrith, 1979					X
SGAlogenius Gebien, 1910					X
= Pedionomus Haag-Rutenberg, 1875
Epiphysa Dejean, 1834					X
Eustolopus Gebien, 1938					X
= Entinopoda Gebien, 1938
Metriopus Solier, 1835					X
SGCeradesmia Gebien, 1920					X
SGCoeladesmia Reitter, 1916					X
= Peltadesmia Kuntzen, 1916
SGMetriopus Solier, 1835					X
= Iphthimera Reitter, 1916
Onymacris Allard, 1885					X
Orientocara Koch, 1952					X
Physadesmia Penrith, 1979					X
Renatiella Koch, 1944					X
= Spongesmima Koch, 1944
Stenocara Solier, 1835					X
SGArenacara Penrith, 1979					X
SGCauricara Penrith, 1979					X
SGStenocara Solier, 1835					X
Stenodesia Reitter, 1916					X
= Cephaladesmia Gebien, 1920
= Karroocara Koch, 1952
Akidini Billberg, 1820
Akis Herbst, 1799				X
= Acidia Illiger, 1804
= Stenopsis Rafinesque, 1815
= Acis Billberg, 1820
Cyphogenia Solier, 1837				X
SGCyphogenia Solier, 1837				X
= Eocyphogenia G.S. Medvedev, 1968
SGLechriomus Morawitz, 1865				X
Morica Dejean, 1834				X
Sarothropus Kraatz, 1865				X
Solskyia Solsky, 1881				X
Anepsiini LeConte, 1862
Anepsius LeConte, 1851		X
Batuliodes Casey, 1907		X
Batuliomorpha Doyen, 1987		X
Batulius LeConte, 1851		X
Asidini Fleming, 1821
Afrasida Wilke, 1922					X
SGAfrasida Wilke, 1922					X
SGArchasida Wilke, 1922					X
Alphasida Escalera, 1905				X
SGAlphasida Escalera, 1905				X
SGBetasida Reitter, 1917				X
= Subalphasida Escalera, 1928
SGElongasida Escalera, 1906				X
= Cribrasida Reitter, 1917
= Pseudoelongasida Escalera, 1922
SGGlabrasida Escalera, 1910				X
= Aplanasida Reitter, 1917
= Aulonasida Reitter, 1917
= Durasida Reitter, 1917
= Gymnetasida Reitter, 1917
= Melambasida Reitter, 1917
= Mimelasida Reitter, 1917
= Pedarasida Reitter, 1917
SGGranasida Reitter, 1917				X
SGMachlasida Escalera, 1907				X
= Protomachlasida Escalera, 1928
Amachla Koch, 1962					X
Andremiopsis Chatanay, 1913					X
Andremius Fairmaire, 1903					X
Ardamimicus Smith, 2013		X
Asida Latreille, 1802				X
SGAsida Latreille, 1802				X
= Dolichasida Reitter, 1917
= Euryasida Reitter, 1917
= Leptasida Reitter, 1917
= Insulasida Escalera, 1922
= Rugasida Escalera, 1922
SGGlobasida Escalera, 1905				X
SGGracilasida Escalera, 1905				X
= Planasida Escalera, 1907
= Trachasida Reitter, 1917
= Granulasida Escalera, 1922
= Pseudoplanasida Escalera, 1921
SGPeltasida Reitter, 1917				X
SGPolasida Reitter, 1917				X
= Opatrasida Escalera, 1922
Asidesthes Fairmaire, 1900					X
Asidomorpha Koch, 1962					X
Bartolozzia Ferrer, 1998					X
Cardigenius Solier, 1836			X
SGCardigenius Solier, 1836			X
= Cardiogenius Agassiz, 1846
SGEllidoneus Wilke, 1922			X
Craniotus LeConte, 1851		X
Cryptasida Koch, 1962					X
Euryprosternum Chatanay, 1914					X
Ferveoventer Smith, 2013		X
Heterasida Casey, 1912		X
Kochotella Bouchard & Bousquet, nom. nov.					X
= Millotella Koch, 1962
Leptasida Chatanay, 1914					X
Litasida Casey, 1912		X
Machla Herbst, 1799					X
= Machloplasta Wilke, 1922
= Pseudomachla Wilke, 1922
Machleida Fåhraeus, 1870					X
= Machloida Rye, 1873
Machlomorpha Péringuey, 1899					X
SGAsidomachla Wilke, 1922					X
SGMachlomorpha Péringuey, 1899					X
Machlophila Wilke, 1924					X
Micrasida Smith, 2013		X
Microschatia Solier, 1836		X
= Pycnonotida Casey, 1912
= Acroschatia Wilke, 1922
Oxyge Chatanay, 1914					X
Pelecyphorus Solier, 1836		X
SGAstrotus LeConte, 1858		X
SGPelecyphorus Solier, 1836		X
SGPleisiasida Smith, 2013		X
= Parasida Casey, 1912
SGPoliorcetes Champion, 1884		X
SGSicharbas Champion, 1884		X
SGStenosides Solier, 1836		X
= Pactostoma LeConte, 1858
= Ologlyptus Lacordaire, 1858
SGUcalegon Champion, 1884		X
SGZaleucus Champion, 1892		X
= Zamolxis Champion, 1884
Philolithus Lacordaire, 1858		X
SGGlyptasida Casey, 1912		X
SGGonasida Casey, 1912		X
SGHerthasida Wilke, 1922		X
SGPhilolithus Lacordaire, 1858		X
SGTisamenes Champion, 1884		X
Prosodidius Fairmaire, 1903					X
Pseudasida Fairmaire, 1895					X
Saeculum Kamiński, Kanda & Smith, 2021					X
Scotinesthes Fairmaire, 1895					X
= Parecatus Fairmaire, 1900
Scotinus W. Kirby, 1819			X
Stenomorpha Solier, 1836		X
SGAsidina Casey, 1912		X
SGAsidopsis Casey, 1912		X
SGBothrasida Casey, 1912		X
SGMegasida Casey, 1912		X
SGNotiasida Casey, 1912		X
SGPlatasida Casey, 1912		X
SGPycnomorpha Motschulsky, 1870		X
SGStenomorpha Solier, 1836		X
= Euschides LeConte, 1851
= Psilomera Motschulsky, 1870
SGStethasida Casey, 1912		X
SGTrichiasida Casey, 1912		X
Tamatasida Koch, 1962					X
Boromorphini Skopin, 1978
Boromorphus Wollaston, 1854				X
Branchini LeConte, 1862
Anectus Horn, 1867			X
Branchus LeConte, 1862		X	X
Oxinthas Champion, 1884			X
Caenocrypticini Koch, 1958
Caenocrypticoides Kaszab, 1969			X
Caenocrypticus Gebien, 1920					X
SGCaenocapicus Endrödy-Younga, 1996					X
SGCaenocrypticus Gebien, 1920					X
= Thorictophasis Koch, 1950
SGCryptocarpes Koch, 1952					X
= Lornamus Koch, 1952
SGPhyloradix Endrödy-Younga, 1996					X
SGPsammotopulus Endrödy-Younga, 1996					X
SGVernayella Koch, 1958					X
Ceratanisini Gebien, 1937
Ceratanisus Gemminger, 1870				X
= Anisocerus Faldermann, 1837
= Apolites Jacquelin du Val, 1861
= Haemerophygus Baudi di Selve, 1876
= Seidlitzellus Reitter, 1920
= Idastrandiella Strand, 1929
Tenebriocephalon Pic, 1925						X
= Klapperichia Kaszab, 1954
Cnemeplatiini Jacquelin du Val, 1861
Actizetina Watt, 1992
Actizeta Pascoe, 1875							X
Alaudina Aalbu, Caterino & Smith, 2018
Alaudes Horn, 1870		X
Cnemeplatiina Jacquelin du Val, 1861
Cnemeplatia Costa, 1847				X	X	X
= Autocera Wollaston, 1857
= Cnemoplatia Wollaston, 1865
Lepidocnemeplatia Bousquet & Bouchard, 2018		X	X			X
Philhammus Fairmaire, 1871				X	X
SGPhilhamellus Kaszab, 1962				X
SGPhilhammus Fairmaire, 1871				X	X
= Psilachnopus Reitter, 1901
= Canariella Uyttenboogaart, 1929
Rondoniellina Ferrer & Moragues, 2000
Durandius Kaszab, 1970						X
Rondoniella Kaszab, 1970						X
Thorictosomatina Watt, 1992
Thorictosoma Lea, 1919							X
Wattiana Matthews & Lawrence, 2005							X
Cnemodinini Gebien, 1910
Cnemodinus Cockerell, 1906		X
= Cnemodus Horn, 1870
Coniontini G.R. Waterhouse, 1858
Coelus Eschscholtz, 1829		X
= Coelomorpha Casey, 1890
= Pseudocoelus Casey, 1908
Coniontis Eschscholtz, 1829		X
= Coelotaxis Horn, 1876
= Coniontellus Casey, 1890
= Brachyontis Casey, 1908
= Coniontides Casey, 1908
= Crypticomorpha Casey, 1908
Conisattus Casey, 1895		X
Eusattus LeConte, 1851		X
= Conipinus LeConte, 1862
= Discodemus LeConte, 1862
= Eusattodes Casey, 1908
= Megasattus Casey, 1908
= Nesostes Casey, 1908
= Sphaeriontis Casey, 1908
= Coelosattus Blaisdell, 1927
Cossyphodini Wasmann, 1899
Cossyphodina Wasmann, 1899
Cossyphodes Westwood, 1851				X	X
= Allocossyphodes Andreae, 1961
= Hypercossyphodes Andreae, 1961
= Metacossyphodes Andreae, 1961
= Pachycossyphodes Andreae, 1961
Paramellops Andreae, 1961					X
Cossyphoditina Basilewsky, 1950
Cossyphodites Brauns, 1901					X
Esemephina Steiner, 1980
Esemephe Steiner, 1980			X
Paramellonina Andreae, 1961
Cossyphodinus Wasmann, 1899					X	X
Paramellon C.O. Waterhouse, 1882				X	X	X
Cossyphodini incertae sedis
Mimocossyphus Pic, 1923				X
Cryptochilini Solier, 1841
Calognathina Lacordaire, 1859
Calognathus Guérin-Méneville, 1836					X
= Callignathus Agassiz, 1846
Cryptochilina Solier, 1841
Cerasoma Endrödy-Younga, 1989					X
Cryptochile Latreille, 1828					X
= Phymatium Billberg, 1820
= Cryptotrophus Gistel, 1848
Cychrochile Koch, 1953					X
Epipagus Haag-Rutenberg, 1872					X
Horatoma Solier, 1841					X
= Horatomodes Haag-Rutenberg, 1872
= Saccophorus Haag-Rutenberg, 1872
= Saccophorella Strand, 1935
= Parapachynotela Koch, 1952
Horatomella Penrith & Endrödy-Younga, 1994					X
Orientochile Penrith & Endrödy-Younga, 1994					X
Pachynotelus Solier, 1841					X
= Fossilochile Koch, 1952
Homebiina Endrödy-Younga, 1989
Homebius Endrödy-Younga, 1989					X
Vansoniina Koch, 1955
Vansonium Koch, 1950					X
Cryptoglossini LeConte, 1862
Asbolus LeConte, 1851		X
Cryptoglossa Solier, 1837		X
= Centrioptera Mannerheim, 1843
= Oochila LeConte, 1862
= Amblycyphus Motschulsky, 1870
Schizillus Horn, 1874		X
Edrotini Lacordaire, 1859
Armalia Casey, 1907		X	X
Arthroconus Solier, 1851			X
= Gymnognathus Solier, 1851
Ascelosodis Redtenbacher, 1868				X		X
Auchmobius LeConte, 1851		X
Chilometopon Horn, 1874		X
= Prometopion Casey, 1907
Cryptadius LeConte, 1851		X
Ditaphronotus Casey, 1907			X
Edrotes LeConte, 1851		X
SGEdrotes LeConte, 1851		X
= Hedrotes Gemminger, 1870
SGOdrotes La Rivers, 1947		X
Emmenastrichus Horn, 1894		X
Emmenides Casey, 1907		X
Eremocantor Smith & Wirth, 2016		X
Eschatomoxys Blaisdell, 1935		X
Eurymetopon Eschscholtz, 1831		X
= Eurymetopum Agassiz, 1846
Falsoarthroconus Kaszab, 1978			X
Garridoa Marcuzzi, 1985			X
Hylithus Guérin-Méneville, 1834			X
= Sciaca Solier, 1835
Hylocrinus Casey, 1907		X	X
SGHylocrinus Casey, 1907		X	X
SGLocrodes Casey, 1907		X	X
SGParavius Casey, 1907		X
Kocakia Kaszab, 1985			X
= Idiopsis Kaszab, 1981
Koneus Giraldo-Mendoza & Flores, 2019			X
Melanastus Casey, 1907		X
Mencheres Champion, 1884			X
Mesabates Champion, 1884		X
Mesabatodes Casey, 1907		X
Metoponium Casey, 1907		X
SGMetoponiopsis Casey, 1907		X
SGMetoponium Casey, 1907		X
Micrarmalia Casey, 1907			X
Micromes Casey, 1907		X
Orthostibia Blaisdell, 1923		X
Oxygonodera Casey, 1907		X
Pachacamacius Flores & Giraldo-Mendoza, 2019			X
Paraguania Marcuzzi, 1953			X
Pescennius Champion, 1884		X
Pimeliopsis Champion, 1892		X
Posides Champion, 1884		X
Prohylithus Kaszab, 1964			X
Sechuranus Flores & Giraldo-Mendoza, 2019			X
Soemias Champion, 1884		X
Steriphanides Casey, 1907		X
Steriphanus Casey, 1907		X
Stibia Horn, 1870		X
= Eutriorophus Casey, 1924
Stictodera Casey, 1907		X
Stomion G.R. Waterhouse, 1845			X
= Stomium Agassiz, 1846
Telabis Casey, 1890		X
Telaponium Blaisdell, 1923		X
Texaponium Thomas, 1984		X
Tlascalinus Casey, 1907		X
Trichiotes Casey, 1907		X
Trientoma Solier, 1835			X
Trimytantron Ardoin, 1977			X
= Bielawskia Marcuzzi, 1985
Trimytis LeConte, 1851		X
= Pimalius Casey, 1907
Triorophus LeConte, 1851		X
Triphalopsis Blaisdell, 1923		X
Triphalopsoides Doyen, 1990			X
Triphalus LeConte, 1866		X
Troglogeneion Aalbu, 1985		X
Vizcainyx Aalbu & Smith, 2020		X
Elenophorini Solier, 1837
Elenophorina Solier, 1837
Leptoderis Billberg, 1820				X
= Elenophorus Dejean, 1821
= Helenophorus Gemminger, 1870
Megelenophorina Ferrer, 2015
Megelenophorus Gebien, 1910			X
= Cacicus Dejean, 1834
Psammetichus Latreille, 1828			X
Epitragini Blanchard, 1845
Aspidolobus Redtenbacher, 1868			X
Bothrotes Casey, 1907		X	X
Conoecus Horn, 1885		X
Cyrtomius Casey, 1907			X
SGCyrtomius Casey, 1907			X
SGGrandicyrtomius Freude, 1967			X
Ecnomosternum Gebien, 1928			X
Epitragella Kulzer, 1958			X
Epitragodes Casey, 1890		X	X
Epitragopsis Casey, 1907			X
Epitragosoma Brown & Triplehorn, 2002		X
Epitragus Latreille, 1802			X
SGEpitragus Latreille, 1802			X
= Lygophilus Rafinesque, 1815
= Aethales Dejean, 1834
SGGobretus Freude, 1967			X
SGSimilepitragus Freude, 1967			X
Eunotiodes Casey, 1907			X
Geoborus Blanchard, 1842			X
= Deroplatus Solier, 1851
Hemasodes Casey, 1907			X
Hypselops Solier, 1851			X
Kaszabus Freude, 1967			X
Lobometopon Casey, 1907		X	X
Metopoloba Casey, 1907		X
Nyctopetus Guérin-Méneville, 1831			X
Omopheres Casey, 1907			X
SGMicroomopheres Freude, 1993			X
SGOmopheres Casey, 1907			X
Ortheolus Casey, 1907			X
Parepitragus Casey, 1907			X
Pechalius Casey, 1907		X	X
= Epitragoma Casey, 1907
Pectinepitragus Pic, 1927			X
Penaus Freude, 1968			X
Phegoneus Casey, 1907		X	X
SGPectphegoneus Freude, 1968			X
SGPhegoneus Casey, 1907		X	X
SGSchoeniphegoneus Freude, 1968			X
Phitophilus Guérin-Méneville, 1831			X
Polemiotus Casey, 1907		X	X
Pseudortheolus Freude, 1968			X
Pseudothinobatis Bouchard & Bousquet, gen. nov.			X
Schoenicus LeConte, 1866		X	X
Stictodere Gebien, 1928			X
= Stictoderia Gebien, 1937
Tapinocomus Gebien, 1928			X
Tydeolus Champion, 1884			X
Erodiini Billberg, 1820
Ammodoides Lesne, 1915					X
Ammozoides Kaszab, 1979				X
Ammozoum Semenov, 1891				X
Amnodeis Miller, 1858				X
Anodesis Solier, 1834					X
Apentanodes Reitter, 1914				X		X
SGApentanodes Reitter, 1914				X
SGRasphytus Kulzer, 1956				X		X
Arthrodeis Solier, 1834				X	X
SGApentanes Reitter, 1914				X
SGArthrodeis Solier, 1834				X	X
= Arthrodes Agassiz, 1846
SGArthrodinus Reitter, 1900				X
SGKocheria Antoine, 1946				X	X
Arthrodibius Lesne, 1915				X	X
SGArthrodibius Lesne, 1915					X
SGErodibius Löbl, Bouchard, Merkl & Bousquet, 2020				X	X
SGHelioarthrodibius Koch, 1960					X
Arthrodion Lesne, 1915					X
Arthrodosis Reitter, 1900				X
Arthrodygmus Reitter, 1914						X
Arthrohyalosis Kaszab, 1979				X
Arthrohyalus Koch, 1943				X
Bulbulus Lesne, 1915				X
Capricephalius Koch, 1943				X
Diaphanidus Reitter, 1900				X
SGDiaphanidus Reitter, 1900				X
= Globularthrodosis Kaszab, 1979
SGPseudodiaphanidus Bogatchev, 1950				X
Diodontes Solier, 1834					X
Erodinus Reitter, 1900*
Erodiontes Reitter, 1914				X
= Iranarthrodosis Kaszab, 1959
Erodius Fabricius, 1775				X
SGDimeriseis Solier, 1834				X
SGDirosis Miller, 1858				X
SGEodirosis Kwieton, 1980				X
SGErodius Fabricius, 1775				X
= Cephacerus Rafinesque, 1815
= Acantophorus Billberg, 1820
= Herodius Agassiz, 1846
SGZophoserodius Reitter, 1914				X
Farsarthrosis Kaszab, 1979				X
Foleya Peyerimhoff, 1916				X
Histeromimus Gahan, 1895				X
Histeromorphus Kraatz, 1865					X
Hyalarthrodosis Kaszab, 1979				X
Hyalerodius Kaszab, 1979				X
Iranerodius Kaszab, 1959				X
Leptonychoides Schawaller, 1990				X
Leptonychus Chevrolat, 1833				X
Piestognathoides Kaszab, 1981				X
Piestognathus P.H. Lucas, 1858				X
Somalammodes Koch, 1943					X
Spyrathus Kraatz, 1865				X		X
Evaniosomini Lacordaire, 1859
Achanius Erichson, 1847			X
SGAchanius Erichson, 1847			X
SGAmbigatus Fairmaire, 1892			X
Aryenis Bates, 1868			X
Chorasmius Bates, 1868			X
Evaniosomus Guérin-Méneville, 1834			X
= Nochelius Gistel, 1848
Evelina J. Thomson, 1860			X
Melaphorus Guérin-Méneville, 1834			X
= Stenholma Solier, 1835
= Raptor Gistel, 1848
= Melanophorus Lacordaire, 1859
Oppenheimeria Koch, 1952					X
Vaniosus Kulzer, 1956			X
Falsomycterini Gebien, 1910
Falsomycterus Pic, 1907			X
Pteroctenus Kirsch, 1866			X
Idisiini G.S. Medvedev, 1973
Idisia Pascoe, 1866				X
Klewariini Gebien, 1910
Klewaria Reitter, 1910				X
Lachnogyini Seidlitz, 1894
Lachnodactylina Reitter, 1904
Lachnodactylus Seidlitz, 1898				X
= Lachnopus Seidlitz, 1894
Lachnogyina Seidlitz, 1894
Lachnogya Ménétriés, 1849				X		X
Netuschiliina Ferrer & Yvinec, 2004
Netuschilia Reitter, 1904				X
Leptodini Lacordaire, 1859
Leptodes Dejean, 1834				X
SGLeptodes Dejean, 1834				X
= Leptodinopsis Kaszab, 1959
SGLeptodopsis Haag-Rutenberg, 1879				X
SGMesoleptodes G.S. Medvedev & Iljina, 2007				X
SGParaleptodes G.S. Medvedev, 1967				X
SGProleptodes G.S. Medvedev, 1967				X
Tapenopsis Solier, 1843				X
= Tapinopsis Agassiz, 1846
Nycteliini Solier, 1834
Auladera Solier, 1836			X
= Aulacodera Agassiz, 1846
Callyntra Solier, 1836			X
Entomobalia Flores & Triplehorn, 2002			X
Entomoderes Solier, 1836			X
Epipedonota Solier, 1836			X
Gyriosomus Guérin-Méneville, 1834			X
= Brachygenius Dejean, 1836
Mitragenius Solier, 1836			X
Nyctelia Berthold, 1827			X
= Nyctelius Guérin-Méneville, 1827
= Nyctelioma Casey, 1908
Patagonogenius Flores, 1999			X
Pilobalia Burmeister, 1875			X
Psectrascelis Solier, 1836			X
= Cerostena Solier, 1836
= Stenocera Agassiz, 1846
Scelidospecta Kulzer, 1954			X
Nyctoporini Lacordaire, 1859
Nyctoporis Eschscholtz, 1831		X
= Emeax Pascoe, 1866
= Enneacoides Fairmaire, 1881
Phrynocarenini Gebien, 1928
Phrynocarenum Gebien, 1928			X
= Pseudoscotobius Kulzer, 1955
Physogasterini Lacordaire, 1859
Entomochilus Gay & Solier, 1843			X
Philorea Erichson, 1834			X
= Polpocara Solier, 1843
Physogaster Lacordaire, 1830			X
Physogasterinus Kaszab, 1981			X
Pimelosomus Burmeister, 1875			X
Pimeliini Latreille, 1802
Afghanopachys Kwieton, 1978				X
Allotadzhikistania Bogatchev, 1960				X
Apatopsis Semenov, 1891				X
Argyradelpha G.S. Medvedev, 2005				X
Argyrophana Semenov, 1889				X
Astorthocnemis Lillig & Pavlíček, 2002				X
Balachowskya Peyerimhoff, 1928				X
Bogatshevia G.S. Medvedev & Iwan, 2006				X
= Achaemenes Bogatchev, 1949
Cyclocnera Leo, 2018				X
Diesia Fischer von Waldheim, 1820				X
= Diesiola Skopin, 1961
Dietomorpha Reymond, 1938				X
Earophanta Semenov, 1903				X
= Earophila Semenov, 1903
= Earophilina Strand, 1917
Euryostola Reitter, 1893				X
Euthriptera Reitter, 1893				X
Gedeon Reiche & Saulcy, 1857				X
Habrobates Semenov, 1903				X
Habrochiton Semenov, 1907				X
Homopsis Semenov, 1893				X
Idiesa Reitter, 1893				X
Iranolasiostola Pierre, 1968				X
Iranopachyscelis Pierre, 1968				X
Kawiria Schuster, 1935				X
Lasiostola Dejean, 1834				X
SGCentrocnemita Strand, 1935				X
= Centrocnemis Kraatz, 1882
SGLasiocnema G.S. Medvedev, 1993				X
SGLasiograna G.S. Medvedev, 1993				X
SGLasiostola Dejean, 1834				X
= Pseudopimelia Gebler, 1859
Leucolaephus P.H. Lucas, 1859				X
= Mecopisthopus Karsch, 1881
Mantichorula Reitter, 1889				X
Meladiesia Reitter, 1909				X
Ocnera Fischer von Waldheim, 1822				X
= Brachycyphus Gebler, 1859
Pachylodera Quedenfeldt, 1890				X
Pachyscelina Kwieton, 1978				X
Pachyscelis Solier, 1836				X
SGPachyscelis Solier, 1836				X
= Brachyscelis Dejean, 1834
SGParapachyscelis Kwieton, 1978				X
Paraplatyope Löbl, Bouchard, Merkl & Bousquet, 2020				X
Pelorocnemis Solsky, 1876				X
Phymatiotris Solier, 1836				X
= Graecopachys Skopin, 1968
Pimelia Fabricius, 1775				X	X	X
SGAmblyptera Solier, 1836				X
SGAmblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 2018				X
SGAphanaspis Wollaston, 1864				X
SGCamphonota Solier, 1836				X
= Eurypimelia Reitter, 1915
SGChaetotoma Motschulsky, 1860				X
SGEcphoroma Solier, 1836				X
SGHispanomelia Mas-Peinado, Buckley, Ruiz & García-París, 2018				X
SGIberomelia Mas-Peinado, Buckley, Ruiz & García-París, 2018				X
SGItalomelia Mas-Peinado, Buckley, Ruiz & García-París, 2018				X
SGMagrebmelia Mas-Peinado, Buckley, Ruiz & García-París, 2018				X
SGMassadraamelia Mas-Peinado, Buckley, Ruiz & García-París, 2018				X
SGMelanostola Solier, 1836				X
= Balius Gistel, 1848
SGPimelia Fabricius, 1775				X	X	X
= Pimidia Rafinesque, 1815
= Agelarches Gistel, 1848
= Doderoella Schuster, 1926
SGPseudamblyptera Pierre, 1985				X
Pimeliocnera Reitter, 1909				X
Pimelipachys Skopin, 1962				X
Pisterotarsa Motschulsky, 1860				X
= Sympiezocnemis Solsky, 1876
= Piesterotarsa Sénac, 1884
Platyesia Skopin, 1971				X
Platyope Fischer von Waldheim, 1820				X
Podhomala Solier, 1836				X
SGPodhomala Solier, 1836				X
= Podomala Agassiz, 1846
= Uriela Reitter, 1887
= Pterocomodes Reitter, 1901
SGUrielina Reitter, 1888				X
Polpogenia Solier, 1836					X
Prionotheca Dejean, 1834				X	X
Przewalskia Semenov, 1893				X
Pseudopachyscelis Skopin, 1968				X
Pseudoplatyope Pierre, 1964				X
Pseudopodhomala Schuster, 1938				X
= Pseudopodhomalina Kaszab, 1960
= Gedrosia Bogatchev, 1961
Pseudostorthocnemis Gridelli, 1952				X
Pterocoma Dejean, 1834				X
SGDzhungaropterocoma Skopin, 1974				X
SGEupterocoma Skopin, 1974				X
SGHemipterocoma Skopin, 1974				X
SGMesopterocoma Skopin, 1974				X
SGMongolopterocoma Skopin, 1974				X
SGNeopterocoma Skopin, 1974				X
SGPachypterocoma Skopin, 1974				X
SGParapterocoma Skopin, 1974				X
SGPoopterocoma Skopin, 1974				X
SGPropterocoma Skopin, 1974				X
SGPseudopterocoma Skopin, 1974				X
SGPterocoma Dejean, 1834				X
SGSubpterocoma Bouchard & Bousquet, nom. nov.				X
= Pseudopimelia Motschulsky, 1860
Pterolasia Solier, 1836				X	X
Scelace Marseul, 1887				X
= Pachyscelodes Sénac, 1887
Spectrocnera Kwieton, 1981					X
Stalagmoptera Solsky, 1876				X
= Arnoldiola Semenov & Bogatchev, 1940
Sternocnera Skopin, 1964				X
Sternodes Fischer von Waldheim, 1837				X
Sternoplax Frivaldszky, 1890				X
SGMesosternoplax Skopin, 1973				X
SGPachysternoplax Skopin, 1973				X
SGParasternoplax Skopin, 1973				X
SGPseudosternoplax Skopin, 1973				X
SGSternoplax Frivaldszky, 1890				X
= Pseudeuthriptera Bogatchev & Kryzhanovskii, 1955
Sternotrigon Skopin, 1973				X
Storthocnemis Karsch, 1881				X	X
Tadzhikistania Bogatchev, 1960				X
Thriptera Solier, 1836				X	X
Trachyderma Latreille, 1828				X	X		X
SGAtrachyderma Skopin, 1962				X
SGTrachyderma Latreille, 1828				X	X		X
Trigonocnera Reitter, 1893				X
Trigonopachys Skopin, 1968				X
Trigonoscelis Dejean, 1834				X
SGChinotrigon Skopin, 1973				X
SGEchinotrigon Skopin, 1973				X
SGTrigonoscelis Dejean, 1834				X
Waterhousia Skopin, 1973				X
= Heinrichesia Carl, 2000
Praociini Eschscholtz, 1829
Antofagapraocis Flores, 2000			X
Asidelia Fairmaire, 1905			X
Calymmophorus Solier, 1841			X
= Calymmatophorus Gemminger, 1870
Eutelocera Solier, 1841			X
= Euteleocera Agassiz, 1846
Falsopraocis Kulzer, 1958			X
Gyrasida Koch, 1962			X
Neopraocis Kulzer, 1958			X
Parapraocis Flores & Giraldo, 2020			X
Patagonopraocis Flores & Chani-Posse, 2005			X
Pilobaloderes Kulzer, 1958			X
Platesthes G.R. Waterhouse, 1845			X
Platyholmus Dejean, 1834			X
= Platyolmus Burmeister, 1875
= Edrotopus Haag-Rutenberg, 1877
Praocidia Fairmaire, 1904			X
Praocis Eschscholtz, 1829			X
SGAnthrasomus Guérin-Méneville, 1834			X
= Anthracosomus Agassiz, 1846
SGFilotarsus Solier, 1841			X
SGHemipraocis Flores & Pizarro-Araya, 2014			X
SGMesopraocis Flores & Pizarro-Araya, 2014			X
SGOrthogonoderes Solier, 1841			X
= Aulacus Gray, 1832
= Eurygona Laporte, 1840
SGPostpraocis Flores & Pizarro-Araya, 2014			X
SGPraocida Flores & Pizarro-Araya, 2014			X
SGPraocis Eschscholtz, 1829			X
= Arctylus Dejean, 1834
SGPraonoda Flores & Pizarro-Araya, 2014			X
Thylacoderes Solier, 1843			X
Sepidiini Eschscholtz, 1829
Hypomelina Koch, 1955
Argenticrinis Louw, 1979					X
Bombocnodulus Koch, 1955					X
Brinckia Koch, 1962					X
Hypomelus Solier, 1843					X
Iugidorsum Louw, 1979					X
Sulcipectus Louw, 1979					X
Trachynotidus Péringuey, 1899					X
Triangulipenna Louw, 1979					X
Uniungulum Koch, 1962					X
Molurina Solier, 1834
Amiantus Fåhraeus, 1870					X
Arturium Koch, 1951					X
Brachyphrynus Fairmaire, 1882					X
Chiliarchum Koch, 1953					X
Dichtha Haag-Rutenberg, 1871					X
Distretus Haag-Rutenberg, 1871					X
SGDistretus Haag-Rutenberg, 1871					X
SGPerdistretus Koch, 1953					X
Euphrynus Fairmaire, 1897					X
Glyptophrynus Fairmaire, 1899					X
Huilamus Koch, 1953					X
Melanolophus Fairmaire, 1882					X
Moluris Latreille, 1802					X
= Physodera Solier, 1843
Ocnodes Fåhraeus, 1870					X
= Phanerotoma Solier, 1843
= Phanerotomea Koch, 1958
Phrynocolus Lacordaire, 1859					X
SGPhrynocolopsis Koch, 1951					X
SGPhrynocolus Lacordaire, 1859					X
= Cryptogenius Solier, 1843
SGSpinophrynus Koch, 1951					X
Phrynophanes Koch, 1951					X
Physophrynus Fairmaire, 1882					X
Psammodes W. Kirby, 1819					X
= Piesomera Solier, 1843
= Psammodophysis Péringuey, 1899
= Parmularia Koch, 1955
Psammophanes Lesne, 1922					X
SGPsammolophus Koch, 1953					X
SGPsammophanes Lesne, 1922					X
SGPsammophrynopsis Koch, 1953					X
SGPsammophrynus Koch, 1953					X
SGPsammostretus Koch, 1953					X
SGPsammotyriopsis Koch, 1953					X
SGSomalarabes Koch, 1953					X
Psammoryssus Kolbe, 1886					X
Psammotyria Koch, 1953					X
Stridulomus Koch, 1955					X
Tarsocnodes Gebien, 1920					X
Tibiocnodes ​Gearner & Kamiński, 2021					X
Toktokkus Kamiński & Gearner, 2020					X
Tuberocnodes ​Gearner & Kamiński, 2021					X
Oxurina Koch, 1955
Decoriplus Louw, 1979					X
Miripronotum Louw, 1979					X
Namibomodes Koch, 1952					X
Oxura W. Kirby, 1819					X
= Oxyura Agassiz, 1846
Palpomodes Koch, 1952					X
SGPalpomodes Koch, 1952					X
SGPygmaeodes Koch, 1952					X
Pterostichula Koch, 1952					X
SGPterostichula Koch, 1952					X
SGRipicolodes Koch, 1952					X
Stenethmus Gebien, 1937					X
Synhimba Koch, 1952					X
Sepidiina Eschscholtz, 1829
Dimoniacis Koch, 1958					X
Echinotus Solier, 1843					X
Peringueyia Koch, 1958					X
Sepidiopsis Gestro, 1892					X
Sepidiostenus Fairmaire, 1884					X
= Sepidiacis Fairmaire, 1884
Sepidium Fabricius, 1775				X	X
= Espidium Rafinesque, 1815
Vieta Laporte, 1840				X	X
= Dymonus Solier, 1843
= Divieta Reitter, 1914
Vietomorpha Fairmaire, 1887					X
Trachynotina Koch, 1955
Cyrtoderes Dejean, 1834					X
= Phligra Laporte, 1840
Epairopsis Koch, 1955					X
Ethmus Haag-Rutenberg, 1873					X
SGEthmomerus Koch, 1954					X
SGEthmophobes Koch, 1954					X
SGEthmus Haag-Rutenberg, 1873					X
= Tynthlobia Fairmaire, 1888
Histrionotus Koch, 1955					X
Microphligra Koch, 1955					X
Ossiporis Pascoe, 1866					X
= Epairops Fåhraeus, 1870
Oxycerus Koch, 1955					X
Somaticus Hope, 1841					X
SGAcromaticus Koch, 1955					X
SGBechuanitis Koch, 1955					X
SGCeromelaephus Koch, 1955					X
SGClinocranion Solier, 1843					X
SGDiacis Koch, 1955					X
SGSomaticus Hope, 1841					X
= Gonopterus Solier, 1843
SGTracheloeum Hope, 1841					X
SGTrachyderes Koch, 1955					X
SGTrichotrachys Koch, 1955					X
SGTrichotrichus Koch, 1955					X
SGTropitrachys Koch, 1955					X
Trachynotus Latreille, 1828					X
= Hipomelus Dejean, 1834
Trichethmus Gebien, 1937					X
Stenosini Schaum, 1859 (1834)
Araeoschizina Casey, 1907
Araeoschizus LeConte, 1851		X
Dichillina Reitter, 1916
Afghanillus Kaszab, 1960				X
Anchomma LeConte, 1858		X
Aspidocephalus Motschulsky, 1839				X
Dichillus Jacquelin du Val, 1860				X
SGDichillesthes Reitter, 1916				X
SGDichillinus Reitter, 1916				X
SGDichillocerus Reitter, 1916				X
SGDichillodontus Reitter, 1916				X
SGDichillomessor Reitter, 1916				X		X
SGDichillus Jacquelin du Val, 1860				X
SGMyrmecodichillus Kaszab, 1960				X
SGPushtunillus G.S. Medvedev, 1995				X
Discopleurus Lacordaire, 1859			X
= Pleurophorus Solier, 1851
Herbertfranzia Kaszab, 1973						X
Herbertfranziella Kaszab, 1973				X		X
Hexagonochilus Solier, 1851			X
Indochillus Koch, 1941						X
Microtelus Solier, 1838				X
Nepalofranziella Fouquè, 2013						X
Oogaster Faldermann, 1837				X
Pseudethas Fairmaire, 1896				X		X
SGPseudethas Fairmaire, 1896				X		X
= Schizillus Wasmann, 1899
= Dischizillus Wasmann, 1902
SGStenillus Blair, 1927				X
Pseudochillus Fouquè, 2015						X
SGKaszabochillus Fouquè, 2015						X
SGMicropseudochillus Fouquè, 2015						X
SGPseudochillus Fouquè, 2015						X
Reitterella Semenov, 1891				X
Harvengiina Ferrer, 2004
Harvengia Ferrer, 2004						X
Platamodina Reitter, 1900
Fitzsimonsium Koch, 1962					X
= Fitzsimonsia Koch, 1955
Microblemma Semenov, 1889				X
Platamodes Ménétriés, 1849				X
Stenosina Schaum, 1859 (1834)
Anethas Jakobson, 1924					X
SGAfrethas Koch, 1962					X
SGAnethas Jakobson, 1924					X
= Pseudethas Fairmaire, 1898
SGTetrethas Koch, 1962					X
Caribanosis Nabozhenko, Kirejtshuk, Merkl, Varela, Aalbu & Smith, 2016			X
Ecnomoderes Gebien, 1928			X
Ethas Pascoe, 1862						X
Eutagenia Reitter, 1886				X
Gebieniella Koch, 1940						X
Grammicus G.R. Waterhouse, 1845			X
Indostola G.S. Medvedev, 1991						X
Itampolis Koch, 1962					X
Microtelopsis Koch, 1940				X		X
SGExtetranosis Koch, 1940						X
SGHypermicrotelopsis Koch, 1940				X
SGMicrotelopsis Koch, 1940						X
SGTetranosis G.S. Medvedev, 1995				X		X
Mitotagenia Reitter, 1916				X	X
Perdicus Fairmaire, 1899					X
Renefouqueosis Aalbu, Smith, Kanda & Bouchard, 2017			X
Schizaraeus Kulzer, 1955			X
Schusteriella Koch, 1940					X
Stenosethas Kaszab, 1975						X
Stenosis Herbst, 1799				X	X	X
SGAfronosis G.S. Medvedev, 1995					X
SGBurmanosis G.S. Medvedev, 1995						X
SGIndianosis Koch, 1941						X
SGStenosidops Koch, 1940					X
SGStenosis Herbst, 1799				X	X	X
= Tagenia Latreille, 1802
Tagenostola Reitter, 1916				X		X
Tetranillus Wasmann, 1899				X		X
Timosmithus Ardoin, 1974					X
Typhlusechina Casey, 1907
Typhlusechus Linell, 1897		X
Stenosini incertae sedis
†Miostenosis Wickham, 1913
Tentyriini Eschscholtz, 1831
Abigopsis Escalera, 1914				X
Afrinus Fairmaire, 1888					X
SGAfrinus Fairmaire, 1888					X
= Nerina Lacordaire, 1859
SGGynandrocera Gebien, 1920					X
SGPalpafrina Koch, 1950					X
Alcinoeta Strand, 1929				X
= Alcinoe Ménétriés, 1849
= Allodengitha Bogatchev, 1963
Amblycarenum Gebien, 1910				X
= Amblycara Fairmaire, 1893
Ammogiton Peyerimhoff, 1920				X
Anatolica Eschscholtz, 1831				X
SGAnatolica Eschscholtz, 1831				X
SGEurepipleura Bogdanov-Katjkov, 1915				X
Aphrotus Péringuey, 1904					X
= Xenus Péringuey, 1899
Archinamibia Koch, 1952					X
Asphaltesthes Kraatz, 1865					X
SGAsphaltesthes Kraatz, 1865					X
SGTagenesthes Koch, 1941					X
Broomium Koch, 1950					X
Calyptopsis Solier, 1835				X
= Choristopsis Kraatz, 1865
Cantopipleurus Koch, 1943					X
Capnisiceps Chatanay, 1914					X
Catomulus Reitter, 1897				X
Cimipsa Peyerimhoff, 1911				X
= Cirta Gemminger, 1870
Colposcelis Dejean, 1834				X
SGColposcelis Dejean, 1834				X
SGColposceloides Schuster, 1940				X
SGColposcythis Reitter, 1889				X
SGScelocolpis Reitter, 1900				X
SGTurcmenicola Bogatchev, 1952				X
Colposphena Semenov, 1889				X
Craniosphena Koch, 1962					X
Cychrachna Koch, 1950					X
Cyphostethe Marseul, 1867				X	X
SGApterocyphostethe Kaszab, 1962				X
SGCyphostethe Marseul, 1867				X	X
= Asphena Semenov, 1889
SGCyphostethoides Löbl & Merkl, 2020				X
SGDerostethe Koch, 1950					X
SGHimastethe Koch, 1950					X
SGTrichostethe Koch, 1950					X
Dailognatha Steven, 1828				X
= Delognatha Agassiz, 1846
Dengitha Reitter, 1887				X
Derosphaerius Westwood, 1881					X
SGApterosphaeria Koch, 1950					X
SGDerosphaerius Westwood, 1881					X
= Derostrophus Fairmaire, 1888
= Oatesius Westwood, 1889
SGEpityria Koch, 1950					X
Dichomma Solier, 1835				X
Dividiopsa Koch, 1944					X
Epitrichia Gebler, 1859				X
Eschatostena Keleinikova, 1977				X
Eulipus Wollaston, 1864				X
= Mogadoria Escalera, 1905
Eusyntelia C.O. Waterhouse, 1881					X
Falsocatomulus Pic, 1914				X
= Microhionthis Blair, 1923
Freudeia Kaszab, 1961						X
Freyitia Koch, 1943						X
Girardius L. Soldati, 2009				X
Gnathosia Fischer von Waldheim, 1821				X
= Capnisa Dejean, 1836
Gnophota Erichson, 1843					X
Hegeter Latreille, 1802				X	X
SGHegeter Latreille, 1802				X	X
SGHomalapipleurus Español, 1957				X
SGPseudotalpophila Reitter, 1900				X
Hegeterocara Reitter, 1900				X
= Fourtaus Pic, 1921
Herlesa Reitter, 1897				X
Hionthis Miller, 1861				X
Homala Eschscholtz, 1831					X
= Omala Agassiz, 1846
Homalinota Koch, 1950					X
= Homalopsis Lesne, 1922
Homoeonota Fairmaire, 1882				X	X
= Isonota Fairmaire, 1887
Hyonthosoma Reitter, 1900				X
= Hionthisoma Gebien, 1937
Hyperops Eschscholtz, 1831				X	X	X
SGBelutschistanops Löbl, Bouchard, Merkl & Bousquet, 2020				X	X	X
SGDebeauxiella Bouchard & Bousquet, subgen. nov.						X
SGHyperops Eschscholtz, 1831				X	X	X
= Pachycera Eschscholtz, 1831
= Tetromma Dejean, 1834
= Oedenocera Reiche, 1862
SGHyperopsis Bouchard & Bousquet, subgen. nov.						X
SGPachycerops Koch, 1943						X
Hypsosoma Ménétriés, 1854				X
Imatismus Dejean, 1834				X	X	X
SGHimatismus Erichson, 1843					X
SGImatismus Dejean, 1834				X	X	X
= Curimosphena Gebien, 1920
Kokeniella Reitter, 1906						X
Leptosphena Semenov, 1891				X
Megagenius Solier, 1835				X
Melanochrus Wollaston, 1864				X
= Melasmocara Reitter, 1900
Melaxumia Reitter, 1895				X
Mesostena Eschscholtz, 1831				X	X
SGMesostena Eschscholtz, 1831				X	X
= Comphosida Macquart, 1850
SGMesostenopa Kraatz, 1865				X	X
SGPlatystena Koch, 1940					X
SGSaxistena Löbl & Merkl, 2020				X	X
Micipsa P.H. Lucas, 1855				X	X
SGCirsa P.H. Lucas, 1857				X
= Cyrta Lacordaire, 1859
SGMicipsa P.H. Lucas, 1855				X	X
Microdera Eschscholtz, 1831				X
SGAmicrodera Kaszab, 1966				X
SGDordanea Reitter, 1887				X
= Adordanea Reitter, 1897
SGFalsomicrodera Kaszab, 1966				X
SGIliodera Skopin, 1961				X
SGMicrodera Eschscholtz, 1831				X
= Rhostax Fischer von Waldheim, 1844
SGTentyrodera Koch, 1943				X
Microderopsis Haag-Rutenberg, 1876					X
Namaquaeon Koch, 1950					X
Namibismus Koch, 1952					X
Neognathosia Kaszab, 1959				X
Nerinodon Koch, 1952					X
Nothrocerus Fairmaire, 1887					X
Orostegastopsis Koch, 1962					X
Oterophloeus Desbrochers des Loges, 1881				X
= Tynteria Reitter, 1897
Oxycara Solier, 1835				X	X	X
SGOxycara Solier, 1835				X	X	X
= Emmenastus Motschulsky, 1845
= Melancrus Reiche & Saulcy, 1857
= Crypticoides Fairmaire, 1898
SGPleuroxycara Koch, 1959					X
SGSymphoxycara Koch, 1943				X	X
Oxycarops Reitter, 1900				X
Pachychila Eschscholtz, 1831				X
SGAnebacis Peyerimhoff, 1927				X
SGHegeteromorpha Escalera, 1913				X
SGNeocisba Reitter, 1900				X
= Neacisba Peyerimhoff, 1927
SGPachychila Eschscholtz, 1831				X
= Acisba Dejean, 1834
= Lophoma Solier, 1835
= Pachychile Lacordaire, 1859
SGPachychilina Reitter, 1900				X
SGTentyromorpha Escalera, 1913				X
= Tentyriomorpha Peyerimhoff, 1927
Paivaea Wollaston, 1864				X
= Paivea Scudder, 1882
Parabigopsis Español, 1946				X
Paracirta Schuster, 1930				X
Paulianesthes Koch, 1962					X
Phaeotribon Kraatz, 1865				X	X
Prochoma Solier, 1835				X		X
SGOxypistoma Löbl, Bouchard, Merkl & Bousquet, 2020				X		X
SGProchoma Solier, 1835				X
Psammocryptus Kraatz, 1865				X
Psammoica Solier, 1835				X
= Psammoeca Agassiz, 1846
Rhammatodes Haag-Rutenberg, 1876					X
SGAngoleantus Koch, 1952					X
SGRhammatodes Haag-Rutenberg, 1876					X
= Euleantus Haag-Rutenberg, 1876
= Tagenodes Haag-Rutenberg, 1876
= Asbolius Fairmaire, 1902
Rhomaleus Chatanay, 1915				X	X
Rhytinota Eschscholtz, 1831					X	X
SGNemapus Solier, 1835						X
= Melarachnica Kraatz, 1865
SGProrhytinota Bouchard & Bousquet, subgen. nov.					X
SGRhydimorpha Koch, 1943						X
SGRhytinota Eschscholtz, 1831					X
= Rhytidonota Agassiz, 1846
= Axumia Reiche, 1848
SGRhytistena Bouchard & Bousquet, subgen. nov.					X
SGSphenariopsis Kraatz, 1865						X
Rozonia Fairmaire, 1888					X
SGPseudorozonia Bouchard & Bousquet, subgen. nov.					X
SGRozonia Fairmaire, 1888					X
Scelosodis Solier, 1835				X
= Cratopus Eschscholtz, 1831
= Sceleoides Agassiz, 1846
= Abiga Guérin-Méneville, 1860
= Sceleodis Gemminger, 1870
Schweinfurthia Andres, 1922				X	X
Scythis Kraatz, 1865				X
= Semenovonymus Bogatchev, 1946
= Megascythis Keleinikova, 1963
Scytosoma Reitter, 1895				X
= Scythodonta Reitter, 1897
Sinoecia Chatanay, 1914				X
Sphenaria Ménétriés, 1849				X
Stegastopsis Kraatz, 1865				X
= Ohyonthis Reitter, 1898
Stenosida Solier, 1835						X	X
= Notioscythis Fairmaire, 1883
= Aprosphaena Reitter, 1916
Syachis Bates, 1879				X
= Orocina Reitter, 1897
Tamena Reitter, 1900				X
Tentyria Latreille, 1802				X
= Heliodromus Brullé, 1832
Tentyrina Reitter, 1900				X	X
= Tentyriina Peyerimhoff, 1907
Tentyronota Reitter, 1900				X
Thalpobia Fairmaire, 1871				X
= Micipsina Reitter, 1900
Thalpophilodes Strand, 1942					X
SGKaszabiella Koch, 1943					X
SGRhytinopsis Bouchard & Bousquet, subgen. nov.					X
SGThalpophilodes Strand, 1942					X
= Thalpophila Solier, 1835
Thraustocolus Kraatz, 1866				X	X
SGLeptoderops Löbl, Bouchard, Merkl & Bousquet, 2020				X	X
SGProthraustocola Kaszab, 1957				X
SGThraustocolus Kraatz, 1866				X
= Calobamon Kraatz, 1865
= Ibnsaudia Koch, 1941
Trichosphaena Reitter, 1916				X	X
Uyttenboogaartia Koch, 1943				X
Thinobatini Lacordaire, 1859
Cordibates Kulzer, 1956			X
Thinobatis Eschscholtz, 1831			X
Trilobocarini Lacordaire, 1859
Derosalax Gebien, 1926			X
Eremoecus Lacordaire, 1859			X
Peltolobus Lacordaire, 1859			X
= Megalophrys G.R. Waterhouse, 1845
= Alhuena Kulzer, 1956
Salax Guérin-Méneville, 1834			X				X
= Pilioloba Erichson, 1846
Trilobocara Solier, 1851			X
= Edrotinus Fairmaire, 1904
= Orthonychius Gebien, 1926
Vacronini Gebien, 1910
Alaephus Horn, 1870		X
= Vacronus Casey, 1907
Eupsophulus Cockerell, 1906		X
= Eupsophus Horn, 1870
Exangeltus Blackburn, 1897							X
Lixionica Blackburn, 1896							X
Zophosini Solier, 1834
Zophosis Latreille, 1802				X	X	X
SGAnacardiosis Endrödy-Younga, 1986					X
SGAnisosis Deyrolle, 1867					X
SGCalosis Deyrolle, 1867					X
SGCardiosis Deyrolle, 1867				X	X
SGCaroliphosis Penrith, 1981					X
SGCarpiella Koch, 1962					X
SGCerosis Gebien, 1920					X
SGCheirosis Deyrolle, 1867				X	X
= Chirosis Gemminger, 1870
SGDactylocalcar Gebien, 1938					X
SGDignathosis Koch, 1958					X
SGGahanosis Penrith, 1983					X
SGGyrosis Gebien, 1920					X
SGHeliophosis Koch, 1952					X
SGHesseosis Koch, 1958					X
SGHologenosis Deyrolle, 1867				X	X
SGLatipleurosis Penrith, 1977					X
SGMicrosis Koch, 1958					X
SGMyrmecophosis Koch, 1958					X
SGNamaphosis Penrith, 1981					X
SGOccidentophosis Penrith, 1977					X
SGOculosis Penrith, 1977				X	X
SGOnychosis Deyrolle, 1867					X
SGOphthalmosis Deyrolle, 1867				X	X
SGPlanirostrosis Penrith, 1977					X
SGPropemicrosis Penrith, 1981					X
SGProtocalosis Penrith, 1977					X
SGProtodactylus Koch, 1952					X
= Zophosodactylus Koch, 1962
SGSabulophosis Penrith, 1977					X
SGScopulophosis Penrith, 1977					X
SGSeptentriophosis Penrith, 1982				X	X	X
SGSulcosis Penrith, 1977					X
SGTarsosis Gebien, 1920					X
SGZophosis Latreille, 1802					X
= Calcarosis Penrith, 1977
= Predactylosis Penrith, 1977
Zolodininae Watt, 1975
†Praezolodinus Bao, 2020
Tanylypa Pascoe, 1869							X
Zolodinus Blanchard, 1853							X
Lagriinae Latreille, 1825 (1820)
Adeliini W. Kirby, 1828
Adelium W. Kirby, 1819							X
= Dystalica Pascoe, 1869
= Rues Casey, 1891
= Tropidopterus Cazurro Ruiz, 1897
Adelodemus Haag-Rutenberg, 1878							X
= Apostethus Pascoe, 1882
Adelozotypus Kaszab, 1982							X
Aoupinia Matthews, 2003							X
Apasis Pascoe, 1869							X
Apocryphodes Matthews, 1998							X
Arcothymus Pascoe, 1866							X
Bellendenum Matthews, 1998							X
Blepegenes Pascoe, 1868							X
= Ceradelium Preudhomme de Borre, 1868
Bluops Carter, 1914							X
Bolusculus Matthews, 1998							X
Brycopia Pascoe, 1869							X
= Dinoria Pascoe, 1869
Cardiothorax Motschulsky, 1860							X
= Thoracophorus Hope, 1841
= Atryphodes Pascoe, 1866
= Otrintus Pascoe, 1866
Coripera Pascoe, 1866							X
Cymbeba Pascoe, 1866							X
Daedrosis Bates, 1868							X
= Macroperas Carter, 1914
Diaspirus Matthews, 1998							X
Dicyrtodes Matthews, 1998							X
Diemenoma Matthews, 1998							X
Dorrigonum Matthews, 1998							X
Epomidus Matthews, 1998							X
Exadelium Watt, 1992							X
Gondvanadelium Kaszab, 1981			X
Isopteron Hope, 1841							X
= Cestrinus Erichson, 1842
= Isopterum Agassiz, 1846
= Apatelus Mulsant & Rey, 1859
= Prionotus Mulsant & Rey, 1859
= Achora Pascoe, 1869
= Priothorax Gebien, 1910
Kaszabadelium Watt, 1992							X
Leptogastrus W.J. MacLeay, 1872							X
Licinoma Pascoe, 1869			X				X
Mesopatrum Broun, 1893							X
Montaguea Kaszab, 1982							X
Monteithium Matthews, 1998							X
Neoadelium Carter, 1908							X
= Pseudadelium Kaszab, 1982
Nolicima Matthews, 1998							X
Nototrintus Carter, 1924							X
Ozotypoides Kaszab, 1982							X
Penadelium Matthews, 1998			X
Periatrum Sharp, 1886							X
Pheloneis Pascoe, 1866							X
= Amarosoma Redtenbacher, 1868
Pseudobyrsax Kaszab, 1982							X
Pseudocilibe Kaszab, 1982							X
Pseudopatrum Sharp, 1886							X
= Mitua Hope, 1848
Seirotrana Pascoe, 1866							X
Stenadelium Watt, 1992							X
Valdivium Matthews, 1998			X
Wattadelium Emberson, 2000							X
= Edalus Broun, 1893
Yarranum Matthews, 1998							X
Zeadelium Watt, 1992							X
Belopini Reitter, 1917
Adelonia Laporte, 1840		X	X
SGAdelonia Laporte, 1840		X	X
= Merotemnus Horn, 1870
= Rhacius Champion, 1885
= Ostorius Fairmaire, 1889
SGLatorhascius Pic, 1925			X
Centorus Mulsant, 1854				X	X
SGBelopomerus Reitter, 1920				X
SGBelopus Gebien, 1911				X	X
= Calcar Dejean, 1821
= Hemeralopius Gistel, 1848
SGCentorus Mulsant, 1854				X
SGNanocalcar Skopin, 1974				X
Doyenia Matthews & Lawrence, 2005							X
Euclarkia Lea, 1919							X
Eulea Carter, 1937							X
Exeniotis Pascoe, 1871			X
Kershawia Lea, 1905							X
Rhypasma Pascoe, 1862			X
= Derosimus Fairmaire, 1904
Thoseus Pic, 1925						X
†Yantaroxenos Nabozhenko, Kirejtshuk & Merkl, 2016
Chaerodini Doyen, Matthews & Lawrence, 1990
Chaerodes White, 1846							X
= Choerodes Gemminger, 1870
Sphargeris Pascoe, 1860							X
Cossyphini Latreille, 1802
Cossyphus G.-A. Olivier, 1791				X	X	X	X
SGAcontodactylus Desbrochers des Loges, 1894				X	X	X	X
= Paracossyphus Viñolas & Cartagena, 2005
SGCossyphus G.-A. Olivier, 1791				X	X	X
Endustomus Brême, 1842					X
= Endostomus Gemminger, 1870
= Endostostomus Jakobson, 1914
Eschatoporiini Blaisdell, 1906
Eschatoporis Blaisdell, 1906		X
Goniaderini Lacordaire, 1859
Acropachia Mäklin, 1875			X
Anaedus Blanchard, 1842		X	X	X	X	X
= Aspisoma Duponchel & Chevrolat, 1841
= Anaedes Agassiz, 1846
= Aspidosoma Agassiz, 1846
Gamaxus Bates, 1868			X
Goniadera Perty, 1832			X
SGAemymone Bates, 1868			X
SGGoniadera Perty, 1832			X
= Goniodera Agassiz, 1846
SGOpatresthes Gebien, 1928			X
Lyprochelyda Fairmaire, 1899					X
= Basanaedus Pic, 1917
Microanaedus Pic, 1923					X
Microgoniadera Pic, 1917			X
Microlyprops Kaszab, 1939						X
Myrmecopeltoides Kaszab, 1973			X
Paratenetus Spinola, 1845		X	X
= Lagriola Kirsch, 1874
= Storthephora Mäklin, 1875
Pengalenganus Pic, 1917						X	X
Phobelius Blanchard, 1842			X
Phymatestes Pascoe, 1866			X
= Phymatodes Dejean, 1834
Prateus LeConte, 1862		X	X
Pseudanaedus Gebien, 1921					X
Pseudolyprops Fairmaire, 1882							X
= Trichulodes Carter, 1914
Spinadaenus Pic, 1921						X
Tithassa Pascoe, 1860			X
Xanthicles Champion, 1886			X
†Gonialaenini Nabozhenko, Bukejs & Telnov, 2019
†Gonialaena Nabozhenko, Bukejs & Telnov, 2019
Laenini Seidlitz, 1895
Afrolaena Endrödy-Younga & Schawaller, 2002					X
Borneolaena Schawaller, 1998						X
Chaetyllus Pascoe, 1860			X
= Polytropus Kirsch, 1866
= Rhicnodus Fairmaire, 1892
Enigmatica Ferrer, 2005					X
Grabulax Kanda, 2016			X
Hovadelium Ardoin, 1962					X
Hypolaenopsis Masumoto, 2001				X
Laena Dejean, 1821				X		X
= Catolaena Reitter, 1900
= Psilolaena Heller, 1923
= Ebertius Jedlička, 1965
Lucidolaena Endrödy-Younga & Schawaller, 2002					X
Mimolaena Ardoin, 1962					X
Nepalolaena Schawaller, 2001						X
Prolaena Kaszab, 1980						X
Rhacolaena Kaszab, 1979						X
Lagriini Latreille, 1825 (1820)
Lagriina Latreille, 1825 (1820)
Acerogria Borchmann, 1936							X
= Birolagria Pic, 1956
Acritolagria Borchmann, 1916					X
Acutogria Merkl, 1988							X
Adosogria Borchmann, 1936					X
Adynata Fåhraeus, 1870				X	X
= Microlagria Seidlitz, 1898
Alagria Borchmann, 1916					X
= Lagriostira Kolbe, 1902
Allogria Borchmann, 1916					X
Anotoma Borchmann, 1936					X
Aulonogria Borchmann, 1929						X
Bequaertiella Pic, 1914					X
Bothrichara Borchmann, 1916							X
Bothrionota Borchmann, 1936						X
Bothynogria Borchmann, 1916				X		X
Calogria Borchmann, 1916							X
Ceratoma Borchmann, 1916*
Cerogria Borchmann, 1911				X	X	X	X
SGCerogria Borchmann, 1911				X	X	X	X
= Cerogriodes Borchmann, 1941
= Aeschrocera Chen & Chou, 1996
SGDrepanomela Borchmann, 1936							X
SGParogria Borchmann, 1936					X
Cerostira Borchmann, 1942					X
= Allocera Borchmann, 1916
Chilenolagria Pic, 1936			X
Chrysolagria Seidlitz, 1898				X	X
Costiferolagria Pic, 1915						X
= Auristira Borchmann, 1916
Ctenogria Borchmann, 1916						X
Derolagria Borchmann, 1916					X
Ecnocera Borchmann, 1936					X
Ecnolagria Borchmann, 1916							X
SGEcnolagria Borchmann, 1916							X
SGOnocera Borchmann, 1936							X
Emydodes Pascoe, 1860			X
Falsolagria Pic, 1927			X
Flabellolagria Pic, 1927					X
Gronophora Borchmann, 1916							X
Helogria Borchmann, 1916						X
Kaindilagria Merkl, 1988							X
Lagria Fabricius, 1775			X	X	X	X	X
SGAmmocera Borchmann, 1941				X		X
SGApteronympha Seidlitz, 1898				X	X
SGGrandelagria Pic, 1940					X
SGLagria Fabricius, 1775			X	X	X	X	X
= Lachna Billberg, 1820
SGLagriella Borchmann, 1916					X	X
Lagriopsis Borchmann, 1916							X
Lopholagria Borchmann, 1916					X
SGLopholagria Borchmann, 1916					X
SGMokalagria Pic, 1953					X
Lorona Borchmann, 1936						X
Mallogria Borchmann, 1936					X
Metriolagria Merkl, 1987							X
Mimolagria Pic, 1927			X
Minasius Pic, 1932			X
Neogria Borchmann, 1911						X
= Lagriomima Pic, 1950 syn. nov.
Nothogria Borchmann, 1916							X
Odontogria Borchmann, 1936						X
Oreogria Merkl, 1988							X
Oroptera Borchmann, 1916							X
Phaedogria Borchmann, 1936						X
Physogria Borchmann, 1916					X
Physolagria Fairmaire, 1891					X
Porrolagria Kolbe, 1883					X
= Lagrimina Fairmaire, 1894
Pseudolagria Champion, 1917			X
Rhaibogria Borchmann, 1936					X
Ruandania Pic, 1955					X
Schevodera Borchmann, 1936						X
Schevogria Borchmann, 1936					X
Sphinctoderus Fairmaire, 1903						X
Stenolagria Merkl, 1987							X
Sulcolagria Pic, 1955					X
Tomogria Merkl, 1988							X
Xenocerogria Merkl, 2007				X		X
= Xenocera Borchmann, 1936
Xenogena Borchmann, 1936					X
Xenolagria Merkl, 1987							X
Statirina Blanchard, 1845
Anisostira Borchmann, 1915				X		X
Arthromacra W. Kirby, 1837		X		X		X
= Macrarthra Agassiz, 1846
Arunogria Merkl, 1991						X
Astatira Borchmann, 1921			X
Barsenis Pascoe, 1887			X
SGBarsenis Pascoe, 1887			X
SGMicrodisema Pic, 1917			X
SGModicodisema Pic, 1917			X
SGNemostiromorpha Pic, 1917			X
= Adisema Borchmann, 1936
Borchmannia Pic, 1912						X
SGBorchmannia Pic, 1912						X
SGPseuduroplatopsis Pic, 1913						X
Bothriostira Borchmann, 1936					X
Casnonidea Fairmaire, 1882					X	X	X
SGCasnonidea Fairmaire, 1882					X	X	X
= Synatractus W.J. MacLeay, 1887
= Falsocasnonidea Pic, 1934
SGPilosocasnonidea Pic, 1934						X
Chlorophila Semenov, 1891				X		X
Colparthrum Kirsch, 1866			X
SGColparthrum Kirsch, 1866			X
SGPseudocolparthrum Borchmann, 1916			X
Costatosora Pic, 1934					X
Cylindrosora Borchmann, 1936						X
Cylindrostira Borchmann, 1936						X
Davaona Borchmann, 1930						X
Derostira Fairmaire, 1897					X
Diorhychina Borchmann, 1936					X
Disema Mäklin, 1875			X
Disemorpha Pic, 1917			X
Donaciolagria Pic, 1914				X		X
Dysodera Borchmann, 1936					X
Dysopinus Borchmann, 1936						X
Eccoptostira Borchmann, 1936					X
Entypodera Gerstaecker, 1871					X
= Loubacantus Bonadona, 1959
Epicydes Champion, 1889			X
SGCybostira Borchmann, 1936			X
SGEpicydes Champion, 1889			X
Exostira Borchmann, 1925						X
Falsonemostira Pic, 1917						X
Gebienia Borchmann, 1921			X
Hosohamudama Masumoto, 1988						X
Hypostatira Fairmaire, 1889			X
Hysterarthron J. Thomson, 1864						X
Impressosora Pic, 1952					X
SGImpressosora Pic, 1952					X
SGNeoeutrapela Bousquet & Bouchard, 2013					X
= Eutrapela Dejean, 1834
Isocera Borchmann, 1909			X
= Isotoma Blanchard, 1842
Lagriodema Borchmann, 1930						X
Lagriogonia Fairmaire, 1891				X
Lagriomima Pic, 1934						X	X
Lagriostira Fairmaire, 1883					X
Leptinostethus Borchmann, 1936					X
Leptosora Borchmann, 1936					X
Lobophilomorphus Pic, 1911					X
Lophophyllus Fairmaire, 1887					X
Macrocasnonidea Pic, 1934						X
Macrolagria Lewis, 1895				X
Malaiseum Borchmann, 1941						X
Meniscophorus Champion, 1889			X
Merklia Chen, 1997						X
Meropria Borchmann, 1921			X
Mimoborchmania Pic, 1934						X
Mimuroplatopsis Borchmann, 1936					X
Natalostira Pic, 1913					X
Nevermanniella Borchmann, 1936			X
Ocularisora Pic, 1934					X
Odontocerostira Merkl, 2007				X		X
= Odontocera Chen & Yuan, 1996
Othryades Champion, 1889			X
Pachystira Chen, 1997						X
Piciella Borchmann, 1936			X
Pseudeutrapela Pic, 1952					X
Pseudocasnonidea Borchmann, 1936						X
Pseudostira Fairmaire, 1903					X
Rhagostira Borchmann, 1936					X
Rhaibodera Borchmann, 1921			X
Rhosaces Champion, 1889			X
Robustosora Pic, 1954					X
Rouyerus Pic, 1911						X
SGBorneostira Pic, 1912						X
SGRouyerus Pic, 1911						X
Sipolisia Fairmaire, 1889			X
Sora Walker, 1859				X	X	X	X
SGHirsutosora Pic, 1934						X
SGNemostiropsis Borchmann, 1936						X
SGSora Walker, 1859				X	X	X	X
= Nemostira Fairmaire, 1869
Sphragidophorus Champion, 1889			X
Splichalia Reitter, 1913				X
Staius Fairmaire, 1896					X
SGDerostirostaius Borchmann, 1936					X
SGStaius Fairmaire, 1896					X
Statira Lepeletier & Audinet-Serville, 1828		X	X
SGFoveostatira Pic, 1918			X
SGPleurostira Borchmann, 1921			X
SGSpinostatira Pic, 1918			X
= Hoplostira Borchmann, 1921
SGStatira Lepeletier & Audinet-Serville, 1828		X	X
SGXenostira Borchmann, 1921			X
Statiropsis Borchmann, 1912			X
Strongylagria Pic, 1915				X
Taiwanolagria Masumoto, 1988				X		X
Thoracostira Borchmann, 1936			X
Uroplatopsis Champion, 1889			X
Xanthalia Fairmaire, 1894				X	X	X
= Xanthia Fairmaire, 1893
= Heterogria Fairmaire, 1896
= Lagriocera Fairmaire, 1896
= Wallardilagria Pic, 1910
= Pachylagria Borchmann, 1912
= Lagriodes Borchmann, 1930
= Diversogria Pic, 1954
Xenostethus Bates, 1868					X
= Coracostira Fairmaire, 1899
Lupropini Lesne, 1926
Antennoluprops Schawaller, 2007					X
Ardoiniellus Schawaller, 2013					X
Bolitrium Gebien, 1914						X
Capeluprops Schawaller, 2011					X
Coxelinus Fairmaire, 1869					X
Curtolyprops Pic, 1917					X
Dichastops Gerstaecker, 1871					X
Enicmosoma Gebien, 1922					X
= Enicmonota Ardoin, 1958
Falsotithassa Pic, 1934						X
= Derispiolina Kaszab, 1979
Indenicmosoma Ardoin, 1964				X		X
Iscanus Fauvel, 1904							X	X
= Araucaricola Lea, 1929
Kuschelus Kaszab, 1982							X
Lorelus Sharp, 1876			X				X	X
= Lorelopsis Champion, 1896
Luprops Hope, 1833				X	X	X	X
= Oligorus Dejean, 1834
= Syggona Fåhraeus, 1870
= Etazeta Fairmaire, 1889
= Syngona Rye, 1873
Mesotretis Bates, 1872							X
Microcalcar Pic, 1925					X
Micropedinus Lewis, 1894				X		X	X
= Notoprataeus Carter, 1924
Mimocellus Wasmann, 1904					X
Paralorelopsis Marcuzzi, 1994			X
Sphingocorse Gebien, 1921					X	X
Spinolagriella Pic, 1955					X
= Spinorhacus Kaszab, 1969
Spinolyprops Pic, 1917				X	X	X
Terametus Motschulsky, 1869					X
Pycnocerini Lacordaire, 1859
Aediatorix Bates, 1868						X
= Sipirocus Fairmaire, 1896
Amorphochirus Gebien, 1904					X
Calostegia Westwood, 1843					X
= Calostega Gemminger, 1870
= Apristopus Kolbe, 1903
Catamerus Fairmaire, 1887					X
Chirocharis Kolbe, 1903					X
Chiroscelis Lamarck, 1804					X
Gabonisca Fairmaire, 1894					X
= Gabonia Fairmaire, 1894
Hemipristula Bouchard & Bousquet, nom. nov.					X
= Hemipristis Kolbe, 1903
Malayoscelis Schawaller, 2003						X
Metallonotus Gray, 1832					X
= Aspidosternum Mäklin, 1867
Passalocharis Koch, 1954					X
Pezodontus Dejean, 1834					X
= Odontopus Silbermann, 1833
= Odontopezus Alluaud, 1889
Pheugonius Fairmaire, 1899						X
Prioproctus Kolbe, 1903					X
Prioscelides Kolbe, 1889					X
Prioscelis Hope, 1841					X
= Iphius Dejean, 1834
= Bovius Gistel, 1848
Pristophilus Kolbe, 1903					X
Pycnocerus Westwood, 1844					X
SGDinoscelis Gerstaecker, 1854					X
SGPycnocerus Westwood, 1844					X
= Pachylocerus Hope, 1841
Stratodemus Gebien, 1921					X
Lagriinae incertae sedis
Asiopus Sharp, 1892			X
Pseudesarcus Champion, 1913			X
Nilioninae Oken, 1843
Nilio Latreille, 1802			X
SGLinio Bouchard & Bousquet, subgen. nov.			X
SGMicronilio Pic, 1936			X
SGNilio Latreille, 1802			X
Phrenapatinae Solier, 1834
Archaeoglenini Watt, 1975
Archaeoglenes Broun, 1893			X	X	X	X	X	X
Sepilokus Iwan & Raś, 2020						X
Penetini Lacordaire, 1859
Afrotagalus Gebien, 1942					X
Archeophthora Kaszab, 1978			X				X
Clamoris Gozis, 1886		X	X	X		X
= Phtora Mulsant, 1854
= Phthora Champion, 1893
Cleolaus Champion, 1886			X
Daochus Champion, 1886			X
Dioedus LeConte, 1862		X	X	X	X	X	X	X
= Arrhabaeus Champion, 1886
= Brachycilibe Carter, 1911
= Tagalus Gebien, 1914
Endroeditagalus Schawaller & Bouchard, 2019					X
Exechophthalmus Ardoin, 1974					X
Falsotagalus Kaszab, 1977							X
Leleupium Kaszab, 1956					X
Madagassa Koch, 1950					X
= Pycna Fairmaire, 1894
Molion Champion, 1886			X
Nanotagalus Gebien, 1942					X
Neotagalus Kaszab, 1955								X
Peneta Lacordaire, 1859			X
Pseudophthora Kaszab, 1970						X	X	X
SGFalsophthora Kaszab, 1977								X
SGPseudophthora Kaszab, 1970						X	X
Pycnochilus C.O. Waterhouse, 1879					X
Scolytocaulus Fairmaire, 1896					X	X	X
= Platycilibe Carter, 1911
= Picnotagalus Kaszab, 1939
Tagalinus Kaszab, 1977							X
Tagalopsis Kaszab, 1955								X
Taiwanotagalus Masumoto, 1982						X
Telchis Champion, 1886			X
Zypoetes Champion, 1893			X			X	X
Phrenapatini Solier, 1834
Delognatha Lacordaire, 1859			X
Phrenapates Gray, 1831			X
Phrenapatinae incertae sedis
Aphtora Bates, 1872							X
Kuhitangiinae G.S. Medvedev, 1962
Foranotini Nabozhenko & Sadeghi, 2017
Foranotum Nabozhenko & Sadeghi, 2017				X
Kuhitangiini G.S. Medvedev, 1962
Kuhitangia G.S. Medvedev, 1962				X
Blaptinae Leach, 1815
Amphidorini LeConte, 1862
Eleodes Eschscholtz, 1829		X	X
SGAmphidora Eschscholtz, 1829		X
SGArdeleodes Blaisdell, 1937		X
SGBlapylis Horn, 1870		X
= Eleodopsis Blaisdell, 1939
SGCaverneleodes Triplehorn, 1975		X
SGChaseleodes Thomas, 2015			X
SGCratidus LeConte, 1862		X
SGDiscogenia LeConte, 1866		X
SGEleodes Eschscholtz, 1829		X	X
= Elaeodes Gemminger, 1870
SGHeteropromus Blaisdell, 1909		X
SGLitheleodes Blaisdell, 1909		X
SGMelaneleodes Blaisdell, 1909		X	X
SGMetablapylis Blaisdell, 1909		X
SGOmegeleodes Triplehorn & Thomas, 2012		X	X
SGPromus LeConte, 1862		X	X
SGPseudeleodes Blaisdell, 1909		X
= Trichoderulus Blaisdell, 1923
SGSteneleodes Blaisdell, 1909		X	X
= Xysta Eschscholtz, 1829
= Holeleodes Blaisdell, 1937
SGTricheleodes Blaisdell, 1909		X
Eleodimorpha Blaisdell, 1909		X
Embaphion Say, 1824		X
Lariversius Blaisdell, 1947		X
Neobaphion Blaisdell, 1925		X
Nycterinus Eschscholtz, 1829			X
Trogloderus LeConte, 1879		X
Blaptini Leach, 1815
Blaptina Leach, 1815
Ablapsis Reitter, 1887				X
Blaps Fabricius, 1775		X		X	X	X	X
SGArenoblaps G.S. Medvedev, 1999				X
SGBlaps Fabricius, 1775		X		X	X	X	X
= Leptomorpha Faldermann, 1835
= Agroblaps Motschulsky, 1860
= Blapimorpha Motschulsky, 1860
= Blapisa Motschulsky, 1860
= Lithoblaps Motschulsky, 1860
= Platyblaps Motschulsky, 1860
= Rhizoblaps Motschulsky, 1860
= Uroblaps Motschulsky, 1860
= Leptocolena Allard, 1880
= Acanthoblaps Reitter, 1889
= Blapidurus Fairmaire, 1891
= Blapidium Bauer, 1921
= Hypoblaps Kolbe, 1928
= Mesooblaps Kolbe, 1928
= Notoblaps Kolbe, 1928
= Opisthoblaps Kolbe, 1928
= Nanoblaps Semenov & Bogatchev, 1936
SGDineria Motschulsky, 1860				X
= Laraliprosodes Bogatchev, 1947
SGGenoblaps Bauer, 1921				X		X
= Prosoblapsia Skopin & Kaszab, 1978
Caraboblaps Bauer, 1921*
Coelocnemodes Bates, 1879						X
= Neoblaps Ren & Li, 2001
Dila Fischer von Waldheim, 1844				X
= Caenoblaps König, 1906
Dilablaps Bogatchev, 1976				X
Holoblaps Bauer, 1921*
Hoplitoblaps Fairmaire, 1889						X
Medvedevia Chigray, 2019				X
Medvedevoblaps Bouchard & Bousquet, nom. nov.				X
= Protoblaps G.S. Medvedev, 1998
Nalepa Reitter, 1887				X
Periblaps Bauer, 1921*
Protoblaps Bauer, 1921*
Thaioblaps Masumoto, 1989						X
Thaumatoblaps Kaszab & G.S. Medvedev, 1984				X
Gnaptorina G.S. Medvedev, 2001
Gnaptor Brullé, 1831				X
SGGnaptor Brullé, 1831				X
SGPlesiognaptor Chigray, Nabozhenko & Keskin, 2015				X
Gnaptorinina G.S. Medvedev, 2001
Agnaptoria Reitter, 1887				X
Asidoblaps Fairmaire, 1886				X		X
Belousovia G.S. Medvedev, 2007				X
Blaptogonia G.S. Medvedev, 1998				X		X
Colasia Koch, 1965				X
Gnaptorina Reitter, 1887				X		X
SGAustroptorina Bai, Li & Ren, 2020						X
SGBoreoptorina G.S. Medvedev, 2009				X
SGGnaptorina Reitter, 1887				X
SGHesperoptorina G.S. Medvedev, 2009				X		X
Itagonia Reitter, 1887				X
Montagona G.S. Medvedev, 1998				X		X
Nepalindia G.S. Medvedev, 1998						X
Pseudognaptorina Kaszab, 1977				X		X
Sintagona G.S. Medvedev, 1998				X
Tagonoides Fairmaire, 1886				X
Viettagona G.S. Medvedev & Merkl, 2003						X
Prosodina Skopin, 1960
Prosodes Eschscholtz, 1829				X		X
SGDilopersina Reitter, 1909				X
SGDiprosodes Reitter, 1909				X
SGEuryprosodes Reitter, 1909				X
SGFerganoprosodes G.S. Medvedev, 1997				X
SGGebleria Motschulsky, 1846				X
= Aulonoscelis Reitter, 1896
SGHypoprosodes Reitter, 1909				X
SGIndoprosodes G.S. Medvedev, 2003						X
SGIranosodes G.S. Medvedev, 1996				X
SGLyprosodes Reitter, 1909				X
SGMegaprosodes Reitter, 1909				X
= Altiprosodes G.S. Medvedev, 1997
SGMeropersina Reitter, 1909				X
SGMesoprosodes G.S. Medvedev, 1995				X
SGMontiprosodes G.S. Medvedev, 2001				X
SGOliprosodes Reitter, 1909				X
SGPeltarium Fischer von Waldheim, 1844				X
= Blaposodes Skopin, 1960
SGPlanoprosodes G.S. Medvedev, 2005				X
SGProsodella Reitter, 1909				X
= Paraprosodes Reitter, 1909
SGProsodes Eschscholtz, 1829				X
= Nyctipates Gebler, 1841
= Eupomeca Solier, 1848
= Blaptoprosodes Reitter, 1909
= Lioprosodes Reitter, 1909
= Platyprosodes Reitter, 1909
= Prosodila Reitter, 1909
= Pseudoprosodes Reitter, 1909
SGProsodestes Reitter, 1909				X
SGProsodinia Reitter, 1909				X
SGProsodopria Reitter, 1909				X
= Nyctipates Solier, 1848
SGProsodoscelis Reitter, 1909				X
SGProsodura Reitter, 1909				X
SGTangiprosodes G.S. Medvedev, 2005				X
SGUroprosodes Reitter, 1909				X
Tagona Fischer von Waldheim, 1820				X
Remipedellina Semenov, 1907
Remipedella Semenov, 1907				X
Dendarini Mulsant & Rey, 1854
Dendarina Mulsant & Rey, 1854
Bioplanes Mulsant, 1854				X
Dendarophylan Español, 1945				X
Dendarus Dejean, 1821				X
SGDendaroscelis Reitter, 1904				X
SGDendarus Dejean, 1821				X
= Pandarus Dejean, 1834
= Microdendarus Escalera, 1944
= Reitterellus Escalera, 1944
SGPandarinus Mulsant & Rey, 1854				X
SGParoderus Mulsant & Rey, 1854				X
= Dichromma Mulsant & Rey, 1855
SGRhizalemus Reitter, 1904				X
SGRizalus Mulsant & Rey, 1854				X
= Rhizalus Gebien, 1938
Heliopates Dejean, 1834				X
SGHeliocrates Reitter, 1904				X
SGHeliopates Dejean, 1834				X
= Heliocaes Bedel, 1906
= Heliopathes Gebien, 1938
Litoboriolus Español, 1945				X
Litororus Reitter, 1904				X
Meglyphus Motschulsky, 1872					X
Microphylacinus Iwan, Kamiński & Aalbu, 2011					X
Micrositus Mulsant & Rey, 1854				X
Neoisocerus Bouchard, Lawrence, Davies & Newton, 2005				X
= Isocerus Dejean, 1821
Phylacinus Fairmaire, 1896					X
Phylan Sturm, 1826				X
SGEumicrositus Viñolas, 1990				X
SGMeladeras Mulsant & Rey, 1854				X
SGMeladocrates Reitter, 1904				X
SGPhylan Sturm, 1826				X
= Heliophilus Dejean, 1821
= Phylax Brullé, 1832
= Olocrates Mulsant, 1854
SGPlatyolus Mulsant & Rey, 1854				X
Phylanmania Ferrer, 2013				X
Pythiopus Koch, 1953					X
Melambiina Mulsant & Rey, 1854
Allophylax Bedel, 1906				X
SGAllophylax Bedel, 1906				X
= Neophylax Bedel, 1906
= Melambiophylax Schuster, 1922
SGLitoboromimus Koch, 1948				X
SGPhylaximon Koch, 1948				X
Bermejoina Español, 1944				X
Gridelliopus Koch, 1956					X
Guildia Antoine, 1957				X
Hadroderus Koch, 1956					X
Haemodus Gebien, 1943					X
= Haemus Péringuey, 1904
Hanstroemium Koch, 1953					X
Hoplarion Mulsant & Rey, 1854				X
SGAtlasion Koch, 1948				X
= Megatlasion Koch, 1948
SGGlyptariobius Koch, 1948				X
SGHoplariobius Reitter, 1904				X
SGHoplarion Mulsant & Rey, 1854				X
SGMentariobius Koch, 1948				X
SGSaharoplarion Koch, 1948				X
Lasioderus Mulsant & Rey, 1854					X
Litoborus Mulsant & Rey, 1854				X
SGLitoborus Mulsant & Rey, 1854				X
SGParalitoborus Antoine, 1931				X
Melambius Mulsant & Rey, 1854				X
SGHadromelambius Koch, 1948				X
SGHoplambius Reitter, 1904				X
SGMelambatlasus Koch, 1948				X
SGMelambius Mulsant & Rey, 1854				X
Melansis Wollaston, 1864				X
Melasmana Strand, 1935				X
SGHeliomelasma Koch, 1948				X
SGMelasmana Strand, 1935				X
= Melasma Wollaston, 1864
Minorus Mulsant & Rey, 1854					X
Orarabion Leo & Liberto, 2011					X
Oreomelasma Español, 1975				X
Otinia Antoine, 1942				X
SGAntoineius Koch, 1948				X
SGOrophylaxus Koch, 1948				X
SGOtinia Antoine, 1942				X
Peyerimhoffius Koch, 1948				X
Psammoardoinellus Leo, 1981				X
Pseudemmallus Koch, 1956					X
Silvestriellum Koch, 1956					X
Selenepistoma Dejean, 1834					X
SGSelenepistoma Dejean, 1834					X
= Euzadenos Koch, 1956
SGSerridenos Koch, 1956					X
SGZadenos Laporte, 1840					X
Tragardhus Koch, 1956					X
SGMitragardhus Koch, 1956					X
SGTragardhus Koch, 1956					X
Zoutpansbergia Koch, 1956					X
Opatrini Brullé, 1832
Ammobiina Desbrochers des Loges, 1902
Adavius Mulsant & Rey, 1859				X		X
= Udebra Reitter, 1896
Ammidium Erichson, 1843					X
= Eremonomus Wollaston, 1861
= Ecripsis Pascoe, 1866
Ammobius Guérin-Méneville, 1844				X		X
= Ammophtorus Lacordaire, 1859
Amphithrixoides Bouchard & Löbl, 2008				X
= Amphithrix Español, 1953
Arabammobius Grimm & Lillig, 2020				X	X
Asiocaedius G.S. Medvedev & Nepesova, 1985				X
= Pseudocaedius G.S. Medvedev, 1966
Brachyidium Fairmaire, 1883						X	X	X
= Cnemodasus Gebien, 1914
Caediexis Lebedev, 1932				X
Caedius Blanchard, 1845				X	X	X	X
= Isarida Pascoe, 1866
Clitobius Mulsant & Rey, 1859				X	X
= Halonomus Wollaston, 1861
= Apithesis C.O. Waterhouse, 1881
= Apteroclitobius Koch, 1960
Coeloecetes Blair, 1929						X
Corinta Koch, 1950					X
Cornopterus Koch, 1950					X
Cyptus Gerstaecker, 1871				X	X
= Neocaedius Pierre, 1972
Diaderma Koch, 1960					X
Dilamus Jacquelin du Val, 1861				X	X
SGChoresmolamus G.S. Medvedev, 1978				X
SGDilamus Jacquelin du Val, 1861				X	X
SGOchrolamus Reitter, 1904				X	X
Emmalus Erichson, 1843					X
= Emmallus Agassiz, 1846
= Uzagaria Ancey, 1881
Falsammidium Koch, 1960					X
Falsocaedius Español, 1943				X
Freyula Koch, 1959					X
Hadrodes Wollaston, 1877				X
Helenomelas Ardoin, 1972				X
Mateuina Español, 1944				X
Messoricolum Koch, 1960					X
Moragacinella Español, 1954				X
= Moralesia Kaszab, 1944
Nesocaedius Kolbe, 1915				X		X
Perithrix Fairmaire, 1879				X
Platyprocnemis Español & Lindberg, 1963					X
Plesioderes Mulsant & Rey, 1859					X	X
= Epeurycaulus Kolbe, 1902
Prodilamus Ardoin, 1969				X	X
Proscheimus Desbrochers des Loges, 1881				X
Psammestus Reichardt, 1936				X
Pseudoleichenum Ardoin, 1972					X
Raynalius Chatanay, 1912					X
Tarphiophasis Wollaston, 1877					X
Trigonopoda Gebien, 1914						X
Weisea Semenov, 1891				X
Blapstinina Mulsant & Rey, 1853
Aconobius Casey, 1895		X
Ammodonus Mulsant & Rey, 1859		X	X
= Pseudonomus Fairmaire, 1884
= Scaptes Champion, 1886
= Trichotoides Marcuzzi, 1954
Austrocaribius Marcuzzi, 1954			X
Blapstinus Dejean, 1821		X	X					X
= Heteropus Laporte, 1840
= Pedonoeces G.R. Waterhouse, 1845
= Pedoeces Agassiz, 1846
= Tessaromma Boheman, 1858
= Aspidius Mulsant & Rey, 1859
= Lachnoderes Mulsant & Rey, 1859
= Lodinus Mulsant & Rey, 1859
= Mecysmus Horn, 1870
Cenophorus Mulsant & Rey, 1859			X
Conibiosoma Casey, 1890		X
Conibius LeConte, 1851		X	X					X
= Euconibius Casey, 1895
= Gondwanodilamus Kaszab, 1969 syn. nov.
= Ooconibius Casey, 1895
Cybotus Casey, 1890			X
Diastolinus Mulsant & Rey, 1859			X
= Sellio Mulsant & Rey, 1859
= Ctesicles Champion, 1896
Goajiria Ivie & Hart, 2016			X
Hummelinckia Marcuzzi, 1954			X
Nevisia Marcuzzi, 1986			X
Nocibiotes Casey, 1895		X
Notibius LeConte, 1851		X
Platylus Mulsant & Rey, 1859			X
Tonibiastes Casey, 1895		X
Tonibius Casey, 1895		X
Trichoton Hope, 1841		X	X
= Epilasium Erichson, 1842
= Trichotum Agassiz, 1846
= Bycrea Pascoe, 1868
Ulus Horn, 1870		X	X
Xerolinus Ivie & Hart, 2016			X
Heterotarsina Blanchard, 1845
Diphyrrhynchus Fairmaire, 1849				X	X	X	X	X
= Acanthosternus Montrouzier, 1860
= Abantis Fairmaire, 1892
= Abantiades Fairmaire, 1894
= Neoabantis Gebien, 1910
Heterocheira Dejean, 1836					X		X
= Heterochira Agassiz, 1846
Heterotarsus Latreille, 1829				X	X	X
= Helopimorphus Desbrochers des Loges, 1881
= Hopatropteron Reitter, 1889
= Oubanghinum Pic, 1933
Scymena Pascoe, 1866							X
Neopachypterina Bouchard, Löbl & Merkl, 2007
Amblysphagus Fairmaire, 1896					X	X
= Trachymetus Reitter, 1904
†Eupachypterus Kirejtshuk, Nabozhenko & Nel, 2010
Neopachypterus Bouchard, Löbl & Merkl, 2007				X		X
= Pachypterus P.H. Lucas, 1847
Pseudolamus Fairmaire, 1874				X
Opatrina Brullé, 1832
Anatrum Reichardt, 1936				X
Brachyesthes Fairmaire, 1868				X		X
Caediomorpha Blackburn, 1888							X
Ephalus LeConte, 1862		X	X
= Pseudephalus Casey, 1924
Eumylada Reitter, 1904				X
Falsolobodera Kaszab, 1967				X
Gonocephalum Solier, 1834		X		X	X	X	X	X
SGGonocephalum Solier, 1834		X		X	X	X	X	X
= Dasus Motschulsky, 1845
= Megadasus Reitter, 1904
= Hasticollinum Kaszab, 1939
SGMyladanesthes Skopin, 1961				X
SGOpatropis Reitter, 1904				X	X	X
Hadrophasis Ferrer, 1992					X
Jintaium Ren, 1999				X
Melanesthes Dejean, 1834				X
SGLesbidana Reitter, 1904				X
SGMelanesthes Dejean, 1834				X
= Hemitrichestes Reitter, 1904
= Miglica Reitter, 1904
SGMongolesthes Reitter, 1904				X
SGOpatronesthes Reitter, 1904				X
Melanocoma Wollaston, 1868					X
Mesomorphus Miedel, 1880				X	X	X	X	X
= Pentholasius Reitter, 1904
= Hopatromorpha Blackburn, 1907
Myladina Reitter, 1889				X
Opatroides Brullé, 1832		X		X	X	X
= Hopatroides Agassiz, 1846
Opatrum Fabricius, 1775				X
SGColpopatrum Reitter, 1904				X
SGColpophorus Mulsant & Rey, 1859				X
= Colpophorinus Escalera, 1914
SGOpatrum Fabricius, 1775				X
= Hopatrum Agassiz, 1846
= Thoracon Gistel, 1848
Penthicinus Reitter, 1896				X
Penthicus Faldermann, 1836				X	X
SGAllomyladion Bogatchev, 1972				X
SGAulonolcus Reitter, 1904				X
SGDiscotus Reitter, 1904				X	X
SGMyladion Reitter, 1887				X
= Penthicomelanesthes Bogatchev, 1972
SGPenthicus Faldermann, 1836				X
= Lobodera Mulsant & Rey, 1859
= Lobothorax Gemminger, 1870
= Penthomegus Reitter, 1904
= Stonavus Reitter, 1904
SGPseudopenthicinus Bogatchev, 1972				X
Phelopatrum Marseul, 1876				X
= Pseudadrus Fairmaire, 1897
Polycoelogastridion Reichardt, 1936				X		X
Reichardtiellina Kaszab, 1982				X
= Reichardtiella Kaszab, 1942
Scleropatroides Löbl & Merkl, 2003				X	X	X
Scleropatrum Reitter, 1887				X		X
= Monatrum Reichardt, 1936
Sinorus Mulsant & Revelière, 1861				X
Sobas Pascoe, 1863							X
= Trigonotarsus Hope, 1843
= Pseudocaedius Blackburn, 1890
Socotropatrum Koch, 1970					X
Tidiguinia Español, 1959				X
Trichosternum Wollaston, 1861					X
= Trichopodus Mulsant & Rey, 1859
= Japetus Reitter, 1904
Wolladrus Iwan & Kamiński, 2016				X
= Hadrus Wollaston, 1854
Sclerina Lacordaire, 1859
Eurycaulus Fairmaire, 1868				X	X
SGEurycaulinus Koch, 1937				X
SGEurycaulus Fairmaire, 1868				X	X
= Ammotrypes Fairmaire, 1879
= Scleronimon Reitter, 1904
= Scleronopsis Koch, 1935
†Palaeosclerum Nabozhenko & Kirejtshuk, 2017
Platynosum Mulsant & Rey, 1859				X
= Melanimon Motschulsky, 1845
Sclerum Dejean, 1834				X	X	X
= Scleron Hope, 1841
= Anticlia Gistel, 1848
= Chlamydion Gistel, 1848
Stizopodina Lacordaire, 1859
Adoryacus Koch, 1963					X
Amathobius Gebien, 1920					X
Blacodatus Koch, 1963					X
Blenosia Laporte, 1840					X
= Blacodes Duponchel, 1842
Calaharena Koch, 1963					X
Crististibes Koch, 1963					X
Eichleria Kamiński, 2015					X
Eremostibes Koch, 1963					X
Helibatus Mulsant & Rey, 1859					X
= Ennychius Fåhraeus, 1870
Luebbertia Koch, 1963					X
Microstizopus Koch, 1963					X
Namazopus Koch, 1963					X
Nemanes Fairmaire, 1888					X
Parastizopus Gebien, 1938					X
= Ennychiatus Koch, 1963
Periloma Gebien, 1938					X
Planostibes Gemminger, 1870					X
= Planodes Mulsant & Rey, 1859
Psammogaster Koch, 1953					X
Sphaerostibes Koch, 1963					X
Stizopus Erichson, 1843					X
= Doryagus Pascoe, 1887
Sulpius Fairmaire, 1906					X
Syntyphlus Koch, 1953					X
Opatrini incertae sedis
Hovarygmus Fairmaire, 1898					X
Pachymastus Fairmaire, 1896					X
Pocadiopsis Fairmaire, 1896						X
Scleroides Fairmaire, 1883							X
Trigonopilus Fairmaire, 1893						X
Pedinini Eschscholtz, 1829
Helopinina Lacordaire, 1859
Amatodes Dejean, 1834					X
SGAmatodes Dejean, 1834					X
= Oncosoma Westwood, 1843
= Oncoosoma Gebien, 1911
SGConophthalmus Quedenfeldt, 1885					X
SGStrophiamixa Robiche, 2005					X
= Strophia Robiche, 2004
Ametrocera Fåhraeus, 1870					X
= Idricus Fairmaire, 1888
Anaxius Fåhraeus, 1870					X
Aptila Fåhraeus, 1870					X
Asidodema Koch, 1958					X
Blastarnodes Koch, 1958					X
Diestecopus Solier, 1848					X
= Blastarnus Fairmaire, 1897
Drosochrus Erichson, 1843				X	X
SGDesertosochrus Koch, 1958					X
SGDrosochrus Erichson, 1843				X	X
SGHelopinus Solier, 1848				X	X
= Pteraulus Solier, 1848
= Emyon Gerstaecker, 1854
Micrantereus Solier, 1848				X	X
= Solenomerus Fåhraeus, 1870
Nicandra Fairmaire, 1888					X
SGCalous Koch, 1958					X
SGHeteronicandra Koch, 1958					X
SGNicandra Fairmaire, 1888					X
SGOncotopsis Koch, 1958					X
Oncopteryx Gebien, 1943					X
= Oncopterus Fairmaire, 1887
Piscicula Robiche, 2004					X
Psectes Hesse, 1935					X
Leichenina Mulsant, 1854
Apsheronellus Bogatchev, 1967				X
= Microleichenum G.S. Medvedev, 1973
Leichenum Dejean, 1834		X	X	X	X	X	X
= Lichenum Agassiz, 1846
= Endothina Carter, 1924
Pedinina Eschscholtz, 1829
Cabirutus Strand, 1929				X		X
SGCabirutus Strand, 1929				X
= Cabirus Mulsant & Rey, 1853
= Asiobirus G.S. Medvedev, 1968
= Dentibirus G.S. Medvedev, 1968
SGNeocabirutus Kulzer, 1964						X
Colpotinus Fairmaire, 1891				X
Loensus R. Lucas, 1920					X	X
SGLoensus R. Lucas, 1920					X
= Pedinopsis Gebien, 1910
SGPseudopedinus Ardoin, 1969						X
Pedinus Latreille, 1797				X		X
SGBlindus Mulsant & Rey, 1853				X		X
SGColpotus Mulsant & Rey, 1853				X
SGPedinus Latreille, 1797				X
= Vadalus Mulsant & Rey, 1853
= Pedinulus Seidlitz, 1893
Platynotini Mulsant & Rey, 1853
Eurynotina Mulsant & Rey, 1854
Byrrhoncus Koch, 1954					X
Capidium Koch, 1954					X
Colophonesthes Koch, 1954					X
Eurynotus W. Kirby, 1819					X
SGBiolus Mulsant & Rey, 1854					X
SGEurynotus W. Kirby, 1819					X
SGNeosolenopistoma Bouchard & Bousquet, subgen. nov.					X
Heteropsectropus Kaszab, 1941					X
Hirtograbies Koch, 1954					X
Isoncophallus Koch, 1954					X
Menederes Solier, 1848					X
SGAnamenederes Koch, 1954					X
SGMenederes Solier, 1848					X
Menederopsis Koch, 1954					X
= Archinamaqua Schawaller, 2012
Ograbies Péringuey, 1899					X
Oncotus Blanchard, 1845					X
SGCilioncotus Koch, 1954					X
SGMenoncotus Koch, 1954					X
SGOncotus Blanchard, 1845					X
SGQuadroncotus Koch, 1954					X
Phaleriderma Koch, 1954					X
Phylacastus Fairmaire, 1897					X
Psectrapus Solier, 1848					X
= Psectropus Gemminger, 1870
Schyzoschelus Koch, 1954					X
Stridigula Koch, 1954					X
Platynotina Mulsant & Rey, 1853
Adamus Iwan, 1997						X
Alaetrinus Iwan, 1995		X	X
Amblychirus Koch, 1956					X
Anchophthalmops Koch, 1956		X			X
= Platykochius Iwan, 2002
Anchophthalmus Gerstaecker, 1854					X
SGAnchophthalmus Gerstaecker, 1854					X
= Oxythorax Fåhraeus, 1870
= Oncotiphallops Koch, 1956
SGKochogaster Kamiński & Raś, 2011					X
= Cosmogaster Koch, 1956
Angolositus Koch, 1955					X
= Aberlencus Iwan, 2002
= Platymedvedevia Iwan & Banaszkiewicz, 2007
Anomalipus Guérin-Méneville, 1831					X
= Heteroscelis Latreille, 1828
= Ectatocnemis Horn, 1867
= Acmoeus Fåhraeus, 1870
= Apodemus Fåhraeus, 1870
Atrocrates Koch, 1956					X
Atrocrypticanus Iwan, 1999					X
Bantodemus Koch, 1955					X
Clastopus Fairmaire, 1898					X
= Hovademulus Iwan, 1996
Claudegirardius Iwan, 1999					X
Colpotinoides Kaszab, 1975					X
Crypticanus Fairmaire, 1897					X
Doyenus Iwan, 1996					X
Ectateus Koch, 1956					X
Eleoselinus Kamiński, 2014					X
Eucolus Mulsant & Rey, 1853						X
= Indeucolus Kaszab, 1975
Eviropodus Koch, 1956					X
Glyptopteryx Gebien, 1910					X
= Microselinus Koch, 1956
= Quadrideres Koch, 1956
= Synquadrideres Iwan, 2003
Gonopus Latreille, 1828					X
SGAgonopus Gebien, 1920					X
SGGonopus Latreille, 1828					X
Hovademus Iwan, 1996					X
Lechius Iwan, 1995					X
Madobalus Fairmaire, 1901					X
Melanocratus Fairmaire, 1895					X
Melanopterus Mulsant & Rey, 1854					X
Menearchus Carter, 1920					X
Monodius Koch, 1956					X
Nesopatrum Gebien, 1921					X
Notocorax Dejean, 1834						X
= Platydendarus Kaszab, 1975
Opatrinus Dejean, 1821			X
= Hopatrinus Agassiz, 1846
= Dema Gistel, 1848
Parabantodemus Iwan, 2000					X
Paraselinus Kamiński, 2013					X
Penthicoides Fairmaire, 1896						X
Phallocentrion Koch, 1956					X
Phymatoplata Koch, 1956					X
Platyburak Iwan, 1990						X
Platyburmanicus Iwan, 2003						X
Platycolpotus Iwan, 1997						X
Platynotoides Kaszab, 1975						X
Platynotus Fabricius, 1801						X
= Ixalus Gistel, 1848
Pokryszkiella Iwan, 1996					X
Pseudoblaps Guérin-Méneville, 1834				X		X
= Nyctalops Gistel, 1848
Pseudonotocorax Iwan, 1997						X
Pteroselinus Kamiński, 2015					X
Rugoplatynotus Kaszab, 1975						X
Schelodontes Koch, 1956					X
= Lawrenceus Iwan, 1998
= Platycharlesus Iwan, 1998
= Warchalowskiellus Iwan, 1998
Sebastianus Iwan, 1996					X
Selinopodus Koch, 1956					X
Selinus Mulsant & Rey, 1853					X
Stenogonopus Gebien, 1938					X
Styphacus Fairmaire, 1901					X
Trigonopus Mulsant & Rey, 1853					X
Upembarus Koch, 1956					X
SGPseudoselinus Iwan, 2002					X
SGUpembarus Koch, 1956					X
Zidalus Mulsant & Rey, 1853				X	X
= Zodinus Mulsant & Rey, 1853
= Apterozidalus Ardoin, 1965
Zophodes Fåhraeus, 1870					X
Platyscelidini Lacordaire, 1859
Bioramix Bates, 1879				X
SGBioramix Bates, 1879				X
SGCardiochianalus Kaszab, 1940				X
SGChianalus Bates, 1879				X
= Botiras Fairmaire, 1891
SGEuryhelops Reitter, 1902				X
= Cardiobioramix Kaszab, 1940
SGFaustia Kraatz, 1882				X
SGLeipopleura Seidlitz, 1893				X
SGNudoplatyscelis Kaszab, 1940				X
SGOvalobioramix Egorov, 2004				X
SGPlanoplatyscelis Kaszab, 1940				X
= Pleioplatyscelis Kaszab, 1940
SGPlatynoscelis Kraatz, 1882				X
SGTrichochianalus Kaszab, 1940				X
SGTricholeipopleura Kaszab, 1940				X
SGTrichoplatyscelis Reinig, 1931				X
= Pseudotrichoplatyscelis Kaszab, 1960
Microplatyscelis Kaszab, 1940				X
Myatis Bates, 1879				X
Oodescelis Motschulsky, 1845				X
SGAcutoodescelis Kaszab, 1940				X
SGClavatoodescelis Kaszab, 1940				X
= Oblongoodescelis Kaszab, 1940
SGConvexoodescelis Egorov, 2004				X
SGLonguloodescelis Kaszab, 1940				X
= Trichoodescelis Kaszab, 1940
SGMontanoodescelis Egorov, 2004				X
SGOodescelis Motschulsky, 1845				X
= Oodeoscelis Agassiz, 1846
SGOvaloodescelis Kaszab, 1940				X
SGPlanoodescelis Egorov, 2004				X
SGSpinoodescelis Kaszab, 1940				X
SGSplenoodescelis Egorov, 2004				X
SGTruncatoodescelis Kaszab, 1940				X
Platyscelis Latreille, 1818				X
SGOblongoplatyscelis Kaszab, 1940				X
SGParaplatyscelis Kaszab, 1940				X
SGPlatyscelis Latreille, 1818				X
= Kaszaboscelis Löbl & Merkl, 2003
SGPleiopleura Seidlitz, 1893				X
Somocoelia Heyden & Kraatz, 1882				X
Somocoeloplatys Skopin, 1968				X
Trichomyatis Schuster, 1931				X
= Trichoplatynoscelis Kaszab, 1940
Blaptinae incertae sedis
Stenolamus Gebien, 1920					X
Tenebrioninae Latreille, 1802
Acropteronini Doyen, 1989
Acropteryx Gistel, 1831			X
= Acropteron Perty, 1832
= Sphenosoma Dejean, 1834
= Acropterum Agassiz, 1846
= Arthroplatus Solier, 1851
Alphitobiini Reitter, 1917
Alphitobius Stephens, 1829		X	X	X	X	X	X
= Heterophaga Dejean, 1834
= Cryptops Solier, 1851
= Proselytus Fåhraeus, 1870
= Microphyes W.J. MacLeay, 1872
= Latetribolium Lepesme, 1943
†Alphitopsis Kirejtshuk, Nabozhenko & Nel, 2011
Ardoinia Kaszab, 1969					X
Diaclina Jacquelin du Val, 1861				X	X	X	X
Epipedodema Gebien, 1921					X
Guanobius Grimm, 2008						X
Hoplopeltis Fairmaire, 1894						X
Peltoides Laporte, 1833					X
SGMicropeltoides Pic, 1916					X
SGPeltoides Laporte, 1833					X
= Oopiestus Chevrolat, 1833
Amarygmini Gistel, 1848
Alienoplonyx Bremer, 2019						X
Alymon Pascoe, 1866					X
= Ghaleca Péringuey, 1899
Amarygmus Dalman, 1823				X		X	X	X
SGAmarygmus Dalman, 1823				X		X	X	X
= Dietysus Pascoe, 1866
= Elixota Pascoe, 1866
= Eurypera Pascoe, 1870
= Dictysus Rye, 1874
= Aphyllocerus Fairmaire, 1881
= Anacycus Fairmaire, 1896
= Platolenes Gebien, 1914
= Pseudamarygmus Pic, 1915
= Apelina Saha, 1988
= Plesiamarygmus Masumoto, 1989
SGBecvaramarygmus Masumoto, 1999						X
SGCornugeton Bremer, 2016							X
SGDryadigmus Bremer, 2007						X
SGHyperamarygmus Kaszab, 1964							X
SGOogeton Kaszab, 1941						X
SGPhaenogeton Bremer, 2016							X
SGPodamarygmus Carter, 1928						X
SGPyanirygmus Pic, 1915						X
SGVarogeton Bremer, 2014						X
Asthenochirus Fairmaire, 1885					X
Asyleptus Péringuey, 1896					X
= Barlacus Fairmaire, 1900
= Termitonebria Wasmann, 1925
= Falsozialeus Pic, 1951
Atropsorodes Ardoin, 1963					X
Axynaon Blackburn, 1897							X
= Catopherus Carter, 1918
Azarelius Fairmaire, 1892						X
Bunamarygmus Masumoto, 1988						X
Cantaloubeus Ardoin, 1959					X
Caudamarygmus Bremer, 2001						X
Cephalamarygmus Bremer, 2001						X
Cerysia Bremer, 2001						X	X
Chalcoplonyx Ardoin, 1963					X
Chalcopteroides Gebien, 1948							X
= Chalcopterus Blessig, 1861
Cheiroplus Ardoin, 1963					X
Cleognathus Gebien, 1921					X
Coccimarygmus Ardoin, 1966					X
SGCoccimarygmus Ardoin, 1966					X
SGLeiochromimus Ardoin, 1966					X
Crypsinous Fairmaire, 1891					X
= Cryptadius Fairmaire, 1894
= Ubangia Gebien, 1914
Cymatothes Dejean, 1834		X	X
= Physignathus Gistel, 1834
= Pyanisia Laporte, 1840
= Cymatodes Agassiz, 1846
Dalmanius Bremer, 2001						X
Dasyplonyx Bremer, 2014						X
Dichotymus Fairmaire, 1891					X
Erycastus Fairmaire, 1897					X
Euglyptonotus Gestro, 1901					X
Eulytus C.O. Waterhouse, 1882					X
Eumolpamarygmus Pic, 1923						X
Eumolparamarygmus Bremer, 2006						X
Eupezoplonyx Pic, 1922						X
Eupezus Dejean, 1834					X
Euspinamarygmus Masumoto, 1989						X
Fahraeus Ardoin, 1963					X
Falsastenochirus Pic, 1938					X
= Falsasthenochirus Ardoin, 1965
Falsoplonyx Ardoin, 1963					X
Falsosynopticus Pic, 1936					X
Garambanus Ardoin, 1964					X
Gonocnemis J. Thomson, 1858					X	X
= Acastus Péringuey, 1896
Gonocnemocistela Pic, 1935					X
Hesseodes Ardoin, 1963					X
Hoplobrachium Fairmaire, 1886					X	X
= Holobrachium Gebien, 1905
= Cephaloplonyx Pic, 1922
Hoplonyx J. Thomson, 1858					X
SGHoplonyx J. Thomson, 1858					X
= Hoplochirus Scudder, 1882
SGHyloplonyx Ardoin, 1963					X
SGNataloplonyx Ardoin, 1963					X
SGNemoplonyx Ardoin, 1963					X
Hypamarygmus Gebien, 1904					X
Insolitoplonyx Bremer, 2014						X
Isopteroplonyx Bremer, 2006							X
Javamarygmus Pic, 1928						X
Lemoultia Chatanay, 1913					X
Lobatopezus Pic, 1952						X
Luzonoplonyx Bremer, 2009						X
Macrosynopticus Pic, 1922						X
= Cyrtostrongylium Blair, 1929
Megacantha Westwood, 1843					X
Meracantha W. Kirby, 1837		X
= Falacer Laporte, 1840
= Physocoelus Haldeman, 1850
Meroxys Ardoin, 1963					X
Mimosynopticus Pic, 1922					X
Neoplonyx Ardoin, 1963					X
Nepaloplonyx Bremer, 2014						X
Nesioticus Westwood, 1843					X
Oplocheirus Lacordaire, 1859					X
Overlaetia Pic, 1937					X
Paragonocnemis Kraatz, 1899					X	X
SGBorneogonocnemis Pic, 1936						X
SGLycogonocnemis Pic, 1915					X
SGMicrogonocnemis Pic, 1936					X
SGParagonocnemis Kraatz, 1899					X
Paramarygmus Quedenfeldt, 1885					X
SGMeracanthoides Linell, 1896					X
SGParamarygmus Quedenfeldt, 1885					X
= Pareupezus Kolbe, 1889
Pilosoplonyx Bremer, 2014						X
Pimelionotus Ardoin, 1963					X
Platypsorodes Ardoin, 1963					X
Plegacerus Gebien, 1921					X
Plesiophthalmus Motschulsky, 1857		X		X		X
SGChaeroplonyx Bremer, 2014						X
SGCyriogeton Pascoe, 1871						X
SGEumolpocyriogeton Pic, 1922						X
SGInspinogeton Pic, 1937						X
SGOpacoplonyx Bremer, 2014				X		X
SGPlesiophthalmus Motschulsky, 1857		X		X		X
SGSpinamarygmus Pic, 1915						X
Plinthochrous Fairmaire, 1891					X
= Lycoscelis Blair, 1929
Podacamptus Ardoin, 1964					X
Pontianacus Fairmaire, 1898						X
Pseudalymon Ardoin, 1969					X
Pseudoogeton Masumoto, 1989				X		X
Psilocastus Ardoin, 1963					X
Psoroderes Ardoin, 1962					X
Psorodes Dejean, 1834					X
= Acanthomera Latreille, 1828
= Acanthomerus Agassiz, 1846
Psorophodes Ardoin, 1963					X
Pterodes Ardoin, 1963					X
Pubamarygmus Pic, 1915							X
Reichenspergeria Wasmann, 1921						X
Seorsoplonyx Bremer, 2010						X
Singapura Gebien, 1925						X
Spathulipezus Gebien, 1921							X
Spinodietysus Pic, 1927						X
Stemmoderus Spinola, 1842					X
= Stemmatoderus Agassiz, 1846
Sylvanoplonyx Bremer, 2010						X
Timogebienus Ardoin, 1963					X
Trichamarygmus Carter, 1913							X
Umslatus Péringuey, 1899					X
Vutsimus Péringuey, 1899					X
Ziaelas Fairmaire, 1892						X
Apocryphini Lacordaire, 1859
Apocrypha Eschscholtz, 1831		X	X
= Compsomorphus Solier, 1851
Diplocyrtus Quedenfeldt, 1887				X
Plastica C.O. Waterhouse, 1903			X
Pseudapocrypha Champion, 1886			X
Bolitophagini W. Kirby, 1837
Afrobyrsax Ardoin, 1973					X
Atasthalomorpha Miyatake, 1964				X
Atasthalus Pascoe, 1871						X
Boletoxenus Motschulsky, 1858				X		X
= Bolitoxenus Gemminger, 1870
Bolitolaemus Gebien, 1921					X
Bolitonaeus Lewis, 1894				X		X
Bolitophagiella Miyatake, 1964				X
Bolitophagus Illiger, 1798		X		X
= Boletophagus Agassiz, 1846
Bolitotherus Candèze, 1861		X
= Phellidius LeConte, 1862
Bolitotrogus Miyatake, 1964				X		X
Byrsax Pascoe, 1860				X		X	X
Eleates Casey, 1886		X
Eledona Latreille, 1797				X
= Heledona Agassiz, 1846
Eledonoprius Reitter, 1911				X
Lanhsia Shibata, 1980						X
Megeleates Casey, 1895		X
Microbolitonaeus Grimm, 2014						X
Microatasthalus Ando, 2010						X
Parabolitophagus Miyatake, 1964				X
†Proteleates Wickham,1914
Rhipidandrus LeConte, 1862		X	X	X	X	X	X	X
= Xyloborus Motschulsky, 1860
= Eutomus Lacordaire, 1865
= Heptaphylla Friedenreich, 1883
= Cherostus C.O. Waterhouse, 1894
= Bolitopertha Gebien, 1910
Sumbawia Gebien, 1925						X
Centronopini Doyen, 1989
Centronopus Solier, 1848		X	X
SGCentronopus Solier, 1848			X
= Centropus Jakobson, 1914
SGMenechides Motschulsky, 1872		X	X
= Scotobates Rye, 1877
= Pyres Champion, 1885
Scotobaenus LeConte, 1859		X
Tauroceras Hope, 1841			X
= Tauroceropedus Pic, 1913
Cerenopini Horn, 1870
Argoporis Horn, 1870		X
= Threnus Motschulsky, 1870
Cerenopus LeConte, 1851		X
Dissonomini G.S. Medvedev, 1968
Bradyus Dejean, 1834				X
= Aphaleria Reitter, 1896
Dissonomus Jacquelin du Val, 1861				X
SGDissonomus Jacquelin du Val, 1861				X
= Heterophylus Mulsant & Rey, 1859
SGEudissonomus G.S. Medvedev, 1968				X
SGNeodissonomus G.S. Medvedev, 1968				X
SGParadissonomus G.S. Medvedev, 1968				X
Eulabini Horn, 1870
Apsena LeConte, 1862		X
Epantius LeConte, 1851		X
Eulabis Eschscholtz, 1829		X
= Heterarthron Gistel, 1848
Falsocossyphini Ferrer, 2006
Blatticephalus Heller, 1918					X
= Catobleps Blair, 1918
Falsocossyphus Pic, 1916						X
Microblattellus Ferrer, 2006						X
Heleini Fleming, 1821
Asphalina Matthews & Lawrence, 2005
Asphalus Pascoe, 1868							X
Bassianus Matthews & Doyen, 1989							X
Meneristes Pascoe, 1869							X
= Asiris Motschulsky, 1872
Sloanea Carter, 1916							X
Cyphaleina Lacordaire, 1859
Agastenes R. Lucas, 1920							X
= Agasthenes Bates, 1873
= Batessia Ponting, 2018
Aglypta Gebien, 1908							X
= Onoglypta Carter, 1926
Amarygmimus Bates, 1873							X
= Amarygmomimus Rye, 1875
Amphianax Bates, 1873							X
Atoreuma Gebien, 1941							X
= Toreuma Carter, 1913
= Eutoreuma Carter, 1914
Bolbophanes Carter, 1913							X
= Pseudobolbophanes Kulzer, 1954
Byallius Pascoe, 1869							X
Cyphaleus Westwood, 1841							X
= Chrysobalus Boisduval, 1835
= Chartopteryx Westwood, 1841
= Oremasis Pascoe, 1866
= Altes Pascoe, 1869
= Anausis Bates, 1873
= Apomestris Bates, 1873
= Trisilus Haag-Rutenberg, 1878
Hemicyclus Westwood, 1841							X
= Cyclophanes Carter, 1913
Mithippia Pascoe, 1869							X
Mitrothorax Carter, 1914							X
= Ctimene Bates, 1873
= Mitrephorus Carter, 1913
= Timeneca Carter, 1914
Nyctozoilus Guérin-Méneville, 1831							X
= Sphenogenius Solier, 1848
= Onosterrhus Pascoe, 1866
= Hypocilibe Bates, 1872
= Aethalides Bates, 1873
= Ononyctus Carter, 1914
Olisthaena Erichson, 1842							X
= Decialma Pascoe, 1869
= Hectus Pascoe, 1869
= Aphectus Carter, 1926
Onotrichus Carter, 1911							X
Pachycoelia Boisduval, 1835							X
= Lepispilus Westwood, 1841
= Lepidospilus Agassiz, 1846
= Tyndarisus Pascoe, 1869
Paraphanes W.J. MacLeay, 1887							X
Phanechloros Matthews & Bouchard, 2008							X
= Chlorophanes Matthews, 1992
Platyphanes Westwood, 1849							X
= Opigenia Pascoe, 1869
= Laonicus Haag-Rutenberg, 1878
Prophanes Westwood, 1849							X
= Lygestira Pascoe, 1866
= Maerodes C.O. Waterhouse, 1877
= Moerodes Rye, 1879
Styrus Bates, 1873							X
Heleina Fleming, 1821
Boreosaragus Matthews, 1993							X
Brises Pascoe, 1869							X
= Ephidonius Pascoe, 1869
Camponotiphilus Lea, 1914							X
Celibe Boisduval, 1835							X
Cillibus Matthews, 1993							X
Dysarchus Pascoe, 1866							X
= Saragodinus Bates, 1872
= Saragella Carter, 1937
Edylius Champion, 1894							X
Emcephalus W. Kirby, 1828							X
= Encephalus Agassiz, 1846
= Ellaemus Pascoe, 1866
= Encara Gemminger, 1870
= Euhelaeus Gebien, 1921
Helea Latreille, 1804							X
= Soradeus Rafinesque, 1815
= Elaeus Gemminger, 1870
Mimopeus Pascoe, 1866							X
Ospidus Pascoe, 1866							X
Pterohelaeus Brême, 1842							X
= Barytipha Pascoe, 1869
= Pterelaeus Gemminger, 1870
= Pezohelaeus Gebien, 1921
Saragus Erichson, 1842							X
= Cyclosattus Casey, 1892
Sympetes Pascoe, 1866							X
Trichosaragus Blackburn, 1890							X
Heleini incertae sedis
Cerodolus Sharp, 1886							X
Pseudhelops Guérin-Méneville, 1841							X
Helopini Latreille, 1802
Cylindrinotina Español, 1956
Armenohelops Nabozhenko, 2002				X
Asialassus Nabozhenko & Ando, 2018				X		X
Ceratopelius Antoine, 1963				X
Cylindrinotus Faldermann, 1837				X
= Cylindronotus Agassiz, 1846
= Stenomacidius Seidlitz, 1895
Ectromopsis Antoine, 1949				X
Eustenomacidius Nabozhenko, 2006				X
SGCaucasohelops Nabozhenko, 2006				X
SGEustenomacidius Nabozhenko, 2006				X
Gunarus Gozis, 1886				X
Idahelops Keskin & Nabozhenko, 2012				X
Microdocnemis Nabozhenko & Keskin, 2010				X
Nalassus Mulsant, 1854		X		X		X
SGCaucasonotus Nabozhenko, 2000				X
SGHoristelops Gozis, 1910				X
= Helopondrus Reitter, 1922
SGNalassus Mulsant, 1854		X		X		X
= Helopocerodes Reitter, 1922
SGNipponalassus Nabozhenko & Ando, 2018				X
Odocnemis Allard, 1876				X
SGHeloponotus Reitter, 1922				X
SGOdocnemis Allard, 1876				X
= Omaleis Allard, 1876
= Isopedus Stein, 1877
= Homalus Rye, 1878
= Homaleis Rye, 1879
= Odontocnemis Rye, 1878
Pseudoprobaticus Nabozhenko, 2001				X
Reitterohelops Skopin, 1960				X
Stenomax Allard, 1876				X
SGPystelops Gozis, 1910				X
= Asyrmatus Canzoneri, 1959 syn. nov.
SGStenomax Allard, 1876				X
Stygohelops Leo & Liberto, 2003				X
Taurohelops Keskin & Nabozhenko, 2015				X
Turkmenohelops G.S. Medvedev, 1987				X
Turkonalassus Keskin, Nabozhenko & Alpagut-Keskin, 2017				X
Xanthohelops Nabozhenko, 2006				X
Xanthomus Mulsant, 1854				X
Zophohelops Reitter, 1902				X
SGZophohelops Reitter, 1902				X
= Euryhelops Reitter, 1902
SGZophondrus Nabozhenko, 2014				X
Enoplopodina Reitter, 1917
Accanthopus Dejean, 1821				X
= Enoplopus Solier, 1848
Helopina Latreille, 1802
Adelphinus Fairmaire & Coquerel, 1866				X
SGAdelphinops Reitter, 1922				X
SGAdelphinus Fairmaire & Coquerel, 1866				X
Allardius Ragusa, 1898				X
= Pseudoparablops Heyden, 1908
Apterotarpela Kaszab, 1954						X
Catomus Allard, 1876				X
SGCatomodontus Löbl & Merkl, 2020				X
SGCatomus Allard, 1876				X	X
= Catomidius Seidlitz, 1895
SGMontanocatomus Nabozhenko, 2006				X
SGSinocatomus Nabozhenko, 2006				X
†Cryptohelops Nabozhenko & Kirejtshuk, 2014
Deretus Gahan, 1900					X
Entomogonus Solier, 1848				X
SGDelonurops Reitter, 1922				X
= Macrophanes Iablokoff-Khnzorian, 1957
SGEntomogonus Solier, 1848				X
SGEutelogonus Reitter, 1922				X
Erionura Reitter, 1903				X
Euboeus Boieldieu, 1865				X
SGEuboeus Boieldieu, 1865				X
= Probaticus Seidlitz, 1895
SGHelopidoxus Reitter, 1922				X
SGHelopostygnus Antoine, 1949				X
SGHelopotrichus Reitter, 1922				X
SGPelorinus Vauloger de Beaupré, 1900				X
Hedyphanes Fischer von Waldheim, 1820				X
SGGranulophanes Nabozhenko, 2013				X
SGHedyphanes Fischer von Waldheim, 1820				X
= Coelophanes Iablokoff-Khnzorian, 1964
SGMicrohedyphanes Nabozhenko & Lillig, 2013				X
Helops Fabricius, 1775		X	X	X
= Hypulus Rafinesque, 1815
= Anteros Laporte, 1840
= Stenotrichus LeConte, 1862
= Biomorphus Motschulsky, 1872
= Coscinoptilix Allard, 1876
= Mesohelops Reitter, 1922
Italohelops Español, 1961				X
Mamorina Antoine, 1951				X
Nautes Pascoe, 1866		X	X
Neohelops Dajoz, 2001		X
Nephodinus Gebien, 1943				X
SGNephodinus Gebien, 1943				X
= Nephodes Blanchard, 1845
SGParanephodes Antoine, 1955				X
Nesotes Allard, 1876				X
SGHelopogonus Reitter, 1922				X
SGNesotes Allard, 1876				X
= Diastixus Allard, 1876
= Gyrinodes Fauvel, 1897 syn. nov.
Nipponohelops Masumoto, Ando & Akita, 2006				X
Physohelops Schuster, 1937				X
Raiboscelis Allard, 1876				X
= Hipponome Laporte, 1840
= Rhaebosceles Rye, 1878
Sabularius Escalera, 1914				X
Socotraphanes Nabozhenko, 2019					X
Stenohelops Reitter, 1922				X
SGHelopelius Reitter, 1922				X
= Stenomaleis Español, 1957
SGStenohelops Reitter, 1922				X
= Gunarellus Reitter, 1922
SG †Stenolassus Nabozhenko, Chigray & Bukejs, 2020
Tarpela Bates, 1870		X	X
= Lamperos Allard, 1876
Helopini incertae sedis
Dolphus Blanchard, 1847			X
Erulipothydemus Pic, 1918						X
Helopidesthes Fairmaire, 1895					X
Microcatomus Pic, 1925					X
Melanimonini Seidlitz, 1894 (1854)
Cheirodes Gené, 1839		X		X	X		X
SGAnemiadena Bouchard & Bousquet, subgen. nov.					X
SGCheirodes Gené, 1839		X		X	X		X
= Anemia Laporte, 1840
= Chirodes Agassiz, 1846
SGHistiaea Fairmaire, 1892				X	X
SGPseudanemia Wollaston, 1864				X	X
= Ammidanemia Reitter, 1904
SGSpinanemia Löbl, Bouchard, Merkl & Bousquet, 2020				X	X
SGTrichanemia Ardoin, 1971					X
Dolamara Reichardt, 1935				X
Melanimon Steven, 1828				X
= Microzoum Dejean, 1834
= Microzoon Hope, 1841
= Fundulus Gistel, 1848
Metaclisini Steiner, 2016
Metaclisa Jacquelin du Val, 1861		X	X	X		X
= Amarantha Motschulsky, 1860
= Tharsus LeConte, 1862
Palorini Matthews, 2003
Astalbus Fairmaire, 1900					X
Austropalorus Halstead, 1967							X
Eutermicola Lea, 1916							X
Palorinus Blair, 1930						X	X
Paloropsis Masumoto & Grimm, 2004						X
Palorus Mulsant, 1854		X	X	X	X	X	X	X
= Caenocorse C.G. Thomson, 1859
= Eba Pascoe, 1863
= Platyotus Gerstaecker, 1871
= Circomus Fleischer, 1900
= Stenopalorus Blair, 1930
Platycotylus Olliff, 1883					X	X	X
= Thurea Ferrer, 1998
Prolabrus Fairmaire, 1897					X
Pseudeba Blackburn, 1903							X
Ulomina Baudi di Selve, 1876		X	X	X	X	X	X	X
= Coelopalorus Blair, 1930
Ulomotypus Broun, 1886							X
†Vabole Alekseev & Nabozhenko, 2015
Paoligenini Ferrer, 2013
Paoligena Pic, 1928					X
Praeugenini De Moor, 1970
Anarmostodera Fairmaire, 1897					X
Dysgena Mäklin, 1863					X
Miltoprepes Gerstaecker, 1871					X
= Anephyctus Fairmaire, 1891
Nesogena Mäklin, 1863					X
SGArmigena Bouchard & Bousquet, subgen. nov.					X
SGHeterogena Froussart, 1961					X
SGNesogena Mäklin, 1863					X
= Bradygena Fairmaire, 1903
SGParagena Bouchard & Bousquet, subgen. nov.					X
Phaeostolus Fairmaire, 1884					X
Praeugena Laporte, 1840					X
= Adelphus Dejean, 1834
= Praogena Agassiz, 1846
= Lamprobothris Fairmaire, 1887
= Tactoderus Fairmaire, 1892
= Ergenna Fairmaire, 1897
Pseudopraeugena De Moor, 1970					X
Rhysopaussini Wasmann, 1896
Mimoxenotermes Pic, 1931						X
Rhysopaussus Wasmann, 1896						X
Rhyzodina Chevrolat, 1873					X
SGApistocerus Fairmaire, 1899					X
SGEurhysodina Wasmann, 1921					X
SGRhyzodina Chevrolat, 1873					X
= Rhysodina Wasmann, 1921
Xenotermes Wasmann, 1896						X
Scaurini Billberg, 1820
Carchares Pascoe, 1887					X
= Podoces Péringuey, 1886
Cephalostenus Solier, 1838				X
= Stenocephalus Agassiz, 1846
Herpiscius Solier, 1838					X
Scaurus Fabricius, 1775				X	X
= Scauris Rafinesque, 1815
Scotobiini Solier, 1838
Ammophorus Guérin-Méneville, 1831			X
= Selenomma Dejean, 1836
Diastoleus Solier, 1838			X
Emmallodera Blanchard, 1842			X
Leptynoderes Solier, 1838			X
Pumiliofossorum Silvestro & Giraldo-Mendoza, 2015			X
Scotobius Germar, 1823			X
= Gonogenius Solier, 1838
Tenebrionini Latreille, 1802
Ariarathus Fairmaire, 1891				X		X
= Teneatopus Reitter, 1920
Athrodactyla Klug, 1833					X
Bius Dejean, 1834		X		X
= Bia Hope, 1841
= Dendroscopius Gistel, 1848
Bouchardandrus Steiner, 2016		X
Bremerus Ferrer, 2004					X
Cedrosius Fairmaire, 1902					X
Falsocalcar Pic, 1925					X
Gridellia Kammerer, 2006					X
= Villiersia Gridelli, 1951
Hipalmus Bates, 1870			X
= Lobetas Motschulsky, 1872
Idiobates Casey, 1891		X
Microzophobas Pic, 1944			X
Neatus LeConte, 1862		X		X
Neozophobas Ferrer, 2006			X
Paratoxicum Champion, 1894							X
= Schizophthalmotribolium Kaszab, 1940
Phanerops Solier, 1851			X
Rhinandrus LeConte, 1866		X	X
= Exerestus Bates, 1870
= Proderops Fairmaire, 1873
Satanocalcar Pic, 1925					X
Tenebrio Linnaeus, 1758		X	X	X	X	X	X
SGAfrotenebrio Gridelli, 1951					X
SGCruracurvamtenebrio Robiche, 2019					X
SGMegatenebrio Gridelli, 1951					X
SGTenebrio Linnaeus, 1758		X	X	X	X	X	X
= Menedrio Motschulsky, 1872
= Tenebrionellus Crotch, 1874
Trichotenebrio Ardoin, 1962					X
Zophobas Dejean, 1834		X	X		X
SGMacrozophobas Pic, 1913		X	X
SGZophobas Dejean, 1834			X		X
= Pythonissus Gistel, 1834
Titaenini Fauvel, 1905
Artystona Bates, 1874							X
Callismilax Bates, 1874							X
Demtrius Broun, 1895							X
Partystona Watt, 1992							X
Titaena Erichson, 1842							X
Toxicini Oken, 1843
Dysantina Gebien, 1922
Calymmus Montrouzier, 1860							X
Cylindrosia Gebien, 1922					X
Diceroderes Solier, 1841			X
= Prosomenes Blanchard, 1845
Dysantes Pascoe, 1869					X	X
= Eudysantes Bouchard, Lawrence, Davies & Newton, 2005
Ilyxerus Pascoe, 1866							X
Mychestes Pascoe, 1870							X
Opostirus Kirsch, 1865			X
Orcopagia Pascoe, 1868							X
Ozolais Pascoe, 1866			X
Wattius Kaszab, 1982			X
Nycteropina Lacordaire, 1859
Chalcostylus Fairmaire, 1898					X
Macellocerus Solier, 1848					X
= Dolichoderus Klug, 1833
= Dillacerus Solier, 1835
= Stierlinius Forel, 1893
Nycteropus Klug, 1833					X
Toxicina Oken, 1843
Cryphaeus Klug, 1833				X	X	X	X
= Anthracias Dejean, 1834
= Ardelio Gistel, 1848
Epitoxicum Bates, 1873						X
Taiwanocryphaeus Masumoto, 1996						X
Toxicum Latreille, 1802				X		X	X
SGMutiloxicum Nabozhenko & Ivanov, 2018				X
SGToxicum Latreille, 1802				X		X	X
= Trestonia Rafinesque, 1815
Trachelostenini Lacordaire, 1859
Leaus Matthews & Lawrence, 1992							X
Myrmecodema Gebien, 1943			X
= Myrmecosoma Germain, 1855
Trachelostenus Solier, 1851			X
Triboliini Gistel, 1848
Aesymnus Champion, 1886			X
Hypogena Dejean, 1834		X	X
= Ulosonia Laporte, 1840
Latheticus C.O. Waterhouse, 1880		X	X	X	X	X	X
Lyphia Mulsant & Rey, 1859		X		X	X	X	X
= Lindia Blackburn, 1888
Metulosonia Bates, 1873			X
Mycotrogus Horn, 1870		X	X
Platybolium Blair, 1938						X
Spelaebiosis Bousquet & Bouchard, 2018			X
= Orghidania Ardoin, 1977
= Ardoinia Özdikmen, 2005
Tribolium W.S. MacLeay, 1825		X	X	X	X	X	X	X
SGAphanotus LeConte, 1862		X
SGTribolium W.S. MacLeay, 1825		X	X	X	X	X	X	X
= Stene Stephens, 1832
= Margus Dejean, 1834
= Tenebrioloma Gebien, 1910
= Leanum Uyttenboogaart, 1934
Xenogloeus Wollaston, 1861					X
Ulomini Blanchard, 1845
Achthosus Pascoe, 1863							X
Alegoria Laporte, 1840			X
= Hylonoma Macquart, 1850
Antimachus Gistel, 1829			X
= Ceratupis Perty, 1830
Apteruleda Gebien, 1928			X
Apteruloma Gebien, 1928			X
Basanopsis Gebien, 1914						X
Brachypophlaeus Fairmaire, 1897				X	X	X
= Leptoscapha Fairmaire, 1886
= Stenoscapha Fairmaire, 1885
Cenoscelis Wollaston, 1868				X	X	X	X
SGAptereutochia Kaszab, 1980						X
SGCenoscelis Wollaston, 1868				X	X	X	X
Cneocnemis Gebien, 1914				X		X	X
Curtopeltoides Pic, 1916						X
Donisiellus Bremer, 1992					X
Eutochia LeConte, 1862		X	X
= Aniara Melsheimer 1853:139
= Delopygus LeConte, 1866
= Aniarus Gemminger, 1870
= Holaniara Fairmaire, 1871
Macruloma Pic, 1921						X
Metabolocerus Bates, 1873			X
Microcenoscelis Schawaller, 2015					X
Neopsectropus Kaszab, 1941					X
Neooligocara Guerrero, Vidal & Moore, 2007			X
Oligocara Solier, 1848			X
Pheres Champion, 1886			X
Pycnuloma Fairmaire, 1896						X
Scotochares Boheman, 1858								X
Semieutochia Kaszab, 1980						X
Typhluloma Lea, 1912							X
Uleda Laporte, 1840			X
Uloma Dejean, 1821		X	X	X	X	X	X	X
SGApterulomoides Kaszab, 1982							X
SGUloma Dejean, 1821		X	X	X	X	X	X	X
= Prioscelida White, 1846
= Melasia Perroud & Mulsant, 1856
Ulomimus Bates, 1873				X		X
= Pseuduloma Fairmaire, 1893
Tenebrioninae incertae sedis
Anophthalmolamus Ferrer, 1993				X
Hangaya Matthews & Merkl, 2015							X
Penichrus Champion, 1885			X
Alleculinae Laporte, 1840
Alleculini Laporte, 1840
Alleculina Laporte, 1840
Aeanes Champion, 1893			X
Alethia Champion, 1888		X	X
Allecula Fabricius, 1801			X	X	X	X
SGAllecula Fabricius, 1801			X	X	X	X
SGAlleculina Pic, 1954					X
SGDietopsis Solier, 1835			X
Alogista Fåhraeus, 1870					X
Amaropsis Champion, 1893			X
Anognathena Ando, 2017						X
Anthracula Fairmaire, 1897						X
Atoichus Carter, 1915							X
Barbora Novák, 2020						X
Barycistela Blackburn, 1891							X
Blepusa Westwood, 1842			X
Bobina Novák, 2015				X		X
Bobisthes Novák, 2019						X
Bolbostetha Fairmaire, 1896						X
= Alleculodes Borchmann, 1925
Borbochara Novák, 2009						X
Borbonalia Novák, 2014				X		X
Borborella Novák, 2020						X
Borboresthes Fairmaire, 1897				X		X
Charisius Champion, 1888			X
= Narses Champion, 1888
Chitwania Novák, 2015						X
Cistelampra Fairmaire, 1897					X
Cisteloida Fairmaire, 1882						X
Cistelopsis Fairmaire, 1896				X		X
Cteisodella Novák, 2020						X
Cteisodes Borchmann, 1932						X
Dimorphochilus Borchmann, 1908							X
Diopoenus Champion, 1888			X
Doranalia Novák, 2020				X		X
Dorota Novák, 2018						X
Erzika Novák, 2020						X
Eucaliga Fairmaire & Germain, 1861			X
Euomma Boheman, 1858							X
= Apellatus Pascoe, 1863
Evaostetha Novák, 2008						X
Fifina Novák, 2018						X
Fifinoides Novák, 2020						X
Gerdacula Novák, 2015						X
Havanalia Novák, 2020				X
Hemicistela Blackburn, 1891							X
Homotrysis Pascoe, 1866							X
= Hybrenia Pascoe, 1866
Houaphanica Novák, 2020						X
Hymenalia Mulsant, 1856				X	X	X
Hymenorus Mulsant, 1852		X	X	X		X
Impressallecula Pic, 1951					X
Indricula Novák, 2016						X
Jaklia Novák, 2010						X
Jophon Champion, 1895							X
†Jurallecula L.N. Medvedev, 1969
Knausia Fall, 1931		X
Kombacula Novák, 2012						X
Ksukolcula Novák, 2017						X
Latacula Campbell, 1971			X
Lepturidea Fauvel, 1862							X
= Atractus Boisduval, 1835
= Aethyssius Pascoe, 1863
= Chromomaea Pascoe, 1866
= Alcmeonis Bates, 1868
= Anaxo Bates, 1868
= Licymnius Bates, 1868
= Neoatractus Borchmann, 1909
Liodocistela Pic, 1930						X
Litopous Matthews, 2012							X
Lobopoda Solier, 1835		X	X
SGFlavipoda Campbell, 1966			X
SGGlabrilobopoda Campbell, 1966			X
SGLobopoda Solier, 1835		X	X
SGMesolobopoda Campbell, 1966		X	X
SGMonoloba Solier, 1835			X
Loriculoides Novák, 2020						X
= Loricula Novák, 2016
Lycula Campbell, 1976			X
Macrocistelopsis Pic, 1956							X
Madreallecula Kanda, 2013		X
Magdanalia Novák, 2020				X
Makicula Novák, 2012						X
Matthewsotys Bouchard & Bousquet, gen. nov.							X
Menes Champion, 1888			X
Menoeceus Champion, 1888		X	X
Metistete Pascoe, 1866							X
= Lisa Haag-Rutenberg, 1879
Microcistelopsis Pic, 1922							X
	= Microcistela Pic, 1919
Microstenogena Pic, 1924					X
Microsthes Novák, 2011						X
Mimopraogena Pic, 1952					X
Mycetocharina Seidlitz, 1890				X
SGAlleculopsis Semenov, 1894				X
SGMycetocharina Seidlitz, 1890				X
= Caristela Fairmaire, 1894
Mycetocula Novák, 2015						X
Narsodes Campbell, 1976			X
Nesogenomorpha Pic, 1917					X
Neocistela Borchmann, 1909							X
= Pseudocistela Blackburn, 1891
Netopha Fairmaire, 1893				X		X
NikomenaliaDubrovina, 1975				X		X
Nocar Blackburn, 1891							X
Notacula Campbell, 1971			X
Notocistela Carter, 1915							X
Nypsius Champion, 1895							X
Obesacula Campbell, 1971			X
Ommatochara Borchmann, 1932						X
Ommatophorus W.J. MacLeay, 1872							X
Omocula Borchmann, 1937			X
Oocistela Borchmann, 1908							X
= Melaps Carter, 1908
Oracula Novák, 2019					X
SGDuocula Novák, 2019					X
SGOracula Novák, 2019					X
Orchesiolobopoda Pic, 1919			X
Palpichara Borchmann, 1932						X
Palpicula Novák, 2018						X
Parahymenorus Campbell, 1971			X
Pemanoa Buck, 1955							X
Petrostetha Novák, 2008						X
Phediodes Campbell, 1976			X
Phedius Champion, 1888		X	X
Pitholaus Champion, 1888			X
Pizura Novák, 2016						X
Platyallecula Blair, 1935					X
Polyidus Champion, 1888			X
Potocula Novák, 2012						X
Prionalia Novák, 2020				X
Prionychus Solier, 1835				X
= Eryx Stephens, 1832
= Pelops Gistel, 1834
Pseudocistelopsis Novák, 2018						X
Psis Novák, 2019						X
Punctacula Campbell, 1971			X
Scaletomerus Blackburn, 1891							X
= Otys Champion, 1895
Scaphinion Matthews, 2012							X
Simarus Borchmann, 1909							X
= Ismarus Haag-Rutenberg, 1878
Socotralia Novák, 2007					X
Spinecula Novák, 2019						X
Stenochidus LeConte, 1862		X
Stenogena Fairmaire, 1895					X
Stilbocistela Borchmann, 1932						X
Synallecula Kolbe, 1883					X
Tanychilus Newman, 1838							X
Taxes Champion, 1895							X
Telesicles Champion, 1888		X
Temnes Champion, 1888			X
Theatetes Champion, 1888			X
Upinella Mulsant, 1857				X		X
SGThornella Novák, 2019						X
SGTibinella Novák, 2019				X		X
SGUpinella Mulsant, 1856				X		X
Vietnalia Novák, 2021						X
Zizu Novák, 2019						X
Gonoderina Seidlitz, 1896
Andrimus Casey, 1891		X
Androchirus LeConte, 1862		X
AsiomiraDubrovina, 1973				X
= Kirgisomira Weise, 1974
Brachycula Fairmaire, 1906					X
†Calcarocistela Nabozhenko, 2016
Capnochroa LeConte, 1862		X
Chromatia LeConte, 1862		X
Copistethus Seidlitz, 1890				X
Cornucistela Campbell, 1980				X
Cryptomysia Pic, 1954					X
Eubalia Laporte, 1840					X
= Plesia Klug, 1833
= Cacoplesia Fairmaire, 1898
Gerandryus Rottenberg, 1873				X
= Parablops Rottenberg, 1871
Gonodera Mulsant, 1856				X
Helopsisomira Pic, 1952					X
Isomira Mulsant, 1856		X	X	X	X	X
SGApteromira Weise, 1974				X
SGDanielomira Weise, 1974				X
SGHeteromira Hölzel, 1958				X
SGIsomira Mulsant, 1856		X	X	X		X
= Tedinus Casey, 1891
SGMucheimira Novák, 2016				X		X
SGParaisomiraDubrovina, 1982				X
SGPubeirosoma Pic, 1954					X
Kralia Novák, 2013				X
Malaymira Novák, 2020						X
Micrisomira Pic, 1930						X
Microcistela Pic, 1904				X
Onychomira Campbell, 1984		X
Paracistela Borchmann, 1941				X		X
Piccula Bousquet & Bouchard, 2015					X
= Gerardia Pic, 1954
Pseudocistela Crotch, 1874		X	X	X	X	X	X	X
Pseudohymenalia Novák, 2008				X		X
Viriathus Fairmaire, 1902					X
Mycetocharina Gistel, 1848
Caulostena Fairmaire, 1896					X
Cylindrothorus Solier, 1843					X
SGCylindrothorus Solier, 1843					X
= Othelecta Pascoe, 1866
SGMicrothelecta Pic, 1952					X
Hymenochara Campbell, 1978		X
Labetis C.O. Waterhouse, 1879								X
Mycetochara Guérin-Méneville, 1827		X		X
SGErnocharis C.G. Thomson, 1859		X		X
= Stigmatoma LeConte, 1862
SGMycetochara Guérin-Méneville, 1827		X		X
= Mycetophila Gyllenhal, 1810
= Mycetocharis Gyllenhal, 1827
= Mycetochares Latreille, 1829
= Bolithophilus Gistel, 1832
SGOculochara Novák, 2020				X
SGPterna Reitter, 1884				X
†Mycetocharoides Schaufuss, 1889
Xystropodina Solier, 1835
Anamphidora Casey, 1924		X
Cteisa Solier, 1835			X
Dasytoxystropus Pic, 1921			X
Erxias Champion, 1888			X
Lystronychus Latreille, 1829		X	X
SGLystronychus Latreille, 1829		X	X
SGXystronia Solier, 1835			X
Microprostenus Pic, 1921			X
Prostenus Klug, 1829			X
= Mecocerus Solier, 1835
Scotobiopsis Brèthes, 1910			X
Tucumana Gebien, 1911			X
= Eustenia Fairmaire, 1905
Xystropus Solier, 1835			X
Alleculini incertae sedis
Omedes Broun, 1893							X
Xylochus Broun, 1880							X
Zomedes Watt, 1992							X
Cteniopodini Solier, 1835
†Amberophlus Novák & Háva, 2019
Balassogloa Semenov, 1891				X
Cistelina Seidlitz, 1896				X		X
Cistelomorpha Redtenbacher, 1868				X		X
Cnecosochara Reitter, 1913				X
Cteniopinus Seidlitz, 1896				X		X
SGCteniopinus Seidlitz, 1896				X		X
SGLechinius Blair, 1922						X
= Lechinius Borchmann, 1930
Cteniopus Solier, 1835				X
SGCtenioposomus Reitter, 1906				X
SGCteniopus Solier, 1835				X
= Cistela Fabricius, 1775
= Cistella Gistel, 1848
= Telacis Poey, 1854
= Sarandonyx Gozis, 1881
SGRhinobarus Reitter, 1906				X
Diastanus Fairmaire, 1902					X
Falsomophlus Pic, 1925						X
Gastrhaema Jacquelin du Val, 1863				X
Heliomophlus Reitter, 1906				X
Heliostrhaema Reitter, 1890				X
Heliotaurus Mulsant, 1856				X	X
SGAtlasotaurus Bouyon, 2011				X
SGHeliotaurus Mulsant, 1856				X	X
SGJulogenius Reitter, 1906				X
Holdhausia Reitter, 1906				X
Hypocistela Bates, 1879				X
Megischia Solier, 1835				X
Megischina Reitter, 1906				X
Nesotaurus Fairmaire, 1896					X
Omophlina Reitter, 1890				X
Omophlus Dejean, 1834				X
SGEuomophlus Iablokoff-Khnzorian, 1983				X
SGMicromophlus Znojko, 1950				X
SGOdontomophlus Seidlitz, 1896				X
= Pleuromophlus Reitter, 1906
SGOmophlus Dejean, 1834				X
SGPaurodontomophlus Muche, 1979				X
SGPhibalus Gistel, 1856				X
= Paromophlus Iablokoff-Khnzorian, 1983
Petria Semenov, 1894				X
†Platycteniopus Chang, Nabozhenko, Pu, Xu, Jia, Li, 2016
Podonta Solier, 1835				X
= Pododonta Agassiz, 1846
Podontinus Seidlitz, 1896				X
Proctenius Reitter, 1890				X
†Sinocistela Zhang, 1989
Stenerophlina Reitter, 1906				X
Steneryx Reitter, 1890				X
Tripolicryptus Strand, 1929				X
= Brachycryptus Quedenfeldt, 1891
Alleculinae incertae sedis
Alogistopsis Borchmann, 1943					X
Amorphopoda Fåhraeus, 1870					X
Apalmia Fairmaire, 1896						X
Aptericula Borchmann, 1937					X
Asticostena Fairmaire, 1897						X
Bancocistela Pic, 1947					X
Bearnicistela Pic, 1909						X
Borchmannius Bousquet & Bouchard, 2015					X
= Glyptothorax Borchmann, 1937
Borneocistela Pic, 1922						X
Bratyna Westwood, 1875					X
= Morocaulus Fairmaire, 1899
Buxela Fairmaire, 1894						X
Cistelodema Borchmann, 1932						X
Compsocula Fairmaire, 1898					X
Costallecula Pic, 1954					X
Dioxycula Fairmaire, 1896						X
Ectatocera Fåhraeus, 1870					X
Ectenostoma Fåhraeus, 1870					X
Eutrapelodes Borchmann, 1929					X
Falsopsilonycha Pic, 1930					X
Flabellalogista Pic, 1954					X
Helopsallecula Pic, 1936					X
Homoropsis Fairmaire, 1886					X
Hovacula Fairmaire, 1898					X
Idatius Fairmaire, 1906					X
Isomiropsis Borchmann, 1942					X
Lagriallecula Pic, 1920					X
Macrocistela Pic, 1941					X
Mayidicistela Pic, 1954					X
Microamarygmus Pic, 1915						X
Mimocistela Borchmann, 1938					X
Omolepta Fåhraeus, 1870					X
Pseudomorocaulus Pic, 1915					X
Rhipidonyx Reitter, 1876						X
Seydelicistela Pic, 1954					X
Stenerula Fairmaire, 1875					X
Stenogenomorpha Pic, 1919					X
Strongyallecula Pic, 1955					X
Diaperinae Latreille, 1802
Crypticini Brullé, 1832
Apteroseriscius Koch, 1950					X
Araeopselaphus Gebien, 1921					X
Capicrypticus Koch, 1950					X
Cechenosternum Gebien, 1921				X	X	X
Crypticus Latreille, 1816				X
SGCrypticopsis Antoine, 1945				X
SGCrypticus Latreille, 1816				X
SGPlatycrypticus Español, 1952				X
= Ulomoides Escalera, 1927
SGSeriscius Motschulsky, 1845				X
Ellipsodes Wollaston, 1854			X	X	X	X	X	X
SGAnthrenopsis Koch, 1950			X	X	X	X	X	X
SGEllipsodes Wollaston, 1854				X
Gondwanocrypticus Español, 1955		X	X
Lamprocrypticus Español, 1950				X
Lineocrypticus Koch, 1950					X
Microcrypticus Gebien, 1921					X	X
SGCrypticocatops Kaszab, 1975						X
SGMicrocrypticus Gebien, 1921					X
Myrmecocatops Wasmann, 1897					X
Oochrotus P.H. Lucas, 1852				X
Poecilocrypticus Gebien, 1928		X	X
Pseudoseriscius Español, 1950				X	X
SGAustraloseriscius Koch, 1950					X
SGPseudoseriscius Español, 1950				X
Diaperini Latreille, 1802
Adelinina LeConte, 1862
Adelina Dejean, 1835		X	X	X		X	X
= Doliema Pascoe, 1860
= Schedarosus Reitter, 1876
Alphitophagus Stephens, 1832		X		X		X	X
= Phyletes Redtenbacher, 1845
Arabcynaeus Schawaller, 2009				X
Cynaeus LeConte, 1862		X		X		X
Doliodesmus Spilman, 1967		X
Doliopines Horn, 1894		X
Gnatocerus Thunberg, 1814		X	X	X	X	X	X
SGEchocerus Horn, 1870		X	X	X	X	X
SGGnatocerus Thunberg, 1814		X	X	X	X	X	X
= Cerandria Dejean, 1834
= Gnathocerus Agassiz, 1846
= Sicinus Champion, 1886
Iccius Champion, 1886		X	X
Mophis Champion, 1886			X
Neoplateia Marcuzzi, 1986			X
Palembomimus Matthews & Lawrence, 2005							X
Sitophagus Mulsant, 1854		X	X
Diaperina Latreille, 1802
Basides Motschulsky, 1873				X		X	X
= Ischnodactylus Chevrolat, 1877
Ceropria Laporte & Brullé, 1831			X	X	X	X	X
= Epilampus Dejean, 1834
= Epilamprus Gistel, 1848
= Dictysomorphus Pic, 1921
Cissides Chatanay, 1915					X
Coelopleurum Gebien, 1921					X
Cosmonota Blanchard, 1842			X
Cyclobiomorphus Pic, 1916						X
Cyclobium Pic, 1916						X
Diaperis Geoffroy, 1762		X	X	X		X
= Allophasia Pascoe, 1871
Espagnolina Kaszab, 1965						X
Exapinaeus Pascoe, 1882			X
Falsocosmonota Kaszab, 1962						X
Gargilius Fairmaire, 1891					X
Gressittiola Kaszab, 1955							X
Heterophylus Klug, 1833					X
= Heterophyllus Gemminger, 1870
Hoplaspis Motschulsky, 1858						X
Lelegeis Champion, 1886			X
Liodema Horn, 1870		X	X
Louwerensia Kaszab, 1964							X
Loxostethus Triplehorn, 1962			X
Neomida Latreille, 1829		X	X	X		X
= Oplocephala Laporte & Brullé, 1831
= Arrhenoplita W. Kirby, 1837
= Hoplocephala Agassiz, 1846
= Evoplus LeConte, 1866
Paniasis Champion, 1886			X
= Pseudapsida Kulzer, 1961
Pentaphyllus Dejean, 1821		X	X	X	X	X	X	X
= Iphicorynus Jacquelin du Val, 1861
Phayllus Champion, 1886			X
Platydema Laporte & Brullé, 1831		X	X	X	X	X	X	X
= Typhobia Pascoe, 1869
= Histeropsis Chevrolat, 1878
= Anisocara Gebien, 1925
Platydemoides Kaszab, 1980						X
Pseudobasides Pic, 1916						X
Saptine Champion, 1886			X
Sciophagus Sharp, 1885							X	X
= Pachycerus Montrouzier, 1860
Stenoscapha Bates, 1873			X
Stomylus Fåhraeus, 1870					X
= Pselaphidion Gebien, 1921
Ulomoides Blackburn, 1888			X	X		X	X	X
= Palembus Casey, 1891
= Martianus Fairmaire, 1893
= Tenebriomimus Kolbe, 1901
= Phayllidius Gebien, 1922
Yamatotakeru Ando, 2015				X		X
Diaperini incertae sedis
Pelleas Bates, 1872				X
Ectychini Doyen, Matthews & Lawrence, 1990
Ectyche Pascoe, 1869							X
Micrectyche Bates, 1873							X
Gnathidiini Gebien, 1921
Anopidiina Jeannel & Paulian, 1945
Anopidium Jeannel & Paulian, 1945					X
Caecophloeus Dajoz, 1972			X
Cryptozoon Schaufuss, 1882			X
Mauritianopidium Dajoz, 1977					X
Menimopsis Champion, 1896			X
= Caecomenimopsis Kaszab, 1970
Nanocaecus Schawaller & Purchart, 2012					X
Neanopidium Dajoz, 1975			X
Paralyreus Grouvelle, 1918					X
Paranopidium Dajoz, 1974					X
Peyrierasia Dajoz, 1975					X
Prototyrtaeus Spiessberger & Ivie, 2020			X
Pseudanopidium Dajoz, 1974					X
Sphaerognathium Dajoz, 1975			X
Tyrtaeus Champion, 1913			X		X		X
Gnathidiina Gebien, 1921
Anommabates Koch, 1956					X
Caecochares Koch, 1956					X
Enanea Lewis, 1894				X		X
Gnathidium Gebien, 1921					X
Menimus Sharp, 1876				X		X	X	X
SGMenimus Sharp, 1876				X		X	X	X
= Ceramba Fauvel, 1904
= Paita Fauvel, 1904
= Microcilibe Carter, 1919
= Tjikoraia Pic, 1921
= Neomenimus Kaszab, 1939
SGSinomenimus G.S. Medvedev, 2007						X
Micropeneta Pic, 1921						X
= Menimoides Kaszab, 1946
Pseudoenanea Pic, 1924						X
Sakaiomenimus Ando, 2003				X
Szentivanya Kaszab, 1958							X
Taiwanomenimus Masumoto, Akita & Lee, 2019						X
Gnathidiini incertae sedis
Betschia Dajoz, 1980					X
Mireanopidium Dajoz, 1977					X
Hyociini G.S. Medvedev & Lawrence, 1982
Brittonina G.S. Medvedev & Lawrence, 1986
Brittona G.S. Medvedev & Lawrence, 1986							X
Magela G.S. Medvedev & Lawrence, 1986							X
Hyociina G.S. Medvedev & Lawrence, 1982
Csiro G.S. Medvedev & Lawrence, 1984							X
SGCsiro G.S. Medvedev & Lawrence, 1984							X
SGMillstreamia G.S. Medvedev & Lawrence, 1984							X
Hyocis Pascoe, 1866							X
SGHyocis Pascoe, 1866							X
SGNannohyocis G.S. Medvedev & Lawrence, 1983							X
SGNeohyocis G.S. Medvedev & Lawrence, 1983							X
Parahyocis Kaszab, 1955							X	X
Uptonina G.S. Medvedev & Lawrence, 1986
Uptona G.S. Medvedev & Lawrence, 1986							X
Hypophlaeini Billberg, 1820
Cheilopoma Murray, 1867					X
Corticeus Piller & Mitterpacher, 1783		X	X	X	X	X	X	X
SG Alienophloeus Bremer, 2018			X
SGCorticeus Piller & Mitterpacher, 1783		X	X	X	X	X		X
= Hypophlaeus Fabricius, 1790
= Paraphloeus Seidlitz, 1894
= Syncolydium Kolbe, 1897
SGMetacorticeus Bremer & Lillig, 2017						X	X
SGNeglectophloeus Bremer & Lillig, 2017			X			X	X	X
SGPogonophloeus Bremer, 1998		X		X	X	X
SGSeorsophloeus Bremer, 1998					X	X
SGStenophloeus Blair, 1921			X		X	X	X
= Cnemophloeus Bremer, 1998
SGTylophloeus Bremer, 1998		X	X	X		X
Ischnarthron Gebien, 1921					X
Myonophloeus Bremer & Lillig, 2017			X
Pogonoxenus Wasmann, 1899					X
Typhlophloeus Jeannel & Paulian, 1945				X	X
Leiochrinini Lewis, 1894
Ades Guérin-Méneville, 1857				X	X	X	X	X
= Leiochrodes Westwood, 1883
Crypsis C.O. Waterhouse, 1877				X		X
= Leiochrotina Westwood, 1883
Derispia Lewis, 1894				X		X	X
Derispiella Kaszab, 1961						X
Derispiola Kaszab, 1946						X
Leiochrinus Westwood, 1883				X		X	X
Leiochrodinus Kaszab, 1961						X
Leiochrodontes Kaszab, 1946						X
Leiochrota Westwood, 1883						X
Pimplema Pascoe, 1887						X	X
= Hades J. Thomson, 1860
Stethotrypes Gebien, 1914						X
= Leichrodomorphus Pic, 1921
Myrmechixenini Jacquelin du Val, 1858
Myrmechixenus Chevrolat, 1835		X		X	X	X	X
= Myrmecoxenus Märkel, 1844
= Myrmechoxenus Aubé, 1850
Phaleriini Blanchard, 1845
Emypsara Pascoe, 1866				X
= Callicomus Motschulsky, 1860
= Obenbergeria Strand, 1929
Halammobia Semenov, 1901				X
Pachyphaleria Gebien, 1920					X
Paranemia Heyden, 1892				X
= Taklamakania Ferrer & Yvinec, 2004
= Taclamacanius Ferrer & Yvinec, 2005
Phaleria Latreille, 1802		X	X	X	X	X	X
SGEpiphaleria Lewis, 1894				X	X	X	X
SGEremophaleria Español, 1951				X
SGNeophaleria Español, 1963					X
SGPhaleria Latreille, 1802		X	X	X			X
= Uria Gistel, 1848
= Sepedonastes Gistel, 1856
= Halophalerus Crotch, 1874
= Phalerisida Kulzer, 1959
= Atahualpina Español, 1960
Phaleromela Reitter, 1916		X		X
Phtora Germar, 1836				X	X
SGClypeophtora F. Soldati & L. Soldati, 2003				X
SGPhtora Germar, 1836				X	X
= Cataphronetis P.H. Lucas, 1846
= Pseudostene Wollaston, 1861
= Phthora Gemminger, 1870
Scaphidemini Reitter, 1922
Basanus Lacordaire, 1859				X		X
Laoscapha Schawaller, 2016						X
†Palaeobasanus Nabozhenko & Kirejtshuk, 2020
Pseudoscaphidema Pic, 1926				X
Scaphidema Redtenbacher, 1848		X		X		X
= Nelites LeConte, 1850
= Microbasanus Pic, 1921
Spiloscapha Bates, 1873				X		X	X
Trachyscelini Blanchard, 1845
Macrotrachyscelis Pic, 1925					X
Taiwanotrachyscelis Masumoto, Akita & Lee, 2012						X
Trachyscelis Latreille, 1809		X	X	X	X	X	X
Diaperinae incertae sedis
Triplehornia Matthews & Lawrence, 2005							X
Stenochiinae W. Kirby, 1837
Cnodalonini Oken, 1843
Acanthobas Gebien, 1928			X
Acanthocamaria Gebien, 1919			X
Achariotheca Kaszab, 1970							X
Achrostus Fairmaire, 1891					X
Actanorie Bates, 1879					X
= Callimaria Fairmaire, 1888
Aesthetus C.O. Waterhouse, 1890			X
Agymnonyx Gebien, 1921							X
Ahexaroptrum Kaszab, 1960						X
Ainu Lewis, 1894				X		X
SGAinu Lewis, 1894				X		X
SGErulipus Fairmaire, 1903						X
Alcyonotus Pascoe, 1882					X
= Adonicus Fairmaire, 1891
= Sterces Champion, 1891
Alobates Motschulsky, 1872		X
Amarsenes Bates, 1879					X
Amenophis J. Thomson, 1858					X
SGAmenophis J. Thomson, 1858					X
= Hemerobates Kolbe, 1884
= Praostetha Fairmaire, 1897
= Pseudamenophis Pic, 1916
SGDeriles Motschulsky, 1872					X
= Anadischidus Kolbe, 1897
Anachayus Bouchard & Bousquet, nom. nov.					X
= Chatanayus Ardoin, 1957
Andocamaria Masumoto, 1993						X
Androsus Gebien, 1921						X	X
Anisophaedis Ando, 1993						X
Annamosdara Kaszab, 1941						X
Apsida Lacordaire, 1859		X	X
= Hapsida Gemminger, 1870
Aptereucyrtus Gebien, 1922						X
Apterobrachys Kaszab, 1986							X
Apteromaia Kulzer, 1952						X
Apteromerus Blair, 1928							X	X
Apterophenus Gebien, 1921							X	X
Apterotheca Gebien, 1921							X
= Austropeus Carter, 1924
= Caxtonana Buck, 1960
Argobrachium Fairmaire, 1899					X
= Aphelus Gebien, 1921
Argutiolana Robiche, 2001					X
= Wahlbergylium Ferrer, 2011
Artactes Pascoe, 1868						X
= Macroartactes Pic, 1924
Asbolodes Fairmaire, 1892						X
Asbolodomimus Pic, 1921						X
= Allopezus Gebien, 1922
Asididius Fairmaire, 1869					X
Asidobothris Fairmaire, 1886					X
Asopidiopsis Kaszab, 1955								X
Asopis Haag-Rutenberg, 1878								X
Astathmetus Bates, 1874			X
Augolesthus Motschulsky, 1872						X
= Chrysomaia Kulzer, 1952
Baratus Fairmaire, 1897						X
Becvarius Masumoto, 1998						X
Biroum Kaszab, 1956						X
Blapida Perty, 1830			X
= Rhyssochiton Gray, 1831
= Metonites Gistel, 1848
Borneocamaria Pic, 1917						X
= Homoeogenus C.O. Waterhouse, 1882
= Homoeocamaria Blair, 1919
= Krollus R. Lucas, 1920
Borneosphaerotus Grimm, 2015						X
Borneosphena Purchart & Grimm, 2016						X
Borneosynopticus Grimm, 2015						X
Bothynocara Gebien, 1928			X
Bothynocephalus Doyen, 1988			X
Bradymerus Perroud & Montrouzier, 1865				X	X	X	X	X
= Isostira Pascoe, 1870
= Bradynocerus Fairmaire, 1883
= Pseudobradymerus Pic, 1926
= Planibates Kaszab, 1939
Bradysphaerotus Kaszab, 1986							X
Brasilius Gebien, 1928			X
Brosimapsida Ferrer & Ødegaard, 2005			X
Byzacnus Pascoe, 1866					X
Calabosca Fairmaire, 1894						X
= Ascalabus Fairmaire, 1893
Calydonella Doyen, 1995			X
Calydoniomorpha Pic, 1917			X
Calydonis Pascoe, 1882			X
Camaria Lepeletier & Audinet-Serville, 1828			X
= Truncatocamaria Pic, 1922
Camarimena Motschulsky, 1863						X
= Sinopium Pascoe, 1866
= Espitomorphus Pic, 1921
Camariocropterum Pic, 1920			X
Camariodes Fairmaire, 1869					X
= Tinophyllus Fairmaire, 1869
Camariomorpha Pic, 1915						X
= Methistamena Gebien, 1919
Campolene Pascoe, 1863							X
Campsia Lepeletier & Audinet-Serville, 1828			X
SGBlapidocampsia Pic, 1919			X
SGCampsia Lepeletier & Audinet-Serville, 1828			X
= Celox Gistel, 1848
Campsiomorpha Pic, 1917				X		X
Camptobrachys Kaszab, 1941						X
Carabelops Fairmaire, 1899					X
= Agraecus Fairmaire, 1900
Caracasa Pic, 1921			X
Cataphanus Gebien, 1921							X
Catapiestus Perty, 1831						X
= Plateia Laporte, 1840
Celebesa Pic, 1921							X
Cephalothydemus Pic, 1923						X
Cerandrosus Gebien, 1921							X
Cerocamptus Gebien, 1919						X
Chaetopsia Gebien, 1925						X
Chalcocyclus Fairmaire, 1884					X
Chalcopauliana Ardoin, 1961					X
Charianus Bates, 1879					X
Chariotheca Pascoe, 1860							X	X
= Chariothes Carter, 1914
Chemolanus Bates, 1879					X
Chlorocamma Bates, 1873							X
Choastes Champion, 1893			X
= Choaspes Champion, 1885
Chrysopeplus Gebien, 1942							X
= Leiopeplus Broun, 1893
Cibdelis Mannerheim, 1843		X
= Scotera Motschulsky, 1845
Cleomis Fairmaire, 1892						X
= Pseudeucyrtus Pic, 1916
= Oyanus Pic, 1921
Cnephalura Doyen, 1988			X
Cnodalon Latreille, 1797			X
= Cnodalum Agassiz, 1846
Coelocnemis Mannerheim, 1843		X
Coelometopus Solier, 1848				X
Cophodema Gebien, 1943			X
= Cophosoma Gebien, 1928
Cryptobates Fairmaire, 1882						X
Cryptobatoides Kaszab, 1941						X
Cryptobrachys Kaszab, 1941						X
Cryptostenophanes Kaszab, 1941						X
Csikiola Kaszab, 1955								X
Cuemus Bouchard, 2000							X
Cybopiestes Reitter, 1917				X
Cyclonesus Fairmaire, 1896						X
Cyrtosoma Perty, 1830			X
Cyrtotyche Pascoe, 1866					X
Cyrtotyctus Kolbe, 1897					X
Damatris Laporte, 1840					X
= Hybonotus Dejean, 1834
= Malacova Fairmaire, 1898
Danodema Gebien, 1925					X
Dechiustes Blair, 1940								X
Dentatoploedipus Kaszab, 1984						X
Deplanchesia Fauvel, 1860			X
Derosphaerus J. Thomson, 1858				X	X	X	X
= Euphron Dejean, 1834
= Encyalesthus Motschulsky, 1860
= Pachyurgus LeConte, 1862
= Cholipus Pascoe, 1866
= Notiolesthus Motschulsky, 1872
= Neandrosus Pic, 1921
= Falsoencyalesthus Pic, 1923
Diachoriops Ando, 2020				X		X	X	X
= Schizomma Gebien, 1921
Dinomus Brême, 1842			X
Diopethes Pascoe, 1882			X
Dioscoridemus Koch, 1970					X
Drocleana Bates, 1879					X
Eccoptostoma Gebien, 1913					X
= Ogoueum Pic, 1923
Ectomopsis Fairmaire, 1905			X
Elomosda Bates, 1870			X
Epicalla Lacordaire, 1859			X
Episopus Bates, 1873							X
Eremobatodes Gebien, 1943					X
= Eremobates Gebien, 1921
Espites Pascoe, 1882							X
Eucyrtus Lacordaire, 1859						X
= Microeucyrtus Pic, 1926
Euhemicera Ando, 1996						X
Euphloeus Pascoe, 1887						X
Eutelonodolinus Robiche, 2007					X
Eutelonotus Fairmaire, 1902					X
= Eutelus Solier, 1843
= Nodotelus Koch, 1950
Euthysternum Chatanay, 1915					X
Exocolena Gebien, 1914						X
Falsandrosus Kaszab, 1980						X
Falsobates Kaszab, 1941						X
Falsobrachys Kulzer, 1954						X
Falsocamaria Pic, 1917				X		X
= Eucamaria Gebien, 1919
Falsocamariodes Ardoin, 1956					X
Falsodiopethes Pic, 1924			X
Falsonannocerus Pic, 1947					X
Falsoperichilus Pic, 1920					X
Falsozotypus Kaszab, 1980						X
Foochounus Pic, 1921						X
= Chariophenus Blair, 1929
= Anobriomaia Kaszab, 1941
= Microcameria Ren, 1998
Gaurobates Gebien, 1928			X
Gauromaia Pascoe, 1866						X
SGFalsogauromaia Pic, 1921						X
SGGauromaia Pascoe, 1866						X
= Cephaleucyrtus Pic, 1923
Gebienella Kaszab, 1941						X
Gebienocamaria Masumoto, 1993						X
Gigantopigeus Kaszab, 1984						X
Girardocamaria Masumoto, 1993						X
Glyptotus LeConte, 1858		X	X
Gnesis Pascoe, 1866						X
= Tromosternus Harold, 1876
Gonespites Gebien, 1921							X
Gonospa Champion, 1886			X
Graptopezus Gebien, 1921							X
Haplandrus LeConte, 1862		X
Haporema Fairmaire, 1892					X
Hegemona Laporte, 1840			X
= Eucamptus Germar, 1842
= Eusarca Chevrolat, 1845
Heliofugus Guérin-Méneville, 1831			X
SGCollariheliofugus Freude, 1960			X
SGHeliofugus Guérin-Méneville, 1831			X
= Heliosteres Hope, 1841
= Heliophygus Agassiz, 1846
= Euschatia Solier, 1851
SGInscutoheliofugus Freude, 1960			X
SGRugosiheliofugus Freude, 1960			X
Hemicera Laporte & Brullé, 1831						X
SGHemicera Laporte & Brullé, 1831						X
= Chrysolinoides Jolivet, 1951
SGNanohemicera Pic, 1923						X
Hemimmedia Gebien, 1928			X
Hesiodus Champion, 1885			X
Hexarhopalus Fairmaire, 1891				X		X
SGHexarhopalus Fairmaire, 1891				X		X
= Hexarhoptrum Fairmaire, 1894
= Laosocryptobates Pic, 1928
= Apteroleprocaulus Kaszab, 1983
SGLeprocaulus Fairmaire, 1896						X
= Pseudocaelophus Pic, 1922
= Pseudoderosphaerus Pic, 1922
Hicetaon Champion, 1885			X
Holobrachys Fairmaire, 1869					X
Hoploedipinus Kaszab, 1984						X
Hoploedipus Fairmaire, 1898						X
Hyboproctus Kolbe, 1897					X
Hydissus Pascoe, 1869							X
= Hydisus Scudder, 1882
Hypaulax Bates, 1868							X
= Chileone Bates, 1868
Hypocalis Dejean, 1834					X
Hypovinsonia Ardoin, 1961					X
Ilus Champion, 1885			X
Immedia Pascoe, 1882			X
Iphthiminus Spilman, 1973		X		X
Iphthimulus Reitter, 1920				X
Irianobates Kaszab, 1986							X
Isaminas Champion, 1887			X
= Pteroglymmius Gebien, 1928
Isicerdes Champion, 1885			X
Isopus Montrouzier, 1860							X
Kabakoviella Kaszab, 1980						X
Kaszaba Matthews & Doyen, 1989							X
Lenkous Kaszab, 1973			X
Lepidocaulinus Schawaller, Masumoto & Merkl, 2013						X
Leprocaulinus Kaszab, 1982						X
= Pigeocaulinus Kaszab, 1984
Lomocnemis Gebien, 1921							X
Lordodera Gebien, 1921					X
Lycidioides Ando, 2003						X
Macropachylesthus Pic, 1923						X
Macrostethus Wollaston, 1854				X
Mahena Gebien, 1922					X
Malayaplamius Masumoto, 1986						X
Malaysphena Bečvář & Purchart, 2008						X
Maracia Gebien, 1919			X
Mariepskopia Schawaller, 2012					X
Mechanetes C.O. Waterhouse, 1887						X
= Diabolicobates Pic, 1930
Melobrachys Kaszab, 1960						X
Menandris Haag-Rutenberg, 1878								X
Menephilus Mulsant, 1854				X		X		X
Merinus LeConte, 1862		X
Metisopus Bates, 1873							X	X
Micreuphlaeus Fairmaire, 1897						X
Microbradymerus Schawaller, 1999						X
Micromenandris Kaszab, 1955								X
Microphenus Gebien, 1921							X
Microsphaerotus Pic, 1928						X
Misolampidius Solsky, 1875				X		X
= Ptilonix Allard, 1877
Misolampomorphus Kaszab, 1941						X
Misolampus Latreille, 1806				X
Mitys Champion, 1885			X
Moeon Champion, 1886			X
Mophon Champion, 1886			X
Moromelas Fairmaire, 1898					X
Morphostenophanes Pic, 1925				X		X
= Promorphostenophanes Kaszab, 1960
Mrazius Pic, 1925			X
Mylaris Pallas, 1781			X
= Iphthinus Dejean, 1834
= Cecrops Gistel, 1834
= Nyctobates Guérin-Méneville, 1834
= Iphthimus Gemminger, 1870
Nannalcyon Koch, 1950					X
= Nannocerus Fairmaire, 1887
Necrobioides Fairmaire, 1882						X
Neoplamius Löbl, Bouchard, Merkl & Bousquet, 2020						X
Neoporphyrhyba Ardoin, 1956					X
Neotheca Carter, 1930							X
Nesocyrtosoma Marcuzzi, 1976			X
= †Hesiodobates Kaszab & Schawaller, 1984
= Pachycyrtosoma Marcuzzi, 1999
= Serrania Garrido, 2003
Nesosphaerotus Gebien, 1921					X
= Nesophaerotus Ardoin, 1962
Nuptis Motschulsky, 1872			X
Oeatus Champion, 1885			X
Oectosis Pascoe, 1869							X
Oedemutes Pascoe, 1860						X
SGOedemutes Pascoe, 1860						X
SGTamdaous Pic, 1923						X
Oenopion Champion, 1885		X	X
Omolipus Pascoe, 1860							X
Osdara Walker, 1858						X	X
SGOsdara Walker, 1858						X
SGSpinosdara Bouchard & Bousquet, subgen. nov.						X	X
Osdaroides Kaszab, 1980						X
Osternus Fairmaire, 1895					X
SGMicrocalydonis Pic, 1923					X
SGOsternus Fairmaire, 1895					X
Othryoneus Champion, 1886			X
Oxidates Champion, 1886			X
Ozaenimorphus Fairmaire, 1882					X
Ozotypus Pascoe, 1862						X
Pachylesthus Fairmaire, 1897						X
Papuamisolampus Kaszab, 1986							X
Paramisolampidius Merkl & Masumoto, 2020				X		X
Parimmedia Gebien, 1928			X
Paroeatus Gebien, 1928			X
Paulianaria Bouchard & Bousquet, gen. nov.					X
Perichilus Quedenfeldt, 1885					X
Periphanodes Gebien, 1943						X
= Periphanes Fairmaire, 1882
Pezomaia Kulzer, 1952						X
Pezophenus Gebien, 1921							X
Phaedis Pascoe, 1866						X
= Pseudeumolpus Kraatz, 1880
= Phaedeucyrtus Pic, 1916
= Microgauromaia Pic, 1921
Phenus Gebien, 1921							X
Phymaeus Pascoe, 1883						X
Picocamaria Masumoto, 1993						X
Pigeostrongylium Kaszab, 1984						X
Pigeus Gebien, 1919						X
= Pseudocamarimena Pic, 1923
Piloxys Fairmaire, 1895					X
Plamius Fairmaire, 1896				X		X
= Cnemandrosus Gebien, 1927
Platycrepis Lacordaire, 1859						X
Poeciltoides Fairmaire, 1896					X
Polopinus Casey, 1924		X
Polposipus Solier, 1848					X
= Pulposipes Gemminger, 1870
= Dysceladus C.O. Waterhouse, 1875
Polypleurus Eschscholtz, 1831		X
Ponapeida Kulzer, 1957								X
Porphyrhyba Fairmaire, 1877					X
= Porphyrohyba Rye, 1879
= Hybocaulus Fairmaire, 1895
Postandrosus Kulzer, 1951						X
Priocamaria Gebien, 1919			X
Promethis Pascoe, 1869				X	X	X	X	X
= Mederis Motschulsky, 1872
= Pediris Motschulsky, 1872
= Setenis Motschulsky, 1872
= Pseudobates Fairmaire, 1882
Proscorus Fairmaire, 1901					X
Pseudabax Kraatz, 1880						X
Pseudamarsenes Ardoin, 1955					X
Pseudandrosus Kulzer, 1951						X
Pseudhadrus Kolbe, 1910					X
= Paradrus Jakobson, 1924
Pseudimmedia Kulzer, 1958			X
Pseudisopus Kulzer, 1957								X
Pseudoblapida Pic, 1917			X
SGBlapidocamaria Pic, 1919			X
SGPseudoblapida Pic, 1917			X
Pseudocamaria Bates, 1879					X
Pseudochrysomela Pic, 1925						X
Pseudoderiles Gebien, 1928			X
Pseudonautes Fairmaire, 1892				X		X
= Thydemus Lewis, 1894
Pseudoperichilus Pic, 1921					X
Pseudopigeus Kaszab, 1984						X
Pseudothryoneus Pic, 1921			X
Psydocamaria Pic, 1923						X
Psydomorphus Pic, 1921						X
Psydus Pascoe, 1868						X
Rehumius Fairmaire, 1893						X
= Melobates Kaszab, 1941
Rhopalobates Fairmaire, 1897						X
Robustocamaria Pic, 1922						X
= Neocamaria Kulzer, 1954
Rhophobas Motschulsky, 1872						X
Sadanaria Ando & Ichiyanagi, 2009						X
Saziches Champion, 1886			X
Scotaeus Hope, 1834						X
Scotoderus Perroud & Montrouzier, 1865							X	X
= Dechius Pascoe, 1866
= Pelecypalpus Hinton, 1947
Scutopiloxys Pic, 1924					X
Simalura Gebien, 1914				X		X
= Microhemicera Pic, 1921
Sophrobates Fairmaire, 1889			X
Sphaerocaulus Fairmaire, 1869					X
Sphaeromatris Fairmaire, 1899					X
Sphaerotidius Kaszab, 1941						X
Sphaerotus W. Kirby, 1819			X
Spheneuphloeus Kaszab, 1941						X
Sphenolampidius Kaszab, 1941						X
Sphenosdara Kaszab, 1941						X
Spinepicalla Pic, 1921			X
Spinoderosphaerus Pic, 1922						X
= Spinogauromaia Pic, 1922
Srilanka Kaszab, 1980						X
Steneucyrtus Fairmaire, 1896						X
= Pseudochariotheca Pic, 1934
Stenochinus Motschulsky, 1860				X	X	X
= Dicraeus Marseul, 1876
= Brachypilium Fairmaire, 1896
= Dicraeosis Gebien, 1911
Stenophanes Solsky, 1876				X
Stenothesilea Kulzer, 1951						X	X
Sternomaia Kulzer, 1952						X
Sthenoboea Champion, 1885			X
Strepsius Fairmaire, 1896					X
Styphloeus Kaszab, 1941						X
Suarezius Fairmaire, 1895					X
Sulpiusoma Ferrer, 2006					X
Sundon Pic, 1923						X
Sycophantes Kirsch, 1866			X
Sycophantomorphus Pic, 1924			X
Tabarus Gebien, 1921							X
Taichius Ando, 1996						X
Taiwanomenephilus Masumoto, 1986						X
Tanchirus Fairmaire, 1897						X
Taphrosoma Kirsch, 1866			X
= Orobychus Pascoe, 1868
Taraxides C.O. Waterhouse, 1876					X
= Dischidus Kolbe, 1886
Tearchus Kraatz, 1880						X
= Heteromerotylus Pic, 1921
Teles Mulsant & Godart, 1876				X
Telethrus Pascoe, 1882			X
Telleus Fairmaire, 1904			X
Temnoaphelus Ferrer, 1988					X
Temnophthalmus Gebien, 1921					X
Tenebriocamaria Pic, 1919			X
Tenebriopsis Gebien, 1928			X
Tenesis Duvivier, 1892					X
Tentyriopsis Gebien, 1928			X
Tetragonomenes Chevrolat, 1878				X		X	X
= Tetragonomecus Rye, 1880
= Obriomaia Gebien, 1927
= Falsoaugolesthus Masumoto, 1993
Tetraphyllus Laporte & Brullé, 1831				X		X
= Adeps Gistel, 1857
= Addia Lewis, 1894
= Adepsion Strand, 1917
Thecacerus Lacordaire, 1859			X
Thesilea Haag-Rutenberg, 1878						X	X	X
Thettea Bates, 1879					X
Thydemorphus Pic, 1918						X
Tonkinius Fairmaire, 1903						X
Toxocnema Fåhraeus, 1870					X
Trichodamatris Chatanay, 1915					X
Uenomisolampidius Masumoto, 1996						X
Upis Fabricius, 1792		X		X
Xanthobates Gebien, 1928			X
Xantusiella Kaszab, 1941						X
Xenius Champion, 1886			X
Xylopinus LeConte, 1862		X
= Taenobates Motschulsky, 1872
Zabroideus Fairmaire, 1894				X
Zophius Dejean, 1834					X
Zophophilus Fairmaire, 1881						X	X
= Sphenothorax Gebien, 1906
= Teremenes Carter, 1914
Stenochiini W. Kirby, 1837
Agissopterus Fairmaire, 1884			X
Asemogena Péringuey, 1904					X
Azonoderus Harold, 1879					X
Bionesus Fairmaire, 1879								X
Bremerianus Masumoto & Bečvář, 2005						X
Cuphotes Champion, 1887			X
= Spheniscus W. Kirby, 1819
= Eucosmus Gistel, 1848
= Phygoscotus Schulz, 1902
Dauresia Ferrer, 2001					X
Dicyrtus Duponchel, 1844			X
Diestesoma Péringuey, 1904					X
Dorelogena Péringuey, 1904					X
Elasmocerella Strand, 1935			X
= Elasmocera Mäklin, 1867
Epiplecta Mäklin, 1867			X
Eucrossoscelis Nakane, 1963						X
Eutherama Carter, 1914							X
Falsocuphotes Pic, 1918			X
Falsonotostrongylium Kaszab, 1955								X
Falsostrongylium Pic, 1915			X
Flabellostrongylium Pic, 1938			X
Freudella Ardoin, 1961					X
Genateropa Bouchard & Bousquet, nom. nov.					X
= Apterogena Ardoin, 1962
Heterostrongylium Kaszab, 1977							X
Holostrongylium Kaszab, 1977						X	X
Hoplostrongylium Ardoin, 1965					X
Hyperchalca Fairmaire, 1869					X
SGHyperchalca Fairmaire, 1869					X
SGMacrohyperchalca Pic, 1935					X
Lophocnemis Mäklin, 1867						X	X
= Pseudostrongylium Kraatz, 1880
= Enganodia Fairmaire, 1898
= Mimothydemus Pic, 1923
Mentes Champion, 1893			X
Microtocerus Pic, 1918			X
Mictopsis Fairmaire, 1899					X
Miotodera Fairmaire, 1901					X
= Cyphelops Fairmaire, 1901
Nodosogylium Pic, 1951					X
Oenomia Pascoe, 1883			X
Oploptera Chevrolat, 1844			X
SGOploptera Chevrolat, 1844			X
= Otocerus Mäklin, 1867
= Hoploptera Gemminger, 1870
SGPlicatocerus Pic, 1918			X
Otoceromorphus Pic, 1915			X
Parastrongylium Kaszab, 1977							X
Phyllechus Bouchard & Bousquet, gen. nov.						X
Phymatosoma Laporte & Brullé, 1831						X
Platyesthus Mäklin, 1878			X
Poecilesthus Dejean, 1834			X
= Dinax Gistel, 1848
= Diestica Pascoe, 1868
Pseudogena Fairmaire, 1899					X
Pseudotocerus Champion, 1888			X
Psilonesogena Bates, 1879					X
= Asthenopoda Chatanay, 1915
Saitostrongylium Masumoto, 1996						X
Strongylacanthus Brèthes, 1925			X
Strongylium W. Kirby, 1819		X	X	X	X	X	X	X
SGAfrostrongylium Robiche, 2019					X
SGStrongylium W. Kirby, 1819		X	X	X	X	X	X	X
= Stenochia W. Kirby, 1819
= Gentinadis Laporte, 1840
= Saerangodes Gistel, 1848
= Anomoearthrum Mäklin, 1867
= Coelolophus Mäklin, 1867
= Xanthothopeia Mäklin, 1867
= Xanthothopia Gemminger, 1870
= Styrax Westwood, 1875
= Messalia Pascoe, 1883
= Strongyliastrum Fairmaire, 1894
= Eustrongylium Kolbe, 1895
= Allostrongylium Kolbe, 1896
= Ebenolus Fairmaire, 1897
= Zuercheria Reitter, 1908
= Crossoscelis Gebien, 1914
= Notostrongylium Carter, 1915
= Pedostrongylium Pic, 1916
= Falsolophocnemis Pic, 1917
= Gibbostrongylium Pic, 1917
= Microstrongylium Pic, 1917
= Poecilesthostrongylium Pic, 1918
= Reminius Casey, 1924
= Mimogoueum Pic, 1952
Theresea Pic, 1917						X
Uenostrongylium Masumoto, 1999						X
Talanini Champion, 1887 (1883)
Talanus Jacquelin du Val, 1857			X
= Dignamptus LeConte, 1878
Stenochiinae incertae sedis
†Anthracohelops Haupt, 1950
†Caryosoma Haupt, 1950
†Eodromus Haupt, 1950
†Mimohelops Haupt, 1950
†Parakeleusticus Haupt, 1950
†Pseudohelops Haupt, 1950
†Pyrochalcaspis Haupt, 1950
Tenebrionidae incertae sedis
Afrohelops Schawaller, 2012					X
Allotriocochabambia Faúndez, Rider & Carvajal, 2014			X
= Cochabambia Marcuzzi, 1985
Ancylopoma Pascoe, 1871			X
Baryscelis Boisduval, 1835							X
Camarothelops Kolbe, 1910					X
†Cretaceites Wang, 1997
†Eoallognosis Haupt, 1950
†Eocallidium Haupt, 1950
†Eohelaeus Haupt, 1950
Erelus Mulsant & Rey, 1853				X
Gnathelops Gebien, 1922					X
Homocyrtus Dejean, 1834			X
= Cyphonotus Guérin-Méneville, 1831
= Schlinkus R. Lucas, 1920
Macrophtalmus Montrouzier, 1855							X
= Macrophthalmata Strand, 1935
†Paropiophorus Haupt, 1950
Physciolagria Pic, 1930					X
†Protoplatycera Wickham, 1914
Pygidiphorus Mulsant, 1856				X
†Rhinohelaeites Haupt, 1950
†Tagenopsis Heer, 1864
†Tenebrionites Cockerell, 1920

Distribution

The world’s major biogeographical realms, as defined by Olson et al. (2001), were used to record the overall distribution of each valid genus and subgenus (Table 2). The maps showing the borders of each realm (Figs 1–5) were produced with the free and open source software QGIS (version 3.10) using the layer Terrestrial Ecoregions of the World (WWF 2012).

Figure 1.

Map showing the biogeographic realms used to record the distribution of each valid genus and subgenus of Tenebrionidae (see Table 2). Blue (dark): Palaearctic; blue (pale): Australasia; green: Neotropic; orange: Afrotropic; red: Nearctic; yellow: Indo-Malay (see Olson et al. (2001: fig. 1) for the boundaries of the Oceania biogeographic realm). Figures 2–5 show closeup maps of boundary areas. The Antarctic realm is excluded since none of the valid Tenebrionidae genera and subgenera are distributed therein.

Figure 5.

Closeup map showing the boundary between the Indo-Malay and Australasia biogeographic realms. Countries with two biogeographic realms are named.

Figure 2.

Closeup map showing the boundary between the Nearctic and Neotropic biogeographic realms. Countries with two biogeographic realms are named.

Figure 3.

Closeup map showing the boundary between the Palaearctic and Afrotropic biogeographic realms. Countries with two biogeographic realms are named.

Figure 4.

Closeup map showing the boundary between the Palaearctic and Indo-Malay biogeographic realms. Countries with two biogeographic realms are named.

Publication dates

We tried to establish the date of publication of works listed in this catalogue as accurately as possible. In the References section, publication dates are listed after each reference in square brackets “[]” and preceded by DP (i.e., date of publication), unless only the year was found. These dates are either specific dates of publication (e.g., extracted from the works themselves or the journal wrappers, title pages, footers) or are the earliest known publication dates (e.g., extracted from recording journals or accounts of society meetings). In the latter case, the date is preceded by the word “by.” Sources of information for the dates of publication from societies, recording journals, and works are mentioned below.

Societies

Acad Nat Sci Phil: Academy of Natural Sciences, Philadelphia (Philadelphia, USA) [Proceedings];

Acad Sci Fr: Académie des Sciences (Paris, France) [Compte Rendus Hebdomadaires];

Acad Sci St. Peters: Académie Impériale des Sciences de St.-Pétersbourg (Saint Petersburg, Russia) [Bulletin Scientifique; Bulletin de la Classe historico-philologique];

Amer Ant Soc: American Antiquarian Society (Worcester, USA) [Proceedings];

Amer Ent Soc: American Entomological Society (Philadelphia, USA) [Proceedings];

Amer Phil Soc: American Philosophical Society (Philadelphia, USA) [Proceedings];

Bost Soc Nat Hist: Boston Society of Natural History (Boston, USA) [Proceedings];

Ent Soc Lond: Entomological Society of London (London, UK) [Transactions, Proceedings];

Ent Ver Stettin: Entomologischer Verein zu Stettin (Stettin, Germany [now Poland]) [Vereinsangelegenheiten];

Nederl Ent Ver: Nederlandsche Entomologische Vereniging (Leiden, Netherlands) [Inhouds-Opgave van Werken];

Roy Soc Queensl: Royal Society of Queensland (Brisbane, Australia) [Proceedings];

Soc Ent Belg: Société Entomologique de Belgique (Brussels, Belgium) [Annales, Comptes-Rendus];

Soc Ent Fr: Société Entomologique de France (Paris, France) [Annales, Bulletin];

Soc Imp Nat Mosc: Société Impériale des Naturalistes de Moscou (Moscow, Russia) [Bulletin].

Recording journals

Allg Bibl Deutsch: Allgemeine Bibliographie für Deutschland (Leipzig, Germany);

Bibl Belg: Bibliographie de la Belgique (Brussels, Belgium);

Bibl Fr: Bibliographie de la France (Paris, France);

Bull Nord: Bulletin du Nord, journal scientifique et littéraire, contenant: des mémoires et notices, des analyses et extraits d’ouvrages nouveaux; des variétés et mélanges, des annonces bibliographiques, etc., etc. (Moscow, Russia);

Ent Nachr: Entomologische Nachrichten (Berlin, Germany);

Lit Centrbl: Literarisches Centralblatt für Deutschland (Leipzig, Germany);

Lit Ztg: Literarische Zeitung (Berlin, Germany);

Nat Nov: Naturae Novitates (Berlin, Germany);

Naturaliste: Le Naturaliste (Paris, France);

Pet Nouv Ent: Petites Nouvelles Entomologiques (Paris, France);

Rev Coleopt: Revue Coléoptérologique (Brussels, Belgium);

Zool Rec: Zoological Record (London, UK).

Works reporting dates of publication

The following works included important information for dating the references: Kielmeyer and Jäger (1835), Guérin-Méneville (1838, 1858), Lefèvre (1885, 1895), Sharp (1892a), Sclater (1893), Oshanin (1910), Wheeler (1912), Sherborn (1934), Duncan (1937), Peavot (1937), F.H. Waterhouse (1937), ICZN (1957, 1958, 1999), F.M. Brown (1964), Raphael (1970), Cowan (1976), Kerzhner (1984), Hayek (1989), Baker (1996), Evenhuis (1997a, b, 2002, 2003, 2012, 2015, 2019a, b), Nagel and Schmidlin (2014), Merkl et al. (2008), Bouchard et al. (2011), Fujioka (2011), Bousquet (2016a, b, 2017), Williams (2017).

Results

Overall diversity and distribution

The family Tenebrionidae contains 4122 nomenclaturally available genus-group names (see Table 2 and List of available genus-group names in Tenebrionidae Latreille, 1802 below). Of the 2307 valid genera, 33 are extinct. A total of 761 subgenera are currently used as valid. The subfamily Pimeliinae contains the highest number of valid genera (n = 578) followed by Stenochiinae (n = 394), Tenebrioninae (n = 349), Blaptinae (n = 300), Lagriinae (n = 273), Alleculinae (n = 231) and Diaperinae (n = 128). The other four subfamilies contain few genera (Phrenapatinae = 28, Zolodininae = 3, Kuhitangiinae = 2 and Nilioninae = 1). Twenty valid genera could not be confidently placed in a subfamily and are included here as Tenebrionidae incertae sedis. The Afrotropic biogeographical realm is the most diverse with 1022 valid extant genus-group taxa (755 genera, 267 subgenera; see Table 2). The Palaearctic, Indo-Malay, and Neotropic realms follow with 925 (545 genera, 380 subgenera), 610 (501, 109) and 518 (428, 90) valid extant genus-group taxa respectively. The lowest number of valid extant genus-group taxa are in Australasia with 348 (310 genera, 38 subgenera), the Nearctic with 279 (206, 73) and Oceania with 62 (52, 10).

List of available genus-group names in Tenebrionidae Latreille, 1802

Abantiades Fairmaire, 1894f: 395 [M]. Type species [automatic]: Abantisaenescens Fairmaire, 1892, by monotypy. Status: junior synonym of Diphyrrhynchus Fairmaire, 1849 in Blaptinae: Opatrini: Heterotarsina. Note: replacement name for Abantis Fairmaire, 1892; junior homonym of Abantiades Herrich-Schäffer, 1856 [Lepidoptera].

Abantis Fairmaire, 1892a: 109 [F]. Type species: Abantisaenescens Fairmaire, 1892, by monotypy. Status: junior synonym of Diphyrrhynchus Fairmaire, 1849 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Gebien (1938a: 407). Note: junior homonym of Abantis Hoppfer, 1855 [Lepidoptera].

Aberlencus Iwan, 2002b: 559 [M]. Type species: Aberlencusangolensis Iwan, 2002, by original designation. Status: junior synonym of Angolositus Koch, 1955 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 90).

Abiga Guérin-Méneville, 1860: clxxxix [F]. Type species: Abigahumilis Guérin-Méneville, 1860, by subsequent designation (Löbl et al. 2008a: 40). Status: junior synonym of Scelosodis Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Guérin-Méneville (1862: 376).

Abigopsis Escalera, 1914: 277 [F]. Type species: Scelosodisustus Fairmaire, 1879, by subsequent designation (Gebien 1937a: 612). Status: valid genus in Pimeliinae: Tentyriini.

Ablapsis Reitter, 1887a: 368 [F]. Type species: Blaps compressipes Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Acanthioides Fairmaire, 1894e: 320 [F]. Type species: Acanthioidesasperula Fairmaire, 1894, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Acanthobas Gebien, 1928: 168, 175 [M]. Type species: Acanthobasangusticollis Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Acanthoblaps Reitter, 1889a: 687 [F]. Type species: Acanthoblapsdentitibia Reitter, 1889, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gebien (1910b: 226).

Acanthocamaria Gebien, 1919: 26, 31 [F]. Type species: Acanthocamariabrunneoopaca Gebien, 1919, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Acanthomera Latreille, 1828: 580 [F]. Type species: Pimeliadentipes Fabricius, 1787, by monotypy. Status: senior synonym of Psorodes Dejean, 1834 in Tenebrioninae: Amarygmini. Note: junior homonym of Acanthomera Wiedemann, 1821 [Diptera].

Acanthomerus Agassiz, 1846b: 2 [M]. Type species [automatic]: Pimeliadentipes Fabricius, 1787, by monotypy. Status: junior synonym of Psorodes Dejean, 1834 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Acanthomera Latreille, 1828, not in prevailing usage.

Acanthosternus Montrouzier, 1860: 289 [M]. Type species: Acanthosternushalorageos Montrouzier, 1860, by monotypy. Status: junior synonym of Diphyrrhynchus Fairmaire, 1849 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Bates (1872a: 97).

Acantophorus Billberg, 1820: 32 [M]. Type species: Erodiusgibbus Fabricius, 1775, by subsequent designation (Bouchard and Bousquet 2020b: 7). Status: junior synonym of Erodius Fabricius, 1775 in Pimeliinae: Erodiini. Synonymy: Bouchard and Bousquet (2020b: 7).

Acastus Péringuey, 1896: 177 [M]. Type species: Acastusagrestis Péringuey, 1896, by subsequent designation (Ardoin 1963a: 87). Status: junior synonym of Gonocnemis J. Thomson, 1858 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1913: 76).

Accanthopus Dejean, 1821: 71 [M]. Type species: fixed herein (ICZN 1999, Article 70.3) as Tenebriovelikensis Piller & Mitterpacher, 1783, misidentified as Tenebriocaraboides Linnaeus, 1758 in the original designation by monotypy in Dejean (1821). Status: valid genus in Tenebrioninae: Helopini: Enoplopodina. Note: the type species “Blaps caraboides Germ.” was first established by monotypy; Seidlitz (1895: 681) first noted that Tenebriocaraboides Linnaeus of Germar (1817) was identical to Helopsdentipes Rossi, 1790, currently a junior synonym of Tenebriovelikensis Piller & Mitterpacher, 1783; we follow currently accepted concepts (e.g., Löbl et al. 2008b: 246) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Tenebriocaraboides Linnaeus, 1758 is a valid species in the genus Cychrus Fabricius, 1794 [Coleoptera: Carabidae].

Acerogria Borchmann, 1936: 19, 137 [F]. Type species: Cerogriadohrni Borchmann, 1911, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Acestophanus Koch, 1950a: 67 [M]. Type species [automatic]: Acestuselongatus Haag-Rutenberg, 1875, by subsequent designation (R. Lucas 1920: 69). Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Acestus Haag-Rutenberg, 1875.

Acestus Haag-Rutenberg, 1875b: 4, 56 [M]. Type species: Acestuselongatus Haag-Rutenberg, 1875, by subsequent designation (R. Lucas 1920: 69). Status: senior synonym of Acestophanus Koch, 1950 in Pimeliinae: Adelostomini. Note: junior homonym of Acestus Leidy, 1851 [Annelida].

Achaemenes Bogatchev, 1949: 39 [M]. Type species: Achaemenesvillosus Bogatchev, 1949 (= Thripterabogatchevi Kwieton, 1982), by original designation. Status: senior synonym of Bogatshevia G.S. Medvedev & Iwan, 2006 in Pimeliinae: Pimeliini. Note: junior homonym of Achaemenes Stål, 1866 [Hemiptera].

Achanius Erichson, 1847a: 118 [M]. Type species: Achaniusanthicoides Erichson, 1847, by monotypy. Status: valid genus and subgenus in Pimeliinae: Evaniosomini. Note: transferred from Edrotini (where it was previously placed by Doyen (1994: 500)) by Flores and Aballay (2015: 167).

Achariotheca Kaszab, 1970a: 273 [F]. Type species: Achariothecabaloghi Kaszab, 1970, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Achora Pascoe, 1869: 279 [F]. Type species: Asidaserricollis Hope, 1843 (= Cestrinusobscurus Erichson, 1842), by monotypy. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Gemminger in Gemminger and Harold (1870: 1929, with Prionotus Mulsant & Rey, 1859, a junior synonym of Isopteron Hope, 1841); Matthews (1998: 788).

Achrostus Fairmaire, 1891b: 256 [M]. Type species: Achrostusrufonitens Fairmaire, 1891, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Achthosus Pascoe, 1863a: 42 [M]. Type species: Achthosuswestwoodii Pascoe, 1863, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Acidia Illiger, 1804: 79 [F]. Type species [automatic]: Pimeliareflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Akis Herbst, 1799 in Pimeliinae: Akidini. Note: unjustified emendation of Akis Herbst, 1799, not in prevailing usage.

Acis Billberg, 1820: 32 [F]. Type species [automatic]: Pimeliareflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Akis Herbst, 1799 in Pimeliinae: Akidini. Note: unjustified emendation of Akis Herbst, 1799, not in prevailing usage.

Acisba Dejean, 1834: 185 [F]. Type species: Tentyriacribrosa Besser, 1832 (= Pimeliapunctata Fabricius, 1798), by subsequent designation (Löbl et al. 2008a: 40). Status: junior synonym of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Solier (1835b: 288). Note: the original combination of the accepted name of the type species, Pimeliapunctata Fabricius, 1798, is a junior primary homonym of Pimeliapunctata Thunberg, 1787.

Acmoeus Fåhraeus, 1870: 293 [M]. Type species: Anomalipuselephas Fåhraeus, 1870, by monotypy. Status: junior synonym of Anomalipus Guérin-Méneville, 1831 in Blaptinae: Platynotini: Platynotina. Synonymy: Gebien (1938a: 409).

Aconobius Casey, 1895: 617 [M]. Type species: Conibiosomalaciniata Casey, 1892, by original designation. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Acontodactylus Desbrochers des Loges, 1894: 5 [M]. Type species: Cossyphusminutissimus Laporte, 1840, by original designation. Status: valid subgenus of Cossyphus G.-A. Olivier, 1791 in Lagriinae: Cossyphini.

Acritolagria Borchmann, 1916a: 48, 98 [F]. Type species: Lagriafulvopilosa Fairmaire, 1887, by subsequent designation (Borchmann 1936: 166). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Acromaticus Koch, 1955a: 143 [M]. Type species: Sepidiumstriatum Quensel, 1806 (= Sepidiumstriatum Thunberg, 1787), by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Acropachia Mäklin, 1875: 656 [F]. Type species: Acropachiabifoveolata Mäklin, 1875, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Acropteron Perty, 1832: 64 [N]. Type species: Acropteronrufipes Perty, 1832 (= Acropteryxrufipes Gistel, 1831), by subsequent designation (Hope 1841: 133). Status: junior synonym of Acropteryx Gistel, 1831 in Tenebrioninae: Acropteronini. Synonymy: Gistel (1857: 551, synonymy of the type species), Martins and Pereira (1966: 159, as “Acropteroxysrufipes Distl., 1831”).

Acropterum Agassiz, 1846b: 6 [N]. Type species [automatic]: Acropteronrufipes Perty, 1832 (= Acropteryxrufipes Gistel, 1831), by subsequent designation (Hope 1841: 133). Status: junior synonym of Acropteryx Gistel, 1831 in Tenebrioninae: Acropteronini. Note: unjustified emendation of Acropteron Perty, 1832, not in prevailing usage.

Acropteryx Gistel, 1831: 308 [F]. Type species: Acropteryxrufipes Gistel, 1831, by original designation. Status: valid genus in Tenebrioninae: Acropteronini. Note: genus previously included in Erotylidae: Languriinae (see Leschen and Węgrzynowicz 1998: 234); reversal of precedence cannot be used to conserve the usage of Acropteron Perty, 1832 because Acropteryx was used as valid after 1899 (e.g., Blackwelder 1945: 427).

Acroschatia Wilke, 1922: 269 [F]. Type species: Microschatiarobusta Horn, 1893, by original designation. Status: junior synonym of Microschatia Solier, 1836 in Pimeliinae: Asidini. Synonymy: K.W. Brown and Doyen (1992: 546).

Actanorie Bates, 1879a: 289 [M]. Type species: Camariaundaticollis Fairmaire, 1875, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Actizeta Pascoe, 1875: 214 [F]. Type species: Actizetaammobioides Pascoe, 1875 (= Actizetaalbata Pascoe, 1875), by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Actizetina.

Acutogria Merkl, 1988a: 137 [F]. Type species: Acutogriafalcata Merkl, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Acutoodescelis Kaszab, 1940b: 941, 951 [F]. Type species: Platyscelispunctatissima Fairmaire, 1886, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Adamus Iwan, 1997: 255 [M]. Type species: Platynotusmicrositoides Kaszab, 1975, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Adavius Mulsant & Rey, 1859a: 126, 138 [M]. Type species: Adaviusclavipes Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Addia Lewis, 1894: 465 [F]. Type species: Addiascatebrae Lewis, 1894, by monotypy. Status: junior synonym of Tetraphyllus Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini. Synonymy: Masumoto (1982a: 23, probable); Ando (1991: 64, confirmed).

Adelina Dejean, 1835: 315 [F]. Type species: Cucujusplanus Fabricius, 1801, by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Adelium W. Kirby, 1819a: 420 [N]. Type species: Adeliumcalosomoides W. Kirby, 1819, by subsequent designation (Meyer 1844: 180). Status: valid genus in Lagriinae: Adeliini. Note: the type species previously recognized for this genus was Carabusporcatus Fabricius, 1775, by subsequent designation by Lacordaire (1859b: 438).

Adelodemus Haag-Rutenberg, 1878: 100 [M]. Type species: Adelodemusasperulus Haag-Rutenberg, 1878 (= Cestrinussqualidus W.J. MacLeay, 1872), by monotypy. Status: valid genus in Lagriinae: Adeliini.

Adelonia Laporte, 1840: 221 [F]. Type species: Ulomafiliforme Laporte, 1840, by monotypy. Status: valid genus and subgenus in Lagriinae: Belopini.

Adelostoma Duponchel, 1827: 342 [N]. Type species: Adelostomasulcatum Duponchel, 1827, by monotypy. Status: valid genus and subgenus in Pimeliinae: Adelostomini.

Adelostomoides Carl, 1991: 24 [M]. Type species: Adelostomagrande Haag-Rutenberg, 1879, by monotypy. Status: junior synonym of Zarudnionymus Semenov-Tjan-Shansky & Bogatchev, 1947 in Pimeliinae: Adelostomini. Synonymy: Purchart (2007: 240).

Adelozotypus Kaszab, 1982b: 224 [M]. Type species: Adelozotypusnovaecaledoniae Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Adeliini.

Adelphinops Reitter, 1922b: 168, 169 [M]. Type species: Adelphinusordubadensis Reitter, 1890, by monotypy. Status: valid subgenus of Adelphinus Fairmaire & Coquerel, 1866 in Tenebrioninae: Helopini: Helopina.

Adelphinus Fairmaire & Coquerel, 1866: 44 [M]. Type species: Eutrapelasuturalis P.H. Lucas, 1847, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina.

Adelphus Dejean, 1834: 208 [M]. Type species: Helopsmarginatus Fabricius, 1792, by subsequent designation (Bousquet and Bouchard 2013a: 58). Status: senior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Synonymy: Chevrolat (1847b: 458). Note: nomen oblitum (see Bouchard and Bousquet, 2020b: 6).

Adeps Gistel, 1857: 63 [M]. Type species: Amarygmuspaykullii Dalman, 1823, by original designation. Status: junior synonym of Tetraphyllus Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini. Synonymy: new synonym [PB]. Note: Adeps Gistel, 1857, and its replacement name Adepsion Strand, 1917, have been forgotten in the literature; the type species of Adeps is currently included in the genus Tetraphyllus Laporte & Brullé, 1831 and for that reason Gistel’s name is considered a junior synonym of Tetraphyllus; junior homonym of Adeps Gistel, 1848 [Crustacea].

Adepsion Strand, 1917: 90 [N]. Type species [automatic]: Amarygmuspaykullii Dalman, 1823, by original designation. Status: junior synonym of Tetraphyllus Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini. Synonymy: new synonym [PB]. Note: replacement name for Adeps Gistel, 1857.

Ades Guérin-Méneville, 1857: 277 [M]. Type species: Adeshemisphericus Guérin-Méneville, 1857, by monotypy. Status: valid genus in Diaperinae: Leiochrinini.

Adesmia Fischer, 1822: 153 [F]. Type species: Adesmialongipes Fischer, 1822 (= Pimeliaanomala Fischer, 1820), by monotypy. Status: valid genus and subgenus in Pimeliinae: Adesmiini.

Adesmina Reitter, 1916a: 5, 29 [F]. Type species: Adesmiaarabica Reitter, 1916, by subsequent designation (Löbl et al. 2008b: 122). Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Adisema Borchmann, 1936: pl. 7 [F]. Type species: Disemaambigua Mäklin, 1875, by monotypy. Status: junior synonym of Nemostiromorpha Pic, 1917 in Lagriinae: Lagriini: Statirina.

Adonicus Fairmaire, 1891b: 258 [M]. Type species: Adonicuspurpuripennis Fairmaire, 1891, by monotypy. Status: junior synonym of Alcyonotus Pascoe, 1882 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1905: 255).

Adordanea Reitter, 1897b: 229 [F]. Type species: Microderasubseriata Reitter, 1889, by monotypy. Status: junior synonym of Dordanea Reitter, 1887 in Pimeliinae: Tentyriini. Synonymy: Kaszab (1966: 304), G.S. Medvedev (1990: 112).

Adoryacus Koch, 1963: 34 [M]. Type species: Stizopusrotundicollis Fairmaire, 1897, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Adosogria Borchmann, 1936: 16, 63 [F]. Type species: Lagriabennigseni Borchmann, 1909, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Adynata Fåhraeus, 1870: 330 [F]. Type species: Adynatatricolor Fåhraeus, 1870, by subsequent designation (Borchmann 1936: 23). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Aeanes Champion, 1893a: 566 [M]. Type species: Aeanesangusticollis Champion, 1893, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Aediatorix Bates, 1868: 315 [M]. Type species: Aediatorixjansoni Bates, 1868, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Aemymone Bates, 1868: 314 [F]. Type species: Goniaderacariosa Bates, 1868, by original designation. Status: valid subgenus of Goniadera Perty, 1832 in Lagriinae: Goniaderini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Aequigula Penrith, 1979: 20, 23 [F]. Type species: Alogeniushughesae Penrith, 1979, by original designation. Status: valid subgenus of Alogenius Gebien, 1910 in Pimeliinae: Adesmiini.

Aeschrocera Chen & Chou, 1996: 265 [F]. Type species: Cerogriabrunneocollis Chen & Chou, 1996 (= Cerogriadiversicornis Pic, 1933), by original designation. Status: junior synonym of Cerogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina. Synonymy: Merkl (2007: 262).

Aesthetus C.O. Waterhouse, 1890: 552 [M]. Type species: Aesthetustuberculatus C.O. Waterhouse, 1890, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Aesymnus Champion, 1886: 168 [M]. Type species: Aesymnusnitidus Champion, 1886, by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Aethales Dejean, 1834: 180 [M]. Type species: Epitragusbrunnicornis Latreille, 1811, by monotypy. Status: junior synonym of Epitragus Latreille, 1802 in Pimeliinae: Epitragini. Synonymy: Bousquet and Bouchard (2013a: 43). Note: as mentioned by Bousquet and Bouchard (2013a: 43) the type species Epitragusbrunnicornis Latreille was listed as a species incertae sedis in the genus Epitragus Latreille, 1802 by Freude (1967: 176).

Aethalides Bates, 1873d: 50 [M]. Type species: Aethalidespunctipennis Bates, 1873, by monotypy. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 473).

Aethyssius Pascoe, 1863a: 45 [M]. Type species [automatic]: Atractusviridis Boisduval, 1835, by subsequent designation (Duponchel and Chevrolat 1841: 312). Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324). Note: replacement name for Atractus Boisduval, 1835.

Afghanillus Kaszab, 1960a: 1 [M]. Type species: Afghanillusklapperichi Kaszab, 1960, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Afghanopachys Kwieton, 1978: 29, 30 [M]. Type species: Pachyscelishaarlovi Gridelli, 1954, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Afrasida Wilke, 1922: 260 [F]. Type species: Asidacaryophyllea Wiedemann, 1823, by original designation. Status: valid genus and subgenus in Pimeliinae: Asidini.

Afrethas Koch, 1962a: 23 [M]. Type species: Stenosispusillima Fairmaire, 1888, by original designation. Status: valid subgenus of Anethas Jakobson, 1924 in Pimeliinae: Stenosini: Stenosina.

Afrinus Fairmaire, 1888a: 189 [M]. Type species: Afrinusgrandicornis Fairmaire, 1888, by subsequent designation (R. Lucas 1920: 79). Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: the designation of Afrinusstriolifrons Fairmaire, 1888 as the type species of Afrinus Fairmaire, 1888 by Koch (1950b: 329) is invalid (ICZN 1999, Article 69.1).

Afrobyrsax Ardoin, 1973: 894 [M]. Type species: Afrobyrsaxgirardi Ardoin, 1973, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Afrohelops Schawaller, 2012a: 75 [M]. Type species: Afrohelopskenyaensis Schawaller, 2012, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: removed from the tribe Helopini and placed as Tenebrionidae incertae sedis by Nabozhenko (2018: 183).

Afrolaena Endrödy-Younga & Schawaller, 2002: 9, 11 [F]. Type species: Afrolaenatibialis Endrödy-Younga & Schawaller, 2002, by original designation. Status: valid genus in Lagriinae: Laenini.

Afronosis G.S. Medvedev, 1995a: 849, 859 [F]. Type species: Stenosisciliaris Gebien, 1920, by original designation. Status: valid subgenus of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina. Note: Stenosisleontjevi G.S. Medvedev, 1995 was listed as a second type species of this genus in the original publication (G.S. Medvedev, 1995: 859); we act as First Revisers and select Stenosisciliaris Gebien, 1920 as the type species for Afronosis G.S. Medvedev, 1995.

Afrostrongylium Robiche, 2019a: 83 [N]. Type species: Strongyliumfrancoisi Robiche, 2019, by original designation. Status: valid subgenus of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini.

Afrotagalus Gebien, 1942b: 111, 120 [M]. Type species: Afrotagaluseidmanni Gebien, 1942, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Afrotenebrio Gridelli, 1951: 223, 230 [M]. Type species: Tenebrioguineensis Imhoff, 1843, by original designation. Status: valid subgenus of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini.

Agastenes R. Lucas, 1920: 79 [M]. Type species [automatic]: Agastheneswestwoodi Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: unjustified emendation of Agasthenes Bates, 1873.

Agasthenes Bates, 1873e: 352 [M]. Type species: Agastheneswestwoodi Bates, 1873, by monotypy. Status: senior synonym of Agastenes R. Lucas, 1920 in Tenebrioninae: Heleini: Cyphaleina. Note: junior homonym of Agasthenes Förster, 1869 [Hymenoptera].

Agelarches Gistel, 1848a: x [M]. Type species [automatic]: Pimeliaangulata Fabricius, 1775, by subsequent designation (Hope 1841: 118). Status: junior synonym of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Pimelia Fabricius, 1775.

Agissopterus Fairmaire, 1884a: 513 [M]. Type species: Agissopterussemipunctatus Fairmaire, 1884, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Aglypta Gebien, 1908a: 329 [F]. Type species: Aglyptaoctocostata Gebien, 1908, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Agnaptoria Reitter, 1887a: 364, 372 [F]. Type species: Agnaptoriarubripes Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Agonopus Gebien, 1920: 110 [M]. Type species: Gonopuspuncticollis Solier, 1848, by subsequent designation (Endrödy-Younga 2000: 7). Status: valid subgenus of Gonopus Latreille, 1828 in Blaptinae: Platynotini: Platynotina.

Agraecus Fairmaire, 1900a: 21 [M]. Type species: Agraecuschalcoides Fairmaire, 1900, by monotypy. Status: junior synonym of Carabelops Fairmaire, 1899 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1956b: 90).

Agroblaps Motschulsky, 1860c: 531 [F]. Type species: Blaps fatidica Sturm, 1807 (= Blaps lethifera Marsham, 1802), by subsequent designation (Nabozhenko 2008: 35). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860).

Agymnonyx Gebien, 1921a: 325, 328 [M]. Type species: Agymnonyxsulciventris Gebien, 1921, by subsequent designation (Gebien 1942a: 334). Status: valid genus in Stenochiinae: Cnodalonini.

Ahexaroptrum Kaszab, 1960b: 291 [N]. Type species: Ahexaroptrumhumeridens Kaszab, 1960, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Ainu Lewis, 1894: 479 [M]. Type species: Ainutenuicornis Lewis, 1894, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Akis Herbst, 1799: 124 [F]. Type species: Pimeliareflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: valid genus in Pimeliinae: Akidini.

Alaephus Horn, 1870: 344, 346 [M]. Type species: Alaephuspallidus Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Vacronini.

Alaetrinus Iwan, 1995a: 14, 24 [M]. Type species: Tenebriopullus Sahlberg, 1823, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Alagria Borchmann, 1916a: 50, 183 [F]. Type species [automatic]: Lagriostirahispida Kolbe, 1902 (= Lagriasubseriata Reitter, 1880), by subsequent designation (Borchmann 1936: 224). Status: valid genus in Lagriinae: Lagriini: Lagriina. Note: replacement name for Lagriostira Kolbe, 1902.

Alaudes Horn, 1870: 361 [M]. Type species: Alaudessingularis Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Cnemeplatiini: Alaudina.

Alcinoe Ménétriés, 1849: 230 [F]. Type species: Alcinoehelopioides Ménétriés, 1849, by monotypy. Status: senior synonym of Alcinoeta Strand, 1929 in Pimeliinae: Tentyriini. Note: junior homonym of Alcinoe Rang, 1828 [Ctenophora].

Alcinoeta Strand, 1929: 23 [F]. Type species [automatic]: Alcinoehelopioides Ménétriés, 1849, by monotypy. Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Alcinoe Ménétriés, 1849.

Alcmeonis Bates, 1868: 270 [F]. Type species: Alcmeonispulchra Bates, 1868, by monotypy. Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324, with Aethyssius Pascoe, 1863, a junior synonym of Lepturidea Fauvel, 1862).

Alcyonotus Pascoe, 1882: 35 [M]. Type species: Alcyonotusiridescens Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Alegoria Laporte, 1840: 221 [F]. Type species: Alegoriadilatata Laporte, 1840, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Alethia Champion, 1888: 417 [F]. Type species: Alethiasallaei Champion, 1888, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Alhuena Kulzer, 1956b: 912 [F]. Type species: Alhuenapenai Kulzer, 1956 (= Peltolobuswaterhousei Bates, 1873), by original designation. Status: junior synonym of Peltolobus Lacordaire, 1859 in Pimeliinae: Trilobocarini. Synonymy: Kulzer (1962: 100).

Alienophloeus Bremer, 2018: 45 [M]. Type species: Corticeuspumilio Bremer, 2018, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Alienoplonyx Bremer, 2019: 59 [M]. Type species: Alienoplonyxalleni Bremer, 2019, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: we act as First Revisers and reject the alternative original spelling Alienolonyx, used by Bremer (2019: 60).

Allardius Ragusa, 1898: 130 [M]. Type species: Parablopsoculatus Baudi di Selve, 1876, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Allecula Fabricius, 1801b: 21 [F]. Type species: Cistelamorio Fabricius, 1787, by subsequent designation (Duponchel 1840: 283). Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Alleculina Pic, 1954: 246 [F]. Type species: Synalleculamacroceps Pic, 1922, by original designation. Status: valid subgenus of Allecula Fabricius, 1801 in Alleculinae: Alleculini: Alleculina.

Alleculodes Borchmann, 1925: 335 [F]. Type species: Alleculodesdiscrepans Borchmann, 1925, by original designation. Status: junior synonym of Bolbostetha Fairmaire, 1896 in Alleculinae: Alleculini: Alleculina. Synonymy: Borchmann (1941a: 23).

Alleculopsis Semenov, 1894: 609 [F]. Type species: Alleculopsisdeserticola Semenov, 1894, by original designation. Status: valid subgenus of Mycetocharina Seidlitz, 1890 in Alleculinae: Alleculini: Alleculina.

Allocera Borchmann, 1916a: 49, 175 [F]. Type species: Porrolagriasubaenea Borchmann, 1908, by subsequent designation (Borchmann 1936: 227). Status: senior synonym of Cerostira Borchmann, 1942 in Lagriinae: Lagriini: Lagriina. Note: junior homonym of Allocera Sichel, 1866 [Hymenoptera].

Allocossyphodes Andreae, 1961: 204, 215 [M]. Type species: Cossyphodesarnoldi Brauns, 1925, by original designation. Status: junior synonym of Cossyphodes Westwood, 1851 in Pimeliinae: Cossyphodini: Cossyphodina. Synonymy: Schawaller (2013c: 362, implied by inclusion of Cossyphodesarnoldi Brauns, 1925 in Cossyphodes Westwood, 1851 without use of a subgenus rank).

Allodengitha Bogatchev, 1963: 57 [F]. Type species: Allodengithadeserta Bogatchev, 1963, by original designation. Status: junior synonym of Alcinoeta Strand, 1929 in Pimeliinae: Tentyriini. Synonymy: G.S. Medvedev (1989: 798).

Allogria Borchmann, 1916a: 48, 100 [F]. Type species: Allogriaspinosa Borchmann, 1916, by subsequent designation (Borchmann 1936: 74). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Allomyladion Bogatchev, 1972: 626 [N]. Type species: Penthicusporicollis Reitter, 1896, by original designation. Status: valid subgenus of Penthicus Faldermann, 1836 in Tenebrioninae: Opatrini: Opatrina.

Allopezus Gebien, 1922a: 504 [M]. Type species: Allopezusmiritarsis Gebien, 1922, by monotypy. Status: junior synonym of Asbolodomimus Pic, 1921 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1948: 543).

Allophasia Pascoe, 1871: 351 [F]. Type species: Allophasiafryi Pascoe, 1871, by monotypy. Status: junior synonym of Diaperis Geoffroy, 1762 in Diaperinae: Diaperini: Diaperina. Synonymy: Triplehorn and Brendell (1985: 14).

Allophylax Bedel, 1906b: 177 [M]. Type species: Phylaxlittoralis Mulsant, 1854 (= Opatrumpicipes G.-A. Olivier, 1812), by subsequent designation (Gebien 1938a: 412). Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina. Note: replacement name for Neophylax Bedel, 1906.

Allostrongylium Kolbe, 1896: 364 [N]. Type species: Allostrongyliumsilvestre Kolbe, 1896, by present designation. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 539, with Anomoearthrum Mäklin, 1867, a junior synonym of Strongylium W. Kirby, 1819).

Allotadzhikistania Bogatchev, 1960a: 43 [F]. Type species: Allotadzhikistaniacomata Bogatchev, 1960, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Allotriocochabambia Faúndez, Rider & Carvajal, 2014: 595 [F]. Type species [automatic]: Cochabambiakulzeri Marcuzzi, 1985, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: replacement name for Cochabambia Marcuzzi, 1985; Marcuzzi (1985: 185) mentioned that his new genus Cochabambia could not be placed in “any given Neotropical tribe so far known” and is therefore included here in Tenebrionidae incertae sedis under its replacement name.

Alobates Motschulsky, 1872: 25 [M]. Type species: Tenebriopensylvanicus DeGeer, 1775, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Alogenius Gebien, 1910a: 91 [M]. Type species [automatic]: Metriopusfavosus Erichson, 1843, by subsequent designation (Gebien 1937a: 661). Status: valid genus and subgenus in Pimeliinae: Adesmiini. Note: replacement name for Pedionomus Haag-Rutenberg, 1875.

Alogista Fåhraeus, 1870: 318 [F]. Type species: Alogistaabnormis Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Alogistopsis Borchmann, 1943: 53 [F]. Type species: Alogistopsispilistriata Borchmann, 1943, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Alphasida Escalera, 1905a: 380 [F]. Type species: Asidagaditana Escalera, 1905 (= Alphasidatypica Gebien 1937), by subsequent designation (Casey 1912: 78). Status: valid genus and subgenus in Pimeliinae: Asidini.

Alphitobius Stephens, 1829: 19 [M]. Type species: Helopspicipes Panzer, 1794 (= Opatrumlaevigatum Fabricius, 1781), by monotypy (see ICZN 1975, Opinion 1039). Status: valid genus in Tenebrioninae: Alphitobiini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1975, Opinion 1039).

Alphitophagus Stephens, 1832b: 12 [M]. Type species: Alphitophagusquadripustulatus Stephens, 1832 (= Diaperisbifasciata Say, 1824), by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

†Alphitopsis Kirejtshuk, Nabozhenko & Nel, 2011: 549 [F]. Type species: Alphitopsisinitialis Kirejtshuk, Nabozhenko & Nel, 2011, by original designation. Status: valid genus in Tenebrioninae: Alphitobiini. Note: described from Lower Cretaceous deposits (China).

Altes Pascoe, 1869: 288, 290 [M]. Type species: Chartopteryxbinodosus Pascoe, 1862, by original designation. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Gemminger in Gemminger and Harold (1870: 1994, with Chartopteryx Westwood, 1841, a synonym of Cyphaleus Westwood, 1841); Matthews (1992: 490).

Altiprosodes G.S. Medvedev, 1997: 580 [M]. Type species: Prosodeskuhistanica G.S. Medvedev, 1996, by original designation. Status: junior synonym of Megaprosodes Reitter, 1909 in Blaptinae: Blaptini: Prosodina. Synonymy: G.S. Medvedev (2001: 83). Note: originally described as a section within a subgenus.

Alymon Pascoe, 1866a: 484 [M]. Type species: Alymonprolatus Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Amachla Koch, 1962a: 117 [F]. Type species: Machlasulcicollis Fåhraeus, 1871, by original designation. Status: valid genus in Pimeliinae: Asidini.

Amarantha Motschulsky, 1860b: 141 [F]. Type species: Amaranthaviridis Motschulsky, 1860, by monotypy. Status: senior synonym of Metaclisa Jacquelin du Val, 1861 in Tenebrioninae: Metaclisini. Synonymy: Lewis (1891: 70). Note: nomen oblitum (see Bouchard et al. 2007: 393).

Amaropsis Champion, 1893a: 567 [F]. Type species: Amaropsisannulicornis Champion, 1893, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Amarosoma Redtenbacher, 1868: 131 [N]. Type species: Amarosomasimulans Redtenbacher, 1868, by monotypy. Status: junior synonym of Pheloneis Pascoe, 1866 in Lagriinae: Adeliini. Synonymy: Pascoe (1876: 52).

Amarsenes Bates, 1879a: 297 [M]. Type species: Tetraphyllusoblongocamelus Fairmaire, 1877, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Amarygmimus Bates, 1873e: 354 [M]. Type species: Amarygmimusduboulayi Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Amarygmomimus Rye, 1875: 290 [M]. Type species [automatic]: Amarygmimusduboulayi Bates, 1873, by monotypy. Status: junior synonym of Amarygmimus Bates, 1873 in Tenebrioninae: Heleini: Cyphaleina. Note: unjustified emendation of Amarygmimus Bates, 1873, not in prevailing usage.

Amarygmus Dalman, 1823: 60 [M]. Type species: Chrysomelamicans Fabricius, 1794, by subsequent designation (Gebien 1921a: 410). Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Amathobius Gebien, 1920: 129 [M]. Type species: Amathobiusglyptopterus Gebien, 1920, by subsequent designation (Koch 1963: 42). Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Amatodes Dejean, 1834: 189 [F]. Type species: Pimeliagemmata Fabricius, 1801, by monotypy. Status: valid genus and subgenus in Blaptinae: Pedinini: Helopinina. Note: the original combination of the name of the type species, Pimeliagemmata Fabricius, 1801, is a junior primary homonym of Pimeliagemmata Herbst, 1799.

†Amberophlus Novák & Háva, 2019: 128 [M]. Type species: Amberophlusniger Novák & Háva, 2019, by original designation. Status: valid genus in Alleculinae: Cteniopodini. Note: described from Eocene Baltic amber.

Ambigatus Fairmaire, 1892b: 246 [M]. Type species: Ambigatusrufonitens Fairmaire, 1892, by subsequent designation (R. Lucas 1920: 87). Status: valid subgenus of Achanius Erichson, 1847 in Pimeliinae: Evaniosomini.

Amblycara Fairmaire, 1893a: cxlvii [N]. Type species: Amblycarabiskrense Fairmaire, 1893 (= Melancrusalutaceum Fairmaire, 1875), by monotypy. Status: senior synonym of Amblycarenum Gebien, 1910 in Pimeliinae: Tentyriini. Note: junior homonym of Amblycara Bergroth, 1891 [Hemiptera].

Amblycarenum Gebien, 1910a: 77 [N]. Type species [automatic]: Amblycarabiskrense Fairmaire, 1893 (= Melancrusalutaceum Fairmaire, 1875), by monotypy. Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Amblycara Fairmaire, 1893.

Amblychirus Koch, 1956a: 87 [M]. Type species: Trigonopusbrevior Fairmaire, 1897, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Amblycyphus Motschulsky, 1870: 401 [M]. Type species: Amblycyphusasperatus Motschulsky, 1870 (= Centriopterapectoralis Blaisdell, 1921), by monotypy. Status: junior synonym of Cryptoglossa Solier, 1837 in Pimeliinae: Cryptoglossini. Synonymy: Aalbu et al. (1995: 483).

Amblyptera Solier, 1836: 195 [F]. Type species: Pimeliascabrosa Solier, 1836, by subsequent designation (Viñolas and Cartagena 2005: 267). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Amblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 2018: 543 [F]. Type species: Pimeliarugosa Fabricius, 1792 (= Pimeliafairmairii Kraatz, 1865), by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Note: we act as First Revisers and reject the alternative original spelling Amplypteraca, used by Mas-Peinado et al. (2018: 531, 543).

Amblysphagus Fairmaire, 1896a: 16 [M]. Type species: Amblysphaguspachyderus Fairmaire, 1896, by monotypy. Status: valid genus in Blaptinae: Opatrini: Neopachypterina.

Amenophis J. Thomson, 1858: 93 [M]. Type species: Amenophisfairmairei J. Thomson, 1858, by subsequent designation (Gebien 1941: 342). Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Ametrocera Fåhraeus, 1870: 260 [F]. Type species: Ametroceraaurita Fåhraeus, 1870, by subsequent designation (R. Lucas 1920: 88). Status: valid genus in Blaptinae: Pedinini: Helopinina.

Amiantus Fåhraeus, 1870: 279 [M]. Type species: Amiantusgibbosus Fåhraeus, 1870, by subsequent designation (Haag-Rutenberg 1871: 45). Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Amicrodera Kaszab, 1966: 282, 292 [F]. Type species: Microderalindbergi Kaszab, 1966, by original designation. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Ammidanemia Reitter, 1904: 132 [F]. Type species: Anemiafausti Solsky, 1882 (= Cheirodesbrevicollis Wollaston, 1864), by subsequent designation (G.S. Medvedev 1990: 212). Status: junior synonym of Pseudanemia Wollaston, 1864 in Tenebrioninae: Melanimonini. Synonymy: Ardoin (1971: 359, through placement of the type species in the subgenusPseudanemia Wollaston, 1864).

Ammidium Erichson, 1843: 250 [N]. Type species: Ammidiumciliatum Erichson, 1843, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Ammobius Guérin-Méneville, 1844: 121 [M]. Type species: Ammobiusrufus Guérin-Méneville, 1844, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6); the original combination of the name of the type species has been given as “Trachyscelisrufus Guérin-Méneville, 1844” in recent literature; however, we use the combination Ammobiusrufus here as it was used by Guérin-Méneville (1844: 560) in his ‘Table alphabétique des genres, sous-genres et espèces figurés ou décrits dans l’iconographie du règne animal. Insectes’.

Ammocera Borchmann, 1941a: 5 [F]. Type species: Ammoceraunicolor Borchmann, 1941, by original designation. Status: valid subgenus of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina.

Ammodoides Lesne, 1915: 227, 233 [M]. Type species: Arthrodeislateripunctatus Fairmaire, 1890, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Ammodonus Mulsant & Rey, 1859a: 141, 143 [M]. Type species: Opatrumfossor J.L. LeConte, 1847, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina. Note: transferred from Ammobiina by Lumen et al. (2020: 341).

Ammogiton Peyerimhoff, 1920: 326 [M]. Type species: Ammogitonviberti Peyerimhoff, 1920, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Ammophorus Guérin-Méneville, 1831a: pl. 4 [M]. Type species: Ammophorusperuvianus Guérin-Méneville, 1831, by monotypy. Status: valid genus in Tenebrioninae: Scotobiini.

Ammophtorus Lacordaire, 1859a: 284 [M]. Type species [automatic]: Ammobiusrufus Guérin-Méneville, 1844, by monotypy. Status: junior synonym of Ammobius Guérin-Méneville, 1844 in Blaptinae: Opatrini: Ammobiina. Note: unnecessary replacement name for Ammobius Guérin-Méneville, 1844.

Ammotrypes Fairmaire, 1879a: 194 [M]. Type species: Ammotrypescrenulicollis Fairmaire, 1879, by monotypy. Status: junior synonym of Eurycaulus Fairmaire, 1868 in Blaptinae: Opatrini: Sclerina. Synonymy: Grimm (2005: 9); Löbl and Smetana (2010: 33).

Ammozoides Kaszab, 1979a: 91 [M]. Type species: Ammozoumvalidicorne Reitter, 1900, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Ammozoum Semenov, 1891: 352 [N]. Type species: Ammozoumhyalinum Semenov, 1891, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Amnodeis Miller, 1858: 117 [M]. Type species: Amnodeisgrandis Miller, 1858, by subsequent designation (R. Lucas 1920: 90). Status: valid genus in Pimeliinae: Erodiini.

Amorphochirus Gebien, 1904a: 339 [M]. Type species: Pycnocerushercules Fairmaire, 1884, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Amorphopoda Fåhraeus, 1870: 320 [F]. Type species: Amorphopodaelateroides Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Amphianax Bates, 1873e: 350 [M]. Type species: Amphianaxsubcoriaceus Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Amphidora Eschscholtz, 1829: 9 [F]. Type species: Amphidoralittoralis Eschscholtz, 1829, by monotypy. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini. Note: Doyen and Lawrence (1979: 367) synonymized this genus with Eleodes Eschscholtz, 1829; however, Smith and Johnston in Bousquet et al. (2018: 143) used Amphidora as a valid subgenus of Eleodes.

Amphithrix Español, 1952b: 306 [F]. Type species: Amphithrixpeyerimhoffi Español, 1952, by original designation. Status: senior synonym of Amphithrixoides Bouchard & Löbl, 2008 in Blaptinae: Opatrini: Ammobiina. Note: junior homonym of Amphithrix Ragonot, 1893 [Lepidoptera].

Amphithrixoides Bouchard & Löbl, 2008: 39 [M]. Type species [automatic]: Amphithrixpeyerimhoffi Español, 1952, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: replacement name for Amphithrix Español, 1952.

Anacardiosis Endrödy-Younga, 1986: 210, 221 [F]. Type species: Cardiosishamiltonuli Koch, 1969, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Anachayus Bouchard & Bousquet, new replacement name [M]. Type species [automatic]: Chemolanusvillosipes Fairmaire, 1884, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Chatanayus Ardoin, 1957.

Anacycus Fairmaire, 1896a: 33 [M]. Type species: Dietysusnavicularis Fairmaire, 1894, by subsequent designation (Löbl et al. 2008: 215). Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Blair (1929a: 245; with Elixota Pascoe, 1866, a junior synonym of Amarygmus Dalman, 1823).

Anadischidus Kolbe, 1897a: 241 [M]. Type species: Nyctobatesiphthinoides Quedenfeldt, 1885, by monotypy. Status: junior synonym of Deriles Motschulsky, 1872 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1911a: 441).

Anaedes Agassiz, 1846b: 20 [M]. Type species [automatic]: Anaeduspunctatissimus Blanchard, 1842, by monotypy. Status: junior synonym of Anaedus Blanchard, 1842 in Lagriinae: Goniaderini. Note: unjustified emendation of Anaedus Blanchard, 1842, not in prevailing usage.

Anaedus Blanchard, 1842: pl. 14 [M]. Type species: Anaeduspunctatissimus Blanchard, 1842, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Anamenederes Koch, 1954a: 68 [M]. Type species: Menederesdigitatus Koch, 1954, by original designation. Status: valid subgenus of Menederes Solier, 1848 in Blaptinae: Platynotini: Eurynotina.

Anamphidora Casey, 1924: 330 [F]. Type species: Anamphidoraparvula Casey, 1924, by original designation. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Anarmostodera Fairmaire, 1897a: 114 [F]. Type species: Anarmostoderacrassicornis Fairmaire, 1897, by monotypy. Status: valid genus in Tenebrioninae: Praeugenini.

Anatolica Eschscholtz, 1831: 5, 7 [F]. Type species: Tentyriasubquadrata Tauscher, 1812, by subsequent designation (Gebien 1937a: 599). Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Anatrum Reichardt, 1936: 84, 209 [N]. Type species: Anatrumsongoricum Reichardt, 1936, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Anausis Bates, 1873e: 355 [F]. Type species: Anausismacleayi Bates, 1873, by monotypy. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 490).

Anaxius Fåhraeus, 1870: 307 [M]. Type species: Anaxiusobesus Fåhraeus, 1870, by monotypy. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Anaxo Bates, 1868: 272 [F]. Type species: Anaxobrevicornis Bates, 1868, by monotypy. Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324, with Aethyssius Pascoe, 1863, a junior synonym of Lepturidea Fauvel, 1862).

Anchomma J.L. LeConte, 1858a: 63 [N]. Type species: Anchommacostatum J.L. LeConte, 1858, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina. Note: new placement [RLA], previously included in Pimeliinae: Anepsiini; genus originally described in Tenebrionoidea: Zopheridae: Colydiinae.

Anchophthalmops Koch, 1956a: 173 [M]. Type species: Anchophthalmopsbrevipleurum Koch, 1956, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Anchophthalmus Gerstaecker, 1854: 533 [M]. Type species: Anchophthalmussilphoides Gerstaecker, 1854, by subsequent designation (Gebien 1938a: 298). Status: valid genus and subgenus in Blaptinae: Platynotini: Platynotina.

Ancylopoma Pascoe, 1871: 354 [N]. Type species: Ancylopomapunctigerum Pascoe, 1871, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: see Bouchard and Bousquet (2020a: 102) for comments regarding the placement of this genus and its associated family-group name.

Andocamaria Masumoto, 1993b: 230, 232 [F]. Type species: Campsiomorphaformosana Pic, 1930, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Andremiopsis Chatanay, 1913a: 406 [F]. Type species: Andremiopsiscostata Chatanay, 1913, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Andremius Fairmaire, 1903b: 364 [M]. Type species: Andremiuscrispatus Fairmaire, 1903, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Andrimus Casey, 1891: 155 [M]. Type species: Cteniopusmurrayi J.L. LeConte, 1866, by subsequent designation (R. Lucas 1920: 96). Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Androchirus J.L. LeConte, 1862: 244 [M]. Type species: Cistelafuscipes Melsheimer, 1846 (= Cistelaerythropa W. Kirby, 1837), by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Androsus Gebien, 1921a: 325, 386 [M]. Type species: Chariothecaviolacea Pascoe, 1887, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Anebacis Peyerimhoff, 1927: 4, 15 [F]. Type species: Tentyriacribricollis Fairmaire, 1875, by subsequent designation (Löbl et al. 2008a: 40). Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Anectus Horn, 1867: 399 [M]. Type species: Anectusvestitus Horn, 1867, by monotypy. Status: valid genus in Pimeliinae: Branchini.

Anemia Laporte, 1840: 218 [F]. Type species: Anemiagranulata Laporte, 1840, by monotypy. Status: junior synonym of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini. Synonymy: Marseul (1863: 183); Spilman (1973: 41).

Anemiadena Bouchard & Bousquet, new subgenus [F]. Type species: Anemiaconvexa Gestro, 1881, by present designation. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini. Note: Ardoin (1971: 362, 402) introduced the new subgenus name Anemiadena for two nominal species, but unfortunately did not designate a type species; the fact that Anemiaconvexa Gestro, 1881 was listed as the “type” of Anemiadena in the Zoological Record for the year 1971 (Anonymous in Commonwealth Institute of Entomology 1975) does not represent a valid type species designation since the nomenclatural act is anonymous (ICZN 1999, Article 14); the subgenusAnemiadena, which has been treated as valid since 1971, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Anemiaconvexa Gestro, 1881 as type species and referring to Ardoin (1971: 362) for the character states that characterise and differentiate Anemiadena.

Anephyctus Fairmaire, 1891b: 257 [M]. Type species: Anephyctushirtulus Fairmaire, 1891, by monotypy. Status: junior synonym of Miltoprepes Gerstaecker, 1871 in Tenebrioninae: Praeugenini. Synonymy: Gridelli (1939: 78, through placement of the type species in Miltoprepes Gerstaecker, 1871).

Anepsius J.L. LeConte, 1851: 147 [M]. Type species: Anepsiusdelicatulus J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Anepsiini.

Anethas Jakobson, 1924: 242 [M]. Type species [automatic]: Pseudethaslongiceps Fairmaire, 1898, by monotypy. Status: valid genus and subgenus in Pimeliinae: Stenosini: Stenosina. Note: replacement name for Pseudethas Fairmaire, 1898.

Angoleantus Koch, 1952a: 128, 133 [M]. Type species: Rhammatodesstriatulus Koch, 1941, by original designation. Status: valid subgenus of Rhammatodes Haag-Rutenberg, 1876 in Pimeliinae: Tentyriini.

Angolositus Koch, 1955c: 448 [M]. Type species: Angolositussadabandeirus Koch, 1955, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Aniara Melsheimer, 1853: 139 [F]. Type species: Ulomapiceum Melsheimer, 1846, by monotypy. Status: senior synonym of Eutochia J.L. LeConte, 1862 in Tenebrioninae: Ulomini. Note: name previously attributed to “Lacordaire, 1859” in the literature; junior homonym of Aniara Hope, 1838 [Coleoptera: Carabidae].

Aniarus Gemminger in Gemminger and Harold, 1870: 1964 [M]. Type species [automatic]: Ulomapiceum Melsheimer, 1846, by monotypy. Status: junior synonym of Eutochia J.L. LeConte, 1862 in Tenebrioninae: Ulomini. Note: unjustified emendation of Aniara Melsheimer, 1853, not in prevailing usage.

Anisocara Gebien, 1925e: 101 [N]. Type species: Anisocaragynandromorphum Gebien, 1925, by monotypy. Status: junior synonym of Platydema Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Synonymy: Schawaller (2004: 5).

Anisocerus Faldermann, 1837: 39 [M]. Type species: Anisocerustristis Faldermann, 1837, by monotypy. Status: senior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Note: junior homonym of Anisocerus Audinet-Serville, 1835 [Coleoptera: Cerambycidae].

Anisophaedis Ando, 1993: 107 [M]. Type species: Anisophaedisohkurai Ando, 1993, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Anisosis Deyrolle, 1867: 81, 232 [F]. Type species: Anisosiscaudata Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Note: the alternative original spelling Urosis, used by Deyrolle (1867: 81), was rejected by Dallas (1868: 265–266) who acted as First Reviser.

Anisostira Borchmann, 1915: 296 [F]. Type species: Anisostiravaricolor Borchmann, 1915 (= Macrolagriarugipennis Lewis, 1896), by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Annamosdara Kaszab, 1941a: 3, 30 [F]. Type species: Annamosdaramultidentata Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Anobriomaia Kaszab, 1941b: 67 [F]. Type species: Anobriomaiasulcata Kaszab, 1941, by original designation. Status: junior synonym of Foochounus Pic, 1921 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 134).

Anodesis Solier, 1834: 508, 594 [F]. Type species: Anodesiscleryi Solier, 1834, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Anognathena Ando in Ando et al., 2017: 148 [F]. Type species: Anognathenaneraida Ando, 2017, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Anomalipus Guérin-Méneville, 1831b: pl. 29 [M]. Type species: Blapsdentipes Fabricius, 1794, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina. Note: nomen protectum (see Bouchard and Bousquet 2020a: 101, 132); unjustified emendation of the original spelling Anomalipes, introduced by Lacordaire (1859a: 257), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1), see Bouchard et al. (2005: 511).

Anommabates Koch, 1956b: 84 [M]. Type species: Anommabatespauliani Koch, 1956, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Anomoearthrum Mäklin, 1867: 482 [N]. Type species: Anomoearthrumdebile Mäklin, 1867, by subsequent designation (Gebien 1948: 540). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gemminger in Gemminger and Harold (1870: 2028).

Anophthalmolamus Ferrer, 1993: 122 [M]. Type species: Anophthalmolamusfuerteventurae Ferrer, 1993, by original designation. Status: valid genus in Tenebrioninae: incertae sedis.

Anopidium Jeannel & Paulian, 1945: 63 [N]. Type species: Anopidiumelgonicum Jeannel & Paulian, 1945, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.Note: originally described in Tenebrionoidea: Zopheridae: Colydiinae.

Anotoma Borchmann, 1936: 18, 81 [F]. Type species: Chrysolagriaheynei Borchmann, 1915, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Antennoluprops Schawaller, 2007: 30 [M]. Type species: Antennolupropsbremeri Schawaller, 2007, by original designation. Status: valid genus in Lagriinae: Lupropini.

Anteros Laporte, 1840: 235 [M]. Type species: Helopschalibeus Rossi, 1790 (= Tenebriocaeruleus Linnaeus, 1758), by subsequent designation (Iablokoff-Khnzorian 1964: 303). Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Erichson in Agassiz (1846a: 12). Note: junior homonym of Anteros Hübner, 1819 [Lepidoptera].

Anthracias Dejean, 1834: 205 [M]. Type species: Ulomacornutum Fischer, 1823, by monotypy. Status: junior synonym of Cryphaeus Klug, 1833 in Tenebrioninae: Toxicini: Toxicina. Synonymy: Gebien (1921b: 238).

†Anthracohelops Haupt, 1950: 114, 128 [M]. Type species: Anthracohelopsgigas Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Anthracosomus Agassiz, 1846b: 26 [M]. Type species [automatic]: Anthrasomuschevrolatii Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Anthrasomus Guérin-Méneville, 1834 in Pimeliinae: Praociini. Note: unjustified emendation of Anthrasomus Guérin-Méneville, 1834, not in prevailing usage.

Anthracula Fairmaire, 1897c: 236 [F]. Type species: Anthraculalatifrons Fairmaire, 1897, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Anthrasomus Guérin-Méneville, 1834: 32 [M]. Type species: Anthrasomuschevrolatii Guérin-Méneville, 1834, by monotypy. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini.

Anthrenopsis Koch, 1950c: 74 [F]. Type species: Platydemascriptipennis Fairmaire, 1875 (= Basidesziczac Motschulsky, 1873), by original designation. Status: valid subgenus of Ellipsodes Wollaston, 1854 in Diaperinae: Crypticini.

Anticlia Gistel, 1848a: x, 125 [F]. Type species [automatic]: Opatrumorientale Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: junior synonym of Sclerum Dejean, 1834 in Blaptinae: Opatrini: Sclerina. Note: unnecessary replacement name for Sclerum Dejean, 1834.

Antimachus Gistel, 1829: 1055 [M]. Type species: Phalariafurcifera Dalman, 1821, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Antofagapraocis Flores, 2000b: 62, 68 [M]. Type species: Falsopraocissubnudus Kulzer, 1959, by original designation. Status: valid genus in Pimeliinae: Praociini.

Antoineius Koch, 1948: 418 [M]. Type species: Micrositusjeanneli Koch, 1945 (= Melambiusinermis Antoine, 1942), by original designation. Status: valid subgenus of Otinia Antoine, 1942 in Blaptinae: Dendarini: Melambiina.

Aoupinia Matthews, 2003a: 441 [F]. Type species: Aoupiniapseudohelea Matthews, 2003, by original designation. Status: valid genus in Lagriinae: Adeliini.

Apalmia Fairmaire, 1896a: 60 [F]. Type species: Apalmiacerambycina Fairmaire, 1896, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Apasis Pascoe, 1869: 139 [F]. Type species: Apasishowittii Pascoe, 1869, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Apatelus Mulsant & Rey, 1859a: 88, 91 [M]. Type species: Apatelushopei Mulsant & Rey, 1859, by monotypy. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Carter (1926: 123).

Apatopsis Semenov, 1891: 368 [F]. Type species: Apatopsisgrombczewskii Semenov, 1891, by subsequent designation (R. Lucas 1920: 107). Status: valid genus in Pimeliinae: Pimeliini.

Apelina Saha, 1988: 429 [F]. Type species: Apelinakeralaensis Saha, 1988, by original designation. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer and Lillig (2014: 12).

Apellatus Pascoe, 1863a: 45 [M]. Type species: Apellatuslateralis Pascoe, 1863 (= Apellatusamoenus Pascoe, 1866), by monotypy. Status: junior synonym of Euomma Boheman, 1858 in Alleculinae: Alleculini: Alleculina. Synonymy: Pascoe (1866a: 491).

Apentanes Reitter, 1914a: 45, 51 [M]. Type species: Arthrodeisoccidentalis Fairmaire, 1868, by subsequent designation (Gebien 1937a: 536). Status: valid subgenus of Arthrodeis Solier, 1834 in Pimeliinae: Erodiini.

Apentanodes Reitter, 1914a: 46, 53 [M]. Type species: Arthrodeisglobosus Reiche & Saulcy, 1857, by subsequent designation (Gebien 1937a: 539). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Aphaleria Reitter, 1896a: 235 [F]. Type species: Aphaleriacapnisoides Reitter, 1896 (= Erodiuspygmaeus Fischer, 1821), by monotypy. Status: junior synonym of Bradyus Dejean, 1834 in Tenebrioninae: Dissonomini. Synonymy: Löbl et al. (2008b: 240).

Aphanaspis Wollaston, 1864: 478 [F]. Type species: Pimeliagranulicollis Wollaston, 1864, by subsequent designation (Löbl et al. 2008a: 40). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Aphanotus J.L. LeConte, 1862: 233 [M]. Type species: Eulabisbrevicornis J.L. LeConte, 1859, by original designation. Status: valid subgenus of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini.

Aphectus Carter, 1926: 127, 128 [M]. Type species [automatic]: Hectusanthracinus Pascoe, 1869, by monotypy. Status: junior synonym of Olisthaena Erichson, 1842 in Tenebrioninae: Heleini: Cyphaleina. Note: unnecessary replacement name for Hectus Pascoe, 1869.

Aphelus Gebien, 1921b: 62, 78 [M]. Type species: Aphelussimplicicollis Gebien, 1921, by subsequent designation (Gebien 1941: 341). Status: junior synonym of Argobrachium Fairmaire, 1899 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1961b: 31).

Aphrotus Péringuey, 1904: 252 [M]. Type species [automatic]: Xenustricorniger Péringuey, 1899, by monotypy. Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Xenus Péringuey, 1899.

Aphtora Bates, 1872b: 265 [F]. Type species: Aphtorarufipes Bates, 1872, by monotypy. Status: valid genus in Phrenapatinae: incertae sedis. Note: Matthews and Lawrence (2019: 623) placed this genus in the subfamily Phrenapatinae (tribal position uncertain).

Aphyllocerus Fairmaire, 1881a: 348 [M]. Type species: Aphyllocerusdecipiens Fairmaire, 1881, by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1921a: 409).

Apistocerus Fairmaire, 1899a: 78 [M]. Type species: Apistoceruswasmanni Fairmaire, 1899, by monotypy. Status: valid subgenus of Rhyzodina Chevrolat, 1873 in Tenebrioninae: Rhysopaussini.

Apithesis C.O. Waterhouse, 1881: 476 [F]. Type species: Apithesisobesa C.O. Waterhouse, 1881, by monotypy. Status: junior synonym of Clitobius Mulsant & Rey, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Purchart and Kamiński (2017: 143).

Aplanasida Reitter, 1917a: 11, 30 [F]. Type species: Asidabrevicosta Solier, 1836, by monotypy. Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Apocrypha Eschscholtz, 1831: 13 [F]. Type species: Apocryphaanthicoides Eschscholtz, 1831, by monotypy. Status: valid genus in Tenebrioninae: Apocryphini.

Apocryphodes Matthews, 1998: 704, 765 [M]. Type species: Apocryphodesthompsoni Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Apodemus Fåhraeus, 1870: 293 [M]. Type species: Anomalipusplanus Fåhraeus, 1870, by subsequent designation (Iwan 2002c: 234). Status: junior synonym of Anomalipus Guérin-Méneville, 1831 in Blaptinae: Platynotini: Platynotina. Synonymy: Gebien (1938a: 409). Note: junior homonym of Apodemus Kaup, 1829 [Mammalia].

Apolites Jacquelin du Val, 1861: 324 [M]. Type species: Helopsmucoreus Waltl, 1838, by monotypy. Status: senior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Synonymy: Allard (1876b: ciii). Note: junior homonym of Apolites Sundevall, 1835 [Aves].

Apomestris Bates, 1873e: 357 [M]. Type species: Apomestriswestwoodi Bates, 1873, by monotypy. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1926: 119, with Altes Pascoe, 1869, a junior synonym of Cyphaleus Westwood, 1841).

Apostethus Pascoe, 1882: 27 [M]. Type species: Apostethusterrenus Pascoe, 1882, by monotypy. Status: junior synonym of Adelodemus Haag-Rutenberg, 1878 in Lagriinae: Adeliini. Synonymy: Carter (1926: 131).

Apristopus Kolbe, 1903: 167 [M]. Type species: Priosceliscrassicornis Westwood, 1842, by monotypy. Status: junior synonym of Calostegia Westwood, 1843 in Lagriinae: Pycnocerini. Synonymy: Gebien (1904a: 169).

Aprosphaena Reitter, 1916c: 140, 142 [F]. Type species: Aprosphaenaadriani Reitter, 1916 (= Tageniastriatopunctata Wiedemann, 1821), by subsequent designation (Löbl et al. 2008a: 40). Status: junior synonym of Stenosida Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Blair (1935a: 103).

Apsena J.L. LeConte, 1862: 228 [F]. Type species: Eulabispubescens J.L. LeConte, 1851, by original designation. Status: valid genus in Tenebrioninae: Eulabini.

Apsheronellus Bogatchev, 1967: 157 [M]. Type species: Apsheronellusarenarius Bogatchev, 1967, by original designation. Status: valid genus in Blaptinae: Pedinini: Leichenina.

Apsida Lacordaire, 1859a: 309 [F]. Type species: Apsidachrysomelina Lacordaire, 1859, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Aptereucyrtus Gebien, 1922a: 477 [M]. Type species: Aptereucyrtushemichalceus Gebien, 1922, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Aptereutochia Kaszab, 1980a: 190 [F]. Type species: Eutochiaaptera Kaszab, 1980, by original designation. Status: valid subgenus of Cenoscelis Wollaston, 1868 in Tenebrioninae: Ulomini. Note: this name was first published by Kaszab (1979a: 92) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article (13.1) and is therefore unavailable from that date.

Aptericula Borchmann, 1937: 219 [F]. Type species: Aptericulanyassensis Borchmann, 1937, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Apterobrachys Kaszab, 1986: 295 [M]. Type species: Apterobrachyswilhelminae Kaszab, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Apteroclitobius Koch, 1960: 391, 404 [M]. Type species: Clitobiusgrandis Fairmaire, 1896, by original designation. Status: junior synonym of Clitobius Mulsant & Rey, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Purchart and Kamiński (2017: 143).

Apterocyphostethe Kaszab, 1962a: 302 [F]. Type species: Himatismuskoltzei Reitter, 1895, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini.

Apterogena Ardoin, 1962a: 67 [F]. Type species: Apterogenacanonnei Ardoin, 1962, by original designation. Status: senior synonym of Genateropa Bouchard & Bousquet, nom. nov. in Stenochiinae: Stenochiini. Note: junior homonym of Apterogena Amyot, 1847 [Hemiptera].

Apteroleprocaulus Kaszab, 1983c: 182 [M]. Type species: Leprocaulusmontanus Kaszab, 1982, by original designation. Status: junior synonym of Hexarhopalus Fairmaire, 1891 in Stenochiinae: Cnodalonini. Synonymy: Bečvář and Purchart (2008: 39).

Apteromaia Kulzer, 1952: 719 [F]. Type species: Eucyrtusovipennis Gebien, 1913, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Apteromerus Blair, 1928: 72 [M]. Type species: Opatrinusconvexus Fairmaire, 1849, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Apteromira Weise, 1974: 123 [F]. Type species: Cistelaovulum Kiesenwetter, 1863, by original designation. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Apteronympha Seidlitz, 1898b: 336, 338 [F]. Type species: Lagriarubida Graells, 1858, by subsequent designation (Merkl 2004: 284). Status: valid subgenus of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina.

Apterophenus Gebien, 1921a: 325, 342 [M]. Type species: Apterophenusevanescens Gebien, 1921, by subsequent designation (Gebien 1941: 1142). Status: valid genus in Stenochiinae: Cnodalonini.

Apteroseriscius Koch, 1950c: 64 [M]. Type species: Pseudoserisciusespanoli Koch, 1950, by original designation. Status: valid genus in Diaperinae: Crypticini.

Apterosphaeria Koch, 1950b: 298 [F]. Type species: Derosphaeriusrugifrons Fairmaire, 1888, by original designation. Status: valid subgenus of Derosphaerius Westwood, 1881 in Pimeliinae: Tentyriini.

Apterotarpela Kaszab, 1954: 262 [F]. Type species: Apterotarpelaklapperichi Kaszab, 1954, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Apterotheca Gebien, 1921a: 348 [F]. Type species: Chariothecabesti Blackburn, 1894, by subsequent designation (Gebien 1942a: 338). Status: valid genus in Stenochiinae: Cnodalonini.

Apterozidalus Ardoin, 1965b: 1315 [M]. Type species: Apterozidalusroyi Ardoin, 1965, by original designation. Status: junior synonym of Zidalus Mulsant & Rey, 1853 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (1995b: 362).

Apteruleda Gebien, 1928: 134 [F]. Type species: Apteruledauncipes Gebien, 1928, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Apteruloma Gebien, 1928: 149 [N]. Type species: Apterulomamagnum Gebien, 1928, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Apterulomoides Kaszab, 1982d: 279 [M]. Type species: Ulomarotundipenne Kaszab, 1982, by monotypy. Status: valid subgenus of Uloma Dejean, 1821 in Tenebrioninae: Ulomini.

Aptila Fåhraeus, 1870: 258 [F]. Type species: Aptilacostata Fåhraeus, 1870, by subsequent designation (R. Lucas 1920: 115). Status: valid genus in Blaptinae: Pedinini: Helopinina.

Arabammobius Grimm & Lillig, 2020: 307 [M]. Type species: Ammobiustarsalis Grimm, 2012, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Arabcynaeus Schawaller, 2009a: 164 [M]. Type species: Arabcynaeusbremeri Schawaller, 2009, by original designation. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Araeopselaphus Gebien, 1921b: 10 [M]. Type species: Araeopselaphusmyrmecophilus Gebien, 1921, by monotypy. Status: valid genus in Diaperinae: Crypticini.

Araeoschizus J.L. LeConte, 1851: 138 [M]. Type species: Araeoschizuscostipennis J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Araeoschizina.

Araucaricola Lea, 1929: 218 [F]. Type species: Araucaricolaebenina Lea, 1929, by monotypy. Status: junior synonym of Iscanus Fauvel, 1904 in Lagriinae: Lupropini. Synonymy: Kaszab (1982a: 59).

Archaeoglenes Broun, 1893a: 188 [M]. Type species: Archaeoglenescostipennis Broun, 1893, by monotypy. Status: valid genus in Phrenapatinae: Archaeoglenini.

Archasida Wilke, 1922: 261 [F]. Type species: Afrasidainnotata Wilke, 1922, by original designation. Status: valid subgenus of Afrasida Wilke, 1922 in Pimeliinae: Asidini.

Archeophthora Kaszab, 1978a: 166 [F]. Type species: Brachycilibetasmanicum Carter, 1919, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Archinamaqua Schawaller, 2012b: 80 [F]. Type species: Archinamaqualyleae Schawaller, 2012 (= Menederopsisconstricta Koch, 1954), by original designation. Status: junior synonym of Menederopsis Koch, 1954 in Blaptinae: Platynotini: Eurynotina. Synonymy: Kamiński and Schawaller (2019: 293).

Archinamibia Koch, 1952a: 157 [F]. Type species: Archinamibiapeezi Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Arcothymus Pascoe, 1866a: 476 [M]. Type species: Arcothymuscoenosus Pascoe, 1866 (= Adeliumtriste Montrouzier, 1860), by monotypy. Status: valid genus in Lagriinae: Adeliini. Note: as mentioned by Matthews and Bouchard (2008: 350) the type species was originally described from Australia in error; the genus Arcothymus Pascoe, 1866 is endemic to New Caledonia.

Arctylus Dejean, 1834: 180 [M]. Type species: Praocispentagonus Lacordaire, 1830, by monotypy. Status: junior synonym of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Synonymy: Bousquet and Bouchard (2013a: 43).

Ardamimicus Smith, 2013: 601 [M]. Type species: Ardamimicuscognatoi Smith, 2013, by original designation. Status: valid genus in Pimeliinae: Asidini.

Ardeleodes Blaisdell, 1937: 128 [M]. Type species: Eleodestibialis Blaisdell, 1909, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini. Note: the alternative original spelling Arpeleodes, used by Blaisdell (1937: 128), was rejected by Gebien (1938a: 63) who acted as First Reviser.

Ardelio Gistel, 1848a: xi [M]. Type species [automatic]: Ulomacornutum Fischer, 1823, by monotypy. Status: junior synonym of Cryphaeus Klug, 1833 in Tenebrioninae: Toxicini: Toxicina. Note: unnecessary replacement name for Anthracias Dejean, 1834.

Ardoinia Kaszab, 1969a: 249 [F]. Type species: Ardoiniadiaclinoides Kaszab, 1969, by original designation. Status: valid genus in Tenebrioninae: Alphitobiini.

Ardoinia Özdikmen, 2005: 202 [F]. Type species [automatic]: Orghidaniatorrei Ardoin, 1977, by monotypy. Status: senior synonym of Spelaebiosis Bousquet & Bouchard, 2018 in Tenebrioninae: Triboliini. Note: replacement name for Orghidania Ardoin, 1977; junior homonym of Ardoinia Kaszab, 1969 [Coleoptera: Tenebrionidae: Tenebrioninae: Alphitobiini].

Ardoiniellus Schawaller, 2013a: 138 [M]. Type species: Ardoiniellusmontanus Schawaller, 2013, by original designation. Status: valid genus in Lagriinae: Lupropini.

Arenacara Penrith, 1979: 43, 46 [N]. Type species: Stenocarabrunnipes Haag-Rutenberg, 1877, by original designation. Status: valid subgenus of Stenocara Solier, 1835 in Pimeliinae: Adesmiini.

Arenoblaps G.S. Medvedev, 1999b: 400 [F]. Type species: Blaps hiemalis Semenov-Tjan-Shansky & Bogatchev, 1940, by original designation. Status: valid subgenus of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina.

Argasidus Péringuey, 1899: 251 [M]. Type species: Argasidussquamosus Péringuey, 1899, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Argenticrinis Louw, 1979: 99, 100 [M]. Type species: Argenticrinishaackei Louw, 1979 (= Psammodeslossowi Koch, 1952), by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Argobrachium Fairmaire, 1899d: 216 [N]. Type species: Argobrachiumimpressifrons Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Argoporis Horn, 1870: 325 [F]. Type species: Cerenopuscostipennis J.L. LeConte, 1851, by subsequent designation (Gebien 1937a: 797). Status: valid genus in Tenebrioninae: Cerenopini. Note: as pointed out by Bousquet et al. (2018: 97, 179), evidence shows that Threnus Motschulsky, 1870 was published before the currently accepted valid name Argoporis Horn, 1870; as Threnus Motschulsky, 1870 was used as valid after 1899 (e.g., Leng 1920: 224), reversal of precedence cannot be used to conserve usage of Argoporis Horn, 1870; an application to the ICZN is necessary to conserve usage of Argoporis Horn, 1870.

Argutiolana Robiche, 2001: 191 [F]. Type species: Argutiolanamaguini Robiche, 2001, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: transferred from Tenebrioninae: Tenebrionini by Robiche (2019a: 97).

Argyradelpha G.S. Medvedev, 2005a: 306 [F]. Type species: Argyradelphalopatini G.S. Medvedev, 2005, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Argyrophana Semenov, 1889: 222, 224 [F]. Type species: Argyrophanadeserti Semenov, 1889, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Ariarathus Fairmaire, 1891e: ccxi [M]. Type species: Ariarathusulomoides Fairmaire, 1891 (= Tenebrioatronitens Fairmaire, 1891), by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Armalia Casey, 1907: 289, 330 [F]. Type species: Emmenastustexanus J.L. LeConte, 1866, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Armenohelops Nabozhenko, 2002a: 42 [M]. Type species: Armenohelopsarmeniacus Nabozhenko, 2002, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Armigena Bouchard & Bousquet, new subgenus [F]. Type species: Nesogenatestaceipes Fairmaire, 1868, by present designation. Status: valid subgenus of Nesogena Mäklin, 1863 in Tenebrioninae: Praeugenini. Note: first proposed by Froussart (1961: 60) without type species designation; the subgenusArmigena, which is currently used as valid, is therefore unavailable (ICZN 1999, Articles 13.3, 16.1); we hereby make the name available by selecting Nesogenatestaceipes Fairmaire, 1868 as type species and referring to Froussart (1961: 60) for the character states that characterise and differentiate Armigena.

Arnoldiola Semenov-Tjan-Shansky & Bogatchev, 1940: 201 [F]. Type species: Arnoldiolapeculiaris Semenov-Tjan-Shansky & Bogatchev, 1940 (= Stalagmopteraruginota Reitter, 1896), by original designation. Status: junior synonym of Stalagmoptera Solsky, 1876 in Pimeliinae: Pimeliini. Synonymy: Löbl et al. (2008b: 168). Note: junior homonym of Arnoldiola Strand, 1928 [Diptera].

Arrhabaeus Champion, 1886: 144 [M]. Type species: Arrhabaeusconvexus Champion, 1886, by monotypy. Status: junior synonym of Dioedus J.L. LeConte, 1862 in Phrenapatinae: Penetini. Synonymy: Kaszab (1977a: 314).

Arrhenoplita W. Kirby, 1837: 235 [F]. Type species: Ips haemorrhoidalis Fabricius, 1787, by original designation. Status: junior synonym of Neomida Latreille, 1829 in Diaperinae: Diaperini: Diaperina. Synonymy: Duponchel and Chevrolat (1841: 157).

Artactes Pascoe, 1868: xii [M]. Type species: Artactesnigritarsis Pascoe, 1868, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Arthrochora Gebien, 1938b: 75 [F]. Type species: Arthrochoraarenicola Gebien, 1938, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Arthroconus Solier, 1851: 238 [M]. Type species: Arthroconuspiceus Solier, 1851 (= Gymnognathusfuscus Solier, 1851), by subsequent designation (Lacordaire 1859a: 67). Status: valid genus in Pimeliinae: Edrotini. Note: the First Reviser (Arthroconus Solier, 1851 versus Gymnognathus Solier, 1851) is Lacordaire (1859a: 67).

Arthrodeis Solier, 1834: 508, 513 [M]. Type species: Arthrodeisrotundatus Solier, 1834, by subsequent designation (Hope 1841: 114). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Arthrodes Agassiz, 1846b: 35 [M]. Type species [automatic]: Arthrodeisrotundatus Solier, 1834, by subsequent designation (Hope 1841: 114). Status: junior synonym of Arthrodeis Solier, 1834 in Pimeliinae: Erodiini. Note: unjustified emendation of Arthrodeis Solier, 1834, not in prevailing usage.

Arthrodibius Lesne, 1915: 227, 235 [M]. Type species: Arthrodeislaxepunctatus Fairmaire, 1884, by subsequent designation (Sharp 1919: 119). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Arthrodinus Reitter, 1900b: 299 [M]. Type species: Erodiusobesus Brullé, 1839, by subsequent designation (Reitter 1914a: 46). Status: valid subgenus of Arthrodeis Solier, 1834 in Pimeliinae: Erodiini.

Arthrodion Lesne, 1915: 227, 234 [N]. Type species: Spyrathusafricanus Fairmaire, 1882, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Arthrodosis Reitter, 1900b: 299 [F]. Type species: Erodiusglobosus Faldermann, 1837, by subsequent designation (R. Lucas 1920: 119). Status: valid genus in Pimeliinae: Erodiini. Note: the original combination of the accepted name of the type species, Erodiusglobosus Faldermann, 1837, is a junior primary homonym of Erodiusglobosus Thunberg, 1787.

Arthrodygmus Reitter, 1914a: 46 [M]. Type species: Arthrodygmusfieberi Reitter, 1914, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Arthrohyalosis Kaszab, 1979a: 75 [F]. Type species: Arthrodosismostofii Pierre, 1975, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Arthrohyalus Koch, 1943a: 490, 503 [M]. Type species: Arthrohyalussarcinipennis Koch, 1943, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Arthromacra W. Kirby, 1837: 238 [F]. Type species: Arthromacradonacioides W. Kirby, 1837 (= Lagriaaenea Say, 1824), by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Arthroplatus Solier, 1851: 246 [M]. Type species: Arthroplatuspallipes Solier, 1851, by monotypy. Status: junior synonym of Acropteryx Gistel, 1831 in Tenebrioninae: Acropteronini. Synonymy: Schaum (1852: 183, with Acropteron Perty, 1832, a junior synonym of Acropteryx Gistel, 1831).

Arturium Koch, 1951: 83 [N]. Type species: Melanolophusater C.O. Waterhouse, 1885, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Artystona Bates, 1874: 104 [F]. Type species: Titaenaerichsonii White, 1846, by subsequent designation (R. Lucas 1920: 120). Status: valid genus in Tenebrioninae: Titaenini. Note: this name was introduced one year earlier by Bates (1873f: 473) but it is unavailable from that date since there is no description nor originally included species (ICZN 1999, Article 12.1).

Arunogria Merkl, 1991: 14 [F]. Type species: Arunogriapubescens Merkl, 1991, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Aryenis Bates, 1868: 309 [F]. Type species: Aryenisrufescens Bates, 1868 (= Statiraunicolor Blanchard, 1843), by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Asbolius Fairmaire, 1902a: 134 [M]. Type species: Asboliusquadricollis Fairmaire, 1902, by monotypy. Status: junior synonym of Rhammatodes Haag-Rutenberg, 1876 in Pimeliinae: Tentyriini. Synonymy: Blair (1935a: 103).

Asbolodes Fairmaire, 1892c: 52 [M]. Type species: Asbolodeshumerosus Fairmaire, 1892, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asbolodomimus Pic, 1921d: 20 [M]. Type species: Asbolodomimussubcarinatus Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asbolus J.L. LeConte, 1851: 129 [M]. Type species: Asbolusverrucosus J.L. LeConte, 1851, by subsequent designation (Aalbu 2005: 721). Status: valid genus in Pimeliinae: Cryptoglossini. Note: the older name Asbolus Voet, 1793 [Coleoptera: Silphidae] was published in a work that did not include consistent application of binominal nomenclature and is therefore unavailable (ICZN 1999, Article 11.4).

Ascalabus Fairmaire, 1893b: 30 [M]. Type species: Ascalabuspedinoides Fairmaire, 1893, by monotypy. Status: senior synonym of Calabosca Fairmaire, 1894 in Stenochiinae: Cnodalonini. Note: junior homonym of Ascalabus Agassiz, 1846 [Pisces].

Ascelosodis Redtenbacher, 1868: 117 [F]. Type species: Ascelosodisserripes Redtenbacher, 1868, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Asemogena Péringuey, 1904: 281 [F]. Type species: Asemogenasimplex Péringuey, 1904, by subsequent designation (R. Lucas 1920: 121). Status: valid genus in Stenochiinae: Stenochiini.

Asialassus Nabozhenko & Ando, 2018: 311 [M]. Type species: Helopscordicollis Marseul, 1876, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Asida Latreille, 1802: 167 [F]. Type species: Opatrumgriseum Fabricius, 1781, by monotypy. Status: valid genus and subgenus in Pimeliinae: Asidini.

Asidelia Fairmaire, 1905: 296 [F]. Type species: Asideliacontracta Fairmaire, 1905, by monotypy. Status: valid genus in Pimeliinae: Praociini.

Asidesthes Fairmaire, 1900c: 246 [F]. Type species: Asidesthesperrieri Fairmaire, 1900, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Asididius Fairmaire, 1869b: 236 [M]. Type species: Asididiuscoquerelii Fairmaire, 1869, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asidina Casey, 1912: 76, 169 [F]. Type species: Pelecyphorusparallelus J.L. LeConte, 1851, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Asidoblaps Fairmaire, 1886d: 342 [F]. Type species: Asidoblapsdavidis Fairmaire, 1886, by subsequent designation (R. Lucas 1920: 122). Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Asidobothris Fairmaire, 1886c: 72 [F]. Type species: Asidobothrisclathrata Fairmaire, 1886, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asidodema Koch, 1958: 139 [F]. Type species: Oncosomaalternicoste Gebien, 1910, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Asidomachla Wilke, 1922: 262 [F]. Type species: Asidabicostata Fåhraeus, 1870, by original designation. Status: junior synonym of Machlomorpha Péringuey, 1899 in Pimeliinae: Asidini. Synonymy: Koch (1962a: 125).

Asidomorpha Koch, 1962a: 121 [F]. Type species: Afrasidaprona Wilke, 1922, by original designation. Status: valid genus in Pimeliinae: Asidini.

Asidopsis Casey, 1912: 77, 185 [F]. Type species: Asidaopaca Say, 1824, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Asiobirus G.S. Medvedev, 1968a: 170, 179 [M]. Type species: Cabirusvalidipes Reitter, 1891, by original designation. Status: junior synonym of Cabirutus Strand, 1929 in Blaptinae: Pedinini: Pedinina. Synonymy: Kamiński and Iwan (2017: 595).

Asiocaedius G.S. Medvedev & Nepesova, 1985: 138 [M]. Type species [automatic]: Pseudocaediuskiseritzkii G.S. Medvedev, 1966, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: replacement name for Pseudocaedius G.S. Medvedev, 1966.

Asiomira Dubrovina, 1973: 367 [F]. Type species: Isomiraophthalmica Seidlitz, 1896, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: changed from subgenus of Isomira Mulsant, 1856 to valid genus by Novák (2016a: 179).

Asiopus Sharp, 1892b: 43 [M]. Type species: Asiopusopatroides Sharp, 1892, by monotypy. Status: valid genus in Lagiinae: incertae sedis. Note: this little-known genus has been treated as a member of the subfamily Lagriinae (e.g., Fery 2013: 66).

Asiris Motschulsky, 1872: 24 [F]. Type species: Asirisangulicollis Motschulsky, 1872 (= Tenebrioaustralis Boisduval, 1835), by original designation. Status: junior synonym of Meneristes Pascoe, 1869 in Tenebrioninae: Heleini: Asphalina. Synonymy: C.O. Waterhouse (1876: 288).

Asopidiopsis Kaszab, 1955a: 511, 515 [F]. Type species: Asopidiopsisovalis Kaszab, 1955, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Asopis Haag-Rutenberg, 1878: 104 [F]. Type species: Asopissuavis Haag-Rutenberg, 1878, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Asphaltesthes Kraatz, 1865: 80, 181 [F]. Type species: Mesostenacostata Erichson, 1843, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Asphalus Pascoe, 1868: xii [M]. Type species: Asphalusebeninus Pascoe, 1868, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Asphalina.

Asphena Semenov, 1889: 218 [F]. Type species: Asphenakomarowi Semenov, 1889, by monotypy. Status: junior synonym of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini. Synonymy: Reitter (1916c: 141).

Aspidius Mulsant & Rey, 1859: 123 [M]. Type species: Blapspunctata Fabricius, 1792, by subsequent designation (Bousquet et al. 2018: 195). Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Champion (1885: 124).

Aspidocephalus Motschulsky, 1839: 63 [M]. Type species: Aspidocephalusdesertus Motschulsky, 1839, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina. Note: unjustified emendation of the original spelling Aspicephalus, introduced by Lacordaire (1859a: 107), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Aspidolobus Redtenbacher, 1868: 118 [M]. Type species: Aspidolobuspiliger Redtenbacher, 1868, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Aspidosoma Agassiz, 1846b: 36, 37 [N]. Type species [automatic]: Aspisomafulvipenne Duponchel & Chevrolat, 1841, by original designation. Status: junior synonym of Anaedus Blanchard, 1842 in Lagriinae: Goniaderini. Note: unjustified emendation of Aspisoma Duponchel & Chevrolat, 1841, not in prevailing usage.

Aspidosternum Mäklin, 1867: 500 [N]. Type species: Tenebriocyaneus Fabricius, 1794, by monotypy. Status: junior synonym of Metallonotus Gray, 1832 in Lagriinae: Pycnocerini. Synonymy: Alluaud (1889: xlvi).

Aspila Fåhraeus, 1870: 251 [F]. Type species: Aspilabicostata Fåhraeus, 1870, by monotypy. Status: senior synonym of Prunaspila Koch, 1950 in Pimeliinae: Adelostomini. Note: junior homonym of Aspila Stephens, 1834 [Lepidoptera].

Aspilomorpha Koch, 1952b: 33, 101 [F]. Type species: Aspilomorphamediolobata Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Aspisoma Duponchel & Chevrolat, 1841: 240 [N]. Type species: Aspisomafulvipenne Duponchel & Chevrolat, 1841, by original designation. Status: senior synonym of Anaedus Blanchard, 1842 in Lagriinae: Goniaderini. Synonymy: Lacordaire (1859a: 396). Note: junior homonym of Aspisoma Laporte, 1833 [Coleoptera: Lampyridae].

Astalbus Fairmaire, 1900d: 484 [M]. Type species: Astalbusscrobicollis Fairmaire, 1900, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Astathmetus Bates, 1874: 23 [M]. Type species: Astathmetusalienus Bates, 1874, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: this name was introduced one year earlier by Bates (1873f: 472) but it is unavailable from that date since there is no description nor originally included species (ICZN 1999, Article 12.1).

Astatira Borchmann, 1921: 218, 322 [F]. Type species: Astatirahumeralis Borchmann, 1921, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Asthenochirus Fairmaire, 1885a: viii [M]. Type species: Asthenochirusnigropunctatus Fairmaire, 1885, by subsequent designation (R. Lucas 1920: 123). Status: valid genus in Tenebrioninae: Amarygmini. Note: redescribed as new by Fairmaire (1887b: 298).

Asthenopoda Chatanay, 1915a: 527, 544 [F]. Type species: Asthenopodafragilis Chatanay, 1915, by original designation. Status: junior synonym of Psilonesogena Bates, 1879 in Stenochiinae: Stenochiini. Synonymy: De Moor (1970: 9).

Asticostena Fairmaire, 1897c: 228 [F]. Type species: Asticostenaalternata Fairmaire, 1897, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Astorthocnemis Lillig & Pavlíček, 2002: 98 [F]. Type species: Storthocnemissaudita Koch, 1965, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Astrotus J.L. LeConte, 1858b: 19 [M]. Type species: Microschatiacontorta J.L. LeConte, 1853, by original designation. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Asyleptus Péringuey, 1896: 179 [M]. Type species: Asyleptusfumosus Péringuey, 1896, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Asyrmatus Canzoneri, 1959: 149 [M]. Type species: Helopspiceus Sturm, 1826, by original designation. Status: junior synonym of Pystelops Gozis, 1910 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: new synonym [YB]. Note: Pystelops Gozis, 1910 has been forgotten in the literature; its type species is currently included in the subgenusAsyrmatus Canzoneri, 1959 and for that reason Canzoneri’s name is considered a junior synonym of Pystelops; the original combination of the accepted name of the type species, Helopspiceus Sturm, 1826, is a junior primary homonym of Helopspiceus G.-A. Olivier, 1792.

Atahualpina Español, 1960: 113 [F]. Type species: Atahualpinaperuviana Español, 1960 (= Phaleriasubparalella Chevrolat, 1878), by original designation. Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Synonymy: Triplehorn (1991: 258).

Atasthalomorpha Miyatake, 1964: 68, 77 [F]. Type species: Atasthalusdentifrons Lewis, 1894, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Atasthalus Pascoe, 1871: 348 [M]. Type species: Atasthalusspectrum Pascoe, 1871, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Athrodactyla Klug, 1833: 90 [F]. Type species: Athrodactylaelongata Klug, 1833, by subsequent designation (Hope 1841: 126). Status: valid genus in Tenebrioninae: Tenebrionini.

Atlasion Koch, 1948: 427 [N]. Type species: Micrositusbedeli Escalera, 1914, by original designation. Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina. Note: the First Reviser (Atlasion Koch, 1948 versus Megatlasion Koch, 1948) is Antoine (1957: 351).

Atlasotaurus Bouyon, 2011: 464 [M]. Type species: Omophlusmaroccanus P.H. Lucas, 1846, by original designation. Status: valid subgenus of Heliotaurus Mulsant, 1856 in Alleculinae: Cteniopodini.

Atoichus Carter, 1915a: 72 [M]. Type species: Licymniusbicolor Blackburn, 1893, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Atoreuma Gebien, 1941: 1132 [N]. Type species [automatic]: Toreumacupreum Carter, 1913, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: replacement name for Eutoreuma Carter, 1914.

Atrachyderma Skopin, 1962: 228 [N]. Type species: Pimeliasetosa Faldermann, 1832, by original designation. Status: valid subgenus of Trachyderma Latreille, 1828 in Pimeliinae: Pimeliini. Note: Pimeliasetosa, the type species, has been attributed to Fischer in the literature; the species was described as “Pimeliasetosa Fald.” which, from the context, indicates that Faldermann was responsible for the description of the species.

Atractus Boisduval, 1835: 283 [M]. Type species: Atractusviridis Boisduval, 1835, by subsequent designation (Duponchel and Chevrolat 1841: 312). Status: senior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324). Note: junior homonym of Atractus Wagler, 1828 [Reptilia].

Atrocrates Koch, 1956a: 82 [M]. Type species: Trigonopusplatyderus Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Atrocrypticanus Iwan, 1999a: 73 [M]. Type species: Atrocrypticanusfraternus Iwan, 1999, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Atropsorodes Ardoin, 1963a: 88 [M]. Type species: Atropsorodesbarnardi Ardoin, 1963, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: name published earlier by Ardoin (1962b: 969) without fixation of a type species in the original publication (ICZN 1999, Article 13.3) and is therefore unavailable from that date.

Atryphodes Pascoe, 1866a: 478 [M]. Type species [automatic]: Thoracophoruswalckenaerii Hope, 1841, by original designation. Status: junior synonym of Cardiothorax Motschulsky, 1860 in Lagriinae: Adeliini. Note: unnecessary replacement name for Thoracophorus Hope, 1841.

Auchmobius J.L. LeConte, 1851: 139 [M]. Type species: Auchmobiussublaevis J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Augolesthus Motschulsky, 1872: 34 [M]. Type species: Augolesthuspurpureofasciatus Motschulsky, 1872, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Aulacodera Agassiz, 1846b: 40, 41 [F]. Type species [automatic]: Nycteliacrenicosta Guérin-Méneville, 1834, by subsequent designation (Gebien 1937a: 752). Status: junior synonym of Auladera Solier, 1836 in Pimeliinae: Nycteliini. Note: unjustified emendation of Auladera Solier, 1836, not in prevailing usage.

Aulacus Gray in Griffith and Pidgeon, 1832: 783 [M]. Type species: Aulacuschilensis Gray, 1832, by monotypy. Status: senior synonym of Orthogonoderes Solier, 1841 in Pimeliinae: Praociini. Synonymy: Kulzer (1958a: 81). Note: junior homonym of Aulacus Jurine, 1807 [Hymenoptera].

Auladera Solier, 1836: 307, 331 [F]. Type species: Nycteliacrenicosta Guérin-Méneville, 1834, by subsequent designation (Gebien 1937a: 752). Status: valid genus in Pimeliinae: Nycteliini.

Aulonasida Reitter, 1917a: 9, 19 [F]. Type species: Asidachauveneti Solier, 1836, by subsequent designation (F. Soldati 2008: 32). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Aulonogria Borchmann, 1929a: 9 [F]. Type species: Lagriarugosa Fabricius, 1801, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina. Note: also published as new in Borchmann (1930a: 405, 424).

Aulonolcus Reitter, 1904: 168 [M]. Type species: Pedinusaltaicus Gebler, 1830, by monotypy. Status: valid subgenus of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina.

Aulonoscelis Reitter, 1896c: 173 [F]. Type species: Platyscelishauseri Reitter, 1895 (= Dilaphilacoides Fischer von Waldheim, 1844), by original designation. Status: junior synonym of Gebleria Motschulsky, 1846 in Blaptinae: Blaptini: Prosodina. Synonymy: G.S. Medvedev (2005b: 99).

Auristira Borchmann, 1916a: 47, 129 [F]. Type species: Auristiraoctocostata Borchmann, 1916, by present designation. Status: junior synonym of Costiferolagria Pic, 1915 in Lagriinae: Lagriini: Lagriina. Synonymy: Borchmann (1936: 176).

Australoseriscius Koch, 1950c: 65 [M]. Type species: Crypticusexplorator Gebien, 1920, by original designation. Status: valid subgenus of Pseudoseriscius Español, 1950 in Diaperinae: Crypticini.

Austrocaribius Marcuzzi, 1954: 18 [M]. Type species: Austrocaribiusvenezuelensis Marcuzzi, 1954, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Austropalorus Halstead, 1967a: 72, 129 [M]. Type species: Austropalorusplanatus Halstead, 1967, by original designation. Status: valid genus in Tenebrioninae: Palorini.

Austropeus Carter, 1924b: 543 [M]. Type species: Austropeuspustulosus Carter, 1924, by monotypy. Status: junior synonym of Apterotheca Gebien, 1921 in Stenochiinae: Cnodalonini. Synonymy: Bouchard (2002: 452).

Austroptorina Bai, Li & Ren, 2020: 166 [F]. Type species: Gnaptorinalongicornis Li & Ren, 2004, by original designation. Status: valid subgenus of Gnaptorina Reitter, 1887 in Blaptinae: Blaptini: Gnaptorinina.

Autocera Wollaston, 1857: 154 [F]. Type species: Autoceralaticeps Wollaston, 1857, by monotypy. Status: junior synonym of Cnemeplatia Costa, 1847 in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Synonymy: Kraatz (1859: 75).

Axumia Reiche, 1850: pl. 22 [F]. Type species: Axumiapraelonga Reiche, 1850, by monotypy. Status: junior synonym of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Kraatz (1865: 170).

Axynaon Blackburn, 1897a: 34 [N]. Type species: Axynaonchampioni Blackburn, 1897, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Azarelius Fairmaire, 1892d: vii [M]. Type species: Azareliussculpticollis Fairmaire, 1892, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: transferred from Rhysopaussini to Amarygmini by Löbl and Smetana (2010: 34).

Azonoderus Harold, 1879: 125 [M]. Type species: Azonoderustristis Harold, 1879, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Balachowskya Peyerimhoff, 1928: 61 [F]. Type species: Balachowskyaportentosa Peyerimhoff, 1928, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Balassogloa Semenov, 1891: 372 [F]. Type species: Balassogloasphenarioides Semenov, 1891, by subsequent designation (R. Lucas 1920: 131). Status: valid genus in Alleculinae: Cteniopodini.

Balius Gistel, 1848a: xi, xiv [M]. Type species [automatic]: Pimeliasimplex Solier, 1836, by monotypy. Status: junior synonym of Melanostola Solier, 1836 in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Melanostola Solier, 1836.

Bancocistela Pic, 1947: 151 [F]. Type species: Bancocistelaivoirensis Pic, 1947, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Bantodemus Koch, 1955c: 428 [M]. Type species: Trigonopuslethaeus Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Baratus Fairmaire, 1897c: 233 [M]. Type species: Baratuscrenulatus Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Barbora Novák, 2020b: 462 [F]. Type species: Barboracastanea Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Barlacus Fairmaire, 1900b: 45 [M]. Type species: Barlacuscostulatus Fairmaire, 1900 (= Asyleptusfumosus Péringuey, 1896), by monotypy. Status: junior synonym of Asyleptus Péringuey, 1896 in Tenebrioninae: Amarygmini. Synonymy: Ardoin (1962b: 957); Bremer (2013a: 77).

Barsenis Pascoe, 1887: 17 [F]. Type species: Barsenisfulvipes Pascoe, 1887, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Bartolozzia Ferrer, 1998a: 369 [F]. Type species: Bartolozziahispida Ferrer, 1998, by original designation. Status: valid genus in Pimeliinae: Asidini.

Barycistela Blackburn, 1891: 327 [F]. Type species: Barycistelarobusta Blackburn, 1891, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Baryscelis Boisduval, 1835: 253 [F]. Type species: Baryscelislaticollis Boisduval, 1835, by subsequent designation (Hope 1841: 126). Status: valid genus in Tenebrionidae: incertae sedis. Note: the type material of the two species included in this genus has not been located (Matthews and Bouchard 2008: 350) and therefore the identity of the genus is uncertain.

Barytipha Pascoe, 1869: 288, 292 [F]. Type species: Barytiphasocialis Pascoe, 1869, by monotypy. Status: junior synonym of Pterohelaeus Brême, 1842 in Tenebrioninae: Heleini: Heleina. Synonymy: Matthews (1993: 1041).

Basanaedus Pic, 1917d: 11 [M]. Type species: Basanaedusluteomaculatus Pic, 1917, by subsequent designation (Gebien 1941: 820). Status: junior synonym of Lyprochelyda Fairmaire, 1899 in Lagriinae: Goniaderini. Synonymy: Ardoin (1969a: 166).

Basanopsis Gebien, 1914a: 21 [F]. Type species: Basanopsiscurvipes Gebien, 1914, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Basanus Lacordaire, 1859a: 306 [M]. Type species: Basanusforticornis Lacordaire, 1859, by monotypy. Status: valid genus in Diaperinae: Scaphidemini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Basides Motschulsky, 1873: 471 [M]. Type species: Basidesbifasciatus Motschulsky, 1873, by subsequent designation (Gebien 1925d: 560). Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: Basides Motschulsky, 1873 was used as valid after 1899 (e.g., Gebien, 1911a: 381) and therefore reversal of precedence cannot be used to conserve usage of Ischnodactylus Chevrolat, 1877.

Basilewskyum Koch, 1952b: 30, 94 [N]. Type species: Basilewskyumstenosinoide Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Bassianus Matthews & Doyen, 1989: 44 [M]. Type species: Tenebriocolydioides Erichson, 1842, by original designation. Status: valid genus in Tenebrioninae: Heleini: Asphalina.

Batessia Ponting, 2018: 131 [F]. Type species [automatic]: Agastheneswestwoodi Bates, 1873, by monotypy. Status: junior synonym of Agastenes R. Lucas, 1920 in Tenebrioninae: Heleini: Cyphaleina. Note: unnecessary replacement name for Agasthenes Bates, 1873.

Batuliodes Casey, 1907: 499 [M]. Type species: Batuliusrotundicollis J.L. LeConte, 1851, by original designation. Status: valid genus in Pimeliinae: Anepsiini.

Batuliomorpha Doyen, 1987: 359 [F]. Type species: Batuliomorphacomata Doyen, 1987, by original designation. Status: valid genus in Pimeliinae: Anepsiini.

Batulius J.L. LeConte, 1851: 148 [M]. Type species: Batuliussetosus J.L. LeConte, 1851, by subsequent designation (Casey 1907: 497). Status: valid genus in Pimeliinae: Anepsiini.

Bearnicistela Pic in E. Olivier and Pic, 1909: 139 [F]. Type species: Bearnicistelaluteicolor Pic, 1909, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Bechuanitis Koch, 1955a: 93 [F]. Type species: Trachynotusbrucki Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Becvaramarygmus Masumoto, 1999c: 369 [M]. Type species: fixed herein (ICZN 1999, Article 70.3) as Dietysusnodicornis Gravely, 1915, misidentified as Dietysusatricolor Pic, 1922 in the original designation in Masumoto (1999). Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Note: the type species “Dietysusatricolor Pic” was first established by original designation; Bremer and Lillig (2014: 12, 49) first noted that Dietysusatricolor Pic of Masumoto (1999) was identical to Dietysusnodicornis Gravely, 1915; we follow the concept of Bremer and Lillig (2014: 12, 49) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Dietysusatricolor Pic, 1922 is a junior synonym of Amarygmusfilicornis (Gravely, 1915); Iwan et al. (2020: 262) erroneously placed Becvaramarygmus Masumoto, 1999 as a junior synonym of the subgenusAmarygmus Dalman, 1823 (M. Lillig and H. Bremer, pers. comm. 2020).

Becvarius Masumoto, 1998: 207 [M]. Type species: Becvariusstanislavi Masumoto, 1998, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bellendenum Matthews, 1998: 704, 794 [N]. Type species: Bellendenumgonyxuthum Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Belopomerus Reitter, 1920a: 3 [M]. Type species: Calcarzoufali Reitter, 1915, by monotypy. Status: valid subgenus of Centorus Mulsant, 1854 in Lagriinae: Belopini.

Belopus Gebien, 1911a: 459 [M]. Type species [automatic]: Tenebrioelongatus Herbst, 1797, by monotypy. Status: valid subgenus of Centorus Mulsant, 1854 in Lagriinae: Belopini. Note: replacement name for Calcar Dejean, 1821; nomen protectum (see Bouchard and Bousquet 2020b: 6).

Belousovia G.S. Medvedev, 2007a: 157 [F]. Type species: Belousoviahelenae G.S. Medvedev, 2007, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Belutschistanops Löbl, Bouchard, Merkl & Bousquet, 2020: 3 [M]. Type species: Hyperopsschusteri Koch, 1940, by original designation. Status: valid subgenus of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: name first proposed by Koch (1943a: 523, 552) without a type species originally included (ICZN 1999, Article 13.3); Löbl et al. (2008a: 40) designated Hyperopsschusteri Koch, 1940 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Bequaertiella Pic, 1914a: 486 [F]. Type species: Bequaertiellacoerulescens Pic, 1914, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Bermejoina Español, 1944: 12 [F]. Type species: Bermejoinaaiunica Español, 1944, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Betasida Reitter, 1917a: 8, 11 [F]. Type species: Asidaluctuosa Boisduval, 1835, by subsequent designation (Viñolas and Cartagena 2005: 110). Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini. Note: some authors (e.g., F. Soldati 2008: 32) have used Asidaargenteolimbata Escalera, 1901 as type species for this genus; however, the older designation by Viñolas and Cartagena (2005: 110) should stand since Asidaluctuosa Boisduval, 1835 was one of the two originally included nominal species in Betasida.

Betschia Dajoz, 1980: 135 [F]. Type species: Betschiaminuta Dajoz, 1980, by original designation. Status: valid genus in Diaperinae: Gnathidiini: incertae sedis. Note: transferred from Tenebrionoidea: Zopheridae (as “Colydiidae”) by Ivie and Ślipiński (1990: 18); placed in “Gnathidiini incertae sedis” by Schawaller and Purchart (2012: 312).

Bia Hope, 1841: 132 [F]. Type species [automatic]: Trogossitathoracica Fabricius, 1792, by monotypy. Status: junior synonym of Bius Dejean, 1834 in Tenebrioninae: Tenebrionini. Note: unjustified emendation of Bius Dejean, 1834, not in prevailing usage.

Bielawskia Marcuzzi, 1985: 179 [F]. Type species: Bielawskiacubana Marcuzzi, 1985 (= Trimytantrondecui Ardoin, 1977), by monotypy. Status: junior synonym of Trimytantron Ardoin, 1977 in Pimeliinae: Edrotini. Synonymy: Marcuzzi (1998: 153).

Biolus Mulsant & Rey, 1854: 25 [M]. Type species: Eurynotusasperipennis Mulsant & Rey, 1854 (= Helopsgranulatus Fabricius, 1787), by subsequent designation (Koch 1953a: 276). Status: valid subgenus of Eurynotus W. Kirby, 1819 in Blaptinae: Platynotini: Eurynotina.

Biomorphus Motschulsky, 1872: 38 [M]. Type species: Biomorphustuberculatus Motschulsky, 1872 (= Amphidoraattenuata J.L. LeConte, 1851), by original designation. Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Aalbu et al. (1995: 485).

Bionesus Fairmaire, 1879b: 70 [M]. Type species: Bionesuscinereosparsus Fairmaire, 1879, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Bioplanes Mulsant, 1854: 144 [M]. Type species: Bioplanesmeridionalis Mulsant, 1854, by monotypy. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Bioramix Bates, 1879b: 478 [M]. Type species: Bioramixovalis Bates, 1879, by subsequent designation (Kaszab 1940a: 175). Status: valid genus and subgenus in Blaptinae: Platyscelidini. Note: nomenclatural stability in this genus is threatened by the discovery of an older type species designation (Bioramixpamirensis Bates, 1879, by subsequent designation by R. Lucas (1920: 138), which is currently included in the valid subgenusPlanoplatyscelis Kaszab, 1940); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Kaszab (1940a: 175); taxon also described as a new taxon with the same species by Bates (1890: 69).

Birolagria Pic, 1956: 86 [F]. Type species: Birolagriacicatricosa Pic, 1956 (= Cerogriaoriunda Borchmann, 1924), by monotypy. Status: junior synonym of Acerogria Borchmann, 1936 in Lagriinae: Lagriini: Lagriina. Synonymy: Merkl (1987: 124).

Biroum Kaszab, 1956a: 104 [N]. Type species: Biroumparadoxum Kaszab, 1956, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bius Dejean, 1834: 205 [M]. Type species: Trogossitathoracicus Fabricius, 1792, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Blacodatus Koch, 1963: 42 [M]. Type species: Blacodesvertagus Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Blacodes Duponchel, 1842a: 590 [M]. Type species: Pedinussulcatus Laporte, 1840, by monotypy. Status: junior synonym of Blenosia Laporte, 1840 in Blaptinae: Opatrini: Stizopodina. Synonymy: Erichson (1843: 245). Note: name previously attributed to Blanchard (1845: 13) in the literature.

Blapida Perty, 1830: 58 [F]. Type species: Blapidaokeni Perty, 1830, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Blapidium Bauer, 1921: 231 [N]. Type species: Blaps ocreata Allard, 1880, by subsequent designation (Nabozhenko and Chigray 2020: 10). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10). Note: originally proposed without included nominal species; Kolbe (1928: 200), by including four species in association with this name, was the first author to subsequently and expressly included nominal species in Blapidium (ICZN 1999, Article 67.2.2).

Blapidocamaria Pic, 1919a: 7 [F]. Type species: Pseudoblapidaincostata Pic, 1919, by monotypy. Status: valid subgenus of Pseudoblapida Pic, 1917 in Stenochiinae: Cnodalonini.

Blapidocampsia Pic, 1919b: 3 [F]. Type species: Campsiapallidipes Pic, 1918, by present designation. Status: valid subgenus of Campsia Lepeletier & Audinet-Serville, 1828 in Stenochiinae: Cnodalonini.

Blapidurus Fairmaire, 1891d: xcvi [M]. Type species: Blapiduruscrassicornis Fairmaire, 1891, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Champion (1895: 48).

Blapimorpha Motschulsky, 1860c: 531 [F]. Type species: Blapsreflexa Gebler, 1832, by subsequent designation (Nabozhenko 2008: 35). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860).

Blapisa Motschulsky, 1860c: 530 [F]. Type species: Blaps jaegeri Hummel, 1827 (= Tenebriomortisagus Linnaeus, 1758), by subsequent designation (Nabozhenko 2008: 35). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860).

Blaposodes Skopin, 1960a: 47 [M]. Type species: Blaps baerii Fischer von Waldheim, 1842, by original designation. Status: junior synonym of Peltarium Fischer von Waldheim, 1844 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Blaps Fabricius, 1775: 254 [F]. Type species: Tenebriomortisagus Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Blaptinae: Blaptini: Blaptina.

Blapstinus Dejean, 1821: 66 [M]. Type species: Blapspunctata Fabricius, 1792, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Blaptogonia G.S. Medvedev, 1998a: 186 [F]. Type species: Tagonoidescostulata Fairmaire, 1901, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Blaptoprosodes Reitter, 1909a: 120 [M]. Type species: Prosodesmucronata Reitter, 1893, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: G.S. Medvedev (2001: 54).

Blapylis Horn, 1870: 304, 315 [F]. Type species: Eleodescordata Eschscholtz, 1829, by subsequent designation (Bousquet et al. 2018: 143). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Blastarnodes Koch, 1958: 154 [M]. Type species: Blastarnodesherero Koch, 1958, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Blastarnus Fairmaire, 1897f: 132 [M]. Type species: Blastarnusgrallator Fairmaire, 1897, by subsequent designation (Oustalet 1898: 972). Status: junior synonym of Diestecopus Solier, 1848 in Blaptinae: Pedinini: Helopinina. Synonymy: Koch (1958: 152).

Blatticephalus Heller, 1918: 377 [M]. Type species: Blatticephalusadelotopus Heller, 1918, by monotypy. Status: valid genus in Tenebrioninae: Falsocossyphini.

Blenosia Laporte, 1840: 209 [F]. Type species: Pedinussulcatus Laporte, 1840, by subsequent designation (R. Lucas 1920: 140). Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Blepegenes Pascoe, 1868: xii [M]. Type species: Blepegenesaruspex Pascoe, 1868, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Blepusa Westwood, 1842: 69 [F]. Type species: Blepusacostata Westwood, 1842, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Blindus Mulsant & Rey, 1853b: 206 [M]. Type species: Pedinusstrigosus Faldermann, 1835, by monotypy. Status: valid subgenus of Pedinus Latreille, 1797 in Blaptinae: Pedinini: Pedinina.

Bluops Carter, 1914a: 227 [M]. Type species: Bluopsverrucosus Carter, 1914, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Bobina Novák, 2015a: 125 [F]. Type species: Bobinajendeki Novák, 2015, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Bobisthes Novák, 2019a: 178 [M]. Type species: Bobisthesbellator Novák, 2019, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Bogatshevia G.S. Medvedev & Iwan, 2006: 618 [F]. Type species [automatic]: Achaemenesvillosus Bogatchev, 1949 (= Thripterabogatchevi Kwieton, 1982), by original designation. Status: valid genus in Pimeliinae: Pimeliini. Note: replacement name for Achaemenes Bogatchev, 1949 (as “Achaemenus”).

Bolbophanes Carter, 1913a: 86 [M]. Type species: Paraphanesdumbrelli Lea, 1895, by subsequent designation (Gebien 1941: 1133). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Bolbostetha Fairmaire, 1896c: 117 [F]. Type species: Bolbostethasoleata Fairmaire, 1896, by subsequent designation (Novák and Pettersson 2008: 320). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Boletophagus Agassiz, 1846b: 48 [M]. Type species [automatic]: Silphareticulata Linnaeus, 1767, by subsequent designation (C.G. Thomson 1859: 115). Status: junior synonym of Bolitophagus Illiger, 1798 in Tenebrioninae: Bolitophagini. Note: unjustified emendation of Bolitophagus Illiger, 1798, not in prevailing usage.

Boletoxenus Motschulsky, 1858a: 63 [M]. Type species: Boletoxenusgibber Motschulsky, 1858, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolithophilus Gistel, 1832: 137 [M]. Type species [automatic]: Cistelascapularis Illiger, 1805 (= Cistelahumeralis Fabricius, 1787), by subsequent designation (Westwood 1838: 32). Status: junior synonym of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: replacement name for Mycetophila Gyllenhal, 1810.

Bolitolaemus Gebien, 1921b: 23 [M]. Type species: Bolitolaemuscatenulatus Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolitonaeus Lewis, 1894: 387 [M]. Type species: Bolitonaeusmergae Lewis, 1894, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolitopertha Gebien, 1910c: 379 [F]. Type species: Bolitoperthanovemcostata Gebien, 1910 (= Bolitophagusborbonicus Fairmaire, 1880), by monotypy. Status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: Gebien (1922b: 268, with Cherostus C.O. Waterhouse, 1894, a junior synonym of Rhipidandrus J.L. LeConte, 1862).

Bolitophagiella Miyatake, 1964: 68, 71 [F]. Type species: Bolitophaguspannosus Lewis, 1894, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolitophagus Illiger, 1798: 100 [M]. Type species: Silphareticulata Linnaeus, 1767, by subsequent designation (C.G. Thomson 1859: 115). Status: valid genus in Tenebrioninae: Bolitophagini. Note: as mentioned by Bouchard and Bousquet (2020b: 8) nomenclatural stability in this genus is threatened by the discovery of an older type species designation (Opatrumagricola Herbst, 1783, by subsequent designation by Curtis (1836: pl. 586), which is currently the type species of the valid genus Eledona Latreille, 1797); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by C.G. Thomson (1859: 115).

Bolitotherus Candèze, 1861: 367 [M]. Type species: Bolitophaguscornutus Fabricius, 1801, by subsequent designation (J.L. LeConte 1862: 236). Status: valid genus in Tenebrioninae: Bolitophagini. Note: the younger species name Bolitophaguscornutus Fabricius, 1801 was given priority over the older synonym Opatrumbifurcum Fabricius, 1798 by the ICZN (2019, Opinion 2438).

Bolitotrogus Miyatake, 1964: 68, 80 [M]. Type species: Bolitotroguskurosonis Miyatake, 1964, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Bolitoxenus Gemminger in Gemminger and Harold, 1870: 1946 [M]. Type species [automatic]: Boletoxenusgibber Motschulsky, 1858, by monotypy. Status: junior synonym of Boletoxenus Motschulsky, 1858 in Tenebrioninae: Bolitophagini. Note: unjustified emendation of Boletoxenus Motschulsky, 1858, not in prevailing usage.

Bolitrium Gebien, 1914b: 390 [N]. Type species: Bolitriumcrenulicolle Gebien, 1914, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Bolusculus Matthews, 1998: 704, 798 [M]. Type species: Bolusculusarcanus Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Bombocnodulus Koch, 1955a: 36 [M]. Type species: Psammodescrinicollis Haag- Rutenberg, 1879, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Borbochara Novák, 2009: 259 [F]. Type species: Borbocharabicolor Novák, 2009, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Borbonalia Novák, 2014: 136 [F]. Type species: Borbonaliabrancuccii Novák, 2014, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Borborella Novák, 2020a: 196 [F]. Type species: Borborellahergovitsi Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Borboresthes Fairmaire, 1897d: 253 [F]. Type species: Alleculacruralis Marseul, 1876, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: see Löbl and Smetana (2013: 39) for comments on the gender of this name.

Borchmannia Pic in Borchmann and Pic, 1912: 35 [F]. Type species: Borchmannialineaticeps Pic, 1912, by subsequent designation (R. Lucas 1920: 144). Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Borchmannius Bousquet & Bouchard in Bousquet et al., 2015: 142 [M]. Type species [automatic]: Glyptothoraxpilosus Borchmann, 1937, by original designation. Status: valid genus in Alleculinae: incertae sedis. Note: replacement name for Glyptothorax Borchmann, 1937.

Boreoptorina G.S. Medvedev, 2009: 417 [F]. Type species: Gnaptorinacordicollis G.S. Medvedev, 1998, by original designation. Status: junior synonym of Hesperoptorina G.S. Medvedev, 2009 in Blaptinae: Blaptini: Gnaptorinina. Synonymy and First Reviser action (Boreoptorina G.S. Medvedev, 2009 versus Hesperoptorina G.S. Medvedev, 2009) by Li et al. (2021: 245).

Boreosaragus Matthews, 1993: 1040, 1067 [M]. Type species: Saraguslugubris Lea, 1897, by original designation. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Borneocamaria Pic, 1917e: 17 [F]. Type species: Borneocamariaatra Pic, 1917, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Borneocistela Pic, 1922d: 18 [F]. Type species: Borneocisteladiversicornis Pic, 1922, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Borneogonocnemis Pic, 1936b: 16 [F]. Type species: Borneogonocnemisruficolor Pic, 1936, by monotypy. Status: valid subgenus of Paragonocnemis Kraatz, 1899 in Tenebrioninae: Amarygmini.

Borneolaena Schawaller, 1998: 2 [F]. Type species: Borneolaenariedeli Schawaller, 1998, by original designation. Status: valid genus in Lagriinae: Laenini.

Borneosphaerotus Grimm, 2015: 218 [M]. Type species: Borneosphaerotussantubongicus Grimm, 2015, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Borneosphena Purchart & Grimm, 2016: 522 [F]. Type species: Borneosphenafouquei Purchart & Grimm, 2016, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Borneostira Pic, 1912: 53 [F]. Type species: Rouyerusbrevilineatus Pic, 1912, by monotypy. Status: valid subgenus of Rouyerus Pic, 1911 in Lagriinae: Lagriini: Statirina.

Borneosynopticus Grimm, 2015: 219 [M]. Type species: Borneosynopticustubericollis Grimm, 2015, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Boromorphus Wollaston, 1854: 492 [M]. Type species: Boromorphusmaderae Wollaston, 1854 (= Borostagenioides P.H. Lucas, 1846), by monotypy. Status: valid genus in Pimeliinae: Boromorphini.

Bothrasida Casey, 1912: 76, 122 [F]. Type species: Asidaclathrata Champion, 1884, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Bothrichara Borchmann, 1916a: 48, 104 [F]. Type species: Lagriapulchella Guérin-Méneville, 1830, by subsequent designation (Borchmann 1936: 68). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Bothrionota Borchmann, 1936: 16, 66 [F]. Type species: Bothrionotafoveata Borchmann, 1930, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Bothriostira Borchmann, 1936: 239, 473 [F]. Type species: Bothriostiracylindracea Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Bothrotes Casey, 1907: 379 [M]. Type species: Epitraguscanaliculatus Say, 1824, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Bothynocara Gebien, 1928: 167, 170 [N]. Type species: Bothynocaratenuepunctatum Gebien, 1928, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bothynocephalus Doyen, 1988: 315 [M]. Type species: Bothynocephaluscristatus Doyen, 1988, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bothynogria Borchmann, 1916a: 128 [F]. Type species: Bothynogriacalcarata Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Botiras Fairmaire, 1891d: xcviii [M]. Type species: Botirasstriatellus Fairmaire, 1891, by subsequent designation (R. Lucas 1920: 145). Status: junior synonym of Chianalus Bates, 1879 in Blaptinae: Platyscelidini. Synonymy: Blair (1923a: 283).

Bouchardandrus Steiner, 2016: 543 [M]. Type species: Haplandrusconcolor J.L. LeConte, 1866, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini.

Bovius Gistel, 1848a: ix [M]. Type species [automatic]: Tenebrioserratus Fabricius, 1775, by monotypy. Status: junior synonym of Prioscelis Hope, 1841 in Lagriinae: Pycnocerini. Note: unnecessary replacement name for Iphius Dejean, 1834.

Brachycilibe Carter, 1911a: 207 [F]. Type species: Brachycilibeantennata Carter, 1911, by monotypy. Status: junior synonym of Dioedus J.L. LeConte, 1862 in Phrenapatinae: Penetini. Synonymy: Kaszab (1978a: 164).

Brachycryptus Quedenfeldt, 1891: 129 [M]. Type species: Brachycryptustripolitanus Quedenfeldt, 1891, by monotypy. Status: senior synonym of Tripolicryptus Strand, 1929 in Alleculinae: Cteniopodini. Note: junior homonym of Brachycryptus C.G. Thomson, 1873 [Hymenoptera].

Brachycula Fairmaire, 1906: 278 [F]. Type species: Brachyculaquadrivittata Fairmaire, 1906, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: the alternative original spelling Brachicula, used by Fairmaire (1906: 278), was rejected by Bousquet et al. (2015: 143) who acted as the First Revisers.

Brachycyphus Gebler, 1859: 473. Type species: Pimeliaimbricata Fischer, 1820, by monotypy. Status: junior synonym of Ocnera Fischer, 1822 in Pimeliinae: Pimeliini. Synonymy: Gemminger in Gemminger and Harold (1870: 1888).

Brachyesthes Fairmaire, 1868: 490 [F]. Type species: Melanesthespilosellus Marseul, 1866, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Brachygenius Dejean, 1836: 206 [M]. Type species [automatic]: Nyctelialuczotii Guérin-Méneville, 1831, by subsequent designation (Duponchel 1845b: 449). Status: junior synonym of Gyriosomus Guérin-Méneville, 1834 in Pimeliinae: Nycteliini. Note: unnecessary replacement name for Gyriosomus Guérin-Méneville, 1834.

Brachyidium Fairmaire, 1883a: 33 [N]. Type species: Brachyidiumbreviusculum Fairmaire, 1883, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Brachymoschium Fairmaire, 1896b: 348 [N]. Type species: Brachymoschiumparvitarse Fairmaire, 1896, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Brachyontis Casey, 1908: 82, 141 [F]. Type species: Coniontisglobulina Casey, 1895, by monotypy. Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Aalbu et al. (2002: 487).

Brachyphrynus Fairmaire in Fairmaire et al., 1882: 71 [M]. Type species: Brachyphrynusspissicornis Fairmaire, 1882, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Brachypilium Fairmaire, 1896a: 23 [N]. Type species: Brachypiliumsculpturatum Fairmaire, 1896, by monotypy. Status: junior synonym of Stenochinus Motschulsky, 1860 in Stenochiinae: Cnodalonini. Synonymy: Löbl et al. (2008b: 349).

Brachypophlaeus Fairmaire, 1897a: 113 [M]. Type species: Hypophlaeusdimidiatipennis Fairmaire, 1880, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Brachyscelis Dejean, 1834: 179 [F]. Type species: Pimeliamusiva Ménétriés, 1832, by subsequent designation (Bouchard et al. 2007: 388). Status: senior synonym of Pachyscelis Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Bouchard et al. (2007: 388). Note: the date of publication of the name Brachyscelis Germar, 1834 (Coleoptera: Chrysomelidae) is unknown whereas Brachyscelis Dejean was published by 30 June 1834; nomen oblitum (see Bouchard et al. 2007: 388).

Bradygena Fairmaire, 1903c: 211 [F]. Type species: Nesogenaviolacea Fairmaire, 1903 (= Nesogenaepiscopalis Fairmaire, 1875), by subsequent designation (Froussart 1961: 91). Status: junior synonym of Nesogena Mäklin, 1863 in Tenebrioninae: Praeugenini. Synonymy: Gebien (1948: 549).

Bradymerus Perroud & Montrouzier, 1865: 110 [M]. Type species: Bradymerustuberculatus Perroud & Montrouzier, 1865, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Bradynocerus Fairmaire, 1883a: 36 [M]. Type species: Bradynocerusaulacopterus Fairmaire, 1883, by monotypy. Status: junior synonym of Bradymerus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1939: 751).

Bradysphaerotus Kaszab, 1986: 293 [M]. Type species: Bradysphaerotuspapuanus Kaszab, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bradyus Dejean, 1834: 190 [M]. Type species: Erodiuspygmaeus Fischer, 1821, by monotypy. Status: valid genus in Tenebrioninae: Dissonomini.

Branchus J.L. LeConte, 1862: 222 [M]. Type species: Branchusfloridanus J.L. LeConte, 1862, by monotypy. Status: valid genus in Pimeliinae: Branchini.

Brasilius Gebien, 1928: 169, 178 [M]. Type species: Nyctobatesexaratus Fairmaire, 1889, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Bratyna Westwood, 1875: 228 [F]. Type species: Bratynaapicalis Westwood, 1875, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Bremerianus Masumoto & Bečvář, 2005: 418 [M]. Type species: Bremerianuscameronensis Masumoto & Bečvář, 2005, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Bremerus Ferrer, 2004a: 234 [M]. Type species: Bremerusfrankkochi Ferrer, 2004, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini.

Brinckia Koch, 1962b: 117 [F]. Type species: Psammodesdebilis Péringuey, 1899, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Brises Pascoe, 1869: 145 [M]. Type species: Brisestrachynotoides Pascoe, 1869, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina. Note: the First Reviser (Brises Pascoe, 1869 versus Ephidonius Pascoe, 1869) is Carter (1914b: 46).

Brittona G.S. Medvedev & Lawrence, 1986: 574 [F]. Type species: Brittonaminuta G.S. Medvedev & Lawrence, 1986, by original designation. Status: valid genus in Diaperinae: Hyociini: Brittonina.

Broomium Koch, 1950b: 338, 342 [N]. Type species: Broomiumnudum Koch, 1950, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Brosimapsida Ferrer & Ødegaard, 2005: 640 [F]. Type species: Brosimapsidagonospoides Ferrer & Ødegaard, 2005, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Brycopia Pascoe, 1869: 141 [F]. Type species: Brycopiapilosella Pascoe, 1869, by monotypy. Status: valid genus in Lagriinae: Adeliini. Note: the First Reviser (Brycopia Pascoe, 1869 versus Dinoria Pascoe, 1869) is Carter (1920a: 238).

Bulbulus Lesne, 1915: 227, 240 [M]. Type species: Arthrodeisbyrrhiformis Fairmaire, 1892, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Bunamarygmus Masumoto, 1988a: 127 [M]. Type species: Bunamarygmusthailandicus Masumoto, 1988, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Burmanosis G.S. Medvedev, 1995a: 845, 859 [F]. Type species: Stenosiskaszabi G.S. Medvedev, 1995, by original designation. Status: valid subgenus of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina.

Buxela Fairmaire, 1894a: 28 [F]. Type species: Buxelasordescens Fairmaire, 1894, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Byallius Pascoe, 1869: 42 [M]. Type species: Byalliusreticulatus Pascoe, 1869, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Bycrea Pascoe, 1868: xii [F]. Type species: Bycreavillosa Pascoe, 1868, by monotypy. Status: junior synonym of Trichoton Hope, 1841 in Blaptinae: Opatrini: Blapstinina. Synonymy: Kamiński et al. (2019c: 360).

Byrrhoncus Koch, 1954a: 49 [M]. Type species: Byrrhoncusmonticola Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Byrsax Pascoe, 1860a: 42 [M]. Type species: Byrsaxcoenosus Pascoe, 1860 (= Boletophagusgibbifer Wesmael, 1836), by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini. Note: genus originally described in Tenebrionoidea: Zopheridae: Colydiinae.

Byzacnus Pascoe, 1866a: 469 [M]. Type species: Byzacnuspicticollis Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cabirus Mulsant & Rey, 1853b: 148, 223 [M]. Type species: Cabirusminutissimus Mulsant & Rey, 1853, by subsequent designation (Gebien 1938a: 314). Status: senior synonym of Cabirutus Strand, 1929 in Blaptinae: Pedinini: Pedinina. Note: junior homonym of Cabirus Hübner, 1819 [Lepidoptera].

Cabirutus Strand, 1929: 24 [M]. Type species [automatic]: Cabirusminutissimus Mulsant & Rey, 1853, by subsequent designation (Gebien 1938a: 314). Status: valid genus and subgenus in Blaptinae: Pedinini: Pedinina. Note: replacement name for Cabirus Mulsant & Rey, 1853.

Cacicus Dejean, 1834: 182 [M]. Type species: Elenophorusamericanus Lacordaire, 1830, by monotypy. Status: senior synonym of Megelenophorus Gebien, 1910 in Pimeliinae: Elenophorini: Megelenophorina. Note: junior homonym of Cacicus Lacépède, 1799 [Aves].

Cacoplesia Fairmaire, 1898a: 237 [F]. Type species [automatic]: Plesiamelanura Klug, 1833, by subsequent designation (Bousquet et al. 2015: 137). Status: junior synonym of Eubalia Laporte, 1840 in Alleculinae: Alleculini: Gonoderina. Synonymy: Borchmann (1909a: 713). Note: replacement name for Plesia Klug, 1833.

Caecochares Koch, 1956b: 91 [M]. Type species: Caecocharesgrjebinei Koch, 1956, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Caecomenimopsis Kaszab, 1970b: 198 [F]. Type species: Caecomenimopsisleleupi Kaszab, 1970, by original designation. Status: junior synonym of Menimopsis Champion, 1896 in Diaperinae: Gnathidiini: Anopidiina. Synonymy: Peck (1990: 370).

Caecophloeus Dajoz, 1972: 278 [M]. Type species: Caecophloeusfranzi Dajoz, 1972, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Caediexis Lebedev, 1932: 125 [F]. Type species: Caediexisarenicola Lebedev, 1932, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Caediomorpha Blackburn, 1888: 272 [F]. Type species: Caediomorphaaustralis Blackburn, 1888 (= Morychusheteromerus King, 1869), by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: moved from the subtribe Ammobiina to Opatrina by Lumen et al. (2020: 346).

Caedius Blanchard, 1845: 13 [M]. Type species: Opatrumsphaeroides Hope, 1843, by subsequent designation (Lacordaire 1859a: 262). Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: Caedius is an incorrect subsequent spelling of the original spelling Coedius, first used by Lacordaire (1859a: 261), and in prevailing usage; Caedius is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Caenoblaps König, 1906: 24 [F]. Type species: Caenoblapsdifformis König, 1906, by monotypy. Status: junior synonym of Dila Fischer von Waldheim, 1844 in Blaptinae: Blaptini: Blaptina. Synonymy: Chigray et al. (2019: 3).

Caenocapicus Endrödy-Younga, 1996: 15, 19 [M]. Type species: Caenocrypticuscapensis Endrödy-Younga, 1996, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini.

Caenocorse C.G. Thomson, 1859: 117 [F]. Type species: Hypophlaeusdepressus Fabricius, 1790, by original designation. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Bedel (1906a: 92).

Caenocrypticoides Kaszab, 1969b: 322 [M]. Type species: Caenocrypticoidesloksai Kaszab, 1969, by original designation. Status: valid genus in Pimeliinae: Caenocrypticini. Note: we act as First Revisers and reject the alternative original spelling Caenocripticoides, used by Kaszab (1969b: 324).

Caenocrypticus Gebien, 1920: 139 [M]. Type species: Caenocrypticusuncinatus Gebien, 1920, by monotypy. Status: valid genus and subgenus in Pimeliinae: Caenocrypticini.

Calabosca Fairmaire, 1894f: 395 [F]. Type species [automatic]: Ascalabuspedinoides Fairmaire, 1893, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Ascalabus Fairmaire, 1893.

Calaharena Koch, 1963: 64 [F]. Type species: Calaharenadutoiti Koch, 1963, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Calcar Dejean, 1821: 67 [N]. Type species: Tenebrioelongatus Herbst, 1797, by monotypy. Status: senior synonym of Belopus Gebien, 1911 in Lagriinae: Belopini. Note: junior homonym of Calcar de Montfort, 1810 [Mollusca].

†Calcarocistela Nabozhenko in Nabozhenko et al., 2016a: 1421 [F]. Type species: Calcarocistelakirejtshuki Nabozhenko, 2016, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: described from Lower Cretaceous deposits (China).

Calcarosis Penrith, 1977: 21, 98 [F]. Type species: Zophosismichaelis Penrith, 1977, by original designation. Status: junior synonym of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Synonymy: Penrith (1981a: 22).

Callicomus Motschulsky, 1860d: 138 [M]. Type species: Diaperisriederii Faldermann, 1833, by subsequent designation (Jakobson 1924: 243). Status: senior synonym of Emypsara Pascoe, 1866 in Diaperinae: Phaleriini. Synonymy: Jakobson (1924: 243). Note: junior homonym of Callicomus Agassiz, 1846 [Coleoptera: Cerambycidae].

Callignathus Agassiz, 1846b: 58, 60 [M]. Type species [automatic]: Calognathuschevrolatii Guérin-Méneville, 1836, by monotypy. Status: junior synonym of Calognathus Guérin-Méneville, 1836 in Pimeliinae: Cryptochilini: Calognathina. Note: unjustified emendation of Calognathus Guérin-Méneville, 1836, not in prevailing usage.

Callimaria Fairmaire, 1888b: 12 [F]. Type species: Callimariaimpressipennis Fairmaire, 1888, by monotypy. Status: junior synonym of Actanorie Bates, 1879 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1956b: 90).

Callismilax Bates, 1874: 105 [F]. Type species: Leptomorphaaenea Montrouzier, 1860, by subsequent designation (Gebien 1942a: 333). Status: valid genus in Tenebrioninae: Titaenini. Note: Matthews and Lawrence (2019: 637) mentioned that this genus is not distinguishable from Titaena Erichson, 1842.

Callyntra Solier, 1836: 307, 335 [F]. Type species: Nycteliamulticosta Guérin-Méneville, 1834, by subsequent designation (Duponchel 1842b: 63). Status: valid genus in Pimeliinae: Nycteliini.

Calobamon Kraatz, 1865: 80, 105 [M]. Type species: Calobamonleptoderus Kraatz, 1865, by monotypy. Status: senior synonym of Thraustocolus Kraatz, 1866 in Pimeliinae: Tentyriini. Note: junior homonym of Calobamon Loew, 1850 [Diptera].

Calognathus Guérin-Méneville, 1836: pl. 172 [M]. Type species: Calognathuschevrolatii Guérin-Méneville, 1836, by monotypy. Status: valid genus in Pimeliinae: Cryptochilini: Calognathina.

Calogria Borchmann, 1916a: 48, 107 [F]. Type species: Calogriacostata Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Calosis Deyrolle, 1867: 81, 222 [F]. Type species: Calosisamabilis Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Note: the alternative original spelling Calliosis (p. 81) was corrected to Calosis in the “Errata” of the same work (p. 248), Calosis is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Calostega Gemminger in Gemminger and Harold, 1870: 1991 [F]. Type species [automatic]: Calostegiapurpuripennis Westwood, 1843, by monotypy. Status: junior synonym of Calostegia Westwood, 1843 in Lagriinae: Pycnocerini. Note: unjustified emendation of Calostegia Westwood, 1843, not in prevailing usage.

Calostegia Westwood, 1843: 117 [F]. Type species: Calostegiapurpuripennis Westwood, 1843, by monotypy. Status: valid genus in Lagriinae: Pycnocerini. Note: redescribed as a new genus by Westwood (1844: 221, as “Calostega”).

Calous Koch, 1958: 151 [M]. Type species: Blastarnusmichaelseni Gebien, 1920, by original designation. Status: valid subgenus of Nicandra Fairmaire, 1888 in Blaptinae: Pedinini: Helopinina.

Calydonella Doyen, 1995: 8 [F]. Type species: Calydonellalisa Doyen, 1995, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Calydoniomorpha Pic, 1917g: 19 [F]. Type species: Calydoniomorphabrevicornis Pic, 1917, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Calydonis Pascoe, 1882: 31 [F]. Type species: Calydonisrefulgens Pascoe, 1882, by subsequent designation (Gebien 1919: 141). Status: valid genus in Stenochiinae: Cnodalonini.

Calymmatophorus Gemminger in Gemminger and Harold, 1870: 1904 [M]. Type species [automatic]: Praociscucullatus Guérin-Méneville, 1834, by subsequent designation (R. Lucas 1920: 161). Status: junior synonym of Calymmophorus Solier, 1841 in Pimeliinae: Praociini. Note: unjustified emendation of Calymmophorus Solier, 1841 (as “Calymmaphorus”), not in prevailing usage.

Calymmophorus Solier, 1841a: 209, 245 [M]. Type species: Praociscucullata Guérin-Méneville, 1834, by subsequent designation (R. Lucas 1920: 161). Status: valid genus in Pimeliinae: Praociini. Note: two spellings were originally used, Calymmaphorus (pp. 209, 245, 246, 247, 250, 370) and Galymmaphorus (p. 247, in note); however, the incorrect subsequent spelling Calymmophorus, first introduced by Burmeister (1875: 494), is in prevailing usage and deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Calymmus Montrouzier, 1860: 289 [M]. Type species: Toxicumberardi Montrouzier, 1860, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina. Note: the name Calymmus was listed as a synonym of Toxicum Latreille, 1802 by Montrouzier (1860: 289); it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Fauvel 1862: 148); therefore, Calymmus was made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Calyptopsis Solier, 1835b: 253, 269 [F]. Type species: Calyptopsisemondi Solier, 1835 (= Hegetercaraboides Brullé, 1832), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Camaria Lepeletier & Audinet-Serville, 1828: 454 [F]. Type species: Camarianitida Lepeletier & Audinet-Serville, 1828 (= Tenebrionitens G.-A. Olivier, 1795), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camarimena Motschulsky, 1863: 473 [F]. Type species: Camarimenaovicauda Motschulsky, 1863, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camariocropterum Pic, 1920a: 16 [N]. Type species: Camariocropterumlaticeps Pic, 1920, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camariodes Fairmaire, 1869b: 232 [M]. Type species: Camariodescoquerelii Fairmaire, 1869 (= Camariahelopioides Klug, 1833), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: the First Reviser (Camariodes Fairmaire, 1869 versus Tinophyllus Fairmaire, 1869) is Fairmaire (1886c: 75).

Camariomorpha Pic, 1915d: 7 [F]. Type species: Camariomorphasingularis Pic, 1915, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camarothelops Kolbe, 1910: 30 [M]. Type species: Camarothelopsbraueri Kolbe, 1910, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: removed from the tribe Helopini and placed as Tenebrionidae incertae sedis by Nabozhenko (2018: 183).

Camphonota Solier, 1836: 195 [F]. Type species: Tenebriosubglobosus Pallas, 1781, by subsequent designation (Lacordaire 1859a: 188). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Campolene Pascoe, 1863a: 44 [F]. Type species: Campolenenitida Pascoe, 1863, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Camponotiphilus Lea, 1914: 257 [M]. Type species: Camponotiphilusfimbricollis Lea, 1914, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Campsia Lepeletier & Audinet-Serville, 1828: 455 [F]. Type species: Cnodalonirroratum Dalman, 1823, by subsequent designation (Hope 1841: 133). Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Campsiomorpha Pic, 1917g: 19 [F]. Type species: Camarialata Pic, 1915, by subsequent designation (Gebien 1942a: 323). Status: valid genus in Stenochiinae: Cnodalonini.

Camptobrachys Kaszab, 1941a: 4, 24 [M]. Type species: Camptobrachyssulcatus Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Canariella Uyttenboogaart, 1929: 341, 342 [F]. Type species: Canariellaarenapta Uyttenboogaart, 1929 (= Philhammussericans Fairmaire, 1871), by monotypy. Status: junior synonym of Philhammus Fairmaire, 1871 in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Synonymy: Uyttenboogaart (1940: 68). Note: junior homonym of Canariella Hesse, 1918 [Mollusca].

Cantaloubeus Ardoin, 1959b: 203 [M]. Type species: Cantaloubeusviridis Ardoin, 1959, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Cantopipleurus Koch, 1943a: 578 [M]. Type species: Tentyriamesostenoides Baudi di Selve, 1881, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Capeluprops Schawaller, 2011: 271, 273 [M]. Type species: Capelupropslaenoides Schawaller, 2011, by original designation. Status: valid genus in Lagriinae: Lupropini.

Capicrypticus Koch, 1950c: 55 [M]. Type species: Lamprocrypticuscapensis Koch, 1950, by original designation. Status: valid genus in Diaperinae: Crypticini.

Capidium Koch, 1954a: 44 [N]. Type species: Oncotustardus Solier, 1848, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Capnisa Dejean, 1836: 197 [F]. Type species: Bradyuskarelini Faldermann, 1836, by monotypy. Status: junior synonym of Gnathosia Fischer, 1821 in Pimeliinae: Tentyriini. Synonymy: Reitter (1896d: 129).

Capnisiceps Chatanay, 1914a: 215 [M]. Type species: Capnisicepsmaindroni Chatanay, 1914, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Capnochroa J.L. LeConte, 1862: 244 [F]. Type species: Cistelafuliginosa Melsheimer, 1846, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Capricephalius Koch, 1943a: 492, 508 [M]. Type species: Arthrodibiusbazmanicus Schuster, 1938, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Carabelops Fairmaire, 1899e: 534 [M]. Type species: Carabelopsaenescens Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Caraboblaps Bauer, 1921: 231 [F]. Type species: none designated. Status: undetermined taxon in Blaptinae: Blaptini: Blaptina. Note: this genus was described before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Caraboblaps and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Caracasa Pic, 1921d: 21 [F]. Type species: Caracasaaeneipennis Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Carchares Pascoe, 1887: 12 [M]. Type species: Carcharesmacer Pascoe, 1887, by monotypy. Status: valid genus in Tenebrioninae: Scaurini.

Cardigenius Solier, 1836: 407, 492 [M]. Type species: Cardigeniuslaticollis Solier, 1836, by subsequent designation (Wilke 1922: 276). Status: valid genus and subgenus in Pimeliinae: Asidini.

Cardiobioramix Kaszab, 1940a: 154, 183 [M]. Type species: Bioramixasidioides Bates, 1879, by original designation. Status: junior synonym of Euryhelops Reitter, 1902 in Blaptinae: Platyscelidini. Synonymy: Nabozhenko and Egorov (2020: 266).

Cardiochianalus Kaszab, 1940a: 150, 202 [M]. Type species: Botirassculptipennis Fairmaire, 1891, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini. Note: the alternative original spelling Chardiochianalus, used by Kaszab (1940a: 202), was rejected by Kaszab (1975a: 4) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Cardiogenius Agassiz, 1846b: 65, 66 [M]. Type species [automatic]: Cardigeniuslaticollis Solier, 1836, by subsequent designation (Wilke 1922: 276). Status: junior synonym of Cardigenius Solier, 1836 in Pimeliinae: Asidini. Note: unjustified emendation of Cardigenius Solier, 1836, not in prevailing usage.

Cardiosis Deyrolle, 1867: 81, 235 [F]. Type species: Cardiosismouffleti Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Cardiothorax Motschulsky, 1860a: 67 [M]. Type species [automatic]: Thoracophoruswalckenaerii Hope, 1841, by original designation. Status: valid genus in Lagriinae: Adeliini. Note: replacement name for Thoracophorus Hope, 1841.

Caribanosis Nabozhenko, Kirejtshuk, Merkl, Varela, Aalbu & Smith, 2016d: 568 [F]. Type species: Rhypasmaquisqueyanus Garrido & Varela, 2011, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Carinosella Purchart, 2010: 254 [F]. Type species: Carinosellamaasaiorum Purchart, 2010, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Caristela Fairmaire, 1894d: 311 [F]. Type species: Caristelamegalops Fairmaire, 1894, by monotypy. Status: junior synonym of Mycetocharina Seidlitz, 1890 in Alleculinae: Alleculini: Alleculina. Synonymy: Bedel (1897: 36).

Caroliphosis Penrith, 1981b: 114, 115 [F]. Type species: Hesseosisadamantina Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Carpiella Koch, 1962b: 148 [F]. Type species: Carpiellalatisternum Koch, 1962, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

†Caryosoma Haupt, 1950: 114, 125 [N]. Type species: Caryosomarugosum Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Casnonidea Fairmaire, 1882a: 264 [F]. Type species: Casnonideaholomelaena Fairmaire, 1882, by subsequent designation (Borchmann 1930a: 444). Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Catamerus Fairmaire, 1887b: 290 [M]. Type species: Catamerusrevoili Fairmaire, 1887, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Cataphanus Gebien, 1921a: 325, 346 [M]. Type species: Cataphanusquadraticollis Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cataphronetis P.H. Lucas, 1846: pl. 30 [F]. Type species: Cataphronetislevaillantii P.H. Lucas, 1846 (= Phtoracrenata Germar, 1836), by monotypy. Status: junior synonym of Phtora Germar, 1836 in Diaperinae: Phaleriini. Synonymy: Lacordaire (1859a: 335).

Catapiestus Perty, 1831: xxxviii [M]. Type species: Catapiestuspiceus Perty, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Catobleps Blair, 1918: 149 [M]. Type species: Catoblepsblattoides Blair, 1918 (= Blatticephalusadelotopus Heller, 1918), by original designation. Status: junior synonym of Blatticephalus Heller, 1918 in Tenebrioninae: Falsocossyphini. Synonymy: Blair (1919c: 102).

Catolaena Reitter, 1900a: 282 [F]. Type species: Laenaturkestanica Reitter, 1897, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Laena Dejean, 1821 in Lagriinae: Laenini. Synonymy: Schawaller (2001b: 4).

Catomidius Seidlitz, 1895: 791, 792 [M]. Type species: Hedyphanesrhynchophorus Seidlitz, 1895, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Catomus Allard, 1876 in Tenebrioninae: Helopini: Helopina. Synonymy: Reitter (1922a: 7).

Catomodontus Löbl & Merkl in Löbl et al., 2020: 2 [M]. Type species: Catomuscoronatus Koch, 1935, by original designation. Status: valid subgenus of Catomus Allard, 1876 in Tenebrioninae: Helopini: Helopina. Note: the name was first proposed by Koch (1935: 107, 108) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 251) designated Catomuscoronatus Koch, 1935 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Catomulus Reitter, 1897a: 302 [M]. Type species: Catomulussubaeneus Reitter, 1897 (= Oxycaraolcesii Fairmaire, 1883), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Catomus Allard, 1876a: 4 [M]. Type species: Catomuspersicus Allard, 1876, by subsequent designation (Gebien 1943: 409). Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina.

Catopherus Carter, 1918: 713 [M]. Type species: Catopheruscorpulentus Carter, 1918 (= Axynaonchampioni Blackburn, 1897), by monotypy. Status: junior synonym of Axynaon Blackburn, 1897 in Tenebrioninae: Amarygmini. Synonymy: Carter (1920a: 249).

Caucasohelops Nabozhenko, 2006: 816 [M]. Type species: Eustenomacidiussvetlanae Nabozhenko, 2006, by original designation. Status: valid subgenus of Eustenomacidius Nabozhenko, 2006 in Tenebrioninae: Helopini: Cylindrinotina.

Caucasonotus Nabozhenko, 2000: 107 [M]. Type species: Cylindronotusdombaicus Nabozhenko, 2000, by original designation. Status: valid subgenus of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina.

Caudamarygmus Bremer, 2001b: 88, 94 [M]. Type species: Caudamarygmusnotabilis Bremer, 2001, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Caulostena Fairmaire, 1896b: 355 [F]. Type species: Caulostenafoveicollis Fairmaire, 1896, by monotypy. Status: valid genus in Alleculinae: Alleculini: Mycetocharina.

Cauricara Penrith, 1979: 7, 42 [N]. Type species: Stenocaravelox Péringuey, 1886, by original designation. Status: valid subgenus of Stenocara Solier, 1835 in Pimeliinae: Adesmiini.

Caverneleodes Triplehorn, 1975: 39 [M]. Type species: Eleodeseasterlai Triplehorn, 1975, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Caxtonana Buck, 1960: 224 [F]. Type species: Caxtonanacostata Buck, 1960, by monotypy. Status: junior synonym of Apterotheca Gebien, 1921 in Stenochiinae: Cnodalonini. Synonymy: Bouchard (2002: 452).

Cechenosternum Gebien, 1921b: 9 [N]. Type species: Cechenosternumnigromaculatum Gebien, 1921, by subsequent designation (Gebien 1939: 748). Status: valid genus in Diaperinae: Crypticini.

Cecrops Gistel, 1834: 21 [M]. Type species: Tenebriogigas Linnaeus, 1763, by subsequent designation (Bousquet and Bouchard 2017: 131). Status: junior synonym of Mylaris Pallas, 1781 in Stenochiinae: Cnodalonini. Synonymy: Bousquet and Bouchard (2017: 131).

Cedrosius Fairmaire, 1902b: 332 [M]. Type species: Cedrosiuscalosomoides Fairmaire, 1902, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Celebesa Pic, 1921d: 23 [F]. Type species: Celebesaelongata Pic, 1921, by subsequent designation (Riley 1923: 126). Status: valid genus in Stenochiinae: Cnodalonini.

Celibe Boisduval, 1835: 262 [F]. Type species: Celibeaustralis Boisduval, 1835, by subsequent designation (Watt 1968: 36). Status: valid genus in Tenebrioninae: Heleini: Heleina.

Celox Gistel, 1848a: 126 [F]. Type species [automatic]: Cnodalonirroratum Dalman, 1823, by subsequent designation (Hope 1841: 133). Status: junior synonym of Campsia Lepeletier & Audinet-Serville, 1828 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Campsia Lepeletier & Audinet-Serville, 1828.

Cenophorus Mulsant & Rey, 1859a: 113 [M]. Type species: Cenophorusviduus Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Cenoscelis Wollaston, 1868: 179 [F]. Type species: Cenoscelistibialis Wollaston, 1868 (= Ulomapullum Erichson, 1843), by monotypy. Status: valid genus and subgenus in Tenebrioninae: Ulomini.

Centorus Mulsant, 1854: 272 [M]. Type species: Calcarprocerum Mulsant, 1854, by monotypy. Status: valid genus and subgenus in Lagriinae: Belopini.

Centrioptera Mannerheim, 1843: 279 [F]. Type species: Centriopteracaraboides Mannerheim, 1843, by monotypy. Status: junior synonym of Cryptoglossa Solier, 1837 in Pimeliinae: Cryptoglossini. Synonymy: Aalbu et al. (2002: 486).

Centrocnemis Kraatz in Heyden and Kraatz, 1882: 330 [F]. Type species: Centrocnemismollis Kraatz, 1882, by monotypy. Status: senior synonym of Centrocnemita Strand, 1935 in Pimeliinae: Pimeliini. Note: junior homonym of Centrocnemis Signoret, 1852 [Hemiptera].

Centrocnemita Strand, 1935a: 284 [F]. Type species [automatic]: Centrocnemismollis Kraatz, 1882, by monotypy. Status: valid subgenus of Lasiostola Dejean, 1834 in Pimeliinae: Pimeliini. Note: replacement name for Centrocnemis Kraatz, 1882.

Centronopus Solier, 1848: 153, 154, 258 [M]. Type species: Centronopusextensicollis Solier, 1848 (= Tenebriosuppressus Say, 1835), by original designation. Status: valid genus and subgenus in Tenebrioninae: Centronopini.

Centropus Jakobson, 1914: 528 [M]. Type species [automatic]: Centronopusextensicollis Solier, 1848 (= Tenebriosuppressus Say, 1835), by original designation. Status: junior synonym of Centronopus Solier, 1848 in Tenebrioninae: Centronopini. Note: unjustified emendation of Centronopus Solier, 1848, not in prevailing usage; junior homonym of Centropus Illiger, 1811 [Aves].

Cephacerus Rafinesque, 1815: 113 [M]. Type species [automatic]: Erodiusgibbus Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Erodius Fabricius, 1775 in Pimeliinae: Erodiini. Note: unnecessary replacement name for Erodius Fabricius, 1775.

Cephaladesmia Gebien, 1920: 49, 61 [F]. Type species: Cephaladesmiathomseni Gebien, 1920, by monotypy. Status: junior synonym of Stenodesia Reitter, 1916 in Pimeliinae: Adesmiini. Synonymy: Penrith (1979: 70, 85).

Cephalamarygmus Bremer, 2001b: 92, 95 [M]. Type species: Amarygmuspreangerensis Pic, 1952, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Cephaleucyrtus Pic, 1923c: 22 [M]. Type species: Cephaleucyrtusviridicollis Pic, 1923, by monotypy. Status: junior synonym of Gauromaia Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1941: 1144).

Cephaloplonyx Pic, 1922c: 12 [M]. Type species: Cephaloplonyxopacus Pic, 1922 (= Helopsdentipes Fabricius, 1781), by monotypy. Status: junior synonym of Hoplobrachium Fairmaire, 1886 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1943: 922).

Cephalostenus Solier, 1838b: 160, 184 [M]. Type species: Cephalostenusdejeanii Solier, 1838 (= Scauruselegans Brullé, 1832), by subsequent designation (Hope 1841: 115). Status: valid genus in Tenebrioninae: Scaurini.

Cephalothydemus Pic, 1923c: 24 [M]. Type species: Cephalothydemustheresae Pic, 1923, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ceradelium Preudhomme de Borre, 1868: 126 [N]. Type species: Ceradeliumarmatum Preudhomme de Borre, 1868 (= Blepegenesaruspex Pascoe, 1868), by monotypy. Status: junior synonym of Blepegenes Pascoe, 1868 in Lagriinae: Adeliini. Synonymy: Pascoe (1869: 41).

Ceradesmia Gebien, 1920: 49, 63 [F]. Type species: Stenocaraalbicolle Haag-Rutenberg, 1878, by original designation. Status: valid subgenus of Metriopus Solier, 1835 in Pimeliinae: Adesmiini.

Ceramba Fauvel, 1904: 206 [F]. Type species: Cerambahydrovatina Fauvel, 1904, by monotypy. Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Gebien (1940: 429).

Cerandria Dejean, 1834: 200 [F]. Type species: Trogossitacornuta Fabricius, 1798, by subsequent designation (Duponchel 1842b: 285). Status: junior synonym of Gnatocerus Thunberg, 1814 in Diaperinae: Diaperini: Adelinina. Synonymy: Erichson (1847a: 119).

Cerandrosus Gebien, 1921a: 325, 394 [M]. Type species: Cerandrosusnodipennis Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cerasoma Endrödy-Younga, 1989: 113 [N]. Type species: Cerasomacerasus Endrödy-Younga, 1989, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Ceratanisus Gemminger in Gemminger and Harold, 1870: 1818 [M]. Type species [automatic]: Anisocerustristis Faldermann, 1837, by monotypy. Status: valid genus in Pimeliinae: Ceratanisini. Note: replacement name for Anisocerus Faldermann, 1837.

Ceratoma Borchmann, 1916a: 49 [F]. Type species: none designated. Status: undetermined taxon in Lagriinae: Lagriini: Lagriina. Note: this genus was included in a key, which fulfils the criteria of availability for new genus-group names proposed before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Ceratoma and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Ceratopelius Antoine, 1963: 52, 54 [M]. Type species: Ceratopeliusmussardi Antoine, 1963, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Ceratupis Perty, 1830: 57 [F]. Type species: Ceratupisnigerrima Perty, 1830, by monotypy. Status: junior synonym of Antimachus Gistel, 1829 in Tenebrioninae: Ulomini. Synonymy: Lacordaire (1859a: 330).

Cerenopus J.L. LeConte, 1851: 143 [M]. Type species: Cerenopusconcolor J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 173). Status: valid genus in Tenebrioninae: Cerenopini.

Cerocamptus Gebien, 1919: 28, 151 [M]. Type species: Camariamalayana Fairmaire, 1893, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cerodolus Sharp, 1886: 410 [M]. Type species: Cerodoluschrysomeloides Sharp, 1886, by monotypy. Status: valid genus in Tenebrioninae: Heleini: incertae sedis. Note: transferred from Titaenini by Matthews and Lawrence (2019: 628).

Cerogria Borchmann, 1911: 210 [F]. Type species: Lagriaanisocera Wiedemann, 1823, by subsequent designation (Borchmann 1936: 119). Status: valid genus and subgenus in Lagriinae: Lagriini: Lagriina. Note: as mentioned by Löbl et al. (2020: 5), discovery of the earlier designation of Cerogriadohrni Borchmann, 1911 as the type species of Cerogria (by subsequent designation by R. Lucas (1920: 174)) threatens the nomenclatural stability of this genus since C.dohrni is the type species of the valid genus Acerogria Borchmann, 1936; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Borchmann (1936: 119).

Cerogriodes Borchmann, 1941b: 25 [M]. Type species: Cerogriaklapperichi Borchmann, 1941, by monotypy. Status: junior synonym of Cerogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina. Synonymy: Merkl (2007: 262).

Ceromelaephus Koch, 1955a: 87 [M]. Type species: Trachynotusbadeni Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Ceropria Laporte & Brullé, 1831: 332, 396 [F]. Type species: Helopsindutus Wiedemann, 1819, by subsequent designation (Duponchel 1844b: 359). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cerosis Gebien, 1920: 33, 35 [F]. Type species: Cerosishereroensis Gebien, 1920, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Cerostena Solier, 1836: 307, 325 [F]. Type species: Cerostenadeplanata Solier, 1836, by present designation. Status: junior synonym of Psectrascelis Solier, 1836 in Pimeliinae: Nycteliini. Synonymy: Fairmaire (1876: 356).

Cerostira Borchmann, 1942a: 36 [F]. Type species [automatic]: Porrolagriasubaenea Borchmann, 1908, by subsequent designation (Borchmann 1936: 227). Status: valid genus in Lagriinae: Lagriina. Note: replacement name for Allocera Borchmann, 1916.

Cerysia Bremer, 2001a: 68 [F]. Type species: Elixotalaevicornis Blair, 1929, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Cestrinus Erichson, 1842a: 172 [M]. Type species: Cestrinustrivialis Erichson, 1842, by subsequent designation (Blair 1919a: 529). Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Champion (1894a: 354), Doyen et al. (1990: 231).

Chaerodes White, 1846: 12 [M]. Type species: Chaerodestrachyscelides White, 1846, by monotypy. Status: valid genus in Lagriinae: Chaerodini.

Chaeroplonyx Bremer, 2014a: 36 [M]. Type species: Plesiophthalmuskimanisensis Masumoto, 2001, by original designation. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Chaetopsia Gebien, 1925c: 567 [F]. Type species: Chaetopsiaangusticollis Gebien, 1925, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Chaetotoma Motschulsky, 1860c: 533 [F]. Type species: Tenebriocephalotes Pallas, 1781, by subsequent designation (Löbl et al. 2008a: 41). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Chaetyllus Pascoe, 1860b: 122 [M]. Type species: Chaetyllusanthicoides Pascoe, 1860, by monotypy. Status: valid genus in Lagriinae: Laenini. Note: placed in the tribe Laenini by Kanda (2016: 562).

Chalcocyclus Fairmaire, 1884d: 74 [M]. Type species: Chalcocyclusspeculifer Fairmaire, 1884, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Chalcopauliana Ardoin, 1961a: 209 [F]. Type species: Chalcopaulianavinsoni Ardoin, 1961, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Chalcoplonyx Ardoin, 1963b: 308, 333 [M]. Type species: Gonocnemisviridis Kraatz, 1899, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Chalcopteroides Gebien, 1948: 497 [M]. Type species [automatic]: Chalcopterusiridicolor Blessig, 1861, by subsequent designation (Gebien 1948: 497). Status: valid genus in Tenebrioninae: Amarygmini. Note: replacement name for Chalcopterus Blessig, 1861; Chalcopteroides was proposed earlier by Strand (1935b: 302); however, this name is unavailable because Strand did not designate a type species for the nominal taxon, a mandatory requirement for replacement name without valid typification proposed after 1930 (ICZN 1999; Article 13.3.1).

Chalcopterus Blessig, 1861: 103 [M]. Type species: Chalcopterusiridicolor Blessig, 1861, by subsequent designation (Gebien 1948: 497). Status: senior synonym of Chalcopteroides Gebien, 1948 in Tenebrioninae: Amarygmini. Note: junior homonym of Chalcopterus Reichenbach, 1852 [Aves].

Chalcostylus Fairmaire, 1898b: 409 [M]. Type species: Chalcostylusperrieri Fairmaire, 1898, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Nycteropina.

Charianus Bates, 1879a: 297 [M]. Type species: Tetraphylluspurpuratus Coquerel, 1851, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Chariophenus Blair, 1929a: 239 [M]. Type species: Chariophenuswasmanni Blair, 1929, by original designation. Status: junior synonym of Foochounus Pic, 1921 in Stenochiinae: Cnodalonini. Synonymy: Schawaller and Ando (2009: 260).

Chariotheca Pascoe, 1860b: 125 [F]. Type species: Chariothecacoruscans Pascoe, 1860, by subsequent designation (Gebien 1942a: 335). Status: valid genus in Stenochiinae: Cnodalonini.

Chariothes Carter, 1914b: 78 [M]. Type species [automatic]: Chariothecacoruscans Pascoe, 1860, by subsequent designation (Gebien 1942a: 335). Status: junior synonym of Chariotheca Pascoe, 1860 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Chariotheca Pascoe, 1860.

Charisius Champion, 1888: 421 [M]. Type species: Charisiusfasciatus Champion, 1888, by subsequent designation (R. Lucas 1920: 178). Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: the First Reviser (Charisius Champion, 1888 versus Narses Champion, 1888) is Campbell (2014: 271).

Chartopteryx Westwood, 1841a: 43 [F]. Type species: Chartopteryxchildrenii Westwood, 1841, by monotypy. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy and First Reviser action (Chartopteryx Westwood, 1841 versus Cyphaleus Westwood, 1841): Matthews (1992: 490).

Chaseleodes Thomas, 2015: 122 [F]. Type species: Eleodescurta Champion, 1884, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Chatanayus Ardoin, 1957: 61 [M]. Type species: Chemolanusvillosipes Fairmaire, 1884, by original designation. Status: senior synonym of Anachayus Bouchard & Bousquet, nom. nov. in Stenochiinae: Cnodalonini. Note: junior homonym of Chatanayus Fleutiaux 1940 [Coleoptera: Elateridae].

Cheilopoma Murray, 1867: 20 [N]. Type species: Cheilopomacastaneum Murray, 1867, by monotypy. Status: valid genus in Diaperinae: Hypophlaeini.

Cheirodes Gené, 1839: 73 [M]. Type species: Cheirodessardous Gené, 1839, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Melanimonini.

Cheiroplus Ardoin, 1963a: 129 [M]. Type species: Cheiroplusfreyi Ardoin, 1963, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: the earlier usage of Cheiroplus by Ardoin (1962b: 970) is unavailable since it was published after 1930 without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Cheirosis Deyrolle, 1867: 81, 220 [F]. Type species: Zophosisovata Faldermann, 1837 (= Pedinusacuminatus Fischer, 1832), by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Chemolanus Bates, 1879a: 296 [M]. Type species: Tetraphyllusconsobrinus Fairmaire, 1868 (= Camariaobscura Klug, 1833), by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cherostus C.O. Waterhouse, 1894: 68 [M]. Type species: Cherostuswalkeri C.O. Waterhouse, 1894, by subsequent designation (Barber 1914: 191). Status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: Gebien (1939: 762).

Chianalus Bates, 1879b: 479 [M]. Type species: Chianaluscostipennis Bates, 1879, by monotypy. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Chilenolagria Pic, 1936a: 28 [F]. Type species: Chilenolagrialimbata Pic, 1936, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Chileone Bates, 1868: 264 [F]. Type species: Chileonedeyrollei Bates, 1868, by monotypy. Status: junior synonym of Hypaulax Bates, 1868 in Stenochiinae: Cnodalonini. Synonymy: Carter (1914b: 46).

Chiliarchum Koch, 1953a: 263 [N]. Type species: Molurisbertolonii Guérin-Méneville, 1844, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina. Note: elevated to rank of genus (from subgenus of Ocnodes Fåhraeus, 1870) by Kamiński et al. (2020: 9, 17)

Chilometopon Horn, 1874: 31 [N]. Type species: Trimytisabnormis Horn, 1870, by subsequent designation (Casey 1907: 367). Status: valid genus in Pimeliinae: Edrotini.

Chinotrigon Skopin, 1973: 170 [M]. Type species: Trigonoscelissublaevigata Reitter, 1887, by original designation. Status: valid subgenus of Trigonoscelis Dejean, 1834 in Pimeliinae: Pimeliini.

Chirocharis Kolbe, 1903: 166, 179 [F]. Type species: Chiroscelisaustralis Westwood, 1845, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Chirodes Agassiz, 1846b: 78, 81 [M]. Type species [automatic]: Cheirodessardous Gené, 1839, by monotypy. Status: junior synonym of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini. Note: unjustified emendation of Cheirodes Gené, 1839, not in prevailing usage.

Chiroscelis Lamarck, 1804: 261 [F]. Type species: Chiroscelisbifenestra Lamarck, 1804 (= Tenebriodigitatus Fabricius, 1801), by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Chirosis Gemminger in Gemminger and Harold, 1870: 1806 [F]. Type species [automatic]: Zophosisovata Faldermann, 1837 (= Pedinusacuminatus Fischer, 1832), by monotypy. Status: junior synonym of Cheirosis Deyrolle, 1867 in Pimeliinae: Zophosini. Note: unjustified emendation of Cheirosis Deyrolle, 1867, not in prevailing usage.

Chitwania Novák, 2015d: 91 [F]. Type species: Chitwaniakejvali Novák, 2015, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Chlamydion Gistel, 1848a: x [N]. Type species [automatic]: Opatrumorientale Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: junior synonym of Sclerum Dejean, 1834 in Blaptinae: Opatrini: Sclerina. Note: unnecessary replacement name for Sclerum Dejean, 1834.

Chlorocamma Bates, 1873e: 371 [N]. Type species: Chlorocammacarenipenne Bates, 1873 (= Leptomorphasulcata Montrouzier, 1860), by subsequent designation (Gebien 1942a: 755). Status: valid genus in Stenochiinae: Cnodalonini.

Chlorophanes Matthews, 1992: 451, 461 [M]. Type species: Platyphanespunctipennis Carter, 1911, by original designation. Status: senior synonym of Phanechloros Matthews & Bouchard, 2008 in Tenebrioninae: Heleini: Cyphaleina. Note: junior homonym of Chlorophanes Reichenbach, 1853 [Aves].

Chlorophila Semenov, 1891: 374 [F]. Type species: Lagriaportschinskii Semenov, 1891, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Choaspes Champion, 1885: 118 [M]. Type species: Choaspespurpureus Champion, 1885, by subsequent designation (Gebien 1941: 338). Status: senior synonym of Choastes Champion, 1893 in Stenochiinae: Cnodalonini. Note: junior homonym of Choaspes Moore, 1881 [Lepidoptera].

Choastes Champion, 1893a: 526 [M]. Type species [automatic]: Choaspespurpureus Champion, 1885, by subsequent designation (Gebien 1941: 338). Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Choaspes Champion, 1885.

Choerodes Gemminger in Gemminger and Harold, 1870: 1944 [M]. Type species [automatic]: Chaerodestrachyscelides White, 1846, by monotypy. Status: junior synonym of Chaerodes White, 1846 in Lagriinae: Chaerodini. Note: unjustified emendation of Chaerodes White, 1846, not in prevailing usage; junior homonym of Choerodes Leidy, 1852 [Mammalia] and Choerodes Guenée, 1857 [Lepidoptera].

Cholipus Pascoe, 1866a: 471 [M]. Type species: Cholipusbrevicornis Pascoe, 1866, by original designation. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Mäklin (1863a: 552, with Encyalesthus Motschulsky, 1860, a junior synonym of Derosphaerus J. Thomson, 1858).

Chorasmius Bates, 1868: 310 [M]. Type species: Evaniosomusprocerus Erichson, 1847, by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Choresmolamus G.S. Medvedev, 1978: 150 [M]. Type species: Dilamusoxianus G.S. Medvedev, 1978, by original designation. Status: valid subgenus of Dilamus Jacquelin du Val, 1861 in Blaptinae: Opatrini: Ammobiina.

Choristopsis Kraatz, 1865: 81, 227 [F]. Type species: Choristopsiscaucasica Kraatz, 1865, by monotypy. Status: junior synonym of Calyptopsis Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Reitter (1900c: 130).

Chromatia J.L. LeConte, 1862: 244 [F]. Type species: Cistelaamoena Say, 1824, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Chromomaea Pascoe, 1866a: 490 [F]. Type species: Chromomaeapicta Pascoe, 1866, by monotypy. Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Fauvel (1905: 225).

Chrysobalus Boisduval, 1835: 267 [M]. Type species: Chrysobalusfulgidipennis Boisduval, 1835, by monotypy. Status: senior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Lacordaire (1859b: 409); although the identity of the type species is uncertain (Matthews 1992: 439, Matthews and Bouchard 2008: 350), it is probably a species of Cyphaleus Westwood, 1841. Note: nomen oblitum; we provide references to support the conservation of Cyphaleus Westwood, 1841 as the valid name for this genus (ICZN 1999, Article 23.9.1) in Appendix 2.

Chrysolagria Seidlitz, 1898b: 336, 339 [F]. Type species: Lagriaviridipennis Fabricius, 1798, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Chrysolinoides Jolivet, 1951: 2 [M]. Type species: Chrysolinoidesphilippinensis Jolivet, 1951 (= Hemicerairidicolor Gebien, 1921), by monotypy. Status: junior synonym of Hemicera Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini. Synonymy: Kulzer (1952: 764). Note: genus originally described in Chrysomeloidea: Chrysomelidae.

Chrysomaia Kulzer, 1952: 755, 756 [F]. Type species: Eucyrtuscarbunculus Fairmaire, 1885, by original designation. Status: junior synonym of Augolesthus Motschulsky, 1872 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 134).

Chrysopeplus Gebien, 1942a: 755 [M]. Type species [automatic]: Helopsexpolitus Broun, 1880, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Leiopeplus Broun, 1893.

Cibdelis Mannerheim, 1843: 282 [F]. Type species: Cibdelisblaschkii Mannerheim, 1843, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cilioncotus Koch, 1954a: 41 [M]. Type species: Oncotusnamaquanus Koch, 1954, by original designation. Status: valid subgenus of Oncotus Blanchard, 1845 in Blaptinae: Platynotini: Eurynotina.

Cillibus Matthews, 1993: 1040, 1078 [M]. Type species: Saragusblackburni W.J. MacLeay, 1888, by original designation. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Cimicia Fairmaire, 1891a: lxxxix [F]. Type species: Cimiciaspinipes Fairmaire, 1891, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Cimicichora Koch, 1952b: 38 [F]. Type species: Cimicichoracrenulata Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Cimiciopsis Koch, 1952b: 21, 87 [F]. Type species: Cimiciopsiscastleae Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Cimipsa Peyerimhoff, 1911: 346 [F]. Type species: Cimipsasergenti Peyerimhoff, 1911, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Circomus Fleischer, 1900: 236 [M]. Type species: Hypophlaeussubdepressus Wollaston, 1864, by monotypy. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Löbl et al. (2008b: 276).

Cirsa P.H. Lucas, 1857: lvi [F]. Type species: Cirsastriaticollis P.H. Lucas, 1857, by monotypy. Status: valid subgenus of Micipsa P.H. Lucas, 1855 in Pimeliinae: Tentyriini.

Cirta Gemminger in Gemminger and Harold, 1870: 1831 [F]. Type species [automatic]: Cirsastriaticollis P.H. Lucas, 1857, by monotypy. Status: junior synonym of Cirsa P.H. Lucas, 1857 in Pimeliinae: Tentyriini. Note: unjustified emendation of Cirsa P.H. Lucas, 1857, not in prevailing usage.

Cissides Chatanay, 1915a: 475, 495 [M]. Type species: Heterophyluspunctatissimus Fairmaire, 1869, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cistela Fabricius, 1775: 116 [F]. Type species: Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: senior synonym of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: junior homonym of Cistela Geoffroy, 1762 [Coleoptera: Byrrhidae]. Note: the older name Cistela Geoffroy, 1762 was suppressed for purposes of the Principle of Priority but not for those of the Principle of Homonymy (ICZN 1994a, Opinion 1754).

Cistelampra Fairmaire, 1897b: 386 [F]. Type species: Cistelamprapurpurina Fairmaire, 1897, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Cistelina Seidlitz, 1896: 195 [F]. Type species: Cisteladavidis Fairmaire, 1878, by subsequent designation (R. Lucas 1920: 187). Status: valid genus in Alleculinae: Cteniopodini.

Cistella Gistel, 1848a: xi [F]. Type species [automatic]: Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: unjustified emendation of Cistela Fabricius, 1775, not in prevailing usage.

Cistelodema Borchmann, 1932a: 380 [F]. Type species: Pseudocistelacyanea Pic, 1930, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Cisteloida Fairmaire, 1882a: 256 [F]. Type species: Cisteloidacastanescens Fairmaire, 1882, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Cistelomorpha Redtenbacher, 1868: 134 [F]. Type species: Cistelomorphastraminea Redtenbacher, 1868, by subsequent designation (Borchmann 1932a: 358). Status: valid genus in Alleculinae: Cteniopodini.

Cistelopsis Fairmaire, 1896a: 39 [F]. Type species: Cistelopsisrufina Fairmaire, 1896, by subsequent designation (R. Lucas 1920: 187). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Clamoris Gozis, 1886: 25 [F]. Type species [automatic]: Phtoracrenata Mulsant, 1854 (= Clamorisinsurgens Gozis, 1886), by monotypy. Status: valid genus in Phrenapatinae: Penetini. Note: replacement name for Phtora Mulsant, 1854.

Clastopus Fairmaire, 1898b: 407 [M]. Type species: Clastopuseurynotoides Fairmaire, 1898, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Claudegirardius Iwan, 1999b: 372 [M]. Type species: Claudegirardiusbertiae Iwan, 1999, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Clavatoodescelis Kaszab, 1940b: 942, 974 [F]. Type species: Platyscelismelas Fischer, 1823, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini. Note: the First Reviser (Clavatoodescelis Kaszab, 1940 versus Oblongoodescelis Kaszab, 1940) is Egorov (2020: 380).

Cleognathus Gebien, 1921b: 154 [M]. Type species: Cleognathusprosternalis Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Cleolaus Champion, 1886: 142 [M]. Type species: Penetasommeri Lacordaire, 1859, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Cleomis Fairmaire, 1892c: 54 [M]. Type species: Cleomisviolaceipes Fairmaire, 1892, by subsequent designation (R. Lucas 1920: 189). Status: valid genus in Stenochiinae: Cnodalonini. Note: removed from synonymy with Psydus Pascoe, 1868 by Kaszab (1983b: 382).

Clinocranion Solier, 1843: 114 [N]. Type species: Clinocranionspinosum Solier, 1843, by subsequent designation (R. Lucas 1920: 190). Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina. Note: the alternative original spelling Clynocranion, used by Solier (1843: 4), was rejected by Koch (1955a: 70) who acted as the First Reviser.

Clitobius Mulsant & Rey, 1859c: 141 [M]. Type species: Clitobiussabulicola Mulsant & Rey, 1859 (= Opatrumovatum Erichson, 1843), by monotypy. Status: valid genus and subgenus in Blaptinae: Opatrini: Ammobiina. Note: the original combination of the accepted name of the type species, Opatrumovatum Erichson, 1843, is a junior primary homonym of Opatrumovatum Fabricius, 1801.

Clypeophtora F. Soldati & L. Soldati, 2003: 4 [F]. Type species: Phtoratronqueti F. Soldati & L. Soldati, 2003, by original designation. Status: valid subgenus of Phtora Germar, 1836 in Diaperinae: Phaleriini.

Cnecosochara Reitter, 1913: 660 [F]. Type species: Cnecosocharapetriiformis Reitter, 1913, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Cnemandrosus Gebien, 1927: 42 [M]. Type species: Eucyrtussemipurpureus Fairmaire, 1896, by original designation. Status: junior synonym of Plamius Fairmaire, 1896 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1941: 1143).

Cnemeplatia Costa, 1847: 146 [F]. Type species: Cnemeplatiaatropos Costa, 1847, by monotypy. Status: valid genus in Pimeliinae: Cnemeplatiini: Cnemeplatiina.

Cnemodasus Gebien, 1914b: 374 [M]. Type species: Cnemodasusrectangulus Gebien, 1914, by subsequent designation (Kaszab 1982b: 43). Status: junior synonym of Brachyidium Fairmaire, 1883 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1921a: 492).

Cnemodinus Cockerell, 1906: 242 [M]. Type species [automatic]: Cnemodustestaceus Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Cnemodinini. Note: replacement name for Cnemodus Horn, 1870.

Cnemodus Horn, 1870: 266 [M]. Type species: Cnemodustestaceus Horn, 1870, by monotypy. Status: senior synonym of Cnemodinus Cockerell, 1906 in Pimeliinae: Cnemodinini. Note: junior homonym of Cnemodus Herrich-Schaeffer, 1850 [Hemiptera].

Cnemophloeus Bremer, 1998: 10, 13 [M]. Type species: Hypophlaeusfilum Fairmaire, 1893, by original designation. Status: junior synonym of Stenophloeus Blair, 1921 in Diaperinae: Hypophlaeini. Synonymy: Löbl et al. (2008b: 312).

Cnemoplatia Wollaston, 1865: 411 [F]. Type species [automatic]: Cnemeplatiaatropos Costa, 1847, by monotypy. Status: junior synonym of Cnemeplatia Costa, 1847 in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Note: unjustified emendation of Cnemeplatia Costa, 1847, not in prevailing usage.

Cneocnemis Gebien, 1914c: 32 [F]. Type species: Ulomahaemorrhoum Fairmaire, 1893, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Cnephalura Doyen, 1988: 313 [F]. Type species: Cnephaluraumbrata Doyen, 1988, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cnodalon Latreille, 1797: 23 [N]. Type species: Cnodalonviride Latreille, 1804, by subsequent monotypy (Latreille 1804: 321). Status: valid genus in Stenochiinae: Cnodalonini. Note: originally proposed without included nominal species; in order to promote nomenclatural stability Bousquet and Bouchard (2019: 114, 116) recommended to the ICZN to use its plenary powers to designate Cnodalonviride Latreille, 1804 as the type species for the nominal genus Cnodalon Latreille, 1797, as currently accepted in the literature, after the recent discovery that this species was in fact not an originally included nominal species in the genus Cnodalon, and so not available as the type species.

Cnodalum Agassiz, 1846b: 91 [N]. Type species [automatic]: Cnodalonviride Latreille, 1804, by subsequent monotypy (Latreille 1804: 321). Status: junior synonym of Cnodalon Latreille, 1797 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Cnodalon Latreille, 1797, not in prevailing usage; in a recent application to the ICZN to fix type species problems with the genus Cnodalon Latreille, 1797, Bousquet and Bouchard (2019: 116) recommended that Cnodalum Agassiz, 1846 be placed on the Official Index of Rejected and Invalid Generic Names in Zoology in order to avoid homonymy problems with the name Cnodalum Emeljanov, 1978 [Hemiptera], which is currently used as valid.

Coccimarygmus Ardoin, 1966: 185, 187 [M]. Type species: Paramarygmusmorychoides Fairmaire, 1894, by original designation. Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Cochabambia Marcuzzi, 1985: 184 [F]. Type species: Cochabambiakulzeri Marcuzzi, 1985, by monotypy. Status: senior synonym of Allotriocochabambia Faúndez, Rider & Carvajal, 2014 in Tenebrionidae: incertae sedis. Note: junior homonym of Cochabambia Pirán, 1959 [Hemiptera].

Coeladesmia Reitter, 1916a: 3 [F]. Type species: Metriopusplatynotus Gerstaecker, 1854, by subsequent designation (Gebien 1937a: 656). Status: valid subgenus of Metriopus Solier, 1835 in Pimeliinae: Adesmiini.

Coelocnemis Mannerheim, 1843: 280 [F]. Type species: Coelocnemisdilaticollis Mannerheim, 1843, by subsequent designation (R. Lucas 1920: 194). Status: valid genus in Stenochiinae: Cnodalonini.

Coelocnemodes Bates, 1879b: 474 [M]. Type species: Coelocnemodesstoliczkanus Bates, 1879, by monotypy. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Coeloecetes Blair, 1929b: 384 [M]. Type species: Coeloecetescavernicola Blair, 1929, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Coelolophus Mäklin, 1867: 502 [M]. Type species: Coelolophusbicarinatus Mäklin, 1867, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Coelometopus Solier, 1848: 154, 278 [M]. Type species: Coelometopusclypeatus Solier, 1848 (= Blaps clypeata Germar, 1813), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Coelomorpha Casey, 1890a: 182 [F]. Type species: Coelomorphamaritima Casey, 1890, by monotypy. Status: junior synonym of Coelus Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 371).

Coelopalorus Blair, 1930: 135 [M]. Type species: Palorusfoveicollis Blair, 1930 (= Ulominacarinata Baudi di Selve, 1876), by monotypy. Status: junior synonym of Ulomina Baudi di Selve, 1876 in Tenebrioninae: Palorini. Synonymy: Scupola (2002: 186).

Coelophanes Iablokoff-Khnzorian, 1964: 309 [M]. Type species: Hedyphanesimpressicollis Faldermann, 1837 (= Hedyphaneslaticollis Fischer, 1832), by original designation. Status: junior synonym of Hedyphanes Fischer, 1820 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko (2005: 351).

Coelopleurum Gebien, 1921b: 35 [N]. Type species: Coelopleurumglabratum Gebien, 1921, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Coelosattus Blaisdell, 1927: 166 [M]. Type species: Coelosattusfortineri Blaisdell, 1927 (= Eusattusdilatatus J.L. LeConte, 1851), by monotypy. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 373).

Coelotaxis Horn, 1876b: 200 [F]. Type species: Coelotaxispunctulata Horn, 1876, by subsequent designation (Gebien 1938a: 289). Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 373).

Coelus Eschscholtz, 1829: 5 [M]. Type species: Coelusciliatus Eschscholtz, 1829, by monotypy. Status: valid genus in Pimeliinae: Coniontini.

Colasia Koch, 1965: 127, 131 [F]. Type species: Colasiaakisoides Koch, 1965, by monotypy. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Collariheliofugus Freude, 1960a: 125, 126 [M]. Type species: Euschatiacollaris Germain, 1855, by original designation. Status: valid subgenus of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini.

Colophonesthes Koch, 1954a: 48 [F]. Type species: Colophonesthesmontisatri Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Colparthrum Kirsch, 1866: 204 [N]. Type species: Colparthrumgerstaeckeri Kirsch, 1866, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Colpopatrum Reitter, 1904: 148 [N]. Type species: Opatrumasperipenne Reitter, 1897, by monotypy. Status: valid subgenus of Opatrum Fabricius, 1775 in Blaptinae: Opatrini: Opatrina.

Colpophorinus Escalera, 1914: 336 [M]. Type species: Opatrumgonocephaloides Escalera, 1914, by monotypy. Status: junior synonym of Colpophorus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Opatrina. Synonymy: Español (1954b: 317).

Colpophorus Mulsant & Rey, 1859c: 44 [M]. Type species: Opatrumemarginatum P.H. Lucas, 1847, by subsequent designation (Iwan and Löbl 2008: 269). Status: valid subgenus of Opatrum Fabricius, 1775 in Blaptinae: Opatrini: Opatrina.

Colposcelis Dejean, 1834: 185 [F]. Type species: Tentyrialongicollis Zubkov, 1833, by subsequent designation (Gebien 1937a: 598). Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Colposceloides Schuster, 1940: 20 [M]. Type species: Colposcelislicenti Schuster, 1940, by monotypy. Status: valid subgenus of Colposcelis Dejean, 1834 in Pimeliinae: Tentyriini.

Colposcythis Reitter, 1889b: 113 [F]. Type species: Colposcythiswalteri Reitter, 1889, by monotypy. Status: valid subgenus of Colposcelis Dejean, 1834 in Pimeliinae: Tentyriini.

Colposphena Semenov, 1889: 217 [F]. Type species: Sphenariabrevicollis Solsky, 1882, by subsequent designation (Gebien 1937a: 577). Status: valid genus in Pimeliinae: Tentyriini.

Colpotinoides Kaszab, 1975b: 282, 354 [M]. Type species: Pseudoblapsgebieni Kaszab, 1942, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Colpotinus Fairmaire, 1891c: xvii [M]. Type species: Colpotinussimulator Fairmaire, 1891, by monotypy. Status: valid genus in Blaptinae: Pedinini: Pedinina.

Colpotus Mulsant & Rey, 1853b: 148, 208 [M]. Type species: Colpotusstrigicollis Mulsant & Rey, 1853, by subsequent designation (Gebien 1938a: 313). Status: valid subgenus of Pedinus Latreille, 1797 in Blaptinae: Pedinini: Pedinina.

Comphosida Macquart, 1850: 174 [F]. Type species: Mesostenapunctipennis Solier, 1835 (= Pimeliaangustata Fabricius, 1775), by monotypy. Status: junior synonym of Mesostena Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Lacordaire (1859a: 52).

Compsocula Fairmaire, 1898a: 236 [F]. Type species: Stenogenaapicata Fairmaire, 1896, by subsequent designation (R. Lucas 1920: 199). Status: valid genus in Alleculinae: incertae sedis. Note: placed in Alleculinae by Chatanay (1915a: 526).

Compsomorphus Solier, 1851: 208 [M]. Type species: Compsomorphuselegans Solier, 1851, by original designation. Status: junior synonym of Apocrypha Eschscholtz, 1831 in Tenebrioninae: Apocryphini. Synonymy: Lacordaire (1859b: 433).

Conibiosoma Casey, 1890b: 476 [N]. Type species: Conibiuselongatus Horn, 1870, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Conibius J.L. LeConte, 1851: 145 [M]. Type species: Conibiusseriatus J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 199). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Coniontellus Casey, 1890b: 388 [M]. Type species: Coniontisobesa J.L. LeConte, 1851, by subsequent designation (Casey 1908: 57). Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 373).

Coniontides Casey, 1908: 57, 78 [M]. Type species: Coniontislata J.L. LeConte, 1866, by original designation. Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Doyen (1972: 373).

Coniontis Eschscholtz, 1829: 7 [F]. Type species: Coniontisviatica Eschscholtz, 1829, by subsequent designation (Casey 1908: 57). Status: valid genus in Pimeliinae: Coniontini.

Conipinus J.L. LeConte, 1862: 223 [M]. Type species: Eusattusdubius J.L. LeConte, 1851, by subsequent designation (Gebien 1938a: 284). Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: J.L. LeConte (1866a: 60).

Conisattus Casey, 1895: 614 [M]. Type species: Conisattusrectus Casey, 1895, by monotypy. Status: valid genus in Pimeliinae: Coniontini.

Conoecus Horn, 1885: 159 [M]. Type species: Conoecusovipennis Horn, 1885, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Conophthalmus Quedenfeldt, 1885: 13 [M]. Type species: Conophthalmussetulosus Quedenfeldt, 1885, by monotypy. Status: valid subgenus of Amatodes Dejean, 1834 in Blaptinae: Pedinini: Helopinina. Note: see Robiche (2013: 522) for placement of the genus.

Convexoodescelis Egorov, 2004: 593 [F]. Type species: Platyscelisbrevipennis Kaszab, 1938, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Cophodema Gebien, 1943: 402 [F]. Type species [automatic]: Cophosomahumeridens Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Cophosoma Gebien, 1928.

Cophosoma Gebien, 1928: 219, 222 [N]. Type species: Cophosomahumeridens Gebien, 1928, by monotypy. Status: senior synonym of Cophodema Gebien, 1943 in Stenochiinae: Cnodalonini. Note: junior homonym of Cophosoma Costa, 1849 [Hemiptera].

Copistethus Seidlitz, 1890: 524 [M]. Type species: Cistelaspadix Kiesenwetter, 1861, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Coracostira Fairmaire, 1899d: 220 [F]. Type species: Coracostiraarmipes Fairmaire, 1899, by present designation. Status: junior synonym of Xenostethus Bates, 1868 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1936: 460).

Cordibates Kulzer, 1956b: 909 [M]. Type species: Cordibateschilensis Kulzer, 1956, by original designation. Status: valid genus in Pimeliinae: Thinobatini.

Corinta Koch, 1950c: 36 [F]. Type species: Corintalitoralis Koch, 1950, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Coripera Pascoe, 1866a: 483 [F]. Type species: Adeliumdeplanatum Boisduval, 1835, by original designation. Status: valid genus in Lagriinae: Adeliini.

Cornopterus Koch, 1950c: 40 [M]. Type species: Cornopteruswykehami Koch, 1950, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Cornucistela Campbell, 1980: 133 [F]. Type species: Cornucistelaserrata Campbell, 1980, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Cornugeton Bremer, 2016: 224 [M]. Type species: Platolenesmonilicornis Gebien, 1920, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Corticeus Piller & Mitterpacher, 1783: 87 [M]. Type species: Corticeusunicolor Piller & Mitterpacher, 1783, by monotypy. Status: valid genus and subgenus in Diaperinae: Hypophlaeini.

Coscinoptilix Allard, 1876a: 15 [M]. Type species: Coscinoptilixgracilicornis Allard, 1876, by monotypy. Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Champion (1887: 312). Note: Allard’s original spelling Coscinopter is corrected to Coscinoptilix in the errata for volume 14 of ‘L’Abeille, Journal d’Entomologie’ (at the end of page 36 of the section “Table alphabétique”) and therefore Coscinoptilix is considered the correct original spelling (ICZN 1999, Article 32.5.1.1); see Bousquet et al. (2018: 183).

Cosmogaster Koch, 1956a: 164 [F]. Type species: Anchophthalmusimpressicollis Fairmaire, 1897, by original designation. Status: senior synonym of Kochogaster Kamiński & Raś, 2011 in Blaptinae: Platynotini: Platynotina. Note: junior homonym of Cosmogaster Faust, 1904 [Coleoptera: Curculionidae].

Cosmonota Blanchard, 1842: pl. 14 [F]. Type species: Cosmonotaangustata Blanchard, 1842, by subsequent designation (Gebien 1928: 127). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cossyphodes Westwood, 1851: 168 [M]. Type species: Cossyphodeswollastonii Westwood, 1851, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: Cossyphodina.

Cossyphodinus Wasmann, 1899a: 163 [M]. Type species: Cossyphodinusindicus Wasmann, 1899, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: Paramellonina.

Cossyphodites Brauns, 1901: 91 [M]. Type species: Cossyphodeswoodrooffei Péringuey, 1885, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: Cossyphoditina.

Cossyphus G.-A. Olivier, 1791: 121 [M]. Type species: Lampyrisdepressa Fabricius, 1781, by monotypy. Status: valid genus and subgenus in Lagriinae: Cossyphini.

Costallecula Pic, 1954: 235 [F]. Type species: Costalleculaluteocostata Pic, 1954, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Costatosora Pic, 1934: 32 [F]. Type species: Nemostiradistincticornis Pic, 1911, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Costiferolagria Pic, 1915b: 5 [F]. Type species: Lagriasemialutacea Pic, 1915, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Coxelinus Fairmaire, 1869b: 202 [M]. Type species: Coxelinusstricticollis Fairmaire, 1869, by subsequent designation (Gebien 1941: 826). Status: valid genus in Lagriinae: Lupropini.

Craniosphena Koch, 1962a: 45, 54, 143 [F]. Type species: Himatismusjusti Fairmaire, 1880, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Craniotus J.L. LeConte, 1851: 142 [M]. Type species: Craniotuspubescens J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Cratidus J.L. LeConte, 1862: 239 [M]. Type species: Amphidoraosculans J.L. LeConte, 1851, by monotypy. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Cratopus Eschscholtz, 1831: 5, 8 [M]. Type species: Cratopuscastaneus Eschscholtz, 1831, by monotypy. Status: senior synonym of Scelosodis Solier, 1835 in Pimeliinae: Tentyriini. Note: junior homonym of Cratopus Schönherr, 1826 [Coleoptera: Curculionidae].

†Cretaceites Wang, 1997: 203 [M]. Type species: Cretaceitesjingxiensis Wang, 1997, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Lower Cretaceous deposits (China).

Cribrasida Reitter, 1917a: 11, 38 [F]. Type species: Asidagrandipalpis Allard, 1869, by subsequent designation (F. Soldati 2008: 32). Status: junior synonym of Elongasida Escalera, 1906 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 290).

Crististibes Koch, 1963: 62 [M]. Type species: Planostibesbinodosus Gebien, 1920, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Crossoscelis Gebien, 1914a: 52 [F]. Type species: Crossoscelisclauda Gebien, 1914, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Masumoto (1999a: 123).

Cruracurvamtenebrio Robiche, 2019b: 97 [M]. Type species: Tenebriokamgangi Robiche, 2019, by monotypy. Status: valid subgenus of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini.

Cryphaeus Klug, 1833: 89 [M]. Type species: Cryphaeusaries Klug, 1833, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Toxicina.

Crypsinous Fairmaire, 1891b: 262 [M]. Type species: Crypsinousacutispina Fairmaire, 1891, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Crypsis C.O. Waterhouse, 1877: 73 [F]. Type species: Crypsisviolaceipennis C.O. Waterhouse, 1877, by monotypy. Status: valid genus in Diaperinae: Leiochrinini.

Cryptadius Fairmaire, 1894f: 395 [M]. Type species [automatic]: Crypsinousacutispina Fairmaire, 1891, by monotypy. Status: junior synonym of Crypsinous Fairmaire, 1891 in Tenebrioninae: Amarygmini. Note: unnecessary replacement name for Crypsinous Fairmaire, 1891 (as “Crypsinon”); junior homonym of Cryptadius J.L. LeConte, 1851 [Coleoptera: Tenebrionidae: Pimeliinae: Edrotini].

Cryptadius J.L. LeConte, 1851: 140 [M]. Type species: Cryptadiusinflatus J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Cryptasida Koch, 1962a: 129 [F]. Type species: Asidanamaqua Péringuey, 1899, by original designation. Status: valid genus in Pimeliinae: Asidini.

Crypticanus Fairmaire, 1897f: 119 [M]. Type species: Crypticanuscuneatus Fairmaire, 1897 (= Melanopterusedwarsii Mulsant & Rey, 1854), by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Crypticocatops Kaszab, 1975c: 102 [M]. Type species: Platydemacatopoides Fairmaire, 1896, by original designation. Status: valid subgenus of Microcrypticus Gebien, 1921 in Diaperinae: Crypticini.

Crypticoides Fairmaire, 1898d: 389 [M]. Type species: Crypticoidesmellyi Fairmaire, 1898, by monotypy. Status: junior synonym of Oxycara Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Koch (1960: 382).

Crypticomorpha Casey, 1908: 81, 140 [F]. Type species: Coniontistenuis Casey, 1908, by monotypy. Status: junior synonym of Coniontis Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Aalbu et al. (2002: 487).

Crypticopsis Antoine, 1945: 270 [F]. Type species: Crypticuscorticeus Fairmaire, 1871, by original designation. Status: valid subgenus of Crypticus Latreille, 1816 in Diaperinae: Crypticini.

Crypticus Latreille, 1816: 298 [M]. Type species: Helopsglaber Fabricius, 1775 (= Tenebrioquisquilius Linnaeus, 1761), by monotypy. Status: valid genus and subgenus in Diaperinae: Crypticini.

Cryptobates Fairmaire, 1882a: 231 [M]. Type species: Cryptobatesrubigineus Fairmaire, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cryptobatoides Kaszab, 1941a: 2, 15 [F]. Type species: Cryptobatoidesopaca Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cryptobrachys Kaszab, 1941a: 4, 14 [M]. Type species: Cryptobatescrassecostatus Fairmaire, 1898, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cryptocarpes Koch, 1952a: 191 [M]. Type species: Cryptocarpeselongatus Koch, 1952, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini. Note: the First Reviser (Cryptocarpes Koch, 1952 versus Lornamus Koch, 1952) is Endrödy-Younga (1996: 21).

Cryptochile Latreille, 1828: 576 [F]. Type species: Pimeliamaculata Fabricius, 1781, by monotypy. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina. Note: discovery of the older unused name Phymatium Billberg, 1820 threatens the validity of Cryptochile Latreille, 1828 and its associated family-group names currently used as valid; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to conserve usage of Cryptochile Latreille, 1828.

Cryptogenius Solier, 1843: 37, 122 [M]. Type species: Cryptogeniusdentatus Solier, 1843, by original designation. Status: senior synonym of Phrynocolus Lacordaire, 1859 in Pimeliinae: Sepidiini: Molurina. Note: junior homonym of Cryptogenius Westwood, 1842 [Coleoptera: Hybosoridae].

Cryptoglossa Solier, 1837a: 638, 680 [F]. Type species: Cryptoglossabicostata Solier, 1837, by monotypy. Status: valid genus in Pimeliinae: Cryptoglossini.

†Cryptohelops Nabozhenko & Kirejtshuk, 2014: 68 [M]. Type species: Cryptohelopsmenaticus Nabozhenko & Kirejtshuk, 2014, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina. Note: described from Middle-Upper Paleocene deposits (France).

Cryptomysia Pic, 1954: 260 [F]. Type species: Cryptomysiaminor Pic, 1954, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Cryptops Solier, 1851: 235 [M]. Type species: Cryptopsulomoides Solier, 1851 (= Tenebriodiaperinus Panzer, 1797), by original designation. Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Gemminger in Gemminger and Harold (1870: 1962). Note: junior homonym of Cryptops Leach, 1814 [Chilopoda].

Cryptostenophanes Kaszab, 1941a: 5, 12 [M]. Type species: Cryptostenophanesborneensis Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cryptotrophus Gistel, 1848a: xi [M]. Type species [automatic]: Pimeliamaculata Fabricius, 1781, by monotypy. Status: junior synonym of Cryptochile Latreille, 1828 in Pimeliinae: Cryptochilini: Cryptochilina. Note: unnecessary replacement name for Cryptochile Latreille, 1828.

Cryptozoon Schaufuss, 1882: 47 [N]. Type species: Cryptozooncivile Schaufuss, 1882, by subsequent designation (Bousquet et al. 2018: 284). Status: valid genus in Diaperinae: Gnathidiini: Anopidiina. Note: transferred from Colydiidae (see Doyen and Lawrence 1979: 366).

Csikiola Kaszab, 1955a: 493 [F]. Type species: Csikiolathesileiformis Kaszab, 1955, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Csiro G.S. Medvedev & Lawrence, 1984: 563 [F]. Type species: Hyocissubparallelus Champion, 1894, by original designation. Status: valid genus and subgenus in Diaperinae: Hyociini: Hyociina.

Cteisa Solier, 1835a: 242 [F]. Type species: Cteisahirta Solier, 1835, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Cteisodella Novák, 2020f: 50 [F]. Type species: Cteisodellaassamica Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Cteisodes Borchmann, 1932a: 307 [F]. Type species: Cteisodessericea Borchmann, 1932, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Cteniopinus Seidlitz, 1896: 200 [M]. Type species: Cistelaaltaica Gebler, 1830, by subsequent designation (R. Lucas 1920: 212). Status: valid genus and subgenus in Alleculinae: Cteniopodini.

Ctenioposomus Reitter, 1906b: 131 [M]. Type species: Cteniopusfrater Reitter, 1903, by subsequent designation (Novák and Pettersson 2008: 330). Status: valid subgenus of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: the alternative original spelling Ctenoposomus, used by Reitter (1906b: 135), was rejected by Bousquet et al. (2015: 140) who acted as the First Revisers.

Cteniopus Solier, 1835: 245, 246 [M]. Type species: Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Westwood 1838: 32). Status: valid genus and subgenus in Alleculinae: Cteniopodini.

Ctenogria Borchmann, 1916a: 48, 101 [F]. Type species: Ctenogriavermiculata Borchmann, 1916, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Ctesicles Champion, 1896: 7 [M]. Type species: Ctesiclesinsularis Champion, 1896, by subsequent designation (R. Lucas 1920: 214). Status: junior synonym of Diastolinus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Blapstinina. Synonymy: Ivie and Hart (2016: 468).

Ctimene Bates, 1873e: 359 [F]. Type species: Ctimenebreweri Bates, 1873, by monotypy. Status: senior synonym of Mitrothorax Carter, 1914 in Tenebrioninae: Heleini: Cyphaleina. Note: junior homonym of Ctimene Boisduval, 1832 [Lepidoptera].

Cuemus Bouchard, 2000: 96 [M]. Type species: Cuemusmonteithi Bouchard, 2000, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Cuphotes Champion, 1887: 332 [F]. Type species [automatic]: Spheniscuserotyloides W. Kirby, 1819, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: replacement name for Spheniscus W. Kirby, 1819.

Curimosphena Gebien, 1920: 42, 43 [F]. Type species: Himatismusvillosus Haag-Rutenberg, 1870, by original designation. Status: junior synonym of Imatismus Dejean, 1834 in Pimeliinae: Tentyriini. Synonymy: Bouchard et al. (2007: 386).

Curtolyprops Pic, 1917a: 11 [M]. Type species: Curtolypropslatipennis Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Curtopeltoides Pic, 1916d: 14 [M]. Type species: Curtopeltoidesrufescens Pic, 1916, by subsequent designation (Gebien 1940: 1062). Status: valid genus in Tenebrioninae: Ulomini.

Cybopiestes Reitter, 1917b: 148, 150 [M]. Type species: Cybopiestescsikii Reitter, 1917, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cybostira Borchmann, 1936: 429, 430 [F]. Type species: Cybostiracaligata Borchmann, 1936, by original designation. Status: valid subgenus of Epicydes Champion, 1889 in Lagriinae: Lagriini: Statirina.

Cybotus Casey, 1890b: 482 [M]. Type species: Blapstinusestriatus J.L. LeConte, 1878, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Cychrachna Koch, 1950b: 341, 361 [F]. Type species: Cychrachnacarcharoides Koch, 1950, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Cychrochile Koch, 1953b: 160 [F]. Type species: Cychrochileerodioides Koch, 1953, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Cyclobiomorphus Pic, 1916c: 1 [M]. Type species: Cyclobiomorphusundulatus Pic, 1916, by subsequent designation (Gebien 1940: 420). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cyclobium Pic, 1916a: 11 [N]. Type species: Cyclobiumvesiculiferum Pic, 1916, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Cyclocnera Leo, 2018: 50 [F]. Type species: Cyclocneraazarovi Leo, 2018, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Cyclonesus Fairmaire, 1896c: 104 [M]. Type species: Cyclonesusparvicollis Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cyclophanes Carter, 1913a: 92 [M]. Type species: Cyclophanesvariegatus Carter, 1913, by subsequent designation (R. Lucas 1920: 217). Status: junior synonym of Hemicyclus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 497).

Cyclosattus Casey, 1892: 710 [M]. Type species: Eusattuswebsteri Casey, 1891 (= Celibecostata Solier, 1848), by monotypy. Status: junior synonym of Saragus Erichson, 1842 in Tenebrioninae: Heleini: Heleina. Synonymy: Doyen et al. (1990: 244). Note: the type species was originally described from Colorado [United States of America] in error, Doyen et al. (1990: 244) pointed out that Eusattuswebsteri Casey, 1891 is a synonym of the Australian species Celibecostata Solier, 1848.

Cylindrinotus Faldermann, 1837: 73 [M]. Type species: Cylindrinotuslugubris Faldermann, 1837 (= Helopsfemoratus Faldermann, 1837), by subsequent designation (Gebien 1943: 425). Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina. Note: Gebien (1943: 425) selected Helopsfemoratus Faldermann, 1837 as type species, which is not an originally included species in Cylindrinotus; however, since Gebien listed Cylindrinotuslugubris Faldermann, 1837 (a species originally included in Cylindrinotus) as a synonym of Helopsfemoratus Faldermann, 1837, he is deemed to have designated the latter as type species (ICZN 1999, Article 69.2.2); the original combination of the accepted name of the type species, Helopsfemoratus Faldermann, 1837, is a junior primary homonym of Helopsfemoratus Fabricius, 1798.

Cylindronotus Agassiz, 1846b: 111 [M]. Type species [automatic]: Cylindrinotuslugubris Faldermann, 1837 (= Helopsfemoratus Faldermann, 1837), by subsequent designation (Gebien 1943: 425). Status: junior synonym of Cylindrinotus Faldermann, 1837 in Tenebrioninae: Helopini: Cylindrinotina. Note: unjustified emendation of Cylindrinotus Faldermann, 1837, not in prevailing usage.

Cylindrosia Gebien, 1922b: 289 [F]. Type species: Cylindrosiafoveifrons Gebien, 1922, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Cylindrosora Borchmann, 1936: 237, 375 [F]. Type species: Soramacer Borchmann, 1930, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Cylindrostira Borchmann, 1936: 237, 374 [F]. Type species: Casnonideacorporaali Borchmann, 1929, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Cylindrothorus Solier, 1843: 4, 50 [M]. Type species: Cylindrothoruspilosus Solier, 1843, by original designation. Status: valid genus and subgenus in Alleculinae: Alleculini: Mycetocharina.

Cymatodes Agassiz, 1846b: 112 [M]. Type species [automatic]: Helopsundatus Fabricius, 1792 (= Erotylusnebulosus Fabricius, 1781), by monotypy. Status: junior synonym of Cymatothes Dejean, 1834 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Cymatothes Dejean, 1834, not in prevailing usage; the older name Cymatodes W. Kirby & Spence, 1828 [Coleoptera: Curculionidae] is not nomenclaturally available.

Cymatothes Dejean, 1834: 208 [M]. Type species: Helopsundatus Fabricius, 1792 (= Erotylusnebulosus Fabricius, 1781), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Cymbeba Pascoe, 1866a: 483 [F]. Type species: Cymbebadissimilis Pascoe, 1866, by monotypy. Status: valid genus in Lagriinae: Adeliini. Note: as mentioned by Matthews and Bouchard (2008: 350) the type species was erroneously described from Australia; the genus Cymbeba Pascoe, 1866 is endemic to New Caledonia.

Cynaeus J.L. LeConte, 1862: 233 [M]. Type species: Platydemaangusta J.L. LeConte, 1851, by original designation. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Cyphaleus Westwood, 1841a: 43 [M]. Type species: Helopsrugosus Gray, 1831, by subsequent designation (Duponchel 1844a: 547). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: nomen protectum (see Appendix 2); Cyphaleus is sometimes attributed to Hope (1841: 126) in the literature (e.g., Matthews 2019: 631); however, Hope did not describe the genus and there is no indication in his publication that “Cyphaleusrugosus Hope”; the only species he included in the genus is the same as Helopsrugosus Gray, 1831 (“Cyphaleus Hope” was treated as an unavailable name by Neave (1939: 939)); the First Reviser (Cyphaleus Westwood, 1841 versus Chartopteryx Westwood, 1841) is Matthews (1992: 490).

Cyphelops Fairmaire, 1901a: 73 [M]. Type species: Cyphelopsinflatus Fairmaire, 1901, by monotypy. Status: junior synonym of Miotodera Fairmaire, 1901 in Stenochiinae: Stenochiini. Synonymy: Chatanay (1915a: 526).

Cyphogenia Solier, 1837a: 638, 677 [F]. Type species: Tenebrioauritus Pallas, 1781, by monotypy. Status: valid genus and subgenus in Pimeliinae: Akidini.

Cyphonotus Guérin-Méneville, 1831a: pl. 5 [M]. Type species: Cyphonotusdromedarius Guérin-Méneville, 1831, by monotypy. Status: senior synonym of Homocyrtus Dejean, 1834 in Tenebrionidae: incertae sedis. Note: junior homonym of Cyphonotus Fischer, 1823 [Coleoptera: Scarabaeidae].

Cyphostethe Marseul, 1866: xxxix [F]. Type species: Himatismusferrugineus Marseul, 1866, by subsequent designation (Gebien 1937a: 573). Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Cyphostethoides Löbl & Merkl in Löbl et al., 2020: 3 [M]. Type species: Cyphostethebrunnea Kaszab, 1962, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini. Note: name first proposed by Kaszab (1979c: 273) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 244) designated Cyphostethebrunnea Kaszab, 1962 as the type species of Kaszab’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Cyptus Gerstaecker, 1871: 61 [M]. Type species: Cyptusscabrosus Gerstaecker, 1871, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Cyriogeton Pascoe, 1871: 356 [M]. Type species: Cyriogetoninsignis Pascoe, 1871, by monotypy. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Cyrta Lacordaire, 1859: 724 [F]. Type species [automatic]: Cirsastriaticollis P.H. Lucas, 1857, by monotypy. Status: junior synonym of Cirsa P.H. Lucas, 1857 in Pimeliinae: Tentyriini. Note: unjustified emendation of Cirsa P.H. Lucas, 1857, not in prevailing usage.

Cyrtoderes Dejean, 1834: 181 [M]. Type species: Sepidium lacunosum Thunberg, 1784 (= Tenebriocristatus DeGeer, 1778), by subsequent designation (Bousquet and Bouchard 2013a: 45). Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Cyrtomius Casey, 1907: 379 [M]. Type species: Cyrtomiuscavicauda Casey, 1907 (= Epitragusplicatus Champion, 1884), by original designation. Status: valid genus and subgenus in Pimeliinae: Epitragini.

Cyrtosoma Perty, 1830: 59 [N]. Type species: Cyrtosomaunicolor Perty, 1830, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cyrtostrongylium Blair, 1929a: 246 [N]. Type species: Cyrtostrongyliumrhysopaussoides Blair, 1929 (= Macrosynopticuscostatus Pic, 1922), by monotypy. Status: junior synonym of Macrosynopticus Pic, 1922 in Tenebrioninae: Amarygmini. Synonymy: Pic (1930d: 12).

Cyrtotyche Pascoe, 1866a: 469 [F]. Type species: Cyrtotychesatanas Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Cyrtotyctus Kolbe, 1897a: 241 [M]. Type species: Cyrtotyctusosdaroides Kolbe, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Dactylocalcar Gebien, 1938b: 46 [N]. Type species: Dactylocalcarcaecus Gebien, 1938, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Daedrosis Bates, 1868: 266 [F]. Type species: Daedrosiscrenatostriata Bates, 1868, by subsequent designation (R. Lucas 1920: 224). Status: valid genus in Lagriinae: Adeliini.

Dailognatha Steven, 1828: 88 [F]. Type species: Tentyriaaequalis Tauscher, 1812, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Dalmanius Bremer, 2001b: 90, 95 [M]. Type species: Dalmaniusperegrinus Bremer, 2001, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Damatris Laporte, 1840: 224 [F]. Type species: Tetraphyllusformosus Laporte & Brullé, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Danielomira Weise, 1974: 119 [F]. Type species: Isomiracantabrica Weise, 1974, by original designation. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Danodema Gebien, 1925c: 570 [F]. Type species: Danodemasubcalva Gebien, 1925, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Daochus Champion, 1886: 139 [M]. Type species: Daochusmandibularis Champion, 1886, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Dasus Motschulsky, 1845a: 78 [M]. Type species: Opatrumfuscum Herbst, 1793 (= Opatrumrusticum G.-A. Olivier, 1812), by original designation. Status: junior synonym of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Bedel (1894: 154).

Dasyplonyx Bremer, 2014a: 37 [M]. Type species: Cyriogetonmaculosus Pic, 1922, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Dasytoxystropus Pic, 1921b: 12 [M]. Type species: Dasytoxystropussubparallelus Pic, 1921, by original designation. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Dauresia Ferrer, 2001: 187 [F]. Type species: Dauresiamontisusti Ferrer, 2001, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Davaona Borchmann, 1930a: 442, 524 [F]. Type species: Casnonideaperforata Borchmann, 1913, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Debeauxiella Bouchard & Bousquet, new subgenus [F]. Type species: Pachyceraangulophthalma Koch, 1943, by present designation. Status: valid subgenus of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: Koch (1943a: 524, 546) introduced the new subgenus name Debeauxiella for three nominal species, but unfortunately did not designate a type species; the subgenusDebeauxiella, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Pachyceraangulophthalma Koch, 1943 as type species and referring to Koch (1943a: 524) for the character states that characterise and differentiate Debeauxiella.

Dechius Pascoe, 1866a: 455 [M]. Type species: Dechiusaphodioides Pascoe, 1866 (= Tenebriocancellatus Montrouzier, 1860), by monotypy. Status: junior synonym of Scotoderus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Bates (1873f: 473).

Dechiustes Blair, 1940: 137 [M]. Type species: Dechiustescarolinensis Blair, 1940, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Decialma Pascoe, 1869: 288, 291 [F]. Type species: Decialmatenuitarsis Pascoe, 1869, by monotypy. Status: junior synonym of Olisthaena Erichson, 1842 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1913a: 62).

Decoriplus Louw, 1979: 117, 120 [M]. Type species: Psammodespictus Haag-Rutenberg, 1871, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Delognatha Agassiz, 1846b: 116, 118 [F]. Type species [automatic]: Tentyriaaequalis Tauscher, 1812, by monotypy. Status: junior synonym of Dailognatha Steven, 1828 in Pimeliinae: Tentyriini. Note: unjustified emendation of Dailognatha Steven, 1828, not in prevailing usage; suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy and placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the ICZN (2010b, Opinion 2250).

Delognatha Lacordaire, 1859a: 315 [F]. Type species: Delognathalacordairei Lacordaire, 1859, by subsequent designation (Gebien 1940: 756). Status: valid genus in Phrenapatinae: Phrenapatini. Note: the senior homonym Delognatha Agassiz, 1846, an unjustified emendation of Dailognatha Steven, 1828 [Coleoptera: Tenebrionidae: Pimeliinae: Tentyriini], was placed on the Official Index of Rejected and Invalid Generic Names in Zoology and suppressed for the purposes of both the Principle of Priority and the Principle of Homonymy by the ICZN (2010b, Opinion 2250).

Delonurops Reitter, 1922a: 24, 25 [M]. Type species: Entomogonusclavimanus Reitter, 1903, by subsequent designation (Iablokoff-Khnzorian 1964: 304). Status: valid subgenus of Entomogonus Solier, 1848 in Tenebrioninae: Helopini: Helopina.

Delopygus J.L. LeConte, 1866b: 129 [M]. Type species: Delopyguscrenatus J.L. LeConte, 1866, by monotypy. Status: junior synonym of Eutochia J.L. LeConte, 1862 in Tenebrioninae: Ulomini. Synonymy: Horn (1870: 372).

Dema Gistel, 1848a: xi [F]. Type species [automatic]: Opatrumclathratum Fabricius, 1787, by monotypy. Status: junior synonym of Opatrinus Dejean, 1821 in Blaptinae: Platynotini: Platynotina. Note: unnecessary replacement name for OpatrinusDejean 1821.

Demtrius Broun, 1895: 243 [M]. Type species: Demtriuscarinulatus Broun, 1895, by monotypy. Status: valid genus in Tenebrioninae: Titaenini. Note: placed in Titaenini by Matthews (2003b: 13).

Dendarophylan Español, 1945: 312, 313 [M]. Type species: Phylan pardoi Español, 1945, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Dendaroscelis Reitter, 1904: 79 [F]. Type species: Dendarusserripes Reitter, 1904, by monotypy. Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Dendarus Dejean, 1821: 65 [M]. Type species: Helopstristis Rossi, 1790, by subsequent designation (Blanchard 1844: pl. 48). Status: valid genus and subgenus in Blaptinae: Dendarini: Dendarina. Note: as pointed out by Silfverberg (1984: 59) and Löbl et al. (2008a: 41) the Helopstristis Herbst of Laporte (1840) (= Pandaruscoarcticollis Mulsant, 1854) was used as the type species of Dendarus (e.g., Gebien 1938a: 299) because Helopstristis Rossi, 1790 was treated as a nomen dubium; however, the Helopstristis Herbst of Laporte (1840), in addition to being nomenclaturally unavailable, was not originally included in Dendarus Dejean, 1821 and therefore cannot be used as the type species; the earliest, and valid, type species designation is that of Blanchard (1844: pl. 48, as “Pedinus (Dendarus) tristis Rossi”).

Dendroscopius Gistel, 1848a: 125 [M]. Type species [automatic]: Trogossitathoracica Fabricius, 1792, by monotypy. Status: junior synonym of Bius Dejean, 1834 in Tenebrioninae: Tenebrionini. Note: unnecessary replacement name for Bius Dejean, 1834.

Dengitha Reitter, 1887b: 516 [F]. Type species: Dengithalutea Reitter, 1887, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Dentatoploedipus Kaszab, 1984: 355, 383 [M]. Type species: Dentatoploedipussembilanicus Kaszab, 1984, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Dentibirus G.S. Medvedev, 1968a: 170, 204 [M]. Type species: Heliopatespusillus Ménétriés, 1849, by original designation. Status: junior synonym of Cabirutus Strand, 1929 in Blaptinae: Pedinini: Pedinina. Synonymy: Kamiński and Iwan (2017: 595).

Deplanchesia Fauvel, 1860: 310 [F]. Type species: Deplanchesiametallescens Fauvel, 1860, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Deretus Gahan, 1900: 10 [M]. Type species: Deretusdenticollis Gahan, 1900, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Deriles Motschulsky, 1872: 24 [M]. Type species: Upisexcavata Herbst, 1797, by original designation. Status: valid subgenus of Amenophis J. Thomson, 1858 in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Derilis, used by Motschulsky (1872: 27).

Derispia Lewis, 1894: 389 [F]. Type species: Diaperismaculipennis Marseul, 1876, by original designation. Status: valid genus in Diaperinae: Leiochrinini.

Derispiella Kaszab, 1961a: 357, 364 [F]. Type species: Derispiellahingstoni Kaszab, 1961, by original designation. Status: valid genus in Diaperinae: Leiochrinini.

Derispiola Kaszab, 1946a: 30, 115 [F]. Type species: Derispiolafruhstorferi Kaszab, 1946, by original designation. Status: valid genus in Diaperinae: Leiochrinini. Note: Derispiola was used earlier by Gebien (1940: 434) but is unavailable from that date because it was published without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1).

Derispiolina Kaszab, 1979b: 279 [F]. Type species: Derispiolinapterolomoides Kaszab, 1979, by original designation. Status: junior synonym of Falsotithassa Pic, 1934 in Lagriinae: Lupropini. Synonymy: Schawaller (2000: 2).

Derolagria Borchmann, 1916a: 61 [F]. Type species: Lagriaplicatula Borchmann, 1909, by subsequent designation (Borchmann 1936: 151). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Deroplatus Solier, 1851: 133 [M]. Type species: Geoboruscostatus Blanchard, 1842, by subsequent designation (Gebien 1937a: 563). Status: junior synonym of Geoborus Blanchard, 1842 in Pimeliinae: Epitragini. Synonymy: Lacordaire (1859a: 77).

Derosalax Gebien, 1926: 85 [M]. Type species: Derosalaxbruchi Gebien, 1926, by monotypy. Status: valid genus in Pimeliinae: Trilobocarini.

Derosimus Fairmaire, 1904a: 62 [M]. Type species: Derosimusquadricollis Fairmaire, 1904, by monotypy. Status: junior synonym of Rhypasma Pascoe, 1862 in Lagriinae: Belopini. Synonymy: Blair (1935a: 104).

Derosphaerius Westwood, 1881: 362 [M]. Type species: Derosphaeriusanthracinus Westwood, 1881, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Derosphaerus J. Thomson, 1858: 99 [M]. Type species: Derosphaerusglobicollis J. Thomson, 1858, by subsequent designation (R. Lucas 1920: 233). Status: valid genus in Stenochiinae: Cnodalonini. Note: nomen protectum (see Bouchard and Bousquet 2020b: 6).

Derostethe Koch, 1950b: 285 [F]. Type species: Himatismuspunctatissimus Haag-Rutenberg, 1870, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini.

Derostira Fairmaire, 1897b: 388 [F]. Type species: Derostiracrenulipennis Fairmaire, 1897, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Derostirostaius Borchmann, 1936: 477 [M]. Type species: Derostiralaticollis Pic, 1913, by original designation. Status: valid subgenus of Staius Fairmaire, 1896 in Lagriinae: Lagriini: Statirina. Note: first proposed by Pic (1934: 32) without original type species designation; it is therefore unavailable from that date (ICZN 1999, Article 13.3).

Derostrophus Fairmaire, 1888a: 186 [M]. Type species [automatic]: Derosphaeriusanthracinus Westwood, 1881, by monotypy. Status: junior synonym of Derosphaerius Westwood, 1881 in Pimeliinae: Tentyriini. Note: unnecessary replacement name for Derosphaerius Westwood, 1881.

Desertosochrus Koch, 1958: 149 [M]. Type species: Drosochruspiligaster Koch, 1958, by original designation. Status: valid subgenus of Drosochrus Erichson, 1843 in Blaptinae: Pedinini: Helopinina.

Diabolicobates Pic, 1930b: 30 [M]. Type species: Diabolicobatescornutus Pic, 1930 (= Mechanetesplatitubera Kaszab, 1941), by monotypy. Status: junior synonym of Mechanetes C.O. Waterhouse, 1887 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1941a: 43).

Diachoriops Ando, 2020: 8 [M]. Type species [automatic]: Schizommacucumericola Gebien, 1921, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Schizomma Gebien, 1921.

Diacis Koch, 1955a: 105 [M]. Type species: Trachynotusregalis Haag-Rutenberg, 1876, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Diaclina Jacquelin du Val, 1861: 296 [F]. Type species: Tenebriochrysomelinus Herbst, 1799 (= Tenebriotestudineus Piller & Mitterpacher, 1783), by original designation. Status: valid genus in Tenebrioninae: Alphitobiini.

Diaderma Koch, 1960: 399, 405 [N]. Type species: Diadermapunticum Koch, 1960, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Diaperis Geoffroy, 1762: 337 [F]. Type species: Chrysomelaboleti Linnaeus, 1758, by subsequent monotypy (Müller 1776: 74; see ICZN 1994a, Opinion 1754). Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: originally proposed without included nominal species; placed on the Official List of Generic names in Zoology by the ICZN (1994a, Opinion 1754).

Diaphanidus Reitter, 1900b: 299, 301 [M]. Type species: Arthrodeisantennatus Reitter, 1894, by subsequent designation (R. Lucas 1920: 235). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Diaspirus Matthews, 1998: 704, 759 [M]. Type species: Diaspirusbellendenus Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Diastanus Fairmaire, 1902b: 338 [M]. Type species: Diastanusnitidiventris Fairmaire, 1902, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Diastixus Allard, 1876a: 5 [M]. Type species: Helopspunctipennis P.H. Lucas, 1846 (= Helopsheteromorphus P.H. Lucas, 1846), by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Nesotes Allard, 1876 in Tenebrioninae: Helopini: Helopina. Synonymy: Antoine (1949: 134).

Diastoleus Solier, 1838b: 8, 67 [M]. Type species: Scotobiuscollaris Guérin-Méneville, 1834, by original designation. Status: valid genus in Tenebrioninae: Scotobiini.

Diastolinus Mulsant & Rey, 1859a: 70, 74 [M]. Type species: Blapsclathrata Fabricius, 1792, by subsequent designation (R. Lucas 1920: 236). Status: valid genus in Blaptinae: Opatrini: Blapstinina. Note: the First Revisers (Diastolinus Mulsant & Rey, 1859 versus Sellio Mulsant & Rey, 1859) are Ivie and Hart (2016: 468).

Diceroderes Solier, 1841b: 30, 46 [M]. Type species: Diceroderesmexicanus Solier, 1841, by original designation. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Dichastops Gerstaecker, 1871: 63 [M]. Type species: Dichastopssubaeneus Gerstaecker, 1871, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Dichillesthes Reitter, 1916d: 156 [F]. Type species: Dichilluscordicollis Reitter, 1886, by monotypy. Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillinus Reitter, 1916d: 156, 161 [M]. Type species: Tageniapusilla Ménétriés, 1849, by subsequent designation (G.S. Medvedev 1975: 599). Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillocerus Reitter, 1916d: 155, 167 [M]. Type species: Hyperopspertusus Kiesenwetter, 1861, by subsequent designation (G.S. Medvedev 1975: 603). Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillodontus Reitter, 1916d: 156, 166 [M]. Type species: Dichillusadriani Reitter, 1916, by monotypy. Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillomessor Reitter, 1916d: 155 [M]. Type species: Dichillushimalayanus Fairmaire, 1896, by subsequent designation (G.S. Medvedev 1975: 594). Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Dichillus Jacquelin du Val, 1860: 253 [M]. Type species: Tageniaminuta Solier, 1838, by original designation. Status: valid genus and subgenus in Pimeliinae: Stenosini: Dichillina.

Dichomma Solier, 1835b: 253, 271 [N]. Type species: Dichommamaillei Solier, 1835 (= Tentyriadardana Steven, 1828), by subsequent designation (Jacquelin du Val 1860: 249). Status: valid genus in Pimeliinae: Tentyriini.

Dichotymus Fairmaire, 1891f: ccxcv [M]. Type species: Dichotymusstriatipennis Fairmaire, 1891, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Dichromma Mulsant & Rey, 1855: 94 [N]. Type species: Pandarinusforaminosus Mulsant & Rey, 1855, by monotypy. Status: junior synonym of Paroderus Mulsant & Rey, 1854 in Blaptinae: Dendarini: Dendarina. Synonymy: Reitter (1904: 89, through placement of the type species in Paroderus Mulsant & Rey, 1854).

Dichtha Haag-Rutenberg, 1871: 39 [F]. Type species: Cryptogeniusinflatus Gerstaecker, 1854, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Dicraeosis Gebien, 1911a: 355 [F]. Type species [automatic]: Dicraeusbacillus Marseul, 1876, by monotypy. Status: junior synonym of Stenochinus Motschulsky, 1860 in Stenochiinae: Cnodalonini. Note: replacement name for Dicraeus Marseul, 1876.

Dicraeus Marseul, 1876: 103 [M]. Type species: Dicraeusbacillus Marseul, 1876, by monotypy. Status: junior synonym of Stenochinus Motschulsky, 1860 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1925a: 79). Note: junior homonym of Dicraeus Loew, 1873 [Diptera].

Dictysomorphus Pic, 1921d: 24 [M]. Type species: Dictysomorphusdentaticornis Pic, 1921, by monotypy. Status: junior synonym of Ceropria Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Synonymy: Kaszab (1983a: 132).

Dictysus Rye, 1874: 281 [M]. Type species [automatic]: Dietysusconfusus Pascoe, 1866 (= Cnodalonmetallicum Perty, 1831), by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Note: unjustified emendation for Dietysus Pascoe, 1866, not in prevailing usage.

Dicyrtodes Matthews, 1998: 706, 740 [M]. Type species: Dicyrtodesarneius Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Dicyrtus Duponchel, 1844a: 5 [M]. Type species: Dicyrtusgibbosus Duponchel, 1844, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Diemenoma Matthews, 1998: 707, 710 [F]. Type species: Adeliumcommodum Pascoe, 1869, by original designation. Status: valid genus in Lagriinae: Adeliini.

Diesia Fischer, 1820: pl. 14 [F]. Type species: Diesiaquadridentata Fischer, 1820, by subsequent designation (Hope 1841: 118). Status: valid genus in Pimeliinae: Pimeliini.

Diesiola Skopin, 1961a: 397, 398 [F]. Type species: Diesiaquadridentata Fischer, 1820, by original designation. Status: junior synonym of Diesia Fischer, 1820 in Pimeliinae: Pimeliini. Synonymy: Skopin (1971: 328).

Diestecopus Solier, 1848: 152, 194 [M]. Type species: Diestecopuserodioides Solier, 1848, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Diestesoma Péringuey, 1904: 271 [N]. Type species: Diestesomapulchrum Péringuey, 1904, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Diestica Pascoe, 1868: xii [F]. Type species: Diesticaviridipennis Pascoe, 1868, by monotypy. Status: junior synonym of Poecilesthus Dejean, 1834 in Stenochiinae: Stenochiini. Synonymy: Gebien (1911b: 589).

Dietomorpha Reymond, 1938: 143 [F]. Type species: Dietomorphapardalis Reymond, 1938, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Dietopsis Solier, 1835a: 236 [F]. Type species: Alleculasolieri Laporte, 1840, by subsequent monotypy (Laporte 1840: 243). Status: valid subgenus of Allecula Fabricius, 1801 in Alleculinae: Alleculini: Alleculina. Note: originally proposed without included nominal species; Laporte (1840: 243), by describing one new species Alleculasolieri Laporte, 1840 in association with the subgenus “Dietopsis Sol.”, was the first author to subsequently and expressly include nominal species in Dietopsis (ICZN 1999, Article 67.2.2).

Dietysus Pascoe, 1866a: 486 [M]. Type species: Dietysusconfusus Pascoe, 1866 (= Cnodalonmetallicum Perty, 1831), by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1921a: 409).

Dignamptus J.L. LeConte, 1878: 421 [M]. Type species: Dignamptusstenochinus J.L. LeConte, 1878, by subsequent designation (Bousquet et al. 2018: 325). Status: junior synonym of Talanus Jacquelin du Val, 1857 in Stenochiinae: Talanini. Synonymy: Champion (1887: 321).

Dignathosis Koch, 1958: 75 [F]. Type species: Dignathosisarcana Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Dila Fischer von Waldheim, 1844: 111 [F]. Type species: Blapslaevicollis Gebler, 1841, by subsequent designation (Motschulsky 1860c: 530). Status: valid genus in Blaptinae: Blaptini: Blaptina.

Dilablaps Bogatchev, 1976: 98 [F]. Type species: Dilablapsparadoxa Bogatchev, 1976, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Dilamus Jacquelin du Val, 1861: 279 [M]. Type species: Borosrufipes P.H. Lucas, 1846, by original designation. Status: valid genus and subgenus in Blaptinae: Opatrini: Ammobiina.

Dillacerus Solier, 1835: 497 [M]. Type species: Dillacerusduponti Solier, 1835 (= Dolichoderusacuminatus Klug, 1833), by monotypy. Status: senior synonym of Macellocerus Solier, 1848 in Tenebrioninae: Toxicini: Nycteropina. Note: discovery of this forgotten name threatens the stability of the junior synonym Macellocerus Solier, 1848; although Dillacerus Solier, 1835 has not been used as valid in the literature after 1899, we could not find usage of Macellocerus Solier, 1848 in at least 25 works, published by at least ten authors in the immediately preceding 50 years, and therefore reversal of precedence cannot be used to treat Dillacerus Solier, 1835 as a nomen oblitum; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to conserve usage of Macellocerus Solier, 1848, a genus that includes approximately 50 valid species.

Dilopersina Reitter, 1909a: 117 [F]. Type species: Prosodesjakowlewi Semenov, 1894, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Dimeriseis Solier, 1834: 530 [F]. Type species: Erodiuslaevigatus G.-A. Olivier, 1792, by subsequent designation (Löbl et al. 2008a: 41). Status: valid subgenus of Erodius Fabricius, 1775 in Pimeliinae: Erodiini.

Dimoniacis Koch, 1958: 44 [F]. Type species: Dimoniacisjacksoni Koch, 1958, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Dimorphochilus Borchmann, 1908: 352 [M]. Type species: Dimorphochilusapicalis Borchmann, 1908, by subsequent designation (R. Lucas 1920: 240). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Dinax Gistel, 1848a: xi [M]. Type species [automatic]: Erotylusfasciatus Fabricius, 1781, by subsequent designation (Hope 1841: 133). Status: junior synonym of Poecilesthus Dejean, 1834 in Stenochiinae: Stenochiini. Note: unnecessary replacement name for Poecilesthus Dejean, 1834.

Dineria Motschulsky, 1860c: 530 [F]. Type species: Blapsconfusa Ménétriés, 1832 (= Blaps halophila Fischer, 1820), by subsequent designation (G.S. Medvedev and Iwan 2006: 617). Status: valid subgenus of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina.

Dinomus Brême, 1842a: 113 [M]. Type species: Dinomusperforatus Brême, 1842, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Dinoria Pascoe, 1869: 141 [F]. Type species: Dinoriapicta Pascoe, 1869, by monotypy. Status: junior synonym of Brycopia Pascoe, 1869 in Lagriinae: Adeliini. Synonymy: Carter (1920a: 238).

Dinoscelis Gerstaecker, 1854: 533 [F]. Type species: Dinoscelispasserinii Gerstaecker, 1854, by monotypy. Status: valid subgenus of Pycnocerus Westwood, 1841 in Lagriinae: Pycnocerini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Diodontes Solier, 1834: 508, 518 [M]. Type species: Diodontesporcatus Solier, 1834, by subsequent designation (Hope 1841: 114). Status: valid genus in Pimeliinae: Erodiini.

Dioedus J.L. LeConte, 1862: 238 [M]. Type species: Dioeduspunctatus J.L. LeConte, 1862, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Diopethes Pascoe, 1882: 32 [M]. Type species: Diopethesarachnoides Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Diopoenus Champion, 1888: 445 [M]. Type species: Diopoenuscompressicornis Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Diorhychina Borchmann, 1936: 239, 472 [F]. Type species: Diorhychinaperforata Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Dioscoridemus Koch, 1970: 115 [M]. Type species: Dioscoridemusvonhayekae Koch, 1970, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Dioxycula Fairmaire, 1896c: 115 [F]. Type species: Dioxyculaaranea Fairmaire, 1896, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Diphyrrhynchus Fairmaire, 1849: 445 [M]. Type species: Diphyrrhynchuschalceus Fairmaire, 1849, by monotypy. Status: valid genus in Blaptinae: Opatrini: Heterotarsina. Note: Diphyrrhynchus is an incorrect subsequent spelling of the original spelling Diphyrhynchus, first used by Gemminger in Gemminger and Harold (1870: 1955), and in prevailing usage; Diphyrrhynchus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Diplocyrtus Quedenfeldt, 1887: 257 [M]. Type species: Diplocyrtusfloccosus Quedenfeldt, 1887, by monotypy. Status: valid genus in Tenebrioninae: Apocryphini.

Diprosodes Reitter, 1909a: 115 [M]. Type species: Prosodesbiformis Semenov, 1894, by subsequent designation (G.S. Medvedev 1997: 570). Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Dirosis Miller, 1858: 115 [F]. Type species: Dirosisnervosa Miller, 1858 (= Erodiusservillei Solier, 1834), by monotypy. Status: valid subgenus of Erodius Fabricius, 1775 in Pimeliinae: Erodiini.

Dischidus Kolbe, 1886: 297 [M]. Type species: Helopssinuatus Fabricius, 1801 (= Tenebriolaevigatus Fabricius, 1781), by present designation. Status: junior synonym of Taraxides C.O. Waterhouse, 1876 in Stenochiinae: Cnodalonini. Synonymy: Champion (1891: 637). Note: the original combination of the accepted name of the type species, Tenebriolaevigatus Fabricius, 1781, is a junior primary homonym of Tenebriolaevigatus Linnaeus, 1767.

Dischizillus Wasmann, 1902: 244 [M]. Type species [automatic]: Schizillusrogersi Wasmann, 1899, by monotypy. Status: junior synonym of Pseudethas Fairmaire, 1896 in Pimeliinae: Stenosini: Dichillina. Note: replacement name for Schizillus Wasmann, 1899.

Discodemus J.L. LeConte, 1862: 223 [M]. Type species: Zophosisreticulata Say, 1824, by monotypy. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: J.L. LeConte (1866a: 60).

Discogenia J.L. LeConte, 1866b: 117 [F]. Type species: Eleodesscabricula J.L. LeConte, 1858, by subsequent designation (Bousquet et al. 2018: 149). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Discopleurus Lacordaire, 1859a: 105 [M]. Type species [automatic]: Pleurophorusquadricollis Solier, 1851, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina. Note: replacement name for Pleurophorus Solier, 1851.

Discotus Reitter, 1904: 161 [M]. Type species: Opatrumdilectans Faldermann, 1836, by subsequent designation (G.S. Medvedev 1990: 185). Status: valid subgenus of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina.

Disema Mäklin, 1875: 646 [F]. Type species: Disemabimaculata Mäklin, 1875, by subsequent designation (R. Lucas 1920: 244). Status: valid genus in Lagriinae: Lagriini: Statirina.

Disemorpha Pic, 1917f: 16 [F]. Type species: Disemorphaciliata Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Dissonomus Jacquelin du Val, 1861: 280 [M]. Type species [automatic]: Heliopatespicipes Faldermann, 1837, by subsequent designation (Gebien 1938a: 397). Status: valid genus and subgenus in Tenebrioninae: Dissonomini. Note: replacement name for Heterophylus Mulsant & Rey, 1859.

Distretus Haag-Rutenberg, 1871: 42 [M]. Type species: Molurisamplipennis Fåhraeus, 1870, by subsequent designation (Rye 1873: 287). Status: valid genus and subgenus in Pimeliinae: Sepidiini: Molurina.

Ditaphronotus Casey, 1907: 289, 341 [M]. Type species: Emmenastusfoveicollis Champion, 1884, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Diversogria Pic, 1954: 229 [F]. Type species: Heterogrialepersonnei Pic, 1954, by original designation. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Dividiopsa Koch, 1944a: 158 [F]. Type species: Gnophotaangolensis Harold, 1879, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Divieta Reitter, 1914c: 390 [F]. Type species: Vietacostata Allard, 1874, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Vieta Laporte, 1840 in Pimeliinae: Sepidiini: Sepidiina. Synonymy: Lesne (1922: 696).

Doderoella Schuster, 1926: 133 [F]. Type species: Doderoellacyrenaica Schuster, 1926, by monotypy. Status: junior synonym of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Synonymy: Kwieton (1982: 35).

Dolamara Reichardt, 1935: 251 [F]. Type species: Melanimoncupreomicans Reitter, 1915, by original designation. Status: valid genus in Tenebrioninae: Melanimonini.

Dolichasida Reitter, 1917a: 40, 58 [F]. Type species: Asidamoraguezi Schaufuss, 1879, by subsequent designation (Wilke 1922: 258). Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: Gebien (1937a: 733; with Insulasida Escalera, 1922, a junior synonym of Asida Latreille, 1802).

Dolichoderus Klug, 1833: 87 [M]. Type species: Dolichoderusacuminatus Klug, 1833, by monotypy. Status: senior synonym of Macellocerus Solier, 1848 in Tenebrioninae: Toxicini: Nycteropina. Note: junior homonym of Dolichoderus Lund, 1831 [Hymenoptera].

Doliema Pascoe, 1860a: 50 [N]. Type species: Doliemaplatisoides Pascoe, 1860, by monotypy. Status: junior synonym of Adelina Dejean, 1835 in Diaperinae: Diaperini: Adelinina. Synonymy: Fleutiaux and Sallé (1890: 428), Spilman (1973: 40).

Doliodesmus Spilman, 1967: 149 [M]. Type species: Doliodesmuscharlesi Spilman, 1967, by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Doliopines Horn, 1894: 427 [M]. Type species: Doliopinescucujinus Horn, 1894, by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Dolphus Blanchard, 1847: pl. 11 [M]. Type species: Dolphusglobipennis Blanchard, 1847, by monotypy. Status: valid genus in Tenebrioninae: Helopini: incertae sedis.

Donaciolagria Pic, 1914b: 14 [F]. Type species: Donaciolagriaimpressipennis Pic, 1914, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Donisiellus Bremer, 1992: 111, 113 [M]. Type species: Donisiellusdecellei Bremer, 1992, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Doranalia Novák, 2020c: 481 [F]. Type species: Cistelarufipennis Marseul, 1876, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Dordanea Reitter, 1887a: 357 [F]. Type species: Dordaneaelegans Reitter, 1887, by monotypy. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Dorelogena Péringuey, 1904: 280 [F]. Type species: Dorelogenacastanea Péringuey, 1904, by subsequent designation (R. Lucas 1920: 248). Status: valid genus in Stenochiinae: Stenochiini.

Dorota Novák, 2018c: 452 [F]. Type species: Allecularufoposticalis Pic, 1944, by original designation. Status: valid genus in Alleculina: Alleculini: Alleculina.

Dorrigonum Matthews, 1998: 708, 752 [N]. Type species: Licinomaumbilicata Carter, 1924, by original designation. Status: valid genus in Lagriinae: Adeliini.

Doryagus Pascoe, 1887: 12 [M]. Type species: Doryagustalpa Pascoe, 1887, by monotypy. Status: junior synonym of Stizopus Erichson, 1843 in Blaptinae: Opatrini: Stizopodina. Synonymy: Gebien (1920: 120).

Doyenia Matthews & Lawrence, 2005: 532 [F]. Type species: Doyeniacrematogastri Matthews & Lawrence, 2005, by original designation. Status: valid genus in Lagriinae: Belopini.

Doyenus Iwan, 1996: 385, 386 [M]. Type species: Doyenusuncus Iwan, 1996, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Drepanomela Borchmann, 1936: 119 [F]. Type species: Lagriacribratula Schaufuss, 1887, by original designation. Status: valid subgenus of Cerogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina.

Drocleana Bates, 1879a: 291 [F]. Type species: Camariachalcoptera Klug, 1833, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Drosochrus Erichson, 1843: 243 [M]. Type species: Drosochrusbrunnipes Erichson, 1843, by subsequent designation (Gebien 1943: 910). Status: valid genus and subgenus in Blaptinae: Pedinini: Helopinina.

Dryadigmus Bremer, 2007: 26 [M]. Type species: Amarygmuscechovskyi Bremer, 2007, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Duocula Novák, 2019f: 56 [F]. Type species: Oraculaclara Novák, 2019, by original designation. Status: valid subgenus of Oracula Novák, 2019 in Alleculinae: Alleculini: Alleculina.

Durandius Kaszab, 1970c: 115 [M]. Type species: Durandiusardoini Kaszab, 1970, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Rondoniellina.

Durasida Reitter, 1917a: 9, 14 [F]. Type species: Tenebriosilphoides Linnaeus, 1767, by subsequent designation (F. Soldati 2008: 32). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Dymonus Solier, 1843: 7, 121 [M]. Type species: Sepidium vestitum Guérin-Méneville, 1831, by original designation. Status: junior synonym of Vieta Laporte, 1840 in Pimeliinae: Sepidiini: Sepidiina. Synonymy: Lacordaire (1859a: 205).

Dysantes Pascoe, 1869: 31 [M]. Type species: Dicerodereselongatus Redtenbacher, 1868, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina. Note: this name is not a junior homonym of Dysantes Forster, 1869 [Hymenoptera], as previously reported in the literature, since the name in Coleoptera was published 1 January 1869 whereas the name in Hymenoptera was issued in May 1869 (see Bouchard and Bousquet 2020a: 101 for additional comments).

Dysarchus Pascoe, 1866a: 449 [M]. Type species: Dysarchusodewahnii Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Dysceladus C.O. Waterhouse, 1875: 411 [M]. Type species: Dysceladustuberculatus C.O. Waterhouse, 1875 (= Polposipusherculeanus Solier, 1848), by monotypy. Status: junior synonym of Polposipus Solier, 1848 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1922b: 268, 311).

Dysgena Mäklin, 1863b: 558 [F]. Type species: Dysgenalugubris Mäklin, 1863, by subsequent designation (R. Lucas 1920: 252). Status: valid genus in Tenebrioninae: Praeugenini.

Dysodera Borchmann, 1936: 238, 470 [F]. Type species: Dysoderamethneri Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Dysopinus Borchmann, 1936: 236, 379 [M]. Type species: Nemostiraserra Borchmann, 1913, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Dystalica Pascoe, 1869: 142 [F]. Type species: Dystalicahomogenea Pascoe, 1869, by monotypy. Status: junior synonym of Adelium W. Kirby, 1819 in Lagriinae: Adeliini. Synonymy: Matthews (1998: 777).

Dzhungaropterocoma Skopin, 1974b: 150 [F]. Type species: Pterocomasubnuda Reitter, 1897, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Earophanta Semenov, 1903b: 172 [F]. Type species [automatic]: Platyopeserrata Semenov, 1893, by original designation. Status: valid genus in Pimeliinae: Pimeliini. Note: replacement name for Earophila Semenov, 1903.

Earophila Semenov, 1903a: 9 [F]. Type species: Platyopeserrata Semenov, 1893, by original designation. Status: senior synonym of Earophanta Semenov, 1903 in Pimeliinae: Pimeliini. Note: junior homonym of Earophila Gumppenberg, 1887 [Lepidoptera].

Earophilina Strand, 1917: 99 [F]. Type species [automatic]: Platyopeserrata Semenov, 1893, by original designation. Status: junior synonym of Earophanta Semenov, 1903 in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Earophila Semenov, 1903.

Eba Pascoe, 1863b: 122, 129 [F]. Type species: Ebacerylonoides Pascoe, 1863, by monotypy. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Carter and Zeck (1937: 194). Note: transferred from “Tenebrionoidea: Colydiidae” by Carter and Zeck (1937: 194).

Ebenolus Fairmaire, 1897c: 227 [M]. Type species: Ebenolusvernicatus Fairmaire, 1897, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Kaszab (1977b: 29).

Ebertius Jedlička, 1965: 98 [M]. Type species: Ebertiusnepalensis Jedlička, 1965 (= Laenalongipilis Schuster, 1926), by original designation. Status: junior synonym of Laena Dejean, 1821 in Lagriinae: Laenini. Synonymy: Kaszab (1970d: 429, through placement of the type species in Laena Dejean, 1821). Note: originally described in the family Carabidae.

Eccoptostira Borchmann, 1936: 236, 377 [F]. Type species: Nemostirarohdei Pic, 1912, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: we act as First Revisers and reject the alternative original spelling Ecoptostira, used by Borchmann (1936: 236).

Eccoptostoma Gebien, 1913: 70 [N]. Type species: Taraxidesruficrus Fairmaire, 1894, by present designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: redescribed as new by Gebien (1921b: 62, 82).

Echinotrigon Skopin, 1973: 170 [M]. Type species: Trigonoscelisgranulata Reitter, 1915, by original designation. Status: valid subgenus of Trigonoscelis Dejean, 1834 in Pimeliinae: Pimeliini.

Echinotus Solier, 1843: 30 [M]. Type species: Sepidium spinicolle Laporte, 1840, by original designation [p. 122]. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Echocerus Horn, 1870: 364, 366 [M]. Type species: Trogossitamaxillosa Fabricius, 1801, by monotypy. Status: valid subgenus of Gnatocerus Thunberg, 1814 in Diaperinae: Diaperini: Adelinina.

Ecnocera Borchmann, 1936: 22, 220 [F]. Type species: Porrolagriagracilis Borchmann, 1909, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Ecnolagria Borchmann, 1916a: 49, 139 [F]. Type species: Lagriagrandis Gyllenhal, 1817, by original designation. Status: valid genus and subgenus in Lagriinae: Lagriini: Lagriina.

Ecnomoderes Gebien, 1928: 109, 110 [M]. Type species: Ecnomoderesbarbatus Gebien, 1928, by subsequent designation (Gebien 1937a: 690). Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Ecnomosternum Gebien, 1928: 104 [N]. Type species: Ecnomosternumvermiculatum Gebien, 1928, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Ecphoroma Solier, 1836: 195 [N]. Type species: Pimeliahemisphaerica Solier, 1836, by subsequent designation (Löbl et al. 2008b: 157). Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Ecripsis Pascoe, 1866a: 456 [F]. Type species: Ecripsispubescens Pascoe, 1866, by monotypy. Status: junior synonym of Ammidium Erichson, 1843 in Blaptinae: Opatrini: Ammobiina. Synonymy: Blair in Carter (1914c: 406). Note: this African genus was originally described from Australia in error (see Matthews and Bouchard 2008: 351).

Ectateus Koch, 1956a: 230 [M]. Type species: Anchophthalmusmodestus Fairmaire, 1887, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Ectatocera Fåhraeus, 1870: 325 [F]. Type species: Ectatoceralongicornis Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Ectatocnemis Horn, 1867: 400 [F]. Type species: Ectatocnemismultilineata Horn, 1867, by monotypy. Status: junior synonym of Anomalipus Guérin-Méneville, 1831 in Blaptinae: Platynotini: Platynotina. Synonymy: Endrödy-Younga (1988: 19). Note: this taxon was originally described from Chile in error, it is native to South Africa (see Endrödy-Younga 1988: 19).

Ectenostoma Fåhraeus, 1870: 317 [N]. Type species: Ectenostomanigriventris Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Ectomopsis Fairmaire, 1905: 299 [F]. Type species: Ectomopsisbruchi Fairmaire, 1905, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ectromopsis Antoine, 1949: 125, 145 [F]. Type species: Catomuspoliticollis Allard, 1876, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Ectyche Pascoe, 1869: 143 [F]. Type species: Ectycheerebea Pascoe, 1869, by monotypy. Status: valid genus in Diaperinae: Ectychini. Note: “Ectyche? nana Pascoe, 1869” was doubtfully included in this genus by Pascoe (1869: 143) and is therefore not an originally included species (ICZN 1999: Article 67.2.5).

Edalus Broun, 1893b: 1159 [M]. Type species: Edalusopacus Broun, 1893, by subsequent designation (R. Lucas 1920: 255). Status: senior synonym of Wattadelium Emberson, 2000 in Lagriinae: Adeliini. Note: junior homonym of Edalus Broun, 1886 [Coleoptera: Zopheridae].

Edrotes J.L. LeConte, 1851: 140 [M]. Type species: Edrotesventricosus J.L. LeConte, 1851, by monotypy. Status: valid genus and subgenus in Pimeliinae: Edrotini.

Edrotinus Fairmaire, 1904b: 461 [M]. Type species: Edrotinustucumanus Fairmaire, 1904, by original designation. Status: junior synonym of Trilobocara Solier, 1851 in Pimeliinae: Trilobocarini. Synonymy: Gebien (1937a: 592).

Edrotopus Haag-Rutenberg, 1877: 129 [M]. Type species: Edrotopusstrigicollis Haag-Rutenberg, 1877, by monotypy. Status: junior synonym of Platyholmus Dejean, 1834 genus in Pimeliinae: Praociini. Synonymy: Fairmaire (1884a: 508).

Edylius Champion, 1894a: 353 [M]. Type species: Edyliuscanescens Champion, 1894, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Eichleria Kamiński, 2015b: 132 [F]. Type species: Eichleriaostrowskii Kamiński, 2015, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Elaeodes Gemminger in Gemminger and Harold, 1870: 1868 [F]. Type species [automatic]: Eleodesdentipes Eschscholtz, 1829, by subsequent designation (Hope 1841: 124). Status: junior synonym of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini. Note: unjustified emendation of Eleodes Eschscholtz, 1829, not in prevailing usage.

Elaeus Gemminger in Gemminger and Harold, 1870: 1969 [M] Type species [automatic]: Heleaperforata Latreille, 1816, by subsequent monotypy (Latreille 1817: 261). Status: junior synonym of Helea Latreille, 1804 in Tenebrioninae: Heleini: Heleina. Note: unjustified emendation of Helea Latreille, 1804 (as “Helaeus Latreille”), not in prevailing usage.

Elasmocera Mäklin, 1867: 504 [F]. Type species: Elasmoceradentipes Mäklin, 1867, by monotypy. Status: senior synonym of Elasmocerella Strand, 1935 in Stenochiinae: Stenochiini. Note: junior homonym of Elasmocera Rondani, 1846 [Diptera].

Elasmocerella Strand, 1935b: 302 [F]. Type species [automatic]: Elasmoceradentipes Mäklin, 1867, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: replacement name for Elasmocera Mäklin, 1867.

Eleates Casey, 1886: 253 [M]. Type species: Eleatesoccidentalis Casey, 1886, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Eledona Latreille, 1797: 19 [F]. Type species: Opatrumagricola Herbst, 1783, by subsequent monotypy (Latreille 1802: 162). Status: valid genus in Tenebrioninae: Bolitophagini. Note: originally proposed without included nominal species; Latreille (1802: 162)) , by including the species “Bolitophagusagricola” in association with this name, was the first author to subsequently and expressly include nominal species in Eledona (ICZN 1999, Article 67.2.2).

Eledonoprius Reitter, 1911: 329, 338 [M]. Type species: Opatrumarmatum Panzer, 1799, by subsequent designation (R. Lucas 1920: 258). Status: valid genus in Tenebrioninae: Bolitophagini.

Elenophorus Dejean, 1821: 64 [M]. Type species: Tenebriocollaris Linnaeus, 1767, by monotypy. Status: junior synonym of Leptoderis Billberg, 1820 in Pimeliinae: Elenophorini: Elenophorina. Synonymy: Silfverberg (1984: 59).

Eleodes Eschscholtz, 1829: 8 [F]. Type species: Eleodesdentipes Eschscholtz, 1829, by subsequent designation (Hope 1841: 124). Status: valid genus and subgenus in Blaptinae: Amphidorini.

Eleodimorpha Blaisdell, 1909: 477 [F]. Type species: Eleodimorphabolcan Blaisdell, 1909, by original designation. Status: valid genus in Blaptinae: Amphidorini.

Eleodopsis Blaisdell, 1939: 52 [F]. Type species: Eleodopsissubvestita Blaisdell, 1939, by original designation. Status: junior synonym of Blapylis Horn, 1870 in Blaptinae: Amphidorini. Synonymy: Spilman (1962a: 59).

Eleoselinus Kamiński, 2014: 101 [M]. Type species: Ectateusvilliersi Ardoin, 1965, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Elixota Pascoe, 1866a: 475 [F]. Type species: Elixotacuprea Pascoe, 1866 (= Amarygmushopei Bremer, 2001), by original designation. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer (2001a: 65).

Ellaemus Pascoe, 1866c: 495 [M]. Type species: Emcephalussubmaculatus Brême, 1842, by present designation. Status: junior synonym of Emcephalus W. Kirby, 1828 in Tenebrioninae: Heleini: Heleina. Synonymy: Matthews and Bouchard (2008: 304).

Ellidoneus Wilke, 1922: 277 [M]. Type species: Cardigeniusgranulatus Fairmaire, 1873, by original designation. Status: valid subgenus of Cardigenius Solier, 1836 in Pimeliinae: Asidini.

Ellipsodes Wollaston, 1854: 485 [M]. Type species: Sphaeridiumglabratum Fabricius, 1781, by monotypy. Status: valid genus and subgenus in Diaperinae: Crypticini.

Elomosda Bates, 1870: 273 [F]. Type species: Elomosdabeltii Bates, 1870, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Elongasida Escalera, 1906: 306 [F]. Type species: Asidagrandipalpis Allard, 1869, by subsequent designation (Viñolas and Cartagena 2005: 164). Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini.

Embaphion Say, 1824: 254 [N]. Type species: Akismuricata Say, 1824, by monotypy. Status: valid genus in Blaptinae: Amphidorini.

Emcephalus W. Kirby, 1828: 524 [M]. Type species: Emcephalusgibbosus W. Kirby, 1828, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Emeacoides Gebien, 1937a: 698 [M]. Type species [automatic]: Enneacoidesvinculiger Fairmaire, 1881 (= Nyctoporiscarinata J.L. LeConte, 1851), by monotypy. Status: junior synonym of Nyctoporis Eschscholtz, 1831 in Pimeliinae: Nyctoporini. Note: unjustified emendation of Enneacoides Fairmaire, 1881, not in prevailing usage.

Emeax Pascoe, 1866a: 450 [M]. Type species: Emeaxsculpturatus Pascoe, 1866 (= Nyctoporiscristata Eschscholtz, 1831), by monotypy. Status: junior synonym of Nyctoporis Eschscholtz, 1831 in Pimeliinae: Nyctoporini. Synonymy: J.L. LeConte (1873: 334). Note: as mentioned by Matthews and Bouchard (2008: 351) the type species was originally described from Australia in error; the genus Nyctoporis Eschscholtz, 1831 is endemic to the Nearctic realm.

Emmallodera Blanchard, 1842: pl. 13 [F]. Type species: Emmalloderacrenaticostata Blanchard, 1842, by monotypy. Status: valid genus in Tenebrioninae: Scotobiini. Note: unjustified emendation of the original spelling Emalodera, introduced by Agassiz (1846b: 137), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Emmallus Agassiz, 1846b: 137 [M]. Type species [automatic]: Emmaluspilosus Erichson, 1843, by monotypy. Status: junior synonym of Emmalus Erichson, 1843 in Blaptinae: Opatrini: Ammobiina. Note: unjustified emendation of Emmalus Erichson, 1843, not in prevailing usage.

Emmalus Erichson, 1843: 251 [M]. Type species: Emmaluspilosus Erichson, 1843, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Emmenastrichus Horn, 1894: 413 [M]. Type species: Emmenastrichuscribratus Horn, 1894, by subsequent designation (Casey 1907: 289). Status: valid genus in Pimeliinae: Edrotini.

Emmenastus Motschulsky, 1845a: 75 [M]. Type species: Emmenastusrugosus Motschulsky, 1845, by subsequent designation (J.L. LeConte 1866b: 106). Status: junior synonym of Oxycara Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Aalbu et al. (1995: 484).

Emmenides Casey, 1907: 329 [M]. Type species: Emmenastuspunctatus J.L. LeConte, 1866, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Emydodes Pascoe, 1860a: 56 [M]. Type species: Emydodescollaris Pascoe, 1860, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Emyon Gerstaecker, 1854: 532 [M]. Type species: Emyoncaelatus Gerstaecker, 1854, by monotypy. Status: junior synonym of Helopinus Solier, 1848 in Blaptinae: Pedinini: Helopinina. Synonymy: Koch (1958: 149).

Emypsara Pascoe, 1866a: 460 [F]. Type species: Emypsaraadamsii Pascoe, 1866 (= Diaperisriederii Faldermann, 1833), by subsequent designation (Löbl et al. 2008a: 41). Status: valid genus in Diaperinae: Phaleriini.

Enanea Lewis, 1894: 467 [F]. Type species: Enaneatestacea Lewis, 1894, by monotypy. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Encara Gemminger, 1870: 124 [N]. Type species [automatic]: Emcephalusgibbosus W. Kirby, 1828, by monotypy. Status: junior synonym of Emcephalus W. Kirby, 1828 in Tenebrioninae: Heleini: Heleina. Note: replacement name for Emcephalus W. Kirby, 1828 (as “Encephalus Brême”).

Encephalus Agassiz, 1846b: 137 [M]. Type species [automatic]: Emcephalusgibbosus W. Kirby, 1828, by monotypy. Status: junior synonym of Emcephalus W. Kirby, 1828 in Tenebrioninae: Heleini: Heleina. Note: unjustified emendation of Emcephalus W. Kirby, 1828, not in prevailing usage; junior homonym of Encephalus Stephens, 1832 [Coleoptera: Staphylinidae].

Encyalesthus Motschulsky, 1860d: 139 [M]. Type species: Encyalesthussubviolaceus Motschulsky, 1860, by monotypy. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1987: 43).

Endostomus Gemminger in Gemminger and Harold, 1870: 1973 [M]. Type species [automatic]: Cossyphussenegalensis Laporte, 1833, by subsequent monotypy (Duponchel 1844b: 315). Status: junior synonym of Endustomus Brême, 1842 in Lagriinae: Cossyphini. Note: unjustified emendation of Endustomus Brême, 1842, not in prevailing usage.

Endostostomus Jakobson, 1914: 528 [M]. Type species [automatic]: Cossyphussenegalensis Laporte, 1833, by subsequent monotypy (Duponchel 1844b: 315). Status: junior synonym of Endustomus Brême, 1842 in Lagriinae: Cossyphini. Note: unjustified emendation of Endustomus Brême, 1842, not in prevailing usage.

Endothina Carter, 1924b: 536 [F]. Type species: Endothinasquamosa Carter, 1924 (= Opatrumcanaliculatum Fabricius, 1798), by monotypy. Status: junior synonym of Leichenum Dejean, 1834 in Blaptinae: Pedinini: Leichenina. Synonymy: Carter (1928: 284).

Endroeditagalus Schawaller & Bouchard, 2019: 192 [M]. Type species: Endroeditagalusntsubanus Schawaller & Bouchard, 2019, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Endustomus Brême, 1842b: 17 [M]. Type species: Cossyphussenegalensis Laporte, 1833, by subsequent monotypy (Duponchel 1844b: 315). Status: valid genus in Lagriinae: Cossyphini. Note: originally proposed without included nominal species; Duponchel (1844b: 315), by including the species “Cossyphussenegalensis” in association with this name, was the first author to subsequently and expressly include nominal species in Endustomus (ICZN 1999, Article 67.2.2).

Enganodia Fairmaire, 1898d: 398 [F]. Type species: Enganodiasanguinicrus Fairmaire, 1898, by monotypy. Status: junior synonym of Lophocnemis Mäklin, 1867 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 543).

Enicmonota Ardoin, 1959a: 61 [F]. Type species: Enicmosomacrassicorne Ardoin, 1959, by monotypy. Status: junior synonym of Enicmosoma Gebien, 1922 in Lagriinae: Lupropini. Synonymy: Schawaller (2013a: 139).

Enicmosoma Gebien, 1922b: 312 [N]. Type species: Enicmosomapunctum Gebien, 1922, by subsequent designation (Gebien 1941: 823). Status: valid genus in Lagriinae: Lupropini.

Enigmatica Ferrer, 2005: 202 [F]. Type species: Enigmaticaendroedyi Ferrer, 2005, by monotypy. Status: valid genus in Lagriinae: Laenini.

Enneacoides Fairmaire, 1881c: 277 [M]. Type species: Enneacoidesvinculiger Fairmaire, 1881 (= Nyctoporiscarinata J.L. LeConte, 1851), by monotypy. Status: junior synonym of Nyctoporis Eschscholtz, 1831 in Pimeliinae: Nyctoporini. Synonymy: Gebien (1908b: 287).

Ennychiatus Koch, 1963: 28 [M]. Type species: Stizopuscaraboides Fairmaire, 1897, by original designation. Status: junior synonym of Parastizopus Gebien, 1938 in Blaptinae: Opatrini: Stizopodina. Synonymy: Iwan and Schimrosczyk (2017: 384).

Ennychius Fåhraeus, 1870: 299 [M]. Type species: Ennychiusmorio Fåhraeus, 1870, by monotypy. Status: junior synonym of Helibatus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Stizopodina. Synonymy: Gebien (1938a: 74).

Enoplopus Solier, 1848: 151, 158 [M]. Type species [automatic]: fixed herein (ICZN 1999, Article 70.3) as Tenebriovelikensis Piller & Mitterpacher, 1783, misidentified as Tenebriocaraboides Linnaeus, 1758 in the original designation by monotypy in Dejean (1821). Status: junior synonym of Accanthopus Dejean, 1821 in Tenebrioninae: Helopini: Enoplopodina. Note: unnecessary replacement name for Accanthopus Dejean, 1821.

Entinopoda Gebien, 1938b: 64 [F]. Type species: Eustolopusoctoseriatus Gebien, 1938, by monotypy. Status: junior synonym of Eustolopus Gebien, 1938 in Pimeliinae: Adesmiini. Synonymy: Penrith (1979: 40).

Entomobalia Flores & Triplehorn, 2002: 607 [F]. Type species: Asidaplatynotos Perty, 1830, by original designation. Status: valid genus in Pimeliinae: Nycteliini.

Entomochilus Gay & Solier in Solier, 1843: 48 [M]. Type species: Entomochiluspilosus Gay & Solier, 1843, by original designation. Status: valid genus in Pimeliinae: Physogasterini.

Entomoderes Solier, 1836: 308, 346 [M]. Type species: Entomodereserebi Solier, 1836, by monotypy. Status: valid genus in Pimeliinae: Nycteliini.

Entomogonus Solier, 1848: 151, 155 [M]. Type species: Entomogonusbarthelemyi Solier, 1848, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina.

Entypodera Gerstaecker, 1871: 66 [F]. Type species: Entypoderaanthicoides Gerstaecker, 1871, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: new placement [OM], previously included in Lagriinae: Lagriini: Lagriina.

†Eoallognosis Haupt, 1950: 115, 137 [F]. Type species: Eoallognosisundulata Haupt, 1950, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

†Eocallidium Haupt, 1950: 143 [N]. Type species: Eocallidiumrugulosum Haupt, 1950, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: originally included in the family Cerambycidae, transferred to Tenebrionidae by Vitali (2008: 8); described from Middle Eocene deposits (Germany).

Eocyphogenia G.S. Medvedev, 1968b: 897, 898 [F]. Type species: Akisrugipennis Faldermann, 1835, by original designation. Status: junior synonym of Cyphogenia Solier, 1837 in Pimeliinae: Akidini. Synonymy: Ren and Yu (2000: 54).

Eodirosis Kwieton, 1980: 25 [F]. Type species: Erodiusquadrilineatus Kraatz, 1865, by original designation. Status: valid subgenus of Erodius Fabricius, 1775 in Pimeliinae: Erodiini. Note: the original combination of the accepted name of the type species, Erodiusquadrilineatus Kraatz, 1865, is a junior primary homonym of Erodiusquadrilineatus G.-A. Olivier, 1792.

†Eodromus Haupt, 1950: 113, 120 [M] Type species: †Ancylochiraagilis Meunier, 1915, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: originally proposed in the family Carabidae by Pongrácz (1935: 538) without type species (ICZN 1999, Article 13.3); we act as First Revisers and reject the alternative original spelling Edromus, used by Haupt (1950: 120); described from Middle Eocene deposits (Germany).

†Eohelaeus Haupt, 1950: 115, 135 [M]. Type species: Eohelaeussublaevis Haupt, 1950, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Epairops Fåhraeus, 1870: 282 [M]. Type species: Epairopsfragilis Fåhraeus, 1870, by monotypy. Status: junior synonym of Ossiporis Pascoe, 1866 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Gebien (1937b: 37).

Epairopsis Koch, 1955a: 47 [F]. Type species: Trachynotusfrontalis Haag-Rutenberg, 1873, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Epantius J.L. LeConte, 1851: 144 [M]. Type species: Epantiusobscurus J.L. LeConte, 1851, by monotypy. Status: valid genus in Tenebrioninae: Eulabini.

Epeurycaulus Kolbe, 1902a: 579 [M]. Type species: Epeurycaulusaldabricus Kolbe, 1902, by present designation. Status: junior synonym of Plesioderes Mulsant, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1922b: 268).

Ephalus J.L. LeConte, 1862: 228 [M]. Type species: Heliopateslatimanus J.L. LeConte, 1847, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: transferred from Ammobiina by Lumen et al. (2020: 344).

Ephidonius Pascoe, 1869: 151 [M]. Type species: Ephidoniusacuticornis Pascoe, 1869, by original designation. Status: junior synonym of Brises Pascoe, 1869 in Tenebrioninae: Heleini: Heleina. Synonymy: Carter (1914b: 46).

Epicalla Lacordaire, 1859a: 309 [F]. Type species: Epicallavaripes Champion, 1886, by subsequent designation (R. Lucas 1920: 268). Status: valid genus in Stenochiinae: Cnodalonini. Note: the authorship of this genus name has been credited to Champion (1886: 249) in the literature; however, Lacordaire (1859a: 309) mentioned that the genus Epicalla has the “mandibules entières au bout”, which fulfils the requirements of availability (ICZN 1999, Article 12.1); Champion (1886: 249–251), by describing three new species in association with the genus Epicalla, was the first author to subsequently and expressly include nominal species in this genus (ICZN 1999, Article 67.2.2).

Epicydes Champion, 1889: 60 [M]. Type species: Epicydesoculatus Champion, 1889, by subsequent designation (Borchmann 1936: 429). Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Epilamprus Gistel, 1848a: xiv [M]. Type species [automatic]: Helopsindutus Wiedemann, 1819, by subsequent designation (Duponchel 1844b: 359). Status: junior synonym of Ceropria Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Note: unjustified emendation of Epilampus Dejean, 1834, not in prevailing usage.

Epilampus Dejean, 1834: 198 [M]. Type species [automatic]: Helopsindutus Wiedemann, 1819, by subsequent designation (Duponchel 1844b: 359). Status: junior synonym of Ceropria Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Note: unnecessary replacement name for Ceropria Laporte & Brullé, 1831 (see Bousquet and Bouchard 2013a: 52).

Epilasium Erichson, 1842a: 237 [N]. Type species: Trichotoncayennense Hope, 1841, by monotypy. Status: junior synonym of Trichoton Hope, 1841 in Blaptinae: Opatrini: Blapstinina. Synonymy: Erichson (1842a: 237). Note: the name Epilasium was listed as synonym of Trichoton Hope, 1841 by Erichson (1842a: 237), it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Curtis 1844: 222), Epilasium was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Epipagus Haag-Rutenberg, 1872: 274, 311 [M]. Type species: Epipagusbenguelensis Haag-Rutenberg, 1872, by monotypy. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Epipedodema Gebien, 1921b: 54 [F]. Type species: Epipedodemadepressa Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Alphitobiini.

Epipedonota Solier, 1836: 307, 342 [F]. Type species: Epipedonotaebenina Solier, 1836, by subsequent designation (Solier 1851: 159). Status: valid genus in Pimeliinae: Nycteliini. Note: the alternative original spelling Epipedonata, used by Solier (1836: 342), was rejected by Solier (1838a: 487) who acted as the First Reviser.

Epiphaleria Lewis, 1894: 382 [F]. Type species: Epiphaleriaatriceps Lewis, 1894, by original designation. Status: valid subgenus of Phaleria Latreille, 1802 in Diaperinae: Phaleriini.

Epiphysa Dejean, 1834: 178 [F]. Type species: Pimeliaflavicollis Fabricius, 1794, by monotypy. Status: valid genus in Pimeliinae: Adesmiini.

Epiplecta Mäklin, 1867: 498 [F]. Type species: Epiplectamaculata Mäklin, 1867, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Episopus Bates, 1873e: 372 [M]. Type species: Episopuspolitus Bates, 1873, by subsequent designation (R. Lucas 1920: 270). Status: valid genus in Stenochiinae: Cnodalonini.

Epitoxicum Bates, 1873d: 46 [N]. Type species: Epitoxicumhaplandroides Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Toxicina.

Epitragella Kulzer, 1958b: 184 [F]. Type species: Epitragelladimorpha Kulzer, 1958, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Epitragodes Casey, 1890b: 365 [M]. Type species: Epitragustomentosus J.L. LeConte, 1866, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Epitragoma Casey, 1907: 386 [N]. Type species: Epitragusvestitus Casey, 1891, by monotypy. Status: junior synonym of Pechalius Casey, 1907 in Pimeliinae: Epitragini. Synonymy: Freude (1968: 61).

Epitragopsis Casey, 1907: 386 [F]. Type species: Epitragusgodmani Champion, 1884, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Epitragosoma K.W. Brown & Triplehorn, 2002: 515 [N]. Type species: Epitragosomaarenaria K.W. Brown & Triplehorn, 2002, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Epitragus Latreille, 1802: 165 [M]. Type species: Epitragusfuscus Latreille, 1804, by subsequent monotypy (Latreille 1804: 322). Status: valid genus and subgenus in Pimeliinae: Epitragini. Note: originally proposed without included nominal species since the species listed (Helopsvariegatus ? Fab.) is conditionally included; Latreille (1804: 322), by including the species Epitragusfuscus Latreille, 1804 in association with this name, was the first author to subsequently and expressly include a nominal species in Epitragus (ICZN 1999, Article 67.2.2).

Epitrichia Gebler, 1859: 475 [F]. Type species: Helopstomentosus Gebler, 1842, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Epityria Koch, 1950b: 298 [F]. Type species: Himatismusstriatopunctatus Haag-Rutenberg, 1877, by original designation. Status: valid subgenus of Derosphaerius Westwood, 1881 in Pimeliinae: Tentyriini.

Epomidus Matthews, 1998: 705, 763 [M]. Type species: Epomidusprionodes Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Erelus Mulsant & Rey, 1853a: 185 [M]. Type species: Erelussulcipennis Mulsant & Rey, 1853, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: included in a list of “Tenebrionidaenomina dubia” by Iwan et al. (2020: 475).

Eremobates Gebien, 1921b: 120 [M]. Type species: Eremobatescrux Gebien, 1921, by monotypy. Status: senior synonym of Eremobatodes Gebien, 1943 in Stenochiinae: Cnodalonini. Note: junior homonym of Eremobates Banks, 1900 [Arachnida].

Eremobatodes Gebien, 1943: 404 [M]. Type species [automatic]: Eremobatescrux Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Eremobates Gebien, 1921.

Eremocantor Smith & Wirth, 2016: 582 [M]. Type species: Eremocantormarioni Smith & Wirth, 2016, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Eremoecus Lacordaire, 1859a: 69 [M]. Type species: Hyperopseschscholtzii Solier, 1851, by monotypy. Status: valid genus in Pimeliinae: Trilobocarini.

Eremonomus Wollaston, 1861: 199 [M]. Type species: Eremonomushuttoni Wollaston, 1861 (= Ammidiumciliatum Erichson, 1843), by monotypy. Status: junior synonym of Ammidium Erichson, 1843 in Blaptinae: Opatrini: Ammobiina. Synonymy: Ardoin (1971: 358).

Eremophaleria Español, 1951: 32, 34 [F]. Type species: Phaleriabedeli Chobaut, 1900, by original designation. Status: valid subgenus of Phaleria Latreille, 1802 in Diaperinae: Phaleriini.

Eremostibes Koch, 1963: 60 [M]. Type species: Eremostibesopacus Koch, 1963, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Ergenna Fairmaire, 1897f: 139 [F]. Type species: Ergennacaerulescens Fairmaire, 1897, by monotypy. Status: junior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Synonymy: De Moor (1970: 42).

Erionura Reitter, 1903: 18 [F]. Type species: Helopsgiganteus Kraatz, 1862, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Ernocharis C.G. Thomson, 1859: 118 [F]. Type species: Cistelabrevis Illiger, 1794 (= Cistelamaura Fabricius, 1792), by original designation. Status: valid subgenus of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina.

Erodibius Löbl, Bouchard, Merkl & Bousquet, 2020: 2 [M]. Type species: Arthrodeiscyphonotus Fairmaire, 1887, by original designation. Status: valid subgenus of Arthrodibius Lesne, 1915 in Pimeliinae: Erodiini. Note: name first proposed by Koch (1960: 354) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl et al. (2008a: 40) designated Arthrodeiscyphonotus Fairmaire, 1887 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Erodinus Reitter, 1900b: 299 [M]. Type species: none designated. Status: undetermined taxon in Pimeliinae: Erodiini. Note: this genus was included in a key, which fulfils the criteria of availability for new genus-group names proposed before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Erodinus and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Erodiontes Reitter, 1914a: 48, 79 [M]. Type species: Erodiontesvermiculatus Reitter, 1914, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Erodius Fabricius, 1775: 258 [M]. Type species: Erodiusgibbus Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Pimeliinae: Erodiini.

Erulipothydemus Pic, 1918b: 19 [M]. Type species: Erulipothydemuslatior Pic, 1918, by subsequent designation (Gebien 1943: 902). Status: valid genus in Tenebrioninae: Helopini: incertae sedis.

Erulipus Fairmaire, 1903a: 14 [M]. Type species: Erulipusfruhstorferi Fairmaire, 1903, by monotypy. Status: valid subgenus of Ainu Lewis, 1894 in Stenochiinae: Cnodalonini.

Erxias Champion, 1888: 460 [M]. Type species: Erxiasbicolor Champion, 1888, by subsequent designation (Bousquet et al. 2015: 138). Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Erycastus Fairmaire, 1897f: 133 [M]. Type species: Erycastusnavicularis Fairmaire, 1897, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Eryx Stephens, 1832b: 27 [M]. Type species: Pyrochroanigra DeGeer, 1775 (= Helopsater Fabricius, 1775), by monotypy. Status: senior synonym of Prionychus Solier, 1835 in Alleculinae: Alleculini: Alleculina. Synonymy: Mulsant (1856a: 12, 61). Note: junior homonym of Eryx Daudin, 1803 [Reptilia].

Erzika Novák, 2020d: 53 [F]. Type species: Erzikatamdaoica Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Eschatomoxys Blaisdell, 1935: 125 [M]. Type species: Eschatomoxyswagneri Blaisdell, 1935, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Eschatoporis Blaisdell, 1906: 76 [F]. Type species: Eschatoporisnunenmacheri Blaisdell, 1906, by monotypy. Status: valid genus in Lagriinae: Eschatoporiini.

Eschatostena Keleinikova, 1977: 654 [F]. Type species: Eschatostenakuznetzovi Keleinikova, 1977, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Esemephe Steiner, 1980: 385 [F]. Type species: Esemephetumi Steiner, 1980, by original designation. Status: valid genus in Pimeliinae: Cossyphodini: Esemephina.

Espagnolina Kaszab, 1965: 117 [F]. Type species: Espagnolinaassamica Kaszab, 1965, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Espidium Rafinesque, 1815: 113 [N]. Type species [automatic]: Sepidium tricuspidatum Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Sepidium Fabricius, 1775 in Pimeliinae: Sepidiini: Sepidiina. Note: unnecessary replacement name for Sepidium Fabricius, 1775.

Espites Pascoe, 1882: 32 [M]. Type species: Espitesbasalis Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Espitomorphus Pic, 1921d: 24 [M]. Type species: Espitomorphusmulticolor Pic, 1921, by monotypy. Status: junior synonym of Camarimena Motschulsky, 1863 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 136).

Etazeta Fairmaire, 1889a: 358 [F]. Type species: Etazetaaeneicolor Fairmaire, 1889, by monotypy. Status: junior synonym of Luprops Hope, 1833 in Lagriinae: Lupropini. Synonymy: Gebien (1914a: 35).

Ethas Pascoe, 1862: 324 [M]. Type species: Ethascarbonarius Pascoe, 1862, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Ethmomerus Koch, 1953a: 243 [M]. Type species: Ethmussubcylindricus Koch, 1953, by original designation. Status: valid subgenus of Ethmus Haag-Rutenberg, 1873 in Pimeliinae: Sepidiini: Trachynotina.

Ethmophobes Koch, 1953a: 244 [M]. Type species: Ethmuslatus Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Ethmus Haag-Rutenberg, 1873 in Pimeliinae: Sepidiini: Trachynotina.

Ethmus Haag-Rutenberg, 1873: 44 [M]. Type species: Ethmusmaculatus Haag-Rutenberg, 1873, by subsequent designation (Gebien 1937a: 778). Status: valid genus and subgenus in Pimeliinae: Sepidiini: Trachynotina. Note: nomenclatural stability is threatened by the discovery of an older type species designation (Ethmuslatus Haag-Rutenberg, 1873, by subsequent designation by R. Lucas (1920: 276), which is the type species of the valid subgenusEthmophobes Koch, 1953); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Gebien (1937a: 778).

Eubalia Laporte, 1840: 257 [F]. Type species: Statiraovalis Laporte, 1840, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Euboeus Boieldieu, 1865: 10 [M]. Type species: Euboeusmimonti Boieldieu, 1865, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Eucaliga Fairmaire & Germain, 1861: 5 [F]. Type species: Eucaligasanguinicollis Fairmaire & Germain, 1861, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Eucamaria Gebien, 1919: 28, 149 [F]. Type species: Camariaspectabilis Pascoe, 1860, by original designation. Status: junior synonym of Falsocamaria Pic, 1917 in Stenochiinae: Cnodalonini. Synonymy: Masumoto (1993a: 142).

Eucamptus Germar, 1842: 444 [M]. Type species: Eucamptusiridis Germar, 1842 (= Hegemonaresplendens Laporte, 1840), by monotypy. Status: junior synonym of Hegemona Laporte, 1840 in Stenochiinae: Cnodalonini. Synonymy: Duponchel (1845b: 498).

Euclarkia Lea, 1919: 180 [F]. Type species: Euclarkiacostata Lea, 1919, by monotypy. Status: valid genus in Lagriinae: Belopini.

Eucolus Mulsant & Rey, 1853b: 39, 67 [M]. Type species: Eucoluspolinierii Mulsant & Rey, 1853, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Euconibius Casey, 1895: 618 [M]. Type species: Notibiusgagates Horn, 1870, by monotypy. Status: junior synonym of Conibius J.L. LeConte, 1851 in Blaptinae: Opatrini: Blapstinina. Synonymy: Aalbu in Bousquet et al. (2018: 202).

Eucosmus Gistel, 1848a: x [M]. Type species [automatic]: Spheniscuserotyloides W. Kirby, 1819, by monotypy. Status: junior synonym of Cuphotes Champion, 1887 in Stenochiinae: Stenochiini. Note: replacement name for Spheniscus W. Kirby, 1819; junior homonym of Eucosmus Agassiz, 1846 [Echinoidea].

Eucrossoscelis Nakane, 1963: 29 [F]. Type species: Eucrossoscelisbroscosomoides Nakane, 1963, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Eucyrtus Lacordaire, 1859b: 417 [M]. Type species: Eucyrtuspretiosus Lacordaire, 1859, by subsequent designation (Gebien 1921a: 335). Status: valid genus in Stenochiinae: Cnodalonini. Note: the name Eucyrtus was listed as synonym of Scotaeus Hope, 1834 by Lacordaire (1859b: 417); as it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Pascoe 1866a: 473), Eucyrtus was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Eudissonomus G.S. Medvedev, 1968a: 218 [M]. Type species: Heterophylussubstriatus Reitter, 1898, by original designation. Status: valid subgenus of Dissonomus Jacquelin du Val, 1861 in Tenebrioninae: Dissonomini.

Eudysantes Bouchard, Lawrence, Davies & Newton, 2005: 508 [M]. Type species [automatic]: Dicerodereselongatus Redtenbacher, 1868, by original designation. Status: junior synonym of Dysantes Pascoe, 1871 in Tenebrioninae: Toxicini: Dysantina. Note: unnecessary replacement name for Dysantes Pascoe, 1871 (see Bouchard and Bousquet 2020a: 101 for additional comments).

Euglyptonotus Gestro, 1901: 744 [M]. Type species: Euglyptonotusmagrettii Gestro, 1901, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Euhelaeus Gebien, 1921a: 281 [M]. Type species: Euhelaeusspeculiferus Gebien, 1921, by monotypy. Status: junior synonym of Emcephalus W. Kirby, 1828 in Tenebrioninae: Heleini: Heleina. Synonymy: Matthews (1993: 1074).

Euhemicera Ando, 1996: 189, 197 [F]. Type species: Epilampuspulcher Hope, 1843, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Eulabis Eschscholtz, 1829: 14 [F]. Type species: Eulabisbicarinata Eschscholtz, 1829, by subsequent designation (Blaisdell 1932: 44). Status: valid genus in Tenebrioninae: Eulabini.

Eulea Carter, 1937: 131 [F]. Type species: Euleacaeca Carter, 1937, by monotypy. Status: valid genus in Lagriinae: Belopini.

Euleantus Haag-Rutenberg, 1876: 85 [M]. Type species: Euleantushumeralis Haag-Rutenberg, 1876, by monotypy. Status: junior synonym of Rhammatodes Haag-Rutenberg, 1876 in Pimeliinae: Tentyriini. Synonymy: Koch (1952a: 133).

Eulipus Wollaston, 1864: 448 [M]. Type species: Tentyriaelongata Brullé, 1839 (= Tentyriabrullaei Wollaston, 1865), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Eulytus C.O. Waterhouse, 1882a: 175 [M]. Type species: Eulytusnodipennis C.O. Waterhouse, 1882, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Eumicrositus Viñolas, 1990: 57, 62 [M]. Type species: Pedinusulissiponensis Germar, 1823, by original designation. Status: valid subgenus of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina. Note: this name was first proposed by Español (1947: 11, 15) without type species designation and is therefore unavailable from that date.

Eumolpamarygmus Pic, 1923b: 11 [M]. Type species: Eumolpamarygmusbigibbosus Pic, 1923, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Eumolparamarygmus Bremer, 2006: 8 [M]. Type species: Eumolpamarygmusnitidus Pic, 1935, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: the earlier usage of Eumolparamarygmus by Pic (1935: 24) is interpreted as an incorrect subsequent spelling of Eumolpamarygmus Pic, 1923 (see Bremer 2005: 210).

Eumolpocyriogeton Pic, 1923f: 305 [M]. Type species: Eumolpocyriogetonconvexus Pic, 1923 (= Plesiophthalmusconcameratus Bremer & Lillig, 2014), by monotypy. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Eumylada Reitter, 1904: 170 [F]. Type species: Myladinapunctifera Reitter, 1889, by subsequent designation (Gebien 1939: 461). Status: valid genus in Blaptinae: Opatrini: Opatrina.

Eunotiodes Casey, 1907: 519 [M]. Type species: Eunotiodesbrevicollis Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Euomma Boheman, 1858: 101 [N]. Type species: Euommalaterale Boheman, 1858, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Euomophlus Iablokoff-Khnzorian, 1983: 134 [M]. Type species: Cistelarugosicollis Brullé, 1832, by original designation. Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini.

†Eupachypterus Kirejtshuk, Nabozhenko & Nel, 2010: 192 [M]. Type species: Eupachypteruseocenicus Kirejtshuk, Nabozhensko & Nel, 2010, by original designation. Status: valid genus in Blaptinae: Opatrini: Neopachypterina. Note: described from Eocene deposits (France).

Eupezoplonyx Pic, 1922c: 12 [M]. Type species: Eupezoplonyxater Pic, 1922, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Eupezus Dejean, 1834: 211 [M]. Type species: Helopslongipes Fabricius, 1781, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Euphloeus Pascoe, 1887: 15 [M]. Type species: Euphloeusverrucosus Pascoe, 1887, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Euphron Dejean, 1834: 206 [M]. Type species: Helopscoerulescens Guérin-Méneville, 1831, by monotypy. Status: senior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Bousquet and Bouchard (2013a: 55). Note: nomen oblitum (see Bouchard and Bousquet 2020b: 6).

Euphrynus Fairmaire, 1897f: 114 [M]. Type species: Euphrynusspinithorax Fairmaire, 1897, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Eupomeca Solier, 1848: 289 [F]. Type species: Blapscylindrica Herbst, 1799 (= Blaps obtusa Fabricius, 1798), by subsequent designation (Nabozhenko 2008: 35). Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Gemminger in Gemminger and Harold (1870: 1866).

Eupsophulus Cockerell, 1906: 242 [M]. Type species [automatic]: Eupsophuscastaneus Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Vacronini. Note: replacement name for Eupsophus Horn, 1870.

Eupsophus Horn, 1870: 344, 347 [M]. Type species: Eupsophuscastaneus Horn, 1870, by monotypy. Status: senior synonym of Eupsophulus Cockerell, 1906 in Pimeliinae: Vacronini. Note: junior homonym of Eupsophus Fitzinger, 1843 [Amphibia].

Eupterocoma Skopin, 1974b: 152 [F]. Type species: Pterocomaganglbaueri Reitter, 1890, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Eurepipleura Bogdanov-Katjkov, 1915: 2 [F]. Type species: Anatolicaminima Bogdanov-Katjkov, 1915, by monotypy. Status: valid subgenus of Anatolica Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Eurhysodina Wasmann, 1921: 18 [F]. Type species: Rhyzodinamarshalli Blair, 1913, by original designation. Status: valid subgenus of Rhyzodina Chevrolat, 1873 in Tenebrioninae: Rhyssopaussini.

Euryasida Reitter, 1917a: 40, 58 [F]. Type species: Asidabarceloi Pérez Arcas, 1868, by monotypy. Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: F. Soldati (2008: 34).

Eurycaulinus Koch, 1937: 468 [M]. Type species: Eurycaulusquedenfeldti Heyden, 1890, by monotypy. Status: valid subgenus of Eurycaulus Fairmaire, 1868 in Blaptinae: Opatrini: Sclerina.

Eurycaulus Fairmaire, 1868: 492 [M]. Type species: Eurycaulusmarmottani Fairmaire, 1868, by monotypy. Status: valid genus and subgenus in Blaptinae: Opatrini: Sclerina.

Eurychora Thunberg, 1789: 9 [F]. Type species: Pimeliaciliata Fabricius, 1781, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: Eurychora is an incorrect subsequent spelling of the original spelling Evrychora, first used by Thunberg (1791: 116), in prevailing usage; Eurychora is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Eurychorula Koch, 1952b: 7 [F]. Type species: Eurychoraacuminata Fairmaire, 1891, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Eurygona Laporte, 1840: 187 [F]. Type species: Aulacuschilensis Gray, 1832, by monotypy. Status: senior synonym of Orthogonoderes Solier, 1841 in Pimeliinae: Praociini. Note: replacement name for Aulacus Gray, 1832; junior homonym of Eurygona Boisduval, 1836 [Lepidoptera].

Euryhelops Reitter, 1902b: 209 [M]. Type species: Helopschampioni Reitter, 1891, by monotypy. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Euryhelops Reitter, 1902b: 214 [M]. Type species: Euryhelopstiro Reitter, 1902, by subsequent designation (Gebien 1943: 424). Status: senior synonym of Zophohelops Reitter, 1902 in Tenebrioninae: Helopini: Cylindrinotina. Note: junior homonym of Euryhelops Reitter, 1902 [Tenebrionidae: Blaptinae: Platyscelidini].

Eurymetopon Eschscholtz, 1831: 5, 8 [N]. Type species: Eurymetoponrufipes Eschscholtz, 1831, by subsequent designation (Casey 1907: 288). Status: valid genus in Pimeliinae: Edrotini.

Eurymetopum Agassiz, 1846b: 151 [N]. Type species [automatic]: Eurymetoponrufipes Eschscholtz, 1831, by subsequent designation (Casey 1907: 288). Status: junior synonym of Eurymetopon Eschscholtz, 1831 in Pimeliinae: Edrotini. Note: unjustified emendation of Eurymetopon Eschscholtz, 1831, not in prevailing usage.

Eurynotus W. Kirby, 1819a: 418 [M]. Type species: Eurynotusmuricatus W. Kirby, 1819 (= Helopscapensis Fabricius, 1794), by monotypy. Status: valid genus and subgenus in Blaptinae: Platynotini: Eurynotina.

Euryostola Reitter, 1893: 202, 207 [F]. Type species: Pachyscelisminor Baudi di Selve, 1875, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Eurypera Pascoe, 1870: 106 [F]. Type species: Euryperacuprea Pascoe, 1870 (= Amarygmuspascoei Gebien, 1911), by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Blackburn (1893a: 92).

Eurypimelia Reitter, 1915: 9, 49 [F]. Type species: Tenebriosubglobosus Pallas, 1781, by subsequent designation (Chernei 2005: 104). Status: junior synonym of Camphonota Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Gebien (1937a: 837).

Euryprosodes Reitter, 1909a: 122 [M]. Type species: Prosodesareolata Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Euryprosternum Chatanay, 1914b: 3 [N]. Type species: Andremiusparallelus Chatanay, 1913, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Eusarca Chevrolat, 1845: 526 [F]. Type species: Eusarcairidipennis Chevrolat, 1845 (= Hegemonaresplendens Laporte, 1840), by monotypy. Status: junior synonym of Hegemona Laporte, 1840 in Stenochiinae: Cnodalonini. Synonymy: Duponchel (1845a: 498). Note: junior homonym of Eusarca Hübner, 1813 [Lepidoptera].

Eusattodes Casey, 1908: 56, 64 [M]. Type species: Eusattuslaevis J.L. LeConte, 1866, by original designation. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: Triplehorn (1968: 379).

Eusattus J.L. LeConte, 1851: 131 [M]. Type species: Eusattusdifficilis J.L. LeConte, 1851, by subsequent designation (Casey 1908: 56). Status: valid genus in Pimeliinae: Coniontini.

Euschatia Solier, 1851: 227 [F]. Type species: Euschatiaproxima Solier, 1851, by present designation. Status: junior synonym of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini. Synonymy: Lacordaire (1859b: 443).

Euschides J.L. LeConte, 1851: 127 [F]. Type species [automatic]: Stenomorphablapsoides Solier, 1836, by subsequent designation (Desmarest 1860: 150). Status: junior synonym of Stenomorpha Solier, 1836 in Pimeliinae: Asidini. Note: unnecessary replacement name for Stenomorpha Solier, 1836.

Euspinamarygmus Masumoto, 1989b: 295 [M]. Type species: Euspinamarygmuskaszabi Masumoto, 1989, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Eustenia Fairmaire, 1905: 303 [F]. Type species: Eusteniatenuimembris Fairmaire, 1905, by monotypy. Status: senior synonym of Tucumana Gebien, 1911 in Alleculinae: Alleculini: Xystropodina. Note: junior homonym of Eustenia Snellen, 1899 [Lepidoptera].

Eustenomacidius Nabozhenko, 2006: 807 [M]. Type species: Helopsluridus Ménétriés, 1849 (= Stenomaxlaevicollis Kraatz, 1882), by original designation. Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina.

Eustolopus Gebien, 1938b: 61 [M]. Type species: Eustolopuscalcaratus Gebien, 1938, by original designation. Status: valid genus in Pimeliinae: Adesmiini. Note: the First Reviser (Eustolopus Gebien, 1938 versus Entinopoda Gebien, 1938) is Penrith (1979: 40).

Eustrongylium Kolbe, 1895: 366 [N]. Type species: Strongyliumepiscopale Kolbe, 1895, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1911b: 590).

Eusyntelia C.O. Waterhouse, 1881: 473 [F]. Type species: Eusynteliabalfouri C.O. Waterhouse, 1881, by subsequent designation (R. Lucas 1920: 290). Status: valid genus in Pimeliinae: Tentyriini.

Eutagenia Reitter, 1886: 100, 125 [F]. Type species: Stenosissmyrnensis Solier, 1848, by subsequent designation (Iwan et al. 2020: 208). Status: valid genus in Pimeliinae: Stenosini: Stenosina. Note: Reitter (1916: 152) first noted that Stenosissmyrnensis Solier of Reitter (1886) was misidentified and proposed the new species Eutageniacribricollis for it; the earlier selection of the taxonomic species Stenosissmyrnensis Solier sensu Reitter, 1886 (= Eutageniacribricollis Reitter, 1916) by Gebien (1937a: 685) as type species is invalid since such an action would have required approval from the International Commission on Zoological Nomenclature at the time; Iwan et al. (2020: 208) fixed the nominal species Stenosissmyrnensis Solier, 1848 as the type species following Article 70.3.1 (ICZN 1999).

Euteleocera Agassiz, 1846b: 152 [F]. Type species [automatic]: Euteloceraviatica Solier, 1841, by original designation. Status: junior synonym of Eutelocera Solier, 1841 in Pimeliinae: Praociini. Note: unjustified emendation of Eutelocera Solier, 1841, not in prevailing usage.

Eutelocera Solier, 1841a: 209, 237 [F]. Type species: Euteloceraviatica Solier, 1841, by original designation. Status: valid genus in Pimeliinae: Praociini.

Eutelogonus Reitter, 1922a: 24 [M]. Type species: Helopsdavidis Fairmaire, 1884, by monotypy. Status: valid subgenus of Entomogonus Solier, 1848 in Tenebrioninae: Helopini: Helopina.

Eutelonodolinus Robiche, 2007: 450 [M]. Type species: Eutelonodolinusjolyi Robiche, 2007, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Eutelonotus Fairmaire in Alluaud, 1902: 456 [M]. Type species [automatic]: Eutelusrequieni Solier, 1843, by subsequent designation (R. Lucas 1920: 291). Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Eutelus Solier, 1843 (see Bousquet 2016a: 44).

Eutelus Solier, 1843: 56 [M]. Type species: Eutelusrequieni Solier, 1843, by subsequent designation (R. Lucas 1920: 291). Status: senior synonym of Eutelonotus Fairmaire in Alluaud, 1902 in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Lutelus, used by Solier (1843: 4); junior homonym of Eutelus Walker, 1834 [Hymenoptera].

Eutermicola Lea, 1916: 273 [M]. Type species: Eutermicolasculpticollis Lea, 1916, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Eutherama Carter, 1914c: 405 [N]. Type species: Eutheramacyaneum Carter, 1914, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Euthriptera Reitter, 1893: 203, 229 [F]. Type species: Thripteragrisescens Fairmaire, 1875, by monotypy. Status: valid genus in Pimeliinae: Pimeliini. Note: we act as First Revisers and reject the alternative original spelling Enthriptera, used by Reitter (1893: 203).

Euthysternum Chatanay, 1915a: 508 [N]. Type species: Chemolanusepiscopalis Fairmaire, 1886, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Eutichus Haag-Rutenberg, 1875b: 4, 58 [M]. Type species: Eutichuswahlbergi Haag-Rutenberg, 1875, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Eutochia J.L. LeConte, 1862: 238 [F]. Type species [automatic]: Ulomapiceum Melsheimer, 1846, by monotypy. Status: valid genus in Tenebrioninae: Ulomini. Note: replacement name for Aniara Melsheimer, 1853.

Eutomus Lacordaire, 1865: 369 [M]. Type species: Eutomusmicrographus Lacordaire, 1865, by subsequent designation (Barber 1914: 191). Status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: J.L. LeConte and Horn (1883: 232). Note: originally described as a member of Curculionoidea: Curculionidae: Scolytinae.

Eutoreuma Carter, 1914b: 78 [N]. Type species [automatic]: Toreumacupreum Carter, 1913, by monotypy. Status: senior synonym of Atoreuma Gebien, 1941 in Tenebrioninae: Heleini: Cyphaleina. Note: replacement name for Toreuma Carter, 1913; junior homonym of Eutoreuma Grote, 1872 [Lepidoptera].

Eutrapela Dejean, 1834: 215 [F]. Type species: Crioceriselongata Fabricius, 1781 (= Chrysomelaunifasciata DeGeer, 1778), by subsequent designation (Duponchel 1845a: 533). Status: senior synonym of Neoeutrapela Bousquet & Bouchard, 2013 in Lagriinae: Lagriini: Statirina. Synonymy: Bousquet and Bouchard (2013a: 61). Note: junior homonym of Eutrapela Hübner, 1809 [Lepidoptera].

Eutrapelodes Borchmann, 1929b: 132 [F]. Type species: Cteniopusgracillimus Fairmaire, 1888, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Eutriorophus Casey, 1924: 296 [M]. Type species: Eutriorophustuckeri Casey, 1924, by original designation. Status: junior synonym of Stibia Horn, 1870 in Pimeliinae: Edrotini. Synonymy: Blaisdell (1933: 210).

Euzadenos Koch, 1956a: 286 [M]. Type species: Eurynotusdelalandii Mulsant & Rey, 1854, by original designation. Status: junior synonym of Selenepistoma Dejean, 1834 in Blaptinae: Dendarini: Melambiina. Synonymy: new synonym [PB]. Note: Euzadenos Koch, 1956 was recently treated as a valid subgenus of Zadenos Laporte, 1840 (Kamiński 2015: 549); however, the type species of the older available genus name Selenepistoma Dejean, 1834 is currently placed in Euzadenos and therefore Selenepistoma has priority.

Evaniosomus Guérin-Méneville, 1834: 14 [M]. Type species: Evaniosomusorbignianus Guérin-Méneville, 1834, by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Evaostetha Novák, 2008a: 208 [F]. Type species: Evaostethapetri Novák, 2008, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Evelina J. Thomson, 1860c: 22 [F]. Type species: Evelinalacordairei J. Thomson, 1860, by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Eviropodus Koch, 1956a: 84 [M]. Type species: Trigonopusalternans Fåhraeus, 1870, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Evoplus J.L. LeConte, 1866b: 128 [M]. Type species: Evoplusferrugineus J.L. LeConte, 1866, by monotypy. Status: junior synonym of Neomida Latreille, 1829 in Diaperinae: Diaperini: Diaperina. Synonymy: Champion (1886: 175, with Arrhenoplita W. Kirby, 1837, a junior synonym of Neomida Latreille, 1829).

Exadelium Watt, 1992: 29 [N]. Type species: Adeliumrufilabrum Broun, 1886, by original designation. Status: valid genus in Lagriinae: Adeliini.

Exangeltus Blackburn, 1897b: 93 [M]. Type species: Exangeltusangustus Blackburn, 1897, by monotypy. Status: valid genus in Pimeliinae: Vacronini.

Exapinaeus Pascoe, 1882: 34 [M]. Type species: Exapinaeuspolitus Pascoe, 1882, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Exechophthalmus Ardoin, 1974a: 168 [M]. Type species: Exechophthalmusguillaumeti Ardoin, 1974, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Exerestus Bates, 1870: 268 [M]. Type species: Exerestusjansonii Bates, 1870 (= Rhinandruselongatus Horn, 1866), by monotypy. Status: junior synonym of Rhinandrus J.L. LeConte, 1866 in Tenebrioninae: Tenebrionini. Synonymy: Bates (1872a: 98).

Exeniotis Pascoe, 1871: 353 [F]. Type species: Exeniotiscollaris Pascoe, 1871, by monotypy. Status: valid genus in Lagriinae: Belopini.

Exocolena Gebien, 1914c: 43 [F]. Type species: Exocolenalongipes Gebien, 1914, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Exostira Borchmann, 1925: 353 [F]. Type species: Exostirasellata Borchmann, 1925, by subsequent designation (Borchmann 1930a: 512). Status: valid genus in Lagriinae: Lagriini: Statirina.

Extetranosis Koch, 1940b: 741 [F]. Type species: Tetranosisethasicornis Koch, 1940, by monotypy. Status: valid subgenus of Microtelopsis Koch, 1940 in Pimeliinae: Stenosini: Stenosina. Note: we act as First Revisers and select Microtelopsis Koch, 1940 as the valid name for this genus instead of Extetranosis Koch, 1940 and Hypermicrotelopsis Koch, 1940.

Fahraeus Ardoin, 1963b: 309, 349 [M]. Type species: Oplocheiruspunctatissimus Fåhraeus, 1870, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Falacer Laporte, 1840: 233 [M]. Type species: Accanthopuscupreus Laporte, 1840 (= Helopscontractus Palisot de Beauvois, 1811), by subsequent designation (Bousquet et al. 2018: 141). Status: junior synonym of Meracantha W. Kirby, 1837 in Tenebrioninae: Amarygmini. Synonymy: Lacordaire (1859b: 466).

Falsammidium Koch, 1960: 395, 405 [N]. Type species: Clitobiuslaevipennis Fairmaire, 1892, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Falsandrosus Kaszab, 1980b: 307 [M]. Type species: Falsandrosustetrops Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: Faslandrosus was introduced earlier by Kaszab (1979a: 104) but the name is unavailable from that date since it was published after 1930 without a description, a definition or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1).

Falsaspila Koch, 1952b: 28 [F]. Type species: Adelostomabatesi Haag-Rutenberg, 1875, by original designation. Status: junior synonym of Zarudnionymus Semenov-Tjan-Shansky & Bogatchev, 1947 in Pimeliinae: Adelostomini. Synonymy: Ardoin (1976: 149).

Falsastenochirus Pic, 1938: 13 [M]. Type species: Asthenochiruscontractus Fairmaire, 1894, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Falsasthenochirus Ardoin, 1965a: 637 [M]. Type species [automatic]: Asthenochiruscontractus Fairmaire, 1894, by monotypy. Status: junior synonym of Falsastenochirus Pic, 1938 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Falsastenochirus Pic, 1938, not in prevailing usage.

Falsoarthroconus Kaszab, 1978b: 52, 57 [M]. Type species: Falsoarthroconusnocturnus Kaszab, 1978, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Falsoaugolesthus Masumoto, 1993c: 38 [M]. Type species: Eucyrtuspulcher Pic, 1927, by original designation. Status: junior synonym of Tetragonomenes Chevrolat, 1878 in Stenochiinae: Cnodalonini. Synonymy: Ando et al. (2016: 65).

Falsobates Kaszab, 1941a: 5, 27 [M]. Type species: Falsobatesxantusi Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Falsobrachys Kulzer, 1954a: 65 [M]. Type species: Falsobrachyslongipes Kulzer, 1954, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Falsocaedius Español, 1943: 140, 142 [M]. Type species: Clitobiusfossulatus Escalera, 1914, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Falsocalcar Pic, 1925: 9 [N]. Type species: Falsocalcarbicolor Pic, 1925, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Falsocamaria Pic, 1917g: 19 [F]. Type species: Falsocamariaobscura Pic, 1917, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Falsocamariodes Ardoin, 1956a: 159 [M]. Type species: Falsocamariodesviettei Ardoin, 1956, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Falsocasnonidea Pic, 1934a: 31 [F]. Type species: Nemostiradiversipennis Pic, 1923, by monotypy. Status: junior synonym of Casnonidea Fairmaire, 1882 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1936: 393).

Falsocatomulus Pic, 1914c: 10 [M]. Type species: Falsocatomuluseuphraticus Pic, 1914, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Falsocosmonota Kaszab, 1962b: 75 [F]. Type species: Falsocosmonotacheni Kaszab, 1962, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Falsocossyphus Pic, 1916b: 4 [M]. Type species: Falsocossyphuspilosus Pic, 1916, by monotypy. Status: valid genus in Tenebrioninae: Falsocossyphini.

Falsocuphotes Pic, 1918b: 22 [F]. Type species: Falsocuphotescurticornis Pic, 1918, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Falsodiopethes Pic, 1924b: 12 [M]. Type species: Falsodiopethesgounellei Pic, 1924, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Falsoencyalesthus Pic, 1923c: 29 [M]. Type species: Falsoencyalesthuslatipennis Pic, 1923, by monotypy. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1969b: 125, with Encyalesthus Motschulsky, 1860, a junior synonym of Derosphaerus J. Thomson, 1858).

Falsogauromaia Pic, 1921d: 22 [F]. Type species: Falsogauromaiaannulipes Pic, 1921, by monotypy. Status: valid subgenus of Gauromaia Pascoe, 1866 in Stenochiinae: Cnodalonini.

Falsolagria Pic, 1927a: 44 [F]. Type species: Falsolagriamarmorata Pic, 1927, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Falsolobodera Kaszab, 1967: 24 [F]. Type species: Falsoloboderaskopini Kaszab, 1967, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Falsolophocnemis Pic, 1917d: 13 [F]. Type species: Falsolophocnemissinuatipes Pic, 1917, by subsequent designation (Gebien 1948: 542). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Kaszab (1977b: 29).

Falsomicrodera Kaszab, 1966: 291, 294 [F]. Type species: Microderaturkestanica Schuster, 1915, by original designation. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Falsomophlus Pic, 1925c: 11 [M]. Type species: Falsomophlusniger Pic, 1925, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Falsomycterus Pic, 1907a: 127 [M]. Type species: Falsomycterusdiversipes Pic, 1907, by monotypy. Status: valid genus in Pimeliinae: Falsomycterini.

Falsonannocerus Pic, 1947: 150 [M]. Type species: Falsonannocerusdentaticeps Pic, 1947, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Falsonemostira Pic, 1917c: 14 [F]. Type species: Nemostiraannulipes Pic, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Falsonotostrongylium Kaszab, 1955a: 538, 552 [N]. Type species: Falsonotostrongyliumbradymeroides Kaszab, 1955, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Falsoperichilus Pic, 1920a: 16 [M]. Type species: Falsoperichilussemirugosus Pic, 1920, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Falsophthora Kaszab, 1977a: 306, 309 [F]. Type species: Tagalusfordi Kulzer, 1957, by monotypy. Status: valid subgenus of Pseudophthora Kaszab, 1970 in Phrenapatinae: Penetini.

Falsoplonyx Ardoin, 1963b: 308, 337 [M]. Type species: Gonocnemisrubripes Fairmaire, 1899, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Falsopraocis Kulzer, 1958a: 5, 102 [M]. Type species: Amphidoraricardae (as “richardae”) Solier, 1851, by original designation. Status: valid genus in Pimeliinae: Praociini.

Falsopsilonycha Pic, 1931c: 303 [F]. Type species: Psilonychausambarana Pic, 1917, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Falsostrongylium Pic, 1915: 11 [N]. Type species: Falsostrongyliumsemirufum Pic, 1915, by subsequent designation (Gebien 1948: 539). Status: valid genus in Stenochiinae: Stenochiini.

Falsosynopticus Pic, 1936b: 17 [M]. Type species: Falsosynopticusater Pic, 1936, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Falsotagalus Kaszab, 1977a: 301, 310 [M]. Type species: Falsotagalussubcoecus Kaszab, 1977, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Falsotithassa Pic, 1934b: 18 [F]. Type species: Falsotithassasumatrana Pic, 1934, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Falsozialeus Pic, 1951: 11 [M]. Type species: Falsozialeusater Pic, 1951 (= Asyleptusfumosus Péringuey, 1896), by monotypy. Status: junior synonym of Asyleptus Péringuey, 1896 in Tenebrioninae: Amarygmini. Synonymy: Ardoin (1962b: 957); Schawaller and Bremer (2013: 81).

Falsozotypus Kaszab, 1980b: 334 [M]. Type species: Falsozotypusopacipennis Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: this name was first published by Kaszab (1979a: 108) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article (13.1) and is therefore unavailable from that date.

Farsarthrosis Kaszab, 1979a: 86 [F]. Type species: Farsarthrosisbenardi Kaszab, 1979, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Faustia Kraatz, 1882: 92 [F]. Type species: Faustiamodesta Kraatz, 1882, by monotypy. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Ferganoprosodes G.S. Medvedev, 1997: 595 [M]. Type species: Prosodesangulicollis Kraatz, 1883, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Ferveoventer Smith, 2013: 604 [M]. Type species: Ferveoventerbrowni Smith, 2013, by original designation. Status: valid genus in Pimeliinae: Asidini.

Fifina Novák, 2018d: 470 [F]. Type species: Fifinaromani Novák, 2018, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Fifinoides Novák, 2020h: 78 [M]. Type species: Fifinoideschinensis Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Filotarsus Solier, 1841a: 209, 239 [M]. Type species: Filotarsustenuicornis Solier, 1841, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini.

Fitzsimonsia Koch, 1955b: 415 [F]. Type species: Fitzsimonsiacymbium Koch, 1955, by original designation. Status: senior synonym of Fitzsimonsium Koch, 1962 in Pimeliinae: Stenosini: Platamodina. Note: junior homonym of Fitzsimonsia Witte, 1943 [Reptilia].

Fitzsimonsium Koch, 1962b: 152 [N]. Type species [automatic]: Fitzsimonsiacymbium Koch, 1955, by original designation. Status: valid genus in Pimeliinae: Stenosini: Platamodina. Note: replacement name for Fitzsimonsia Koch, 1955.

Flabellalogista Pic, 1954: 256 [F]. Type species: Flabellalogistaminuta Pic, 1954, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Flabellolagria Pic, 1927c: 27 [F]. Type species: Flabellolagrialuteovittata Pic, 1927, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Flabellostrongylium Pic, 1938: 18 [N]. Type species: Flabellostrongyliumatronitidum Pic, 1938, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Flavipoda Campbell, 1966: 21 [F]. Type species: Helopsflavipes Fabricius, 1792 [as “Alleculaflavipes Jacquelin du Val, 1857”], by original designation. Status: valid subgenus of Lobopoda Solier, 1835 in Alleculinae: Alleculini: Alleculina.

Foleya Peyerimhoff, 1916: 71 [F]. Type species: Foleyabrevicornis Peyerimhoff, 1916, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Foochounus Pic, 1921d: 22 [M]. Type species: Foochounusconvexipennis Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Foranotum Nabozhenko & Sadeghi, 2017: 165 [N]. Type species: Foranotumperforatum Nabozhenko & Sadeghi, 2017, by original designation. Status: valid genus in Kuhitangiinae: Foranotini.

Fossilochile Koch, 1952c: 63 [F]. Type species: Fossilochilerufa Koch, 1952, by original designation. Status: junior synonym of Pachynotelus Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Synonymy: Penrith and Endrödy-Younga (1994: 54).

Fourtaus Pic in Alfieri, 1921: 47 [M]. Type species: Fourtausbrevicornis Pic, 1921 (= Hegeterocaraarabicum Reitter, 1900), by monotypy. Status: junior synonym of Hegeterocara Reitter, 1900 in Pimeliinae: Tentyriini. Synonymy: Koch (1941: 294).

Foveostatira Pic, 1918b: 24 [F]. Type species: Statirafoveicollis Pic, 1918, by subsequent designation (Borchmann 1936: 244). Status: valid subgenus of Statira Lepeletier & Audinet-Serville, 1828 in Lagriinae: Lagriini: Statirina.

Freudeia Kaszab, 1961b: 216 [F]. Type species: Freudeianepalica Kaszab, 1961, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Freudella Ardoin, 1961b: 30 [F]. Type species: Freudellaauripunctata Ardoin, 1961, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Freyitia Koch, 1943a: 522, 525 [F]. Type species: Freyitiaornatipes Koch, 1943, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Freyula Koch, 1959: 591, 593 [F]. Type species: Freyulapsammarina Koch, 1959, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Fundulus Gistel, 1848a: 125 [M]. Type species [automatic]: Opatrumtibiale Fabricius, 1781, by monotypy. Status: junior synonym of Melanimon Steven, 1828 in Tenebrioninae: Melanimonini. Note: unnecessary replacement name for Microzoum Dejean, 1834; junior homonym of Fundulus Lacepède, 1803 [Pisces].

Gabonia Fairmaire, 1894e: 326 [F]. Type species: Gaboniadenticulata Fairmaire, 1894, by monotypy. Status: senior synonym of Gabonisca Fairmaire, 1894 in Lagriinae: Pycnocerini. Note: junior homonym of Gabonia Jacoby, 1893 [Coleoptera: Chrysomelidae].

Gabonisca Fairmaire, 1894f: 395 [F]. Type species [automatic]: Gaboniadenticulata Fairmaire, 1894, by monotypy. Status: valid genus in Lagriinae: Pycnocerini. Note: replacement name for Gabonia Fairmaire, 1894.

Gahanosis Penrith, 1983: 371, 379 [F]. Type species: Zophosisundulata Gahan, 1900, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Gamaxus Bates, 1868: 315 [M]. Type species: Gamaxushauxwellii Bates, 1868, by monotypy. Status: valid genus in Lagriinae: Goniaderini. Note: combined description of a new genus and single new species (ICZN 1999, Article 12.2.6).

Garambanus Ardoin, 1964a: 847 [M]. Type species: Garambanusverschureni Ardoin, 1964, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Gargilius Fairmaire, 1891b: 251 [M]. Type species: Gargiliustrispinosus Fairmaire, 1891, by subsequent designation (Gebien 1940: 421). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Garridoa Marcuzzi, 1985: 180 [F]. Type species: Garridoakaszabi Marcuzzi, 1985, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Gastrhaema Jacquelin du Val, 1863: 353 [N]. Type species: Cistelarufiventris Waltl, 1835, by subsequent designation (Novák and Pettersson 2008: 331). Status: valid genus in Alleculinae: Cteniopodini.

Gaurobates Gebien, 1928: 170, 184 [M]. Type species: Gaurobatespictus Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Gauromaia Pascoe, 1866a: 473 [F]. Type species: Gauromaiadives Pascoe, 1866, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Gebienella Kaszab, 1941a: 4, 21 [F]. Type species: Gebienellainterrumpens Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Gebienia Borchmann, 1921: 219, 352 [F]. Type species: Gebieniarimulosa Borchmann, 1921, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Gebieniella Koch, 1940b: 736 [F]. Type species: Ethasstenosides Pascoe, 1862, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Gebienocamaria Masumoto, 1993b: 223, 232 [F]. Type species: Camariaangulicollis Fairmaire, 1896, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Gebleria Motschulsky, 1846: 410 [F]. Type species: Dilaphilacoides Fischer von Waldheim, 1844, by monotypy. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Gedeon Reiche & Saulcy, 1857: 219 [M]. Type species: Gedeonhierichonticus Reiche & Saulcy, 1857, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Gedrosia Bogatchev, 1961: 111 [F]. Type species: Gedrosiamonstrosa Bogatchev, 1961 (= Pseudopodhomalagabrieli Schuster, 1938), by original designation. Status: junior synonym of Pseudopodhomala Schuster, 1938 in Pimeliinae: Pimeliini. Synonymy: Löbl et al. (2008b: 166). Note: junior homonym of Gedrosia Stål, 1862 [Hemiptera].

Genateropa Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Apterogenacanonnei Ardoin, 1962, by original designation. Status: valid genus in Stenochiinae: Stenochiini. Note: replacement name for Apterogena Ardoin, 1962.

Genoblaps Bauer, 1921: 230 [F]. Type species: Blaps tentyroides Seidlitz, 1893 (= Blapssocia Seidlitz, 1893), by monotypy. Status: valid subgenus of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina.

Gentinadis Laporte, 1840: 240 [M]. Type species: Stenochiacaerulea Laporte, 1840 (= Strongyliumazureum Germar, 1823), by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Lacordaire (1859b: 484).

Geoborus Blanchard, 1842: pl. 13 [M]. Type species: Geoboruscostatus Blanchard, 1842 (= Gyriosomuslineatus Guérin-Méneville, 1834), by subsequent designation (Pizarro-Araya and Flores 2006: 95). Status: valid genus in Pimeliinae: Epitragini.

Geophanus Haag-Rutenberg, 1875b: 4, 46 [M]. Type species: Psaryphisconfusa Fåhraeus, 1870, by subsequent designation (Gebien 1937a: 675). Status: valid genus in Pimeliinae: Adelostomini.

Gerandryus Rottenberg, 1873: 217 [M]. Type species [automatic]: Parablopsaetnensis Rottenberg, 1871, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: replacement name for Parablops Rottenberg, 1871.

Gerardia Pic, 1954: 258 [F]. Type species: Gerardiasublineata Pic, 1954, by monotypy. Status: senior synonym of Piccula Bousquet & Bouchard, 2015 in Alleculinae: Alleculini: Gonoderina. Note: junior homonym of Gerardia Lacaze-Duthiers, 1864 [Cnidaria].

Gerdacula Novák, 2015b: 145 [F]. Type species: Gerdaculafujianica Novák, 2015, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Ghaleca Péringuey, 1899: 316 [F]. Type species: Ghalecalaeta Péringuey, 1899 (= Alymonprolatus Pascoe, 1866), by monotypy. Status: junior synonym of Alymon Pascoe, 1866 in Tenebrioninae: Amarygmini. Synonymy: Péringuey (1904: 297, through synonymy of its type species with Alymonprolatus Pascoe, 1866).

Gibbostrongylium Pic, 1917d: 18 [N]. Type species: Strongyliummedanense Pic, 1917, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Gigantopigeus Kaszab, 1984: 355, 374 [M]. Type species: Gigantopigeusmirabilis Kaszab, 1984, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Girardius L. Soldati, 2009: 73 [M]. Type species: Micipsapersica Baudi, 1874, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Girardocamaria Masumoto, 1993b: 227, 232 [F]. Type species: Girardocamariaardoini Masumoto, 1993, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Glabrasida Escalera, 1910: 408 [F]. Type species: Glabrasidaconspuata Escalera, 1910, by subsequent designation (F. Soldati 2008: 32). Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini.

Glabrilobopoda Campbell, 1966: 46 [F]. Type species: Lobopodaglabrata Champion, 1888, by original designation. Status: valid subgenus of Lobopoda Solier, 1835 in Alleculinae: Alleculini: Alleculina.

Globasida Escalera, 1905b: 430 [F]. Type species: Asidaoblonga Rambur, 1838, by subsequent designation (Viñolas and Cartagena 2005: 209). Status: valid subgenus of Asida Latreille, 1802 in Pimeliinae: Asidini.

Globularthrodosis Kaszab, 1979a: 95 [F]. Type species: Diaphanidussemenowi Reitter, 1900, by original designation. Status: junior synonym of Diaphanidus Reitter, 1900 in Pimeliinae: Erodiini. Synonymy: G.S. Medvedev and Nepesova (1985: 52).

Glyptariobius Koch, 1948: 423 [M]. Type species: Hoplariobiusexacavatus Koch, 1948, by original designation. Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Glyptasida Casey, 1912: 75, 95 [F]. Type species: Ophryastessordidus J.L. LeConte, 1853, by original designation. Status: valid subgenus of Philolithus Lacordaire, 1858 in Pimeliinae: Asidini.

Glyptophrynus Fairmaire, 1899e: 532 [M]. Type species: Glyptophrynustenuesculptus Fairmaire, 1899, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Glyptopteryx Gebien, 1910c: 376 [F]. Type species: Glyptopteryxforticostis Gebien, 1910 (= Selinusquadricollis Fairmaire, 1887), by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Glyptothorax Borchmann, 1937: 223 [M]. Type species: Glyptothoraxpilosus Borchmann, 1937, by original designation. Status: senior synonym of Borchmannius Bousquet & Bouchard, 2015 in Alleculinae: incertae sedis. Note: junior homonym of Glyptothorax Blyth, 1860 [Pisces]; taxon also described as new by Borchmann (1938: 122).

Glyptotus J.L. LeConte, 1858a: 75 [M]. Type species: Glyptotuscribratus J.L. LeConte, 1858, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Gnaptor Brullé, 1831: 254 [M]. Type species: Tenebriospinimanus Pallas, 1781, by monotypy. Status: valid genus and subgenus in Blaptinae: Blaptini: Gnaptorina. Note: this name was previously attributed to Brullé (1832: 202 as Petrobius, corrected to Gnaptor in the “Errata” of the same work) in the literature.

Gnaptorina Reitter, 1887a: 364 [F]. Type species: Gnaptorinafelicitana Reitter, 1887, by monotypy. Status: valid genus and subgenus in Blaptinae: Blaptini: Gnaptorinina.

Gnathelops Gebien, 1922b: 320 [M]. Type species: Gnathelopschatanayi Gebien, 1922, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: removed from the tribe Helopini and placed as Tenebrionidae incertae sedis by Nabozhenko (2018: 183).

Gnathidium Gebien, 1921b: 41 [N]. Type species: Gnathidiumcephalotes Gebien, 1921, by monotypy. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Gnathocerus Agassiz, 1846b: 164 [M]. Type species [automatic]: Gnatocerusruber Thunberg, 1814 (= Trogossitacornuta Fabricius, 1798), by monotypy (see ICZN 1975, Opinion 1039). Status: junior synonym of Gnatocerus Thunberg, 1814 in Diaperinae: Diaperini: Adelinina. Note: unjustified emendation of Gnatocerus Thunberg, 1814, not in prevailing usage.

Gnathosia Fischer, 1821: 13 [F]. Type species: Gnathosiaglabra Fischer, 1821, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Gnatocerus Thunberg, 1814: 47 [M]. Type species: Gnatocerusruber Thunberg, 1814 (= Trogossitacornuta Fabricius, 1798), by monotypy (see ICZN 1975, Opinion 1039). Status: valid genus and subgenus in Diaperinae: Diaperini: Adelinina. Note: placed on the Official List of Generic Names in Zoology by the ICZN (1975, Opinion 1039).

Gnesis Pascoe, 1866a: 477 [M]. Type species: Gnesishelopioides Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Gnophota Erichson, 1843: 237 [F]. Type species: Gnophotaantracina Erichson, 1843, by subsequent designation (R. Lucas 1920: 306). Status: valid genus in Pimeliinae: Tentyriini.

Goajiria Ivie & Hart, 2016: 474 [F]. Type species: Diastolinuscurtus Mulsant & Rey, 1859, by original designation. Status: valid subgenus of Diastolinus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Blapstinina. Note: name first introduced by Marcuzzi (1986: 180) without a type species designation.

Gobretus Freude, 1967: 148, 152 [M]. Type species: Epitraguscephalotes Freude, 1967, by monotypy. Status: valid subgenus of Epitragus Latreille, 1802 in Pimeliinae: Epitragini.

Gonasida Casey, 1912: 75, 117 [F]. Type species: Pelecyphoruselatus J.L. LeConte, 1853, by original designation. Status: valid subgenus of Philolithus Lacordaire, 1858 in Pimeliinae: Asidini.

Gondvanadelium Kaszab, 1981a: 81 [N]. Type species: Gondvanadeliumseirotranoides Kaszab, 1981, by original designation. Status: valid genus in Lagriinae: Adeliini.

Gondwanocrypticus Español, 1955: 10 [M]. Type species: Crypticusplatensis Fairmaire, 1884, by original designation. Status: valid genus in Diaperinae: Crypticini. Note: this name was introduced earlier by Koch (1950c: 64) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Gondwanodilamus Kaszab, 1969b: 320 [M]. Type species: Conibiusfranzi Kaszab, 1969, by original designation. Status: junior synonym of Conibius J.L. LeConte, 1851 in Blaptinae: Opatrini: Blapstinina. Synonymy: new synonym [RLA]. Note: the new synonymy is based on taxonomic studies of all known species in the genus Conibius J.L. LeConte, 1851 and follows the previous synonymization of the two other subgenera previously treated as valid, Ooconibius Casey, 1895 and Euconibius Casey, 1895, by Aalbu in Bousquet et al. (2018: 202).

Gonespites Gebien, 1921a: 325, 396 [M]. Type species: Gonespitessubcrenatus Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Goniadera Perty, 1832: 62 [F]. Type species: Goniaderacrenata Perty, 1832, by monotypy. Status: valid genus and subgenus in Lagriinae: Goniaderini. Note: the alternative original spelling Gonyodera, used by Perty (1832: 63), was rejected by Perty (1833: 14) who acted as the First Reviser.

†Gonialaena Nabozhenko, Bukejs & Telnov, 2019: 254 [F]. Type species: Gonialaenagroehni Nabozhenko, Bukejs & Telnov, 2019, by original designation. Status: valid genus in Lagriinae: Gonialaenini. Note: described from Eocene Baltic amber.

Goniodera Agassiz, 1846b: 165 [F]. Type species [automatic]: Goniaderacrenata Perty, 1832, by monotypy. Status: junior synonym of Goniadera Perty, 1832 in Lagriinae: Goniaderini. Note: unjustified emendation of Goniadera Perty, 1832, not in prevailing usage.

Gonocephalum Solier, 1834: 498 [N]. Type species: Opatrumfuscum Herbst, 1793 (= Opatrumrusticum G.-A. Olivier, 1812), by subsequent designation (Gebien 1939: 443). Status: valid genus and subgenus in Blaptinae: Opatrini: Opatrina.

Gonocnemis J. Thomson, 1858: 101 [F]. Type species: Gonocnemisstrigipennis J. Thomson, 1858, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Gonocnemocistela Pic, 1935a: 22 [F]. Type species: Gonocnemocistelalutea Pic, 1935, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Gonodera Mulsant, 1856a: 41 [F]. Type species: Cistelafulvipes Fabricius, 1792 (= Cistelaluperus Herbst, 1783), by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Gonogenius Solier, 1838b: 8, 48 [M]. Type species: Scotobiusvulgaris Guérin-Méneville, 1834, by original designation. Status: junior synonym of Scotobius Germar, 1823 in Tenebrioninae: Scotobiini. Synonymy: Lacordaire (1859a: 129).

Gonopterus Solier, 1843: 101 [M]. Type species: Sepidiumrugosum Fabricius, 1781, by monotypy. Status: junior synonym of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Gemminger in Gemminger and Harold (1870: 1900).

Gonopus Latreille, 1828: 580 [M]. Type species: Blapstibialis Fabricius, 1798, by monotypy. Status: valid genus and subgenus in Blaptinae: Platynotini: Platynotina.

Gonospa Champion, 1886: 216 [F]. Type species: Gonospaphaedonoides Champion, 1886, by subsequent designation (Gebien 1940: 426). Status: valid genus in Stenochiinae: Cnodalonini.

Grabulax Kanda, 2016: 558 [M]. Type species: Grabulaxdarlingtoni Kanda, 2016, by original designation. Status: valid genus in Lagriinae: Laenini.

Gracilasida Escalera, 1905b: 445 [F]. Type species: Asidaariasi Escalera, 1909, by subsequent designation (F. Soldati 2008: 32). Status: valid subgenus of Asida Latreille, 1802 in Pimeliinae: Asidini. Note: this genus was established without included nominal species; Asidaariasi Escalera, 1909 and A.pusillima Kraatz, 1874 were first subsequently and expressly included in Gracilasida by Escalera (1909: 135–136).

Graecopachys Skopin, 1968a: 99 [M]. Type species: Tentyriaquadricollis Brullé, 1832, by original designation. Status: junior synonym of Phymatiotris Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Löbl et al. (2008a: 43).

Grammicus G.R. Waterhouse, 1845b: 323 [M]. Type species: Grammicuschilensis G.R. Waterhouse, 1845, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Granasida Reitter, 1917a: 9, 14 [F]. Type species: Asidagranulifera Chevrolat, 1840, by monotypy. Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini.

Grandelagria Pic, 1940: 3 [F]. Type species: Grandelagriabicoloripes Pic, 1940, by monotypy. Status: valid subgenus of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina.

Grandicyrtomius Freude, 1967: 225 [M]. Type species: Epitragusgrandis Champion, 1884, by original designation. Status: valid subgenus of Cyrtomius Casey, 1907 in Pimeliinae: Epitragini.

Granulasida Escalera, 1922a: 427 [F]. Type species: Asidasetipennis Allard, 1869, by subsequent designation (Viñolas and Cartagena 2005: 192). Status: junior synonym of Gracilasida Escalera, 1905 in Pimeliinae: Asidini. Synonymy: F. Soldati (2008: 34, with Planasida Escalera, 1907, a junior synonym of Gracilasida Escalera, 1905).

Granulophanes Nabozhenko, 2013: 2 [M]. Type species: Hedyphaneslutosus Allard, 1877, by original designation. Status: valid subgenus of Hedyphanes Fischer, 1820 in Tenebrioninae: Helopini: Helopina.

Graptopezus Gebien, 1921a: 296 [M]. Type species: Seteniscostipennis Blair, 1915 (= Nyctozoiluscrenaticollis W.J. MacLeay, 1886), by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Gressittiola Kaszab, 1955a: 462, 464 [F]. Type species: Gressittiolaplatydemoides Kaszab, 1955, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Gridellia Kammerer, 2006: 270 [F]. Type species [automatic]: Tenebrioclypealis Gebien, 1920, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini. Note: replacement name for Villiersia Gridelli, 1951.

Gridelliopus Koch, 1956a: 358 [M]. Type species: Gridelliopussubsquamosus Koch, 1956, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Gronophora Borchmann, 1916a: 48, 103 [F]. Type species: Gronophoragravida Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Guanobius Grimm, 2008: 375 [M]. Type species: Guanobiusborneensis Grimm, 2008, by original designation. Status: valid genus in Tenebrioninae: Alphitobiini.

Guildia Antoine, 1957: 346, 359 [F]. Type species: Microsituspunctistriatus Escalera, 1925, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Gunarellus Reitter, 1922a: 22 [M]. Type species: Helopsgratus Frivaldszky, 1894, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Stenohelops Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko et al. (2020b: 297). Note: Gunarellus was first described by Reitter (1922a: 22, in key; issued 30 March 1922) without originally included species; Reitter (1922b: 114–115; issued 25 October 1922) was the first author to subsequently and expressly include nominal species in Gunarellus (ICZN 1999, Article 67.2.2).

Gunarus Gozis, 1886: 25 [M]. Type species: Helopshirtulus Reiche, 1862, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Gymnetasida Reitter, 1917a: 10, 22 [F]. Type species: Asidatricostata Allard, 1869, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Gymnognathus Solier, 1851: 136 [M]. Type species: Gymnognathusfuscus Solier, 1851, by monotypy. Status: junior synonym of Arthroconus Solier, 1851 in Pimeliinae: Edrotini. Synonymy: Lacordaire (1859a: 67). Note: junior homonym of Gymnognathus Schönherr, 1823 [Coleoptera: Anthribidae].

Gynandrocera Gebien, 1920: 46 [F]. Type species: Gynandroceracephalotes Gebien, 1920 (= Afrinusminor Péringuey, 1908), by subsequent designation (Koch 1950b: 328). Status: valid subgenus of Afrinus Fairmaire, 1888 in Pimeliinae: Tentyriini.

Gyrasida Koch, 1962a: 127, 143 [F]. Type species: Afrasidapropensa Wilke, 1922, by original designation. Status: valid genus in Pimeliinae: Praociini. Note: elevated to the rank of genus and transferred from Pimeliinae: Asidini by Flores and Vidal (2009: 52); as mentioned by Flores and Vidal (2009: 48, 56) the type species was originally described from South Africa in error: the genus Gyrasida Koch, 1962 is endemic to Chile.

Gyrinodes Fauvel, 1897: 61 [M]. Type species: Helopsgagatinus Küster, 1850, by monotypy. Status: junior synonym of Nesotes Allard, 1876 in Tenebrioninae: Helopini: Helopina. Synonymy: new synonym [YB]. Note: this genus-group taxon has been forgotten in the literature; its type species is currently placed in the nominotypical subgenus of Nesotes Allard, 1876 and for that reason Fauvel’s name is regarded as a new junior synonym of Nesotes.

Gyriosomus Guérin-Méneville, 1834: 6 [M]. Type species: Nyctelialuczotii Guérin-Méneville, 1831, by subsequent designation (Duponchel 1845b: 449). Status: valid genus in Pimeliinae: Nycteliini.

Gyrosis Gebien, 1920: 33, 37 [F]. Type species: Zophosisorbicularis Deyrolle, 1867, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Habrobates Semenov, 1903a: 11 [M]. Type species: Habrobatesvernalis Semenov, 1903, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Habrochiton Semenov-Tjan-Shansky, 1907a: 177, 179 [M]. Type species: Habrochitonvernus Semenov-Tjan-Shansky, 1907, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Hades J. Thomson, 1860a: 13 [M]. Type species: Hadestenebrosus J. Thomson, 1860 (= Platydemahemisphaerica Laporte & Brullé, 1831), by monotypy. Status: senior synonym of Pimplema Pascoe, 1887 in Diaperinae: Leiochrinini. Synonymy: Gebien (1940: 755). Note: junior homonym of Hades Westwood, 1851 [Lepidoptera].

Hadroderus Koch, 1956a: 347 [M]. Type species: Hadroderustuberculiferus Koch, 1956, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Hadrodes Wollaston, 1877: 226 [M]. Type species: Hadrodeshelenensis Wollaston, 1877, by monotypy. Status: valid genus in Blaptinae: Opatrini: AMMOBIINA.

Hadromelambius Koch, 1948: 432 [M]. Type species: Melambiustelueticus Escalera, 1914, by original designation. Status: valid subgenus of Melambius Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Hadrophasis Ferrer, 1992: 87 [F]. Type species: Hadrophasisangolensis Ferrer, 1992, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Hadrus Wollaston, 1854: 502 [M]. Type species: Hadrusalpinus Wollaston, 1854, by subsequent designation (R. Lucas 1920: 313). Status: senior synonym of Wolladrus Iwan & Kamiński, 2016 in Blaptinae: Opatrini: Opatrina. Note: junior homonym of Hadrus Perty, 1833 [Diptera].

Haemerophygus Baudi di Selve, 1876b: 266 [M]. Type species: Helopsmucoreus Waltl, 1838, by monotypy. Status: junior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Synonymy: Nabozhenko et al. (2016b: 608).

Haemodus Gebien, 1943: 904 [M]. Type species [automatic]: Haemuscarinatipennis Péringuey, 1904, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina. Note: replacement name for Haemus Péringuey, 1904.

Haemus Péringuey, 1904: 228 [M]. Type species: Haemuscarinatipennis Péringuey, 1904, by monotypy. Status: senior synonym of Haemodus Gebien, 1943 in Blaptinae: Dendarini: Melambiina. Note: junior homonym of Haemus Stål, 1862 [Hemiptera].

Halammobia Semenov, 1901: 92 [F]. Type species: Tenebriopellucidus Herbst, 1799, by monotypy. Status: valid genus in Diaperinae: Phaleriini.

Halonomus Wollaston, 1861: 201 [M]. Type species: Halonomusgrayii Wollaston, 1861 (= Opatrumovatum Erichson, 1843), by subsequent designation (Iwan and Löbl 2007: 734). Status: junior synonym of Clitobius Mulsant & Rey, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Fairmaire (1886b: ccv). Note: the original combination of the accepted name of the type species, Opatrumovatum Erichson, 1843, is a junior primary homonym of Opatrumovatum Fabricius, 1801.

Halophalerus Crotch, 1874: 107 [M]. Type species: Phaleriarotundata J.L. LeConte, 1851, by subsequent designation (Bousquet et al. 2018: 289). Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Synonymy: Austin (1880: 38).

Hangaya Matthews & Merkl, 2015: 479 [F]. Type species: Hangayaenigmatica Matthews & Merkl, 2015, by original designation. Status: valid genus in Tenebrioninae: incertae sedis. Note: according to Matthews and Lawrence (2019: 628) the closest relative of this genus within the subfamily Tenebrioninae remains to be determined.

Hanstroemium Koch, 1953c: 19 [N]. Type species: Hanstroemiumadelostomoide Koch, 1953, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Haplandrus J.L. LeConte, 1862: 230 [M]. Type species: Helopsfemoratus Fabricius, 1798 (= Upisfulvipes Herbst, 1797), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Haporema Fairmaire, 1892a: 109 [F]. Type species: Haporemadecipiens Fairmaire, 1892, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hapsida Gemminger in Gemminger and Harold, 1870: 1955 [F]. Type species [automatic]: Apsidachrysomelina Lacordaire, 1859, by original designation. Status: junior synonym of Apsida Lacordaire, 1859 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Apsida Lacordaire, 1859, not in prevailing usage.

Harvengia Ferrer, 2004b: 367 [F]. Type species: Harvengiavietnamita Ferrer, 2004, by original designation. Status: valid genus in Pimeliinae: Stenosini: Harvengiina.

Hasticollinum Kaszab, 1939a: 96 [N]. Type species: Hasticollinumpodagrarium Kaszab, 1939, by original designation. Status: junior synonym of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Ferrer (1995a: 72).

Havanalia Novák, 2020c: 489 [F]. Type species: Havanaliaqazvinica Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Hectus Pascoe, 1869: 288, 289 [M]. Type species: Hectusanthracinus Pascoe, 1869, by monotypy. Status: junior synonym of Olisthaena Erichson, 1842 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1913a: 62).

Hedrotes Gemminger in Gemminger and Harold, 1870: 1816 [M]. Type species [automatic]: Edrotesventricosus J.L. LeConte, 1851, by monotypy. Status: junior synonym of Edrotes J.L. LeConte, 1851 in Pimeliinae: Edrotini. Note: unjustified emendation of Edrotes J.L. LeConte, 1851, not in prevailing usage.

Hedyphanes Fischer, 1820: pl.15 [M]. Type species: Hedyphanescoerulescens Fischer, 1820, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina.

Hegemona Laporte, 1840: 230 [F]. Type species: Hegemonaresplendens Laporte, 1840, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hegeter Latreille, 1802: 172 [M]. Type species: Hegeterstriatus Latreille, 1804 (= Blapstristis Fabricius, 1792), by subsequent monotypy (Latreille 1804: 276). Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: originally proposed without included nominal species; Latreille (1804: 276), by including the species Hegeterstriatus Latreille, 1804 in association with this name, was the first author to subsequently and expressly include nominal species in Hegeter (ICZN 1999, Article 67.2.2); the previously accepted type species for this genus was Blapselongata G.-A. Olivier, 1795 (= Blapstristis Fabricius, 1792), by subsequent designation (Latreille 1810: 429); Latreille (1804: 276) mentioned “Le type de ce genre paroît être le blaps alongé d’Olivier” however he did not use a Latin name for the species.

Hegeterocara Reitter, 1900c: 94, 190 [N]. Type species: Hegeterocaraarabicum Reitter, 1900, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Hegeteromorpha Escalera, 1913: 38 [F]. Type species: Pachychilaexternecostata Haag-Rutenberg, 1875, by subsequent monotypy (Escalera 1914: 281). Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: originally proposed without included nominal species; Escalera (1914: 281), by including the species Pachychilaexternecostata Haag-Rutenberg, 1875 in association with this name, was the first author to subsequently and expressly include nominal species in Hegeteromorpha (ICZN 1999, Article 67.2.2).

Heinrichesia Carl, 2000: 258 [F]. Type species: Heinrichesiaschaeferi Carl, 2000, by monotypy. Status: junior synonym of Waterhousia Skopin, 1973 in Pimeliinae: Pimeliini. Synonymy: G.S. Medvedev (2005a: 309). Note: combined description of new genus-group taxon and new species (ICZN 1999, Article 13.4).

Helea Latreille, 1804: 326 [F]. Type species: Heleaperforata Latreille, 1817, by subsequent monotypy (Latreille 1817: 261). Status: valid genus in Tenebrioninae: Heleini: Heleina. Note: the name Helea Meigen, 1800 (Diptera) was suppressed for the purposes of zoological nomenclature by the ICZN (1963, Opinion 678); originally proposed without included nominal species; Latreille (1817: 261), by including the new species Heleaperforata Latreille, 1817 [as Heleusperforatus] in association with this name, was the first author to subsequently and expressly include nominal species in Helea (ICZN 1999, Article 67.2.2).

Heledona Agassiz, 1846b: 135, 174 [F]. Type species [automatic]: Opatrumagricola Herbst, 1783, by subsequent monotypy (Latreille 1802: 162). Status: junior synonym of Eledona Latreille, 1797 in Tenebrioninae: Bolitophagini. Note: unjustified emendation of Eledona Latreille, 1797, not in prevailing usage.

Helenomelas Ardoin, 1972: 190 [M]. Type species: Helenomelasbasilewskyi Ardoin, 1972, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Helenophorus Gemminger in Gemminger and Harold, 1870: 1851 [M]. Type species [automatic]: Tenebriocollaris Linnaeus, 1767, by monotypy. Status: junior synonym of Leptoderis Billberg, 1820 in Pimeliinae: Elenophorini: Elenophorina. Note: unjustified emendation of Elenophorus Dejean, 1821, not in prevailing usage.

Helibatus Mulsant & Rey, 1859c: 94, 100 [M]. Type species: Helibatusmorio Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Helioarthrodibius Koch, 1960: 353 [M]. Type species: Arthrodeisruguliventris Fairmaire, 1884, by monotypy. Status: valid subgenus of Arthrodibius Lesne, 1915 in Pimeliinae: Erodiini.

Heliocaes Bedel, 1906a: 92 [M]. Type species: Blaps emarginata Fabricius, 1792, by present designation. Status: junior synonym of Heliopates Dejean, 1834 in Blaptinae: Dendarini: Dendarina. Synonymy: Gebien (1938a: 304). Note: this genus was proposed as a replacement name for “Heliopathes” Dejean sensu Mulsant, 1854, an incorrect subsequent spelling of Heliopates Dejean, 1834, which is unavailable (ICZN 1999, Article 33.3); Heliocaes Bedel, 1906 was made available by indication (ICZN 1999, Article 12.2.1); Escalera (1914: 326) was the first author to subsequently and expressly include nominal species in Heliocaes (ICZN 1999, Article 67.2.2).

Heliocrates Reitter, 1904: 98 [M]. Type species: Heliophilushumerangulus Reitter, 1904, by subsequent designation (Iwan and Löbl 2007: 734). Status: valid subgenus of Heliopates Dejean, 1834 in Blaptinae: Dendarini: Dendarina.

Heliodromus Brullé, 1832: 196 [M]. Type species: Heliodromusrotundatus Brullé, 1832, by subsequent designation (Löbl et al. 2008a: 41). Status: junior synonym of Tentyria Latreille, 1802 in Pimeliinae: Tentyriini. Synonymy: Solier (1835b: 314).

Heliofugus Guérin-Méneville, 1831a: pl. 4 [M]. Type species: Heliofugusarenosus Guérin-Méneville, 1831, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Heliomelasma Koch, 1948: 408 [N]. Type species: Melasmanaappenhageni Koch, 1948, by original designation. Status: valid subgenus of Melasmana Strand, 1935 in Blaptinae: Dendarini: Melambiina.

Heliomophlus Reitter, 1906b: 147 [M]. Type species: Heliotaurusscabriusculus Fairmaire, 1866, by subsequent designation (R. Lucas 1920: 319). Status: valid genus in Alleculinae: Cteniopodini.

Heliopates Dejean, 1834: 191 [M]. Type species: Tenebriolusitanicus Herbst, 1797, by subsequent designation (Gebien 1938a: 304). Status: valid genus and subgenus in Blaptinae: Dendarini: Dendarina. Note: this genus was originally proposed as a replacement name for the junior homonym Heliophilus Dejean, 1821 (see Bousquet and Bouchard 2013a: 46) and should have the same type species as Heliophilus Dejean, 1821 (i.e., Pedinushybridus Latreille, 1804); however, recent authors (e.g., Silfverberg 1984: 59, Iwan and Löbl 2008: 279, Iwan and Löbl 2020: 302) have followed the type species designation proposed by Gebien (1938a: 304) and treated Heliopates as a separate genus with Tenebriolusitanicus Herbst, 1797 as its type species; in order to maintain nomenclatural stability we recommend that an application be submitted to the International Commission on Zoological Nomenclature to request that the type species designation proposed by Gebien (1938a: 304) be used for Heliopates Dejean, 1834.

Heliopathes Gebien, 1938a: 304 [M]. Type species: Tenebriolusitanicus Herbst, 1797, by subsequent designation (Gebien 1938a: 304). Status: junior synonym of Heliopates Dejean, 1834 in Blaptinae: Dendarini: Dendarina. Note: unjustified emendation of Heliopates Dejean, 1834, not in prevailing usage; see comments regarding the type species of Heliopates Dejean, 1834 under that name.

Heliophilus Dejean, 1821: 65 [M]. Type species: Pedinushybridus Latreille, 1804, by subsequent designation (Duponchel 1845b: 517). Status: senior synonym of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina. Synonymy: Iwan and Löbl (2008: 282). Note: junior homonym of Heliophilus Meigen, 1803 [Diptera]; the currently accepted valid name of the type species, “Phylan abbreviatus (G.-A. Olivier, 1795)”, should not be considered as an available name since G.-A. Olivier (1795: [57] 17) clearly refers to Tenebrioabbreviatus Fabricius, 1775.

Heliophosis Koch, 1952a: 95 [F]. Type species: Heliophosiskalaharica Koch, 1952, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Heliophygus Agassiz, 1846b: 175 [M]. Type species [automatic]: Heliofugusarenosus Guérin-Méneville, 1831, by monotypy. Status: junior synonym of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Heliofugus Guérin-Méneville, 1831, not in prevailing usage.

Heliosteres Hope, 1841: 124 [M]. Type species [automatic]: Heliofugusarenosus Guérin-Méneville, 1831, by monotypy. Status: junior synonym of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Heliofugus Guérin-Méneville, 1831.

Heliostrhaema Reitter, 1890a: 34 [N]. Type species: Heliotaurusrolphii Fairmaire, 1867, by subsequent designation (R. Lucas 1920: 319). Status: valid genus in Alleculinae: Cteniopodini. Note: Heliostrhaema is an unjustified emendation of the original spelling Heliosthraema, introduced by Seidlitz (1896: 223), in prevailing usage and attributed to the original author and date; it is considered here to be a justified emendation (ICZN 1999, Article 33.2.3.1).

Heliotaurus Mulsant, 1856a: 73 [M]. Type species: Cisteladistincta Laporte, 1840, by subsequent designation (Novák and Pettersson 2008: 332). Status: valid genus and subgenus in Alleculinae: Cteniopodini.

Helogria Borchmann, 1916a: 48, 110 [F]. Type species: Lagriapruinosa Chevrolat, 1841, by subsequent designation (Borchmann 1936: 143). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Helopelius Reitter, 1922b: 123, 152 [M]. Type species: Stenomaxaeneipennis Allard, 1876, by subsequent designation (Nabozhenko 2008: 38). Status: valid subgenus of Stenohelops Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Note: downgraded to a subgenus of Stenohelops Reitter, 1922 by Nabozhenko et al. (2020b: 293).

Helopidesthes Fairmaire, 1895b: 446 [F]. Type species: Helopidesthescoriaria Fairmaire, 1895, by monotypy. Status: valid genus in Tenebrioninae: Helopini: incertae sedis.

Helopidoxus Reitter, 1922a: 32, 44 [M]. Type species: Helopssuperbus Mulsant & Godart, 1855, by monotypy. Status: valid subgenus of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina.

Helopimorphus Desbrochers des Loges, 1881: 140 [M]. Type species: Helopimorphusangulipennis Desbrochers des Loges, 1881, by monotypy. Status: junior synonym of Heterotarsus Latreille, 1829 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Seidlitz (1896: 24).

Helopinus Solier, 1848: 152, 197 [M]. Type species: Helopinuscostatus Solier, 1848, by monotypy. Status: valid subgenus of Drosochrus Erichson, 1843 in Blaptinae: Pedinini: Helopinina. Note: the First Reviser (Helopinus Solier, 1848 versus Pteraulus Solier, 1848) is Koch (1958: 149)

Helopocerodes Reitter, 1922b: 122, 144 [M]. Type species: Helopsfaldermanni Faldermann, 1837, by subsequent designation (Nabozhenko 2001a: 633). Status: junior synonym of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: Keskin et al. (2017: 726).

Helopogonus Reitter, 1922b: 122, 150 [M]. Type species: Helopsviridicollis Schaufuss, 1869, by monotypy. Status: valid subgenus of Nesotes Allard, 1876 in Tenebrioninae: Helopini: Helopina.

Helopondrus Reitter, 1922b: 123, 153 [M]. Type species: Stenomaxsareptanus Allard, 1876, by subsequent designation (Nabozhenko 2001a: 652). Status: junior synonym of Horistelops Gozis, 1910 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: new synonym [YB]. Note: Horistelops Gozis, 1910 has been forgotten in the literature; its type species is currently included in the subgenusHelopondrus Reitter, 1922 and for that reason Reitter’s name is considered a new junior synonym of Horistelops.

Helopostygnus Antoine, 1949: 133, 153 [M]. Type species: Helopsatlantis Antoine, 1926, by original designation. Status: valid subgenus of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina.

Helopotrichus Reitter, 1922a: 32, 44 [M]. Type species: Helopsvillosipennis P.H. Lucas, 1846, by subsequent designation (Viñolas and Cartagena 2005: 38). Status: valid subgenus of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina.

Helops Fabricius, 1775: 257 [M]. Type species: Tenebriocaeruleus Linnaeus, 1758, by subsequent designation (Hope 1841: 133; see ICZN 2009, Opinion 2237). Status: valid genus in Tenebrioninae: Helopini: Helopina. Note: as mentioned in Bousquet et al. (2018: 183) and Vela et al. (2020: 63) the type of the nominal species Tenebriocaeruleus Linnaeus, 1758 is actually a leaf beetle that belongs to the genus Timarcha Samouelle, 1819 (Coleoptera: Chrysomelidae) based on examination of a syntype; since Tenebriocaeruleus Linnaeus, 1758 was placed on the Official List of Specific Names in Zoology and fixed as the type species of Helops Fabricius, 1775 in Opinion 2237 (ICZN 2009), an application to the International Commission on Zoological Nomenclature is necessary to request that the type species of Helops be changed to Tenebriocaeruleus Linnaeus sensu Fabricius, 1775 (= Helopschalibaeus Rossi, 1790).

Helopsallecula Pic, 1936a: 33 [F]. Type species: Helopsalleculaminutissima Pic, 1936, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Helopsisomira Pic, 1952a: 63 [F]. Type species: Helopsisomirakochi Pic, 1952, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Hemasodes Casey, 1907: 378 [M]. Type species: Schoenicusvestitus Champion, 1884, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Hemeralopius Gistel, 1848a: viii, 125 [M]. Type species [automatic]: Tenebrioelongatus Herbst, 1797, by monotypy. Status: senior synonym of Belopus Gebien, 1911 in Lagriinae: Belopini. Note: replacement name for Calcar Dejean, 1821; nomen oblitum (see Bouchard and Bousquet 2020b: 6).

Hemerobates Kolbe, 1884: 189 [M]. Type species: Nyctobatesmechowi Kolbe, 1884, by monotypy. Status: junior synonym of Amenophis J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1911a: 441).

Hemicera Laporte & Brullé, 1831: 332, 393 [F]. Type species: Cnodalonsplendens Wiedemann, 1823, by subsequent designation (Gebien 1921a: 335). Status: valid genus and subgenus in Stenochiinae: Cnodalonini. Note: nomenclatural stability in this genus is threatened by the discovery of an older type species designation (Hemiceraarcuata Laporte & Brullé, 1831 [misspelled “Hemiceraarmata”], by subsequent designation by Duponchel (1845b: 528), which is currently the type species of the valid genus Hypocalis Dejean, 1834); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Gebien (1921a: 335).

Hemicistela Blackburn, 1891: 331 [F]. Type species: Hemicisteladiscoidalis Blackburn, 1891, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Hemicyclus Westwood, 1841a: 44 [M]. Type species: Hemicyclusgrandis Westwood, 1841 (= Tetraphyllusreaumuri Laporte, 1840), by subsequent designation (Gebien 1941: 1133). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Hemimmedia Gebien, 1928: 220, 229 [F]. Type species: Hemimmediacorpulenta Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hemipraocis Flores & Pizarro-Araya, 2014: 66 [M]. Type species: Praocissellatus Berg, 1889, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 13, 60) without type species designation; the fact that Praocissellatus Berg, 1889 was listed as the “type” of Hemipraocis in the Zoological Record for the year 1958 (Anonymous in Commonwealth Institute of Entomology 1960) does not represent a valid type species designation since the nomenclatural act is anonymous (ICZN 1999, Article 14).

Hemipristis Kolbe, 1903: 165, 177 [F]. Type species: Hemipristisukamia Kolbe, 1903, by present designation. Status: senior synonym of Hemipristula Bouchard & Bousquet, nom. nov. in Lagriinae: Pycnocerini. Note: junior homonym of Hemipristis Agassiz, 1833 [Pisces].

Hemipristula Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Hemipristisukamia Kolbe, 1903, by present designation. Status: valid genus in in Lagriinae: Pycnocerini. Note: replacement name Hemipristis Kolbe, 1903; Hemipristula was proposed earlier by Strand (1935a: 291); however, this name is unavailable because Strand did not designate a type species for the nominal taxon, a mandatory requirement for replacement name without valid typification proposed after 1930 (ICZN 1999; Article 13.3.1).

Hemipterocoma Skopin, 1974b: 159 [F]. Type species: Pterocomanikolskii Semenov-Tjan-Shansky, 1910, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Hemitrichestes Reitter, 1904: 169 [M]. Type species: Penthicushirsutus Reitter, 1896, by monotypy. Status: junior synonym of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Löbl et al. (2008b: 267).

Heptaphylla Friedenreich, 1883: 375 [F]. Type species: Heptaphyllafungicola Friedenreich, 1883, by monotypy. Status: junior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: Arrow (1904: 31).

Herbertfranzia Kaszab, 1973a: 26, 28 [F]. Type species: Herbertfranzianepalica Kaszab, 1973, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Herbertfranziella Kaszab, 1973a: 26, 28 [F]. Type species: Herbertfranziaeutagenoides Kaszab, 1973, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Herlesa Reitter, 1897a: 298, 301 [F]. Type species: Herlesaglobicollis Reitter, 1897 (= Micipsacavifrons Fairmaire, 1863), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Herodius Agassiz, 1846b: 143,179 [M]. Type species [automatic]: Erodiusgibbus Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Erodius Fabricius, 1775 in Pimeliinae: Erodiini. Note: unjustified emendation of Erodius Fabricius, 1775, not in prevailing usage.

Herpiscius Solier, 1838b: 160, 188 [M]. Type species: Herpisciusspinolae Solier, 1838, by subsequent designation (Hope 1841: 115). Status: valid genus in Tenebrioninae: Scaurini.

Herpsis Haag-Rutenberg, 1875b: 4, 66 [F]. Type species: Adelostomarugosum Guérin-Méneville, 1831, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Herthasida Wilke, 1922: 269 [F]. Type species: Asidaingens Champion, 1892, by monotypy. Status: valid subgenus of Philolithus Lacordaire, 1858 in Pimeliinae: Asidini.

†Hesiodobates Kaszab & Schawaller, 1984: 1 [M]. Type species: Hesiodobatesantiquus Kaszab & Schawaller, 1984, by original designation. Status: junior synonym of Nesocyrtosoma Marcuzzi, 1976 in Stenochiinae: Cnodalonini. Synonymy: Doyen and Poinar (1994: 45). Note: described from Early Miocene amber (Dominican Republic).

Hesiodus Champion, 1885: 115 [M]. Type species: Hesioduslongitarsis Champion, 1885, by subsequent designation (R. Lucas 1920: 323). Status: valid genus in Stenochiinae: Cnodalonini.

Hesperoptorina G.S. Medvedev, 2009: 417 [F]. Type species: Gnaptorinabrucei Blair, 1923, by original designation. Status: valid subgenus of Gnaptorina Reitter, 1887 in Blaptinae: Blaptini: Gnaptorinina.

Hesseodes Ardoin, 1963a: 92 [M]. Type species: Hoplonyxkalaharica Hesse, 1935, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: the name Hesseodes used earlier by Ardoin (1962b: 969) is unavailable since it was published after 1930 without a type species designation.

Hesseosis Koch, 1958: 76 [F]. Type species: Hesseosispurpurascens Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Heterarthron Gistel, 1848a: 190 [N]. Type species [automatic]: Eulabisbicarinata Eschscholtz, 1829, by subsequent designation (Blaisdell 1932: 44). Status: junior synonym of Eulabis Eschscholtz, 1829 in Tenebrioninae: Eulabini. Note: junior homonym of Heterarthron Dejean, 1836 [Coleoptera: Bostrichidae]; unnecessary replacement name for Eulabis Eschscholtz, 1829 (as “Eulabes”).

Heterasida Casey, 1912: 76, 165 [F]. Type species: Pelecyphorusbifurcus J.L. LeConte, 1861, by original designation. Status: valid genus in Pimeliinae: Asidini.

Heterocheira Dejean, 1836: 220 [F]. Type species: Ulomaaustrale Boisduval, 1835, by monotypy. Status: valid genus in Blaptinae: Opatrini: Heterotarsina.

Heterochira Agassiz, 1846b: 180 [F]. Type species [automatic]: Ulomaaustrale Boisduval, 1835, by monotypy. Status: junior synonym of Heterocheira Dejean, 1836 in Blaptinae: Opatrini: Heterotarsina. Note: unjustified emendation of Heterocheira Dejean, 1836, not in prevailing usage.

Heterogena Froussart, 1961: 60, 105 [F]. Type species: Nesogenagoudotii Fairmaire, 1868, by original designation. Status: valid subgenus of Nesogena Mäklin, 1863 in Tenebrioninae: Praeugenini.

Heterogria Fairmaire, 1896a: 42 [F]. Type species: Heterogriapunctatissima Fairmaire, 1896, by monotypy. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Heteromerotylus Pic, 1921b: 11 [M]. Type species: Heteromerotylusbicoloripes Pic, 1921, by monotypy. Status: junior synonym of Tearchus Kraatz, 1880 in Stenochiinae: Cnodalonini. Synonymy: Kulzer (1954a: 71).

Heteromira Hölzel, 1958: 19 [F]. Type species: Isomiramoroi Hölzel, 1958, by monotypy. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Heteronicandra Koch, 1958: 151 [F]. Type species: Nicandrazumpti Kulzer, 1951, by original designation. Status: valid subgenus of Nicandra Fairmaire, 1888 in Blaptinae: Pedinini: Helopinina.

Heterophaga Dejean, 1834: 199 [F]. Type species: fixed herein (ICZN 1999, Article 70.3) as Opatrumlaevigatum Fabricius, 1781, misidentified as Tenebriomauritanicus Linnaeus, 1764 in the subsequent designation of Duponchel (1845b: 601). Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Wollaston (1854: 498). Note: see Spilman (1972: 32) for the history of the misidentification.

Heterophyllus Gemminger in Gemminger and Harold, 1870: 1955 [M]. Type species [automatic]: Heterophyluschrysomelinus Klug, 1833, by monotypy. Status: junior synonym of Heterophylus Klug, 1833 in Diaperinae: Diaperini: Diaperina. Note: unjustified emendation of Heterophylus Klug, 1833, not in prevailing usage.

Heterophylus Klug, 1833: 91 [M]. Type species: Heterophyluschrysomelinus Klug, 1833, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Heterophylus Mulsant & Rey, 1859c: 7 [M]. Type species: Heliopatespicipes Faldermann, 1837, by subsequent designation (Gebien 1938a: 397). Status: senior synonym of Dissonomus Jacquelin du Val, 1861 in Tenebrioninae: Dissonomini. Note: junior homonym of Heterophylus Klug, 1833 [Tenebrionidae: Diaperinae: Diaperini: Diaperina].

Heteropromus Blaisdell, 1909: 179 [M]. Type species: Eleodesvetorator Horn, 1874, by monotypy. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Heteropsectropus Kaszab, 1941c: 33, 34 [M]. Type species: Heteropsectropusaenescens Kaszab, 1941, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Heteropus Laporte, 1840: 221 [M]. Type species: Heteropusholosericeus Laporte, 1840, by monotypy. Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Lacordaire (1859a: 250). Note: junior homonym of Heteropus Palisot de Beauvois, 1820 [Mammalia].

Heteroscelis Agassiz, 1846b: 181, 266 [F]. Type species [automatic]: Adesmiapulcherrima Solier, 1835 (= Adesmiaaudouini Solier, 1835), by subsequent designation (Hope 1841: 118). Status: junior synonym of Oteroscelis Solier, 1835 in Pimeliinae: Adesmiini. Note: unjustified emendation of Oteroscelis Solier, 1835, not in prevailing usage; junior homonym of Heteroscelis Latreille, 1828 [Coleoptera: Tenebrionidae: Blaptinae: Platynotini: Platynotina].

Heteroscelis Latreille, 1828: 574 [F]. Type species: Blapsdentipes Fabricius, 1794, by subsequent designation (Blanchard 1844: pl. 48). Status: senior synonym of Anomalipus Guérin-Méneville, 1831 in Blaptinae: Platynotini: Platynotina. Synonymy: Lacordaire (1859a: 257). Note: nomen oblitum (see Bouchard and Bousquet 2020a: 101); this genus used to be credited to Latreille (1829a: 18) and treated as a junior homonym of Heteroscelis Latreille, 1829 [Hemiptera] until bibliographic research determined that the tenebrionid name was made available earlier by Latreille (1828: 574) and is in fact the senior homonym (see Bouchard and Bousquet 2020a: 101).

Heterostrongylium Kaszab, 1977b: 10, 27 [N]. Type species: Strongyliumweiskei Gebien, 1921, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Heterotarsus Latreille, 1829a: 26 [M]. Type species: Heterotarsustenebrioides Guérin-Méneville, 1831, by subsequent monotypy (Guérin-Méneville 1831b: pl. 30). Status: valid genus in Blaptinae: Opatrini: Heterotarsina. Note: originally proposed without included nominal species; Guérin-Méneville (1831b: pl. 30), by including the new species Heterotarsustenebrioides Guérin-Méneville, 1831 in association with this name, was the first author to subsequently and expressly include nominal species in Heterotarsus (ICZN 1999, Article 67.2.2).

Hexagonochilus Solier, 1851: 168 [M]. Type species: Hexagonochilusdilaticollis Solier, 1851, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina. Note: Hexagonochilus is an incorrect subsequent spelling of the original spelling Hexagonocheilus, first used by Imhoff (1856: 234), and is in prevailing usage; Hexagonochilus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Hexarhopalus Fairmaire, 1891c: xix [M]. Type species: Hexarhopalussculpticollis Fairmaire, 1891, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Hexarhoptrum Fairmaire, 1894a: 38 [N]. Type species [automatic]: Hexarhopalussculpticollis Fairmaire, 1891, by monotypy. Status: junior synonym of Hexarhopalus Fairmaire, 1891 in Stenochiinae: Cnodalonini. Note: the alternative original spelling Hexaroptrum, used by Fairmaire (1894a: 38), was rejected by Bečvář and Purchart (2008: 39) who acted as the First Revisers; unnecessary replacement name for Hexarhopalus Fairmaire, 1891.

Hicetaon Champion, 1885: 111 [M]. Type species: Hicetaonfrontalis Champion, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hidrosella Koch, 1952b: 32 [F]. Type species: Hidrosisincostata Haag-Rutenberg, 1875, by original designation. Status: valid subgenus of Machlopsis Pomel, 1871 in Pimeliinae: Adelostomini.

Hidrosis Haag-Rutenberg, 1875a: 120 [F]. Type species: Steiracrenatocostata Redtenbacher, 1868, by subsequent designation (Löbl et al. 2008b: 121). Status: junior synonym of Machlopsis Pomel, 1871 in Pimeliinae: Adelostomini. Synonymy: Bedel (1887: 199).

Himastethe Koch, 1950b: 286 [F]. Type species: Cyphostethegigantea Koch, 1950, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini.

Himatismus Erichson, 1843: 253 [M]. Type species: Himatismusmandibularis Erichson, 1843, by subsequent designation (Gebien 1937a: 574). Status: valid subgenus of Imatismus Dejean, 1834 in Pimeliinae: Tentyriini.

Hionthis Miller, 1861: 174 [F]. Type species: Hionthistentyrioides Miller, 1861, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Hionthisoma Gebien, 1937a: 609 [N]. Type species [automatic]: Hionthisoccidentalis Fairmaire, 1897, by monotypy. Status: junior synonym of Hyonthosoma Reitter, 1900 in Pimeliinae: Tentyriini. Note: unjustified emendation of Hyonthosoma Reitter, 1900, not in prevailing usage.

Hipalmus Bates, 1870: 269 [M]. Type species: Tenebriocostatus Guérin-Méneville, 1831, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini. Note: the original combination of the name of the type species, Tenebriocostatus Guérin-Méneville, 1831, is a junior primary homonym of Tenebriocostatus Pallas, 1781.

Hipomelus Dejean, 1834: 181 [M]. Type species: Sepidium vittatum Fabricius, 1781, by subsequent designation (Hope 1841: 116). Status: junior synonym of Trachynotus Latreille, 1828 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Bousquet and Bouchard (2013a: 47).

Hipponome Laporte, 1840: 235 [F]. Type species: Helopsazureus Brullé, 1832, by monotypy. Status: senior synonym of Raiboscelis Allard, 1876 in Tenebrioninae: Helopini: Helopina. Synonymy: Seidlitz (1895: 754). Note: nomen oblitum (see Nabozhenko and Löbl 2009: 194).Note: the original combination of the name of the type species, Helopsazureus Brullé, 1832, is a junior primary homonym of Helopsazureus Germar, 1823.

Hirsutosora Pic, 1934a: 32 [F]. Type species: Nemostirafortithorax Pic, 1922, by monotypy. Status: valid subgenus of Sora Walker, 1859 in Lagriinae: Lagriini: Statirina.

Hirtograbies Koch, 1954a: 23 [M]. Type species: Hirtograbiesoograbiesensis Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Hispanomelia Mas-Peinado, Buckley, Ruiz & García-París, 2018: 541 [F]. Type species: Pimeliamanchega Lauffer, 1905, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Histeromimus Gahan, 1895: 288 [M]. Type species: Histeromimusarabicus Gahan, 1895, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Histeromorphus Kraatz, 1865: 6, 11 [M]. Type species: Histeromorphusplicatus Kraatz, 1865, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Histeropsis Chevrolat, 1878a: 221 [F]. Type species: Platydemaamericana Laporte & Brullé, 1831, by subsequent designation (Löbl et al. 2008a: 42). Status: junior synonym of Platydema Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Synonymy: Champion (1886: 181).

Histiaea Fairmaire, 1892a: 107 [F]. Type species: Histiaeabidentula Fairmaire, 1892, by monotypy. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini.

Histrionotus Koch, 1955a: 44 [M]. Type species: Trachynotuslightfooti Péringuey, 1892, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Holaniara Fairmaire, 1871b: 43 [F]. Type species [automatic]: Ulomapiceum Melsheimer, 1846, by monotypy. Status: junior synonym of Eutochia J.L. LeConte, 1862 in Tenebrioninae: Ulomini. Note: replacement name for Aniara Melsheimer, 1853.

Holdhausia Reitter, 1906b: 125 [F]. Type species: Cteniopuscrassus Fairmaire, 1892, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Holeleodes Blaisdell, 1937: 132 [M]. Type species: Eleodesbeameri Blaisdell, 1937 (= Eleodeshepburni Champion, 1884), by original designation. Status: junior synonym of Steneleodes Blaisdell, 1909 in Blaptinae: Amphidorini. Synonymy: Johnston (2015: 12).

Holoblaps Bauer, 1921: 233 [F]. Type species: none designated. Status: undetermined taxon in Blaptinae: Blaptini: Blaptina. Note: this genus was described before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Holoblaps and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Holobrachium Gebien, 1905: 256 [N]. Type species [automatic]: Hoplobrachiumasperipenne Fairmaire, 1886 (= Helopsdentipes Fabricius, 1781), by monotypy. Status: junior synonym of Hoplobrachium Fairmaire, 1886 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Hoplobrachium Fairmaire, 1886, not in prevailing usage.

Holobrachys Fairmaire, 1869b: 233 [M]. Type species: Holobrachysheterocerus Fairmaire, 1869, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hologenosis Deyrolle, 1867: 81, 82 [F]. Type species: Hologenosislacerata Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Holostrongylium Kaszab, 1977b: 10, 20 [N]. Type species: Strongyliumgravidum Mäklin, 1867, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Homala Eschscholtz, 1831: 5, 6 [F]. Type species: Homalapolita Eschscholtz, 1831, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Homalapipleurus Español, 1957a: 166, 167 [M]. Type species: Hegetergonzalezi Español, 1957, by original designation. Status: valid subgenus of Hegeter Latreille, 1802 in Pimeliinae: Tentyriini.

Homaleis Rye, 1879: 62 [M]. Type species [automatic]: Helopscongener Reiche, 1861, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Note: unjustified emendation of Omaleis Allard, 1877, not in prevailing usage.

Homalinota Koch, 1950a: 66 [F]. Type species [automatic]: Homalaagona Fairmaire, 1884, by subsequent designation (Gebien 1937a: 610). Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Homalopsis Lesne, 1922.

Homalopsis Lesne, 1922: 675 [F]. Type species: Homalaagona Fairmaire, 1884, by subsequent designation (Gebien 1937a: 610). Status: senior synonym of Homalinota Koch, 1950 in Pimeliinae: Tentyriini. Note: junior homonym of Homalopsis Kuhl & Hasselt, 1822 [Reptilia].

Homalus Rye, 1878: 69 [M]. Type species [automatic]: Helopscongener Reiche, 1861, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Note: unjustified emendation of Omaleis Allard, 1876 (as “Omalus”); junior homonym of Homalus Agassiz, 1846 [Hymenoptera].

Homebius Endrödy-Younga, 1989: 124 [M]. Type species: Homebiuskaszabi Endrödy-Younga, 1989, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Homebiina.

Homocyrtus Dejean, 1834: 211 [M]. Type species [automatic]: Cyphonotusdromedarius Guérin-Méneville, 1831, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: replacement name for Cyphonotus Guérin-Méneville, 1831; based on a recent assessment of adult and larval morphological characters, Matthews and Lawrence (2015: 290) determined that this genus could not be placed in any known tribe and that the status of Homocyrtus should be regarded as Tenebrionidae incertae sedis.

Homoeocamaria Blair, 1919b: 75 [F]. Type species [automatic]: Homoeogenuslaticornis C.O. Waterhouse, 1882, by monotypy. Status: junior synonym of Borneocamaria Pic, 1917 in Stenochiinae: Cnodalonini. Note: replacement name for Homoeogenus C.O. Waterhouse, 1882.

Homoeogenus C.O. Waterhouse, 1882a: 174 [M]. Type species: Homoeogenuslaticornis C.O. Waterhouse, 1882, by monotypy. Status: senior synonym of Borneocamaria Pic, 1917 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1942a: 323). Note: junior homonym of Homoeogenus C.O. Waterhouse, 1880 [Coleoptera: Psephenidae].

Homoeonota Fairmaire in Fairmaire et al., 1882: 63 [F]. Type species: Homoeonotasubopaca Fairmaire, 1882, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Homopsis Semenov, 1893: 258, 263 [F]. Type species: Homopsisgrumi Semenov, 1893, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Homoropsis Fairmaire, 1886a: 450 [F]. Type species: Homoropsisustulata Fairmaire, 1886, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Homotrysis Pascoe, 1866a: 489 [F]. Type species: Alleculatristis Germar, 1848 (= Alleculacarbonaria Germar, 1848), by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: the First Reviser (Homotrysis Pascoe, 1866 versus Hybrenia Pascoe, 1866) is Champion (1897: 158).

Hopatrinus Agassiz, 1846b: 185, 260 [M]. Type species [automatic]: Opatrumclathratum Fabricius, 1787, by monotypy. Status: junior synonym of Opatrinus Dejean, 1821 in Blaptinae: Platynotini: Platynotina. Note: unjustified emendation of Opatrinus Dejean, 1821, not in prevailing usage.

Hopatroides Agassiz, 1846b: 185, 260 [M]. Type species [automatic]: Opatroidespunctulatus Brullé, 1832, by monotypy. Status: junior synonym of Opatroides Brullé, 1832 in Blaptinae: Opatrini: Opatrina. Note: unjustified emendation of Opatroides Brullé, 1832, not in prevailing usage.

Hopatromorpha Blackburn, 1907: 286 [F]. Type species [automatic]: Opatrummurinum Baudi di Selve, 1876 (= Opatrinussetosus Mulsant & Rey, 1853), by subsequent designation (Gebien 1938a: 399). Status: junior synonym of Mesomorphus Miedel, 1880 in Blaptinae: Opatrini: Opatrina. Note: unnecessary replacement name for Mesomorphus Miedel, 1880 (incorrectly attributed to “Reitter, 1904”).

Hopatropteron Reitter, 1889a: 701 [N]. Type species: Hopatropteronsubcostatum Reitter, 1889 (= Heterotarsuscarinula Marseul, 1876), by monotypy. Status: junior synonym of Heterotarsus Latreille, 1829 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Seidlitz (1894: 413), Gebien (1911a: 473).

Hopatrum Agassiz, 1846b: 185, 260 [N]. Type species [automatic]: Silphasabulosa Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Opatrum Fabricius, 1775 in Blaptinae: Opatrini: Opatrina. Note: unjustified emendation of Opatrum Fabricius, 1775, not in prevailing usage.

Hoplambius Reitter, 1904: 114 [M]. Type species: Hoplarionmelambioides (as “melambioide”) Fairmaire, 1893, by monotypy. Status: valid subgenus of Melambius Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Hoplariobius Reitter, 1904: 115 [M]. Type species: Micrositusdecurtatus Fairmaire, 1884, by subsequent designation (Antoine 1942: 25). Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Hoplarion Mulsant & Rey, 1854: 150 [N]. Type species: Micrositustumidus Mulsant & Rey, 1854, by monotypy. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Hoplaspis Motschulsky, 1858a: 113 [F]. Type species: Hoplaspislamellicornis Motschulsky, 1858, by subsequent designation (Crotch 1876: 418). Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: the alternative original spelling Hoplapsis, used by Motschulsky (1858: 113), was rejected by Motschulsky (1868: 69) who acted as the First Reviser (ICZN 1999, Article 24.2.4); transferred from Cucujoidea: Erotylidae by Arrow (1909: 196).

Hoplitoblaps Fairmaire, 1889b: 26 [F]. Type species: Hoplitoblapsfallaciosa Fairmaire, 1889, by monotypy. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Hoplobrachium Fairmaire, 1886c: 74 [N]. Type species: Hoplobrachiumasperipenne Fairmaire, 1886 (= Helopsdentipes Fabricius, 1781), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Hoplocephala Agassiz, 1846b: 185, 262 [F]. Type species [automatic]: Ips haemorrhoidalis Fabricius, 1787, by subsequent designation (Motschulsky 1845a: 80). Status: junior synonym of Neomida Latreille, 1829 in Diaperinae: Diaperini: Diaperina. Note: unjustified emendation of Oplocephala Laporte & Brullé, 1831, not in prevailing usage.

Hoplochirus Scudder, 1882: 153 [M]. Type species [automatic]: Hoplonyxalleculoides J. Thomson, 1858, by subsequent designation (Gebien 1943: 919). Status: junior synonym of Hoplonyx J. Thomson, 1858 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Oplocheirus Klug, 1835, not in prevailing usage.

Hoploedipinus Kaszab, 1984: 355, 356 [M]. Type species: Hoploedipusheterodoxus Fairmaire, 1898, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Hoploedipus Fairmaire, 1898d: 395 [M]. Type species: Camarimenaarmipes Fairmaire, 1882, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Hoplonyx J. Thomson, 1858: 98 [M]. Type species: Hoplonyxalleculoides J. Thomson, 1858, by subsequent designation (Gebien 1943: 919). Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Hoplopeltis Fairmaire, 1894a: 22 [F]. Type species: Hoplopeltistricornis Fairmaire, 1894, by monotypy. Status: valid genus in Tenebrioninae: Alphitobiini.

Hoploptera Gemminger in Gemminger and Harold, 1870: 2037 [F]. Type species [automatic]: Strongyliumserraticorne Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Oploptera Chevrolat, 1844 in Stenochiinae: Stenochiini. Note: unjustified emendation of Oploptera Chevrolat, 1844, not in prevailing usage.

Hoplostira Borchmann, 1921: 217, 225 [F]. Type species: Hoplostirafemoralis Borchmann, 1921, by original designation. Status: junior synonym of Spinostatira Pic, 1918 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1936: 247).

Hoplostrongylium Ardoin, 1965c: 1326 [N]. Type species: Hoplostrongyliumghesquierei Ardoin, 1965, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Horatoma Solier, 1841a: 210, 264 [364] [F]. Type species: Horatomaparvula Solier, 1841, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Horatomella Penrith & Endrödy-Younga, 1994: 6, 7 [F]. Type species: Parapachynotelajohni Koch, 1957, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Horatomodes Haag-Rutenberg, 1872: 274, 305 [M]. Type species: Horatomodesbatesi Haag-Rutenberg, 1872, by monotypy. Status: junior synonym of Horatoma Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Synonymy: Penrith and Endrödy-Younga (1994: 10).

Horistelops Gozis, 1910: 112 [M]. Type species: Helopsassimilis Küster, 1850, by original designation. Status: valid subgenus of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina.

Hosohamudama Masumoto, 1988b: 43 [F]. Type species: Hosohamudamasasajii Masumoto, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Houaphanica Novák, 2020b: 470 [F]. Type species: Houaphanicafera Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Hovacula Fairmaire, 1898a: 236 [F]. Type species: Hovaculalineolata Fairmaire, 1898, by monotypy. Status: valid genus in Alleculinae: incertae sedis. Note: placed in Alleculinae by Chatanay (1915a: 526).

Hovadelium Ardoin, 1961c: 34 [N]. Type species: Hovadeliumdiscoidale Ardoin, 1961, by original designation. Status: valid genus in Lagriinae: Laenini.

Hovademulus Iwan, 1996: 385, 390 [M]. Type species: Selinuspunctipennis Fairmaire, 1902, by original designation. Status: junior synonym of Clastopus Fairmaire, 1898 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (2001: 500).

Hovademus Iwan, 1996: 385, 395 [M]. Type species: Hovademusandringitrensis Ardoin, 1974, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina. Note: the name Hovademus was proposed earlier by Ardoin (1974a: 165) without a type species designation; the fact that Hovademusandringitrensis Ardoin, 1974 was listed as the “type” of Hovademus in the Zoological Record for the year 1974 (Anonymous in Staff of the Zoological Society of London 1979) does not represent a valid type species designation since the nomenclatural act is anonymous (ICZN 1999, Article 14).

Hovarygmus Fairmaire, 1898a: 234 [M]. Type species: Hovarygmusinsularis Fairmaire, 1898, by monotypy. Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Huilamus Koch, 1953d: 79 [M]. Type species: Huilamuswelwitschi Koch, 1953, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Hummelinckia Marcuzzi, 1954: 19 [F]. Type species: Hummelinckiacaraibica Marcuzzi, 1954, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Hyalarthrodosis Kaszab, 1979a: 74 [F]. Type species: Arthrodosismonodi Pierre, 1974, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Hyalerodius Kaszab, 1979a: 80 [M]. Type species: Hyalerodiusjirofti Kaszab, 1979, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Hybocaulus Fairmaire, 1895a: 27 [M]. Type species: Hybocauluslaticornis Fairmaire, 1895 (= Porphyrhybaviolaceicolor Fairmaire, 1877), by monotypy. Status: junior synonym of Porphyrhyba Fairmaire, 1877 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1956b: 89).

Hybonotus Dejean, 1834: 211 [M]. Type species: Tetraphyllusformosus Laporte & Brullé, 1831, by monotypy. Status: senior synonym of Damatris Laporte, 1840 in Stenochiinae: Cnodalonini. Synonymy: new synonym [PB]. Note: the type species of Hybonotus Dejean, 1834 is the same as the type species of Damatris Laporte, 1840 and therefore the two genera are objective synonyms; junior homonym of Hybonotus Klug, 1803 [Hymenoptera].

Hyboproctus Kolbe, 1897a: 241 [M]. Type species: Hyboproctusnodifer Kolbe, 1897, by subsequent designation (Gebien 1940: 1089). Status: valid genus in Stenochiinae: Cnodalonini.

Hybrenia Pascoe, 1866a: 489 [F]. Type species: Hybreniavittata Pascoe, 1866, by subsequent designation (Matthews and Bouchard 2008: 328). Status: junior synonym of Homotrysis Pascoe, 1866 in Alleculinae: Alleculini: Alleculina. Synonymy: Champion (1897: 158).

Hydissus Pascoe, 1869: 148 [M]. Type species: Nyctobatesferonioides Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hydisus Scudder, 1882: 167 [M]. Type species [automatic]: Nyctobatesferonioides Pascoe, 1866, by monotypy. Status: junior synonym of Hydissus Pascoe, 1869 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Hydissus Pascoe, 1869, not in prevailing usage.

Hylithus Guérin-Méneville, 1834: 12 [M]. Type species: Hylithustentyrioides Guérin-Méneville, 1834, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Hylocrinus Casey, 1907: 289, 331 [M]. Type species: Eurymetoponlungulum J.L. LeConte, 1851, by original designation. Status: valid genus and subgenus in Pimeliinae: Edrotini.

Hylonoma Macquart, 1850: 183 [F]. Type species: Alegoriadilatata Laporte, 1840, by monotypy. Status: junior synonym of Alegoria Laporte, 1840 in Tenebrioninae: Ulomini. Synonymy: Lacordaire (1859a: 325).

Hyloplonyx Ardoin, 1963c: 716 [M]. Type species: Hoplonyxmonophthalmus J. Thomson, 1858, by original designation. Status: valid subgenus of Hoplonyx J. Thomson, 1858 in Tenebrioninae: Amarygmini.

Hymenalia Mulsant, 1856a: 48 [F]. Type species: Cistelafusca Illiger, 1794 (= Cistelarufipes Fabricius, 1792), by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Hymenochara Campbell, 1978: 435 [F]. Type species: Mycetophilarufipes J.E. LeConte, 1824, by original designation. Status: valid genus in Alleculinae: Alleculini: Mycetocharina.

Hymenorus Mulsant, 1852: 68 [M]. Type species: Hymenorusdoublieri Mulsant, 1852, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: the alternative original spelling Hymenophorus was corrected to Hymenorus in the “Emendanda” of the same work (p. 188), Hymenorus is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1); see Bousquet et al. (2015: 133).

Hyocis Pascoe, 1866a: 457 [M]. Type species: Hyocisbakewellii Pascoe, 1866, by original designation. Status: valid genus and subgenus in Diaperinae: Hyociini: Hyociina.

Hyonthosoma Reitter, 1900c: 89, 142 [N]. Type species: Hionthisoccidentalis Fairmaire, 1897, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Hypamarygmus Gebien, 1904b: 27 [M]. Type species: Hypamarygmuscoccinelloides Gebien, 1904, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Hypaulax Bates, 1868: 259 [F]. Type species: Hypaulaxmarginata Bates, 1868, by subsequent designation (Gebien 1941: 333). Status: valid genus in Stenochiinae: Cnodalonini. Note: the First Reviser (Hypaulax Bates, 1868 versus Chileone Bates, 1868) is Carter (1914b: 46).

Hyperamarygmus Kaszab, 1964a: 291 [M]. Type species: Hyperamarygmusantennalis Kaszab, 1964, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Hyperchalca Fairmaire, 1869b: 238 [F]. Type species: Hyperchalcaaenescens Fairmaire, 1869, by monotypy. Status: valid genus and subgenus in Stenochiinae: Stenochiini.

Hypercossyphodes Andreae, 1961: 205, 215 [M]. Type species: Cossyphodesvandami Andreae, 1961, by original designation. Status: junior synonym of Cossyphodes Westwood, 1851 in Pimeliinae: Cossyphodini: Cossyphodina. Synonymy: Schawaller (2013c: 362, implied by inclusion of Cossyphodesvandami Andreae, 1961 in Cossyphodes Westwood, 1851 without use of a subgenus rank).

Hypermicrotelopsis Koch, 1940b: 743 [F]. Type species: Microtelopsisthibetana Koch, 1940, by monotypy. Status: valid subgenus of Microtelopsis Koch, 1940 in Pimeliinae: Stenosini: Stenosina. Note: combined description of new genus-group taxon and a single new species (ICZN 1999, Article 13.4); we act as First Revisers and select Microtelopsis Koch, 1940 as the valid name for this genus instead of Extetranosis Koch, 1940 and Hypermicrotelopsis Koch, 1940.

Hyperops Eschscholtz, 1831: 5, 9 [M]. Type species: Hyperopstagenioides Eschscholtz, 1831, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Hyperopsis Bouchard & Bousquet, new subgenus [F]. Type species: Pachyceracolasi Koch, 1943, by present designation. Status: valid subgenus of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: Koch (1943a: 524, 536) introduced the new subgenus name Hyperopsis for five nominal species, but unfortunately did not designate a type species; the subgenusHyperopsis, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Pachyceracolasi Koch, 1943 as type species and referring to Koch (1943a: 524) for the character states that characterise and differentiate Hyperopsis.

Hypoblaps Kolbe, 1928: 200 [F]. Type species: Blapsrotundata Solier, 1848, by original designation. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Hypocalis Dejean, 1834: 206 [F]. Type species: Hemiceraarcuata Laporte & Brullé, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hypocilibe Bates, 1872b: 275 [F]. Type species: Hypocilibemacleayi Bates, 1872, by monotypy. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1911b: 139, with Onosterrhus Pascoe, 1866, a junior synonym of Nyctozoilus Guérin-Méneville, 1831).

Hypocistela Bates, 1879b: 482 [F]. Type species: Hypocistelatenuipes Bates, 1879, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Hypogena Dejean, 1834: 199 [F]. Type species: Tenebriobiimpressus Latreille, 1833 (= Peltisbrasilica Perty, 1830), by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Hypolaenopsis Masumoto, 2001: 45 [F]. Type species: Hypolaenopsisuenoi Masumoto, 2001 (= Laenananpingica Schawaller, 2001), by original designation. Status: valid genus in Lagriinae: Laenini.

Hypomelus Solier, 1843: 4, 93, 126 [M]. Type species: Hypomelusbicolor Solier, 1843 (= Helopsperonatus Germar, 1823), by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Hypophlaeus Fabricius, 1790: 222 [M]. Type species: Hypophlaeuscastaneus Fabricius, 1790 (= Corticeusunicolor Piller & Mitterpacher, 1783), by subsequent designation (Curtis 1832: pl. 430). Status: junior synonym of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini. Synonymy: Crotch (1870a: 47).

Hypoprosodes Reitter, 1909a: 122 [M]. Type species: Prosodesminuta Kraatz, 1881, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Hypostatira Fairmaire, 1889c: xlix [F]. Type species: Hypostatiravariicolor Fairmaire, 1889, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Hypovinsonia Ardoin, 1961a: 207 [F]. Type species: Hypovinsoniaalbopilosa Ardoin, 1961, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Hypselops Solier, 1851: 135 [M]. Type species: Hypselopsoblongus Solier, 1851, by subsequent designation (R. Lucas 1920: 347). Status: valid genus in Pimeliinae: Epitragini.

Hypsosoma Ménétriés, 1854: 30 [N]. Type species: Hypsosomamongolicum Ménétriés, 1854, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Hypulus Rafinesque, 1815: 114 [M]. Type species [automatic]: Tenebriocaeruleus Linnaeus, 1758, by subsequent designation (Hope 1841: 133; see ICZN 2009, Opinion 2237). Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Note: unnecessary replacement name for Helops Fabricius, 1775; junior homonym of Hypulus Paykull, 1798 [Coleoptera: Melandryidae]; see entry for Helops Fabricius, 1775 for information regarding the identity of the type species.

Hysterarthron J. Thomson, 1864: 224 [N]. Type species: Hysterarthroncollare J. Thomson, 1864, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Iberomelia Mas-Peinado, Buckley, Ruiz & García-París, 2018: 539 [F]. Type species: Pimeliacastellana Pérez Arcas, 1865, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Ibnsaudia Koch, 1941: 290 [F]. Type species: Ibnsaudiapropheta Koch, 1941, by monotypy. Status: junior synonym of Thraustocolus Kraatz, 1866 in Pimeliinae: Tentyriini. Synonymy: Kaszab (1981b: 341).

Iccius Champion, 1886: 147 [M]. Type species: Icciuscephalotes Champion, 1886, by subsequent designation (Gebien 1940: 760). Status: valid genus in Diaperinae: Diaperini: Adelinina.

Idahelops Keskin & Nabozhenko, 2012: 63 [M]. Type species: Idahelopsalpagutae Keskin & Nabozhenko, 2012, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Idastrandiella Strand, 1929: 23 [F]. Type species [automatic]: Helopsmucoreus Waltl, 1838, by monotypy. Status: junior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Note: replacement name for Apolites Jacquelin du Val, 1861.

Idatius Fairmaire, 1906: 276 [M]. Type species: Idatiusophtalmicus Fairmaire, 1906, by monotypy. Status: valid genus in Alleculinae: incertae sedis. Note: placed in Alleculinae by Chatanay (1915a: 526).

Idiesa Reitter, 1893: 203, 245 [F]. Type species: Diesiafischeri Ménétriés, 1849, by subsequent designation (Gebien 1937a: 807). Status: valid genus in Pimeliinae: Pimeliini.

Idiobates Casey, 1891: 62 [M]. Type species: Tenebriocastaneus Knoch, 1801, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Idiopsis Kaszab, 1981a: 78 [F]. Type species: Idiopsisopaca Kaszab, 1981, by original designation. Status: senior synonym of Kocakia Kaszab, 1985 in Pimeliinae: Edrotini. Note: junior homonym of Idiopsis Brauer & Bergenstamm, 1890 [Diptera].

Idisia Pascoe, 1866a: 452 [F]. Type species: Idisiaornata Pascoe, 1866, by monotypy. Status: valid genus in Pimeliinae: Idisiini.

Idricus Fairmaire, 1888c: 199 [M]. Type species: Idricusdiabolicus Fairmaire, 1888, by monotypy. Status: junior synonym of Ametrocera Fåhraeus, 1870 in Blaptinae: Pedinini: Helopinina. Synonymy: Péringuey (1904: 296).

Iliodera Skopin, 1961a: 396 [F]. Type species: Microderadesertoides Skopin, 1961, by original designation. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Ilus Champion, 1885: 117 [M]. Type species: Ilusapicicornis Champion, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ilyxerus Pascoe, 1866a: 458 [M]. Type species: Ilyxerusasper Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Imatismus Dejean, 1834: 202 [M]. Type species: Helopsfasciculatus Fabricius, 1798, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Immedia Pascoe, 1882: 33 [F]. Type species: Immediaocculta Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Impressallecula Pic, 1951: 12 [F]. Type species: Impressalleculapurpureipes Pic, 1951, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Impressosora Pic, 1952b: 254 [F]. Type species: Impressosoranotaticollis Pic, 1952, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina. Note: new placement [OM], previously included in Lagriinae: Lagriini: Lagriina.

Indenicmosoma Ardoin, 1964b: 688, 689 [N]. Type species: Enicmosomaindochinense Kaszab, 1940, by original designation. Status: valid genus in Lagriinae: Lupropini.

Indeucolus Kaszab, 1975b: 280, 282 [M]. Type species: Indeucoluscostatus Kaszab, 1975, by original designation. Status: junior synonym of Eucolus Mulsant & Rey, 1853 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (1997: 258).

Indianosis Koch, 1941: 297 [F]. Type species: Stenosiscapitata Koch, 1941, by monotypy. Status: valid subgenus of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina.

Indochillus Koch, 1941: 300 [M]. Type species: Indochilluscristatus Koch, 1941, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Indoprosodes G.S. Medvedev, 2003: 690 [M]. Type species: Prosodesboorpi Kaszab, 1956, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Indostola G.S. Medvedev, 1991: 557 [F]. Type species: Indostolapulchella G.S. Medvedev, 1991, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Indricula Novák, 2016c: 47 [F]. Type species: Indriculaargynnis Novák, 2016, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Inscutoheliofugus Freude, 1960a: 126, 130 [M]. Type species: Heliofuguskuscheli Freude, 1960, by original designation. Status: valid subgenus of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini.

Insolitoplonyx Bremer, 2014b: 178, 181 [M]. Type species: Insolitoplonyxseorsus Bremer, 2014, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Inspinogeton Pic, 1937a: 174 [M]. Type species: Cyriogetonimpressipennis Pic, 1937, by monotypy. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Insulasida Escalera, 1922b: 64 [F]. Type species: Asidamoraguezi Schaufuss, 1879, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 291).

Iphicorynus Jacquelin du Val, 1861: 299 [M]. Type species: Pentaphyllusmelanophthalmus Mulsant, 1854 (= Nitidulachrysomeloides Rossi, 1792), by monotypy. Status: junior synonym of Pentaphyllus Dejean, 1821 in Diaperinae: Diaperini: Diaperina. Synonymy: Gemminger in Gemminger and Harold (1870: 1956).

Iphius Dejean, 1834: 203 [M]. Type species: Tenebrioserratus Fabricius, 1775, by monotypy. Status: senior synonym of Prioscelis Hope, 1841 in Lagriinae: Pycnocerini. Synonymy: Imhoff (1856: 238). Note: junior homonym of Iphius Schönherr, 1823 [Coleoptera: Curculionidae].

Iphthimera Reitter, 1916a: 4 [F]. Type species: Stenocararuficorne Solier, 1835, by present designation. Status: junior synonym of Metriopus Solier, 1835 in Pimeliinae: Adesmiini. Synonymy: new synonym [PB]. Note: the type species of Iphthimera Reitter, 1916 is currently included in the genus Metriopus Solier, 1835 and for that reason Iphthimera is considered a junior synonym of Solier’s name.

Iphthiminus Spilman, 1973: 42 [M]. Type species: Iphthinusitalicus Truqui, 1857, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Iphthimulus Reitter, 1920a: 16, 17 [M]. Type species: Iphthinustruquii Marseul, 1869, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Iphthimus Gemminger in Gemminger and Harold, 1870: 1977 [M]. Type species [automatic]: Tenebriogigas Linnaeus, 1763, by subsequent designation (Spilman 1973: 42). Status: junior synonym of Mylaris Pallas, 1781 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Iphthinus Dejean, 1834, not in prevailing usage.

Iphthinus Dejean, 1834: 203 [M]. Type species: Tenebriogigas Linnaeus, 1763, by subsequent designation (Spilman 1973: 42). Status: junior synonym of Mylaris Pallas, 1781 in Stenochiinae: Cnodalonini. Synonymy: Chevrolat (1845b: 106, with Nyctobates Guérin-Méneville, 1834, a junior synonym of Mylaris Pallas, 1781), Spilman (1973: 42).

Iranarthrodosis Kaszab, 1959a: 334 [F]. Type species: Arthrodosispfaundleri Schuster, 1935, by original designation. Status: junior synonym of Erodiontes Reitter, 1914 in Pimeliinae: Erodiini. Synonymy: Kaszab (1979a: 88).

Iranerodius Kaszab, 1959a: 334 [M]. Type species: Arthrodosisrichteri Kaszab, 1957, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Iranolasiostola Pierre, 1968: 1020 [F]. Type species: Iranolasiostoladavatchii Pierre, 1968, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Iranopachyscelis Pierre, 1968: 1027 [F]. Type species: Iranopachysceliseghbali Pierre, 1968 (= Thripterapersica Redtenbacher, 1850), by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Iranosodes G.S. Medvedev, 1996: 605 [M]. Type species: Prosodeskaszabi G.S. Medvedev & Kabakov, 1996, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Irianobates Kaszab, 1986: 291 [M]. Type species: Irianobateskrikkeni Kaszab, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Isaminas Champion, 1887: 266 [M]. Type species: Isaminasgibbipennis Champion, 1887, by subsequent designation (Gebien 1943: 401). Status: valid genus in Stenochiinae: Cnodalonini.

Isarida Pascoe, 1866a: 456 [F]. Type species: Isaridatestacea Pascoe, 1866 (= Caediusfulvus Mulsant & Rey, 1859), by monotypy. Status: junior synonym of Caedius Blanchard, 1845 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1939: 466).

Iscanus Fauvel, 1904: 176 [M]. Type species: Iscanuskuniensis Fauvel, 1904, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Ischnarthron Gebien, 1921b: 47 [N]. Type species: Ischnarthronlongipes Gebien, 1921, by monotypy. Status: valid genus in Diaperinae: Hypophlaeini.

Ischnodactylus Chevrolat, 1877: 173 [M]. Type species: Ischnodactylusquadrioculatus Chevrolat, 1877, by monotypy. Status: junior synonym of Basides Motschulsky, 1873 in Diaperinae: Diaperini: Diaperina. Synonymy: Schawaller (2004: 47, implied through placement of the type species of Basides Motschulsky, 1873 in Ischnodactylus Chevrolat, 1877). Note: the original spelling of the type species, Ischnodactylusquadridentatus, was corrected to Ischnodactylusquadrioculatus in the “Errata & Notes” of the same work (p. 178), Ischnodactylusquadrioculatus is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Isicerdes Champion, 1885: 113 [M]. Type species: Isicerdesoccultus Champion, 1885, by subsequent designation (R. Lucas 1920: 353). Status: valid genus in Stenochiinae: Cnodalonini.

Ismarus Haag-Rutenberg, 1878: 104 [M]. Type species: Ismarusgodeffroyi Haag-Rutenberg, 1878, by monotypy. Status: senior synonym of Simarus Borchmann, 1909 in Alleculinae: Alleculini: Alleculina. Note: junior homonym of Ismarus Haliday, 1835 [Hymenoptera].

Isocera Borchmann, 1909a: 713 [F]. Type species [automatic]: Isotomaemarginaticollis Blanchard, 1842, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: replacement name for Isotoma Blanchard, 1842.

Isocerus Dejean, 1821: 66 [M]. Type species: Tenebriopurpurascens Herbst, 1799 (= Helopsferrugineus Fabricius, 1798), by monotypy. Status: senior synonym of Neoisocerus Bouchard, Lawrence, Davies & Newton, 2005 in Blaptinae: Dendarini: Dendarina. Note: junior homonym of Isocerus Illiger, 1802 [Coleoptera: Cerambycidae].

Isomira Mulsant, 1856a: 52 [F]. Type species: Chrysomelamurina Linnaeus, 1758, by subsequent designation (C.G. Thomson 1859: 119). Status: valid genus and subgenus in Alleculinae: Alleculini: Gonoderina.

Isomiropsis Borchmann, 1942: 48 [F]. Type species: Isomiropsiswittei Borchmann, 1942, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Isoncophallus Koch, 1954a: 55 [M]. Type species: Isoncophalluszabroides Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Isonota Fairmaire, 1887a: 171 [F]. Type species: Isonotaopaca Fairmaire, 1887, by monotypy. Status: junior synonym of Homoeonota Fairmaire, 1882 in Pimeliinae: Tentyriini. Synonymy: Koch (1962c: 254).

Isopedus Stein in Stein and Weise, 1877: 121 [M]. Type species: Helopstenebrioides Germar, 1813, by present designation. Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: Heyden et al. (1883: 136, with Omaleis Allard, 1876, a synonym of Odocnemis Allard, 1876). Note: the original combination of the name of the type species, Helopstenebrioides Germar, 1813, is a junior primary homonym of Helopstenebrioides Palisot de Beauvois, 1812.

Isopteron Hope, 1841: 112 [N]. Type species: Isopteronaustrale Hope, 1841, by original designation. Status: valid genus in Lagriinae: Adeliini. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999, Article 12.2.6).

Isopteroplonyx Bremer, 2006: 7 [M]. Type species: Amarygmustermitophilus Lea, 1910, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Isopterum Agassiz, 1846b: 197 [N]. Type species [automatic]: Isopteronaustrale Hope, 1841, by original designation. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Note: unjustified emendation of Isopteron Hope, 1841, not in prevailing usage.

Isopus Montrouzier, 1860: 299 [M]. Type species: Isopusblanchardi Montrouzier, 1860, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Isostira Pascoe, 1870: 97 [F]. Type species: Isostiracrenata Pascoe, 1870, by monotypy. Status: junior synonym of Bradymerus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1921a: 253).

Isotoma Blanchard, 1842: pl. 15 [F]. Type species: Isotomaemarginaticollis Blanchard, 1842, by monotypy. Status: senior synonym of Isocera Borchmann, 1909 in Lagriinae: Lagriini: Statirina. Note: junior homonym of Isotoma Bourlet, 1839 [Collembola].

Itagonia Reitter, 1887a: 362 [F]. Type species: Itagoniagnaptorinoides Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Italohelops Español, 1961b: 295 [M]. Type species: Parablopssubchalybaeus Reitter, 1907, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Italomelia Mas-Peinado, Buckley, Ruiz & García-París, 2018: 539 [F]. Type species: Pimeliarugulosa Germar, 1823, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Itampolis Koch, 1962a: 31, 143 [F]. Type species: Itampolisoceanica Koch, 1962, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Iugidorsum Louw, 1979: 99, 102 [N]. Type species: Iugidorsumcumstriis Louw, 1979, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Ixalus Gistel, 1848a: x [M]. Type species [automatic]: Blapsexcavata Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: junior synonym of Platynotus Fabricius, 1801 in Blaptinae: Platynotini: Platynotina. Note: unnecessary replacement name for Platynotus Fabricius, 1801.

Jaklia Novák, 2010: 180 [F]. Type species: Jakliaserraticornis Novák, 2010, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Japetus Reitter, 1904: 160 [M]. Type species: Japetusmelanarius Reitter, 1904 (= Opatrummelanarium Erichson, 1843), by monotypy. Status: junior synonym of Trichosternum Wollaston, 1861 in Blaptinae: Opatrini: Opatrina. Synonymy: Gebien (1905: 253, with Trichopodus [as Trichopodum] Mulsant & Rey, 1859, a senior synonym of Trichosternum Wollaston, 1861). Note: junior homonym of Japetus Stål, 1863 [Hemiptera].

Javamarygmus Pic, 1928a: 22 [M]. Type species: Javamarygmustristis Pic, 1928, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Jintaium Ren in Ren and Yu, 1999: 228 [N]. Type species: Jintaiumsulcatum Ren, 1999, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Jophon Champion, 1895a: 224 [M]. Type species: Jophonmyrmecophilus Champion, 1895, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: Jophon is an incorrect subsequent spelling of the original spelling Iophon, first used by Kolbe (1898: 601), in prevailing usage; Jophon is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1), see Matthews and Bouchard (2008: 330).

Julogenius Reitter, 1906b: 138 [M]. Type species: Heliotaurusreichii Mulsant, 1856, by subsequent designation (Novák and Pettersson 2008: 333). Status: valid subgenus of Heliotaurus Mulsant, 1856 in Alleculinae: Cteniopodini.

†Jurallecula L.N. Medvedev, 1969: 124 [F]. Type species: Juralleculagrossa L.N. Medvedev, 1969, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: described from Upper Jurassic deposits (Kazakhstan).

Kabakoviella Kaszab, 1980c: 205 [F]. Type species: Kabakoviellamenephiloides Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Kaindilagria Merkl, 1988a: 138 [F]. Type species: Kaindilagriaforcipata Merkl, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Karroocara Koch, 1952a: 176 [N]. Type species: Stenocaragibbipenne Haag-Rutenberg, 1875, by original designation. Status: junior synonym of Stenodesia Reitter, 1916 in Pimeliinae: Adesmiini. Synonymy: Penrith (1979: 70, 85).

Kaszaba Matthews & Doyen, 1989: 40 [F]. Type species: Tenebriocorvinus Erichson, 1842, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Kaszabadelium Watt, 1992: 29 [N]. Type species: Adeliumaucklandicum Broun, 1880, by original designation. Status: valid genus in Lagriinae: Adeliini.

Kaszabiella Koch, 1943b: 774, 885 [F]. Type species: Thalpophilasubhemisphaerica Koch, 1943, by monotypy. Status: valid subgenus of Thalpophilodes Strand, 1942, in Pimeliinae: Tentyriini.

Kaszabochillus Fouquè, 2015: 228, 240 [M]. Type species: Indochillusandamanus Kaszab, 1981, by original designation. Status: valid subgenus of Pseudochillus Fouquè, 2015 in Pimeliinae: Stenosini: Dichillina.

Kaszaboscelis Löbl & Merkl, 2003: 246 [F]. Type species: Tenebriohypolithus Pallas, 1781, by original designation. Status: junior synonym of Platyscelis Latreille, 1818 in Blaptinae: Platyscelidini. Synonymy: Egorov (2004: 605).

Kaszabus Freude, 1967: 194 [M]. Type species: Kaszabusaurulentiformis Freude, 1967, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Kawiria Schuster, 1935: 26 [F]. Type species: Kawiriagabrieli Schuster, 1935, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Kershawia Lea, 1905: 379 [F]. Type species: Kershawiarugiceps Lea, 1905, by monotypy. Status: valid genus in Lagriinae: Belopini.

Kirgisomira Weise, 1974: 71 [F]. Type species: Isomiraophthalmica Seidlitz, 1896, by original designation. Status: junior synonym of Asiomira Dubrovina, 1973 in Alleculinae: Alleculini: Gonoderina. Synonymy: Muche (1981: 157).

Klapperichia Kaszab, 1954: 249 [F]. Type species: Klapperichiamirabilis Kaszab, 1954 (= Tenebriocephalonthoracicum Pic, 1925), by original designation. Status: junior synonym of Tenebriocephalon Pic, 1925 in Pimeliinae: Ceratanisini. Synonymy: Kaszab (1983a: 130).

Klewaria Reitter, 1910: 20 [F]. Type species: Klewariacolydiiformis Reitter, 1910, by monotypy. Status: valid genus in Pimeliinae: Klewariini.

Knausia Fall, 1931: 15 [F]. Type species: Knausiacrassicornis Fall, 1931, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Kocakia Kaszab, 1985: 54 [F]. Type species [automatic]: Idiopsisopaca Kaszab, 1981, by original designation. Status: valid genus in Pimeliinae: Edrotini. Note: replacement name for Idiopsis Kaszab, 1981.

Kocheria Antoine, 1946: 25, 30 [F]. Type species: Arthrodeisrungsi Español, 1943, by monotypy. Status: valid subgenus of Arthrodeis Solier, 1834 in Pimeliinae: Erodiini.

Kochogaster Kamiński & Raś, 2011: 654 [F]. Type species [automatic]: Anchophthalmusimpressicollis Fairmaire, 1897, by original designation. Status: valid subgenus of Anchophthalmus Gerstaecker, 1854 in Blaptinae: Platynotini: Platynotina. Note: replacement name for Cosmogaster Koch, 1956.

Kochotella Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Millotellamicrocornis Koch, 1962, by original designation. Status: valid genus in Pimeliinae: Asidini. Note: replacement name for Millotella Koch, 1962.

Kokeniella Reitter, 1906a: 41 [F]. Type species: Kokeniellamesostenoides Reitter, 1906, by subsequent designation (Gebien 1937a: 626). Status: valid genus in Pimeliinae: Tentyriini.

Kombacula Novák, 2012: 271 [F]. Type species: Kombaculakantneri Novák, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Koneus Giraldo-Mendoza & Flores, 2019: 94 [M]. Type species: Prohylithusperuanus Kaszab, 1981, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Kralia Novák, 2013: 500 [F]. Type species: Kraliaminshanica Novák, 2013, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Krollus R. Lucas, 1920: 357 [M]. Type species [automatic]: Homoeogenuslaticornis C.O. Waterhouse, 1882, by monotypy. Status: junior synonym of Borneocamaria Pic, 1917 in Stenochiinae: Cnodalonini. Note: replacement name for Homoeogenus C.O. Waterhouse, 1882.

Ksukolcula Novák, 2017a: 168 [F]. Type species: Ksukolculahesperia Novák, 2017, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Kuhitangia G.S. Medvedev, 1962: 1184 [F]. Type species: Kuhitangiakryzhanovskii G.S. Medvedev, 1962, by original designation. Status: valid genus in Kuhitangiinae: Kuhitangiini.

Kuschelus Kaszab, 1982b: 112 [M]. Type species: Kuscheluslathridioides Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Lupropini.

Labetis C.O. Waterhouse, 1879a: 267 [F]. Type species: Labetistibialis C.O. Waterhouse, 1879, by monotypy. Status: valid genus in Alleculinae: Alleculini: Mycetocharina.

Lachna Billberg, 1820: 35 [F]. Type species: Chrysomelahirta Linnaeus, 1758, by subsequent designation (Merkl 2006: 223). Status: junior synonym of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina. Synonymy: Lawrence and Newton (1995: 893).

Lachnodactylus Seidlitz, 1898a: 838 [M]. Type species [automatic]: Lachnopusdigitalis Seidlitz, 1894, by monotypy. Status: valid genus in Pimeliinae: Lachnogyini: Lachnodactylina. Note: replacement name for Lachnopus Seidlitz, 1894.

Lachnoderes Mulsant & Rey, 1859a: 70, 102 [M]. Type species: Pedonoecespubescens G.R. Waterhouse, 1845, by monotypy. Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Gemminger in Gemminger and Harold (1870: 1923, with Pedonoeces G.R. Waterhouse, 1845, a junior synonym of Blapstinus Dejean, 1821); Aalbu and Triplehorn (1991: 170).

Lachnogya Ménétriés, 1849: 228 [F]. Type species: Lachnogyasquamosa Ménétriés, 1849, by monotypy. Status: valid genus in Pimeliinae: Lachnogyini: Lachnogyina.

Lachnopus Seidlitz, 1894: 476 [M]. Type species: Lachnopusdigitalis Seidlitz, 1894, by monotypy. Status: senior synonym of Lachnodactylus Seidlitz, 1898 in Pimeliinae: Lachnogyini: Lachnodactylina. Note: junior homonym of Lachnopus Schönherr, 1840 [Coleoptera: Curculionidae].

Laena Dejean, 1821: 64 [F]. Type species: fixed herein (ICZN 1999, Article 70.3) as Scaurusviennensis Sturm, 1807, misidentified as Helopspimelia Fabricius, 1787 in the original designation by monotypy in Dejean (1821). Status: valid genus in Lagriinae: Laenini. Note: the type species “Helopspimelia Fabricius” was first established by monotypy (ICZN 1999, Article 68.3); Bouchard and Bousquet (2020b: 7) noted that the only valid species originally included in Laena, Helopspimelia Fabricius, 1787, is currently considered a valid species in the genus Penthe Newman, 1838 [Coleoptera: Tetratomidae] and therefore the taxonomic species involved, Helopspimelia Fabricius sensu Dejean 1821 (= Scaurusviennensis Sturm, 1807) is fixed here as the type species of Laena Dejean, 1821 according to the requirements of Article 70.3.2 (ICZN 1999).

Lagria Fabricius, 1775: 124 [F]. Type species: Chrysomelahirta Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Lagriinae: Lagriini: Lagriina.

Lagriallecula Pic, 1920b: 19 [F]. Type species: Lagrialleculaaeneipennis Pic, 1920, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Lagriella Borchmann, 1916a: 61 [F]. Type species: Lagriamima Borchmann, 1916, by original designation. Status: valid subgenus of Lagria Fabricius, 1775 in Lagriinae: Lagriini: Lagriina.

Lagrimina Fairmaire, 1894g: 675 [F]. Type species: Lagriminastrigipennis Fairmaire, 1894 (= Porrolagrianuda Kolbe, 1883), by monotypy. Status: junior synonym of Porrolagria Kolbe, 1883 in Lagriinae: Lagriini: Lagriina. Synonymy: Kolbe (1897a: 250).

Lagriocera Fairmaire, 1896a: 41 [F]. Type species: Lagrioceracavicornis Fairmaire, 1896, by monotypy. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Lagriodema Borchmann, 1930a: 442, 524 [F]. Type species: Nemostiragestroi Borchmann, 1910, by subsequent designation (Borchmann 1936: 458). Status: valid genus in Lagriinae: Lagriini: Statirina.

Lagriodes Borchmann, 1930a: 405, 432 [M]. Type species: Heterogriaarmigera Borchmann, 1930, by monotypy. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Lagriogonia Fairmaire, 1891e: ccxvii [F]. Type species: Lagriogoniahumerosa Fairmaire, 1891, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lagriola Kirsch, 1874: 409 [F]. Type species: Lagriolaoperosa Kirsch, 1874, by subsequent designation (Bousquet et al. 2018: 33). Status: junior synonym of Paratenetus Spinola, 1845 in Lagriinae: Goniaderini. Synonymy: Gebien (1922b: 268), Matthews and Lawrence (2015: 311).

Lagriomima Pic, 1934a: 31 [F]. Type species: Nemostiramaxima Pic, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lagriomima Pic, 1950: 11 [F]. Type species: Lagriomimaalbolineata Pic, 1950, by monotypy. Status: junior synonym of Neogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina. Synonymy: new synonym [OM]. Note: based on the description, the type species belongs to Neogria Borchmann, 1911 and is probably synonymous with the valid species Neogriacyanipennis Borchmann, 1911, therefore, Lagriomima Pic, 1950 is proposed as a new synonym of Neogria; junior homonym of Lagriomima Pic, 1934 [Coleoptera: Tenebrionidae: Lagriinae: Lagriini: Statirina].

Lagriopsis Borchmann, 1916a: 49, 138 [F]. Type species: Lagriopsisinsularis Borchmann, 1916, by subsequent designation (Borchmann 1936: 146). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Lagriostira Fairmaire, 1883b: 103 [F]. Type species: Statirarufonitens Fairmaire, 1883, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lagriostira Kolbe, 1902b: 550 [F]. Type species: Lagriostirahispida Kolbe, 1902 (= Lagriasubseriata Reitter, 1880), by subsequent designation (Borchmann 1936: 224). Status: senior synonym of Alagria Borchmann, 1916 in Lagriinae: Lagriini: Lagriina. Note: junior homonym of Lagriostira Fairmaire, 1883 [Coleoptera: Tenebrionidae: Lagriinae: Lagriini: Statirina].

Lamperos Allard, 1876a: 4 [M]. Type species: Helopsmicans Fabricius, 1798, by subsequent designation (Nabozhenko and Löbl 2008: 256). Status: junior synonym of Tarpela Bates, 1870 in Tenebrioninae: Helopini: Helopina. Synonymy: Champion (1887: 288).

Lamprobothris Fairmaire, 1887b: 302 [F]. Type species: Lamprobothrisattenuata Fairmaire, 1887 (= Praogenafossulata Müller, 1887), by monotypy. Status: junior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Synonymy: De Moor (1970: 42).

Lamprocrypticus Español, 1950: 127 [M]. Type species: Crypticusalpinus Comolli, 1837, by original designation. Status: valid genus in Diaperinae: Crypticini.

Lanhsia Shibata, 1980: 63, 64 [F]. Type species: Lanhsiabucca Shibata, 1980, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Laonicus Haag-Rutenberg, 1878: 100 [M]. Type species: Laonicuspilosus Haag-Rutenberg, 1878, by subsequent designation (R. Lucas 1920: 361). Status: junior synonym of Platyphanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 454).

Laoscapha Schawaller, 2016: 440 [F]. Type species: Laoscaphaphoupanensis Schawaller, 2016, by original designation. Status: valid genus in Diaperinae: Scaphidemini.

Laosocryptobates Pic, 1928a: 25 [M]. Type species: Laosocryptobatestuberculatus Pic, 1928, by monotypy. Status: junior synonym of Hexarhopalus Fairmaire, 1891 in Stenochiinae: Cnodalonini. Synonymy: Bečvář and Purchart (2008: 39).

Laraliprosodes Bogatchev, 1947: 513 [M]. Type species: Prosodeslar Bogatchev, 1947, by original designation. Status: junior synonym of Dineria Motschulsky, 1860 in Blaptinae: Blaptini: Blaptina. Synonymy: G.S. Medvedev and Iwan (2006: 617).

Lariversius Blaisdell, 1947: 59 [M]. Type species: Lariversiustibialis Blaisdell, 1947, by original designation. Status: valid genus in Blaptinae: Amphidorini.

Lasiocnema G.S. Medvedev, 1993: 110 [F]. Type species: Lasiostolaheterogena Fischer von Waldheim, 1844, by original designation. Status: valid subgenus of Lasiostola Dejean, 1834 in Pimeliinae: Pimeliini.

Lasioderus Mulsant & Rey, 1854: 13, 42 [M]. Type species: Lasioderussulcipennis Mulsant & Rey, 1854, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Lasiograna G.S. Medvedev, 1993: 111 [F]. Type species: Lasiostolainterrupta Reitter, 1901, by original designation. Status: valid subgenus of Lasiostola Dejean, 1834 in Pimeliinae: Pimeliini.

Lasiostola Dejean, 1834: 179 [F]. Type species: Tenebriopubescens Pallas, 1781, by subsequent designation (Hope 1841: 118). Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Latacula Campbell, 1971: 103 [F]. Type species: Lataculabeckeri Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Latetribolium Lepesme, 1943: 46 [N]. Type species: Triboliumrisbeci Lepesme, 1943 (= Opatrumlaevigatum Fabricius, 1781), by monotypy. Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Hinton (1948: 13, through synonymy of its type species with Opatrumlaevigatum Fabricius, 1781).

Latheticus C.O. Waterhouse, 1880: 147 [M]. Type species: Latheticusoryzae C.O. Waterhouse, 1880, by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Latipleurosis Penrith, 1977: 19, 204 [F]. Type species: Zophosisbenguelensis Deyrolle, 1867, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Latorhascius Pic, 1925b: 8 [M]. Type species: Rhaciusbaeri Pic, 1925, by monotypy. Status: valid subgenus of Adelonia Laporte, 1840 in Lagriinae: Belopini.

Lawrenceus Iwan, 1998b: 307 [M]. Type species: Lawrenceuscapensis Iwan, 1998, by original designation. Status: junior synonym of Schelodontes Koch, 1956 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan and Kamiński (2014: 171).

Leanum Uyttenboogaart, 1934: 29, 31 [N]. Type species: Triboliummyrmecophilum Lea, 1904, by monotypy. Status: junior synonym of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini. Synonymy: Hinton (1948: 21).

Leaus Matthews & Lawrence, 1992: 312 [M]. Type species: Leaustasmanicus Matthews & Lawrence, 1992, by original designation. Status: valid genus in Tenebrioninae: Trachelostenini. Note: transferred from Titaenini by Matthews and Lawrence (2015: 293).

Lechinius Blair, 1922: 561 [M]. Type species: Lechiniuscatenulatus Blair, 1922 (= Cistelomorphafossulata Pic, 1913), by monotypy. Status: valid subgenus of Cteniopinus Seidlitz, 1896 in Alleculinae: Cteniopodini.

Lechinius Borchmann, 1930b: 151 [M]. Type species: Cistelomorphafossulata Pic, 1913, by subsequent designation (Novák and Pettersson 2008: 330). Status: junior synonym of Lechinius Blair, 1922 in Alleculinae: Cteniopodini. Note: junior homonym of Lechinius Blair, 1922 [Coleoptera: Alleculinae: Cteniopodini].

Lechius Iwan, 1995c: 404 [M]. Type species: Selinusabacoides Fairmaire, 1902, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Lechriomus Morawitz, 1865: 21 [M]. Type species: Akislucifuga Adams, 1817, by subsequent designation (Chernei 2005: 102). Status: valid subgenus of Cyphogenia Solier, 1837 in Pimeliinae: Akidini.

Leichenum Dejean, 1834: 194 [N]. Type species: Opatrumpictum Fabricius, 1801, by monotypy. Status: valid genus in Blaptinae: Pedinini: Leichenina.

Leichrodomorphus Pic, 1921c: 6 [M]. Type species: Leichrodomorphusbrevicornis Pic, 1921, by subsequent designation (Pic 1922e: 135). Status: junior synonym of Stethotrypes Gebien, 1914 in Diaperinae: Leiochrinini. Synonymy: Gebien (1940: 431).

Leiochrinus Westwood, 1883: 68 [M]. Type species: Leiochrinusfulvicollis Westwood, 1883, by subsequent designation (W.F. Kirby 1885a: 89). Status: valid genus in Diaperinae: Leiochrinini.

Leiochrodes Westwood, 1883: 69 [M]. Type species: Leiochrodesdiscoidalis Westwood, 1883, by subsequent designation (W.F. Kirby 1885a: 89). Status: junior synonym of Ades Guérin-Méneville, 1857 in Diaperinae: Leiochrinini. Synonymy: Gebien (1911a: 388).

Leiochrodinus Kaszab, 1961a: 358, 365 [M]. Type species: Leiochrodinustetraphyllus Kaszab, 1961, by original designation. Status: valid genus in Diaperinae: Leiochrinini. Note: the alternative original spelling Leichrodinus, used by Kaszab (1961a: 365), was rejected by Papp and Seeno (1981: 53, 71) who acted as the First Revisers.

Leiochrodontes Kaszab, 1946a: 30, 200 [M]. Type species: Leiochrodesmadurensis Pic, 1918, by original designation. Status: valid genus in Diaperinae: Leiochrinini. Note: the earlier usage of Leiochrodontes by Gebien (1940: 755) was not accompanied by a description, a definition or a bibliographic reference to such a published statement and therefore is not available from that date.

Leiochromimus Ardoin, 1966: 187, 199 [M]. Type species: Coccimarygmuspunctatus Ardoin, 1966, by monotypy. Status: valid subgenus of Coccimarygmus Ardoin, 1966 in Tenebrioninae: Amarygmini.

Leiochrota Westwood, 1883: 70 [F]. Type species: Leiochrinusuniformis Westwood, 1883, by subsequent designation (W.F. Kirby 1885a: 90). Status: valid genus in Diaperinae: Leiochrinini.

Leiochrotina Westwood, 1883: 70, 76 [F]. Type species: Leiochrinusindicus Westwood, 1883, by monotypy. Status: junior synonym of Crypsis C.O. Waterhouse, 1877 in Diaperinae: Leiochrinini. Synonymy: Gebien (1940: 436).

Leiopeplus Broun, 1893b: 1160 [M]. Type species: Helopsexpolitus Broun, 1880, by original designation. Status: senior synonym of Chrysopeplus Gebien, 1942 in Stenochiinae: Cnodalonini. Note: junior homonym of Leiopeplus Murray, 1864 [Coleoptera: Nitidulidae].

Leipopleura Seidlitz, 1893: 342, 345 [F]. Type species: Faustiaintegra Reitter, 1887, by subsequent designation (Kaszab 1940a: 167). Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Lelegeis Champion, 1886: 209 [M]. Type species: Lelegeisaeneipennis Champion, 1886, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Leleupium Kaszab, 1956b: 106 [N]. Type species: Leleupiumsubcoecum Kaszab, 1956, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Lemoultia Chatanay, 1913b: 313 [F]. Type species: Lemoultiascabripennis Chatanay, 1913, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Lenkous Kaszab, 1973b: 315 [M]. Type species: Lenkousmyrmecophilus Kaszab, 1973, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Lepidocaulinus Schawaller, Masumoto & Merkl, 2013: 378 [M]. Type species: Lepidocaulinusmirabilis Schawaller, Masumoto & Merkl, 2013, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Lepidochora Koch, 1952b: 36 [F]. Type species: Lepidochoraeberlanzi Gebien, 1938, by original designation. Status: valid genus in Pimeliinae: Adelostomini. Note: the earlier usage of Lepidochora by Gebien (1938b: 71) is unavailable since it was published after 1930 without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Lepidocnemeplatia Bousquet & Bouchard in Bousquet et al., 2018: 20 [F]. Type species: Cnemeplatiasericea Horn, 1870, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Note: Lepidocnemeplatia was previously described by Kaszab (1938: 80) without original type species fixation and is therefore unavailable from that date (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 245) designated Cnemeplatiasericea Horn, 1870 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Lepidospilus Agassiz, 1846b: 205 [M]. Type species [automatic]: Pachycoeliasulcicollis Boisduval, 1835, by monotypy. Status: junior synonym of Pachycoelia Boisduval, 1835 in Tenebrioninae: Heleini: Cyphaleina. Note: unjustified emendation of Lepispilus Westwood, 1841, not in prevailing usage.

Lepispilus Westwood, 1841a: 44 [M]. Type species: Pachycoeliasulcicollis Boisduval, 1835, by monotypy. Status: junior synonym of Pachycoelia Boisduval, 1835 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Lacordaire (1859b: 413).

Leprocaulinus Kaszab, 1982c: 75 [M]. Type species: Leprocaulinuskrikkeni Kaszab, 1982, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: this genus-group name has been considered a junior homonym of Leprocaulinus Uvarov, 1940 [Orthoptera] in literature on Tenebrionidae; however, no type species has yet been designated for orthopteran name Leprocaulus Redtenbacher, 1908 nor its replacement name Leprocaulinus Uvarov, 1940 (see Herwaarden 1998: 65) and therefore Leprocaulinus Uvarov, 1940 is unavailable since it has been proposed as a replacement name after 1930, without a type species designation, for a generic taxon without valid typification (ICZN 1999: Article 13.3.1).

Leprocaulus Fairmaire, 1896c: 95 [M]. Type species: Leprocaulusclavipes Fairmaire, 1896, by monotypy. Status: valid subgenus of Hexarhopalus Fairmaire, 1891 in Stenochiinae: Cnodalonini.

Leptasida Chatanay, 1914b: 3 [F]. Type species: Leptasidatenuipes Chatanay, 1914, by original designation. Status: valid genus in Pimeliinae: Asidini.

Leptasida Reitter, 1917a: 40, 60 [F]. Type species: Asidadiecki Allard, 1869, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 291). Note: junior homonym of Leptasida Chatanay, 1914 [Coleoptera: Tenebrionidae: Pimeliinae: Asidini].

Leptinostethus Borchmann, 1936: 238, 466 [M]. Type species: Leptinostethusmethneri Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Leptocolena Allard, 1880: 74 [F]. Type species: Blapsmucronata Latreille, 1804, by subsequent designation (Harold 1882: 194). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gebien (1910b: 226).

Leptoderis Billberg, 1820: 31 [F]. Type species: Tenebriocollaris Linnaeus, 1767, by monotypy. Status: valid genus in Pimeliinae: Elenophorini: Elenophorina.

Leptoderops Löbl, Bouchard, Merkl & Bousquet, 2020: 5 [M]. Type species: Thraustocoluspriesneri Koch, 1934, by original designation. Status: valid subgenus of Thraustocolus Kraatz, 1866 in Pimeliinae: Tentyriini. Note: name first proposed by Koch (1934: 96) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl et al. (2008a: 40) designated Thraustocoluspriesneri Koch, 1934 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Leptodes Dejean, 1834: 181 [M]. Type species: Sepidium boisduvalii Zubkov, 1833, by monotypy. Status: valid genus and subgenus in Pimeliinae: Leptodini.

Leptodinopsis Kaszab, 1959b: 353, 359 [F]. Type species: Sepidium boisduvalii Zubkov, 1833, by original designation. Status: junior synonym of Leptodes Dejean, 1834 in Pimeliinae: Leptodini. Synonymy: G.S. Medvedev and Iljina (2007: 882).

Leptodopsis Haag-Rutenberg, 1879a: 408 [F]. Type species: Leptodopsisinsignis Haag-Rutenberg, 1879, by monotypy. Status: valid subgenus of Leptodes Dejean, 1834 in Pimeliinae: Leptodini.

Leptogastrus W.J. MacLeay, 1872: 293 [M]. Type species: Leptogastrusmastersii W.J. MacLeay, 1872, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Leptomorpha Faldermann, 1835: 406 [F]. Type species: Leptomorphachinensis Faldermann, 1835, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Champion (1895: 48).

Leptonychoides Schawaller, 1990: 51 [M]. Type species: Leptonychoidesjuengeri Schawaller, 1990, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Leptonychus Chevrolat, 1833a: 26, pl. 1 [M]. Type species: Leptonychuserodioides Chevrolat, 1833, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Leptoscapha Fairmaire, 1886c: 73 [F]. Type species [automatic]: Stenoscaphaspissicornis Fairmaire, 1885, by monotypy. Status: senior synonym of Brachypophlaeus Fairmaire, 1897 in Tenebrioninae: Ulomini. Synonymy: Alluaud (1899: 342, as “Brachypophloeus”). Note: replacement name for Stenoscapha Fairmaire, 1885; junior homonym of Leptoscapha Fischer, 1883 [Mollusca].

Leptosora Borchmann, 1936: 238, 471 [F]. Type species: Leptosorahamata Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Leptosphena Semenov, 1891: 356, 358 [F]. Type species: Sphenariatomentosa Semenov, 1889 (= Sphenariarubripes Reitter, 1889), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Lepturidea Fauvel, 1862: 150 [F]. Type species: Lepturideadeplanchei Fauvel, 1862, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Leptynoderes Solier, 1838b: 8, 44 [M]. Type species: Scotobiusvaricosus Germar, 1823, by original designation. Status: valid genus in Tenebrioninae: Scotobiini.

Lesbidana Reitter, 1904: 173 [F]. Type species: Melanesthessimplex Reitter, 1897, by subsequent designation (G.S. Medvedev 1990: 203). Status: valid subgenus of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina.

Leucolaephus P.H. Lucas, 1859: xxii [M]. Type species: Leucolaephusperrisii P.H. Lucas, 1859 (= Pimelialiliputana P.H. Lucas, 1857), by subsequent designation (Gebien 1937a: 803). Status: valid genus in Pimeliinae: Pimeliini. Note: Leucolaephus is an incorrect subsequent spelling of the original spelling Leucoloephus, first used by Kraatz (1865: 314), and is in prevailing usage; Leucolaephus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1), see Bouchard et al. (2005: 506).

Lichenum Agassiz, 1846b: 203, 209 [N]. Type species [automatic]: Opatrumpictum Fabricius, 1801, by monotypy. Status: junior synonym of Leichenum Dejean, 1834 in Blaptinae: Pedinini: Leichenina. Note: unjustified emendation of Leichenum Dejean, 1834, not in prevailing usage.

Licinoma Pascoe, 1869: 140 [F]. Type species: Licinomanitida Pascoe, 1869, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Licymnius Bates, 1868: 271 [M]. Type species: Licymniusfoveicollis Bates, 1868, by monotypy. Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Carter (1915a: 60, with Chromomaea Pascoe, 1866, a junior synonym of Lepturidea Fauvel, 1862).

Lindia Blackburn, 1888: 275 [F]. Type species: Lindiaangusta Blackburn, 1888, by monotypy. Status: junior synonym of Lyphia Mulsant & Rey, 1859 in Tenebrioninae: Triboliini. Synonymy: Champion (1894a: 351, 370). Note: junior homonym of Lindia Dujardin, 1841 [Rotifera].

Lineocrypticus Koch, 1950c: 52 [M]. Type species: Lineocrypticushessei Koch, 1950, by original designation. Status: valid genus in Diaperinae: Crypticini.

Linio Bouchard & Bousquet, new subgenus [M]. Type species: Niliolanatus Germar, 1823, by present designation. Status: valid subgenus of Nilio Latreille, 1802 in Nilioninae. Note: Mader (1936: 94) introduced the new subgenus name Linio for several nominal species, but unfortunately did not designate a type species; the subgenusLinio, which has been treated as valid since 1936, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Niliolanatus Germar, 1823 as type species and referring to Mader (1936: 94) for the character states that characterise and differentiate Linio.

Liodema Horn, 1870: 378, 385 [F]. Type species: Platydemalaevis Haldeman, 1848, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Liodocistela Pic, 1930a: 28 [F]. Type species: Cistelopsisrufomarginata Pic, 1930, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Lioprosodes Reitter, 1909a: 121 [M]. Type species: Prosodesdilaticollis Motschulsky, 1859, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Skopin (1960a: 46).

Lisa Haag-Rutenberg, 1879c: 134 [F]. Type species: Lisasingularis Haag-Rutenberg, 1879 (= Alleculaomophloides Hope, 1843), by monotypy. Status: junior synonym of Metistete Pascoe, 1866 in Alleculinae: Alleculini: Alleculina. Synonymy: Carter (1915a: 78).

Litasida Casey, 1912: 77, 184 [F]. Type species: Litasidatownsendi Casey, 1912, by original designation. Status: valid genus in Pimeliinae: Asidini.

Litheleodes Blaisdell, 1909: 114 [M]. Type species: Blaps extricata Say, 1824, by subsequent designation (Triplehorn and Thomas 2015: 11). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Lithoblaps Motschulsky, 1860c: 532 [F]. Type species: Tenebriogigas Linnaeus, 1767, by subsequent designation (Skopin, 1960a: 44). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860). Note: the original combination of the name of the type species, Tenebriogigas Linnaeus, 1767, is a junior primary homonym of Tenebriogigas Linnaeus, 1763.

Litoboriolus Español, 1945: 310, 313 [M]. Type species: Olocratescollaris Mulsant & Rey, 1854, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Litoboromimus Koch, 1948: 413 [M]. Type species: Litoborusparallelus Schuster, 1919, by original designation. Status: valid subgenus of Allophylax Bedel, 1906 in Blaptinae: Dendarini: Melambiina.

Litoborus Mulsant & Rey, 1854: 124, 126 [M]. Type species: Phylaxmoreletii P.H. Lucas, 1846, by subsequent designation (Antoine 1931: 181). Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Litopous Matthews, 2012: 1 [M]. Type species: Litopousbaehri Matthews, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: placed in the subtribe Alleculina by Matthews and Lawrence (2019: 645).

Litororus Reitter, 1904: 96 [M]. Type species: Micrositussemicostatus Mulsant & Rey, 1854, by monotypy. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Lixionica Blackburn, 1896: 280 [F]. Type species: Lixionicacostatipennis Blackburn, 1896, by monotypy. Status: valid genus in Pimeliinae: Vacronini.

Lobatopezus Pic, 1952c: 2 [M]. Type species: Lobatopezusdiversecostatus Pic, 1952, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: Bremer (2005: 211) transferred this genus to Chrysomeloidea: Chrysomelidae, but later retained this genus in Tenebrioninae: Amarygmini after further study (Bremer 2013b: 127–128).

Lobetas Motschulsky, 1872: 26 [M]. Type species: Tenebriocostatus Guérin-Méneville, 1831, by original designation. Status: junior synonym of Hipalmus Bates, 1870 in Tenebrioninae: Tenebrionini. Synonymy: C.O. Waterhouse (1876: 288). Note: the original combination of the name of the type species, Tenebriocostatus Guérin-Méneville, 1831, is a junior primary homonym of Tenebriocostatus Pallas, 1781.

Lobodera Mulsant & Rey, 1859c: 12, 18 [F]. Type species: Loboderarufescens Mulsant & Rey, 1859, by monotypy. Status: junior synonym of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Synonymy: Gebien (1910b: 333).

Lobometopon Casey, 1907: 379, 385 [N]. Type species: Epitragusfusiformis Casey, 1890, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Lobophilomorphus Pic, 1911a: 183 [M]. Type species: Lobophilomorphusrubicollis Pic, 1911, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lobopoda Solier, 1835a: 233 [F]. Type species: Lobopodastriata Solier, 1835, by subsequent designation (Bousquet et al. 2015: 134). Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Lobothorax Gemminger, 1870: 124 [M]. Type species [automatic]: Loboderarufescens Mulsant & Rey, 1859, by monotypy. Status: junior synonym of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Note: unnecessary replacement name for Lobodera [as “Loboderus”] Mulsant & Rey, 1859.

Locrodes Casey, 1907: 332 [M]. Type species: Emmenastuspiceus Casey, 1890, by subsequent designation (Bousquet et al. 2018: 104). Status: valid subgenus of Hylocrinus Casey, 1907 in Pimeliinae: Edrotini.

Lodinus Mulsant & Rey, 1859a: 113, 131 [M]. Type species: Lodinusnigroaeneus Mulsant & Rey, 1859 (= BlapstinuspunctulatusSolier 1851), by monotypy. Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Gemminger in Gemminger and Harold (1870: 1923).

Loensus R. Lucas, 1920: 380, 489 [M]. Type species [automatic]: Pedinopsispilipes Gebien, 1910, by monotypy. Status: valid genus and subgenus in Blaptinae: Pedinini: Pedinina. Note: replacement name for Pedinopsis Gebien, 1910.

Lomocnemis Gebien, 1921a: 287 [F]. Type species: Lomocnemispolita Gebien, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Longuloodescelis Kaszab, 1940b: 940, 957 [F]. Type species: Platyscelishirta Seidlitz, 1893, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini. Note: the First Reviser (Longuloodescelis Kaszab, 1940 versus Trichoodescelis Kaszab, 1940) is Egorov (2004: 595).

Lophocnemis Mäklin, 1867: 505 [F]. Type species: Lophocnemisamabilis Mäklin, 1867, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Lopholagria Borchmann, 1916a: 48, 97 [F]. Type species: Lagriaamoena Fåhraeus, 1870, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Lagriina.

Lophoma Solier, 1835b: 253, 285 [N]. Type species: Pimeliapunctata Fabricius, 1798, by monotypy. Status: junior synonym of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Lacordaire (1859a: 46). Note: the original combination of the name of type species, Pimeliapunctata Fabricius, 1798, is a junior primary homonym of Pimeliapunctata Thunberg, 1787.

Lophophyllus Fairmaire, 1887c: 71 [M]. Type species: Lophophylluscostipennis Fairmaire, 1887, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Lordodera Gebien, 1921b: 64 [F]. Type species: Tenebrioquadrihamatus Fairmaire, 1875, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Lorelopsis Champion, 1896: 15 [F]. Type species: Lorelopsispilosa Champion, 1896, by monotypy. Status: junior synonym of Lorelus Sharp, 1876 in Lagriinae: Lupropini. Synonymy: Doyen (1993: 295).

Lorelus Sharp, 1876: 76 [M]. Type species: Loreluspriscus Sharp, 1876, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Loricula Novák, 2016d: 45 [F]. Type species: Alleculasubaeneipennis Pic, 1922, by original designation. Status: senior synonym of Loriculoides Novák, 2020 in Alleculinae: Alleculini: Alleculina. Note: junior homonym of Loricula Curtis, 1833 [Hemiptera].

Loriculoides Novák, 2020i: 9 [M]. Type species [automatic]: Alleculasubaeneipennis Pic, 1922, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: replacement name for Loricula Novák, 2016.

Lornamus Koch, 1952a: 191 [M]. Type species: Lornamusdividiopsis Koch, 1952, by original designation. Status: junior synonym of Cryptocarpes Koch, 1952 in Pimeliinae: Caenocrypticini. Synonymy: Endrödy-Younga (1996: 21).

Lorona Borchmann, 1936: 16, 62 [F]. Type species: Lagriabakeri Borchmann, 1930, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Loubacantus Bonadona, 1959: 1033 [M]. Type species: Loubacantusgiganteus Bonadona, 1959, by original designation. Status: junior synonym of Entypodera Gerstaecker, 1871 in Lagriinae: Lagriini: Statirina. Synonymy: Bonadona (1984: 469).

Louwerensia Kaszab, 1964b: 104 [F]. Type species: Louwerensiapapuana Kaszab, 1964, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Loxostethus Triplehorn, 1962: 504 [M]. Type species: Loxostethusfasciatus Triplehorn, 1962, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: new placement [RLA], previously included in Diaperinae: Diaperini: Adelinina.

Lucidolaena Endrödy-Younga & Schawaller, 2002: 9, 20 [F]. Type species: Lucidolaenaamatolensis Endrödy-Younga & Schawaller, 2002, by original designation. Status: valid genus in Lagriinae: Laenini.

Luebbertia Koch, 1963: 20, 44 [F]. Type species: Luebbertiaplana Koch, 1963, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Luprops Hope, 1833: 63 [M]. Type species: Lupropschrysophthalmus Hope, 1833 (= Tageniaindica Wiedemann, 1823), by monotypy. Status: valid genus in Lagriinae: Lupropini.

Luzonoplonyx Bremer, 2009: 331 [M]. Type species: Luzonoplonyxgrandis Bremer, 2009, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Lycanthropa J. Thomson, 1860b: 20 [F]. Type species: Eurychoracimicoides Quensel, 1806, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Lycidioides Ando, 2003a: 107 [M]. Type species: Lycidioideskaniei Ando, 2003, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Lycogonocnemis Pic, 1915a: 13 [F]. Type species: Lycogonocnemisrufa Pic, 1915, by monotypy. Status: valid subgenus of Paragonocnemis Kraatz, 1899 in Tenebrioninae: Amarygmini.

Lycoscelis Blair, 1929a: 242 [F]. Type species: Lycoscelisfulva Blair, 1929 (= Plinthochrousgounellei Fairmaire, 1891), by original designation. Status: junior synonym of Plinthochrous Fairmaire, 1891 in Tenebrioninae: Amarygmini. Synonymy: Ardoin (1962b: 957).

Lycula Campbell, 1976: 30 [F]. Type species: Lyculachilensis Campbell, 1976, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Lygestira Pascoe, 1866a: 470 [F]. Type species: Prophanessimplex Westwood, 1849, by subsequent designation (R. Lucas 1920: 384). Status: junior synonym of Prophanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 488).

Lygophilus Rafinesque, 1815: 114 [M]. Type species [automatic]: Epitragusfuscus Latreille, 1804, by subsequent monotypy (Latreille 1804: 322). Status: junior synonym of Epitragus Latreille, 1802 in Pimeliinae: Epitragini. Note: unnecessary replacement name for Epitragus Latreille, 1802.

Lyphia Mulsant & Rey, 1859b: 166 [F]. Type species: Lyphiaficicola Mulsant & Rey, 1859 (= Biustetraphyllus Fairmaire, 1857), by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Lyprochelyda Fairmaire, 1899d: 214 [F]. Type species: Lyprochelydapurpurina Fairmaire, 1899, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Lyprosodes Reitter, 1909a: 116 [M]. Type species: Prosodesquadricostata Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Lystronychus Latreille, 1829a: 41 [M]. Type species: Helopsequestris Fabricius, 1775, by subsequent designation (Saunders 1837: 154). Status: valid genus and subgenus in Alleculinae: Alleculini: Xystropodina. Note: Lystronychus is an incorrect subsequent spelling of the original spelling Lystronichus, first used by Perty (1832: 63), in prevailing usage; Lystronychus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Macellocerus Solier, 1848: 154, 263 [M]. Type species [automatic]: Dolichoderusacuminatus Klug, 1833, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Nycteropina. Note: replacement name for Dolichoderus Klug, 1833; discovery of the older unused name Dillacerus Solier, 1835 threatens the stability of Macellocerus Solier, 1848; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to conserve usage of Macellocerus Solier, 1848.

Machla Herbst, 1799: 152 [F]. Type species: Opatrumvillosum G.-A. Olivier, 1795, by subsequent designation (R. Lucas 1920: 386). Status: valid genus in Pimeliinae: Asidini. Note: the older name Machla Lichtenstein, 1796 was published in a work that was suppressed for nomenclatural purposes (ICZN 1995, Opinion 1820).

Machlasida Escalera, 1907: 336 [F]. Type species: Asidamuleyhafidi Escalera, 1907, by subsequent designation (Escalera 1928: 137). Status: valid subgenus of Alphasida Escalera, 1905 in Pimeliinae: Asidini.

Machleida Fåhraeus, 1870: 256 [F]. Type species: Machleidanodulosa Fåhraeus, 1870, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Machloida Rye, 1873: 286 [F]. Type species [automatic]: Machleidanodulosa Fåhraeus, 1870, by monotypy. Status: junior synonym of Machleida Fåhraeus, 1870 in Pimeliinae: Asidini. Note: unjustified emendation of Machleida Fåhraeus, 1870, not in prevailing usage.

Machlomorpha Péringuey, 1899: 257 [F]. Type species: Machlomorphaaltitudinis Péringuey, 1899, by subsequent designation (Gebien 1937a: 739). Status: valid genus and subgenus in Pimeliinae: Asidini.

Machlophila Wilke, 1924: 521 [F]. Type species: Machlophilavogti Wilke, 1924, by monotypy. Status: junior synonym of Machla Herbst, 1799 in Pimeliinae: Asidini. Synonymy: Koch (1962a: 121, 144).

Machloplasta Wilke, 1922: 263 [F]. Type species: Opatrumvillosum G.-A. Olivier, 1795, by original designation. Status: junior synonym of Machla Herbst, 1799 in Pimeliinae: Asidini. Synonymy: Koch (1962a: 119).

Machlopsis Pomel, 1871: 236 [F]. Type species: Eurychoralevaillantii P.H. Lucas, 1850, by original designation. Status: valid genus and subgenus in Pimeliinae: Adelostomini. Note: Machlopsis is an incorrect subsequent spelling of the original spelling Maclopsis, first used by Bedel (1887: 199), in prevailing usage; Machlopsis is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Macradesmia Kaszab, 1959a: 402 [F]. Type species: Adesmiaservillei Solier, 1835, by monotypy. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: name first proposed by Koch (1944b: 146) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Macradesmia Löbl & Merkl in Löbl et al., 2020: 1 [F]. Type species: Pimeliacancellata Klug, 1830, by original designation. Status: junior synonym of Macradesmia Kaszab, 1959 in Pimeliinae: Adesmiini. Note: name first proposed by Koch (1944b: 146) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); not knowing about the availability of Macradesmia Kaszab, 1959, Löbl & Merkl (2003: 244) designated Pimeliacancellata Klug, 1830 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1) and subsequently Macradesmia was proposed as new by Löbl and Merkl in Löbl et al. (2020: 1); junior homonym of Macradesmia Kaszab, 1959 [Coleoptera: Tenebrionidae: Pimeliinae: Adesmiini].

Macrarthra Agassiz, 1846b: 35, 219 [F]. Type species [automatic]: Arthromacradonacioides W. Kirby, 1837 (= Lagriaaenea Say, 1824), by monotypy. Status: junior synonym of Arthromacra W. Kirby, 1837 in Lagriinae: Lagriini: Statirina. Note: unjustified emendation of Arthromacra W. Kirby, 1837, not in prevailing usage; the original combination of the accepted name of the type species, Lagriaaenea Say, 1824, is a junior primary homonym of Lagriaaenea Fabricius, 1775.

Macroartactes Pic, 1924a: 24 [M]. Type species: Macroartactescostulatus Pic, 1924, by subsequent designation (Gebien 1935: 64). Status: junior synonym of Artactes Pascoe, 1868 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1935: 64).

Macrocasnonidea Pic, 1934a: 31 [F]. Type species: Nemostiraquadrimaculata Pic, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Macrocistela Pic, 1941: 13 [F]. Type species: Macrocistelastriata Pic, 1941, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Macrocistelopsis Pic, 1956: 89 [F]. Type species: Macrocistelopsistestaceicolor Pic, 1956, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Macrohyperchalca Pic, 1935b: 98 [F]. Type species: Hyperchalcastriata Pic, 1935, by monotypy. Status: valid subgenus of Hyperchalca Fairmaire, 1869 in Stenochiinae: Stenochiini.

Macrolagria Lewis, 1895: 422 [F]. Type species: Statirarufobrunnea Marseul, 1876, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Macropachylesthus Pic, 1923c: 28 [M]. Type species: Macropachylesthusgigas Pic, 1923, by subsequent designation (Gebien 1941: 350). Status: valid genus in Stenochiinae: Cnodalonini.

Macroperas Carter, 1914c: 393 [N]. Type species: Macroperasantennale Carter, 1914, by monotypy. Status: junior synonym of Daedrosis Bates, 1868 in Lagriinae: Adeliini. Synonymy: Matthews (1998: 756).

Macrophanes Iablokoff-Khnzorian, 1957: 165 [M]. Type species: Hedyphanescorax Iablokoff-Khnzorian, 1957 (= Helopsamandanus Reitter, 1902), by original designation. Status: junior synonym of Delonurops Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko (2002b: 689).

Macrophtalmus Montrouzier, 1855: 33 [M]. Type species: Macrophtalmuscoeruleus Montrouzier, 1855, by monotypy. Status: senior synonym of Macrophthalmata Strand, 1935 in Tenebrionidae: incertae sedis. Note: this genus has been treated as a junior homonym of Macrophtalmus Laporte, 1832 [Hemiptera] in the literature; however, the hemipteran name was originally proposed with two spellings, Macrophthalmus (Laporte, 1832: 6, 11, 87) and Macrophtalmus (Laporte, 1832: 78); the alternative original spelling Macrophtalmus of the hemipteran name, used by Laporte (1832: 78), was rejected by Neave (1940: 17) who acted as First Reviser; therefore Macrophtalmus Montrouzier, 1855 is the valid name for the tenebrionid genus since it is not a junior homonym.

Macrophthalmata Strand, 1935b: 302 [F]. Type species [automatic]: Macrophtalmuscoeruleus Montrouzier, 1855, by monotypy. Status: junior synonym of Macrophtalmus Montrouzier, 1855 in Tenebrionidae: incertae sedis. Note: replacement name for Macrophtalmus Montrouzier, 1855.

Macropoda Solier, 1835b: 512, 515 [F]. Type species: Pimeliavariolaris G.-A. Olivier, 1795, by subsequent designation (Hope 1841: 118). Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Macropodesmia Löbl & Merkl in Löbl et al., 2020: 1 [F]. Type species: Pimeliareticulata Klug, 1830, by original designation. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: name first proposed by Koch (1944b: 147) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 244) designated Pimeliareticulata Klug, 1830 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Macrostethus Wollaston, 1854: 504 [M]. Type species: Macrostethustuberculatus Wollaston, 1854, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Macrosynopticus Pic, 1922d: 25 [M]. Type species: Macrosynopticuscostatus Pic, 1922, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Macrotrachyscelis Pic, 1925b: 12 [F]. Type species: Macrotrachyscelisrufa Pic, 1925, by monotypy. Status: valid genus in Diaperinae: Trachyscelini.

Macrozophobas Pic, 1913b: 6 [M]. Type species: Macrozophobasgracilicornis Pic, 1913, by monotypy. Status: valid subgenus of Zophobas Dejean, 1834 in Tenebrioninae: Tenebrionini.

Macruloma Pic, 1921d: 20 [N]. Type species: Macrulomagigas Pic, 1921, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Madagassa Koch, 1950a: 67 [F]. Type species [automatic]: Pycnaaphodina Fairmaire, 1894, by monotypy. Status: valid genus in Phrenapatinae: Penetini. Note: replacement name for Pycna Fairmaire, 1894.

Madobalus Fairmaire, 1901a: 73 [M]. Type species: Madobalusrotundicollis Fairmaire, 1901, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Madreallecula Kanda, 2013: 587 [F]. Type species: Madrealleculamcclevei Kanda, 2013, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Maerodes C.O. Waterhouse, 1877: 72 [M]. Type species: Prophaneswestwoodi W.J. MacLeay, 1872 (= Prophanesaculeatus Westwood, 1849), by monotypy. Status: junior synonym of Prophanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1918: 717).

Magdanalia Novák, 2020c: 496 [F]. Type species: Prionychusdenticulatus Muche, 1982, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Magela G.S. Medvedev & Lawrence, 1986: 578 [F]. Type species: Magelauptoni G.S. Medvedev & Lawrence, 1986, by original designation. Status: valid genus in Diaperinae: Hyociini: Brittonina.

Magrebmelia Mas-Peinado, Buckley, Ruiz & García-París, 2018: 540 [F]. Type species: Pimeliaxauenensis Escalera, 1923, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Mahena Gebien, 1922b: 315 [F]. Type species: Mahenacuprea Gebien, 1922, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Makicula Novák, 2012: 275 [F]. Type species: Makiculaphoupaneica Novák, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Malacova Fairmaire, 1898c: 479 [F]. Type species: Malacovabicolor Fairmaire, 1898, by present designation. Status: junior synonym of Damatris Laporte, 1840 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1942a: 316).

Malaiseum Borchmann, 1941a: 13 [N]. Type species: Malaiseumsingulare Borchmann, 1941, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Malayaplamius Masumoto, 1986a: 17 [M]. Type species: Malayaplamiussakaii Masumoto, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Malaymira Novák, 2020g: 56 [F]. Type species: Malaymirajenisi Novák, 2020, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Malayoscelis Schawaller, 2003: 198 [F]. Type species: Malayoscelisgebieni Schawaller, 2003, by original designation. Status: valid genus in Lagriinae: Pycnocerini.

Malaysphena Bečvář & Purchart, 2008: 38 [F]. Type species: Laosocryptobatesrotundipennis Kaszab 1960, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Mallogria Borchmann, 1936: 20, 165 [F]. Type species: Lagrialongipilis Fairmaire, 1875, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Mamorina Antoine, 1951: 98 [F]. Type species: Mamorinasulcaticeps Antoine, 1951, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Mantichorula Reitter, 1889a: 695 [F]. Type species: Mantichorulasemenowi Reitter, 1889, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Maracia Gebien, 1919: 27, 34 [F]. Type species: Camariafemoralis Kirsch, 1866, by subsequent designation (Gebien 1942a: 319). Status: valid genus in Stenochiinae: Cnodalonini.

Margus Dejean, 1834: 200 [M]. Type species: fixed herein (ICZN 1999, Article 70.3) as Colydiumcastaneum Herbst, 1797, misidentified as Tenebrioferrugineus Fabricius, 1781 in the original designation by monotypy in Dejean (1834). Status: junior synonym of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini. Synonymy: Guérin-Méneville (1846: cxvii). Note: the type species “Tenebrioferrugineus Fabricius” was first established by monotypy (ICZN 1999, Article 68.3); as noted by C.O. Waterhouse (1896: 230) and Blair (1913: 223) the Tenebrioferrugineus Fabricius of authors, including Stephens (1829: 19, as “ferruginea, Oliv.”), was misidentified; Blair (1913: 223) noted that the species to which the authors referred is in fact Colydiumcastaneum Herbst, 1797; we follow currently accepted concepts (e.g., Bousquet et al. 2018: 224) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Tenebrioferrugineus Fabricius, 1781 is a valid species in the genus Tribolioides Blair, 1913 [Coleoptera: Cucujidae].

Mariepskopia Schawaller, 2012c: 218 [F]. Type species: Mariepskopiaalbomaculata Schawaller, 2012, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Martianus Fairmaire, 1893d: 540 [M]. Type species: Martianuscastaneus Fairmaire, 1893 (= Palembusocularis Casey, 1891), by original designation. Status: junior synonym of Ulomoides Blackburn, 1888 in Diaperinae: Diaperini: Diaperina. Synonymy: Halstead (1975: 241, with Palembus Casey, 1891, a junior synonym of Ulomoides Blackburn, 1888).

Massadraamelia Mas-Peinado, Buckley, Ruiz & García-París, 2018: 541 [F]. Type species: Pimeliagranulithorax Escalera, 1914, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Mateuina Español, 1944: 22, 30 [F]. Type species: Mateuinakaszabi Español, 1944, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Matthewsotys Bouchard & Bousquet, new genus [M]. Type species: Otysarmatus Champion, 1895, by present designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: Matthews and Bouchard (2008: 333) selected Otysarmatus Champion, 1895 as the type species of Otys Champion, 1895 and treated it as a valid genus; however, an earlier type species designation by R. Lucas (1920: 468) confirms Otys Champion, 1895 as a junior synonym of Scaletomerus Blackburn, 1891; we hereby establish Matthewsotys as a new genus and refer to Matthews and Bouchard (2008: 205) for the character states that characterise and differentiate this taxon.

Mauritianopidium Dajoz, 1977: 244 [N]. Type species: Mauritianopidiumoculatum Dajoz, 1977, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Mayidicistela Pic, 1954: 259 [F]. Type species: Mayidicistelapaulostriata Pic, 1954, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Mechanetes C.O. Waterhouse, 1887: 448 [M]. Type species: Mechanetescornutus C.O. Waterhouse, 1887, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mecocerus Solier, 1835a: 241 [M]. Type species: Xystropusdejeanii Solier, 1835, by monotypy. Status: junior synonym of Prostenus Klug, 1829 in Alleculinae: Alleculini: Xystropodina. Synonymy: Lacordaire (1859b: 513). Note: junior homonym of Mecocerus Schönherr, 1833 [Coleoptera: Anthribidae].

Mecopisthopus Karsch, 1881: 46 [M]. Type species: Mecopisthopusrohlfsi Karsch, 1881, by original designation. Status: junior synonym of Leucolaephus P.H. Lucas, 1859 in Pimeliinae: Pimeliini. Synonymy: Semenov (1893: 261).

Mecysmus Horn, 1870: 349 [M]. Type species: Blapstinusangustus J.L. LeConte, 1851, by monotypy. Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Lumen et al. (2020: 342).

Mederis Motschulsky, 1872: 24 [M]. Type species: Upisangulata Erichson, 1842, by original designation. Status: junior synonym of Promethis Pascoe, 1869 in Stenochiinae: Cnodalonini. Synonymy: C.O. Waterhouse (1876: 288).

Medvedevia Chigray, 2019: 915 [F]. Type species: Medvedeviaglebi Chigray, 2019, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Medvedevoblaps Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Protoblapskashkarovi G.S. Medvedev, 1998, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina. Note: replacement name for Protoblaps G.S. Medvedev, 1998.

Megacantha Westwood, 1843: 121 [F]. Type species: Megacanthatenebrosa Westwood, 1843 (= Helopsdentatus Fabricius, 1801), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: genus also described as new by Westwood (1844: 228).

Megadasus Reitter, 1904: 146 [M]. Type species: Opatrumlefranci Fairmaire, 1863, by original designation. Status: junior synonym of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Gebien (1939: 443).

Megagenius Solier, 1835b: 512, 513 [M]. Type species: Megageniusfrioli Solier, 1835, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Megalophrys G.R. Waterhouse, 1845b: 321, 322 [F]. Type species: Megalophryspatagonica G.R. Waterhouse, 1845, by monotypy. Status: senior synonym of Peltolobus Lacordaire, 1859 in Pimeliinae: Trilobocarini. Note: although the older name Megalophrys Wagler, 1830 [Amphibia] was placed on the Official Index of Rejected and Invalid Generic Names in Zoology by the ICZN (1994b, Opinion 1763), it was not suppressed for the Principle of Homonymy and therefore remains available and the senior homonym.

Megaprosodes Reitter, 1909a: 118 [M]. Type species: Prosodesstriata Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Megasattus Casey, 1908: 56, 62 [M]. Type species: Eusattuserosus Horn, 1870, by original designation. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: Triplehorn (1968: 379).

Megascythis Keleinikova, 1963: 622 [F]. Type species: Megascythispanfilovi Keleinikova, 1963, by monotypy. Status: junior synonym of Scythis Kraatz, 1865 in Pimeliinae: Tentyriini. Synonymy: Skopin (1979: 171).

Megasida Casey, 1912: 77, 202 [F]. Type species: Asidaobliterata Champion, 1892, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Megatenebrio Gridelli, 1951: 221 [M]. Type species: Tenebriogiganteus Fairmaire, 1897, by monotypy. Status: valid subgenus of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini.

Megatlasion Koch, 1948: 427 [N]. Type species: Micrositusatlantis Escalera, 1914, by original designation. Status: junior synonym of Atlasion Koch, 1948 in Blaptinae: Dendarini: Melambiina. Synonymy: Antoine (1957: 351).

Megeleates Casey, 1895: 623 [M]. Type species: Megeleatessequoiarum Casey, 1895, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Megelenophorus Gebien, 1910a: 121 [M]. Type species [automatic]: Elenophorusamericanus Lacordaire, 1830, by monotypy. Status: valid genus in Pimeliinae: Elenophorini: Megelenophorina. Note: replacement name for Cacicus Dejean, 1834.

Megischia Solier, 1835a: 245, 247 [F]. Type species: Cistelacurvipes Brullé, 1832, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Megischina Reitter, 1906b: 118, 171 [F]. Type species: Cistelaarmillata Brullé, 1832, by subsequent designation (R. Lucas 1920: 399). Status: valid genus in Alleculinae: Cteniopodini.

Meglyphus Motschulsky, 1872: 38 [M]. Type species: Meglyphuslaenoides Motschulsky, 1872, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina. Note: we act as First Revisers and reject the alternative original spelling Megliphus, used by Motschulsky (1872: 41).

Meladeras Mulsant & Rey, 1854: 191, 219 [N]. Type species: Meladerasquadratulum Mulsant & Rey, 1854, by subsequent designation (Viñolas 1990: 61). Status: valid subgenus of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina.

Meladiesia Reitter, 1909b: 309 [F]. Type species: Meladiesiamiritarsis Reitter, 1909, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Meladocrates Reitter, 1904: 96 [M]. Type species: Olocratesplaniusculus Mulsant & Rey, 1854, by subsequent designation (Viñolas 1990: 60). Status: valid subgenus of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina.

Melambasida Reitter, 1917a: 10, 21 [F]. Type species: Alphasidainterjecta Reitter, 1917, by monotypy. Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Melambatlasus Koch, 1948: 433 [M]. Type species: Micrositushebes Antoine, 1933, by original designation. Status: valid subgenus of Melambius Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Melambiophylax Schuster, 1922: 48, 49 [M]. Type species: Phylaxsardous Baudi di Selve, 1876, by monotypy. Status: junior synonym of Allophylax Bedel, 1906 in Blaptinae: Dendarini: Melambiina. Synonymy: Leo (1994: 134).

Melambius Mulsant & Rey, 1854: 124 [M]. Type species: Opatrumbarbarum Erichson, 1841, by monotypy. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Melanastus Casey, 1907: 289, 353 [M]. Type species: Eurymetoponatrum J.L. LeConte, 1851, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Melancrus Reiche & Saulcy, 1857: 190 [N]. Type species: Melancruslaevigatum Reiche & Saulcy, 1857, by subsequent designation (Löbl et al. 2008a: 42). Status: junior synonym of Oxycara Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Lacordaire (1859a: 57). Note: authors have assigned a masculine gender for this name in the literature; however, the correct gender is neuter; Melancrus has two roots: melan- and -crus, the first root is clearly Greek: melas, melaina, melan (an irregular adjective meaning dark or black) while the second is Latin: crus (the name for a leg, shank, or shin); the second word is a neuter noun and therefore the gender of Melancrus is neuter (ICZN 1999, Article 30.1.1).

Melaneleodes Blaisdell, 1909: 36 [M]. Type species: Blapscarbonaria Say, 1824, by subsequent designation (Triplehorn and Thomas 2012: 254). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Melanesthes Dejean, 1834: 191 [F]. Type species: Opatrumsibiricum Faldermann, 1833, by subsequent designation (Gebien 1939: 462). Status: valid genus and subgenus in Blaptinae: Opatrini: Opatrina.

Melanimon Motschulsky, 1845a: 78 [N]. Type species: Microzoumcollare Motschulsky, 1839, by monotypy. Status: senior synonym of Platynosum Mulsant & Rey, 1859 in Blaptinae: Opatrini: Sclerina. Synonymy: Seidlitz (1894: 413, 454). Note: junior homonym of Melanimon Steven, 1828 [Coleoptera: Tenebrionidae: Tenebrioninae: Melanimonini].

Melanimon Steven, 1828: 98 [N]. Type species: Opatrumtibiale Fabricius, 1781, by monotypy. Status: valid genus in Tenebrioninae: Melanimonini.

Melanochrus Wollaston, 1864: 467 [M]. Type species: Melanochruslacordairii Wollaston, 1864, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Melanocoma Wollaston, 1868: 181 [F]. Type species: Melanocomavestita Wollaston, 1868, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Melanocratus Fairmaire, 1895a: 21 [M]. Type species: Melanocratusvalidipes Fairmaire, 1895, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Melanolophus Fairmaire in Fairmaire et al., 1882: 69 [M]. Type species: Melanolophusseptemcostatus Fairmaire, 1882, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Melanophorus Lacordaire, 1859a: 74 [M]. Type species [automatic]: Melaphorusreichii Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Melaphorus Guérin-Méneville, 1834 in Pimeliinae: Evaniosomini. Note: unjustified emendation of Melaphorus Guérin-Méneville, 1834, not in prevailing usage.

Melanopterus Mulsant & Rey, 1854: 13, 14 [M]. Type species: Melanopterusporcatus Mulsant & Rey, 1854, by subsequent designation (Gebien 1938a: 292). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Melanostola Solier, 1836: 123 [F]. Type species: Pimeliasimplex Solier, 1836, by monotypy. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Note: the name Melanostola was listed as synonym of Pimelia Fabricius, 1775 by Solier (1836; published by 16 May), being treated before 1961 as an available name and adopted as the name of a taxon (e.g., Dejean 1836 (published by 30 July), Hope 1841: 118); Melanostola was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Melansis Wollaston, 1864: 491 [F]. Type species: Phylaxcostatus Brullé, 1839, by subsequent designation (Dallas 1865: 401). Status: valid genus in Blaptinae: Dendarini: Melambiina.

Melaphorus Guérin-Méneville, 1834: 13 [M]. Type species: Melaphorusreichii Guérin-Méneville, 1834, by monotypy. Status: valid genus in Pimeliinae: Evaniosomini.

Melaps Carter, 1908b: 409 [M]. Type species: Melapscisteloides Carter, 1908, by monotypy. Status: junior synonym of Oocistela Borchmann, 1908 in Alleculinae: Alleculini: Alleculina. Synonymy: Carter (1915a: 78). Note: although this name has been used as valid, with Oocistela Borchmann, 1908 as a synonym, in recent literature (e.g., Matthews and Lawrence 2019: 645), data on the date of publication of both names reveals that Oocistela was published first and has priority over Melaps.

Melarachnica Kraatz, 1865: 80, 174 [F]. Type species: Melarachnicawestermanni Kraatz, 1865 (= Microderacoromandelensis Solier, 1835), by monotypy. Status: junior synonym of Nemapus Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Koch (1943b: 790).

Melasia Perroud & Mulsant, 1856: 160 [F]. Type species: Melasiagagatina Perroud & Mulsant, 1856 (= Tenebrioculinaris Linnaeus, 1758), by subsequent designation (G.S. Medvedev 1990: 227). Status: junior synonym of Uloma Dejean, 1821 in Tenebrioninae: Ulomini. Synonymy: Lacordaire (1859a: 332).

Melasma Wollaston, 1864: 484 [N]. Type species: Phylaxlineatus Brullé, 1838, by monotypy. Status: senior synonym of Melasmana Strand, 1935 in Blaptinae: Dendarini: Melambiina. Note: junior homonym of Melasma Adams, 1854 [Mollusca].

Melasmana Strand, 1935a: 297 [F]. Type species [automatic]: Phylaxlineatus Brullé, 1838, by monotypy. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina. Note: replacement name for Melasma Wollaston, 1864.

Melasmocara Reitter, 1900c: 95 [N]. Type species: Melanochruslacordairii Wollaston, 1864, by subsequent monotypy (Löbl et al. 2008a: 42). Status: junior synonym of Melanochrus Wollaston, 1864 in Pimeliinae: Tentyriini. Synonymy: Jakobson (1924: 243), Löbl et al. (2008a: 42). Note: originally proposed without included nominal species; Löbl et al. (2008a: 42), by including the species Melanochruslacordairii Wollaston, 1864 in association with this name, were the first authors to subsequently and expressly include nominal species in Melasmocara (ICZN 1999, Article 67.2.2).

Melaxumia Reitter, 1895: 280 [F]. Type species: Melaxumiaacutangula Reitter, 1895 (= Tentyriaangulosa Gebler, 1832), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Melobates Kaszab, 1941a: 4, 23 [M]. Type species: Melobatesbiroi Kaszab, 1941, by original designation. Status: junior synonym of Rehumius Fairmaire, 1893 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983d: 84).

Melobrachys Kaszab, 1960b: 273 [M]. Type species: Melobrachyssarawakensis Kaszab, 1960, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Menandris Haag-Rutenberg, 1878: 103 [F]. Type species: Menandrisaenea Haag-Rutenberg, 1878, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mencheres Champion, 1884: 5 [M]. Type species: Mencheresnicaraguensis Champion, 1884, by subsequent designation (R. Lucas 1920: 403). Status: valid genus in Pimeliinae: Edrotini.

Menearchus Carter, 1920a: 229 [M]. Type species: Menearchusimpressosulcatus Carter, 1920 (= Helopsnigritus Fabricius, 1777), by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina. Note: as mentioned by Matthews and Bouchard (2008: 351) the type species was originally described from Australia in error, as the genus Menearchus Carter, 1920 is endemic to India and Sri Lanka.

Menechides Motschulsky, 1872: 26 [M]. Type species: Helopscalcaratus Fabricius, 1798, by original designation. Status: valid subgenus of Centronopus Solier, 1848 in Tenebrioninae: Centronopini.

Menederes Solier, 1848: 152, 203 [M]. Type species: Menederesrufilabris Solier, 1848, by original designation. Status: valid genus and subgenus in Blaptinae: Platynotini: Eurynotina.

Menederopsis Koch, 1954a: 16 [F]. Type species: Menederopsisconstricta Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Menedrio Motschulsky, 1872: 27 [M]. Type species: Tenebrioobscurus Fabricius, 1792, by original designation. Status: junior synonym of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini. Synonymy: Heyden in Heyden et al. (1883: 134).

Menephilus Mulsant, 1854: 291 [M]. Type species: Tenebriocurvipes Fabricius, 1792 (= Tenebriocylindricus Herbst, 1784), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Meneristes Pascoe, 1869: 150 [M]. Type species: Meneristeslaticollis Pascoe, 1869 (= Tenebrioaustralis Boisduval, 1835), by original designation. Status: valid genus in Tenebrioninae: Heleini: Asphalina.

Menes Champion, 1888: 442 [M]. Type species: Menesmeridanus Champion, 1888, by subsequent designation (Bousquet et al. 2015: 134). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Menimoides Kaszab, 1946b: 19 [M]. Type species: Menimoidestarandus Kaszab, 1946, by original designation. Status: junior synonym of Micropeneta Pic, 1921 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Kaszab (1978a: 172).

Menimopsis Champion, 1896: 16 [F]. Type species: Menimopsisexcaecus Champion, 1896, by monotypy. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Menimus Sharp, 1876: 73 [M]. Type species: Menimusbatesi Sharp, 1876, by subsequent designation (Gebien 1940: 429). Status: valid genus and subgenus in Diaperinae: Gnathidiini: Gnathidiina.

Meniscophorus Champion, 1889: 64 [M]. Type species: Meniscophorusamazonicus Champion, 1889, by subsequent designation (R. Lucas 1920: 404). Status: valid genus in Lagriinae: Lagriini: Statirina.

Menoeceus Champion, 1888: 443 [M]. Type species: Menoeceuscrassicornis Champion, 1888, by subsequent designation (Casey 1891: 122). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Menoncotus Koch, 1954a: 34 [M]. Type species: Oncotuschaetotaxicus Koch, 1954, by original designation. Status: valid subgenus of Oncotus Blanchard, 1845 in Blaptinae: Platynotini: Eurynotina.

Mentariobius Koch, 1948: 425 [M]. Type species: Micrositusdistinguendus Mulsant & Rey, 1854, by original designation. Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Mentes Champion, 1893a: 559 [M]. Type species: Mentesruficollis Champion, 1893, by subsequent designation (R. Lucas 1920: 404). Status: valid genus in Stenochiinae: Stenochiini.

Meracantha W. Kirby, 1837: 237 [F]. Type species: Meracanthacanadensis W. Kirby, 1837 (= Helopscontractus Palisot de Beauvois, 1811), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Meracanthoides Linell, 1896: 698 [M]. Type species: Meracanthoidescupreolineatus Linell, 1896, by monotypy. Status: valid subgenus of Paramarygmus Quedenfeldt, 1885 in Tenebrioninae: Amarygmini.

Merinus J.L. LeConte, 1862: 230 [M]. Type species: Tenebriolaevis G.-A. Olivier, 1795, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: the original combination of the name of the type species, Tenebriolaevis G.-A. Olivier, 1795, is a junior primary homonym of Tenebriolaevis Forskål, 1775.

Merklia Chen, 1997: 307 [F]. Type species: Merkliabimaculata Chen, 1997, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Meropersina Reitter, 1909a: 117 [F]. Type species: Prosodescordicollis Allard, 1884, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Meropria Borchmann, 1921: 217, 228 [F]. Type species: Statiraglabrata Mäklin, 1863, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Merotemnus Horn, 1870: 364, 367 [M]. Type species: Merotemnuselongatus Horn, 1870 (= Ulomafiliforme Laporte, 1840), by monotypy. Status: junior synonym of Adelonia Laporte, 1840 in Lagriinae: Belopini. Synonymy: Spilman (1961: 49).

Meroxys Ardoin, 1963a: 132 [M]. Type species: Meroxyshaafi Ardoin, 1963, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: name first proposed by Ardoin (1962b: 970) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Mesabates Champion, 1884: 3 [M]. Type species: Mesabateslatifrons Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Mesabatodes Casey, 1907: 517 [M]. Type species: Mesabatesinaequalis Champion, 1892, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Mesoblaps Bauer, 1921: 231 [F]. Type species: Blaps rugulipennis Fairmaire, 1891, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Mesohelops Reitter, 1922a: 31 [M]. Type species: Helopscyanipes Allard, 1877, by subsequent designation (Nabozhenko and Löbl 2008: 251). Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko and Keskin (2017: 41). Note: the original combination of the name of the type species, Helopscyanipes Allard, 1877, is a junior primary homonym of Helopscyanipes Fabricius, 1801.

Mesoleptodes G.S. Medvedev & Iljina, 2007: 881 [M]. Type species: Leptodessemenowi Reitter, 1892, by original designation. Status: valid subgenus of Leptodes Dejean, 1834 in Pimeliinae: Leptodini.

Mesolobopoda Campbell, 1966: 34 [F]. Type species: Alleculasocia J.L. LeConte, 1854, by original designation. Status: valid subgenus of Lobopoda Solier, 1835 in Alleculinae: Alleculini: Alleculina.

Mesomorphus Miedel, 1880: 140 [M]. Type species: Opatrummurinum Baudi di Selve, 1876 (= Opatrinussetosus Mulsant & Rey, 1853), by subsequent designation (Gebien 1938a: 399). Status: valid genus in Blaptinae: Opatrini: Opatrina.

Mesopatrum Broun, 1893b: 1355 [N]. Type species: Mesopatrumgranulosum Broun, 1893, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Mesopraocis Flores & Pizarro-Araya, 2014: 60 [M]. Type species: Praociscalderanus Kulzer, 1958, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 12, 31) without type species designation.

Mesoprosodes G.S. Medvedev, 1995b: 827 [M]. Type species: Prosodesnuratensis Semenov, 1894, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Mesopterocoma Skopin, 1974b: 147 [F]. Type species: Pterocomasemicarinata Bates, 1879, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Mesostena Eschscholtz, 1831: 5, 9 [F]. Type species: Mesostenapunctata Eschscholtz, 1831 (= Pimeliaangustata Fabricius, 1775), by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Mesostenopa Kraatz, 1865: 80, 179 [F]. Type species: Mesostenopapicea Kraatz, 1865, by subsequent designation (Gebien 1937a: 608). Status: valid subgenus of Mesostena Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Mesosternoplax Skopin, 1973: 110, 142 [F]. Type species: Trigonoscelislaeviuscula Kraatz, 1882, by original designation. Status: valid subgenus of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini.

Mesotretis Bates, 1872c: 151 [F]. Type species: Mesotretisferruginea Bates, 1872, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Messalia Pascoe, 1883: 442 [F]. Type species: Messaliavarians Pascoe, 1883, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Messoricolum Koch, 1960: 384 [N]. Type species: Messoricolumscotti Koch, 1960, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Metablapylis Blaisdell, 1909: 391 [F]. Type species: Eleodesnigrina J.L. LeConte, 1858, by subsequent designation (Bousquet et al. 2018: 159). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Metabolocerus Bates, 1873c: 259 [M]. Type species: Metaboloceruspilosus Bates, 1873, by subsequent designation (R. Lucas 1920: 407). Status: valid genus in Tenebrioninae: Ulomini.

Metaclisa Jacquelin du Val, 1861: 296 [F]. Type species: Platydemaparallela Fairmaire, 1855 (= Diaperisazurea Waltl, 1838), by original designation. Status: valid genus in Tenebrioninae: Metaclisini. Note: nomen protectum (see Bouchard et al. 2007: 393).

Metacorticeus Bremer & Lillig, 2017a: 71 [M]. Type species: Corticeusornatus Bremer, 1993, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Metacossyphodes Andreae, 1961: 205, 215 [M]. Type species: Cossyphodeskundelunguensis Basilewsky, 1950, by original designation. Status: junior synonym of Cossyphodes Westwood, 1851 in Pimeliinae: Cossyphodini: Cossyphodina. Synonymy: Schawaller (2013c: 362, implied by inclusion of Cossyphodeskundelunguensis Basilewsky, 1950 in Cossyphodes Westwood, 1851 without use of a subgenus rank).

Metallonotus Gray in Griffith and Pidgeon, 1832: 790 [M]. Type species: Metallonotusdenticollis Gray, 1832, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Methistamena Gebien, 1919: 28, 151 [F]. Type species: Methistamenaclavipes Gebien, 1919, by original designation. Status: junior synonym of Camariomorpha Pic, 1915 in Stenochiinae: Cnodalonini. Synonymy: Kulzer (1954a: 71).

Metisopus Bates, 1873e: 371 [M]. Type species: Metisopuspurpureipennis Bates, 1873, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Metistete Pascoe, 1866a: 489 [F]. Type species: Tanychilusgibbicollis Newman, 1838, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Metonites Gistel, 1848a: viii, 126 [M]. Type species [automatic]: Blapidaokeni Perty, 1830, by monotypy. Status: junior synonym of Blapida Perty, 1830 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Blapida Perty, 1830.

Metopoloba Casey, 1907: 379 [F]. Type species: Epitraguspruinosus Horn, 1870, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Metoponiopsis Casey, 1907: 290 [F]. Type species: Eurymetoponbicolor Horn, 1870, by monotypy. Status: valid subgenus of Metoponium Casey, 1907 in Pimeliinae: Edrotini.

Metoponium Casey, 1907: 288 [N]. Type species: Eurymetoponabnorme J.L. LeConte, 1851, by original designation. Status: valid genus and subgenus in Pimeliinae: Edrotini.

Metriolagria Merkl, 1987: 124, 138 [F]. Type species: Lagriaaffinis Boisduval, 1835, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Metriopus Solier, 1835b: 512, 570 [M]. Type species: Metriopushoffmanseggii Solier, 1835, by monotypy. Status: valid genus and subgenus in Pimeliinae: Adesmiini.

Metulosonia Bates, 1873c: 261 [F]. Type species: Metulosoniahorni Bates, 1873, by subsequent designation (Gebien 1940: 1061). Status: valid genus in Tenebrioninae: Triboliini.

Micipsa P.H. Lucas, 1855: xxxiv [M]. Type species: Micipsarufitarsis P.H. Lucas, 1855 (= Pimeliamulsanti Levrat, 1853), by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Micipsina Reitter, 1900c: 94, 188 [F]. Type species: Micipsinarolphi Reitter, 1900, by monotypy. Status: junior synonym of Thalpobia Fairmaire, 1871 in Pimeliinae: Tentyriini. Synonymy: Escalera (1914: 279).

Micrantereus Solier, 1848: 151, 175 [M]. Type species: Acanthomerusanomalus Guérin-Méneville, 1834, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Micrarmalia Casey, 1907: 516 [F]. Type species: Emmenastusconstrictus Champion, 1892, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Micrasida Smith, 2013: 608 [F]. Type species: Micrasidaobrienorum Smith, 2013, by original designation. Status: valid genus in Pimeliinae: Asidini.

Micrectyche Bates, 1873e: 362 [F]. Type species: Micrectycheintermedia Bates, 1873, by subsequent designation (R. Lucas 1920: 412). Status: valid genus in Diaperinae: Ectychini.

Micreuphlaeus Fairmaire, 1897e: 223 [M]. Type species: Micreuphlaeusasperipellis Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Micrisomira Pic, 1930a: 30 [F]. Type species: Micrisomiraruficollis Pic, 1930, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Microamarygmus Pic, 1915d: 8 [M]. Type species: Microamarygmusmadurensis Pic, 1915, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Microanaedus Pic, 1923b: 16 [M]. Type species: Microanaedusnotatus Pic, 1923, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Microatasthalus Ando, 2010: 153 [M]. Type species: Microatasthalushadrocerus Ando, 2010, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Microbasanus Pic, 1921a: 1 [M]. Type species: Microbasanusjureceki Pic, 1921, by monotypy. Status: junior synonym of Scaphidema Redtenbacher, 1848 in Diaperinae: Scaphidemini. Synonymy: Löbl et al. (2008b: 318), see also Schawaller (2008: 384).

Microblattellus Ferrer, 2006a: 78 [M]. Type species: Microblattelluslecongmani Ferrer, 2006, by original designation. Status: valid genus in Tenebrioninae: Falsocossyphini.

Microblemma Semenov, 1889: 213 [N]. Type species: Microblemmasimplex Semenov, 1889, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Platamodina.

Microbolitonaeus Grimm, 2014: 194 [M]. Type species: Microbolitonaeusarmatus Grimm, 2014, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Microbradymerus Schawaller, 1999a: 144 [M]. Type species: Microbradymerusmerkli Schawaller, 1999, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Microcalcar Pic, 1925b: 9 [N]. Type species: Belopusinstriatus Pic, 1925, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Microcalydonis Pic, 1923d: 17 [F]. Type species: Microcalydonismetallica Pic, 1923, by monotypy. Status: valid subgenus of Osternus Fairmaire, 1895 in Stenochiinae: Cnodalonini.

Microcameria Ren, 1998: 108, 113 [F]. Type species: Microcameriapygmaea Ren, 1998, by original designation. Status: junior synonym of Foochounus Pic, 1921 in Stenochiinae: Cnodalonini. Synonymy: Ando (2008: 39).

Microcatomus Pic, 1925b: 7 [M]. Type species: Microcatomuslongipilis Pic, 1925, by monotypy. Status: valid genus in Tenebrioninae: Helopini: incertae sedis. Note: Nabozhenko (2018: 183) mentioned that the position of this genus within the tribe Helopini is unclear.

Microcenoscelis Schawaller, 2015: 438 [F]. Type species: Microcenosceliscaeca Schawaller, 2015, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Microcilibe Carter, 1919: 147 [F]. Type species: Microcilibecastanea Carter, 1919, by monotypy. Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Kaszab (1978a: 173).

Microcistela Pic, 1904: 26 [F]. Type species: Microcistelarosinae Pic, 1904, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Microcistela Pic, 1919b: 6 [F]. Type species: Microcistelaobscura Pic, 1919, by monotypy. Status: senior synonym of Microcistelopsis Pic, 1922 in Alleculinae: Alleculini: Alleculina. Note: junior homonym of Microcistela Pic, 1904 [Coleoptera: Tenebrionidae: Alleculinae: Alleculini: Gonoderina].

Microcistelopsis Pic, 1922d: 20 [F]. Type species [automatic]: Microcistelaobscura Pic, 1919, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: replacement name for Microcistela Pic, 1919.

Microcrypticus Gebien, 1921b: 7 [M]. Type species: Diaperisvariegata Klug, 1833, by original designation. Status: valid genus and subgenus in Diaperinae: Crypticini.

Microdendarus Escalera, 1944: 84 [M]. Type species: Dendarusschusteri Español, 1937, by original designation. Status: junior synonym of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina. Synonymy: Español (1961a: 44).

Microdera Eschscholtz, 1831: 5, 6 [F]. Type species: Tentyriadeserta Tauscher, 1812, by subsequent designation (Gebien 1937a: 619). Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Microderopsis Haag-Rutenberg, 1876: 86 [F]. Type species: Microderopsisbenguelensis Haag-Rutenberg, 1876, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Microdisema Pic, 1917b: 2, 3 [F]. Type species: Disemagounellei Pic, 1912, by monotypy. Status: valid subgenus of Barsenis Pascoe, 1887 in Lagriinae: Lagriini: Statirina. Note: this genus was established without included nominal species; Disemagounellei Pic, 1912 was first and expressly included in Microdisema by Pic (1937c: 36).

Microdocnemis Nabozhenko & Keskin, 2010: 841 [F]. Type species: Microdocnemisxerophilica Nabozhenko & Keskin, 2010, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Microeucyrtus Pic, 1926a: 47 [M]. Type species: Microeucyrtuscoomani Pic, 1926, by monotypy. Status: junior synonym of Eucyrtus Lacordaire, 1859 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1941: 1139).

Microgauromaia Pic, 1921d: 23 [F]. Type species: Microgauromaiaminuta Pic, 1921, by monotypy. Status: junior synonym of Phaedis Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Ando (2007: 164).

Microgoniadera Pic, 1917d: 9 [F]. Type species: Microgoniaderahirsuta Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Microgonocnemis Pic, 1936b: 16 [F]. Type species: Microgonocnemiscarinata Pic, 1936, by monotypy. Status: valid subgenus of Paragonocnemis Kraatz, 1899 in Tenebrioninae: Amarygmini.

Microhedyphanes Nabozhenko & Lillig, 2013: 189 [M]. Type species: Hedyphaneschikatunovi Nabozhenko & Lillig, 2013, by original designation. Status: valid subgenus of Hedyphanes Fischer, 1820 in Tenebrioninae: Helopini: Helopina.

Microhemicera Pic, 1921d: 30 [F]. Type species: Hemicerahumeralis Pic, 1921, by monotypy. Status: junior synonym of Simalura Gebien, 1914 in Stenochiinae: Cnodalonini. Synonymy: Löbl et al. (2008b: 347).

Microhionthis Blair, 1923b: 120 [F]. Type species: Microhionthispatriciae Blair, 1923 (= Falsocatomuluseuphraticus Pic, 1914), by monotypy. Status: junior synonym of Falsocatomulus Pic, 1914 in Pimeliinae: Tentyriini. Synonymy: Koch (1941: 267).

Microlagria Seidlitz, 1898b: 336, 339 [F]. Type species: Lagriapoupillieri Reiche, 1864, by monotypy. Status: junior synonym of Adynata Fåhraeus, 1870 in Lagriinae: Lagriini: Lagriina. Synonymy: Borchmann (1936: 23).

Microleichenum G.S. Medvedev, 1973: 648 [N]. Type species: Microleichenumchoresmense G.S. Medvedev, 1973, by original designation. Status: junior synonym of Apsheronellus Bogatchev, 1967 in Blaptinae: Pedinini: Leichenina. Synonymy: G.S. Medvedev and Iwan (2006: 614).

Microlyprops Kaszab, 1939a: 108 [M]. Type species: Microlypropsceylonicus Kaszab, 1939, by original designation. Status: valid genus in Lagriinae: Goniaderini.

Micromenandris Kaszab, 1955a: 511, 513 [F]. Type species: Micromenandrismirabilis Kaszab, 1955, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Micromes Casey, 1907: 432, 441 [M]. Type species: Stibiaovipennis Horn, 1874, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Micromophlus Znojko in Ogloblin and Znojko, 1950: 124 [M]. Type species: Omophlussubtilis Solsky, 1881, by monotypy. Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini.

Micronilio Pic, 1936c: 198 [M]. Type species: Niliopunctatus Pic, 1918 (= Niliogounellei Ihering, 1914), by monotypy. Status: valid subgenus of Nilio Latreille, 1802 in Nilioninae.

Microomopheres Freude, 1993: 214 [M]. Type species: Omopheresbrendelli Freude, 1993, by monotypy. Status: valid subgenus of Omopheres Casey, 1907 in Pimeliinae: Epitragini.

Micropedinus Lewis, 1894: 379 [M]. Type species: Micropedinusalgae Lewis, 1894 (= Heterophagapullulus Boheman, 1858), by subsequent designation (R. Lucas 1920: 415). Status: valid genus in Lagriinae: Lupropini.

Micropeltoides Pic, 1916d: 16 [M]. Type species: Micropeltoidescrypticoides Pic, 1916, by monotypy. Status: valid subgenus of Peltoides Laporte, 1833 in Tenebrioninae: Alphitobiini.

Micropeneta Pic, 1921d: 19 [F]. Type species: Micropenetatestacea Pic, 1921, by subsequent designation (Riley 1923: 128). Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Microphenus Gebien, 1921a: 324, 338 [M]. Type species: Microphenuscordicollis Gebien, 1921 (= Espitesobscurus Blair, 1915), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Microphligra Koch, 1955a: 47 [F]. Type species: Phligraminuta Péringuey, 1904, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Microphyes W.J. MacLeay, 1872: 286 [M]. Type species: Microphyesrufipes W.J. MacLeay, 1872 (= Opatrumlaevigatum Fabricius, 1781), by monotypy. Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Blair (1914: 486).

Microphylacinus Iwan, Kamiński & Aalbu, 2011: 2 [M]. Type species: Microphylacinusverendus Iwan, Kamiński & Aalbu, 2011, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Microplatyscelis Kaszab, 1940a: 142, 144 [F]. Type species: Faustiaseriepunctata Reitter, 1890, by original designation. Status: valid genus in Blaptinae: Platyscelidini.

Microprostenus Pic, 1921d: 14 [M]. Type species: Microprostenuslongicornis Pic, 1921, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Micropseudochillus Fouquè, 2015: 230, 240 [M]. Type species: Pseudochilluspalawanus Fouquè, 2015, by original designation. Status: valid subgenus of Pseudochillus Fouquè, 2015 in Pimeliinae: Stenosini: Dichillina.

Microschatia Solier, 1836: 406, 474 [F]. Type species: Microschatiapunctata Solier, 1836, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Microselinus Koch, 1956a: 214 [M]. Type species: Microselinusmuelleri Koch, 1956, by original designation. Status: junior synonym of Glyptopteryx Gebien, 1910 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 91).

Microsis Koch, 1958: 76, 82 [F]. Type species: Microsisvilhenai Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Micrositus Mulsant & Rey, 1854: 131, 148 [M]. Type species: Micrositusorbicularis Mulsant & Rey, 1854, by subsequent designation (Gebien 1938a: 415). Status: valid genus in Blaptinae: Dendarini: Dendarina.

Microsphaerotus Pic, 1928b: 10 [M]. Type species: Microsphaerotusruficornis Pic, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Microstenogena Pic, 1924a: 30 [F]. Type species: Microstenogenaruficornis Pic, 1924, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Microsthes Novák, 2011: 322 [M]. Type species: Microsthesbarborae Novák, 2011, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Microstizopus Koch, 1963: 36 [M]. Type species: Microstizopusciliatus Koch, 1963, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Microstrongylium Pic, 1917d: 13 [N]. Type species: Microstrongyliumcyanicolle Pic, 1917, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Kaszab (1977b: 30).

Microtelopsis Koch, 1940b: 742 [F]. Type species: Tetranillussimplicifrons Gridelli, 1934, by monotypy. Status: valid genus and subgenus in Pimeliinae: Stenosini: Stenosina. Note: we act as First Revisers and select Microtelopsis Koch, 1940 as the valid name for this genus instead of Extetranosis Koch, 1940 and Hypermicrotelopsis Koch, 1940.

Microtelus Solier, 1838b: 7, 9 [M]. Type species: Microtelusasiaticus Solier, 1838, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Microthelecta Pic, 1952a: 65 [F]. Type species: Cylindrothorusbraunsi Pic, 1952, by monotypy. Status: valid subgenus of Cylindrothorus Solier, 1843 in Alleculinae: Alleculini: Mycetocharina.

Microtocerus Pic, 1918a: 11 [M]. Type species: Microtocerusgrandicornis Pic, 1918, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Microzoon Hope, 1841: 112 [N]. Type species [automatic]: Opatrumtibiale Fabricius, 1781, by monotypy. Status: junior synonym of Melanimon Steven, 1828 in Tenebrioninae: Melanimonini. Note: unjustified emendation of Microzoum Dejean, 1834, not in prevailing usage.

Microzophobas Pic, 1944: 7 [M]. Type species: Microzophobasluteomaculatus Pic, 1944, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Microzoum Dejean, 1834: 193 [N]. Type species: Opatrumtibiale Fabricius, 1781, by monotypy. Status: junior synonym of Melanimon Steven, 1828 in Tenebrioninae: Melanimonini. Synonymy: Seidlitz (1894: 452).

Mictopsis Fairmaire, 1899e: 538 [F]. Type species: Mictopsislaticollis Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Miglica Reitter, 1904: 171 [F]. Type species: Pedinuslaticollis Gebler, 1830, by subsequent designation (Iwan and Löbl 2008: 267). Status: junior synonym of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina. Synonymy: Reichardt (1936: 175).

Millotella Koch, 1962a: 116, 130 [F]. Type species: Millotellamicrocornis Koch, 1962, by original designation. Status: senior synonym of Kochotella Bouchard & Bousquet, nom. nov. in Pimeliinae: Asidini. Note: junior homonym of Millotella Poisson, 1949 [Hemiptera].

Millstreamia G.S. Medvedev & Lawrence, 1984: 579 [F]. Type species: Csiroparodoxa G.S. Medvedev & Lawrence, 1984, by original designation. Status: valid subgenus of Csiro G.S. Medvedev & Lawrence, 1984 in Diaperinae: Hyociini: Hyociina.

Miltoprepes Gerstaecker, 1871: 65 [M]. Type species: Miltoprepeslaetus Gerstaecker, 1871, by monotypy. Status: valid genus in Tenebrioninae: Praeugenini.

Mimelasida Reitter, 1917a: 10, 21 [F]. Type species: Asidapuncticollis Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Mimoborchmania Pic, 1934a: 31 [F]. Type species: Nemostiracoloripes Pic, 1922, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Mimocellus Wasmann, 1904: 11 [M]. Type species: Mimocellustrechoides Wasmann, 1904, by subsequent designation (R. Lucas 1920: 419). Status: valid genus in Lagriinae: Lupropini.

Mimocistela Borchmann, 1938: 124 [F]. Type species: Mimocistelazumpti Borchmann, 1938, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Mimocossyphus Pic, 1923a: 5 [M]. Type species: Mimocossyphusminor Pic, 1923, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: incertae sedis. Note: Schawaller (2013c: 362) mentioned that the position of this taxon within Cossyphodini is doubtful.

Mimogoueum Pic, 1952d: 10 [N]. Type species: Mimogoueumhutteli Pic, 1952, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Ardoin (1962a: 64).

†Mimohelops Haupt, 1950: 114, 130 [M]. Type species: Mimohelopsvenosus Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Mimolaena Ardoin, 1961c: 36 [F]. Type species: Mimolaenapauliani Ardoin, 1961, by monotypy. Status: valid genus in Lagriinae: Laenini.

Mimolagria Pic, 1927a: 44 [F]. Type species: Mimolagriabruchi Pic, 1927, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Mimopeus Pascoe, 1866a: 477 [M]. Type species: Mimopeusamaroides Pascoe, 1866 (= Celibeelongata Brême, 1842), by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina. Note: as mentioned by Matthews and Bouchard (2008: 350) the type species was originally described from Australia in error, the genus Mimopeus Pascoe, 1866 is endemic to New Zealand; Matthews and Lawrence (2019: 629) mentioned that this genus seems indistinguishable from the Australian genus Celibe Boisduval, 1835.

Mimopraogena Pic, 1952d: 10 [F]. Type species: Alleculametallicipennis Pic, 1942, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Mimosynopticus Pic, 1922d: 25 [M]. Type species: Mimosynopticusparvulus Pic, 1922, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Mimothydemus Pic, 1923c: 20 [M]. Type species: Mimothydemusangustatus Pic, 1923 (= Lophocnemiscyaneus Kraatz, 1880), by monotypy. Status: junior synonym of Lophocnemis Mäklin, 1867 in Stenochiinae: Stenochiini. Synonymy: Kulzer (1966: 352).

Mimoxenotermes Pic, 1931a: 106 [M]. Type species: Mimoxenotermesduporti Pic, 1931, by monotypy. Status: valid genus in Tenebrioninae: Rhysopaussini.

Mimuroplatopsis Borchmann, 1936: 485 [F]. Type species: Mimuroplatopsisatricolor Pic, 1931, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: this genus was proposed earlier by Pic (1931b: 32) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Minasius Pic, 1932: 18 [M]. Type species: Minasiusopacipennis Pic, 1932, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Minorus Mulsant & Rey, 1854: 41 [M]. Type species: Eurynotusrugicollis Mulsant & Rey, 1854, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

†Miostenosis Wickham,1913: 297 [F]. Type species: Miostenosislacordairei Wickham, 1913, by original designation. Status: valid genus in Pimeliinae: Stenosini: incertae sedis. Note: described from Upper Eocene deposits (USA).

Miotodera Fairmaire, 1901b: 190 [F]. Type species: Miotoderafuneraria Fairmaire, 1901, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Mireanopidium Dajoz, 1977: 240 [N]. Type species: Mireanopidiumcamerunense Dajoz, 1977, by original designation. Status: valid genus in Diaperinae: Gnathidiini: incertae sedis. Note: placed in “Gnathidiini incertae sedis” by Schawaller and Purchart (2012: 312).

Miripronotum Louw, 1979: 117, 118 [N]. Type species: Miripronotumprominoculatum Louw, 1979, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Misolampidius Solsky, 1876: 292 [M]. Type species: Misolampidiustentyrioides Solsky, 1876, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Misolampomorphus Kaszab, 1941a: 2, 6 [M]. Type species: Misolampomorphuskochi Kaszab, 1941 (= Leprocaulusreitteri Pic, 1934), by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Misolampus Latreille, 1806: 160 [M]. Type species: Misolampushoffmannseggii Latreille, 1806 (= Pimeliagibbula Herbst, 1799), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mithippia Pascoe, 1869: 288, 293 [F]. Type species: Mithippiaaurita Pascoe, 1869, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Mitotagenia Reitter, 1916d: 138, 153 [F]. Type species: Stenosisarabs Baudi di Selve, 1881, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Mitragardhus Koch, 1956a: 379 [M]. Type species: Tragardhusnodosus Koch, 1956, by monotypy. Status: valid subgenus of Tragardhus Koch, 1956 in Blaptinae: Dendarini: Melambiina. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999, Article 13.4).

Mitragenius Solier, 1836: 307, 328 [M]. Type species: Mitrageniusdejeanii Solier, 1836, by monotypy. Status: valid genus in Pimeliinae: Nycteliini. Note: the alternative original spelling Mitrogenius, used by Solier (1836: 330), was rejected by Solier (1838a: 487) who acted as First Reviser (ICZN 1999, Article 24.2.4).

Mitrephorus Carter, 1913a: 83 [M]. Type species: Mitrephorusconvexicollis Carter, 1913, by monotypy. Status: senior synonym of Mitrothorax Carter, 1914 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1914b: 78, with Ctimene Bates, 1873, a senior synonym of Mitrothorax Carter, 1914). Note: junior homonym of Mitrephorus Schönherr, 1837 [Coleoptera: Curculionidae].

Mitrothorax Carter, 1914b: 78 [M]. Type species [automatic]: Mitrephorusconvexicollis Carter, 1913, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: replacement name for Mitrephorus Carter, 1913. Note: the First Reviser (Mitrothorax Carter, 1914 versus Timeneca Carter, 1914) is Matthews (1992: 456)

Mitua Hope, 1848: 56 [F]. Type species: Mituabidwelli Hope, 1848 (= Opatrumtuberculicostatum White, 1846), by original designation. Status: senior synonym of Pseudopatrum Sharp, 1886 in Lagriinae: Adeliini. Synonymy: Blair (1919a: 531). Note: Mitua Hope, 1848 has been used as valid in recent literature despite the fact that it is a junior homonym of Mitua Strickland, 1841, an unjustified emendation for Mitu Lesson, 1831 [Aves]; Strickland's name has been used as valid since 1899 (e.g., Heinroth 1931: 278) and therefore reversal of precedence cannot be used to conserve the younger name as valid in Tenebrionidae.

Mitys Champion, 1885: 97 [M]. Type species: Mitysinflatus Champion, 1885, by subsequent designation (Gebien 1943: 402). Status: valid genus in Stenochiinae: Cnodalonini.

Modicodisema Pic, 1917b: 3 [F]. Type species: Disemasubopaca Pic, 1912, by present designation. Status: valid subgenus of Barsenis Pascoe, 1887 in Lagriinae: Lagriini: Statirina. Note: originally proposed without included nominal species; based on personal communications with Maurice Pic, Borchmann (1936: 510) was the first author to subsequently and expressly include nominal species in Modicodisema (ICZN 1999, Article 67.2.2), by including the species Disemaserraticornis Mäklin, 1875 and Disemasubopaca Pic, 1912, in association with this name.

Moeon Champion, 1886: 251 [N]. Type species: Moeonisthmicum Champion, 1886, by subsequent designation (Gebien 1942a: 330). Status: valid genus in Stenochiinae: Cnodalonini. Note: as pointed out by M.A. Alonso-Zarazaga (in Bousquet et al. 2018: 307) the name Moeon is the Latinization of the Greek noun moion (privy parts) and is neuter.

Moerodes Rye, 1879: 62 [M]. Type species [automatic]: Prophaneswestwoodi W.J. MacLeay, 1872 (= Prophanesaculeatus Westwood, 1849), by monotypy. Status: junior synonym of Prophanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Note: unjustified emendation of Maerodes C.O. Waterhouse, 1877, not in prevailing usage.

Mogadoria Escalera, 1905c: 467 [F]. Type species: Tentyriasubelegans Fairmaire, 1871, by monotypy. Status: junior synonym of Eulipus Wollaston, 1864 in Pimeliinae: Tentyriini. Synonymy: Gebien (1937a: 634).

Mokalagria Pic, 1953: 163 [F]. Type species: Lopholagriaangustata Pic, 1953, by original designation. Status: valid subgenus of Lopholagria Borchmann, 1916 in Lagriinae: Lagriini: Lagriina.

Molion Champion, 1886: 142 [M]. Type species: Penetataurus Lacordaire, 1859, by subsequent designation (Gebien 1940: 758). Status: valid genus in Phrenapatinae: Penetini.

Moluris Latreille, 1802: 169 [F]. Type species: Pimeliagibba Fabricius, 1787 (= Tenebriogibbus Pallas, 1781), by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina. Note: the original combination of the accepted name of the type species, Tenebriogibbus Pallas, 1781, is a junior primary homonym of both Tenebriogibbus Linnaeus, 1767 and Tenebriogibbus DeGeer, 1778.

Monatrum Reichardt, 1936: 81, 208 [N]. Type species: Opatrumcarinatum Gebler, 1830, by original designation. Status: junior synonym of Scleropatrum Reitter, 1887 in Blaptinae: Opatrini: Opatrina. Synonymy: Löbl and Merkl (2003: 250).

Mongolesthes Reitter, 1904: 174 [F]. Type species: Melanesthesheydeni Csiki, 1901, by monotypy. Status: valid subgenus of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina.

Mongolopterocoma Skopin, 1974b: 146 [F]. Type species: Pterocomareitteri Frivaldszky, 1890, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Monodius Koch, 1956a: 180 [M]. Type species: Selinusconvexipennis Gebien, 1904, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Monoloba Solier, 1835a: 235 [F]. Type species: Lobopodadircaeoides Solier, 1835, by monotypy. Status: valid subgenus of Lobopoda Solier, 1835 in Alleculinae: Alleculini: Alleculina.

Montagona G.S. Medvedev, 1998a: 176 [F]. Type species: Tagonoidesasperula Fairmaire, 1896, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Montaguea Kaszab, 1982b: 227 [F]. Type species: Montagueacaledonica Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Adeliini.

Montanocatomus Nabozhenko, 2006: 842 [M]. Type species: Catomusgrandis G.S. Medvedev, 1978, by original designation. Status: valid subgenus of Catomus Allard, 1876 in Tenebrioninae: Helopini: Helopina.

Montanoodescelis Egorov, 2004: 592 [F]. Type species: Platyscelissahlbergi Reitter, 1900, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Monteithium Matthews, 1998: 704, 803 [N]. Type species: Monteithiumascetum Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Montiprosodes G.S. Medvedev, 2001: 83 [M]. Type species: Prosodesalaiensis Kraatz, 1885, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Mophis Champion, 1886: 168 [M]. Type species: Mophismarginicollis Champion, 1886, by subsequent designation (Gebien 1940: 1061). Status: valid genus in Diaperinae: Diaperini: Adelinina.

Mophon Champion, 1886: 247 [M]. Type species: Mophontinctipennis Champion, 1886, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Moragacinella Español, 1954a: 162 [F]. Type species [automatic]: Moralesialongepilosa Kaszab, 1944, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina. Note: replacement name for Moralesia Kaszab, 1944.

Moralesia Kaszab in Español, 1944: 18, 29 [F]. Type species: Moralesialongepilosa Kaszab, 1944, by original designation. Status: senior synonym of Moragacinella Español, 1954 in Blaptinae: Opatrini: Ammobiina. Note: junior homonym of Moralesia Fowler, 1943 [Pisces].

Morica Dejean, 1834: 182 [F]. Type species: Akisplanata Fabricius, 1801, by subsequent designation (Hope 1841: 122). Status: valid genus in Pimeliinae: Akidini.

Morocaulus Fairmaire, 1899d: 215 [M]. Type species: Morocaulusremipes Fairmaire, 1899 (= Bratynaapicalis Westwood, 1875), by monotypy. Status: junior synonym of Bratyna Westwood, 1875 in Alleculinae: incertae sedis. Synonymy: Borchamann (1938: 120, through synonymy of the type species with the type species of Bratyna Westwood, 1875).

Moromelas Fairmaire, 1898c: 481 [M]. Type species: Moromelasfoveipennis Fairmaire, 1898, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Morphostenophanes Pic, 1925b: 7 [M]. Type species: Morphostenophanesaenescens Pic, 1925, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mrazius Pic, 1925d: 86 [M]. Type species: Mraziusnodulosus Pic, 1925, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Mucheimira Novák, 2016a: 188 [F]. Type species: Isomirastoetzneri Muche, 1981, by original designation. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Mutiloxicum Nabozhenko & Ivanov, 2018: 546 [N]. Type species: Toxicumelvirae Nabozhenko & Ivanov, 2018, by original designation. Status: valid subgenus of Toxicum Latreille, 1802 in Tenebrioninae: Toxicini: Toxicina.

Myatis Bates, 1879b: 480 [F]. Type species: Myatishumeralis Bates, 1879, by subsequent designation (R. Lucas 1920: 425). Status: valid genus in Blaptinae: Platyscelidini.

Mycetochara Guérin-Méneville, 1827a: 346 [F]. Type species [automatic]: Cistelaflavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118). Status: valid genus and subgenus in Alleculinae: Alleculini: Mycetocharina. Note: replacement name for Mycetophila Gyllenhal, 1810; Mycetochara was also proposed later in the same year by Berthold (1827: 371), see Bouchard and Bousquet (2020a: 101); nomenclatural stability in this genus is threatened by the discovery that 1) the accepted type species, Cistelascapularis Illiger, 1805 (= Cistelahumeralis Fabricius, 1787) by subsequent designation by Westwood (1838: 32), is currently placed in the valid subgenusErnocharis C.G. Thomson, 1859 (e.g., Novák 2020i: 436) and 2) there are two other species currently placed in subgenusErnocharis C.G. Thomson, 1859 that were selected as type species of Mycetochara before the currently accepted designation in Westwood (1838: 32): Cistelahumeralis Fabricius, 1787 was chosen as the type species by Guérin-Méneville (1827: 346) when he proposed the replacement name Mycetochara, and Cistelalinearis Illiger, 1794 (= Cistelamaura Fabricius, 1792) was selected as the type species for Mycetocharis Gyllenhal, 1827, another replacement name for Mycetophila Gyllenhal, 1810, by Stephens (1832a: 375); of the nominal species originally included in Mycetophila Gyllenhal, 1810 that are currently placed in the subgenusMycetochara Guérin-Méneville, 1827, C.G. Thomson (1859: 118) was the first to select Cistelaflavipes Fabricius, 1792 as the type species; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to set aside previous type species designations for Mycetochara Guérin-Méneville, 1827 and select the type species designation proposed by C.G. Thomson (1859: 118).

Mycetochares Latreille, 1829a: 42 [M]. Type species [automatic]: Cistelaflavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118). Status: junior synonym of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: replacement name for Mycetophila Gyllenhal, 1810.

Mycetocharina Seidlitz, 1890: 136 [F]. Type species: Alleculaorientalis Faust, 1877, by monotypy. Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Mycetocharis Gyllenhal, 1827: 510 [F]. Type species [automatic]: Cistelaflavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118). Status: junior synonym of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: replacement name for Mycetophila Gyllenhal, 1810.

†Mycetocharoides Schaufuss, 1889: 269 [M]. Type species: Mycetocharoidesbaumeisteri Schaufuss, 1889, by monotypy. Status: valid genus in Alleculinae: Alleculini: Mycetocharina. Note: described from Eocene Baltic amber.

Mycetocula Novák, 2015c: 78 [F]. Type species: Mycetocharinasubcruciata Pic, 1922, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Mycetophila Gyllenhal, 1810: 541 [F]. Type species: Cistelaflavipes Fabricius, 1792, by subsequent designation (C.G. Thomson 1859: 118). Status: senior synonym of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: junior homonym of Mycetophila Meigen, 1803 [Diptera].

Mychestes Pascoe, 1870: 96 [M]. Type species: Mychesteslignarius Pascoe, 1870, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Mycotrogus Horn, 1870: 364, 367 [M]. Type species: Mycotroguspiceus Horn, 1870, by subsequent designation (R. Lucas 1920: 427). Status: valid genus in Tenebrioninae: Triboliini.

Myladanesthes Skopin, 1961b: 202 [F]. Type species: Myladinafortidens Reitter, 1915, by original designation. Status: valid subgenus of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina. Note: the alternative original spelling Myladansthes, used by Skopin (1961b: 202), was rejected by Skopin (1967: 205) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Myladina Reitter, 1889a: 706 [F]. Type species: Myladinaunguiculina Reitter, 1889, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Myladion Reitter, 1887a: 386 [N]. Type species: Myladionacuticolle Reitter, 1887, by monotypy. Status: valid subgenus of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Note: the alternative original spelling Miladion, used by Reitter (1887a: 385), was rejected by Reitter (1896b: 166) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Mylaris Pallas, 1781: 37 [F]. Type species: Tenebriogigas Linnaeus, 1763, by subsequent designation (Guérin-Méneville 1844: 120). Status: valid genus in Stenochiinae: Cnodalonini. Note: see Bousquet et al. (2018: 307, footnote 82) regarding a possible nomenclatural problem with this genus name.

Myonophloeus Bremer & Lillig, 2017a: 68 [M]. Type species: Corticeustuberculatus Triplehorn, 1979, by original designation. Status: valid genus in Diaperinae: Hypophlaeini.

Myrmechixenus Chevrolat, 1835: 267 [M]. Type species: Myrmechixenussubterraneus Chevrolat, 1835, by monotypy. Status: valid genus in Diaperinae: Myrmechixenini.

Myrmechoxenus Gaubil, 1849: 288 [M]. Type species [automatic]: Myrmechixenussubterraneus Chevrolat, 1835, by monotypy. Status: junior synonym of Myrmechixenus Chevrolat, 1835 in Diaperinae: Myrmechixenini. Note: unjustified emendation of Myrmechixenus Chevrolat, 1835, not in prevailing usage; while both Myrmechixenus (p. 71) and Myrmechoxenus (p. 266) were used by Gaubil (1849), the fact that Myrmechixenus was written in italics in the Index (p. 282) indicates that the original spelling was treated as a synonym.

Myrmecocatops Wasmann, 1897: 268 [M]. Type species: Myrmecocatopslatus Wasmann, 1897, by monotypy. Status: valid genus in Diaperinae: Crypticini.

Myrmecodema Gebien, 1943: 402 [F]. Type species [automatic]: Myrmecosomanycterinoides Germain, 1855, by monotypy. Status: valid genus in Tenebrioninae: Trachelostenini. Note: replacement name for Myrmecosoma Germain, 1855. Note: transferred from Stenochiinae: Cnodalonini by Matthews and Lawrence (2015: 293).

Myrmecodichillus Kaszab, 1960a: 6, 7, 14 [M]. Type species: Dichillusreichenspergeri Kaszab, 1960, by original designation. Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Myrmecopeltoides Kaszab, 1973b: 318 [M]. Type species: Myrmecopeltoidescamponoti Kaszab, 1973, by original designation. Status: valid genus in Lagriinae: Goniaderini.

Myrmecophosis Koch, 1958: 72 [F]. Type species: Zophosispedinoides Gebien, 1920, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Myrmecosoma Germain, 1855: 403 [N]. Type species: Myrmecosomanycterinoides Germain, 1855, by monotypy. Status: senior synonym of Myrmecodema Gebien, 1943 in Tenebrioninae: Trachelostenini. Note: junior homonym of Myrmecosoma Mannerheim, 1846 [Coleoptera: Anthicidae].

Myrmecoxenus Märkel, 1844: 253 [M]. Type species [automatic]: Myrmechixenussubterraneus Chevrolat, 1835, by monotypy. Status: junior synonym of Myrmechixenus Chevrolat, 1835 in Diaperinae: Myrmechixenini. Note: unjustified emendation of Myrmechixenus Chevrolat, 1835, not in prevailing usage.

Nalassus Mulsant, 1854: 323 [M]. Type species: Helopsdryadophilus Mulsant, 1854, by subsequent designation (Nabozhenko 2001a: 630). Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina.

Nalepa Reitter, 1887a: 366 [F]. Type species: Blapscylindracea Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Namaphosis Penrith, 1981c: 127, 143 [F]. Type species: Zophosissolivaga Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Namaquaeon Koch, 1950b: 305, 341 [N]. Type species: Phaeotribonaustralis Péringuey, 1908, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Namazopus Koch, 1963: 34 [M]. Type species: Namazopusarachnipes Koch, 1963, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Namibismus Koch, 1952a: 107 [M]. Type species: Namibismuscastaneus Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Namibomodes Koch, 1952d: 223 [M]. Type species: Psammodesserrimargo Gebien, 1938, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Nannalcyon Koch, 1950a: 68 [F]. Type species [automatic]: Nannoceruscylindrus Fairmaire, 1887, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Nannocerus Fairmaire, 1887.

Nannocerus Fairmaire, 1887b: 292 [M]. Type species: Nannoceruscylindrus Fairmaire, 1887, by monotypy. Status: senior synonym of Nannalcyon Koch, 1950 in Stenochiinae: Cnodalonini. Note: junior homonym of Nannocerus Mayr, 1885 [Hymenoptera].

Nannohyocis G.S. Medvedev & Lawrence, 1983: 569 [M]. Type species: Hyocisinquilinus Carter, 1921, by original designation. Status: valid subgenus of Hyocis Pascoe, 1866 in Diaperinae: Hyociini: Hyociina.

Nanoblaps Semenov-Tjan-Shansky & Bogatchev, 1936: 565 [F]. Type species: Blaps jakovlevi Semenov-Tjan-Shansky & Bogatchev, 1936, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Löbl et al. (2008c: 219).

Nanocaecus Schawaller & Purchart, 2012: 305, 310 [M]. Type species: Nanocaecushlavaci Schawaller & Purchart, 2012, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Nanocalcar Skopin, 1974a: 67, 102 [N]. Type species: Nanocalcarnanum Skopin, 1974, by monotypy. Status: valid subgenus of Centorus Mulsant, 1854 in Lagriinae: Belopini.

Nanohemicera Pic, 1923c: 25 [F]. Type species: Hemicerarufomaculata Pic, 1923, by monotypy. Status: valid subgenus of Hemicera Laporte & Brullé, 1831 in Stenochiinae: Cnodalonini.

Nanotagalus Gebien, 1942b: 118, 120 [M]. Type species: Afrotagalususambaricus Gebien, 1942, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Narses Champion, 1888: 423 [M]. Type species: Narsessubalatus Champion, 1888, by monotypy. Status: junior synonym of Charisius Champion, 1888 in Alleculinae: Alleculini: Alleculina. Synonymy: Campbell (2014: 271).

Narsodes Campbell, 1976: 32 [M]. Type species: Narsodesbrachypterus Campbell, 1976, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Nataloplonyx Ardoin, 1963c: 716 [M]. Type species: Hoplonyxmicans Schaufuss, 1870, by original designation. Status: valid subgenus of Hoplonyx J. Thomson, 1858 in Tenebrioninae: Amarygmini.

Natalostira Pic, 1913c: 143 [F]. Type species: Natalostirabrevithorax Pic, 1913, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Nautes Pascoe, 1866a: 475 [M]. Type species: Nautesfervidus Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Neacisba Peyerimhoff, 1927: 53 [F]. Type species [automatic]: Pachychiladissecta Kraatz, 1865, by monotypy. Status: junior synonym of Neocisba Reitter, 1900 in Pimeliinae: Tentyriini. Note: unjustified emendation of Neocisba Reitter, 1900, not in prevailing usage.

Neandrosus Pic, 1921b: 12 [M]. Type species: Neandrosussingularipes Pic, 1921, by monotypy. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1987: 44). Note: we act as First Revisers and reject the alternative original spelling Neoandrosus, used by Pic (1921b: 12).

Neanopidium Dajoz, 1975a: 93 [N]. Type species: Neanopidiummexicanum Dajoz, 1975, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Neatus J.L. LeConte, 1862: 233 [M]. Type species: Helopstenebrioides Palisot de Beauvois, 1812, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Necrobioides Fairmaire, 1882a: 234 [M]. Type species: Necrobioidescoeruleatus Fairmaire, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Neglectophloeus Bremer & Lillig, 2017a: 71 [M]. Type species: Hypophlaeusluteomaculatus Pic, 1914, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Nelites J.L. LeConte, 1850: 232 [M]. Type species: Nelitesaeneolus J.L. LeConte, 1850, by monotypy. Status: junior synonym of Scaphidema Redtenbacher, 1848 in Diaperinae: Scaphidemini. Synonymy: J.L. LeConte (1862: 237).

Nemanes Fairmaire, 1888a: 195 [M]. Type species: Nemanesexpansicollis Fairmaire, 1888, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Nemapus Solier, 1835b: 313 [M]. Type species: Microderacoromandelensis Solier, 1835, by original designation. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: the name Nemapus was listed as synonym of Microdera Eschscholtz, 1831 by Solier (1835b: 313); it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Koch 1943b: 860); therefore, Nemapus was made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Nemoplonyx Ardoin, 1963c: 715, 734 [M]. Type species: Hoplonyxgallanus Gridelli, 1939, by original designation. Status: valid subgenus of Hoplonyx J. Thomson, 1858 in Tenebrioninae: Amarygmini.

Nemostira Fairmaire, 1869a: 815 [F]. Type species: Nemostiracoquerelii Fairmaire, 1869, by subsequent designation (R. Lucas 1920: 434). Status: junior synonym of Sora Walker, 1859 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1929a: 5).

Nemostiromorpha Pic, 1917b: 2,3 [F]. Type species: Disemalongicornis Mäklin, 1875, by subsequent monotypy (Borchmann 1936: 510). Status: valid subgenus of Barsenis Pascoe, 1887 in Lagriinae: Lagriini: Statirina. Note: originally proposed without included nominal species; based on personal communications with Maurice Pic, Borchmann (1936: 510) was the first author to subsequently and expressly include nominal species in Nemostiromorpha (ICZN 1999, Article 67.2.2), by including the species Disemalongicornis Mäklin, 1875, in association with this name.

Nemostiropsis Borchmann, 1936: 341 [F]. Type species: Sorapurpureipennis Borchmann, 1930, by original designation. Status: valid subgenus of Sora Walker, 1859 in Lagriinae: Lagriini: Statirina.

Neoabantis Gebien, 1910b: 341 [F]. Type species [automatic]: Abantisaenescens Fairmaire, 1892, by monotypy. Status: junior synonym of Diphyrrhynchus Fairmaire, 1849 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Gebien (1938a: 407). Note: replacement name for Abantiades Fairmaire, 1894.

Neoadelium Carter, 1908a: 259 [N]. Type species: Adeliumfairmairei Bates, 1873, by subsequent designation (Gebien 1942a: 742). Status: valid genus in Lagriinae: Adeliini.

Neoatractus Borchmann, 1909a: 713 [M]. Type species [automatic]: Atractusviridis Boisduval, 1835, by subsequent designation (Duponchel and Chevrolat 1841: 312). Status: junior synonym of Lepturidea Fauvel, 1862 in Alleculinae: Alleculini: Alleculina. Synonymy: Matthews and Bouchard (2008: 324). Note: unnecessary replacement name for Atractus Boisduval, 1835.

Neobaphion Blaisdell, 1925: 390 [N]. Type species: Eleodesplanipennis J.L. LeConte, 1866, by monotypy. Status: valid genus in Blaptinae: Amphidorini.

Neoblaps Ren & Li, 2001: 310 [F]. Type species: Neoblapshuizensis Ren & Li, 2001, by monotypy. Status: junior synonym of Coelocnemodes Bates, 1879 in Blaptinae: Blaptini: Blaptina. Synonymy: Ren et al. (2016: 432).

Neocabirutus Kulzer, 1964: 221 [M]. Type species: Neocabirutusindicus Kulzer, 1964, by monotypy. Status: valid subgenus of Cabirutus Strand, 1929 in Blaptinae: Pedinini: Pedinina.

Neocaedius Pierre, 1972: 960, 967 [M]. Type species: Caediushalli Kaszab, 1949, by original designation. Status: junior synonym of Cyptus Gerstaecker, 1871 in Blaptinae: Opatrini: Ammobiina. Synonymy: Ferrer (1996: 254, 2003: 297).

Neocamaria Kulzer, 1954a: 52 [F]. Type species: Neocamariatibialis Kulzer, 1954, by original designation. Status: junior synonym of Robustocamaria Pic, 1922 in Stenochiinae: Cnodalonini. Synonymy: Masumoto (1993b: 222).

Neocisba Reitter, 1900c: 154 [F]. Type species: Pachychiladissecta Kraatz, 1865, by monotypy. Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Neocistela Borchmann, 1909a: 713 [F]. Type species [automatic]: Pseudocistelaovalis Blackburn, 1891, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: replacement name for Pseudocistela Blackburn, 1891.

Neodissonomus G.S. Medvedev, 1968a: 235 [M]. Type species: Heterophylusangustitarsis Reitter, 1896, by original designation. Status: valid subgenus of Dissonomus Jacquelin du Val, 1861 in Tenebrioninae: Dissonomini.

Neoeutrapela Bousquet & Bouchard, 2013a: 61 [F]. Type species [automatic]: Crioceriselongata Fabricius, 1781 (= Chrysomelaunifasciata DeGeer, 1778), by subsequent designation (Duponchel 1844b: 533). Status: valid subgenus (stat. nov. [OM]) of Impressosora Pic, 1952 in Lagriinae: Lagriini: Statirina. Note: replacement name for Eutrapela Dejean, 1834; new placement [OM], previously included in Lagriinae: Lagriini: Lagriina.

Neognathosia Kaszab, 1959a: 383 [F]. Type species: Gnathosiapseudanemia Reitter, 1915, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Neogria Borchmann, 1911: 222 [F]. Type species: Neogriasulcipennis Borchmann, 1911, by subsequent designation (R. Lucas 1920: 436). Status: valid genus in Lagriinae: Lagriini: Lagriina.

Neohelops Dajoz, 2001: 356 [M]. Type species: Neohelopstexanus Dajoz, 2001, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina. Note: placed in the subtribe Helopina by Nabozhenko and Purchart (2019: 156).

Neohyocis G.S. Medvedev & Lawrence, 1983: 571 [M]. Type species: Hyociswaterhousei G.S. Medvedev & Lawrence, 1983, by original designation. Status: valid subgenus of Hyocis Pascoe, 1866 in Diaperinae: Hyociini: Hyociina.

Neoisocerus Bouchard, Lawrence, Davies & Newton, 2005: 510 [M]. Type species [automatic]: Tenebriopurpurascens Herbst, 1799 (= Helopsferrugineus Fabricius, 1798), by monotypy. Status: valid genus in Blaptinae: Dendarini: Dendarina. Note: replacement name for Isocerus Dejean, 1821.

Neomenimus Kaszab, 1939b: 190 [M]. Type species: Neomenimusclavatus Kaszab, 1939, by original designation. Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: G.S. Medvedev (2007b: 665).

Neomida Latreille, 1829a: 29 [F]. Type species: Ips haemorrhoidalis Fabricius, 1787, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: Neomida was used earlier by Dahl (1823: 44) but Dahl’s work was suppressed for the purposes of zoological nomenclature by the ICZN (1964, Opinion 710).

Neooligocara Guerrero, Vidal & Moore, 2007: 407 [N]. Type species: Oligocarabucki Kulzer, 1962, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Neopachypterus Bouchard, Löbl & Merkl, 2007: 386 [M]. Type species [automatic]: Pachypterusmauritanicus P.H. Lucas, 1847, by monotypy. Status: valid genus in Blaptinae: Opatrini: Neopachypterina. Note: replacement name for Pachypterus P.H. Lucas, 1847.

Neophaleria Español, 1963a: 74, 76 [F]. Type species: Phaleriaardoini Español, 1963, by monotypy. Status: valid subgenus of Phaleria Latreille, 1802 in Diaperinae: Phaleriini.

Neophylax Bedel, 1906a: 92 [M]. Type species [automatic]: Phylaxlittoralis Mulsant, 1854 (= Opatrumpicipes G.-A. Olivier, 1812), by subsequent designation (Gebien 1938a: 412). Status: senior synonym of Allophylax Bedel, 1906 in Blaptinae: Dendarini: Melambiina. Note: junior homonym of Neophylax McLachlan, 1871 [Trichoptera].

Neoplamius Löbl, Bouchard, Merkl & Bousquet, 2020: 4 [M]. Type species: Neoplamiuszoltani Masumoto, 1981, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: name first proposed by Masumoto (1981: 15) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Neoplateia Marcuzzi, 1986: 177 [F]. Type species: Neoplateiakaszabi Marcuzzi, 1986, by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Neoplonyx Ardoin, 1963b: 309, 352 [M]. Type species: Gonocnemissulcicollis Fairmaire, 1899, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Neoporphyrhyba Ardoin, 1956b: 89 [F]. Type species: Porphyrhybacyaneocuprea Fairmaire, 1894, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Neopraocis Kulzer, 1958a: 3, 6 [M]. Type species: Praocisreflexicollis Solier, 1851, by monotypy. Status: valid genus in Pimeliinae: Praociini.

Neopsectropus Kaszab, 1941c: 30 [M]. Type species: Neopsectropusgebieni Kaszab, 1941, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Neopterocoma Skopin, 1974b: 158 [F]. Type species: Pterocomabalchashensis Skopin, 1974, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Neosolenopistoma Bouchard & Bousquet, new subgenus. [N]. Type species: Eurynotusdenticosta Mulsant & Rey, 1854, by present designation. Status: valid subgenus of Eurynotus W. Kirby, 1819 in Blaptinae: Platynotini: Eurynotina. Note: taxon first proposed by Mulsant and Rey (1854: 29; as “Solenopistoma Solier”); however, this is treated as an incorrect subsequent spelling of Selenepistoma Dejean, 1834 (see Bousquet and Bouchard 2013: 50); the subgenusSolenopistoma Mulsant & Rey, 1854, which is currently used as valid, is therefore unavailable (ICZN 1999, Articles 33.3); we hereby make the name Neosolenopistoma available by selecting Eurynotusdenticosta Mulsant & Rey, 1854 as type species and referring to Mulsant and Rey (1854: 29) for the character states that characterise and differentiate Neosolenopistoma.

Neotagalus Kaszab, 1955a: 471, 477 [M]. Type species: Neotagalustuberculiger Kaszab, 1955, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Neotheca Carter, 1930: 540 [F]. Type species: Neothecafusca Carter, 1930, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Neozophobas Ferrer, 2006b: 235 [M]. Type species: Zophobaslaticollis Kraatz, 1880, by original designation. Status: valid genus in Tenebrioninae: Tenebrionini.

Nepalindia G.S. Medvedev, 1998a: 187 [F]. Type species: Tagonoidesalpina Kaszab, 1965, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Nepalofranziella Fouquè, 2013: 194, 196 [F]. Type species: Nepalofranziellakaszabi Fouquè, 2013, by original designation. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Nepalolaena Schawaller, 2001a: 277 [F]. Type species: Nepalolaenakira Schawaller, 2001, by original designation. Status: valid genus in Lagriinae: Laenini.

Nepaloplonyx Bremer, 2014b: 175, 181 [M]. Type species: Nepaloplonyxcaelebs Bremer, 2014, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Nephodes Blanchard, 1845: 34 [M]. Type species: Nephodesvilliger Rosenhauer, 1856, by subsequent monotypy (Rosenhauer 1856: 218). Status: senior synonym of Nephodinus Gebien, 1943 in Tenebrioninae: Helopini: Helopina. Note: originally proposed without included nominal species; Rosenhauer (1856: 218), by including the new species Nephodesvilliger Rosenhauer, 1856 in association with this name, was the first author to subsequently and expressly include nominal species in Nephodes (ICZN 1999, Article 67.2.2); the older name Nephodes Schönherr, 1840 [Coleoptera: Curculionidae] is available as it was originally proposed as a synonym and subsequently treated as a senior homonym by Gebien (1943: 900; see ICZN 1999, Article 11.6.1), therefore Nephodes Blanchard, 1845 is a junior homonym of Nephodes Schönherr, 1840.

Nephodinus Gebien, 1943: 900 [M]. Type species [automatic]: Nephodesvilliger Rosenhauer, 1856, by subsequent monotypy (Rosenhauer 1856: 218). Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina. Note: replacement name for Nephodes Blanchard, 1845.

Nerina Lacordaire, 1859a: 70 [F]. Type species: Nerinadispar Lacordaire, 1859, by monotypy. Status: senior synonym of Afrinus Fairmaire, 1888 in Pimeliinae: Tentyriini. Synonymy: Blair (1935a: 103). Note: junior homonym of Nerina Robineau-Desvoidy, 1830 [Diptera].

Nerinodon Koch, 1952a: 138 [M]. Type species: Nerinodoncaviceps Koch, 1952, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Nesioticus Westwood, 1843: 120 [M]. Type species: Nesioticusflavopictus Westwood, 1843, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: redescribed as new by Westwood (1844: 227).

Nesocaedius Kolbe, 1915: 262 [M]. Type species: Nesocaediusschultzei Kolbe, 1915, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Nesocyrtosoma Marcuzzi, 1976: 137 [N]. Type species: Cyrtosomainflatum Marcuzzi, 1976, by plenary powers (ICZN 2017, Opinion 2398). Status: valid genus in Stenochiinae: Cnodalonini. Note: following an application by Hopp et al. (2014) to conserve the generic name Nesocyrtosoma Marcuzzi, 1976, the ICZN (2017, Opinion 2398) determined that this name is nomenclaturally available despite not having been accompanied by a type species fixation in the original publication, placed it on the Official List of Generic Names in Zoology and, designated Cyrtosomainflatum Marcuzzi, 1976 as its type species.

Nesogena Mäklin, 1863b: 557 [F]. Type species: Nesogenahybrida Mäklin, 1863, by subsequent designation (Gebien 1948: 549). Status: valid genus and subgenus in Tenebrioninae: Praeugenini.

Nesogenomorpha Pic, 1917c: 18 [F]. Type species: Nesogenomorphasemiviolacea Pic, 1917, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Nesopatrum Gebien, 1921b: 20 [N]. Type species: Opatrinusjosephi Karsch, 1881, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Nesophaerotus Ardoin, 1962a: 66, 67 [M]. Type species [automatic]: Nesosphaerotusaeneus Gebien, 1921, by subsequent designation (Gebien 1943: 403). Status: junior synonym of Nesosphaerotus Gebien, 1921 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Nesosphaerotus Gebien, 1921, not in prevailing usage.

Nesosphaerotus Gebien, 1921b: 115 [M]. Type species: Nesosphaerotusaeneus Gebien, 1921, by subsequent designation (Gebien 1943: 403). Status: valid genus in Stenochiinae: Cnodalonini.

Nesostes Casey, 1908: 56, 58 [M]. Type species: Eusattusrobustus J.L. LeConte, 1866, by original designation. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: Triplehorn (1968: 379).

Nesotaurus Fairmaire, 1896b: 354 [M]. Type species: Nesotaurussericans Fairmaire, 1896, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Nesotes Allard, 1876a: 5 [M]. Type species: Helopsasper Küster, 1850, by subsequent designation (Nabozhenko 2008: 38). Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina. Note: the First Reviser (Nesotes Allard, 1876 versus Diastixus Allard, 1876) is Antoine (1949: 134).

Netopha Fairmaire, 1893c: 299 [F]. Type species: Netophapallidipes Fairmaire, 1893, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Netuschilia Reitter, 1904: 34, 35 [F]. Type species: Lachnopushauseri Reitter, 1897, by monotypy. Status: valid genus in Pimeliinae: Lachnogyini: Netuschiliina.

Nevermanniella Borchmann, 1936: 235, 332 [F]. Type species: Statiraalbolineata Champion, 1889, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: the alternative original spelling Nevermannia (pp. 11, 235) was corrected to Nevermanniella in the “Corrigenda” of the same work (p. 541), Nevermanniella is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1)

Nevisia Marcuzzi, 1986: 179 [F]. Type species: Diastolinusbardudensis Marcuzzi, 1962, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Nicandra Fairmaire, 1888a: 191 [F]. Type species: Nicandracostulipennis Fairmaire, 1888, by monotypy. Status: valid genus and subgenus in Blaptinae: Pedinini: Helopinina.

Nikomenalia Dubrovina, 1975: 166 [F]. Type species: Hymenaliakaszabi Muche, 1972, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: elevated from subgenus of Hymenalia Mulsant, 1856 by Novák (2020c: 507).

Nilio Latreille, 1802: 179 [M]. Type species: Coccinellavillosa Fabricius, 1787, by monotypy. Status: valid genus and subgenus in Nilioninae. Note: Nilio is an incorrect subsequent spelling of the original spelling Nilion, first used by Latreille (1804: 333), in prevailing usage; Nilio is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1); the original combination of the name of the type species, Coccinellavillosa Fabricius, 1787, is a junior primary homonym of Coccinellavillosa Fourcroy, 1785.

Nipponalassus Nabozhenko & Ando, 2018: 306 [M]. Type species: Tarpelaandoi Masumoto, 1993, by original designation. Status: valid subgenus of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina.

Nipponohelops Masumoto, Ando & Akita, 2006: 33 [M]. Type species: Nipponohelopsishikawai Masumoto, Ando & Akita, 2006, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Nocar Blackburn, 1891: 328 [N]. Type species: Cisteladepressiuscula W.J. MacLeay, 1872, by subsequent designation (R. Lucas 1920: 441). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Nochelius Gistel, 1848a: xi [M]. Type species [automatic]: Evaniosomusorbignianus Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Evaniosomus Guérin-Méneville, 1834 in Pimeliinae: Evaniosomini. Note: unnecessary replacement name for Evaniosomus Guérin-Méneville, 1834, not in prevailing usage.

Nocibiotes Casey, 1895: 617 [M]. Type species: Notibiusgranulatus J.L. LeConte, 1851, by subsequent designation (Gebien 1938a: 407). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Nodosogylium Pic, 1951: 12 [N]. Type species: Nodosogyliuminaequale Pic, 1951, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: we act as First Revisers and reject the alternative original spelling Nodosogilium, used by Pic (1951: 12).

Nodotelus Koch, 1950a: 67 [M]. Type species [automatic]: Eutelusrequieni Solier, 1843, by subsequent designation (R. Lucas 1920: 291). Status: junior synonym of Eutelonotus Fairmaire in Alluaud, 1902 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Eutelus Solier, 1843.

Nolicima Matthews, 1998: 708, 714 [F]. Type species: Cardiothoraxangusticollis Carter, 1906, by original designation. Status: valid genus in Lagriinae: Adeliini.

Notacula Campbell, 1971: 107 [F]. Type species: Notaculahowdenae Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Nothogria Borchmann, 1916a: 49, 142 [F]. Type species: Nothogrianodipennis Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Nothrocerus Fairmaire, 1887a: 174 [M]. Type species: Nothroceruscylindricornis Fairmaire, 1887, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Notiasida Casey, 1912: 76, 124 [F]. Type species: Notiasidaabstrusa Casey, 1912, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Notibius J.L. LeConte, 1851: 144 [M]. Type species: Notibiuspuberulus J.L. LeConte, 1851, by subsequent designation (Gebien 1938a: 406). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Notiolesthus Motschulsky, 1872: 25 [M]. Type species: Notiolesthusnatalensis Motschulsky, 1872, by original designation. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1921b: 69). Note: we act as First Revisers and reject the alternative original spelling Notiolosthus, used by Motschulsky (1872: 34).

Notioscythis Fairmaire, 1883a: 31 [F]. Type species: Notioscythispunctoseriata Fairmaire, 1883, by monotypy. Status: junior synonym of Stenosida Solier, 1835 in Pimeliinae: Tentyriini. Synonymy: Chatanay (1917: 231).

Notoblaps Bauer, 1921: 232 [F]. Type species: Blaps juliae Allard, 1881, by monotypy. Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Notocistela Carter, 1915a: 78, 102 [F]. Type species: Notocistelatibialis Carter, 1915, by subsequent designation (Matthews and Bouchard 2008: 333). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Notocorax Dejean, 1834: 191 [M]. Type species: Opatrumjavanum Wiedemann, 1819, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Notoprataeus Carter, 1924a: 37 [M]. Type species: Notoprataeuslitoralis Carter, 1924 (= Mesotertisinconstans Lea, 1917), by monotypy. Status: junior synonym of Micropedinus Lewis, 1894 in Lagriinae: Lupropini. Synonymy: G.S. Medvedev (1992: 656), Matthews and Lawrence (2005: 534).

Notostrongylium Carter, 1915b: 523 [N]. Type species: Notostrongyliumrugosicolle Carter, 1915, by subsequent designation (Kulzer 1966: 387). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Kaszab (1977b: 3, 29).

Nototrintus Carter, 1924a: 40 [M]. Type species: Otrintusjacksoni Carter, 1905, by original designation. Status: valid genus in Lagriinae: Adeliini.

Nudoplatyscelis Kaszab, 1940a: 149, 222 [F]. Type species: Platynoscelisturanica Reitter, 1896, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Nuptis Motschulsky, 1872: 25 [M]. Type species: Nuptistenuis Motschulsky, 1872, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Nyctalops Gistel, 1848a: 125 [M]. Type species [automatic]: Pseudoblapssubstriata Guérin-Méneville, 1834, by subsequent designation (Hope 1841: 124). Status: junior synonym of Pseudoblaps Guérin-Méneville, 1834 in Blaptinae: Platynotini: Platynotina. Note: unnecessary replacement name for Pseudoblaps Guérin-Méneville, 1834; junior homonym of Nyctalops Wagler, 1832 [Aves].

Nyctelia Berthold, 1827: 367 [F]. Type species: Zophosisnodosa Germar, 1823 (= Zophosispicipes Billberg, 1815), by monotypy. Status: valid genus in Pimeliinae: Nycteliini. Note: nomen protectum (see Silvestro and Flores 2016: 658).

Nyctelioma Casey, 1908: 163 [F]. Type species: Nycteliomaexplanata Casey, 1908, by original designation. Status: junior synonym of Nyctelia Berthold, 1827 in Pimeliinae: Nycteliini. Synonymy: Gebien (1910a: 141).

Nyctelius Guérin-Méneville, 1827b: 21 [M]. Type species: Zophosisnodosa Germar, 1823 (= Zophosispicipes Billberg, 1815), by monotypy. Status: senior synonym of Nyctelia Berthold, 1827 in Pimeliinae: Nycteliini. Synonymy: Silvestro and Flores (2016: 658). Note: nomen oblitum (see Silvestro and Flores 2016: 658).

Nycterinus Eschscholtz, 1829: 9.13 [M]. Type species: Nycterinusthoracicus Eschscholtz, 1829, by subsequent designation (Hope 1841: 124). Status: valid genus in Blaptinae: Amphidorini.

Nycteropus Klug, 1833: 89 [M]. Type species: Nycteropusebeninus Klug, 1833, by subsequent designation (Hope 1841: 124). Status: valid genus in Tenebrioninae: Toxicini: Nycteropina.

Nyctipates Gebler, 1841: 373 [M]. Type species: Nyctipatesrugulosus Gebler, 1841, by monotypy. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Erichson (1845a: 116).

Nyctipates Solier, 1848: 154, 285 [M]. Type species: Nyctipatescoriaceus Solier, 1848 (= Blapsangustata Zubkov, 1833), by subsequent designation (Löbl et al. 2008c: 236). Status: senior synonym of Prosodopria Reitter, 1909 in Blaptinae: Blaptini: Prosodina. Synonymy: Löbl et al. (2008a: 43). Note: junior homonym of Nyctipates Gebler, 1841 [Coleoptera: Tenebrionidae: Blaptinae: Blaptini: Prosodina].

Nyctobates Guérin-Méneville, 1834: 33 [M]. Type species: Tenebriogigas Linnaeus, 1763, by original designation. Status: junior synonym of Mylaris Pallas, 1781 in Stenochiinae: Cnodalonini. Synonymy: Spilman (1973: 42), Ferrer and Siliansky (2008: 186).

Nyctopetus Guérin-Méneville, 1831a: pl. 4 [M]. Type species: Nyctopetustenebrioides Guérin-Méneville, 1831, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Nyctoporis Eschscholtz, 1831: 10, 11 [F]. Type species: Nyctoporiscristata Eschscholtz, 1831, by subsequent designation (Hope 1841: 124). Status: valid genus in Pimeliinae: Nyctoporini.

Nyctozoilus Guérin-Méneville, 1831a: pl. 4 [M]. Type species: Nyctozoilusobesus Guérin-Méneville, 1831, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Nypsius Champion, 1895a: 219 [M]. Type species: Nypsiusaeneopiceus Champion, 1895, by subsequent designation (R. Lucas 1920: 447). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Oatesius Westwood, 1889: 376 [M]. Type species [automatic]: Derosphaeriusanthracinus Westwood, 1881, by monotypy. Status: junior synonym of Derosphaerius Westwood, 1881 in Pimeliinae: Tentyriini. Note: unnecessary replacement name for Derosphaerius Westwood, 1881.

Obenbergeria Strand, 1929: 24 [F]. Type species [automatic]: Diaperisriederii Faldermann, 1833, by subsequent designation (Jakobson 1924: 243). Status: junior synonym of Emypsara Pascoe, 1866 in Diaperinae: Phaleriini. Synonymy: Gebien (1939: 743). Note: replacement name for Callicomus Motschulsky, 1860.

Obesacula Campbell, 1971: 109 [F]. Type species: Obesaculaaptera Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Oblongoodescelis Kaszab, 1940b: 941, 958 [F]. Type species: Platyscelisoblonga Ballion, 1878, by original designation. Status: junior synonym of Clavatoodescelis Kaszab, 1940 in Blaptinae: Platyscelidini. Synonymy: Egorov (2020: 380).

Oblongoplatyscelis Kaszab, 1940b: 910, 916 [F]. Type species: Platyscelisganglbaueri Seidlitz, 1893, by original designation. Status: valid subgenus of Platyscelis Latreille, 1818 in Blaptinae: Platyscelidini.

Obriomaia Gebien, 1927: 45 [F]. Type species: Eucyrtussubcostatus Fairmaire, 1893, by original designation. Status: junior synonym of Tetragonomenes Chevrolat, 1878 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 133).

Occidentophosis Penrith, 1977: 18, 157 [F]. Type species: Zophosisdamarina Péringuey, 1908, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Ochrolamus Reitter, 1904: 73 [M]. Type species: Dilamuspictus Baudi di Selve, 1881, by monotypy. Status: valid subgenus of Dilamus Jacquelin du Val, 1861 in Blaptinae: Opatrini: Ammobiina.

Ocnera Fischer, 1822: 169 [F]. Type species: Pimeliaimbricata Fischer, 1820, by subsequent designation (Wilke 1922: 260). Status: valid genus in Pimeliinae: Pimeliini. Note: as mentioned by Bouchard and Bousquet (2020b: 7) the nomenclatural stability of this name is threatened by the discovery of an older type species designation (Tenebriocephalotes Pallas, 1781, by subsequent designation by Crotch (1870b: 241), currently the type species of the valid subgenusChaetotoma Motschulsky, 1860 in Pimelia Fabricius, 1775); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Gebien (1937a: 814).

Ocnodes Fåhraeus, 1870: 270 [F]. Type species: Ocnodesscrobicollis Fåhraeus, 1870, by subsequent designation (Kamiński et al. 2019b: 55). Status: valid genus and subgenus in Pimeliinae: Sepidiini: Molurina.

Ocularisora Pic, 1934a: 32 [F]. Type species: Nemostirabenitensis Pic, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Oculochara Novák, 2020e: 80 [F]. Type species: Mycetocharaocularis Reitter, 1884, by original designation. Status: valid subgenus of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina.

Oculosis Penrith, 1977: 18, 126 [F]. Type species: Zophosisboei Solier, 1834, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Odocnemis Allard, 1876a: 4 [F]. Type species: Odocnemiscaudata Allard, 1876 (= Helopspraelongus Baudi di Selve, 1876), by subsequent designation (Nabozhenko 2001a: 662). Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina. Note: the First Reviser (Odocnemis Allard, 1876 versus Omaleis Allard, 1876) is Nabozhenko (2001a: 628).

Odontocera Chen & Yuan, 1996: 183 [F]. Type species: Odontoceraqinlingensis Chen & Yuan, 1996, by original designation. Status: senior synonym of Odontocerostira Merkl, 2007 in Lagriinae: Lagriini: Statirina. Note: junior homonym of Odontocera Audinet-Serville, 1834 [Coleoptera: Cerambycidae].

Odontocerostira Merkl, 2007: 269 [F]. Type species [automatic]: Odontoceraqinlingensis Chen & Yuan, 1996, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: replacement name for Odontocera Chen & Yuan, 1996.

Odontocnemis Rye, 1878: 69 [F]. Type species [automatic]: Odocnemiscaudata Allard, 1876 (= Helopspraelongus Baudi di Selve, 1876), by subsequent designation (Nabozhenko 2001a: 662). Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Note: unjustified emendation of Odocnemis Allard, 1876, not in prevailing usage.

Odontogria Borchmann, 1936: 16, 62 [F]. Type species: Lagriablairi Borchmann, 1925, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Odontomophlus Seidlitz, 1896: 240 [M]. Type species: Omophlusochraceipennis Faldermann, 1837, by subsequent designation (Bousquet et al. 2015: 141). Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini. Note: as pointed out by Bousquet et al. (2015: 141) the type species of this genus was first designated by Novák and Pettersson (2008: 335) who selected Cistelaarmillata Brullé, 1832, a species originally included in Odontomophlus Seidlitz, 1896 but also the type species of the valid genus Megischina Reitter, 1906; Löbl and Smetana (2011: 34) “retracted” the act proposed earlier by Novák and Pettersson (2008: 335); in order to conserve nomenclatural stability Bousquet et al. (2015: 141) recommended usage of Omophlusochraceipennis Faldermann, 1837 as the type species of this genus until an application be submitted to the Commission to rule on the Case.

Odontopezus Alluaud, 1889: xlv [M]. Type species [automatic]: Odontopuscostatus Silbermann, 1833, by subsequent designation (Hope 1841: 126). Status: junior synonym of Pezodontus Dejean, 1834 in Lagriinae: Pycnocerini. Note: unnecessary replacement name for Odontopus Silbermann, 1833.

Odontopus Silbermann, 1833: no 3 [M]. Type species: Odontopuscostatus Silbermann, 1833, by subsequent designation (Hope 1841: 126). Status: senior synonym of Pezodontus Dejean, 1834 in Lagriinae: Pycnocerini. Note: junior homonym of Odontopus Say, 1831 [Coleoptera: Curculionidae].

Odrotes La Rivers, 1947: 320 [M]. Type species: Edrotesarens La Rivers, 1947, by monotypy. Status: valid subgenus of Edrotes J.L. LeConte, 1851 in Pimeliinae: Edrotini.

Oeatus Champion, 1885: 111 [M]. Type species: Oeatuschevrolati Champion, 1885, by subsequent designation (Gebien 1941: 342). Status: valid genus in Stenochiinae: Cnodalonini.

Oectosis Pascoe, 1869: 149 [F]. Type species: Upiscylindrica Germar, 1848, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Oedemutes Pascoe, 1860a: 51 [M]. Type species: Oedemutestumidus Pascoe, 1860, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Oedenocera Reiche, 1862: 372 [F]. Type species [automatic]: fixed herein (ICZN 1999, Article 70.3) as Tenebriobuprestoides Fabricius, 1781, misidentified as Akislaevigata Fabricius, 1801 in the original designation by monotypy in Eschscholtz (1831). Status: junior synonym of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Löbl et al. (2008b: 192). Note: replacement name for Pachycera Eschscholtz, 1831. Note: see additional information in the entry for Pachycera Eschscholtz, 1831.

Oenomia Pascoe, 1883: 441 [F]. Type species: Oenomiafemorata Pascoe, 1883, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Oenopion Champion, 1885: 98 [M]. Type species: Oenopiongibbosus Champion, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ogoueum Pic, 1923c: 27 [N]. Type species: Ogoueumsemirufum Pic, 1923, by monotypy. Status: junior synonym of Eccoptostoma Gebien, 1913 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1962a: 64).

Ograbies Péringuey, 1899: 293 [M]. Type species: Ograbiessingularis Péringuey, 1899 (= Oncotustestaceus Solier, 1848), by monotypy. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Ohyonthis Reitter, 1898: 347 [F]. Type species: Ohyonthismicroderoides Reitter, 1898, by monotypy. Status: junior synonym of Stegastopsis Kraatz, 1865 in Pimeliinae: Tentyriini. Synonymy: Reitter (1900c: 89, 139).

Oligocara Solier, 1848: 153, 224 [N]. Type species: Oligocaranitidum Solier, 1848, by original designation. Status: valid genus in Tenebrioninae: Ulomini.

Oligorus Dejean, 1834: 206 [M]. Type species: Tageniaindica Wiedemann, 1823, by monotypy. Status: junior synonym of Luprops Hope, 1833 in Lagriinae: Lupropini. Synonymy: Lacordaire (1859a: 397, as “Lyprops”).

Oliprosodes Reitter, 1909a: 118 [M]. Type species: Prosodestrisulcata Bates, 1879, by subsequent designation (G.S. Medvedev 1999a: 851). Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Olisthaena Erichson, 1842a: 177 [F]. Type species: Olisthaenanitida Erichson, 1842, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Olocrates Mulsant, 1854: 150, 383 [M]. Type species: Opatrumgibbum Fabricius, 1775, by subsequent designation (C.G. Thomson 1859: 115, as “Omocrates”). Status: junior synonym of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina. Synonymy: Seidlitz (1898a: 828). Note: the original spelling Omocrates (p. 150) was corrected to Olocrates in the “Errata” of the same work (p. 383), Olocrates is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1); see Bouchard and Bousquet (2020a: 100).

Ologlyptus Lacordaire [in LeConte], 1858: 19 [M]. Type species [automatic]: Stenosidesgraciliformis Solier, 1836, by monotypy. Status: junior synonym of Stenosides Solier, 1836 in Pimeliinae: Asidini. Note: unnecessary replacement name for Stenosides Solier, 1836 (as Stenorides).

Omala Agassiz, 1846b: 184 [F]. Type species [automatic]: Homalapolita Eschscholtz, 1831, by monotypy. Status: junior synonym of Homala Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: unjustified emendation of Homala Eschscholtz, 1831, not in prevailing usage.

Omaleis Allard, 1876a: 4 [M]. Type species: Helopscongener Reiche, 1861, by subsequent designation (Nabozhenko 2008: 38). Status: junior synonym of Odocnemis Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: Nabozhenko (2001a: 628). Note: Allard’s original spelling Omalus, which is a junior homonym of Omalus Panzer, 1801 [Hymenoptera], was corrected to “Omaloïs” [= Omalois] in the errata for volume 14 of L’Abeille, Journal d’Entomologie (at the end of page 36 of the section “Table alphabétique”) and therefore Omalois should be considered the correct original spelling (ICZN 1999, Article 32.5.1.1); however, the incorrect subsequent spelling Omaleis, which was introduced by Allard (1877: 36), is in prevailing usage and attributed to the original author, it is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1).

Omandelostoma Purchart, 2017: 282 [N]. Type species: Adelostomamuqalensis Purchart, 2017, by original designation. Status: valid subgenus of Adelostoma Duponchel, 1827 in Pimeliinae: Adelostomini.

Omedes Broun, 1893b: 1169 [M]. Type species: Omedesnitidus Broun, 1893, by subsequent designation (R. Lucas 1920: 456). Status: valid genus in Alleculinae: Alleculini: incertae sedis.

Omegeleodes Triplehorn & Thomas, 2012: 253 [M]. Type species: Eleodesdebilis J.L. LeConte, 1858, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Ommatochara Borchmann, 1932a: 347 [F]. Type species: Ommatocharatibialis Borchmann, 1932, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Ommatophorus W.J. MacLeay, 1872: 304 [M]. Type species: Ommatophorusmastersii W.J. MacLeay, 1872, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Omocula Borchmann, 1937: 222 [F]. Type species: Alleculacollaris Borchmann, 1930, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Omolepta Fåhraeus, 1870: 320 [F]. Type species: Omoleptaelegans Fåhraeus, 1870, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Omolipus Pascoe, 1860b: 127 [M]. Type species: Omolipuscorvus Pascoe, 1860, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Omopheres Casey, 1907: 519 [M]. Type species: Omopheresfarctus Casey, 1907, by original designation. Status: valid genus and subgenus in Pimeliinae: Epitragini.

Omophlina Reitter, 1890a: 34 [F]. Type species: Omophluspodontoides Reitter, 1890, by subsequent designation (Novák and Pettersson 2008: 334). Status: valid genus in Alleculinae: Cteniopodini.

Omophlus Dejean, 1834: 213 [M]. Type species: Cistelalepturoides Fabricius, 1787, by subsequent designation (Solier 1835a: 246). Status: valid genus and subgenus in Alleculinae: Cteniopodini. Note: the earlier usage of the name Omophlus by Dahl (1823: 46) was suppressed for the purposes of zoological nomenclature by the ICZN (1964, Opinion 710).

Oncopterus Fairmaire, 1887a: 178 [M]. Type species: Oncopterusacantholophus Fairmaire, 1887, by monotypy. Status: senior synonym of Oncopteryx Gebien, 1943 in Blaptinae: Pedinini: Helopinina. Note: junior homonym of Oncopterus Steindachner, 1875 [Pisces].

Oncopteryx Gebien, 1943: 905 [F]. Type species [automatic]: Oncopterusacantholophus Fairmaire, 1887, by monotypy. Status: valid genus in Blaptinae: Pedinini: Helopinina. Note: replacement name for Oncopterus Fairmaire, 1887.

Oncosoma Westwood, 1843: 121 [N]. Type species: Oncosomagranulare Westwood, 1843 (= Pimeliagemmata Fabricius, 1801), by monotypy. Status: junior synonym of Amatodes Dejean, 1834 in Blaptinae: Pedinini: Helopinina. Synonymy: Erichson (1844: 282). Note: unjustified emendation of the original spelling Ogcosoma, introduced by Agassiz (1846b: 257, 259), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1), see Bouchard et al. (2005: 509); redescribed by Westwood (1844: 227) under the name Ogcoosoma; the original combination of the accepted name of the type species, Pimeliagemmata Fabricius, 1801, is a junior primary homonym of Pimeliagemmata Herbst, 1799.

Oncoosoma Gebien, 1911a: 563 [N]. Type species [automatic]: Oncosomagranulare Westwood, 1843 (= Pimeliagemmata Fabricius, 1801), by monotypy. Status: junior synonym of Amatodes Dejean, 1834 in Blaptinae: Pedinini: Helopinina. Note: unjustified emendation of Oncosoma Westwood, 1843 (as “Ogcosoma”), not in prevailing usage.

Oncotiphallops Koch, 1956a: 162 [M]. Type species: Oncotiphallopsbarbosai Koch, 1956, by original designation. Status: junior synonym of Anchophthalmus Gerstaecker, 1854 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (2002a: 53).

Oncotopsis Koch, 1958: 152 [F]. Type species: Nicandrabicolor Kulzer, 1951, by original designation. Status: valid subgenus of Nicandra Fairmaire, 1888 in Blaptinae: Pedinini: Helopinina.

Oncotus Blanchard, 1845: 13, 24 [M]. Type species: Oncotusfarctus Solier, 1848, by subsequent designation (Gebien 1938a: 72). Status: valid genus and subgenus in Blaptinae: Platynotini: Eurynotina. Note: originally proposed without included nominal species; Solier (1848: 216–222), by including five new species in association with the genus “Oncotus Dejean”, was the first author to subsequently and expressly include nominal species in Oncotus (ICZN 1999, Article 67.2.2).

Onocera Borchmann, 1936: 139 [F]. Type species: Ecnolagriasecurigera Borchmann, 1916, by original designation. Status: valid subgenus of Ecnolagria Borchmann, 1916 in Lagriinae: Lagriini: Lagriina.

Onoglypta Carter, 1926: 144 [F]. Type species [automatic]: Aglyptaoctocostata Gebien, 1908, by monotypy. Status: junior synonym of Aglypta Gebien, 1908 in Tenebrioninae: Heleini: Cyphaleina. Note: unnecessary replacement name for Aglypta Gebien, 1908.

Ononyctus Carter, 1914c: 382 [M]. Type species: Ononyctussulcatus Carter, 1914, by monotypy. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 473).

Onosterrhus Pascoe, 1866a: 451 [M]. Type species: Onosterrhuslaevis Pascoe, 1866, by monotypy. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 473).

Onotrichus Carter, 1911b: 138, 164 [M]. Type species: Onotrichuslateralis Carter, 1911, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Onychomira Campbell, 1984: 289 [F]. Type species: Onychomirafloridensis Campbell, 1984, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Onychosis Deyrolle, 1867: 226 [F]. Type species: Onychosisgracilipes Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Note: the name Anisosis, also described in the same publication, was used in error for Onychosis in the “Tableau des genres” on p. 81 (see Dallas 1868: 265–266).

Onymacris Allard, 1885: 157 [F]. Type species: Adesmiacandidipennis Brême, 1840, by subsequent designation (Gebien 1937a: 655). Status: valid genus in Pimeliinae: Adesmiini.

Oochila J.L. LeConte, 1862: 220 [F]. Type species: Asbolusinfaustus J.L. LeConte, 1854, by original designation. Status: junior synonym of Cryptoglossa Solier, 1837 in Pimeliinae: Cryptoglossini. Synonymy: Horn (1870: 278, with Centrioptera Mannerheim, 1843, a junior synonym of Cryptoglossa Solier, 1837).

Oochrotus P.H. Lucas, 1852: xxix [M]. Type species: Oochrotusunicolor P.H. Lucas, 1852, by monotypy. Status: valid genus in Diaperinae: Crypticini.

Oocistela Borchmann, 1908: 356 [F]. Type species: Oocistelaconvexa Borchmann, 1908, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Ooconibius Casey, 1895: 618 [M]. Type species: Notibiusopacus J.L. LeConte, 1866, by monotypy. Status: junior synonym of Conibius J.L. LeConte, 1851 in Blaptinae: Opatrini: Blapstinina. Synonymy: Aalbu in Bousquet et al. (2018: 202).

Oodeoscelis Agassiz, 1846b: 260 [F]. Type species [automatic]: Blapspolita Sturm, 1807, by subsequent designation (Kaszab 1940b: 937; see ICZN 1993b, Opinion 1729). Status: junior synonym of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini. Note: unjustified emendation of Oodescelis Motschulsky, 1845, not in prevailing usage.

Oodescelis Motschulsky, 1845a: 76 [F]. Type species: Blapspolita Sturm, 1807, by subsequent designation (Kaszab 1940b: 937; see ICZN 1993b, Opinion 1729). Status: valid genus and subgenus in Blaptinae: Platyscelidini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1993b, Opinion 1729).

Oogaster Faldermann, 1837: 30 [F]. Type species: Oogastermenetriesii Faldermann, 1837 (= Tageniapicea Ménétriés, 1832), by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Oogeton Kaszab, 1941b: 69 [M]. Type species: Oogetonnigrocoeruleus Kaszab, 1941 (= Amarygmusmakii Miwa, 1939), by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Note: name first used by Miwa (1939: 412) without type species fixation (see Masumoto 1989a: 116; Löbl et al. 2008a: 44).

Oopiestus Chevrolat, 1833b: 30, pl. 2 [M]. Type species: Oopiestusovalis Chevrolat, 1833 (= Peltoidessenegalensis Laporte, 1833), by monotypy. Status: senior synonym of Peltoides Laporte, 1833 in Tenebrioninae: Alphitobiini. Synonymy: Audoin and Milne-Edward (1835: 188). Note: the alternative original spelling Opiestus, used by Chevrolat (1833b: 30), was rejected by Chevrolat (1847a: 118) who acted as the First Reviser (ICZN 1999, Article 24.2.4); bibliographic evidence indicates that Oopiestus Chevrolat, 1833 was published by 16 March 1833 before Peltoides Laporte, 1833, issued by 1 April 1833, and should be treated as the valid name instead; an application to the ICZN is necessary to conserve usage of Peltoides Laporte, 1833 as the valid name.

Opacoplonyx Bremer, 2014a: 35 [M]. Type species: Plesiophthalmusdavidis Fairmaire, 1878, by original designation. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Opatrasida Escalera, 1922b: 69 [F]. Type species: Asidajurinei Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Polasida Reitter, 1917 in Pimeliinae: Asidini. Synonymy: Gebien (1937a: 727).

Opatresthes Gebien, 1928: 192 [F]. Type species: Opatresthesbinodosa Gebien, 1928, by subsequent designation (Gebien 1941: 817). Status: valid subgenus of Goniadera Perty, 1832 in Lagriinae: Goniaderini.

Opatrinus Dejean, 1821: 66 [M]. Type species: Opatrumclathratum Fabricius, 1787, by monotypy. Status: valid genus and subgenus in Blaptinae: Platynotini: Platynotina.

Opatroides Brullé, 1832: 219 [M]. Type species: Opatroidespunctulatus Brullé, 1832, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Opatronesthes Reitter, 1904: 174 [F]. Type species: Melanesthespunctipennis Reitter, 1889, by subsequent designation (Iwan and Löbl 2007: 734). Status: valid subgenus of Melanesthes Dejean, 1834 in Blaptinae: Opatrini: Opatrina.

Opatropis Reitter, 1904: 134, 159 [F]. Type species: Opatrumhispidum Brullé, 1839 (= Opatrumaffine Billberg, 1815), by monotypy. Status: valid subgenus of Gonocephalum Solier, 1834 in Blaptinae: Opatrini: Opatrina.

Opatrum Fabricius, 1775: 76 [N]. Type species: Silphasabulosa Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Blaptinae: Opatrini: Opatrina.

Ophthalmosis Deyrolle, 1867: 81, 229 [F]. Type species: Ophthalmosislongipes Deyrolle, 1867, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Note: the original spelling Ophtalmosis (pp. 77, 229, 231) was corrected to Ophthalmosis in a footnote in the “Table des matières” of the same work (p. clxiv), Ophthalmosis is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Opigenia Pascoe, 1869: 288 [F]. Type species: Opigeniairidescens Pascoe, 1869 (= Platyphanesvittatus Westwood, 1849), by monotypy. Status: junior synonym of Platyphanes Westwood, 1849 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 454).

Opisthoblaps Kolbe, 1928: 201 [F]. Type species: Blaps sulcifera Seidlitz, 1893, by subsequent designation (Nabozhenko and Chigray 2020: 10). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Nabozhenko and Chigray (2020: 10).

Oplocephala Laporte & Brullé, 1831: 332, 338 [F]. Type species: Ips haemorrhoidalis Fabricius, 1787, by subsequent designation (Motschulsky 1845a: 80). Status: junior synonym of Neomida Latreille, 1829 in Diaperinae: Diaperini: Diaperina. Synonymy: Dejean (1834: 197).

Oplocheirus Klug, 1835: 40 [M]. Type species: Helopscarbonarius Klug, 1835 (= Acanthomerusstriatus Guérin-Méneville, 1834), by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Oploptera Chevrolat in Guérin-Méneville, 1844: 126 [F]. Type species: Strongyliumserraticorne Guérin-Méneville, 1834, by monotypy. Status: valid genus and subgenus in Stenochiinae: Stenochiini.

Opostirus Kirsch, 1865: 45 [M]. Type species: Opostirusexsectus Kirsch, 1865, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina. Note: transferred from Zopheridae: Colydiinae by Ivie et al. (2016: 759).

Oppenheimeria Koch, 1952a: 110 [F]. Type species: Oppenheimeriabombophthalma Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Evaniosomini.

Oracula Novák, 2019f: 54 [F]. Type species: Oraculabicolor Novák, 2019, by original designation. Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Orarabion Leo & Liberto, 2011: 157 [N]. Type species: Orarabiondominici Leo & Liberto, 2011, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Orchesiolobopoda Pic, 1919: 6 [F]. Type species: Orchesiolobopodaminutissima Pic, 1919, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Orcopagia Pascoe, 1868: xii [F]. Type species: Orcopagiamonstrosa Pascoe, 1868, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Oremasis Pascoe, 1866a: 470 [M]. Type species: Adeliumcupreum Gray, 1831, by original designation. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 490).

Oreogria Merkl, 1988b: 248 [F]. Type species: Oreogriakaszabi Merkl, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Oreomelasma Español, 1975: 238, 240 [N]. Type species: Oreomelasmaoromii Español, 1975, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Orghidania Ardoin, 1977: 383 [F]. Type species: Orghidaniatorrei Ardoin, 1977, by monotypy. Status: senior synonym of Spelaebiosis Bousquet & Bouchard, 2018 in Tenebrioninae: Triboliini. Note: junior homonym of Orghidania Capuse, 1971 [Lepidoptera].

Orientocara Koch, 1952a: 176 [N]. Type species: Stenocaraarachnoides Gerstaecker, 1854, by original designation. Status: valid genus in Pimeliinae: Adesmiini.

Orientochile Penrith & Endrödy-Younga, 1994: 82 [F]. Type species: Cryptochileelegans Gerstaecker, 1854, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Orobychus Pascoe, 1868: xii [M]. Type species: Orobychuslacordairii Pascoe, 1868 (= Taphrosomadohrnii Kirsch, 1866), by monotypy. Status: junior synonym of Taphrosoma Kirsch, 1866 in Stenochiinae: Cnodalonini. Synonymy: Champion (1885: 108).

Orocina Reitter, 1897a: 303 [F]. Type species: Orocinacapnisiceps Reitter, 1897, by subsequent designation (R. Lucas 1920: 463). Status: junior synonym of Syachis Bates, 1879 in Pimeliinae: Tentyriini. Synonymy: Bogdanov-Katjkov (1916: 68).

Orophylaxus Koch, 1948: 417 [M]. Type species: Phylaxincertus Mulsant & Godart, 1866, by original designation. Status: valid subgenus of Otinia Antoine, 1942 in Blaptinae: Dendarini: Melambiina.

Oroptera Borchmann, 1916a: 48, 104 [F]. Type species: Oropteraphysoptera Borchmann, 1916, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Orostegastopsis Koch, 1962c: 255 [F]. Type species: Stegastopsisscorteccii Koch, 1962, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Orphelops Gozis, 1910: 102 [M]. Type species: Helopsimpressicollis Faldermann, 1837 (= Helopsfaldermanni Faldermann, 1837), by original designation. Status: junior synonym of Nalassus Mulsant, 1854 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: new synonym [YB]. Note: this genus-group taxon has been forgotten in the literature; its type species is currently placed in the nominotypical subgenus of Nalassus Mulsant, 1854 and for that reason Gozis’s name is regarded as a new junior synonym of Nalassus.

Ortheolus Casey, 1907: 380 [M]. Type species: Schoenicusoculatus Champion, 1884, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Orthogonoderes Solier, 1841a: 233 [M]. Type species: Praocissubreticulatus Solier, 1841, by subsequent designation (Flores and Pizarro-Araya 2014: 68). Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini.

Orthonychius Gebien, 1926: 83 [M]. Type species: Orthonychiusdigitatus Gebien, 1926, by monotypy. Status: junior synonym of Trilobocara Solier, 1851 in Pimeliinae: Trilobocarini. Synonymy: Doyen (1994: 501).

Orthostibia Blaisdell, 1923: 235 [F]. Type species: Orthostibiafrontalis Blaisdell, 1923, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Osdara Walker, 1858: 284 [F]. Type species: Osdarapicipes Walker, 1858, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Osdaroides Kaszab, 1980b: 324 [M]. Type species: Osdaroidesmetallicus Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: Osdaroides was used earlier by Kaszab (1979a: 108) without a description, a definition or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Ospidus Pascoe, 1866a: 467 [M]. Type species: Ospiduschrysomeloides Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Ossiporis Pascoe, 1866a: 451 [F]. Type species: Ossiporisterrena Pascoe, 1866, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Osternus Fairmaire, 1895b: 446 [M]. Type species: Osternusopacicollis Fairmaire, 1895, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Ostorius Fairmaire, 1889c: xxxiii [M]. Type species: Ostoriusmulticostatus Fairmaire, 1889, by monotypy. Status: junior synonym of Adelonia Laporte, 1840 in Blaptinae: Pedinini: Leichenina. Synonymy: Kamiński et al. (2019a: 92).

Oterophloeus Desbrochers des Loges, 1881: 52 [M]. Type species: Oterophloeuspicipes Desbrochers des Loges, 1881 (= Pachychilahumerosus Fairmaire, 1875), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Oteroscelis Solier, 1835b: 546 [F]. Type species: Adesmiapulcherrima Solier, 1835 (= Adesmiaaudouini Solier, 1835), by subsequent designation (Hope 1841: 118). Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Oteroscelopsis Löbl & Merkl in Löbl et al., 2020: 1 [F]. Type species: Pimeliadilatata Klug, 1830, by original designation. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: name first proposed by Koch (1944b: 147) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 244) designated Pimeliadilatata Klug, 1830 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Othelecta Pascoe, 1866a: 488 [F]. Type species: Othelectatorrida Pascoe, 1866, by monotypy. Status: junior synonym of Cylindrothorus Solier, 1843 in Alleculinae: Alleculini: Mycetocharina. Synonymy: Haag-Rutenberg (1879c: 412).

Othryades Champion, 1889: 72 [M]. Type species: Othryadesfragilicornis Champion, 1889, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Othryoneus Champion, 1886: 245 [M]. Type species: Othryoneuserotyloides Champion, 1886, by subsequent designation (Gebien 1942a: 315). Status: valid genus in Stenochiinae: Cnodalonini.

Otinia Antoine, 1942: 22, 44 [F]. Type species: Otiniaiblanensis Antoine, 1942, by monotypy. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Otoceromorphus Pic, 1915d: 11 [M]. Type species: Otoceromorphusgounellei Pic, 1915, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Otocerus Mäklin, 1867: 484 [M]. Type species [automatic]: Strongyliumserraticorne Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Oploptera Chevrolat, 1844 in Stenochiinae: Stenochiini. Note: unnecessary replacement name for Oploptera Chevrolat, 1844 (as “Hoploptera”).

Otrintus Pascoe, 1866a: 483 [M]. Type species: Prosodesbehrii Germar, 1848, by original designation. Status: junior synonym of Cardiothorax Motschulsky, 1860 in Lagriinae: Adeliini. Synonymy: Bates (1879c: 30).

Otys Champion, 1895a: 221 [M]. Type species: Otysharpalinus Champion, 1895 (= Scaletomerusproximus Blackburn, 1891), by subsequent designation (R. Lucas 1920: 468). Status: junior synonym of Scaletomerus Blackburn, 1891 in Alleculinae: Alleculini: Alleculina. Synonymy: Carter (1915a: 78). Note: Matthews and Bouchard (2008: 333) selected Otysarmatus Champion, 1895 as the type species of this genus (and treated Otys Champion, 1895 as valid) not knowing of the earlier valid designation by R. Lucas (1920: 468).

Oubanghinum Pic, 1933: 4 [N]. Type species: Oubanghinumatrum Pic, 1933, by monotypy. Status: junior synonym of Heterotarsus Latreille, 1829 in Blaptinae: Opatrini: Heterotarsina. Synonymy: Ardoin (1969b: 126).

Ovalobioramix Egorov, 2004: 603 [F]. Type species: Platyscelismolesta Bogatchev, 1947, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Ovaloodescelis Kaszab, 1940b: 940, 947 [F]. Type species: Platyscelisaffinis Seidlitz, 1893, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Overlaetia Pic, 1937b: 304 [F]. Type species: Overlaetiagracilitarsis Pic, 1937, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: as pointed out by Bremer and Lillig (2014: 118) the older name Overlaetia Schouteden, 1932 [Hemiptera] is not nomenclaturally available and therefore Overlaetia Pic, 1937 is not a junior homonym.

Oxidates Champion, 1886: 263 [M]. Type species: Oxidatesplanicollis Champion, 1886, by subsequent designation (Gebien 1943: 402). Status: valid genus in Stenochiinae: Cnodalonini.

Oxinthas Champion, 1884: 72 [M]. Type species: Oxinthaspraocioides Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Branchini.

Oxura W. Kirby, 1819a: 413 [F]. Type species: Oxurasetosa W. Kirby, 1819, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Oxycara Solier, 1835b: 253, 254 [N]. Type species: Oxycarablapsoides Solier, 1835, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Oxycarops Reitter, 1900c: 94 [M]. Type species: Hegeterfuscipes Brullé, 1839, by subsequent designation (Gebien 1937a: 640). Status: valid genus in Pimeliinae: Tentyriini.

Oxycerus Koch, 1955a: 46 [M]. Type species: Trachynotusresolutus Péringuey, 1904, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Oxyge Chatanay, 1914b: 7 [F]. Type species: Oxygerugosa Chatanay, 1914, by original designation. Status: valid genus in Pimeliinae: Asidini.

Oxygonodera Casey, 1907: 433, 444 [F]. Type species: Oxygonoderavillosa Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Oxypistoma Löbl, Bouchard, Merkl & Bousquet, 2020: 5 [N]. Type species: Prochomabucculentum Koch, 1940, by original designation. Status: valid subgenus of Prochoma Solier, 1835 in Pimeliinae: Tentyriini. Note: name first proposed with two alternative spellings, as Oxypistoma (Koch 1940a: 258, 259, 260, 262) and Oxipistoma (Koch 1940a: 259), without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl et al. (2008a: 40) designated Prochomabucculenta Koch, 1940 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Oxythorax Fåhraeus, 1870: 288 [M]. Type species: Oxythoraxclathratus Fåhraeus, 1870, by monotypy. Status: junior synonym of Anchophthalmus Gerstaecker, 1854 in Blaptinae: Platynotini: Platynotina. Synonymy: Péringuey (1904: 238).

Oxyura Agassiz, 1846b: 267, 268 [F]. Type species [automatic]: Oxurasetosa W. Kirby, 1819, by monotypy. Status: junior synonym of Oxura W. Kirby, 1819 in Pimeliinae: Sepidiini: Oxurina. Note: unjustified emendation of Oxura W. Kirby, 1819, not in prevailing usage; junior homonym of Oxyura Bonaparte, 1831 [Aves].

Oyanus Pic, 1921: 23 [M]. Type species: Oyanuscurticornis Pic, 1921, by monotypy. Status: junior synonym of Cleomis Fairmaire, 1892 in Stenochiinae: Cnodalonini. Synonymy: Löbl et al. (2008b: 340).

Ozaenimorphus Fairmaire, 1882b: 127 [M]. Type species: Ozaenimorphuscostulipennis Fairmaire, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Ozolais Pascoe, 1866a: 457 [F]. Type species: Ozolaisscruposa Pascoe, 1866, by monotypy. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Ozotypoides Kaszab, 1982b: 222 [M]. Type species: Ozotypoidesgranulatus Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Adeliini.

Ozotypus Pascoe, 1862: 328 [M]. Type species: Ozotypussetosus Pascoe, 1862, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pachacamacius Flores & Giraldo-Mendoza in Giraldo-Mendoza and Flores, 2019: 84 [M]. Type species: Pachacamaciusaguilari Giraldo-Mendoza & Flores, 2019, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Pachycera Eschscholtz, 1831: 5, 7 [F]. Type species: fixed herein (ICZN 1999, Article 70.3) as Tenebriobuprestoides Fabricius, 1781, misidentified as Akislaevigata Fabricius, 1801 in the original designation by monotypy in Eschscholtz (1831). Status: junior synonym of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Löbl et al. (2008b: 192). Note: junior homonym of Pachycera Billberg, 1820 [Hemiptera]; the type species “Akislaevigata Fabricius” was first established by monotypy; Koch (1943: 530) noted that Akislaevigata Fabricius of Eschscholtz (1831) was misidentified and corresponded to the species Tenebriobuprestoides Fabricius, 1781; we follow currently accepted concepts (e.g., Löbl et al. 2008b: 192) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Akislaevigata Fabricius, 1801 also belongs to the genus Hyperops Eschscholtz, 1831; the original combination of the accepted name of the type species, Tenebriobuprestoides Fabricius, 1781, is a junior primary homonym of Tenebriobuprestoides Scopoli, 1763 and Tenebriobuprestoides Pallas, 1773.

Pachycerops Koch, 1943a: 524, 533 [M]. Type species: Pachycerainsidiosa Fairmaire, 1896, by monotypy. Status: valid subgenus of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Pachycerus Montrouzier, 1860: 291 [M]. Type species: Pachycerusdomesticus Montrouzier, 1860, by monotypy. Status: senior synonym of Sciophagus Sharp, 1885 in Diaperinae: Diaperini: Diaperina. Synonymy: Fauvel (1904: 185). Note: junior homonym of Pachycerus Schönherr, 1823 [Coleoptera: Curculionidae].

Pachychila Eschscholtz, 1831: 5 [F]. Type species: Pimeliapunctata Fabricius, 1798, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: the original combination of the name of the type species, Pimeliapunctata Fabricius, 1798, is a junior primary homonym of Pimeliapunctata Thunberg, 1787.

Pachychile Lacordaire, 1859a: 46 [F]. Type species [automatic]: Pimeliapunctata Fabricius, 1798, by monotypy. Status: junior synonym of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: unjustified emendation of Pachychila Eschscholtz, 1831, not in prevailing usage. Note: the original combination of the name of the type species, Pimeliapunctata Fabricius, 1798, is a junior primary homonym of Pimeliapunctata Thunberg, 1787.

Pachychilina Reitter, 1900c: 91, 145 [F]. Type species: Tentyriadejeani Besser, 1832, by subsequent designation (R. Lucas 1920: 473). Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Pachycoelia Boisduval, 1835: 248 [F]. Type species: Pachycoeliasulcicollis Boisduval, 1835, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: name incorrectly listed as preoccupied in the literature (e.g., Matthews and Bouchard 2008: 296) but used as valid recently (Matthews and Lawrence 2019: 632).

Pachycossyphodes Andreae, 1961: 203, 214 [M]. Type species: Cossyphodesmachadoi Basilewsky, 1952, by original designation. Status: junior synonym of Cossyphodes Westwood, 1851 in Pimeliinae: Cossyphodini: Cossyphodina. Synonymy: Schawaller (2013c: 362, implied by inclusion of Cossyphodesmachadoi Basilewsky, 1952 in Cossyphodes Westwood, 1851 without use of a subgenus rank).

Pachycyrtosoma Marcuzzi, 1999: 81 [N]. Type species: Cyrtosomamerkli Marcuzzi, 1999, by original designation. Status: junior synonym of Nesocyrtosoma Marcuzzi, 1976 in Stenochiinae: Cnodalonini. Synonymy: Hopp and Ivie (2009: 13).

Pachylagria Borchmann, 1912a: 17 [F]. Type species: Pachylagriaovata Borchmann, 1912, by subsequent designation (Merkl 2008: 115). Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Borchmann (1916a: 163, with Heterogria Fairmaire, 1896, a junior synonym of Xanthalia Fairmaire, 1894).

Pachylesthus Fairmaire, 1897e: 219 [M]. Type species: Pachylesthusvalidus Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pachylocerus Hope, 1841: 186 [M]. Type species: Pachyloceruswestermanni Hope, 1841, by monotypy. Status: senior synonym of Pycnocerus Westwood, 1841 in Lagriinae: Pycnocerini. Note: junior homonym of Pachylocerus Hope, 1834 [Coleoptera: Cerambycidae].

Pachylodera Quedenfeldt, 1890: 399 [F]. Type species: Pachyloderabrevicornis Quedenfeldt, 1890, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Pachymastus Fairmaire, 1896b: 350 [M]. Type species: Pachymastusasperulus Fairmaire, 1896, by monotypy. Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Pachynotelus Solier, 1841a: 210, 267 [367] [M]. Type species: Pachynotelusalbiventris Solier, 1841, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Cryptochilina.

Pachyphaleria Gebien, 1920: 136 [F]. Type species: Phaleriacapensis Laporte, 1840, by monotypy. Status: valid genus in Diaperinae: Phaleriini.

Pachypterocoma Skopin, 1974b: 157 [F]. Type species: Pterocomapallasi Semenov-Tjan-Shansky, 1910, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Pachypterus P.H. Lucas, 1847: pl. 29 [M]. Type species: Pachypterusmauritanicus P.H. Lucas, 1847, by monotypy. Status: senior synonym of Neopachypterus Bouchard, Löbl & Merkl, 2007 in Blaptinae: Opatrini: Neopachypterina. Note: junior homonym of Pachypterus Swainson, 1839 [Pisces].

Pachyscelina Kwieton, 1978: 29, 30 [F]. Type species: Pachyscelismicros Kaszab, 1970, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Pachyscelis Solier, 1836: 9, 54 [F]. Type species [automatic]: Pimeliamusiva Ménétriés, 1832, by subsequent designation (Bouchard et al. 2007: 388). Status: valid genus and subgenus in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Brachyscelis Dejean, 1834. Note: nomen protectum (see Bouchard et al. 2007: 388).

Pachyscelodes Sénac, 1887: 189 [M]. Type species: Pachyscelishenoni Sénac, 1887, by subsequent designation (Gebien 1937a: 823). Status: junior synonym of Scelace Marseul, 1887 in Pimeliinae: Pimeliini. Synonymy: Reitter (1915: 2).

Pachysternoplax Skopin, 1973: 110, 154 [F]. Type species: Trigonoscelisarmeniaca Faldermann, 1837, by original designation. Status: valid subgenus of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini.

Pachystira Chen, 1997: 308 [F]. Type species: Pachystiraimpressipennis Chen, 1997, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Pachyurgus J.L. LeConte, 1862: 230 [M]. Type species: Iphthinusaereus Melsheimer, 1846, by original designation. Status: junior synonym of Derosphaerus J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: J.L. LeConte (1873: 335, with Encyalesthus Motschulsky, 1860, a junior synonym of Derosphaerus J. Thomson, 1858). Note: the type locality for Iphthinusaereus Melsheimer, 1846 was incorrectly given as “Pennsylvania” originally (see Bousquet et al. 2018: 384).

Pactostoma J.L. LeConte, 1858b: 19 [N]. Type species: Asidaanastomosis Say, 1824, by original designation. Status: junior synonym of Stenosides Solier, 1836 in Pimeliinae: Asidini. Synonymy: J.L. LeConte (1862: 222, with Ologlyptus Lacordaire, 1858, a junior synonym of Stenosides Solier, 1836).

Paita Fauvel, 1904: 173 [F]. Type species: Paitasetosella Fauvel, 1904, by monotypy. Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Kaszab (1982b: 56).

Paivaea Wollaston, 1864: 449 [F]. Type species: Tentyriahispida Brullé, 1839, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Paivea Scudder, 1882: 213 [F]. Type species [automatic]: Tentyriahispida Brullé, 1839, by monotypy. Status: junior synonym of Paivaea Wollaston, 1864 in Pimeliinae: Tentyriini. Note: unjustified emendation of Paivaea Wollaston, 1864, not in prevailing usage.

†Palaeobasanus Nabozhenko & Kirejtshuk, 2020: 25 [M]. Type species: Palaeobasanusneli Nabozhenko & Kirejtshuk, 2020, by original designation. Status: valid genus in Diaperinae: Scaphidemini. Note: described from Middle-Upper Paleocene deposits (France).

†Palaeosclerum Nabozhenko & Kirejtshuk, 2017: 308 [N]. Type species: Palaeosclerumpohli Nabozhenko & Kirejtshuk, 2017, by original designation. Status: valid genus in Blaptinae: Opatrini: Sclerina. Note: described from Middle-Upper Paleocene deposits (France).

Palembomimus Matthews & Lawrence, 2005: 539 [M]. Type species: Platydemadeplanata Champion, 1894, by original designation. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Palembus Casey, 1891: 65 [M]. Type species: Palembusocularis Casey, 1891, by monotypy. Status: junior synonym of Ulomoides Blackburn, 1888 in Diaperinae: Diaperini: Diaperina. Synonymy: Doyen et al. (1990: 237).

Palorinus Blair, 1930: 135 [M]. Type species: Palorushumeralis Gebien, 1914, by original designation. Status: valid genus in Tenebrioninae: Palorini.

Paloropsis Masumoto & Grimm, 2004: 127 [F]. Type species: Paloropsisirei Masumota & Grimm, 2004, by original designation. Status: valid genus in Tenebrioninae: Palorini.

Palorus Mulsant, 1854: 250 [M]. Type species: Hypophlaeusdepressus Fabricius, 1790, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Palpafrina Koch, 1950b: 329 [F]. Type species: Afrinuswatsoni Koch, 1950, by original designation. Status: valid subgenus of Afrinus Fairmaire, 1888 in Pimeliinae: Tentyriini.

Palpichara Borchmann, 1932a: 355 [F]. Type species: Palpicharaserricornis Borchmann, 1932, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Palpicula Novák, 2018a: 168 [F]. Type species: Alleculafiliola Borchmann, 1925, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Palpomodes Koch, 1952d: 223 [M]. Type species: Psammodesphysopterus Gebien, 1920, by monotypy. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Oxurina.

Pandarinus Mulsant & Rey, 1854: 50, 103 [M]. Type species: Pandarinustenellus Mulsant & Rey, 1854, by subsequent designation (Chatzimanolis and Löbl 2003: 260). Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Pandarus Dejean, 1834: 191 [M]. Type species [automatic]: Type species: Helopstristis Rossi, 1790, by subsequent designation Blanchard (1844: pl. 48). Status: junior synonym of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina. Note: unjustified emendation of Dendarus Dejean, 1821, not in prevailing usage (see Bousquet and Bouchard 2013a: 48); junior homonym of Pandarus Leach, 1816 [Crustacea].

Paniasis Champion, 1886: 208 [M]. Type species: Paniasisdilatipes Champion, 1886, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Paoligena Pic, 1928c: 42 [F]. Type species: Praogenainhumeralis Pic, 1928, by monotypy. Status: valid genus in Tenebrioninae: Paoligenini. Note: originally described as a subgenus of Praeugena Laporte, 1840, elevated to the rank of genus by De Moor (1970: 15).

Papuamisolampus Kaszab, 1986: 288 [M]. Type species: Papuamisolampustoxopeusi Kaszab, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Parabantodemus Iwan, 2000: 345 [M]. Type species: Trigonopusspinipes Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Parabigopsis Español, 1946: 111, 114 [F]. Type species: Parabigopsispeyerimhoffi Español, 1946, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Parablops Rottenberg, 1871: 254 [M]. Type species: Parablopsaetnensis Rottenberg, 1871, by monotypy. Status: senior synonym of Gerandryus Rottenberg, 1873 in Alleculinae: Alleculini: Gonoderina. Note: junior homonym of Parablops Schönherr, 1839 [Coleoptera: Anthribidae].

Parabolitophagus Miyatake, 1964: 67, 70 [M]. Type species: Bolitophagusfelix Lewis, 1894, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini.

Paracirta Schuster, 1930: 66 [F]. Type species: Paracirtaschatzmayri Schuster, 1930, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Paracistela Borchmann, 1941a: 30 [F]. Type species: Paracistelavariabilis Borchmann, 1941, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Paracossyphus Viñolas & Cartagena, 2005: 29, 55 [M]. Type species [automatic]: Cossyphusminutissimus Laporte, 1840, by original designation. Status: junior synonym of Acontodactylus Desbrochers des Loges, 1894 in Lagriinae: Cossyphini. Note: unnecessary replacement name for Acontodactylus Desbrochers des Loges, 1894 (as “Acanthodactylus”).

Paradissonomus G.S. Medvedev, 1968a: 229 [M]. Type species: Dissonomuslongulus Bogatchev & Kryzhanovsky, 1960, by original designation. Status: valid subgenus of Dissonomus Jacquelin du Val, 1861 in Tenebrioninae: Dissonomini.

Paradrus Jakobson, 1924: 242 [M]. Type species [automatic]: Pseudhadrusseriatus Kolbe, 1910, by subsequent designation (Gebien 1941: 332). Status: junior synonym of Pseudhadrus Kolbe, 1910 in Stenochiinae: Cnodalonini. Note: unnecessary replacement name for Pseudhadrus Kolbe, 1910.

Paragena Bouchard & Bousquet, new subgenus [F]. Type species: Nesogenaviridicuprea Fairmaire, 1868, by present designation. Status: valid subgenus of Nesogena Mäklin, 1863 in Tenebrioninae: Praeugenini. Note: first proposed by Froussart (1961: 60) without type species designation; the subgenusParagena, which is currently used as valid, is therefore unavailable (ICZN 1999, Articles 13.3, 16.1); we hereby make the name available by selecting Nesogenaviridicuprea Fairmaire, 1868 as type species and referring to Froussart (1961: 60) for the character states that characterise and differentiate Paragena.

Paragonocnemis Kraatz, 1899: 118 [F]. Type species: Paragonocnemissculpticollis Kraatz, 1899 (= Gonocnemisfoveicollis Fairmaire, 1891), by subsequent designation (Gebien 1943: 918). Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Paraguania Marcuzzi, 1953: 31 [F]. Type species: Paraguaniarelicta Marcuzzi, 1953, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Parahymenorus Campbell, 1971: 100 [M]. Type species: Parahymenorusmetallicus Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Parahyocis Kaszab, 1955b: 650 [M]. Type species: Hyocischampioni Fauvel, 1904, by original designation. Status: valid genus in Diaperinae: Hyociini: Hyociina.

Paraisomira Dubrovina, 1982: 138 [F]. Type species: Cistelaoculata Marseul, 1876, by original designation. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

†Parakeleusticus Haupt, 1950: 114, 126 [M]. Type species: Parakeleusticuspostumus Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Paraleptodes G.S. Medvedev, 1967: 354 [M]. Type species: Leptodeslindbergi Kaszab, 1959, by original designation. Status: valid subgenus of Leptodes Dejean, 1834 in Pimeliinae: Leptodini.

Paralitoborus Antoine, 1931: 190 [M]. Type species: Litoborussternalis Fairmaire, 1884, by original designation. Status: valid subgenus of Litoborus Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Paralorelopsis Marcuzzi, 1994: 117 [F]. Type species: Paralorelopsisbordoni Marcuzzi, 1994, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Paralyreus Grouvelle, 1918: 24 [M]. Type species: Paralyreusscotti Grouvelle, 1918, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Paramarygmus Quedenfeldt, 1885: 25 [M]. Type species: Paramarygmusnigroaeneus Quedenfeldt, 1885 (= Hybonotusfemoralis Imhoff, 1843), by monotypy. Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Paramellon C.O. Waterhouse, 1882b: iv [N]. Type species: Paramellonsociale C.O. Waterhouse, 1882, by monotypy. Status: valid genus in Pimeliinae: Cossyphodini: Paramellonina.

Paramellops Andreae, 1961: 201 [M]. Type species: Cossyphodesbewicki Wollaston, 1861, by original designation. Status: valid genus in Pimeliinae: Cossyphodini: Cossyphodina.

Paramisolampidius Merkl & Masumoto in Löbl et al., 2020: 4 [M]. Type species: Paramisolampidiuskagoshimensis Nakane, 1968, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: name first proposed by Nakane (1968: 82) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Merkl and Masumoto (2007: 1) designated Paramisolampidiuskagoshimensis Nakane, 1968 as the type species of Nakane’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Paranemia Heyden, 1892: 103 [F]. Type species: Paranemiaschroederi Heyden, 1892, by monotypy. Status: valid genus in Diaperinae: Phaleriini.

Paranephodes Antoine, 1955: 208 [M]. Type species: Nephodescamusi Antoine, 1955, by monotypy. Status: valid subgenus of Nephodinus Gebien, 1943 in Tenebrioninae: Helopini: Helopina.

Paranopidium Dajoz, 1974: 429, 434 [N]. Type species: Paranopidiumafricanum Dajoz, 1974, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Parapachynotela Koch, 1952c: 54 [F]. Type species: Parapachynotelabushmanica Koch, 1952, by original designation. Status: junior synonym of Horatoma Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Synonymy: Penrith and Endrödy-Younga (1994: 10).

Parapachyscelis Kwieton, 1978: 29 [F]. Type species: Pimeliavillosa Drapiez, 1820, by original designation. Status: valid subgenus of Pachyscelis Solier, 1836 in Pimeliinae: Pimeliini.

Paraphanes W.J. MacLeay, 1887: 308 [M]. Type species: Paraphanesnitidus W.J. MacLeay, 1887, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Paraphloeus Seidlitz, 1894: 553, 557 [M]. Type species: Hypophlaeuslongulus Gyllenhal, 1827, by subsequent designation (G.S. Medvedev 1990: 231). Status: junior synonym of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini. Synonymy: Bremer (1995: 3).

Paraplatyope Löbl, Bouchard, Merkl & Bousquet, 2020: 4 [F]. Type species: Leucolaephusarabicus Blair, 1931, by original designation. Status: valid genus in Pimeliinae: Pimeliini. Note: name first proposed by Gridelli (1953: 45) without fixation of a type species in the original publication (ICZN 1999, Article 13.3).

Paraplatyscelis Kaszab, 1940b: 912, 936 [F]. Type species: Platyscelissinuata Seidlitz, 1893, by original designation. Status: valid subgenus of Platyscelis Latreille, 1818 in Blaptinae: Platyscelidini.

Parapraocis Flores & Giraldo, 2020: 35 [M]. Type species: Praocisvagecostatus Fairmaire, 1902, by original designation. Status: valid genus in Pimeliinae: Praociini. Note: name originally proposed by Kulzer (1958a: 13, 58) as subgenus of Praocis Eschscholtz, 1829 without type species designation; Flores and Pizarro-Araya (2012: 4) designated Praocisvagecostatus Fairmaire, 1902 as type species, but did not explicitely propose the taxon as new (ICZN 1999, Article 16.1).

Paraprosodes Reitter, 1909a: 119 [M]. Type species: Prosodestriangulipes Reitter, 1909, by original designation. Status: junior synonym of Prosodella Reitter, 1909 in Blaptinae: Blaptini: Prosodina. Synonymy: G.S. Medvedev (1997: 597).

Parapterocoma Skopin, 1974b: 149 [F]. Type species: Pterocomavittata Frivaldszky, 1890, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Paraselinus Kamiński, 2013: 705, 711 [M]. Type species: Paraselinusiwani Kamiński, 2013, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Parasida Casey, 1912: 76, 126 [F]. Type species: Parasidalaciniata Casey, 1912, by original designation. Status: senior synonym of Pleisiasida Smith, 2013 in Pimeliinae: Asidini. Note: junior homonym of Parasida Daday, 1904 [Crustacea].

Parasternoplax Skopin, 1973: 110, 148 [F]. Type species: Pimeliadeplanata Krynicki, 1832, by original designation. Status: valid subgenus of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini.

Parastizopus Gebien, 1938b: 93 [M]. Type species: Parastizopusdiehli Gebien, 1938, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Parastrongylium Kaszab, 1977b: 10, 24 [N]. Type species: Strongyliumlorentzi Gebien, 1921, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Paratenetus Spinola, 1845: 116 [M]. Type species: Paratenetuspunctatus Spinola, 1845, by subsequent designation (R. Lucas 1920: 483). Status: valid genus in Lagriinae: Goniaderini.

Paratoxicum Champion, 1894a: 380 [N]. Type species: Paratoxicumiridescens Champion, 1894, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Paravius Casey, 1907: 332 [M]. Type species: Emmenastusmarginatus Casey, 1890, by monotypy. Status: valid subgenus of Hylocrinus Casey, 1907 in Pimeliinae: Edrotini.

Parecatus Fairmaire, 1900c: 245 [M]. Type species: Parecatusplicatulus Fairmaire, 1900, by subsequent designation (Gebien 1937a: 741). Status: junior synonym of Scotinesthes Fairmaire, 1895 in Pimeliinae: Asidini. Synonymy: Koch (1962a: 67).

Parepitragus Casey, 1907: 518 [M]. Type species: Parepitragussolieri Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Pareupezus Kolbe, 1889: 129 [M]. Type species: Pareupezusglaber Kolbe, 1889, by monotypy. Status: junior synonym of Paramarygmus Quedenfeldt, 1885 in Tenebrioninae: Amarygmini. Synonymy: Gebien (1921b: 148).

Parimmedia Gebien, 1928: 220, 226 [F]. Type species: Parimmediaepipleuralis Gebien, 1928, by subsequent designation (Gebien 1943: 404). Status: valid genus in Stenochiinae: Cnodalonini.

Parmularia Koch, 1955a: 35 [F]. Type species: Psammodescaffer Fåhraeus, 1870, by monotypy. Status: junior synonym of Psammodes W. Kirby, 1819 in Pimeliinae: Sepidiini: Molurina. Synonymy: Kamiński et al. (2019b: 31). Note: junior homonym of Parmularia MacGillivray, 1887 [Bryozoa].

Paroderus Mulsant & Rey, 1854: 111 [M]. Type species: Pandarinuselongatus Mulsant & Rey, 1854, by subsequent designation (Chatzimanolis and Löbl 2003: 260). Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Paroeatus Gebien, 1928: 169, 178 [M]. Type species: Paroeatusopacus Gebien, 1928, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Parogria Borchmann, 1936: 119 [F]. Type species: Lagriaatrata Borchmann, 1912, by original designation. Status: valid subgenus of Cerogria Borchmann, 1911 in Lagriinae: Lagriini: Lagriina.

Paromophlus Iablokoff-Khnzorian, 1983: 134, 139 [M]. Type species: Cistelapicipes Fabricius, 1792, by original designation. Status: junior synonym of Phibalus Gistel, 1856 in Alleculinae: Cteniopodini. Synonymy: Bousquet et al. (2015: 142). Note: the name Paromophlus was first introduced by Znojko in Ogloblin and Znojko (1950: 125) without designation of a type species and is therefore unavailable from that date.

†Paropiophorus Haupt, 1950: 114, 132 [M]. Type species: Paropiophorusnitidus Haupt, 1950, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Middle Eocene deposits (Germany); this genus was previously considered to be “definitely not a tenebrionid” by Watt (1975: 389) but is included here following Nabozhenko (2019: 8).

Partystona Watt, 1992: 47 [F]. Type species: Partystonametallica Watt, 1992, by original designation. Status: valid genus in Tenebrioninae: Titaenini.

Passalocharis Koch, 1954b: 13 [F]. Type species: Chirocharisintermedius Gebien, 1911, by original designation. Status: valid genus in Lagriinae: Pycnocerini.

Patagonogenius Flores, 2000a: 371, 390 [M]. Type species: Mitrageniusquadricollis Fairmaire, 1876, by original designation. Status: valid genus in Pimeliinae: Nycteliini.

Patagonopraocis Flores & Chani-Posse, 2005: 576 [M]. Type species: Patagonopraocismagellanicus Flores & Chani-Posse, 2005, by original designation. Status: valid genus in Pimeliinae: Praociini.

Paulianaria Bouchard & Bousquet, new genus [F]. Type species: Paulianariastrongylioides Ardoin, 1961, by present designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: Ardoin (1961d: 97) introduced the new genus name Paulianaria for five nominal species, but unfortunately did not designate a type species; the genus Paulianaria, which has been treated as valid since 1961, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Paulianariastrongylioides Ardoin, 1961 as type species and referring to Ardoin (1971: 362) for the character states that characterise and differentiate Paulianaria.

Paulianesthes Koch, 1962a: 39, 57 [F]. Type species: Paulianesthesamplipennis Koch, 1962, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Paurodontomophlus Muche, 1979: 171 [M]. Type species: Omophluswittmeri Muche, 1979, by original designation. Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini.

Pechalius Casey, 1907: 379, 420 [M]. Type species: Pechaliussubvittatus Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Epitragini. Note: the First Reviser (Pechalius Casey, 1907 versus Epitragoma Casey, 1907) is Freude (1968: 61).

Pectinepitragus Pic, 1927a: 44 [M]. Type species: Pectinepitraguspubescens Pic, 1927, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Pectphegoneus Freude, 1968: 90, 98 [M]. Type species: Schoenicuspectoralis Champion, 1884, by monotypy. Status: valid subgenus of Phegoneus Casey, 1907 in Pimeliinae: Epitragini.

Pedarasida Reitter, 1917a: 11, 28 [F]. Type species: Asidacariosicollis Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Glabrasida Escalera, 1910 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Pedinopsis Gebien, 1910d: 157 [F]. Type species: Pedinopsispilipes Gebien, 1910, by monotypy. Status: senior synonym of Loensus R. Lucas, 1920 in Blaptinae: Pedinini: Pedinina. Note: junior homonym of Pedinopsis Cotteau, 1863 [Echinoidea].

Pedinulus Seidlitz, 1893: 364, 373 [M]. Type species: Pedinusragusae Baudi di Selve, 1875, by subsequent designation (Iwan and Löbl 2008: 288). Status: junior synonym of Pedinus Latreille, 1797 in Blaptinae: Pedinini: Pedinina. Synonymy: Kamiński and Iwan (2017: 599).

Pedinus Latreille, 1797: 20 [M]. Type species: Tenebriofemoralis Linnaeus, 1767, by subsequent designation (Stephens 1835: 159). Status: valid genus and subgenus in Blaptinae: Pedinini: Pedinina. Note: originally proposed without included nominal species; Latreille (1802: 175), by listing Tenebriofemoralis Linnaeus, 1767 and Tenebriodermestoides Fabricius, 1787 in association with this genus name, was the first author to subsequently and expressly include nominal species in Pedinus (ICZN 1999, Article 67.2.2).

Pedionomus Haag-Rutenberg, 1875c: 42 [M]. Type species: Metriopusfavosus Erichson, 1843, by subsequent designation (Gebien 1937a: 661). Status: senior synonym of Alogenius Gebien, 1910 in Pimeliinae: Adesmiini. Note: junior homonym of Pedionomus Gould, 1841 [Aves].

Pediris Motschulsky, 1872: 24 [F]. Type species: Pedirislongipes Motschulsky, 1872, by original designation. Status: junior synonym of Promethis Pascoe, 1869 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1921a: 289, with Setenis Motschulsky, 1872, a junior synonym of Promethis Pascoe, 1869).

Pedoeces Agassiz, 1846b: 276 [M]. Type species [automatic]: Pedonoecesgalapagoensis G.R. Waterhouse, 1845, by subsequent designation (Aalbu and Triplehorn 1991: 170). Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Note: unjustified emendation of Pedonoeces G.R. Waterhouse, 1845, not in prevailing usage.

Pedonoeces G.R. Waterhouse, 1845a: 32, 34 [M]. Type species: Pedonoecesgalapagoensis G.R. Waterhouse, 1845, by subsequent designation (Aalbu and Triplehorn 1991: 170). Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Aalbu and Triplehorn (1991: 170).

Pedostrongylium Pic, 1916e: 11 [N]. Type species: Strongyliumluteonotatum Pic, 1916 (= Stenochiaxanthozona J. Thomson, 1858), by original designation. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Pelecypalpus Hinton, 1947: 91 [M]. Type species: Pelecypalpusmedon Hinton, 1947, by original designation. Status: junior synonym of Scotoderus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1973c: 258).

Pelecyphorus Solier, 1836: 406, 467 [M]. Type species: Pelecyphorusmexicanus Solier, 1836, by subsequent designation (Hope 1841: 110). Status: valid genus and subgenus in Pimeliinae: Asidini.

Pelleas Bates, 1872a: 98 [M]. Type species: Tenebriocrotchii Wollaston, 1865, by original designation. Status: valid genus in Diaperinae: Diaperini: incertae sedis.

Pelops Gistel, 1834: 22 [M]. Type species: Helopsater Fabricius, 1775, by monotypy. Status: senior synonym of Prionychus Solier, 1835 in Alleculinae: Alleculini: Alleculina. Note: nomen oblitum (see Bousquet and Bouchard 2017: 132).

Pelorinus Vauloger de Beaupré, 1900: 675, 678 [M]. Type species: Helopsanthracinus Germar, 1813, by subsequent designation (Cherney 2005: 382). Status: valid subgenus of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina.

Pelorocnemis Solsky, 1876: 283 [F]. Type species: Pimeliapunctigera Ménétriés, 1849, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Peltadesmia Kuntzen, 1916: 149 [F]. Type species: Metriopusplatynotus Gerstaecker, 1854, by present designation. Status: junior synonym of Coeladesmia Reitter, 1916 in Pimeliinae: Adesmiini. Synonymy: Gebien (1937a: 656).

Peltarium Fischer von Waldheim, 1844: 106 [N]. Type species: Peltariummarginatum Fischer von Waldheim, 1844 (= Blaps baerii Fischer von Waldheim, 1842), by subsequent designation (G.S. Medvedev 1995b: 831). Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Peltasida Reitter, 1917a: 39, 41 [F]. Type species: Asidafavieri Fairmaire, 1880, by monotypy. Status: valid subgenus of Asida Latreille, 1802 in Pimeliinae: Asidini.

Peltoides Laporte, 1833a: 401 [M]. Type species: Peltoidessenegalensis Laporte, 1833, by subsequent designation (Lacordaire 1859a: 337). Status: valid genus and subgenus in Tenebrioninae: Alphitobiini. Note: bibliographic evidence indicates that Oopiestus Chevrolat, 1833 was published before Peltoides Laporte, 1833 and should be treated as the valid name for this genus and subgenus instead; an application to the ICZN is necessary to conserve usage of Peltoides Laporte, 1833.

Peltolobus Lacordaire, 1859a: 71 [M]. Type species [automatic]: Megalophryspatagonica G.R. Waterhouse, 1845, by monotypy. Status: valid genus in Pimeliinae: Trilobocarini. Note: replacement name for Megalophrys G.R. Waterhouse, 1845.

Pemanoa Buck, 1955: 269 [F]. Type species: Pemanoamixta Buck, 1955, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Penadelium Matthews, 1998: 710, 754 [N]. Type species: Penadeliumaraucanum Matthews, 1998, by original designation. Status: valid genus in Lagriinae: Adeliini.

Penaus Freude, 1968: 130 [M]. Type species: Penauspenai Freude, 1968, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Peneta Lacordaire, 1859a: 319 [F]. Type species: Penetalebasii Lacordaire, 1859, by subsequent designation (R. Lucas 1920: 492). Status: valid genus in Phrenapatinae: Penetini.

Pengalenganus Pic, 1917d: 10 [M]. Type species: Pengalenganusinaequalis Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Penichrus Champion, 1885: 134 [M]. Type species: Penichrusblapstinoides Champion, 1885, by monotypy. Status: valid genus in Tenebrioninae: incertae sedis. Note: placed in Tenebrioninae incertae sedis by Johnston et al. (2020: 771).

Pentaphyllus Dejean, 1821: 68 [M]. Type species: Mycetophagustestaceus Hellwig, 1792, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Penthicinus Reitter, 1896b: 161 [M]. Type species: Penthicinuskoltzei Reitter, 1896, by subsequent designation (Gebien 1939: 458). Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: Reitter (1896b) used two different spellings for this genus, including Penticinus (Reitter 1896b: 171); subsequently, Reitter (1904: 135, 170) used Penthicinus as the valid name of the genus and so acted as First Reviser (ICZN 1999: Article 24.2.4).

Penthicoides Fairmaire, 1896a: 20 [M]. Type species: Penthicoidesseriatoporus Fairmaire, 1896, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Penthicomelanesthes Bogatchev, 1972: 631 [F]. Type species: Heliophilusgibbulus Faldermann, 1835, by original designation. Status: junior synonym of Myladion Reitter, 1887 in Blaptinae: Opatrini: Opatrina. Synonymy: G.S. Medvedev (1990: 186).

Penthicus Faldermann, 1836: 384 [M]. Type species: Penthicuspinguis Faldermann, 1836, by subsequent designation (Hope 1841: 126). Status: valid genus and subgenus in Blaptinae: Opatrini: Opatrina.

Pentholasius Reitter, 1904: 178 [M]. Type species: Halonomusvariolatus Allard, 1884, by monotypy. Status: junior synonym of Mesomorphus Miedel, 1880 in Blaptinae: Opatrini: Opatrina. Synonymy: Ferrer (2002: 376).

Penthomegus Reitter, 1904: 161 [M]. Type species: Penthicuscorpulentus Reitter, 1896, by subsequent designation (Iwan and Löbl 2007: 734). Status: junior synonym of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Synonymy: Gebien (1939: 459, with Lobodera Mulsant & Rey, 1859, a junior synonym of Penthicus Faldermann, 1836).

Perdicus Fairmaire, 1899c: 386 [M]. Type species: Perdicusantrophilus Fairmaire, 1899, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Perdistretus Koch, 1953d: 65 [M]. Type species: Distretusvilhenai Koch, 1953, by original designation. Status: valid subgenus of Distretus Haag-Rutenberg, 1871 in Pimeliinae: Sepidiini: Molurina.

Periatrum Sharp, 1886: 407 [N]. Type species: Periatrumhelmsi Sharp, 1886, by monotypy. Status: valid genus in Lagriinae: Adeliini.

Periblaps Bauer, 1921: 32 [F]. Type species: none designated. Status: undetermined taxon in Blaptinae: Blaptini: Blaptina. Note: this genus was described before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Periblaps and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Perichilus Quedenfeldt, 1885: 18 [M]. Type species: Perichilusbrevicornis Quedenfeldt, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Periloma Gebien, 1938b: 100 [N]. Type species: Perilomaalfkeni Gebien, 1938, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Peringueyia Koch, 1958: 44 [F]. Type species: Echinotusdispar Péringuey, 1899, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Periphanes Fairmaire, 1882a: 245 [M]. Type species: Periphanesorichalceus Fairmaire, 1882, by monotypy. Status: senior synonym of Periphanodes Gebien, 1943 in Stenochiinae: Cnodalonini. Note: junior homonym of Periphanes Hübner, 1821 [Lepidoptera].

Periphanodes Gebien, 1943: 902 [M]. Type species [automatic]: Periphanesorichalceus Fairmaire, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: replacement name for Periphanes Fairmaire, 1882; new placement [OM], previously included in Tenebrioninae: Helopini.

Peristeptus Haag-Rutenberg, 1875b: 3, 24 [M]. Type species: Pogonobasislaevigata Gerstaecker, 1854, by subsequent designation (R. Lucas 1920: 495). Status: junior synonym of Pogonobasis Solier, 1837 in Pimeliinae: Adelostomini. Synonymy: Koch (1952b: 17).

Perithrix Fairmaire, 1879a: 193 [F]. Type species: Perithrixgranidorsis Fairmaire, 1879, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Pescennius Champion, 1884: 3 [M]. Type species: Pescenniusvillosus Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Petria Semenov, 1894: 611 [F]. Type species: Petriatachyptera Semenov, 1894, by subsequent designation (R. Lucas 1920: 496). Status: valid genus in Alleculinae: Cteniopodini.

Petrostetha Novák, 2008a: 212 [F]. Type species: Petrostethatibialis Novák, 2008, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Peyerimhoffius Koch, 1948: 420 [M]. Type species: Phylaxplicatus P.H. Lucas, 1846, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Peyrierasia Dajoz, 1975b: 113, 114 [F]. Type species: Peyrierasiasechellensis Dajoz, 1975, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Pezodontus Dejean, 1834: 203 [M]. Type species [automatic]: Odontopuscostatus Silbermann, 1833, by subsequent designation (Hope 1841: 126). Status: valid genus in Lagriinae: Pycnocerini. Note: replacement name for Odontopus Silbermann, 1833 (see Bousquet and Bouchard 2013a: 57).

Pezohelaeus Gebien, 1921a: 282 [M]. Type species: Pterohelaeushirtus W.J. MacLeay, 1888, by monotypy. Status: junior synonym of Pterohelaeus Brême, 1842 in Tenebrioninae: Heleini: Heleina. Synonymy: Carter (1927: 230).

Pezomaia Kulzer, 1952: 752 [F]. Type species: Pezomaiafemoralis Kulzer, 1952, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pezophenus Gebien, 1921a: 325, 339 [M]. Type species: Pezophenusrutilans Gebien, 1921 (= Prophanessubmetallicus W.J. MacLeay, 1887), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Phaedeucyrtus Pic, 1916e: 14 [M]. Type species: Phaedeucyrtusobscuripes Pic, 1916, by monotypy. Status: junior synonym of Phaedis Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Ando (2016: 70).

Phaedis Pascoe, 1866a: 474 [M]. Type species: Phaediselysius Pascoe, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Phaedogria Borchmann, 1936: 16, 57 [F]. Type species: Lagriaionoptera Erichson, 1834, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Phaenogeton Bremer, 2016: 220 [M]. Type species: Amarygmusvaricolor Gebien, 1921, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Phaeostolus Fairmaire, 1884c: cxlvi [M]. Type species: Phaeostolusgrandicornis Fairmaire, 1884, by monotypy. Status: valid genus in Tenebrioninae: Praeugenini. Note: redescribed as new by Fairmaire (1887b: 293).

Phaeotribon Kraatz, 1865: 81, 242 [M]. Type species: Phaeotribonpulchellus Kraatz, 1865, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Phaleria Latreille, 1802: 162 [F]. Type species: Tenebriocadaverinus Fabricius, 1792, by plenary powers (ICZN 1975, Opinion 1039). Status: valid genus and subgenus in Diaperinae: Phaleriini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1975, Opinion 1039).

Phaleriderma Koch, 1954a: 25 [N]. Type species: Oncotusobscuricollis Solier, 1848, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Phalerisida Kulzer, 1959: 563 [F]. Type species: Phalerisidamaculata Kulzer, 1959, by original designation. Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Synonymy: Triplehorn (1991: 258).

Phaleromela Reitter, 1916b: 4 [F]. Type species: Phaleriasubhumeralis Marseul, 1876, by monotypy. Status: valid genus in Diaperinae: Phaleriini.

Phallocentrion Koch, 1956a: 166 [N]. Type species: Selinusedentatus Gebien, 1914, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Phanechloros Matthews & Bouchard, 2008: 300 [M]. Type species [automatic]: Platyphanespunctipennis Carter, 1911, by original designation. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina. Note: replacement name for Chlorophanes Matthews, 1992.

Phanerops Solier, 1851: 233 [M]. Type species: Phaneropselongatus Solier, 1851, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Phanerotoma Solier, 1843: 82, 126 [F]. Type species: Phanerotomaelongata Solier, 1843 (= Pimelialaevigata G.-A. Olivier, 1795), by original designation. Status: senior synonym of Ocnodes Fåhraeus, 1870 in Pimeliinae: Sepidiini: Molurina. Synonymy: Gebien (1937a: 759), Kamiński et al. (2019b: 55). Note: junior homonym of Phanerotoma Wesmael, 1838 [Hymenoptera].

Phanerotomea Koch, 1958: 58 [F]. Type species [automatic]: Phanerotomaelongata Solier, 1843 (= Pimelialaevigata G.-A. Olivier, 1795), by original designation. Status: junior synonym of Ocnodes Fåhraeus, 1870 in Pimeliinae: Sepidiini: Molurina. Note: replacement name for Phanerotoma Solier, 1843.

Phayllidius Gebien, 1922a: 451 [M]. Type species: Phayllidiusdispar Gebien, 1922, by monotypy. Status: junior synonym of Ulomoides Blackburn, 1888 in Diaperinae: Diaperini: Diaperina. Synonymy: Grimm (2018: 103).

Phayllus Champion, 1886: 167 [M]. Type species: Phayllusminutus Champion, 1886, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Phediodes Campbell, 1976: 26 [M]. Type species: Phediodesapterus Campbell, 1976, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Phedius Champion, 1888: 447 [M]. Type species: Phediuschevrolati Champion, 1888, by subsequent designation (R. Lucas 1920: 500). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Phegoneus Casey, 1907: 380, 426 [M]. Type species: Epitragodesjulichi Casey, 1891, by original designation. Status: valid genus and subgenus in Pimeliinae: Epitragini.

Phellidius J.L. LeConte, 1862: 236 [M]. Type species: Bolitophaguscornutus Fabricius, 1801, by original designation. Status: junior synonym of Bolitotherus Candèze, 1861 in Tenebrioninae: Bolitophagini. Synonymy: J.L. LeConte (1866a: 62). Note: the younger species name Bolitophaguscornutus Fabricius, 1801 was given priority over the older synonym Opatrumbifurcum Fabricius, 1798 by the ICZN (2019, Opinion 2438).

Pheloneis Pascoe, 1866a: 483 [M]. Type species: Adeliumharpaloides White, 1846 (= Adeliumamaroides Bates, 1874), by original designation. Status: valid genus in Lagriinae: Adeliini.

Phelopatrum Marseul, 1876: 100 [N]. Type species: Hadrusscaphoides Marseul, 1876, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Phenus Gebien, 1921a: 324, 325 [M]. Type species: Phenuslatitarsis Gebien, 1921, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pheres Champion, 1886: 150 [M]. Type species: Pheresbatesi Champion, 1886, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Pheugonius Fairmaire, 1899b: 313 [M]. Type species: Pheugoniusborneensis Fairmaire, 1899, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Phibalus Gistel, 1856: 384 [M]. Type species: Chrysomelapubescens Linnaeus, 1767, by monotypy. Status: valid subgenus of Omophlus Dejean, 1834 in Alleculinae: Cteniopodini.

Philhamellus Kaszab, 1962b: 83, 84 [M]. Type species: Philhammusmyrmecophilus Kaszab, 1960, by original designation. Status: valid subgenus of Philhammus Fairmaire, 1871 in Pimeliinae: Cnemeplatiini: Cnemeplatiina.

Philhammus Fairmaire, 1871a: 393 [M]. Type species: Philhammussericans Fairmaire, 1871, by monotypy. Status: valid genus and subgenus in Pimeliinae: Cnemeplatiini: Cnemeplatiina.

Philolithus Lacordaire in J.L. LeConte, 1858b: 18 [M]. Type species: Pelecyphoruscarinatus J.L. LeConte, 1851, by subsequent designation (Casey 1912: 79). Status: valid genus and subgenus in Pimeliinae: Asidini.

Philorea Erichson, 1834: 242 [F]. Type species: Philoreapicipes Erichson, 1834, by monotypy. Status: valid genus in Pimeliinae: Physogasterini.

Phligra Laporte, 1840: 197 [F]. Type species: Sepidium degeeri Laporte, 1840 (= Tenebriocristatus DeGeer, 1778), by monotypy. Status: junior synonym of Cyrtoderes Dejean, 1834 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Lacordaire (1859a: 202).

Phobelius Blanchard, 1842: pl. 14 [M]. Type species: Phobeliuscrenatus Blanchard, 1842, by monotypy. Status: valid genus in Lagriinae: Goniaderini. Note: see Bouchard and Bousquet (2020a: 99) for comments about the date of publication of this genus.

Phrenapates Gray in Griffith and Pidgeon, 1831: pl. 50 [M]. Type species: Phrenapatesbennettii Gray, 1831, by monotypy. Status: valid genus in Phrenapatinae: Phrenapatini.

Phrynocarenum Gebien, 1928: 106 [N]. Type species: Phrynocarenumbruchianum Gebien, 1928 (= Emmalloderastrangulata Fairmaire, 1905), by monotypy. Status: valid genus in Pimeliinae: Phrynocarenini.

Phrynocolopsis Koch, 1951: 93 [F]. Type species: Phrynocolusfrondosus Gerstaecker, 1871, by original designation. Status: valid subgenus of Phrynocolus Lacordaire, 1859 in Pimeliinae: Sepidiini: Molurina.

Phrynocolus Lacordaire, 1859a: 201 [M]. Type species [automatic]: Cryptogeniusdentatus Solier, 1843, by original designation. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Molurina. Note: replacement name for Cryptogenius Solier, 1843.

Phrynophanes Koch, 1951: 92 [M]. Type species: Molurisgredleri Haag-Rutenberg, 1877, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Phthora Champion, 1893a: 531 [F]. Type species [automatic]: Phtoracrenata Mulsant, 1854 (= Clamorisinsurgens Gozis, 1886), by monotypy. Status: junior synonym of Clamoris Gozis, 1886 in Phrenapatinae: Penetini. Note: unjustified emendation of Phtora Mulsant, 1854, not in prevailing usage; junior homonym of Phthora Gemminger, 1870 [Coleoptera: Tenebrionidae: Diaperinae: Phaleriini].

Phthora Gemminger in Gemminger and Harold, 1870: 1959 [F]. Type species [automatic]: Phtoracrenata Germar, 1836, by monotypy. Status: junior synonym of Phtora Germar, 1836 in Diaperinae: Phaleriini. Note: unjustified emendation of Phtora Germar, 1836, not in prevailing usage.

Phtora Germar, 1836: pl. 11 [F]. Type species: Phtoracrenata Germar, 1836, by monotypy. Status: valid genus and subgenus in Diaperinae: Phaleriini. Note: the alternative original spelling Phthora, used by Germar (1836: explanation for pl. 11), was rejected by Neave (1940: 736) who acted as First Reviser.

Phtora Mulsant, 1854: 228 [F]. Type species: Phtoracrenata Mulsant, 1854 (= Clamorisinsurgens Gozis, 1886), by monotypy. Status: senior synonym of Clamoris Gozis, 1886 in Phrenapatinae: Penetini. Note: junior homonym of Phtora Germar, 1836 [Coleoptera: Tenebrionidae: Diaperinae: Phaleriini].

Phygoscotus Schulz, 1902: 134 [M]. Type species [automatic]: Spheniscuserotyloides W. Kirby, 1819, by monotypy. Status: junior synonym of Cuphotes Champion, 1887 in Stenochiinae: Stenochiini. Note: replacement name for Spheniscus W. Kirby, 1819.

Phylacastus Fairmaire, 1897f: 116 [M]. Type species: Phylacastusstriolatus Fairmaire, 1897, by monotypy. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Phylacinus Fairmaire, 1896b: 349 [M]. Type species: Phylacinusasperipennis Fairmaire, 1896, by monotypy. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Phylan Sturm, 1826: 23 [M]. Type species: Opatrumgibbum Fabricius, 1775, by subsequent monotypy (Steven 1828: 99). Status: valid genus and subgenus in Blaptinae: Dendarini: Dendarina. Note: originally proposed without included nominal species; Steven (1828: 99), by including the species Opatrumgibbum Fabricius, 1775 in association with this name, was the first author to subsequently and expressly include nominal species in Phylan (ICZN 1999, Article 67.2.2).

Phylanmania Ferrer, 2013: 38 [F]. Type species: Phylan ilerdensis Español & Viñolas, 1981, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Phylax Brullé, 1832: 209 [M]. Type species [automatic]: Opatrumgibbum Fabricius, 1775, by subsequent monotypy (Steven 1828: 99). Status: junior synonym of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina. Note: unnecessary replacement name of Phylan Sturm, 1826 (see Bouchard and Bousquet 2020a: 100).

Phylaximon Koch, 1948: 414 [M]. Type species: Opatrumvariolosum G.-A. Olivier, 1812, by original designation. Status: valid subgenus of Allophylax Bedel, 1906 in Blaptinae: Dendarini: Melambiina.

Phyletes Redtenbacher, 1845: 128 [M]. Type species: Phyletespopuli Chevrolat, 1847 (= Diaperisbifasciata Say, 1824), by subsequent monotypy (Chevrolat 1847b: 57). Status: junior synonym of Alphitophagus Stephens, 1832 in Diaperinae: Diaperini: Adelinina. Synonymy: Heyden et al. (1883: 133). Note: genus originally proposed without included nominal species; Chevrolat (1847b: 57) included the species “Phylethuspopuli Még.” as a synonym of Alphitophagusquadripustulatus Stephens, 1832 in the genus “Phylethus” and Redtenbacher (1848: 589) used “Phylethuspopuli” as valid subsequently; therefore, the author of the type species dates back to its first publication as a synonym (ICZN 1999, Article 11.6.1).

Phyllechus Bouchard & Bousquet, new genus [M]. Type species: Phyllechusboettcheri Kulzer, 1966, by present designation. Status: valid genus in Stenochiinae: Stenochiini. Note: Kulzer (1966: 374) introduced the new genus name Phyllechus for two new species, but unfortunately did not designate a type species; the genus Phyllechus, which has been treated as valid since 1966, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Phyllechusboettcheri Kulzer, 1966 as type species and referring to Kulzer (1966: 374) for the character states that characterise and differentiate Phyllechus.

Phyloradix Endrödy-Younga, 1996: 15, 16 [M]. Type species: Caenocrypticussoror Endrödy-Younga, 1996, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini.

Phymaeus Pascoe, 1883: 439 [M]. Type species: Phymaeuspustulosus Pascoe, 1883, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Phymatestes Pascoe, 1866b: 142 [M]. Type species [automatic]: Lagriatuberculata Fabricius, 1787, by monotypy (see ICZN 1989, Opinion 1525). Status: valid genus in Lagriinae: Goniaderini. Note: replacement name for Phymatodes Dejean, 1834; placed on the Official List of Generic Names in Zoology by the ICZN (1989, Opinion 1525).

Phymatiotris Solier, 1836: 63 [F]. Type species: Tentyriaquadricollis Brullé, 1832, by subsequent designation (Hope 1841: 118). Status: valid genus in Pimeliinae: Pimeliini.

Phymatium Billberg, 1820: 31 [N]. Type species: Pimeliamaculata Fabricius, 1781, by present designation. Status: senior synonym of Cryptochile Latreille, 1828 in Pimeliinae: Cryptochilini: Calognathina. Synonymy: new synonym [PB]. Note: discovery of this forgotten name threatens the stability of the junior objective synonym Cryptochile Latreille, 1828; although Phymatium Billberg, 1820 has not been used as valid in the literature after 1899, we could not find usage of Cryptochile Latreille, 1828 in at least 25 works, published by at least ten authors in the immediately preceding 50 years and therefore reversal of precedence cannot be used to treat Phymatium Billberg, 1820 as a nomen oblitum; we recommend that an application be submitted to the International Commission on Zoological Nomenclature to conserve usage of Cryptochile Latreille, 1828, the type genus of the valid tribe Cryptochilini and the subtribe Cryptochilina.

Phymatodes Dejean, 1834: 203 [M]. Type species: Lagriatuberculata Fabricius, 1787, by monotypy. Status: senior synonym of Phymatestes Pascoe, 1866 in Lagriinae: Goniaderini. Note: name suppressed for the purposes of the Principle of Homonymy and the Principle of Priority by the ICZN (1989, Opinion 1525).

Phymatoplata Koch, 1956a: 269 [F]. Type species: Selinusasperulus Fairmaire, 1897, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Phymatosoma Laporte & Brullé, 1831: 332, 408 [N]. Type species: Phymatosomatuberculatum Laporte & Brullé, 1831, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: unjustified emendation of the original spelling Phymatisoma, introduced by Agassiz (1846b: 290), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Physadesmia Penrith, 1979: 7, 35 [F]. Type species: Adesmiaglobosa Haag-Rutenberg, 1875, by original designation. Status: valid genus in Pimeliinae: Adesmiini.

Physciolagria Pic, 1930c: 7 [F]. Type species: Physciolagrialiturata Pic, 1930, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: we could not assign this genus to any particular group in Tenebrionidae based on the description.

Physignathus Gistel, 1834: 22 [M]. Type species: Helopsundatus Fabricius, 1792 (= Erotylusnebulosus Fabricius, 1781), by monotypy. Status: junior synonym of Cymatothes Dejean, 1834 Tenebrioninae: Amarygmini. Synonymy: Bousquet and Bouchard (2017: 132). Note: junior homonym of Physignathus Cuvier, 1829 [Reptilia].

Physocoelus Haldeman, 1850: 347 [M]. Type species: Psorodesinflata Solier, 1848 (= Helopscontractus Palisot de Beauvois, 1812), by monotypy. Status: junior synonym of Meracantha W. Kirby, 1837 in Tenebrioninae: Amarygmini. Synonymy: Schaum (1850: 181, through synonymy of the type species).

Physodera Solier, 1843: 78, 125 [F]. Type species: Pimeliagibba Fabricius, 1787, by original designation. Status: junior synonym of Moluris Latreille, 1802 in Pimeliinae: Sepidiini: Molurina. Synonymy: Lacordaire (1859a: 194). Note: junior homonym of Physodera Eschscholtz, 1829 [Coleoptera: Carabidae].

Physogaster Lacordaire, 1830a: 276 [F]. Type species: Physogastermendocina Lacordaire, 1830, by monotypy. Status: valid genus in Pimeliinae: Physogasterini.

Physogasterinus Kaszab, 1981a: 79 [M]. Type species: Physogasterinuslanuginosus Kaszab, 1981, by original designation. Status: valid genus in Pimeliinae: Physogasterini.

Physogria Borchmann, 1916a: 48, 108 [F]. Type species: Lagriagibbosa Kolbe, 1901, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Physohelops Schuster, 1937: 50 [M]. Type species: Physohelopsfreyi Schuster, 1937, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Physolagria Fairmaire, 1891g: 114 [F]. Type species: Physolagriamolleri Fairmaire, 1891, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Physophrynus Fairmaire, 1882c: l [M]. Type species: Physophrynusburdoi Fairmaire, 1882, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Physosterna Dejean, 1834: 179 [F]. Type species: Pimeliaovata G.-A. Olivier, 1795 (= Tenebriotorulosus Pallas, 1781), by monotypy. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Phytolostoma Koch, 1952b: 34 [N]. Type species: Phytolostomalimpopoana Koch, 1952, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Phitophilus Guérin-Méneville, 1831a: pl. 4 [M]. Type species: Phitophilushelopioides Guérin-Méneville, 1831, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Piccula Bousquet & Bouchard in Bousquet et al., 2015: 137 [F]. Type species [automatic]: Gerardiasublineata Pic, 1954, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina. Note: replacement name for Gerardia Pic, 1954.

Piciella Borchmann, 1936: 237, 435 [F]. Type species: Piciellahelopioides Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Picnotagalus Kaszab, 1939a: 102 [M]. Type species: Picnotagalushorni Kaszab, 1939, by original designation. Status: junior synonym of Scolytocaulus Fairmaire, 1896 in Phrenapatinae: Penetini. Synonymy: Schawaller (1999b: 422).

Picocamaria Masumoto, 1993b: 226, 232 [F]. Type species: Camariageniculata Pic, 1915, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Piesomera Solier, 1843: 77 [F]. Type species: Pimeliascabra Fabricius, 1775, by monotypy. Status: junior synonym of Psammodes W. Kirby, 1819 in Pimeliinae: Sepidiini: Molurina. Synonymy: Gebien (1937a: 759).

Piesterotarsa Sénac, 1884: 8 [F]. Type species [automatic]: Pimeliagigantea Fischer, 1820, by subsequent designation (Gebien 1937a: 840). Status: junior synonym of Pisterotarsa Motschulsky, 1860 in Pimeliinae: Pimeliini. Note: unjustified emendation of Pisterotarsa Motschulsky, 1860, not in prevailing usage.

Piestognathoides Kaszab, 1981b: 305 [M]. Type species: Piestognathoidesbahrainicus Kaszab, 1981, by original designation. Status: valid genus in Pimeliinae: Erodiini.

Piestognathus P.H. Lucas, 1858: clxxxviii [M]. Type species: Piestognathusdouei P.H. Lucas, 1858, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Pigeocaulinus Kaszab, 1984: 355, 388 [M]. Type species: Pigeocaulinussumatranus Kaszab, 1984 (= Leprocaulinuskrikkeni Kaszab, 1982), by original designation. Status: junior synonym of Leprocaulinus Kaszab, 1982 in Stenochiinae: Cnodalonini. Synonymy: Masumoto (2003: 60); see notes in the entry for Leprocaulinus Kaszab, 1982 regarding the validity of this older name.

Pigeostrongylium Kaszab, 1984: 355, 385 [N]. Type species: Pigeostrongyliumkedahense Kaszab, 1984, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pigeus Gebien, 1919: 28, 153 [M]. Type species: Camarimenanitidipes Fairmaire, 1893, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pilioloba Erichson in Agassiz, 1846a: 144 [F]. Type species: Salaxlacordairii Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Salax Guérin-Méneville, 1834 in Pimeliinae: Trilobocarini. Synonymy: Erichson in Agassiz (1846a: 144). Note: the name Pilioloba was listed as synonym of Salax Guérin-Méneville, 1834 by Erichson in Agassiz (1846a: 144), it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Cazurro Ruiz 1894b: 451), Pilioloba was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Pilobalia Burmeister, 1875: 487 [F]. Type species: Nycteliadecorata Erichson, 1834, by subsequent designation (Rye 1877: 338). Status: valid genus in Pimeliinae: Nycteliini.

Pilobaloderes Kulzer, 1958b: 192 [M]. Type species: Pilobaloderesgebieni Kulzer, 1958, by original designation. Status: valid genus in Pimeliinae: Praociini.

Pilosocasnonidea Pic, 1934a: 31 [F]. Type species: Nemostirabipartita Pic, 1911, by monotypy. Status: valid subgenus of Casnonidea Fairmaire, 1882 in Lagriinae: Lagriini: Statirina.

Pilosoplonyx Bremer, 2014a: 37 [M]. Type species: Plesiophthalmusbremeri Masumoto, 1999, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Piloxys Fairmaire, 1895a: 23 [M]. Type species: Piloxysfoveatus Fairmaire, 1895, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pimalius Casey, 1907: 367 [M]. Type species: Trimytispulverea Horn, 1870, by original designation. Status: junior synonym of Trimytis J.L. LeConte, 1851 in Pimeliinae: Edrotini. Synonymy: MacLachlan and Olson (1990: 79).

Pimelia Fabricius, 1775: 251 [F]. Type species: Pimeliaangulata Fabricius, 1775, by subsequent designation (Hope 1841: 118). Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Pimeliocnera Reitter, 1909b: 310 [F]. Type species: Pimeliocneragebieni Reitter, 1909, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Pimelionotus Ardoin, 1963a: 86 [M]. Type species: Psorodeslugens Fåhraeus, 1870, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Pimeliopsis Champion, 1892: 477 [F]. Type species: Pimeliopsisgranulata Champion, 1892, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Pimelipachys Skopin, 1962: 232 [M]. Type species: Pachyscelislaevicollis Reitter, 1893, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Pimelosomus Burmeister, 1875: 488, 489 [M]. Type species: Pimelosomussphaericus Burmeister, 1875, by monotypy. Status: valid genus in Pimeliinae: Physogasterini.

Pimidia Rafinesque, 1815: 113 [F]. Type species [automatic]: Pimeliaangulata Fabricius, 1775, by subsequent designation (Hope 1841: 118). Status: junior synonym of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini. Note: unjustified emendation of Pimelia Fabricius, 1775, not in prevailing usage.

Pimplema Pascoe, 1887: 16 [F]. Type species: Pimplemaampliata Pascoe, 1887 (= Platydemahemisphaerica Laporte & Brullé, 1831), by monotypy. Status: valid genus in Diaperinae: Leiochrinini.

Piscicula Robiche, 2004a: 736 [F]. Type species: Pisciculasprecherae Robiche, 2004, by monotypy. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Pisterotarsa Motschulsky, 1860c: 532 [F]. Type species: Pimeliagigantea Fischer, 1820, by subsequent designation (Gebien 1937a: 840). Status: valid genus in Pimeliinae: Pimeliini.

Pitholaus Champion, 1888: 446 [M]. Type species: Pitholaushelopioides Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Pizura Novák, 2016b: 436 [F]. Type species: Pizurabarbucha Novák, 2016, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Plamius Fairmaire, 1896a: 30 [M]. Type species: Plamiustenuestriatus Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Planasida Escalera, 1907: 337 [F]. Type species: Asidapaulinoi Pérez Arcas, 1868, by subsequent designation (Viñolas and Cartagena 2005: 223). Status: junior synonym of Gracilasida Escalera, 1905 in Pimeliinae: Asidini. Synonymy: F. Soldati (2008: 137). Note: Planasida Escalera, 1907 was used as valid in the literature recently (e.g., F. Soldati 2008: 137); however, the name Gracilasida Escalera, 1905 is older and should be used as valid (e.g., F. Soldati 2020: 152–153).

Planibates Kaszab, 1939b: 221 [M]. Type species: Planibatespapuanus Kaszab, 1939, by monotypy. Status: junior synonym of Bradymerus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Schawaller (2013b: 163).

Planirostrosis Penrith, 1977: 18, 172 [F]. Type species: Zophosishimba Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Planodes Mulsant & Rey, 1859c: 94, 117 [M]. Type species: Planodesbyrroides Mulsant & Rey, 1859, by subsequent designation (R. Lucas 1920: 517). Status: senior synonym of Planostibes Gemminger, 1870 in Blaptinae: Opatrini: Stizopodina. Note: junior homonym of Planodes Newman, 1842 [Coleoptera: Cerambycidae].

Planoodescelis Egorov, 2004: 591 [F]. Type species: Oodesceliskansouensis Kaszab, 1940, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Planoplatyscelis Kaszab, 1940a: 157, 206 [F]. Type species: Platyscelismargelanica Kraatz, 1882 (= Bioramixpamirensis Bates, 1879), by plenary powers (ICZN 1993a, Opinion 1728). Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1993a, Opinion 1728); the First Reviser (Planoplatyscelis Kaszab, 1940 versus Pleioplatyscelis Kaszab, 1940) is Egorov (2004: 601).

Planoprosodes G.S. Medvedev, 2005b: 90 [M]. Type species: Prosodesreitteri Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Note: originally described as a section within a subgenus.

Planostibes Gemminger in Gemminger and Harold, 1870: 1926 [M]. Type species [automatic]: Planodesbyrroides Mulsant & Rey, 1859, by subsequent designation (R. Lucas 1920: 517). Status: valid genus in Blaptinae: Opatrini: Stizopodina. Note: replacement name for Planodes Mulsant & Rey, 1859.

Plastica C.O. Waterhouse, 1903: 563 [F]. Type species: Plasticapolita C.O. Waterhouse, 1903, by monotypy. Status: valid genus in Tenebrioninae: Apocryphini.

Platamodes Ménétriés, 1849: 233 [M]. Type species: Platamodesdentipes Ménétriés, 1849, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Platamodina.

Platasida Casey, 1912: 77, 182 [F]. Type species: Asidaembaphionides Horn, 1894, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Plateia Laporte, 1840: 215 [F]. Type species: Plateiaorientalis Laporte, 1840, by monotypy. Status: junior synonym of Catapiestus Perty, 1831 in Stenochiinae: Cnodalonini. Synonymy: Blanchard (1845: 16). Note: junior homonym of Plateia Hübner, 1820 [Lepidoptera].

Platesthes G.R Waterhouse, 1845b: 317 [F]. Type species: Platesthessilphoides G.R. Waterhouse, 1845, by monotypy. Status: valid genus in Pimeliinae: Praociini.

Platolenes Gebien, 1914b: 420 [M]. Type species: Platolenesrufipes Gebien, 1914, by monotypy. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer (2001a: 69).

Platyallecula Blair, 1935b: 266 [F]. Type species: Cistelabrunnea C.O. Waterhouse, 1876, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Platyblaps Motschulsky, 1860c: 531 [F]. Type species: Blaps holconota Fischer von Waldheim, 1844, by subsequent designation (Nabozhenko 2008: 36). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860)

Platybolium Blair, 1938: 222 [N]. Type species: Platyboliumalvearium Blair, 1938, by original designation. Status: valid genus in Tenebrioninae: Triboliini.

Platyburak Iwan, 1990: 124 [M]. Type species: Notocoraxnervosus Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platyburmanicus Iwan, 2003b: 715 [M]. Type species: Platyburmanicusignotus Iwan, 2003, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platycharlesus Iwan, 1998b: 307 [M]. Type species: Trigonopusmorosus Mulsant & Rey, 1853, by original designation. Status: junior synonym of Schelodontes Koch, 1956 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan and Kamiński (2014: 171).

Platycilibe Carter, 1911a: 205 [F]. Type species: Platycilibebrevis Carter, 1911, by monotypy. Status: junior synonym of Scolytocaulus Fairmaire, 1896 in Phrenapatinae: Penetini. Synonymy: Kaszab (1978a: 168).

Platycolpotus Iwan, 1997: 255, 266 [M]. Type species: Platydendarusdendaroides Kaszab, 1975, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platycotylus Olliff, 1883: 182 [M]. Type species: Platycotylusinusitatus Olliff, 1883 (= Ipsaphesnitidulus W.J. MacLaeay, 1873), by monotypy. Status: valid genus in Tenebrioninae: Palorini. Note: originally described in the family Cucujidae, transferred to Tenebrionidae by Crowson (1955: 103).

Platycrepis Lacordaire, 1859b: 418 [F]. Type species: Platycrepisviolacea Lacordaire, 1859, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Platycrypticus Español, 1955: 9, 17 [M]. Type species: Crypticusviaticus Fairmaire, 1851, by original designation. Status: valid subgenus of Crypticus Latreille, 1816 in Diaperinae: Crypticini. Note: replacement name for Ulomoides Escalera, 1927. Note: Español (1952a: 117) introduced the replacement name Platycrypticus earlier; however, the name is unavailable from that date since the author did not fix a type species (ICZN 1999, Article 13.3.1).

†Platycteniopus Chang, Nabozhenko, Pu, Xu, Jia & Li, 2016: 291 [M]. Type species: Platycteniopusdiversoculatus Chang, Nabozhenko, Pu, Xu, Jia & Li, 2016, by original designation. Status: valid genus in Alleculinae: Cteniopodini. Note: described from Upper Cretaceous deposits (China).

Platydema Laporte & Brullé, 1831: 332, 350 [F]. Type species: Diaperisviolacea Fabricius, 1790, by subsequent designation (Westwood 1838: 32). Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: see Löbl and Smetana (2010: 34) for comments on the gender of this name.

Platydemoides Kaszab, 1980a: 161 [M]. Type species: Platydemoidesbrincki Kaszab, 1980, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina. Note: Platydemoides was used earlier by Kaszab (1979a: 77) without a description, a definition or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Platydendarus Kaszab, 1975b: 281, 312 [M]. Type species: Opatrumjavanum Wiedemann, 1819, by original designation. Status: junior synonym of Notocorax Dejean, 1834 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan (1990: 123).

Platyesia Skopin, 1971: 335 [F]. Type species: Diesiakarelini Fischer von Waldheim, 1844, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Platyesthus Mäklin, 1878: 92 [M]. Type species: Platyesthuspallidipennis Mäklin, 1878, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Platyholmus Dejean, 1834: 180 [M]. Type species: Praocisdilaticollis Lacordaire, 1830, by subsequent designation (Solier 1841a: 270 [370]). Status: valid genus in Pimeliinae: Praociini.

Platykochius Iwan, 2002a: 48, 88 [M]. Type species: Selinusplatessa Fairmaire, 1887, by original designation. Status: junior synonym of Anchophthalmops Koch, 1956 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 89).

Platylus Mulsant & Rey, 1859a: 70 [M]. Type species: Blapsdilatata Fabricius, 1798, by monotypy. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Platymedvedevia Iwan & Banaszkiewicz, 2007: 725 [F]. Type species: Platymedvedeviademeyeri Iwan & Banaszkiewicz, 2007, by monotypy. Status: junior synonym of Angolositus Koch, 1955 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 90).

Platynoscelis Kraatz, 1882: 91 [F]. Type species: Platynoscelishelopioides Kraatz, 1882, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Platynosum Mulsant & Rey, 1859c: 73, 81, 150 [N]. Type species: Platynosumpaulinae Mulsant & Rey, 1859, by monotypy. Status: valid genus in Blaptinae: Opatrini: Sclerina. Note: the spelling Platysum (pp. 73, 81) was corrected to Platynosum in the “Errata” of the same work (p. 150), Platynosum is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Platynotoides Kaszab, 1975b: 281, 296 [M]. Type species: Platynotusbellii Fairmaire, 1896, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platynotus Fabricius, 1801a: 138 [M]. Type species: Blapsexcavata Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Platyolmus Burmeister, 1875: 492 [M]. Type species [automatic]: Praocisdilaticollis Lacordaire, 1830, by subsequent designation (Solier 1841a: 270 [370]). Status: junior synonym of Platyholmus Dejean, 1834 in Pimeliinae: Praociini. Note: unjustified emendation of Platyholmus Dejean, 1834, not in prevailing usage.

Platyolus Mulsant & Rey, 1854: 153 [M]. Type species: Microsituslongulus Mulsant & Rey, 1854, by subsequent designation (Viñolas 1990: 65). Status: valid subgenus of Phylan Sturm, 1826 in Blaptinae: Dendarini: Dendarina.

Platyope Fischer, 1820: pl. 15 [F]. Type species: Tenebrioleucographus Pallas, 1781, by subsequent designation (Jacquelin du Val 1860: 261). Status: valid genus in Pimeliinae: Pimeliini. Note: Pallas (1773) used two different spellings for this type species, Tenebrioleucographus (p. 474) and Tenebrioleucogrammus (p. 719); subsequently, Pallas (1781: 54) used T.leucographus as the valid name of the species and so acted as First Reviser (ICZN 1999: Article 24.2.4).

Platyotus Gerstaecker, 1871: 62 [M]. Type species: Platyotusglabratus Gerstaecker, 1871, by monotypy. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Halstead (1967a: 72). Note: name also described as new in Gerstaecker (1873: 185).

Platyphanes Westwood, 1849: 206 [M]. Type species: Platyphanesgibbosus Westwood, 1849, by subsequent designation (Lacordaire 1859b: 410). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Platyphanus Koch, 1952b: 26 [M]. Type species: Acestussimilis Haag-Rutenberg, 1878, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Platyprocnemis Español & Lindberg, 1963: 19, 21 [F]. Type species: Trichosternumgranulosum Wollaston, 1868, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Platyprosodes Reitter, 1909a: 121 [M]. Type species: Nyctipatesrugulosus Gebler, 1841, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Skopin (1960a: 46).

Platypsorodes Ardoin, 1963b: 307, 310 [M]. Type species: Acanthomerushelopioides Guérin-Méneville, 1834, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Platyscelis Latreille, 1818: 23 [F]. Type species: Tenebriohypolithus Pallas, 1781, by plenary powers (ICZN 1993b, Opinion 1729). Status: valid genus and subgenus in Blaptinae: Platyscelidini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1993b, Opinion 1729).

Platysemodes Strand, 1935b: 302 [M]. Type species [automatic]: Platysemusbenguelensis Haag-Rutenberg, 1875, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Platysemus Haag-Rutenberg, 1875.

Platysemus Haag-Rutenberg, 1875b: 4, 55 [M]. Type species: Platysemusbenguelensis Haag-Rutenberg, 1875, by monotypy. Status: senior synonym of Platysemodes Strand, 1935 in Pimeliinae: Adelostomini. Note: junior homonym of Platysemus Middendorff, 1848 [Mollusca].

Platystena Koch, 1940c: 95 [F]. Type species: Mesostenablairi Koch, 1940, by monotypy. Status: valid subgenus of Mesostena Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: combined description of a new genus-group taxon and a single new species (ICZN 1999, Article 13.4).

Plegacerus Gebien, 1921b: 142 [M]. Type species: Plegacerussensitivus Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Pleioplatyscelis Kaszab, 1940a: 153, 220 [F]. Type species: Platynoscelislederi Seidlitz, 1893, by original designation. Status: junior synonym of Planoplatyscelis Kaszab, 1940 in Blaptinae: Platyscelidini. Synonymy: Egorov (2004: 601).

Pleiopleura Seidlitz, 1893: 342, 343 [F]. Type species: Platyscelisstriatus Motschulsky, 1859, by monotypy. Status: valid subgenus of Platyscelis Latreille, 1818 in Blaptinae: Platyscelidini.

Pleisiasida Smith, 2013: 610 [F]. Type species [automatic]: Parasidalaciniata Casey, 1912, by original designation. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini. Note: replacement name for Parasida Casey, 1912.

Plesia Klug, 1833: 25 [F]. Type species: Plesiamelanura Klug, 1833, by subsequent designation (Bousquet et al. 2015: 137). Status: senior synonym of Eubalia Laporte, 1840 in Alleculinae: Alleculini: Gonoderina. Synonymy: Borchmann (1909a: 713). Note: junior homonym of Plesia Jurine, 1807 [Hymenoptera].

Plesiamarygmus Masumoto, 1989b: 314 [M]. Type species: Dietysusovoideus Fairmaire, 1882, by original designation. Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer (2005: 205).

Plesioderes Mulsant & Rey, 1859c: 126 [M]. Type species: Caediuscoriaceus Mulsant & Rey, 1859, by subsequent designation (Gebien 1939: 466). Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Plesiognaptor Chigray, Nabozhenko & Keskin, 2015: 1277 [M]. Type species: Gnaptorprolixus Fairmaire, 1866, by original designation. Status: valid subgenus of Gnaptor Brullé, 1831 in Blaptinae: Blaptini: Gnaptorina.

Plesiophthalmus Motschulsky, 1857: 34 [M]. Type species: Plesiophthalmusnigrocyaneus Motschulsky, 1857, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Amarygmini.

Pleuromophlus Reitter, 1906b: 138, 146 [M]. Type species: Omophlusbaudueri Baudi, 1877, by subsequent designation (Bousquet et al. 2015: 141). Status: junior synonym of Odontomophlus Seidlitz, 1896 in Alleculinae: Cteniopodini. Synonymy: Bousquet et al. (2015: 141).

Pleurophorus Solier, 1851: 162 [M]. Type species: Pleurophorusquadricollis Solier, 1851, by monotypy. Status: senior synonym of Discopleurus Lacordaire, 1859 in Pimeliinae: Stenosini: Dichillina. Note: junior homonym of Pleurophorus Mulsant, 1842 [Coleoptera: Scarabaeidae].

Pleurostira Borchmann, 1921: 217, 230 [F]. Type species: Pleurostiraepipleuralis Borchmann, 1921, by original designation. Status: valid subgenus of Statira Lepeletier & Audinet-Serville, 1828 in Lagriinae: Lagriini: Statirina.

Pleuroxycara Koch, 1959: 582 [N]. Type species: Oxycaraproblematicum Koch, 1959, by original designation. Status: valid subgenus of Oxycara Solier, 1835 in Pimeliinae: Tentyriini.

Plicatocerus Pic, 1918a: 11 [M]. Type species: Otocerusimpressipennis Champion, 1888, by monotypy. Status: valid subgenus of Oploptera Chevrolat, 1844 in Stenochiinae: Stenochiini.

Plinthochrous Fairmaire, 1891b: 263 [M]. Type species: Plinthochrousgounellei Fairmaire, 1891, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Pocadiopsis Fairmaire, 1896a: 17 [F]. Type species: Pocadiopsissimulator Fairmaire, 1896, by subsequent designation (Gebien 1939: 457). Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Podacamptus Ardoin, 1964a: 849 [M]. Type species: Podacamptusruficolor Ardoin, 1964, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Podamarygmus Carter, 1928: 287 [M]. Type species: Podamarygmusalternatus Carter, 1928 (= Amarygmusviridipes Gebien, 1927), by monotypy. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Podhomala Solier, 1836: 9, 72 [F]. Type species: Podhomalasuturalis Solier, 1836, by monotypy. Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Podoces Péringuey, 1886: 122 [F]. Type species: Podocesgranosula Péringuey, 1886, by present designation. Status: senior synonym of Carchares Pascoe, 1887 in Tenebrioninae: Scaurini. Synonymy: Péringuey (1892b: [135]). Note: junior homonym of Podoces Fischer, 1821 [Aves].

Pododonta Agassiz, 1846b: 300 [F]. Type species [automatic]: Cistelanigrita Fabricius, 1794, by subsequent designation (R. Lucas 1920: 527). Status: junior synonym of Podonta Solier, 1835 in Alleculinae: Cteniopodini. Note: unjustified emendation of Podonta Solier, 1835, not in prevailing usage.

Podomala Agassiz, 1846b: 300 [F]. Type species [automatic]: Podhomalasuturalis Solier, 1836, by monotypy. Status: junior synonym of Podhomala Solier, 1836 in Pimeliinae: Pimeliini. Note: unjustified emendation of Podhomala Solier, 1836, not in prevailing usage.

Podonta Solier, 1835a: 247 [F]. Type species: Cistelanigrita Fabricius, 1794, by subsequent designation (Cazurro Ruiz 1894b: 883). Status: valid genus in Alleculinae: Cteniopodini.

Podontinus Seidlitz, 1896: 197 [M]. Type species: Cteniopuspunctatissimus Kiesenwetter, 1861, by subsequent designation (R. Lucas 1920: 527). Status: valid genus in Alleculinae: Cteniopodini.

Poecilesthostrongylium Pic, 1918a: 12 [N]. Type species: Strongyliumamazonicum Pic, 1918, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Poecilesthus Dejean, 1834: 207 [M]. Type species: Erotylusfasciatus Fabricius, 1781, by subsequent designation (Hope 1841: 133). Status: valid genus in Stenochiinae: Stenochiini. Note: Poecilesthus is an incorrect subsequent spelling of the original spelling Paecilesthus, first used by Dejean (1836: 229), and in prevailing usage; Poecilesthus is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1), see Bousquet and Bouchard (2013a: 60).

Poecilocrypticus Gebien, 1928: 121 [M]. Type species: Poecilocrypticusformicophilus Gebien, 1928, by monotypy. Status: valid genus in Diaperinae: Crypticini.

Poeciltoides Fairmaire, 1896b: 352 [M]. Type species: Poeciltoidesalternans Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pogonobasis Solier, 1837b: 153, 161 [F]. Type species: Pogonobasisopatroides Solier, 1837 (= Eurychorarugosula Guérin-Méneville, 1831), by subsequent designation (Hope 1841: 121). Status: valid genus in Pimeliinae: Adelostomini.

Pogonocanta Koch, 1952b: 22 [F]. Type species: Pogonobasislongipilis Fairmaire, 1894, by original designation. Status: valid genus in Pimeliinae: Adelostomini. Note: the alternative original spelling Pogonacanta, used by Koch (1952b: 123), was rejected by Koch (1953c: 3) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Pogonophloeus Bremer, 1998: 9 [M]. Type species: Hypophlaeusthoracicus Melsheimer, 1846, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Pogonoxenus Wasmann, 1899b: 172 [M]. Type species: Pogonoxenuskraatzi Wasmann, 1899, by monotypy. Status: valid genus in Diaperinae: Hypophlaeini.

Pokryszkiella Iwan, 1996: 385, 414 [F]. Type species: Pokryszkiellacornuta Iwan, 1996, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Polasida Reitter, 1917a: 39, 41 [F]. Type species: Opatrumsericeum G.-A. Olivier, 1795, by subsequent designation (Viñolas and Cartagena 2005: 188). Status: valid subgenus of Asida Latreille, 1802 in Pimeliinae: Asidini.

Polemiotus Casey, 1907: 379, 381 [M]. Type species: Epitragussubmetallicus J.L. LeConte, 1854, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Poliorcetes Champion, 1884: 70 [M]. Type species: Poliorcetesplatesthoides Champion, 1884, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Polopinus Casey, 1924: 326 [M]. Type species: Polypleurusnitidus J.L. LeConte, 1866, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Polpocara Solier, 1843: 46, 123 [N]. Type species: Polpocarapicipes Solier, 1843, by original designation. Status: junior synonym of Philorea Erichson, 1834 in Pimeliinae: Physogasterini. Synonymy: Erichson (1847a: 116).

Polpogenia Solier, 1836: 9, 70 [F]. Type species: Polpogeniaasidioides Solier, 1836, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Polposipus Solier, 1848: 154, 260 [M]. Type species: Polposipusherculeanus Solier, 1848, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Polycoelogastridion Reichardt, 1936: 85, 208 [N]. Type species: Sclerumsexcostatum Motschulsky, 1858, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Polyidus Champion, 1888: 441 [M]. Type species: Polyidusmeridionalis Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Polypleurus Eschscholtz, 1831: 10, 11 [M]. Type species: Polypleurusgeminatus Eschscholtz, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Polyscopus Waltl, 1835: 73 [M]. Type species: Polyscopuscostatus Waltl, 1835 (= Adelostomasulcatum Duponchel, 1827), by monotypy. Status: junior synonym of Adelostoma Duponchel, 1827 in Pimeliinae: Adelostomini. Synonymy: Erichson in Agassiz (1846a: 133).

Polytropus Kirsch, 1866: 201 [M]. Type species: Polytropuslaenoides Kirsch, 1866, by monotypy. Status: junior synonym of Chaetyllus Pascoe, 1860 in Lagriinae: Laenini. Synonymy: Kaszab (1983a: 138).

Ponapeida Kulzer, 1957: 242, 248 [F]. Type species: Ponapeidarufitarsis Kulzer, 1957, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pontianacus Fairmaire, 1898d: 397 [M]. Type species: Pontianacusrubricrus Fairmaire, 1898, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Poopterocoma Skopin, 1974b: 145 [F]. Type species: Pterocomazaidamica Skopin, 1974, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Porphyrhyba Fairmaire, 1877a: 137 [F]. Type species: Porphyrhybaviolaceicolor Fairmaire, 1877, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Porphyrohyba Rye, 1879: 62 [F]. Type species [automatic]: Porphyrhybaviolaceicolor Fairmaire, 1877, by monotypy. Status: junior synonym of Porphyrhyba Fairmaire, 1877 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Porphyrhyba Fairmaire, 1877, not in prevailing usage.

Porrolagria Kolbe, 1883: 26 [F]. Type species: Porrolagrianuda Kolbe, 1883, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Posides Champion, 1884: 6 [M]. Type species: Posidesdissidens Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Postandrosus Kulzer, 1951b: 490 [M]. Type species: Postandrosusmaculipennis Kulzer, 1951, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Postpraocis Flores & Pizarro-Araya, 2014: 60 [M]. Type species: Praocispentachorda Burmeister, 1875, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 12, 33) without type species designation.

Potocula Novák, 2012: 290 [F]. Type species: Potoculakubani Novák, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Praeugena Laporte, 1840: 241 [F]. Type species: Helopsmarginatus Fabricius, 1792, by subsequent designation (Hope 1841: 133). Status: valid genus in Tenebrioninae: Praeugenini. Note: nomen protectum (see Bouchard and Bousquet 2020b: 6).

†Praezolodinus Bao in Bao and Antunes-Carvalho, 2020: 2 [M]. Type species: Praezolodinuspilosus Bao, 2020, by original designation. Status: valid genus in Zolodininae. Note: described from Cretaceous Burmese amber (Myanmar).

Praocida Flores & Pizarro-Araya, 2014: 70 [F]. Type species: Praociszischkai Kulzer, 1958, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 13, 88) without type species designation.

Praocidia Fairmaire, 1904b: 463 [F]. Type species: Praocisnervosus Fairmaire, 1902, by original designation. Status: valid genus in Pimeliinae: Praociini.

Praocis Eschscholtz, 1829: 6 [M]. Type species: Praocisrufipes Eschscholtz, 1829, by subsequent designation (Guérin-Méneville 1834: 8–9). Status: valid genus and subgenus in Pimeliinae: Praociini. Note: although Guérin-Méneville (1834: 8–9) did not specifically mention the scientific name of the species he considered the type of this genus, his comment “J’ai donné une nouvelle figure de l’espèce qui a servi de type à Eschscholtz, dans la pl. 4, fig. 1, du Voyage de la Coquille” [translated to “I gave a new figure of the species that served as the type by Eschscholtz, in pl. 4, fig. 1, of the Voyage de la Coquille”] leaves no ambiguity regarding the identity of the species, i.e., Praocisrufipes Eschscholtz, 1829, to which he was referring.

Praogena Agassiz, 1846b: 306 [F]. Type species [automatic]: Helopsmarginatus Fabricius, 1792, by subsequent designation (Hope 1841: 133). Status: junior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Note: unjustified emendation of Praeugena Laporte, 1840, not in prevailing usage.

Praonoda Flores & Pizarro-Araya, 2014: 68 [F]. Type species: Praocisbicarinatus Burmeister, 1875, by original designation. Status: valid subgenus of Praocis Eschscholtz, 1829 in Pimeliinae: Praociini. Note: this name was first proposed by Kulzer (1958a: 13, 66) without type species designation.

Praostetha Fairmaire, 1897f: 138 [F]. Type species: Praostethaimpressifrons Fairmaire, 1897, by monotypy. Status: junior synonym of Amenophis J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1921b: 64).

Prateus J.L. LeConte, 1862: 238 [M]. Type species: Prateusfusculus J.L. LeConte, 1862, by original designation. Status: valid genus in Lagriinae: Goniaderini.

Predactylosis Penrith, 1977: 19, 243 [F]. Type species: Predactylosisholmi Penrith, 1977, by original designation. Status: junior synonym of Zophosis Latreille, 1802 in Pimeliinae: Zophosini. Synonymy: Penrith (1981a: 22).

Priocamaria Gebien, 1919: 28, 143 [F]. Type species: Priocamariamacilenta Gebien, 1919, by subsequent designation (Gebien 1942a: 323). Status: valid genus in Stenochiinae: Cnodalonini.

Prionalia Novák, 2020c: 511 [F]. Type species: Gonoderaatronitens Fairmaire, 1892, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Prionotheca Dejean, 1834: 179 [F]. Type species: Pimeliacoronata G.-A. Olivier, 1795, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Prionotus Mulsant & Rey, 1859c: 88 [M]. Type species: Opatrumdenticolle Blanchard, 1846, by monotypy. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Gemminger in Gemminger and Harold (1870: 1929, with Achora Pascoe, 1869, a junior synonym of Isopteron Hope, 1841). Note: junior homonym of Prionotus Lacepède, 1802 [Pisces].

Prionychus Solier, 1835a: 231, 237 [M]. Type species: Helopsater Fabricius, 1775, by subsequent designation (R. Lucas 1920: 536). Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: nomen protectum (see Bousquet and Bouchard 2017: 132); see Löbl and Smetana (2011: 33) for comments on the gender of this genus-group name.

Prioproctus Kolbe, 1903: 165, 174 [M]. Type species: Prioproctusoertzeni Kolbe, 1903, by original designation. Status: valid genus in Lagriinae: Pycnocerini.

Prioscelida White, 1846: 11 [F]. Type species: Prioscelidatenebrionoides White, 1846, by monotypy. Status: junior synonym of Uloma Dejean, 1821 in Tenebrioninae: Ulomini. Synonymy: Broun (1880: 365).

Prioscelides Kolbe, 1889: 128 [M]. Type species: Prioscelidesrugosus Kolbe, 1889, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Prioscelis Hope, 1841: 128 [F]. Type species: Prioscelisfabricii Hope, 1841, by original designation. Status: valid genus in Lagriinae: Pycnocerini. Note: the alternative original spelling Priopus, used by Hope (1841: 73), was rejected by Westwood (1844: 211) who acted as First Reviser.

Priothorax Gebien, 1910b: 318 [M]. Type species [automatic]: Opatrumdenticolle Blanchard, 1846, by monotypy. Status: junior synonym of Isopteron Hope, 1841 in Lagriinae: Adeliini. Synonymy: Gebien (1910b: 318, as a valid synonym of Achora Pascoe, 1869), Blair (1919a: 529, as a synonym of Achora Pascoe, 1869, a junior synonym of Isopteron Hope, 1841). Note: replacement name for Prionotus Mulsant & Rey, 1859.

Pristophilus Kolbe, 1903: 165, 174 [M]. Type species: Chiroscelispassaloides Westwood, 1842, by original designation. Status: valid genus in Lagriinae: Pycnocerini.

Probaticus Seidlitz, 1895: 697, 704, 764 [M]. Type species: Helopsmori Brullé, 1832, by subsequent designation (Gebien 1943: 419). Status: junior synonym of Euboeus Boieldieu, 1865 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko et al. (2017: 496). Note: the alternative original spelling Probatius (pp. 697, 704, 765) was corrected to Probaticus in the “Amendments and corrections” of the same work (p. 849), Probaticus is considered to be the correct original spelling (ICZN 1999, Article 32.5.1.1).

Prochoma Solier, 1835b: 253, 393 [N]. Type species: Prochomaaudouini Solier, 1835, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Proctenius Reitter, 1890b: 256 [M]. Type species: Cistelagranatensis Rosenhauer, 1856, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Proderops Fairmaire, 1873: 393 [M]. Type species: Proderopsforaminosus Fairmaire, 1873 (= Rhinandruselongatus Horn, 1867), by monotypy. Status: junior synonym of Rhinandrus J.L. LeConte, 1866 in Tenebrioninae: Tenebrionini. Synonymy: Kraatz (1880a: 132, with Exerestus Bates, 1870, a junior synonym of Rhinandrus J.L. LeConte, 1866).

Prodilamus Ardoin, 1969e: 258 [M]. Type species: Dilamusbrevicollis Fairmaire, 1894, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Prohylithus Kaszab, 1964c: 382 [M]. Type species: Prohylithuskulzeri Kaszab, 1964, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Prolabrus Fairmaire, 1897a: 111 [M]. Type species: Prolabrusparallelus Fairmaire, 1897, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Prolaena Kaszab, 1980b: 322 [F]. Type species: Laenaceylonica Motschulsky, 1858, by original designation. Status: valid genus in Lagriinae: Laenini. Note: Prolaena was used earlier by Kaszab (1979a: 107) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore not available from that date.

Proleptodes G.S. Medvedev, 1967: 354 [M]. Type species: Leptodessulcicollis Reitter, 1889, by original designation. Status: valid subgenus of Leptodes Dejean, 1834 in Pimeliinae: Leptodini.

Promethis Pascoe, 1869: 148 [M]. Type species: Upisangulata Erichson, 1842, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Prometopion Casey, 1907: 366, 370 [N]. Type species: Prometopionamplipenne Casey, 1907 (= Chilometoponhelopioides Horn, 1874), by original designation. Status: junior synonym of Chilometopon Horn, 1874 in Pimeliinae: Edrotini. Synonymy: MacLachlan and Olson (1990: 72).

Promorphostenophanes Kaszab, 1960b: 277 [M]. Type species: Promorphostenophanesatavus Kaszab, 1960, by original designation. Status: junior synonym of Morphostenophanes Pic, 1925 in Stenochiinae: Cnodalonini. Synonymy: Masumoto and Bečvář (2008: 206).

Promus J.L. LeConte, 1862: 226 [M]. Type species: Blapsopaca Say, 1824, by original designation. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Propemicrosis Penrith, 1981c: 127, 152 [F]. Type species: Microsistransbechuana Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Prophanes Westwood, 1849: 203 [M]. Type species: Prophanesaculeatus Westwood, 1849, by subsequent designation (Lacordaire 1859b: 411). Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Propterocoma Skopin, 1974b: 144 [F]. Type species: Pterocomatibialis Bates, 1879, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Prorhytinota Bouchard & Bousquet, new subgenus [F]. Type species: Rhytinotaoxyoma Fairmaire, 1884, by present designation. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: Koch (1943b: 761, 795) introduced the new subgenus name Prorhytinota for several nominal species, but unfortunately did not designate a type species; the subgenusProrhytinota, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Rhytinotaoxyoma Fairmaire, 1884 as type species and referring to Koch (1943b: 761, 795) for the character states that characterise and differentiate Prorhytinota.

Proscheimus Desbrochers des Loges, 1881: 127 [M]. Type species: Proscheimusarabicus Desbrochers des Loges, 1881, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Proscorus Fairmaire, 1901b: 188 [M]. Type species: Proscoruscyaneostriatus Fairmaire, 1901, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Proselytus Fåhraeus, 1870: 302 [M]. Type species: Proselytuscaffer Fåhraeus, 1870 (= Alphitobiusdiaperinus Panzer, 1796), by monotypy. Status: junior synonym of Alphitobius Stephens, 1829 in Tenebrioninae: Alphitobiini. Synonymy: Ferrer (1999: 266; through synonymy of the type species).

Prosoblapsia Skopin & Kaszab, 1978: 208 [F]. Type species: Leptocolenaallardiana Reitter, 1889, by original designation. Status: junior synonym of Genoblaps Bauer, 1921 in Blaptinae: Blaptini: Blaptina. Synonymy: new synonym [YB]. Note: Genoblaps Bauer, 1921 has been forgotten in the literature; its type species is currently included in the subgenusProsoblapsia Skopin & Kaszab, 1978 and for that reason Skopin & Kaszab’s name is considered a junior synonym of Genoblaps.

Prosodella Reitter, 1909a: 120 [F]. Type species: Prosodesbactriana Semenov, 1894, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Note: the First Reviser (Prosodella Reitter, 1909 versus Paraprosodes Reitter, 1909) is G.S. Medvedev (1997: 597).

Prosodes Eschscholtz, 1829: 9 [F]. Type species: Blapsattenuata Fischer, 1820 (= Blaps obtusa Fabricius, 1798), by subsequent designation (Gebien 1937a: 846). Status: valid genus and subgenus in Blaptinae: Blaptini: Prosodina. Note: the gender of Prosodes is feminine by indication of the author himself, who states: “von προσώδης, foetida”, the latter word being Latin and explaining the Greek meaning (stinking); the word foetida is feminine and therefore Prosodes is feminine also; note, however, that all other genus-group names in this list with the ending –prosodes are masculine.

Prosodestes Reitter, 1909a: 114 [M]. Type species: Prosodesgrandicollis Kraatz, 1883, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosodidius Fairmaire, 1903d: 69 [M]. Type species: Prosodidiusperrieri Fairmaire, 1903, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Prosodila Reitter, 1909a: 121 [F]. Type species: Prosodesstrigiventris Reitter, 1893, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Löbl et al. (2008c: 219).

Prosodinia Reitter, 1909a: 115 [F]. Type species: Prosodescalcarata Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosodopria Reitter, 1909a: 116 [F]. Type species: Blapsangustata Zubkov, 1833, by monotypy. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosodoscelis Reitter, 1909a: 117 [F]. Type species: Prosodessolskyi Faust, 1875, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosodura Reitter, 1909a: 118 [F]. Type species: Prosodessemenowi Reitter, 1893, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Prosomenes Blanchard, 1845: 10 [M]. Type species: Diceroderesmexicanus Solier, 1841, by subsequent monotypy (Chevrolat 1847b: 562). Status: junior synonym of Diceroderes Solier, 1841 in Tenebrioninae: Toxicini: Dysantina. Synonym: Lacordaire (1859a: 356, as “Prosomenes Dejean”), Bousquet et al. (2018: 219). Note: the name Prosomenes was originally introduced as a synonym of “Dicérodères” by Blanchard (1845: 10); however, because the name was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Chevrolat 1847b: 562) we treat it as available and dating from its first publication as a synonym (ICZN 1999, Article 11.6.1); originally proposed without included nominal species; Chevrolat (1847b: 562), by including the species “Pros.mexicanus” in association with this name, was the first author to subsequently and expressly include nominal species in Prosomenes (ICZN 1999, Article 67.2.2).

Prostenus Klug, 1829: 5 [M]. Type species: Prostenusperiscelis Perty, 1830, by subsequent designation (Bousquet et al. 2015: 139, but see Note). Status: valid genus in Alleculinae: Alleculini: Xystropodina. Note: as pointed out by Bousquet et al. (2015: 139) this genus name had been attributed to other authors in the literature (e.g., Latreille, 1825: 377, Berthold, 1827: 369) but it was first made available by Klug (1829: 5) who described two new species in it, Prostenusfemoratus and Prostenuspilosus, neither species being included in Prostenus in its currently accepted sense; Bousquet et al. (2015: 139) suggested using Prostenusperiscelis Perty, 1830 as the type species of Prostenus to promote nomenclatural stability; however, an application to the Commission is needed to confirm the type species of Prostenus.

†Proteleates Wickham,1914a: 267 [M]. Type species: Proteleatescentralis Wickham, 1914, by original designation. Status: valid genus in Tenebrioninae: Bolitophagini. Note: described from Upper Eocene deposits (USA).

Prothraustocola Kaszab, 1957: 293 [F]. Type species: Ibnsaudiabelutschistanica Kaszab, 1957, by original designation. Status: valid subgenus of Thraustocolus Kraatz, 1866 in Pimeliinae: Tentyriini.

Protoblaps Bauer, 1921: 230, 231 [F]. Type species: none designated. Status: undetermined taxon in Blaptinae: Blaptini: Blaptina. Note: this genus was described before 1931 (ICZN 1999, Article 12.1); however, we could not find any nominal species that were subsequently and expressly included in Protoblaps and therefore no “originally included nominal species” could be used to fix the type species (ICZN 1999, Article 67.2.2).

Protoblaps G.S. Medvedev, 1998a: 200 [F]. Type species: Protoblapskashkarovi G.S. Medvedev, 1998, by original designation. Status: senior synonym of Medvedevoblaps Bouchard & Bousquet, nom. nov. in Blaptinae: Blaptini: Blaptina. Note: junior homonym of Protoblaps Bauer, 1921 [Coleoptera: Tenebrionidae: Blaptinae: Blaptini: Blaptina].

Protocalosis Penrith, 1977: 21, 116 [F]. Type species: Zophosisbalti Penrith, 1977, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Protodactylus Koch, 1952a: 83 [M]. Type species: Protodactylusopticus Koch, 1952, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Protomachlasida Escalera, 1928: 137 [F]. Type species: Machlasidaliouvillei Escalera, 1925, by original designation. Status: junior synonym of Machlasida Escalera, 1907 in Pimeliinae: Asidini. Synonymy: Gebien (1937a: 717).

†Protoplatycera Wickham, 1914b: 484 [F]. Type species: Protoplatyceralaticornis Wickham, 1914, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Upper Eocene deposits (USA).

Prototyrtaeus Spiessberger & Ivie, 2020: 669 [M]. Type species: Prototyrtaeusdarlingtoni Spiessberger & Ivie, 2020, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Prunaspila Koch, 1950a: 67 [F]. Type species [automatic]: Aspilabicostata Fåhraeus, 1870, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Aspila Fåhraeus, 1870.

Przewalskia Semenov, 1893: 262 [F]. Type species: Platyopedilatata Reitter, 1887, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Psammestus Reichardt, 1936: 194, 216 [M]. Type species: Ammobiusdilatatus Reitter, 1893, by original designation. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Psammetichus Latreille, 1828: 578 [M]. Type species: Psammetichuscostatus Guérin-Méneville, 1831, by subsequent monotypy (Guérin-Méneville 1831b: pl. 28bis). Status: valid genus in Pimeliinae: Elenophorini: Megelenophorina. Note: originally proposed without included nominal species; Guérin-Méneville (1831b: pl. 28bis), by including the new species “Psammeticuscostatus Guérin-Méneville, 1831” in association with this name, was the first author to subsequently and expressly include nominal species in Psammetichus (ICZN 1999, Article 67.2.2).

Psammoardoinellus Leo, 1981: 34 [M]. Type species: Isocerussardiniensis Ardoin, 1972, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Psammocryptus Kraatz, 1865: 81, 239 [M]. Type species: Tentyriaminuta Tauscher, 1812, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Psammodes W. Kirby, 1819a: 412 [M]. Type species: Psammodeslongicornis W. Kirby, 1819, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Psammodophysis Péringuey, 1899: 296 [F]. Type species: Psammodophysisprobes Péringuey, 1899, by subsequent designation (Kamiński et al. 2019b: 31). Status: junior synonym of Psammodes W. Kirby, 1819 in Pimeliinae: Sepidiini: Molurina. Synonymy: Gebien (1910a: 154).

Psammoeca Agassiz, 1846b: 311 [F]. Type species [automatic]: Microderalucida Solier, 1835, by subsequent designation (Gebien 1937a: 611). Status: junior synonym of Psammoica Solier, 1835 in Pimeliinae: Tentyriini. Note: unjustified emendation of Psammoica Solier, 1835, not in prevailing usage.

Psammogaster Koch, 1953e: 237 [F]. Type species: Psammogastermalani Koch, 1953, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Psammoica Solier, 1835b: 307 [F]. Type species: Microderalucida Solier, 1835, by subsequent designation (Gebien 1937a: 611). Status: valid genus in Pimeliinae: Tentyriini.

Psammolophus Koch, 1953f: 154 [M]. Type species: Psammodesacuticosta Fairmaire, 1884, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psammophanes Lesne, 1922: 690 [M]. Type species: Moluriscatenata Reiche, 1850, by original designation. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Molurina.

Psammophrynopsis Koch, 1953f: 157 [F]. Type species: Phrynocolusfrommi Wilke, 1921, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psammophrynus Koch, 1953f: 146 [M]. Type species: Psammophanesjokli Koch, 1953, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psammoryssus Kolbe, 1886: 289 [M]. Type species: Psammoryssustitanus Kolbe, 1886, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Psammostretus Koch, 1953f: 145 [M]. Type species: Psammodesbisbicostatus Gebien, 1910, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psammotopulus Endrödy-Younga, 1996: 15, 29 [M]. Type species: Caenocrypticuspenrithae Endrödy-Younga, 1996, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini.

Psammotyria Koch, 1953f: 137 [F]. Type species: Psammodesertli Kolbe, 1904, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Psammotyriopsis Koch, 1953f: 144 [F]. Type species: Psammophanesbredoi Koch, 1953, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Psaryphis Erichson, 1843: 241 [F]. Type species: Psaryphisnana Erichson, 1843, by monotypy. Status: valid genus in Pimeliinae: Adelostomini.

Psaryphulum Koch, 1952b: 32 [N]. Type species: Adelostomaabbreviatum Haag-Rutenberg, 1875, by original designation. Status: valid subgenus of Adelostoma Duponchel, 1827 in Pimeliinae: Adelostomini.

Psectes Hesse, 1935: 572 [M]. Type species: Psectesbechuanus Hesse, 1935, by original designation. Status: valid genus in Blaptinae: Pedinini: Helopinina.

Psectrapus Solier, 1848: 153, 213 [M]. Type species: Psectrapusbipartitus Solier, 1848, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Psectrascelis Solier, 1836: 307, 311 [F]. Type species: Nycteliapilipes Guérin-Méneville, 1834, by subsequent designation (Gebien 1937a: 750). Status: valid genus in Pimeliinae: Nycteliini. Note: the First Reviser (Psectrascelis Solier, 1836 versus Cerostena Solier, 1836) is Fairmaire (1876: 356).

Psectropus Gemminger in Gemminger and Harold, 1870: 1911 [M]. Type species [automatic]: Psectrapusbibartitus Solier, 1848, by original designation. Status: junior synonym of Psectrapus Solier, 1848 in Blaptinae: Platynotini: Eurynotina. Note: unjustified emendation of Psectrapus Solier, 1848, not in prevailing usage.

Pselaphidion Gebien, 1921b: 29, 229 [N]. Type species: Platydemamacularia Gemminger, 1870, by subsequent designation (Gebien 1940: 405). Status: junior synonym of Stomylus Fåhraeus, 1870 in Diaperinae: Diaperini: Diaperina. Synonymy: Koch (1953g: 23). Note: the alternative original spelling Pselaphidium, used by Gebien (1921b: 28, 29), was rejected by Gebien (1940: 405) who acted as the First Reviser (ICZN 1999, Article 24.2.4).

Pseudabax Kraatz, 1880b: 107 [M]. Type species: Pseudabaxformosus Kraatz, 1880, by subsequent designation (Gebien 1942a: 308). Status: valid genus in Stenochiinae: Cnodalonini.

Pseudadelium Kaszab, 1982b: 234 [N]. Type species: Adeliumpustulosum Fauvel, 1904, by original designation. Status: junior synonym of Neoadelium Carter, 1908 in Lagriinae: Adeliini. Synonymy: Matthews (1998: 781).

Pseudadrus Fairmaire, 1897g: 39 [M]. Type species: Hadrusscaphoides Marseul, 1876, by monotypy. Status: junior synonym of Phelopatrum Marseul, 1876 in Blaptinae: Opatrini: Opatrina. Synonymy: Reichardt (1936: 90).

Pseudalymon Ardoin, 1969c: 543, 544 [M]. Type species: Asthenochirusfoveolatus Péringuey, 1899, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Pseudamarsenes Ardoin, 1955: 142 [M]. Type species: Amarsenesviridistriatus Fairmaire, 1894, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudamarygmus Pic, 1915d: 9 [M]. Type species: Pseudamarygmustestaceipes Pic, 1915, by subsequent designation (Gebien 1948: 509). Status: junior synonym of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini. Synonymy: Bremer (2001a: 67).

Pseudamblyptera Pierre, 1985: 295 [F]. Type species: Pimeliafrigida Escalera, 1925, by original designation. Status: valid subgenus of Pimelia Fabricius, 1775 in Pimeliinae: Pimeliini.

Pseudamenophis Pic, 1916: 13 [M]. Type species: Pseudamenophisaeneus Pic, 1916 (= Amenophisepipleuralis Gebien, 1904), by monotypy. Status: junior synonym of Amenophis J. Thomson, 1858 in Stenochiinae: Cnodalonini. Synonymy: Ardoin (1962a: 64).

Pseudanaedus Gebien, 1921b: 107, 111 [M]. Type species: Pseudanaedusbiangulatus Gebien, 1921, by subsequent designation (Gebien 1941: 820). Status: valid genus in Lagriinae: Goniaderini.

Pseudandrosus Kulzer, 1951b: 486 [M]. Type species: Chariothecaneomidina Fairmaire, 1881, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudanemia Wollaston, 1864: 492 [F]. Type species: Pseudanemiabrevicollis Wollaston, 1864, by monotypy. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini.

Pseudanopidium Dajoz, 1974: 431, 434 [N]. Type species: Pseudanopidiumpunctatum Dajoz, 1974, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Pseudapocrypha Champion, 1886: 260 [F]. Type species: Pseudapocryphalacordairii Champion, 1886, by monotypy. Status: valid genus in Tenebrioninae: Apocryphini.

Pseudapsida Kulzer, 1961: 219 [F]. Type species: Pseudapsidabrasiliensis Kulzer, 1961, by original designation. Status: junior synonym of Paniasis Champion, 1886 in Diaperinae: Diaperini: Diaperina. Synonymy: Ferrer and Ødegaard (2005: 637).

Pseudasida Fairmaire, 1895b: 444 [F]. Type species: Pseudasidaobesa Fairmaire, 1895, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Pseudeba Blackburn, 1903: 119 [F]. Type species: Pseudebanovica Blackburn, 1903, by monotypy. Status: valid genus in Tenebrioninae: Palorini. Note: transferred from “Tenebrionoidea: Colydiidae” by Carter and Zeck (1937: 194).

Pseudeleodes Blaisdell, 1909: 146 [M]. Type species: Eleodesgranosa J.L. LeConte, 1866, by monotypy. Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Pseudemmallus Koch, 1956a: 355 [M]. Type species: Pseudemmallusaspericollis Koch, 1956, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Pseudephalus Casey, 1924: 333 [M]. Type species: Pseudephalusbrevicornis Casey, 1924, by original designation. Status: junior synonym of Ephalus J.L. LeConte, 1862 in Blaptinae: Opatrini: Opatrina. Synonymy: Lumen et al. (2020: 344).

Pseudesarcus Champion, 1913: 115 [M]. Type species: Pseudesarcusvillosus Champion, 1913, by original designation. Status: valid genus in Lagriinae: incertae sedis. Note: originally described in Mycetophagidae, transferred to Tenebrionidae: Lagriinae by Lawrence and Newton (1995: 886).

Pseudethas Fairmaire, 1896a: 57 [M]. Type species: Pseudethasquadraticeps Fairmaire, 1896, by monotypy. Status: valid genus and subgenus in Pimeliinae: Stenosini: Dichillina.

Pseudethas Fairmaire, 1898c: 477 [M]. Type species: Pseudethaslongiceps Fairmaire, 1898, by monotypy. Status: senior synonym of Anethas Jakobson, 1924 in Pimeliinae: Stenosini: Stenosina. Note: junior homonym of Pseudethas Fairmaire, 1896 [Coleoptera: Tenebrionidae: Pimeliinae: Stenosini: Dichillina].

Pseudeucyrtus Pic, 1916e: 14 [M]. Type species: Pseudeucyrtusniasensis Pic, 1916 (= Cleomisviolaceipes Fairmaire, 1892), by subsequent designation (Gebien 1942a: 308). Status: junior synonym of Cleomis Fairmaire, 1892 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983b: 382). Note: we act as First Revisers and reject the alternative original spelling Pseudocyrtus, used by Pic (1916e: 14), in order to avoid homonymy with Pseudocyrtus Salmon, 1956 [Collembola].

Pseudeumolpus Kraatz, 1880b: 111 [M]. Type species: Pseudeumolpusbicolor Kraatz, 1880, by subsequent designation (Löbl et al. 2008b: 345). Status: junior synonym of Phaedis Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1914d: 71).

Pseudeuthriptera Bogatchev & Kryzhanovsky, 1955: 240 [F]. Type species: Trigonoscelisuzboica Bogatchev & Kryzhanovsky, 1955, by monotypy. Status: junior synonym of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini. Synonymy: Skopin (1964: 395).

Pseudeutrapela Pic, 1952e: 110 [F]. Type species: Pseudeutrapelanigra Pic, 1952, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Pseudhadrus Kolbe, 1910: 31 [M]. Type species: Pseudhadrusseriatus Kolbe, 1910, by subsequent designation (Gebien 1941: 332). Status: valid genus in Stenochiinae: Cnodalonini.

Pseudhelops Guérin-Méneville, 1841: 124 [M]. Type species: Pseudhelopstuberculatus Guérin-Méneville, 1841, by monotypy. Status: valid genus in Tenebrioninae: Heleini: incertae sedis. Note: transferred from Titaenini by Matthews and Lawrence (2019: 628).

Pseudimmedia Kulzer, 1958b: 213 [F]. Type species: Pseudimmediafreyi Kulzer, 1958, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudisopus Kulzer, 1957: 254 [M]. Type species: Pseudisopusgressitti Kulzer, 1957, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudobasides Pic, 1916c: 2 [M]. Type species: Pseudobasidescornutus Pic, 1916, by subsequent designation (Gebien 1940: 419). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Pseudobates Fairmaire, 1882a: 231 [M]. Type species: Nyctobatescoracinus Fairmaire, 1882 (= Nyctobatessubrobustus Motschulsky, 1872), by monotypy. Status: junior synonym of Promethis Pascoe, 1869 in Stenochiinae: Cnodalonini. Synonymy: Kolbe (1900: 74, with Setenis Motschulsky, 1872, a junior synonym of Promethis Pascoe, 1869).

Pseudoblapida Pic, 1917: 18 [F]. Type species: Blapidaboliviensis Pic, 1912, by monotypy. Status: valid genus and subgenus in Stenochiinae: Cnodalonini.

Pseudoblaps Guérin-Méneville, 1834: 28 [F]. Type species: Pseudoblapssubstriata Guérin-Méneville, 1834, by subsequent designation (Hope 1841: 124). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Pseudobolbophanes Kulzer, 1954a: 29 [M]. Type species: Pseudobolbophanesmalaisei Kulzer, 1954, by original designation. Status: junior synonym of Bolbophanes Carter, 1913 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 490).

Pseudobradymerus Pic, 1926b: 16 [M]. Type species: Bradymerussimplicithorax Pic, 1926, by monotypy. Status: junior synonym of Bradymerus Perroud & Montrouzier, 1865 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1939: 751).

Pseudobyrsax Kaszab, 1982b: 214 [M]. Type species: Pseudobyrsaxcurtus Kaszab, 1982, by original designation. Status: valid genus in Lagriinae: Adeliini.

Pseudocaedius Blackburn, 1890a: 91 [M]. Type species: Pseudocaediussquamosus Blackburn, 1890 (= Trigonotarsusaustralis Hope, 1843), by monotypy. Status: junior synonym of Sobas Pascoe, 1863 in Blaptinae: Opatrini: Opatrina. Synonymy: Carter (1921: 307).

Pseudocaedius G.S. Medvedev, 1966: 98 [M]. Type species: Pseudocaediuskiseritzkii G.S. Medvedev, 1966, by original designation. Status: senior synonym of Asiocaedius G.S. Medvedev & Nepesova, 1985 in Blaptinae: Opatrini: Ammobiina. Note: junior homonym of Pseudocaedius Blackburn, 1890 [Coleoptera: Tenebrionidae: Blaptinae: Opatrini: Ammobiina].

Pseudocaelophus Pic, 1922d: 28 [M]. Type species: Strongyliumdifforme Pic, 1922, by monotypy. Status: junior synonym of Leprocaulus Fairmaire, 1896 in Stenochiinae: Cnodalonini. Synonymy: Bečvář and Purchart (2008: 39).

Pseudocamaria Bates, 1879a: 287 [F]. Type species: Camariaalternata Fairmaire, 1875, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudocamarimena Pic, 1923e: 21 [F]. Type species: Pseudocamarimenastriata Pic, 1923, by monotypy. Status: junior synonym of Pigeus Gebien, 1919 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1984: 363).

Pseudocasnonidea Borchmann, 1936: 240, 489 [F]. Type species: Pseudocasnonideaceylanica Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Pseudochariotheca Pic, 1934c: 32 [F]. Type species: Pseudochariothecaminutissima Pic, 1934, by original designation. Status: junior synonym of Steneucyrtus Fairmaire, 1896 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 136).

Pseudochillus Fouquè, 2015: 226, 240 [M]. Type species: Indochillusbangaloreanus Kaszab, 1981, by original designation. Status: valid genus and subgenus in Pimeliinae: Stenosini: Dichillina.

Pseudochrysomela Pic, 1925a: 7 [F]. Type species: Pseudeumolpusseriatoporus Fairmaire, 1888, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: the older name Pseudochrysomela Voet, 1806 [Coleoptera: Erotylidae] was published in a work that did not include consistent application of binominal nomenclature and is therefore unavailable (ICZN 1999, Article 11.4).

Pseudocilibe Kaszab, 1982b: 231 [F]. Type species: Celibeasidaeformis Fauvel, 1904, by original designation. Status: valid genus in Lagriinae: Adeliini. Note: we act as First Revisers and reject the alternative original spelling Pseudocylibe, used by Kaszab (1982b: 291).

Pseudocistela Blackburn, 1891: 316 [F]. Type species: Pseudocistelaovalis Blackburn, 1891, by monotypy. Status: senior synonym of Neocistela Borchmann, 1909 in Alleculinae: Alleculini: Alleculina. Note: junior homonym of Pseudocistela Crotch, 1874 [Coleoptera: Tenebrionidae: Alleculinae: Alleculini: Gonoderina].

Pseudocistela Crotch, 1874: 108 [F]. Type species: Cistelabrevis Say, 1824, by subsequent designation (Novák and Pettersson 2008: 327). Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Pseudocistelopsis Novák, 2018b: 176 [M]. Type species: Pseudocistelopsisjakli Novák, 2018, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Pseudocoelus Casey, 1908: 152 [M]. Type species: Coeluspacificus Fall, 1897, by subsequent designation (Doyen 1976: 608). Status: junior synonym of Coelus Eschscholtz, 1829 in Pimeliinae: Coniontini. Synonymy: Blaisdell (1919: 322).

Pseudocolparthrum Borchmann, 1916b: 230, 236 [N]. Type species: Colparthrumcalcaratum Champion, 1889, by subsequent designation (Borchmann 1936: 452). Status: valid subgenus of Colparthrum Kirsch, 1866 in Lagriinae: Lagriini: Statirina.

Pseudoderiles Gebien, 1928: 169, 181 [M]. Type species: Pseudoderilesdentipennis Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudoderosphaerus Pic, 1922d: 24 [M]. Type species: Leprocaulusrotundicollis Pic, 1922, by monotypy. Status: junior synonym of Leprocaulus Fairmaire, 1896 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983c: 177).

Pseudodiaphanidus Bogatchev, 1950: 234 [M]. Type species: Diaphanidusrobustus Bogatchev, 1950, by monotypy. Status: valid subgenus of Diaphanidus Reitter, 1900 in Pimeliinae: Erodiini.

Pseudoelongasida Escalera, 1922c: 173 [F]. Type species: Asidasilvestrei Escalera, 1922, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Elongasida Escalera, 1906 in Pimeliinae: Asidini. Synonymy: Pérez-Vera et al. (2017: 3).

Pseudoenanea Pic, 1924a: 25 [F]. Type species: Pseudoenanearobusta Pic, 1924, by monotypy. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Pseudogena Fairmaire, 1899e: 539 [F]. Type species: Pseudogenapolyzona Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Pseudognaptorina Kaszab, 1977c: 250 [F]. Type species: Pseudognaptorinanepalica Kaszab, 1977, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

†Pseudohelops Haupt, 1950: 130 [M]. Type species: Pseudohelopsgroenlandicus Haupt, 1950, by monotypy. Status: valid genus in Stenochiinae: incertae sedis. Note: combined description of new genus-group taxon and new species (ICZN 1999, Article 13.4); described from Middle Paleocene deposits (Greenland).

Pseudohymenalia Novák, 2008b: 213 [F]. Type species: Pseudohymenaliayunnanica Novák, 2008, by original designation. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Pseudolagria Champion, 1917: 218 [F]. Type species: Pseudolagriamutabilis Champion, 1917, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Pseudolamus Fairmaire, 1874: 388 [M]. Type species: Pseudolamusseriatoporus Fairmaire, 1874, by monotypy. Status: valid genus in Blaptinae: Opatrini: Neopachypterina.

Pseudoleichenum Ardoin, 1972: 205 [N]. Type species: Pseudoleichenumbenoiti Ardoin, 1972, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Pseudolyprops Fairmaire, 1882a: 236 [M]. Type species: Pseudolypropsdilaticollis Fairmaire, 1882, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Pseudomachla Wilke, 1922: 260 [F]. Type species [automatic]: Opatrumvillosum G.-A. Olivier, 1795, by subsequent designation (R. Lucas 1920: 386). Status: junior synonym of Machla Herbst, 1799 in Pimeliinae: Asidini. Note: unnecessary replacement name for Machla Herbst, 1799.

Pseudomorocaulus Pic, 1915c: 6 [M]. Type species: Pseudomorocaulusrufonotatus Pic, 1915, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Pseudonautes Fairmaire, 1892c: 52 [M]. Type species: Pseudonautesvagevittatus Fairmaire, 1892, by subsequent designation (R. Lucas 1920: 551). Status: valid genus in Stenochiinae: Cnodalonini.

Pseudonomus Fairmaire, 1884a: 510 [M]. Type species: Pseudonomusdermestiformis Fairmaire, 1884, by monotypy. Status: junior synonym of Ammodonus Mulsant, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1939: 470).

Pseudonotocorax Iwan, 1997: 255, 269 [M]. Type species: Pseudonotocoraxmroczkowskii Iwan, 1997, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Pseudoogeton Masumoto, 1989b: 304 [M]. Type species: Plesiophthalmusamplipennis Fairmaire, 1897, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Pseudopachyscelis Skopin, 1968a: 99 [F]. Type species: Trigonoscelispygmaea Ménétriés, 1849, by original designation. Status: valid genus in Pimeliinae: Pimeliini. Note: first proposed by Skopin (1962: 225) without a type species originally designated.

Pseudoparablops Heyden, 1908: 132 [M]. Type species: Parablopssardiniensis Allard, 1877, by monotypy. Status: junior synonym of Allardius Ragusa, 1898 in Tenebrioninae: Helopini: Helopina. Synonymy: Reitter (1922b: 167).

Pseudopatrum Sharp, 1886: 406 [N]. Type species: Pseudopatrumsordidum Sharp, 1886, by subsequent designation (Watt 1992: 30). Status: valid genus in Lagriinae: Adeliini.

Pseudopedinus Ardoin, 1969d: 178 [M]. Type species: Pseudopedinuslaosensis Ardoin, 1969, by original designation. Status: valid subgenus of Loensus R. Lucas, 1920 in Blaptinae: Pedinini: Pedinina.

Pseudopenthicinus Bogatchev, 1972: 628 [M]. Type species: Penthicusmedvedevi Bogatchev, 1972, by original designation. Status: valid subgenus of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina.

Pseudoperichilus Pic, 1921d: 21 [M]. Type species: Pseudoperichilusolivaceus Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudophthora Kaszab, 1970a: 267 [F]. Type species: Pseudophthoralaeana Kaszab, 1970, by original designation. Status: valid genus and subgenus in Phrenapatinae: Penetini.

Pseudopigeus Kaszab, 1984: 355, 359 [M]. Type species: Pseudopigeusunidentatus Kaszab, 1984, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudopimelia Gebler, 1859: 473 [F]. Type species: Lasiostolavariolaris Gebler, 1841, by monotypy. Status: junior synonym of Lasiostola Dejean, 1834 in Pimeliinae: Pimeliini. Synonymy: new synonym [YB]. Note: Pseudopimelia Gebler, 1859 has been forgotten in the literature; its type species is currently included in the nominotypical subgenus of Lasiostola Dejean, 1834 and for that reason Gebler’s name is considered a new junior synonym of Lasiostola.

Pseudopimelia Motschulsky, 1860c: 536 [F]. Type species: Pterocomatuberculata Motschulsky, 1845, by subsequent designation (Skopin 1974b: 159). Status: senior synonym of Subpterocoma Bouchard & Bousquet, nom. nov. in Pimeliinae: Pimeliini. Note: junior homonym of Pseudopimelia Gebler, 1859 [Coleoptera: Tenebrionidae: Pimeliinae: Pimeliini].

Pseudoplanasida Escalera, 1921: 360 [F]. Type species: Asidapygmaea Rosenhauer, 1856, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Gracilasida Escalera, 1905 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 192, with Planasida Escalera, 1907, a junior synonym of Gracilasida Escalera, 1905).

Pseudoplatyope Pierre, 1964: 867, 873 [F]. Type species: Storthocnemisantoinei Español, 1951, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Pseudopodhomala Schuster, 1938: 88 [F]. Type species: Pseudopodhomalagabrieli Schuster, 1938, by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Pseudopodhomalina Kaszab, 1960a: 22 [F]. Type species: Diesiacostifera C.O. Waterhouse, 1889, by original designation. Status: junior synonym of Pseudopodhomala Schuster, 1938 in Pimeliinae: Pimeliini. Synonymy: Kwieton (1982: 35).

Pseudopraeugena De Moor, 1970: 8,40 [F]. Type species: Pseudopraeugenarufa De Moor, 1970, by original designation. Status: valid genus in Tenebrioninae: Praeugenini.

Pseudoprobaticus Nabozhenko, 2001b: 513 [M]. Type species: Helopsgranipennis Allard, 1876, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Pseudoprosodes Reitter, 1909a: 120 [M]. Type species: Prosodestransfuga Reitter, 1893, by original designation. Status: junior synonym of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Synonymy: Skopin (1960a: 46).

Pseudopterocoma Skopin, 1974b: 145 [F]. Type species: Pterocomatrapezicollis Skopin, 1974, by original designation. Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini.

Pseudorozonia Bouchard & Bousquet, new subgenus [F]. Type species: Rozoniaconophthalma Koch, 1944, by present designation. Status: valid subgenus of Rozonia Fairmaire, 1888 in Pimeliinae: Tentyriini. Koch (1944a: 162) introduced the new subgenus name Pseudorozonia for three nominal species, but unfortunately did not designate a type species; the subgenusPseudorozonia, which has been treated as valid since 1944, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Rozoniaconophthalma Koch, 1944 as type species and referring to Koch (1944a: 162) for the character states that characterise and differentiate Anemiadena.

Pseudortheolus Freude, 1968: 110 [M]. Type species: Epitragusminutissimus Fairmaire, 1892, by original designation. Status: valid genus in Pimeliinae: Epitragini.

Pseudoscaphidema Pic, 1926c: 2 [F]. Type species: Pseudoscaphidemarufonotata Pic, 1926, by monotypy. Status: valid genus in Diaperinae: Scaphidemini.

Pseudoscotobius Kulzer, 1955a: 384, 393 [M]. Type species: Emmalloderastrangulata Fairmaire, 1905, by original designation. Status: junior synonym of Phrynocarenum Gebien, 1928 in Tenebrioninae: Phrynocarenini. Synonymy: Marcuzzi (1976: 117).

Pseudoselinus Iwan, 2002a: 48, 94 [M]. Type species: Eurynotuspunctatostriatus Gerstaecker, 1854, by original designation. Status: valid subgenus of Upembarus Koch, 1956 in Blaptinae: Platynotini: Platynotina.

Pseudoseriscius Español, 1950: 125 [M]. Type species: Pedinuspruinosus Dufour, 1820, by original designation. Status: valid genus and subgenus in Diaperinae: Crypticini.

Pseudostene Wollaston, 1861: 247 [F]. Type species: Pseudosteneangusta Wollaston, 1861, by subsequent designation (Löbl et al. 2008b: 317). Status: junior synonym of Phtora Germar, 1836 in Diaperinae: Phaleriini. Synonymy: Escalera (1914: 347).

Pseudosternoplax Skopin, 1973: 110, 141 [F]. Type species: Trigonoscelislacerta Bates, 1879, by original designation. Status: valid subgenus of Sternoplax Frivaldszky, 1890 in Pimeliinae: Pimeliini.

Pseudostira Fairmaire, 1903c: 213 [F]. Type species: Pseudostiralaevipennis Fairmaire, 1903, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Pseudostorthocnemis Gridelli, 1952: 83 [F]. Type species: Storthocnemispatrizii Gridelli, 1933, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Pseudostrongylium Kraatz, 1880b: 114 [N]. Type species: Pseudostrongyliumsemperi Kraatz, 1880, by subsequent designation (Kaszab 1977b: 11). Status: junior synonym of Lophocnemis Mäklin, 1867 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 543).

Pseudotalpophila Reitter, 1900c: 95 [F]. Type species: Thalpophilaplicifrons Wollaston, 1864, by subsequent designation (Gebien 1937a: 641). Status: valid subgenus of Hegeter Latreille, 1802 in Pimeliinae: Tentyriini.

Pseudothinobatis Bouchard & Bousquet, new genus [F]. Type species: Thinobatisohausi Kulzer, 1956, by present designation. Status: valid genus in Pimeliinae: Epitragini. Note: Freude (1960b: 32) introduced the new genus name Pseudothinobatis for two nominal species, but unfortunately did not designate a type species; the genus Pseudothinobatis, which has been treated as valid since 1960, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Thinobatisohausi Kulzer, 1956 as type species and referring to Freude (1960b: 32) for the character states that characterise and differentiate Pseudothinobatis.

Pseudothryoneus Pic, 1921b: 12 [M]. Type species: Pseudothryoneusbicoloripes Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pseudotocerus Champion, 1888: 383 [M]. Type species: Stenochialongipes P.H. Lucas, 1859, by subsequent designation (Gebien 1948: 542). Status: valid genus in Stenochiinae: Stenochiini.

Pseudotrichoplatyscelis Kaszab, 1960a: 83, 111 [F]. Type species: Platynoscelisbadakschanica Kaszab, 1960 (= Bioramixlapidicola Kaszab, 1940), by original designation. Status: junior synonym of Trichoplatyscelis Reinig, 1931 in Blaptinae: Platyscelidini. Synonymy: Egorov (1990: 402). Note: we act as First Revisers and reject the alternative original spellings Pseudotrichoplatynoscelis and Pseudotrichoplatycelis, used by Kaszab (1960a: 82, 83).

Pseuduloma Fairmaire, 1893b: 27 [N]. Type species: Pseudulomacribricolle Fairmaire, 1893 (= Ulomimusindicus Bates, 1873), by monotypy. Status: junior synonym of Ulomimus Bates, 1873 in Tenebrioninae: Ulomini. Synonymy: Gebien (1940: 770).

Pseuduroplatopsis Pic, 1913a: 16 [F]. Type species: Borchmanniajavana Pic, 1913, by present designation. Status: valid subgenus of Borchmannia Pic, 1912 in Lagriinae: Lagriini: Statirina.

Psilachnopus Reitter, 1901: 161 [M]. Type species: Psilachnopuscribratellus Reitter, 1901, by monotypy. Status: junior synonym of Philhammus Fairmaire, 1871 in Pimeliinae: Cnemeplatiini: Cnemeplatiina. Synonymy: Gebien (1938a: 420), Löbl and Smetana (2010: 30).

Psilocastus Ardoin, 1963a: 149 [M]. Type species: Psilocastusletestui Ardoin, 1963, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: Psilocastus was used earlier by Ardoin (1962b: 970) without designation of a type species and is therefore unavailable from that date.

Psilolaena Heller, 1923: 70 [F]. Type species: Psilolaenaschusteri Heller, 1923, by monotypy. Status: junior synonym of Laena Dejean, 1821 in Lagriinae: Laenini. Synonymy: Schawaller (2001b: 34).

Psilomera Motschulsky, 1870: 400 [F]. Type species: Pelecyphorusangulatus J.L. LeConte, 1851, by monotypy. Status: junior synonym of Stenomorpha Solier, 1836 in Pimeliinae: Asidini. Synonymy: Bousquet in Bousquet et al. (2018: 74).

Psilonesogena Bates, 1879a: 305 [F]. Type species: Psilonesogenahybrida Bates, 1879, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Psis Novák, 2019d: 71 [M]. Type species: Psisnanensis Novák, 2019, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Psoroderes Ardoin, 1962b: 969, 1017 [M]. Type species: Psorodeshottentottus Péringuey, 1899, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Psorodes Dejean, 1834: 189 [M]. Type species [automatic]: Pimeliadentipes Fabricius, 1787, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: replacement name for Acanthomera Latreille, 1828.

Psorophodes Ardoin, 1963a: 83 [F]. Type species: Pimeliaarmata Herbst, 1799, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini. Note: Psorophodes was used earlier by Ardoin (1962b: 969) without designation of a type species and is therefore unavailable from that date.

Psydocamaria Pic, 1923d: 17 [F]. Type species: Psydocamariarobusta Pic, 1923, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Psydomorphus Pic, 1921d: 24 [M]. Type species: Psydomorphusdiversipes Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Psydus Pascoe, 1868: xii [M]. Type species: Psydusplantaris Pascoe, 1868, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Pteraulus Solier, 1848: 152, 200 [M]. Type species: Pteraulussulcatipennis Solier, 1848, by present designation. Status: junior synonym of Helopinus Solier, 1848 in Blaptinae: Pedinini: Helopinina. Synonymy: Koch (1958: 149).

Pterelaeus Gemminger in Gemminger and Harold, 1870: 1968 [M]. Type species [automatic]: Pterohelaeuswalkerii Brême, 1842, by subsequent designation (Gebien 1940: 1069). Status: junior synonym of Pterohelaeus Brême, 1842 in Tenebrioninae: Heleini: Heleina. Note: unjustified emendation of Pterohelaeus Brême, 1842, not in prevailing usage.

Pterna Reitter, 1884: 249 [F]. Type species: Ernocharisauricoma Reitter, 1884, by monotypy. Status: valid subgenus of Mycetochara Guérin-Méneville, 1827 in Alleculinae: Alleculini: Mycetocharina. Note: as mentioned by Bousquet et al. (2015: 138) this name was originally proposed as a synonym of Mycetochares Latreille, 1829 but was used as valid subsequently and adopted as the name of a taxon (e.g., Seidlitz 1896: 132, 137) before 1961; therefore, it is available and dates from its publication as a synonym (ICZN 1999, Article 11.6.1).

Pterocoma Dejean, 1834: 178 [F]. Type species: Pimeliapiligera Gebler, 1830, by monotypy. Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Pterocomodes Reitter, 1901: 159 [M]. Type species: Pterocomodesacutus Reitter, 1901, by monotypy. Status: junior synonym of Podhomala Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Skopin (1962: 247).

Pteroctenus Kirsch, 1866: 193 [M]. Type species: Pteroctenuspexus Kirsch, 1866, by monotypy. Status: valid genus in Pimeliinae: Falsomycterini.

Pterodes Ardoin, 1963a: 97 [M]. Type species: Hoplonyxasper Péringuey, 1899, by original designation. Status: valid genus in Tenebrioninae: Amarygmini. Note: Pterodes was used earlier by Ardoin (1962: 970) without a type species designation and is therefore unavailable from that date.

Pteroglymmius Gebien, 1928: 219, 223 [M]. Type species: Pteroglymmiuserotyloides Gebien, 1928, by monotypy. Status: junior synonym of Isaminas Champion, 1887 in Stenochiinae: Cnodalonini. Synonymy: Doyen (1988: 301).

Pterohelaeus Brême, 1842b: 17, 27 [M]. Type species: Pterohelaeuswalkerii Brême, 1842, by subsequent designation (Gebien 1940: 1069). Status: valid genus in Tenebrioninae: Heleini: Heleina.

Pterolasia Solier, 1836: 9, 66 [F]. Type species: Pterolasiasqualida Solier, 1836, by subsequent designation (Hope 1841: 118). Status: valid genus in Pimeliinae: Pimeliini.

Pteroselinus Kamiński, 2015a: 92, 94 [M]. Type species: Opatrinusinsularis Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Pterostichula Koch, 1952d: 224 [F]. Type species: Pterostichulacalathoides Koch, 1952, by original designation. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Oxurina.

Ptilonix Allard, 1877: 62 [M]. Type species: Helopsclavicrus Marseul, 1876, by subsequent designation (Löbl et al. 2008a: 42). Status: junior synonym of Misolampidius Solsky, 1876 in Stenochiinae: Cnodalonini. Synonymy: Lewis (1894: 476).

Pubamarygmus Pic, 1915d: 8 [M]. Type species: Pubamarygmusviridipennis Pic, 1915, by subsequent designation (Gebien 1948: 510). Status: valid genus in Tenebrioninae: Amarygmini.

Pubeirosoma Pic, 1954: 257 [N]. Type species: Isomiradiscoglabrata Pic, 1954, by monotypy. Status: valid subgenus of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina.

Pulposipes Gemminger in Gemminger and Harold, 1870: 1974 [M]. Type species [automatic]: Polposipusherculeanus Solier, 1848, by original designation. Status: junior synonym of Polposipus Solier, 1848 in Stenochiinae: Cnodalonini. Note: unjustified emendation of Polposipus Solier, 1848, not in prevailing usage.

Pumiliofossorum Silvestro & Giraldo-Mendoza in Silvestro et al., 2015: 462, 470 [N]. Type species: Pumiliofossorummoche Silvestro & Flores, 2015, by original designation. Status: valid genus in Tenebrioninae: Scotobiini.

Punctacula Campbell, 1971: 112 [F]. Type species: Punctaculahowdeni Campbell, 1971, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Pushtunillus G.S. Medvedev, 1995a: 855, 865 [M]. Type species: Dichillusskopini G.S. Medvedev & Kabakov, 1995, by original designation. Status: valid subgenus of Dichillus Jacquelin du Val, 1860 in Pimeliinae: Stenosini: Dichillina.

Pyanirygmus Pic, 1915d: 9 [M]. Type species: Pyanirygmuscorinthius Pic, 1915, by monotypy. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Pyanisia Laporte, 1840: 235 [F]. Type species: Helopsundatus Fabricius, 1792 (= Erotylusnebulosus Fabricius, 1781), by subsequent designation (Lacordaire 1859b: 476). Status: junior synonym of Cymatothes Dejean, 1834 in Tenebrioninae: Amarygmini. Synonymy: Chevrolat (1847b: 643).

Pycna Fairmaire, 1894d: 68 [F]. Type species: Pycnaaphodina Fairmaire, 1894, by monotypy. Status: senior synonym of Madagassa Koch, 1950 in Phrenapatinae: Penetini. Note: redescribed as new by Fairmaire (1894b: 141); junior homonym of Pycna Amyot & Audinet-Serville, 1843 [Hemiptera].

Pycnocerus Westwood, 1841b: 67 [M]. Type species [automatic]: Pachyloceruswestermanni Hope, 1841, by monotypy. Status: valid genus and subgenus in Lagriinae: Pycnocerini. Note: replacement name for Pachylocerus Hope, 1841.

Pycnochilus C.O. Waterhouse, 1879b: 263 [M]. Type species: Pycnochilusadvenus C.O. Waterhouse, 1879, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Pycnomorpha Motschulsky, 1870: 398 [F]. Type species: Pycnomorphacalifornica Motschulsky, 1870, by monotypy. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Pycnonotida Casey, 1912: 75, 89 [F]. Type species: Microschatiainaequalis J.L. LeConte, 1851, by original designation. Status: junior synonym of Microschatia Solier, 1836 in Pimeliinae: Asidini. Synonymy: K.W. Brown and Doyen (1992: 546).

Pycnuloma Fairmaire, 1896c: 99 [N]. Type species: Pycnulomaraffrayi Fairmaire, 1896, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Pygidiphorus Mulsant, 1856b: [1, supplement] [M]. Type species: Pygidiphoruscaroli Mulsant, 1856, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: included in a list of “Tenebrionidaenomina dubia” by Iwan et al. (2020: 475).

Pygmaeodes Koch, 1952d: 223 [M]. Type species: Namibomodesrudebecki Koch, 1952, by monotypy. Status: valid subgenus of Palpomodes Koch, 1952 in Pimeliinae: Sepidiini: Oxurina.

Pyres Champion, 1885: 100 [M]. Type species: Pyresmetallicus Champion, 1885 (= Centronopusspeciosus Pascoe, 1883), by subsequent designation (Gebien 1941: 336). Status: junior synonym of Menechides Motschulsky, 1872 in Tenebrioninae: Centronopini. Synonymy: Spilman (1962b: 3).

†Pyrochalcaspis Haupt, 1950: 113, 115 [F]. Type species: Pyrochalcaspisgiseltalensis Haupt, 1950, by original designation. Status: valid genus in Stenochiinae: incertae sedis. Note: described from Middle Eocene deposits (Germany).

Pystelops Gozis, 1910: 103 [M]. Type species: Helopsmeridianus Mulsant, 1854, by original designation. Status: valid subgenus of Stenomax Allard, 1876 in Tenebrioninae: Helopini: Cylindrinotina.

Pythiopus Koch, 1953e: 245 [M]. Type species: Pythiopuscornutipectus Koch, 1953, by original designation. Status: valid genus in Blaptinae: Dendarini: Dendarina.

Pythonissus Gistel, 1834: 21 [M]. Type species: Helopsmorio Fabricius, 1777 (= Tenebrioatratus Fabricius, 1775), by subsequent designation (Bousquet and Bouchard 2017: 132). Status: junior synonym of Zophobas Dejean, 1834 in Tenebrioninae: Tenebrionini. Synonymy: Bousquet and Bouchard (2017: 132).

Quadrideres Koch, 1956a: 189 [M]. Type species: Anchophthalmusscutatus Gerstaecker, 1871, by original designation. Status: junior synonym of Glyptopteryx Gebien, 1910 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 91).

Quadroncotus Koch, 1954a: 40 [M]. Type species: Oncotusirrepertus Koch, 1954, by original designation. Status: valid subgenus of Oncotus Blanchard, 1845 in Blaptinae: Platynotini: Eurynotina.

Raiboscelis Allard, 1876a: 5 [F]. Type species: Helopscorvinus Küster, 1850, by subsequent designation (Gebien 1943: 415). Status: valid genus in Tenebrioninae: Helopini: Helopina. Note: nomen protectum (see Nabozhenko and Löbl 2009: 194); Raiboscelis is an incorrect subsequent spelling of the original spelling Raibosceles and is in prevailing usage; Raiboscelis is deemed to be the correct original spelling (ICZN 1999, Article 33.3.1), see Nabozhenko and Löbl (2009: 189).

Raptor Gistel, 1848a: ix, xi [M]. Type species [automatic]: Melaphorusreichii Guérin-Méneville, 1834, by monotypy. Status: junior synonym of Melaphorus Guérin-Méneville, 1834 in Pimeliinae: Evaniosomini. Note: unnecessary replacement name for Melaphorus Guérin-Méneville, 1834.

Rasphytus Kulzer, 1956a: 639 [M]. Type species: Rasphytusfreyi Kulzer, 1956, by original designation. Status: valid subgenus of Apentanodes Reitter, 1914 in Pimeliinae: Erodiini.

Raynalius Chatanay, 1912a: 297 [M]. Type species: Raynaliushispidus Chatanay, 1912, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Rehumius Fairmaire, 1893b: 32 [M]. Type species: Rehumiusamplithorax Fairmaire, 1893, by subsequent designation (Gebien 1942a: 312). Status: valid genus in Stenochiinae: Cnodalonini.

Reichardtiella Kaszab, 1942: 18 [F]. Type species: Reichardtiellaarmata Kaszab, 1942, by original designation. Status: senior synonym of Reichardtiellina Kaszab, 1982 in Blaptinae: Opatrini: Opatrina. Note: junior homonym of Reichardtiella Filipjev, 1928 [Lepidoptera].

Reichardtiellina Kaszab, 1982c: 79 [F]. Type species [automatic]: Reichardtiellaarmata Kaszab, 1942, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: replacement name for Reichardtiella Kaszab, 1942 (as “Reichardtia Kaszab, 1940”; see Iwan and Löbl 2007: 735).

Reichenspergeria Wasmann, 1921: 18, 19 [F]. Type species: Reichenspergeriaaurocincta Wasmann, 1921, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Reitterella Semenov, 1891: 362 [F]. Type species: Reitterellafusiformis Semenov, 1891, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Dichillina.

Reitterellus Escalera, 1944: 91 [M]. Type species: Dendarusdepressus Reitter, 1915, by monotypy. Status: junior synonym of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina. Synonymy: Español (1961a: 44).

Reitterohelops Skopin, 1960b: 308, 309 [M]. Type species: Zophohelopslazarus Reitter, 1922, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Reminius Casey, 1924: 321 [M]. Type species: Reminiusocularis Casey, 1924 (= Tenebrioterminatus Say, 1824), by original designation. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Spilman (1959: 63).

Remipedella Semenov-Tjan-Shansky, 1907b: 257 [F]. Type species: Remipedelladeserti Semenov-Tjan-Shansky, 1907, by monotypy. Status: valid genus in Blaptinae: Blaptini: Remipedellina.

Renatiella Koch, 1944b: 155 [F]. Type species: Macropodareticulata Gerstaecker, 1854, by monotypy. Status: valid genus in Pimeliinae: Adesmiini. Note: the First Reviser (Renatiella Koch, 1944 versus Spongesmima Koch, 1944) is Penrith (1979: 27).

Renefouqueosis Aalbu, Smith, Kanda & Bouchard, 2017: 314 [F]. Type species: Renefouqueosisperuviensis Aalbu, Smith, Kanda & Bouchard, 2017, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Rhacius Champion, 1885: 120 [M]. Type species: Rhaciussulcatulus Champion, 1885, by subsequent designation (Gebien 1941: 805). Status: junior synonym of Adelonia Laporte, 1840 in Lagriinae: Belopini. Synonymy: Spilman (1961: 50).

Rhacolaena Kaszab, 1979b: 299 [F]. Type species: Rhacolaenatarsalis Kaszab, 1979, by original designation. Status: valid genus in Lagriinae: Laenini.

Rhaebosceles Rye, 1878: 69 [F]. Type species [automatic]: Helopscorvinus Küster, 1850, by subsequent designation (Gebien 1943: 415). Status: junior synonym of Raiboscelis Allard, 1876 in Tenebrioninae: Helopini: Helopina. Note: unjustified emendation of Raiboscelis Allard, 1876, not in prevailing usage.

Rhagostira Borchmann, 1936: 238, 468 [F]. Type species: Rhagostiracollarti Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Rhaibodera Borchmann, 1921: 217, 219 [F]. Type species: Rhaiboderapachycera Borchmann, 1921 (= Statiraeurydera Champion, 1917), by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Rhaibogria Borchmann, 1936: 17, 144 [F]. Type species: Lagriaampla Fairmaire, 1887, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Rhammatodes Haag-Rutenberg, 1876: 83 [M]. Type species: Rhammatodeslongicornis Haag-Rutenberg, 1876, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: the First Reviser (Rhammatodes Haag-Rutenberg, 1876 versus Euleantus Haag-Rutenberg, 1876) is Koch (1952a: 133).

Rhicnodus Fairmaire, 1892f: 87 [M]. Type species: Rhicnodusasper Fairmaire, 1892, by subsequent designation (R. Lucas 1920: 568). Status: junior synonym of Chaetyllus Pascoe, 1860 in Lagriinae: Laenini. Synonymy: Ardoin (1969b: 126), Kaszab (1983a: 132).

Rhinandrus J.L. LeConte, 1866b: 119 [M]. Type species: Rhinandrusgracilis J.L. LeConte, 1866, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Rhinobarus Reitter, 1906b: 131 [M]. Type species: Cistelasulphuripes Germar, 1823, by subsequent designation (Iablokoff-Khnzorian 1983: 133). Status: valid subgenus of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini.

†Rhinohelaeites Haupt, 1950: 115, 140 [M]. Type species: Rhinohelaeiteslongipes Haupt, 1950, by original designation. Status: valid genus in Tenebrionidae: incertae sedis. Note: described from Middle Eocene deposits (Germany); this genus was previously considered to be “definitely not a tenebrionid” by Watt (1975: 389) but is included here following Nabozhenko (2019: 8).

Rhipidandrus J.L. LeConte, 1862: 236 [M]. Type species: Xylotinusflabellicornis Sturm, 1826 (= Melolonthaparadoxa Palisot de Beauvois, 1818), by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Rhipidonyx Reitter, 1876b: 304 [M]. Type species: Rhipidonyxadustus Reitter, 1876, by monotypy. Status: valid genus in Alleculinae: incertae sedis. Note: taxon originally described in Mycetophagidae, transferred to Tenebrionidae by Lawrence et al. (2014: 209).

Rhizalemus Reitter, 1904: 79 [M]. Type species: Dendarusreitteri Seidlitz, 1893, by subsequent designation (Chatzimanolis and Löbl 2003: 260). Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Rhizalus Gebien, 1938a: 303 [M]. Type species [automatic]: Opatrumpiceum G.-A. Olivier, 1812, by monotypy. Status: junior synonym of Rizalus Mulsant & Rey, 1854 in Blaptinae: Dendarini: Dendarina. Note: unjustified emendation of Rizalus Mulsant & Rey, 1854, not in prevailing usage.

Rhizoblaps Motschulsky, 1860c: 532 [F]. Type species: Blapspruinosa Faldermann, 1833, by subsequent designation (Nabozhenko 2008: 36). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860)

Rhomaleus Chatanay, 1915b: 64 [M]. Type species: Rhomaleusscauroides Chatanay, 1915, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Rhopalobates Fairmaire, 1897c: 230 [M]. Type species: Rhopalobatesvillardi Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Rhophobas Motschulsky, 1872: 36 [M]. Type species: Rhophobasasperatus Motschulsky, 1872, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Rophobas, used by Motschulsky (1872: 26).

Rhosaces Champion, 1889: 73 [M]. Type species: Rhosacesclavipes Champion, 1889, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Rhostax Fischer von Waldheim, 1844: 67 [M]. Type species: Rhostaxmenetriesii Fischer von Waldheim, 1844 (= Microderagracilis Eschscholtz, 1831), by subsequent designation (G.S. Medvedev 1990: 109). Status: junior synonym of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Kraatz (1865: 184).

Rhydimorpha Koch, 1943b: 768, 863 [F]. Type species: Tentyriasubfossulata Solier, 1835, by monotypy. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: we act as First Revisers and reject the alternative original spelling Rhytimorpha, used by Koch (1943b: 888), since Rhytimorpha Szépligeti, 1901 is available in Hymenoptera.

Rhypasma Pascoe, 1862: 325 [N]. Type species: Rhypasmapusillum Pascoe, 1862, by monotypy. Status: valid genus in Lagriinae: Belopini.

Rhysodina Wasmann, 1921: 16 [F]. Type species [automatic]: Rhyzodinamniszechii Chevrolat, 1873, by monotypy. Status: junior synonym of Rhyzodina Chevrolat, 1873 in Tenebrioninae: Rhysopaussini. Note: unjustified emendation of Rhyzodina Chevrolat, 1873, not in prevailing usage.

Rhysopaussus Wasmann, 1896: 616 [M]. Type species: Rhysopaussusdohertyi Wasmann, 1896, by monotypy. Status: valid genus in Tenebrioninae: Rhysopaussini.

Rhytidonota Agassiz, 1846b: 327 [F]. Type species [automatic]: Rhytinotascabriuscula Eschscholtz, 1831, by subsequent designation (Chevrolat 1848: 279, as “Rytinota”). Status: junior synonym of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: unjustified emendation of Rhytinota Eschscholtz, 1831 (as “Rytinota”), not in prevailing usage.

Rhytinopsis Bouchard & Bousquet, new subgenus [F]. Type species: Rhytinotafossulata Kraatz, 1880, by present designation. Status: valid subgenus of Thalpophilodes Strand, 1942, in Pimeliinae: Tentyriini. Note: Koch (1943b: 774, 870) introduced the new subgenus name Rhytinopsis for several nominal species, but unfortunately did not designate a type species; the subgenusRhytinopsis, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Rhytinotafossulata Kraatz, 1880 as type species and referring to Koch (1943b: 774, 870) for the character states that characterise and differentiate Rhytinopsis.

Rhytinota Eschscholtz, 1831: 5, 7 [F]. Type species: Rhytinotascabriuscula Eschscholtz, 1831, by subsequent designation (Chevrolat 1848: 279). Status: valid genus and subgenus in Pimeliinae: Tentyriini. Note: this name is an unjustified emendation of the original spelling Rytinota, introduced by Chevrolat (1848: 279), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Rhytistena Bouchard & Bousquet, new subgenus [F]. Type species: Rhytistenagridelli Koch, 1943, by present designation. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: Koch (1943b: 769) introduced the new subgenus name Rhytistena (also spelled Rhytisten on page 888 of the same work) for several nominal species, but unfortunately did not designate a type species; the subgenusRhytistena, which has been treated as valid since 1943, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Rhytistenagridelli Koch, 1943 as type species and referring to Koch (1943b: 769) for the character states that characterise and differentiate Rhytistena.

Rhyzodina Chevrolat, 1873: 208 [F]. Type species: Rhyzodinamniszechii Chevrolat, 1873, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Rhysopaussini.

Ripicolodes Koch, 1952d: 225, 230 [M]. Type species: Pterostichulamisanthropa Koch, 1952, by original designation. Status: valid subgenus of Pterostichula Koch, 1952 in Pimeliinae: Sepidiini: Oxurina.

Rizalus Mulsant & Rey, 1854: 104 [M]. Type species: Opatrumpiceum G.-A. Olivier, 1812, by monotypy. Status: valid subgenus of Dendarus Dejean, 1821 in Blaptinae: Dendarini: Dendarina.

Robustocamaria Pic, 1922b: 25 [F]. Type species: Camariafortipes Pic, 1922, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Robustosora Pic, 1954: 232 [F]. Type species: Robustosorafreynei Pic, 1954, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Rondoniella Kaszab, 1970c: 112 [F]. Type species: Rondoniellacostata Kaszab, 1970, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Rondoniellina.

Rouyerus Pic, 1911b: 3 [M]. Type species: Rouyerusbimaculatus Pic, 1911, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Rozonia Fairmaire, 1888a: 184 [F]. Type species: Rozoniastrigicollis Fairmaire, 1888, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Ruandania Pic, 1955: 182 [F]. Type species: Ruandaniarufescens Pic, 1955, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Rues Casey, 1891: 66 [M]. Type species: Helopsovipennis Casey, 1890, by monotypy. Status: junior synonym of Adelium W. Kirby, 1819 in Lagriinae: Adeliini. Synonymy: Spilman (1959: 63). Note: as pointed out by Spilman (1959: 63) the type species, which was erroneously described from the United States of America, is in fact Australiana.

Rugasida Escalera, 1922b: 65 [F]. Type species: Asidareticulata Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Asida Latreille, 1802 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 291).

Rugoplatynotus Kaszab, 1975b: 281, 288 [M]. Type species: Pseudoblapsandrewesii Fairmaire, 1896, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Rugosiheliofugus Freude, 1960a: 125, 128 [M]. Type species: Heliophygussulcatulus Gemminger, 1870, by original designation. Status: valid subgenus of Heliofugus Guérin-Méneville, 1831 in Stenochiinae: Cnodalonini.

Rhyssochiton Gray in Griffith and Pidgeon, 1831: pl. 50 [M]. Type species: Rhyssochitonpolitus Gray, 1831, by monotypy. Status: junior synonym of Blapida Perty, 1830 in Stenochiinae: Cnodalonini. Synonymy: Lacordaire (1859b: 425). Note: two spellings originally appeared in plates of the same work, Ryssocheton (Gray in Griffith and Pidgeon, 1831: pl. 50) and Rhyssochiton (Gray in Griffith and Pidgeon, 1831: pl. 69); the third spelling Ryssochiton was also used in the “Index for plates” of Volume 15 of the same work (Gray in Griffith and Pidgeon, 1832: 793); we act as First Revisers and select the alternative original spelling Rhyssochiton.

Sabularius Escalera, 1914: 353 [M]. Type species: Helopsfossor Escalera, 1914, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Sabulophosis Penrith, 1977: 19, 227 [F]. Type species: Zophosisgaerdesi Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Saccophorella Strand, 1935b: 303 [F]. Type species [automatic]: Saccophoruscrenulatus Haag-Rutenberg, 1872, by monotypy. Status: junior synonym of Horatoma Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Note: replacement name for Saccophorus Haag-Rutenberg, 1872.

Saccophorus Haag-Rutenberg, 1872: 274, 303 [M]. Type species: Saccophoruscrenulatus Haag-Rutenberg, 1872, by monotypy. Status: junior synonym of Horatoma Solier, 1841 in Pimeliinae: Cryptochilini: Cryptochilina. Synonymy: Penrith and Endrödy-Younga (1994: 10). Note: junior homonym of Saccophorus Kuhl, 1820 [Mammalia].

Sadanaria Ando & Ichiyanagi, 2009: 80 [F]. Type species: Sadanariasakaii Ando & Ichiyanagi, 2009, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Saeculum Kamiński, Kanda & Smith, 2021: 196 [N]. Type species: Saeculumzoologicum Kamiński, Kanda & Smith, 2021, by original designation. Status: valid genus in Pimeliinae: Asidini.

Saerangodes Sturm, 1843: 163 [M]. Type species: Helopsinterpunctatus Germar, 1823, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Blanchard (1845: 33, with Stenochia W. Kirby, 1819, a synonym of Styrongylium W. Kirby, 1819).

Saharoplarion Koch, 1948: 431 [N]. Type species: Microsituscompactus Fairmaire, 1880, by original designation. Status: valid subgenus of Hoplarion Mulsant & Rey, 1854 in Blaptinae: Dendarini: Melambiina.

Saitostrongylium Masumoto, 1996a: 34 [N]. Type species: Saitostrongyliumacco Masumoto, 1996, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Sakaiomenimus Ando, 2003b: 135 [M]. Type species: Sakaiomenimustodai Ando, 2003, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Salax Guérin-Méneville, 1834: 11 [M]. Type species: Salaxlacordairii Guérin-Méneville, 1834, by monotypy. Status: valid genus in Pimeliinae: Trilobocarini.

Saptine Champion, 1886: 180 [F]. Type species: Saptineovata Champion, 1886, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Sarachus Gistel, 1848a: viii [M]. Type species [automatic]: Adesmialongipes Fischer, 1822 (= Pimeliaanomala Fischer, 1820), by monotypy. Status: junior synonym of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: unnecessary replacement name for Adesmia Fischer, 1822.

Saragella Carter, 1937: 136 [F]. Type species: Saragellapalpalis Carter, 1937, by monotypy. Status: junior synonym of Dysarchus Pascoe, 1866 in Tenebrioninae: Heleini: Heleina. Synonymy: Matthews (1993: 1059).

Saragodinus Bates, 1872b: 269 [M]. Type species: Saragodinusduboulayi Bates, 1872, by subsequent designation (R. Lucas 1920: 579). Status: junior synonym of Dysarchus Pascoe, 1866 in Tenebrioninae: Heleini: Heleina. Synonymy: Carter (1921: 307).

Saragus Erichson, 1842a: 171 [M]. Type species: fixed herein (ICZN 1999, Article 70.3) as Celibecostata Solier, 1848, misidentified as Silphalaevicollis Fabricius, 1775 in the original designation by monotypy in Erichson (1842a). Status: valid genus in Tenebrioninae: Heleini: Heleina. Note: the type species “Silphalaevicollis Fabricius” was first established by monotypy; Schaum (1850: 183) first noted that Silphalaevicollis Fabricius of Erichson (1842a) was misidentified and corresponded to Celibecostata Solier, 1848 (as “Cilibecostatus”); we follow currently accepted concepts (e.g., Matthews and Bouchard 2008: 310) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Silphalaevicollis Fabricius, 1775 is a valid species in the genus Boreosaragus Matthews, 1993 [Coleoptera: Tenebrionidae].

Sarandonyx Gozis, 1881: 151 [M]. Type species [automatic]: Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Westwood 1838: 32). Status: junior synonym of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: unnecessary replacement name for Cteniopus Solier, 1835.

Sarothropus Kraatz, 1865: 268 [M]. Type species: Akisdepressa Zoubkoff, 1837, by monotypy. Status: valid genus in Pimeliinae: Akidini.

Satanocalcar Pic, 1925b: 9 [N]. Type species: Satanocalcarcornutum Pic, 1925, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Saxistena Löbl & Merkl in Löbl et al., 2020: 3 [F]. Type species: Mesostenalongicornis Kraatz, 1865, by original designation. Status: valid subgenus of Mesostena Eschscholtz, 1831 in Pimeliinae: Tentyriini. Note: name first proposed by Koch (1940c: 65) without fixation of a type species in the original publication (ICZN 1999, Article 13.3); Löbl and Merkl (2003: 252) designated Mesostenalongicornis Kraatz, 1865 as the type species of Koch’s name but did not explicitly indicate the genus-group name as intentionally new (ICZN 1999, Article 16.1).

Saziches Champion, 1886: 261 [M]. Type species: Sazichessubcaudatus Champion, 1886, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Scaletomerus Blackburn, 1891: 330 [M]. Type species: Scaletomerusharpaloides Blackburn, 1891 (= Cistelapolitus W.J. MacLeay, 1872), by subsequent designation (R. Lucas 1920: 580). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Scaphidema Redtenbacher, 1848: 591 [F]. Type species: Scaphidiumbicolor Fabricius, 1798, by monotypy. Status: valid genus in Diaperinae: Scaphidemini. Note: see Kerzhner (2003: 322) and Löbl and Smetana (2010: 34) for comments on the gender of this name.

Scaphinion Matthews, 2012: 2 [M]. Type species: Scaphinionclavatus Matthews, 2012, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Scaptes Champion, 1886: 222 [M]. Type species: Scaptessquamulatus Champion, 1886 (= Asidatropica Kirsch, 1866), by subsequent designation (Bousquet et al. 2018: 194). Status: junior synonym of Ammodonus Mulsant, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Fall (1912: 48).

Scauris Rafinesque, 1815: 113 [F]. Type species [automatic]: Scaurusatratus Fabricius, 1775, by monotypy. Status: junior synonym of Scaurus Fabricius, 1775 in Tenebrioninae: Scaurini. Note: unjustified emendation of Scaurus Fabricius, 1775, not in prevailing usage.

Scaurus Fabricius, 1775: 253 [M]. Type species: Scaurusatratus Fabricius, 1775, by monotypy. Status: valid genus in Tenebrioninae: Scaurini.

Scelace Marseul, 1887: 325 [F]. Type species: Pimeliatuberculifera P.H. Lucas, 1858, by subsequent designation (Bouchard and Bousquet 2020b: 7). Status: valid genus in Pimeliinae: Pimeliini. Note: Scelace was published by Marseul on June 9, 1887 while its junior synonym Pachyscelodes was published by Sénac (1887: 189) on August 15 of the same year.

Sceleodis Gemminger in Gemminger and Harold, 1870: 1836 [M]. Type species [automatic]: Cratopuscastaneus Eschscholtz, 1831, by monotypy. Status: junior synonym of Scelosodis Solier, 1835 in Pimeliinae: Tentyriini. Note: unjustified emendation of Scelosodis Solier, 1835, not in prevailing usage.

Sceleoides Agassiz, 1846b: 333 [M]. Type species [automatic]: Cratopuscastaneus Eschscholtz, 1831, by monotypy. Status: junior synonym of Scelosodis Solier, 1835 in Pimeliinae: Tentyriini. Note: unjustified emendation of Scelosodis Solier, 1835, not in prevailing usage.

Scelidospecta Kulzer, 1954b: 204 [F]. Type species: Entomodereslobatus Burmeister, 1875, by original designation. Status: valid genus in Pimeliinae: Nycteliini. Note: Scelidospecta is an incorrect subsequent spelling of the original name Scelidopsecta in prevailing usage and attributed to the publication of the original spelling; Scelidospecta is treated as the correct original spelling (ICZN 1999, Article 33.3.1).

Scelocolpis Reitter, 1900c: 105 [M]. Type species: Colposcelisdamone Reitter, 1900, by subsequent designation (G.S. Medvedev 1990: 64). Status: valid subgenus of Colposcelis Dejean, 1834 in Pimeliinae: Tentyriini.

Scelosodis Solier, 1835b: 253, 283 [M]. Type species [automatic]: Cratopuscastaneus Eschscholtz, 1831, by monotypy. Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Cratopus Eschscholtz, 1831.

Schedarosus Reitter, 1876a: 42 [M]. Type species: Schedarosuscucujiformis Reitter, 1876 (= Pythopallidus Say, 1823), by subsequent designation (Löbl et al. 2008a: 42). Status: junior synonym of Adelina Dejean, 1835 in Diaperinae: Diaperini: Adelinina. Synonymy: Champion (1886: 157, with Doliema Pascoe, 1860, a junior synonym of Adelina Dejean, 1835).

Schelodontes Koch, 1956a: 81 [M]. Type species: Trigonopusimmundus Mulsant & Rey, 1853, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Schevodera Borchmann, 1936: 16, 59 [F]. Type species: Lagriahirticollis Borchmann, 1911, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Schevogria Borchmann, 1936: 20, 164 [F]. Type species: Schevogriamethneri Borchmann, 1936, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Schizaraeus Kulzer, 1955b: 479 [M]. Type species: Schizaraeusacuticosta Kulzer, 1955, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Schizillus Horn, 1874: 33 [M]. Type species: Schizilluslaticeps Horn, 1874, by monotypy. Status: valid genus in Pimeliinae: Cryptoglossini.

Schizillus Wasmann, 1899a: 166 [M]. Type species: Schizillusrogersi Wasmann, 1899, by monotypy. Status: junior synonym of Pseudethas Fairmaire, 1896 in Pimeliinae: Stenosini: Dichillina. Synonymy: Gebien (1937a: 679). Note: junior homonym of Schizillus Horn, 1874 [Coleoptera: Tenebrionidae: Pimeliinae: Cryptoglossini].

Schizomma Gebien, 1921a: 325, 392 [N]. Type species: Schizommacucumericola Gebien, 1921, by original designation. Status: senior synonym of Diachoriops Ando, 2020 in Stenochiinae: Cnodalonini. Note: junior homonym of Schizomma Ehrenberg, 1861 [Protista].

Schizophthalmotribolium Kaszab, 1940c: 173 [N]. Type species: Schizophthalmotriboliumaustraliae Kaszab, 1940, by original designation. Status: junior synonym of Paratoxicum Champion, 1894 in Tenebrioninae: Tenebrionini. Synonymy: Matthews (2004: 261).

Schlinkus R. Lucas, 1920: 584 [M]. Type species [automatic]: Cyphonotusdromedarius Guérin-Méneville, 1831, by monotypy. Status: junior synonym of Homocyrtus Dejean, 1834 in Tenebrionidae: incertae sedis. Synonymy: Gebien (1948: 514). Note: replacement name for Cyphonotus Guérin-Méneville, 1831.

Schoenicus J.L. LeConte, 1866b: 109 [M]. Type species: Schoenicuspuberulus J.L. LeConte, 1866, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Schoeniphegoneus Freude, 1968: 99 [M]. Type species: Epitragussemicastaneus Curtis, 1845, by original designation. Status: valid subgenus of Phegoneus Casey, 1907 in Pimeliinae: Epitragini.

Schusteriella Koch, 1940b: 745, 746 [F]. Type species: Stenosisruficornis Reitter, 1886, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Schweinfurthia Andres in Kneucker, 1922: 26 [F]. Type species: Schweinfurthiasinaitica Andres, 1922, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Schyzoschelus Koch, 1954a: 72 [M]. Type species: Schyzoscheluskaszabi Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Sciaca Solier, 1835b: 408 [F]. Type species: Hylithusdisctinctus Solier, 1835, by present designation. Status: junior synonym of Hylithus Guérin-Méneville, 1834 in Pimeliinae: Edrotini. Synonymy: Chevrolat (1848: 423). Note: the name Sciaca was originally listed as synonym of Hylithus Guérin-Méneville, 1834; it was treated before 1961 as an available name and adopted as the name of a taxon (e.g., Dejean 1836: 204), Sciaca was therefore made available from its first publication as a synonym (ICZN 1999, Article 11.6.1).

Sciophagus Sharp in Blackburn and Sharp, 1885: 167 [M]. Type species: Ulomapandanicola Boisduval, 1835, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Scleroides Fairmaire, 1883a: 32 [M]. Type species: Scleroidespluricostatus Fairmaire, 1883, by monotypy. Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Scleron Hope, 1841: 111 [N]. Type species: Opatrumorientale Fabricius, 1775, by original designation. Status: junior synonym of Sclerum Dejean, 1834 in Blaptinae: Opatrini: Sclerina. Synonymy: Erichson (1842b: 237); Bouchard et al. (2005: 511).

Scleronimon Reitter, 1904: 127 [M]. Type species: Eurycaulusgranulatus Reitter, 1904, by subsequent designation (Löbl and Merkl 2003: 249). Status: junior synonym of Eurycaulus Fairmaire, 1868 in Blaptinae: Opatrini: Sclerina. Synonymy: Grimm (2005: 9).

Scleronopsis Koch, 1935: 86, 87 [F]. Type species: Scleronhirsutus Miller, 1861, by monotypy. Status: junior synonym of Eurycaulus Fairmaire, 1868 in Blaptinae: Opatrini: Sclerina. Synonymy: Grimm (2005: 9).

Scleropatroides Löbl & Merkl, 2003: 250 [M]. Type species: Opatrumhirtulum Baudi di Selve, 1876, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Scleropatrum Reitter, 1887a: 388 [N]. Type species: Scleropatrumtuberculatum Reitter, 1887, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Sclerum Dejean, 1834: 193 [N]. Type species: Opatrumorientale Fabricius, 1775, by subsequent designation (Hope 1841: 110). Status: valid genus in Blaptinae: Opatrini: Sclerina.

Scolytocaulus Fairmaire, 1896c: 98 [M]. Type species: Scolytocaulusbouchardi Fairmaire, 1896, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Scopulophosis Penrith, 1977: 19, 215 [F]. Type species: Zophosiscacozela Koch, 1958, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Scotaeus Hope, 1834: 14 [M]. Type species: Scotaeuscorallipes Hope, 1834, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Scotera Motschulsky, 1845b: 365 [F]. Type species: Scoteragibbosa Motschulsky, 1845, by monotypy. Status: junior synonym of Cibdelis Mannerheim, 1843 in Stenochiinae: Cnodalonini. Synonymy: Motschulsky (1845b: 365). Note: as pointed out by Bousquet et al. (2018: 296) the generic name Scotera and the specific name gibbosa, proposed as synonyms of Cibdelisblaschkii by Motschulsky (1845b: 365, 366), are available from Motschulsky’s work (ICZN 1999, Article 11.6.1) because they were treated as valid names before 1961 (e.g., Chevrolat 1848: 454).

Scotinesthes Fairmaire, 1895a: 19 [F]. Type species: Scotinesthesacuticosta Fairmaire, 1895, by subsequent designation (Gebien 1937a: 741). Status: valid genus in Pimeliinae: Asidini.

Scotinus W. Kirby, 1819a: 415 [M]. Type species: Scotinuscrenicollis W. Kirby, 1819, by monotypy. Status: valid genus in Pimeliinae: Asidini.

Scotobaenus J.L. LeConte, 1859: 87 [M]. Type species: Scotobaenusparallelus J.L. LeConte, 1859, by monotypy. Status: valid genus in Tenebrioninae: Centronopini.

Scotobates Rye, 1877: 341 [M]. Type species: Helopscalcaratus Fabricius, 1798, by subsequent designation (R. Lucas 1920: 587). Status: junior synonym of Menechides Motschulsky, 1872 in Tenebrioninae: Centronopini. Synonymy: R. Lucas (1920: 587). Note: see Spilman (1962b: 2–3) regarding availability of this genus name; Scotobates was used earlier by Horn (1876a: 151) without a description, a definition, or an indication (ICZN 1999, Article 12.1) and is therefore unavailable from that date.

Scotobiopsis Brèthes, 1910: 207 [F]. Type species: Scotobiopsisbreyeri Brèthes, 1910, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina.

Scotobius Germar, 1823: 135 [M]. Type species: Scotobiuspilularius Germar, 1823, by subsequent designation (Lacordaire 1830a: 282). Status: valid genus in Tenebrioninae: Scotobiini.

Scotochares Boheman, 1858: 95 [M]. Type species: Scotocharesinsularis Boheman, 1858, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Scotoderus Perroud & Montrouzier, 1865: 114 [M]. Type species: Scotoderuscancellatus Perroud & Montrouzier, 1865 (= Tenebriocancellatus Montrouzier, 1860), by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Scutopiloxys Pic, 1924b: 13 [M]. Type species: Scutopiloxysperroti Pic, 1924, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Scymena Pascoe, 1866a: 455 [F]. Type species: Scymenavariabilis Pascoe, 1866, by original designation. Status: valid genus in Blaptinae: Opatrini: Heterotarsina.

Scythis Schaum in Kraatz, 1865: 80, 102 [F]. Type species: Tentyriamacrocephala Tauscher, 1812, by subsequent designation (Gebien 1937a: 622). Status: valid genus in Pimeliinae: Tentyriini.

Scythodonta Reitter, 1897a: 300 [F]. Type species: Scythishumeridens Reitter, 1887, by monotypy. Status: junior synonym of Scytosoma Reitter, 1895 in Pimeliinae: Tentyriini. Synonymy: Skopin (1979: 171).

Scytosoma Reitter, 1895: 281 [N]. Type species: Scytosomaarcibasis Reitter, 1895 (= Tentyriapygmaea Gebler, 1832), by subsequent designation (Gebien 1937a: 622). Status: valid genus in Pimeliinae: Tentyriini. Note: redescribed as new by Reitter (1897a: 299).

Sebastianus Iwan, 1996: 385, 416 [M]. Type species: Melanocratusmajor Fairmaire, 1899, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Sechuranus Flores & Giraldo-Mendoza in Giraldo-Mendoza and Flores, 2019: 93 [M]. Type species: Prohylithusbarbatus Kaszab, 1964, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Seidlitzellus Reitter, 1920b: 221 [M]. Type species [automatic]: Anisocerustristis Faldermann, 1837, by monotypy. Status: junior synonym of Ceratanisus Gemminger, 1870 in Pimeliinae: Ceratanisini. Note: unnecessary replacement name for Anisocerus Faldermann, 1837, a senior synonym of Ceratanisus Gemminger, 1870.

Seirotrana Pascoe, 1866a: 483 [F]. Type species: Adeliumcatenulatum Boisduval, 1835, by original designation. Status: valid genus in Lagriinae: Adeliini.

Selenomma Dejean, 1836: 203 [F]. Type species [automatic]: Ammophorusperuvianus Guérin-Méneville, 1831, by monotypy. Status: junior synonym of Ammophorus Guérin-Méneville, 1831 in Tenebrioninae: Scotobiini. Note: unnecessary replacement name for Ammophorus Guérin-Méneville, 1831.

Selenepistoma Dejean, 1834: 190 [N]. Type species: Opatrumacutum Wiedemann, 1823, by subsequent designation (Bousquet and Bouchard 2013a: 50). Status: valid genus in Blaptinae: Dendarini: Melambiina.

Selinopodus Koch, 1956a: 79 [M]. Type species: Selinopodusgiganteus Koch, 1956, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Selinus Mulsant & Rey, 1853b: 70, 97 [M]. Type species: Opatrumplanum Fabricius, 1792, by subsequent designation (Cazurro Ruiz 1896: 963). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Sellio Mulsant & Rey, 1859a: 70, 105 [M]. Type species: Blaps tibidens Quensel, 1806, by subsequent designation (Gebien 1938a: 407). Status: junior synonym of Diastolinus Mulsant & Rey, 1859 in Blaptinae: Opatrini: Blapstinina. Synonymy: Ivie and Hart (2016: 468).

Semenovonymus Bogatchev, 1946: 391 [M]. Type species: Semenovonymustenuis Bogatchev, 1946, by monotypy. Status: junior synonym of Scythis Kraatz, 1865 in Pimeliinae: Tentyriini. Synonymy: G.S. Medvedev (1990: 101).

Semieutochia Kaszab, 1980a: 187 [F]. Type species: Semieutochiaooidea Kaszab, 1980, by original designation. Status: valid genus in Tenebrioninae: Ulomini. Note: Semieutochia was used earlier by Kaszab (1979a: 91) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Seorsophloeus Bremer, 1998: 10, 12 [M]. Type species: Corticeusbirmanicus Blair, 1921, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Seorsoplonyx Bremer, 2010: 144, 156 [M]. Type species: Seorsoplonyxantennatus Bremer, 2010, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Sepedonastes Gistel, 1856: 382 [M]. Type species: Tenebriobimaculatus Herbst, 1799 (= Dytiscusbimaculatus Linnaeus, 1767), by subsequent designation (Bouchard et al. 2005: 501). Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Synonymy: Bouchard et al. (2005: 513).

Sepidiacis Fairmaire, 1884c: cxlvi [F]. Type species: Sepidiaciscompressa Fairmaire, 1884, by subsequent designation (W.F. Kirby 1885b: 83). Status: junior synonym of Sepidiostenus Fairmaire, 1884 in Pimeliinae: Sepidiini: Sepidiina. Synonymy: Gestro (1892: 775). Note: redescribed as new by Fairmaire (1887a: 185).

Sepidiopsis Gestro, 1892: 771 [F]. Type species: Sepidiopsiscornigera Gestro, 1892, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Sepidiostenus Fairmaire, 1884b: lxxv [M]. Type species: Sepidiostenuserinaceus Fairmaire, 1884, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina. Note: redescribed as new, under the spelling Sepidostenus, by Fairmaire (1887a: 184).

Sepidium Fabricius, 1775: 250 [N]. Type species: Sepidium tricuspidatum Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Sepilokus Iwan & Raś, 2020: 776 [M]. Type species: Sepilokustenenbaumi Iwan & Raś, 2020, by original designation. Status: valid genus in Phrenapatinae: Archaeoglenini.

Septentriophosis Penrith, 1982: 167 [F]. Type species: Erodiusplanus Fabricius, 1775, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Seriscius Motschulsky, 1845a: 77 [M]. Type species: Serisciuspubescens Motschulsky, 1845 (= Crypticusrufipes Gebler, 1830), by monotypy. Status: valid subgenus of Crypticus Latreille, 1816 in Diaperinae: Crypticini.

Serrania Garrido, 2003: 50 [F]. Type species: Diaperisviridula Zayas, 1988 (= Platydemavirens Laporte & Brullé, 1831), by original designation. Status: junior synonym of Nesocyrtosoma Marcuzzi, 1976 in Stenochiinae: Cnodalonini. Synonymy: Hopp and Ivie (2009: 13).

Serrichora Koch, 1952b: 15 [F]. Type species: Eurychoracrenata Solier, 1837, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Serridenos Koch, 1956a: 325 [M]. Type species: Zadenossolenopistoma Koch, 1956, by original designation. Status: valid subgenus of Selenepistoma Dejean, 1834 in Blaptinae: Dendarini: Melambiina.

Setenis Motschulsky, 1872: 24 [M]. Type species: Tenebriovalgus Wiedemann, 1823, by original designation. Status: junior synonym of Promethis Pascoe, 1869 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1988: 70).

Seydelicistela Pic, 1954: 260 [F]. Type species: Seydelicistelarubrithorax Pic, 1954, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Sicharbas Champion, 1884: 67 [M]. Type species: Sicharbaslobatus Champion, 1884, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Sicinus Champion, 1886: 146 [M]. Type species: Sicinusguatemalensis Champion, 1886, by subsequent designation (Löbl et al. 2008a: 43). Status: junior synonym of Gnatocerus Thunberg, 1814 in Diaperinae: Diaperini: Adelinina. Synonymy: Leng (1920: 233).

Silvestriellum Koch, 1956a: 362 [N]. Type species: Silvestriellumalatum Koch, 1956, by original designation. Status: valid genus in Blaptinae: Dendarini: Melambiina.

Simalura Gebien, 1914d: 71 [F]. Type species: Simalurajacobsoni Gebien, 1914, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Simarus Borchmann, 1909a: 713 [M]. Type species [automatic]: Ismarusgodeffroyi Haag-Rutenberg, 1878, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: replacement name for Ismarus Haag-Rutenberg, 1878.

Similepitragus Freude, 1967: 148, 167 [M]. Type species: Epitragussimilis Steinheil, 1872, by original designation. Status: valid subgenus of Epitragus Latreille, 1802 in Pimeliinae: Epitragini.

Singapura Gebien, 1925f: 325 [F]. Type species: Singapuraquadrihamata Gebien, 1925, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Sinocatomus Nabozhenko, 2006: 852 [M]. Type species: Catomussolitarius Nabozhenko, 2006, by original designation. Status: valid subgenus of Catomus Allard, 1876 in Tenebrioninae: Helopini: Helopina.

†Sinocistela Zhang, 1989: 145 [F]. Type species: Sinocistelasiphla Zhang, 1989, by original designation. Status: valid genus in Alleculinae: Cteniopodini. Note: described from Lower Miocene deposits (China).

Sinoecia Chatanay, 1914a: 220 [F]. Type species: Sinoeciapuncticollis Chatanay, 1914, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Sinomenimus G.S. Medvedev, 2007b: 675 [M]. Type species: Menimuskabaki G.S. Medvedev, 2007, by original designation. Status: valid subgenus of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina.

Sinopium Pascoe, 1866a: 487 [N]. Type species: Strongyliumvariabile Walker, 1858, by original designation. Status: junior synonym of Camarimena Motschulsky, 1863 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1919: 155).

Sinorus Mulsant & Revelière, 1861: 153 [M]. Type species: Sinorusciliaris Mulsant & Revelière, 1861 (= Opatrumcolliardi Fairmaire, 1860), by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Sintagona G.S. Medvedev, 1998b: 585 [F]. Type species: Sintagonamiranda G.S. Medvedev, 1998, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Sipirocus Fairmaire, 1896c: 103 [M]. Type species: Sipirocusritsemae Fairmaire, 1896, by monotypy. Status: junior synonym of Aediatorix Bates, 1868 in Lagriinae: Pycnocerini. Synonymy: Gebien (1905: 258).

Sipolisia Fairmaire, 1889c: xlix [F]. Type species: Sipolisiaserricornis Fairmaire, 1889, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Sitophagus Mulsant, 1854: 264 [M]. Type species: Sitophagussolieri Mulsant, 1854 (= Ulomahololeptoides Laporte, 1840), by monotypy. Status: valid genus in Diaperinae: Diaperini: Adelinina.

Sloanea Carter, 1916: 209 [F]. Type species: Sloaneacostata Carter, 1916, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Asphalina.

Smiliophanus Koch, 1950a: 66 [M]. Type species [automatic]: Smiliotussteiroides Haag-Rutenberg, 1875, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Smiliotus Haag-Rutenberg, 1875.

Smiliotus Haag-Rutenberg, 1875b: 4, 52 [M]. Type species: Smiliotussteiroides Haag-Rutenberg, 1875, by monotypy. Status: senior synonym of Smiliophanus Koch, 1950 in Pimeliinae: Adelostomini. Note: junior homonym of Smiliotus Loew, 1857 [Diptera].

Sobas Pascoe, 1863a: 45 [F]. Type species [automatic]: Trigonotarsusaustralis Hope, 1843, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: replacement name for Trigonotarsus Hope, 1843. Note: moved from the subtribe Ammobiina to Opatrina by Kamiński et al. (2021b: 151).

Socotralia Novák, 2007: 321 [F]. Type species: Socotraliamajor Novák, 2007, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Socotraphanes Nabozhenko in Nabozhenko and Purchart, 2019: 150 [M]. Type species: Socotraphaneskrali Nabozhenko, 2019, by original designation. Status: valid genus in Tenebrioninae: Helopini: Helopina.

Socotropatrum Koch, 1970: 109 [N]. Type species: Opatrumcostiferum C.O. Waterhouse, 1881, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Soemias Champion, 1884: 4 [F]. Type species: Soemiasminuta Champion, 1884, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Solenomerus Fåhraeus, 1870: 306 [M]. Type species: Solenomeruslongipes Fåhraeus, 1870, by monotypy. Status: junior synonym of Micrantereus Solier, 1848 in Blaptinae: Pedinini: Helopinina. Synonymy: Fairmaire (1897f: 131).

Solskyia Solsky, 1881: 48 [F]. Type species: Solskyiaperegrina Solsky, 1881, by monotypy. Status: valid genus in Pimeliinae: Akidini.

Somaladesmia Koch, 1944b: 147 [F]. Type species: Adesmiaconsimilis Gahan, 1896, by monotypy. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini.

Somalammodes Koch, 1943a: 500, 510 [M]. Type species: Somalammodesdelaruei Koch, 1943, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Somalarabes Koch, 1953f: 155 [M]. Type species: Psammodesgracilentus Fairmaire, 1882, by original designation. Status: valid subgenus of Psammophanes Lesne, 1922 in Pimeliinae: Sepidiini: Molurina.

Somaticus Hope, 1841: 117 [M]. Type species: Sepidiumrugosum Fabricius, 1781, by original designation. Status: valid genus and subgenus in Pimeliinae: Sepidiini: Trachynotina. Note: Somaticus, introduced by Lacordaire (1859a: 197), is an incorrect subsequent spelling of the original name Somaticum in prevailing usage and attributed to the publication of the original spelling; we follow Kamiński et al. (2019b: 86) and treat Somaticus as the correct original spelling (ICZN 1999, Article 33.3.1); the First Reviser (Somaticus Hope, 1841 versus Tracheloeum Hope, 1841) is Koch (1955a: 53).

Somocoelia Heyden & Kraatz, 1882: 331 [F]. Type species: Somocoeliapinguis Heyden & Kraatz, 1882, by monotypy. Status: valid genus in Blaptinae: Platyscelidini.

Somocoeloplatys Skopin, 1968a: 82 [M]. Type species: Platynoscelisboroldaica Skopin, 1965, by original designation. Status: valid genus in Blaptinae: Platyscelidini.

Sophrobates Fairmaire, 1889c: xxxvi [M]. Type species: Sophrobatesarcadii Fairmaire, 1889, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Sora Walker, 1859: 259 [F]. Type species: Soramarginata Walker, 1859, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Soradeus Rafinesque, 1815: 114 [M]. Type species [automatic]: Heleaperforata Latreille, 1816, by subsequent monotypy (Latreille 1816b: 261). Status: junior synonym of Helea Latreille, 1804 in Tenebrioninae: Heleini: Heleina. Note: unnecessary replacement name for Helea Latreille, 1804 (as “Heleus Latr.”).

Spathulipezus Gebien, 1921a: 398, 458 [M]. Type species: Spathulipezusmiritaris Gebien, 1921, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Spectrocnera Kwieton, 1981: 402 [F]. Type species: Spectrocneraanguliceps Kwieton, 1981, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Spelaebiosis Bousquet & Bouchard in Bousquet et al., 2018: 223 [F]. Type species [automatic]: Orghidaniatorrei Ardoin, 1977, by monotypy. Status: valid genus in Tenebrioninae: Triboliini. Note: replacement name for Ardoinia Özdikmen, 2005.

Sphaeriontis Casey, 1908: 56, 75 [F]. Type species: Eusattusmuricatus J.L. LeConte, 1851, by original designation. Status: junior synonym of Eusattus J.L. LeConte, 1851 in Pimeliinae: Coniontini. Synonymy: La Rivers (1949: 180).

Sphaerocaulus Fairmaire, 1869b: 235 [M]. Type species: Sphaerocaulusgraniger Fairmaire, 1869, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Sphaerognathium Dajoz, 1975a: 112 [N]. Type species: Sphaerognathiumglobosum Dajoz, 1975, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Sphaeromatris Fairmaire, 1899e: 535 [F]. Type species: Sphaeromatrisaurovittata Fairmaire, 1899, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Sphaerostibes Koch, 1963: 64 [M]. Type species: Sphaerostibessabulicola Koch, 1963, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 13.4).

Sphaerotidius Kaszab, 1941a: 3, 38 [M]. Type species: Sphaerotidiusduplicatus Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Spaerotidius, used by Kaszab (1941a: 39, 40).

Sphaerotus W. Kirby, 1819a: 416 [M]. Type species: Sphaerotuscurvipes W. Kirby, 1819, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Sphargeris Pascoe, 1860b: 122 [M]. Type species: Sphargerisphysodes Pascoe, 1860, by monotypy. Status: valid genus in Lagriinae: Chaerodini.

Sphenaria Ménétriés, 1849: 240 [F]. Type species: Sphenariaelongata Ménétriés, 1849, by subsequent designation (Gebien 1937a: 576). Status: valid genus in Pimeliinae: Tentyriini.

Sphenariopsis Kraatz, 1865: 80, 175 [F]. Type species: Sphenariopsistristis Kraatz, 1865, by monotypy. Status: valid subgenus of Rhytinota Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Spheneuphloeus Kaszab, 1941a: 5, 35 [M]. Type species: Osdarametallica Pic, 1931, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Spheniscus W. Kirby, 1819a: 421 [M]. Type species: Spheniscuserotyloides W. Kirby, 1819, by monotypy. Status: senior synonym of Cuphotes Champion, 1887 in Stenochiinae: Stenochiini. Note: junior homonym of Spheniscus Moehring, 1758 [Aves].

Sphenogenius Solier, 1848: 154, 351 [M]. Type species: Sphenogeniusclathratus Solier, 1848 (= Nyctozoilusobesus Guérin-Méneville, 1831), by original designation. Status: junior synonym of Nyctozoilus Guérin-Méneville, 1831 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Lacordaire (1859a: 350).

Sphenolampidius Kaszab, 1941a: 4, 40 [M]. Type species: Sphenolampidiushemisphaericus Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Sphenosdara Kaszab, 1941a: 2, 28 [F]. Type species: Sphenosdarasachtlebeni Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Sphenosoma Dejean, 1834: 212 [N]. Type species [automatic]: Acropteronrufipes Perty, 1832, by subsequent designation (Hope 1841: 133). Status: junior synonym of Acropteryx Gistel, 1831 in Tenebrioninae: Acropteronini. Note: unnecessary replacement name for Acropteron Perty, 1832.

Sphenothorax Gebien, 1906: 232 [M]. Type species: Tenebrionitidulus Fabricius, 1801, by monotypy. Status: junior synonym of Zophophilus Fairmaire, 1881 in Stenochiinae: Cnodalonini. Synonymy: Carter (1930: 547).

Sphinctoderus Fairmaire, 1903e: 301 [M]. Type species: Sphinctoderusstrangulatus Fairmaire, 1903, by monotypy. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Sphingocorse Gebien, 1921b: 110, 111 [F]. Type species: Sphingocorseangulicollis Gebien, 1921, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Sphragidophorus Champion, 1889: 61 [M]. Type species: Statiracyanipennis Mäklin, 1863, by subsequent designation (R. Lucas 1920: 603). Status: valid genus in Lagriinae: Lagriini: Statirina.

Spiloscapha Bates, 1873a: 202 [F]. Type species: Spiloscaphacrassicornis Bates, 1873 (= Platydemathallioides Pascoe, 1869), by monotypy. Status: valid genus in Diaperinae: Scaphidemini.

Spinadaenus Pic, 1921d: 18 [M]. Type species: Spinadaenussingularis Pic, 1921, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Spinamarygmus Pic, 1915d: 7 [M]. Type species: Spinamarygmusindicus Pic, 1915, by monotypy. Status: valid subgenus of Plesiophthalmus Motschulsky, 1857 in Tenebrioninae: Amarygmini.

Spinanemia Löbl, Bouchard, Merkl & Bousquet, 2020: 2 [F]. Type species: Anemiacornuta Pic, 1898, by original designation. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini. Note: subgenus first proposed by Ardoin (1971: 362, 378) without type species originally designated.

Spinecula Novák, 2019c: 437 [F]. Type species: Spineculahouaphanica Novák, 2019, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Spinepicalla Pic, 1921d: 21 [F]. Type species: Spinepicallaarmata Pic, 1921, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Spinoderosphaerus Pic, 1922d: 26 [M]. Type species: Spinoderosphaerusbrevicornis Pic, 1922, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: the First Reviser (Spinoderosphaerus Pic, 1922 versus Spinogauromaia Pic, 1922) is Kaszab (1983a: 132).

Spinodietysus Pic, 1927b: 21 [M]. Type species: Cyriogetonconvexipennis Pic, 1927, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Spinogauromaia Pic, 1922a: 23 [F]. Type species: Spinogauromaiarufescens Pic, 1922 (= Spinoderosphaerusbrevicornis Pic, 1922), by monotypy. Status: junior synonym of Spinoderosphaerus Pic, 1922 in Stenochiinae: Cnodalonini. Synonymy: Kaszab (1983a: 132).

Spinolagriella Pic, 1955: 183 [F]. Type species: Spinolagriellaminutissima Pic, 1955, by original designation. Status: valid genus in Lagriinae: Lupropini.

Spinolyprops Pic, 1917d: 12 [M]. Type species: Spinolypropsrufithorax Pic, 1917, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Spinoodescelis Kaszab, 1940b: 938, 966 [F]. Type species: Platyscelissomocoeloides Seidlitz, 1893, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Spinophrynus Koch, 1951: 90 [M]. Type species: Phrynocolusspinipennis Gebien, 1910, by original designation. Status: valid subgenus of Phrynocolus Lacordaire, 1859 in Pimeliinae: Sepidiini: Molurina.

Spinorhacus Kaszab, 1969a: 262 [M]. Type species: Spinorhacusbaloghi Kaszab, 1969, by original designation. Status: junior synonym of Spinolagriella Pic, 1955 in Lagriinae: Lupropini. Synonymy: Kaszab (1976: 453).

Spinosdara Bouchard & Bousquet, new subgenus [F]. Type species: Osdarabiroi Kaszab, 1939, by present designation. Status: valid subgenus of Osdara Walker, 1858 in Stenochiinae: Cnodalonini. Note: Kaszab (1941a: 33) introduced the new subgenus name Spinosdara for three nominal species, but unfortunately did not designate a type species; the subgenusSpinosdara, which has been treated as valid since 1941, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Osdarabiroi Kaszab, 1939 as type species and referring to Kaszab (1941a: 33) for the character states that characterise and differentiate Spinosdara.

Spinostatira Pic, 1918b: 22 [F]. Type species: Statiraspinipes Pic, 1918, by subsequent designation (Borchmann 1936: 247). Status: valid subgenus of Statira Lepeletier & Audinet-Serville, 1828 in Lagriinae: Lagriini: Statirina.

Splenoodescelis Egorov, 2004: 593 [F]. Type species: Platyscelisturkestanica Seidlitz, 1893, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Splichalia Reitter, 1913: 664 [F]. Type species: Splichaliatigrinella Reitter, 1913, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Spongesmia Bouchard & Bousquet, new subgenus [F]. Type species: Adesmianassata Erichson, 1843, by present designation. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: Koch (1944b: 149) introduced the new subgenus name Spongesmia for four nominal species, but unfortunately did not designate a type species; the subgenusSpongesmia, which has been treated as valid since 1944, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Adesmianassata Erichson, 1843 as type species and referring to Koch (1944b: 149) for the character states that characterise and differentiate Spongesmia.

Spongesmima Koch, 1944b: 157 [F]. Type species: Adesmiascrobipennis Haag-Rutenberg, 1875, by monotypy. Status: junior synonym of Renatiella Koch, 1944 in Pimeliinae: Adesmiini. Synonymy: Penrith (1979: 27).

Spyrathus Kraatz, 1865: 6, 9 [M]. Type species: Spyrathusindicus Kraatz, 1865, by monotypy. Status: valid genus in Pimeliinae: Erodiini.

Srilanka Kaszab, 1980b: 319 [F]. Type species: Srilankamirabilis Kaszab, 1980, by original designation. Status: valid genus in Stenochiinae: Cnodalonini. Note: Srilanka was used earlier by Kaszab (1979a: 107) without a description, a definition, or a bibliographic reference to such a published statement (ICZN 1999, Article 13.1) and is therefore unavailable from that date.

Staius Fairmaire, 1896b: 359 [M]. Type species: Staiusmiricornis Fairmaire, 1896, by monotypy. Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Stalagmoptera Solsky, 1876: 286 [F]. Type species: Stalagmopteratuberculatocostata Solsky, 1876, by subsequent designation (Gebien 1937a: 821). Status: valid genus in Pimeliinae: Pimeliini.

Statira Lepeletier & Audinet-Serville, 1828: 479 [F]. Type species: Statiraagroides Lepeletier & Audinet-Serville, 1828, by subsequent designation (Blanchard 1844: pl. 53bis). Status: valid genus and subgenus in Lagriinae: Lagriini: Statirina.

Statiropsis Borchmann, 1912b: 389 [F]. Type species: Statiropsisaenea Borchmann, 1912, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Stegastopsis Kraatz, 1865: 80, 176 [F]. Type species: Stegastopsisbabylonica Kraatz, 1865, by monotypy. Status: valid genus and subgenus in Pimeliinae: Tentyriini.

Steira Westwood, 1837: pl. 176 [F]. Type species: Steiracostata Westwood, 1837, by monotypy. Status: senior synonym of Stips Koch, 1950 in Pimeliinae: Adelostomini. Note: junior homonym of Steira Eschscholtz, 1825 [Mollusca].

Stemmatoderus Agassiz, 1846b: 351 [M]. Type species [automatic]: Stemmoderussingularis Spinola, 1842, by monotypy. Status: junior synonym of Stemmoderus Spinola, 1842 in Tenebrioninae: Amarygmini. Note: unjustified emendation of Stemmoderus Spinola, 1842, not in prevailing usage.

Stemmoderus Spinola, 1842: pl. 91 (p. 1) [M]. Type species: Stemmoderussingularis Spinola, 1842, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Stenadelium Watt, 1992: 32 [N]. Type species: Stenadeliumstriatum Watt, 1992, by original designation. Status: valid genus in Lagriinae: Adeliini.

Stene Stephens, 1829: 19 [F]. Type species: fixed herein (ICZN 1999, Article 70.3) as Colydiumcastaneum Herbst, 1797, misidentified as Tenebrioferrugineus Fabricius, 1781 in the original designation by monotypy in Stephens (1829). Status: junior synonym of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini. Synonymy: Shuckard (1840: vii). Note: the type species “Tenebrioferrugineus Fabricius” was first established by monotypy; as noted by C.O. Waterhouse (1896: 230) and Blair (1913: 223) the Tenebrioferrugineus Fabricius of authors, including Stephens (1829: 19, as “ferruginea, Oliv.”), was misidentified; Blair (1913: 223) noted that the species the authors referred to is in fact Colydiumcastaneum Herbst, 1797; we follow currently accepted concepts (e.g., Bousquet et al. 2018: 224) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Tenebrioferrugineus Fabricius, 1781 is a valid species in the genus Tribolioides Blair, 1913 [Coleoptera: Cucujidae].

Steneleodes Blaisdell, 1909: 409 [M]. Type species: Eleodeslongicollis J.L. LeConte, 1851, by subsequent designation (Bousquet et al. 2018: 166). Status: valid subgenus in Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Stenerophlina Reitter, 1906b: 130 [F]. Type species: Omophlinahauseri Reitter, 1894, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Stenerula Fairmaire, 1875: 42 [F]. Type species: Stenerulasubopaca Fairmaire, 1875, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Steneryx Reitter, 1890b: 256 [M]. Type species: Cisteladejeanii Faldermann, 1836, by monotypy. Status: valid genus in Alleculinae: Cteniopodini.

Stenethmus Gebien, 1937b: 41 [M]. Type species: Psammodestentyriniformis Hesse, 1935, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Steneucyrtus Fairmaire, 1896a: 31 [M]. Type species: Steneucyrtuspexicollis Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stenholma Solier, 1835b: 253, 412 [F]. Type species: Stenholmatentyrioides Solier, 1835, by monotypy. Status: junior synonym of Melaphorus Guérin-Méneville, 1834 in Pimeliinae: Evaniosomini. Synonymy: Solier (1835b: 412).

Stenillus Blair, 1927: 245 [M]. Type species: Stenillusmonticola Blair, 1927, by monotypy. Status: valid subgenus of Pseudethas Fairmaire, 1896 in Pimeliinae: Stenosini: Dichillina.

Stenocara Solier, 1835b: 512, 553 [N]. Type species: Pimelialongipes Fabricius, 1775, by subsequent designation (Desmarest 1860: 141). Status: valid genus and subgenus in Pimeliinae: Adesmiini.

Stenocephalus Agassiz, 1846b: 71, 351 [M]. Type species [automatic]: Cephalostenusdejeanii Solier, 1838 (= Scauruselegans Brullé, 1832), by subsequent designation (Hope 1841: 115). Status: junior synonym of Cephalostenus Solier, 1838 in Tenebrioninae: Scaurini. Note: unjustified emendation of Cephalostenus Solier, 1838; junior homonym of Stenocephalus Tschudi, 1838 [Amphibia].

Stenocera Agassiz, 1846b: 74, 351 [F]. Type species [automatic]: Cerostenadeplanata Solier, 1836, by present designation. Status: junior synonym of Psectrascelis Solier, 1836 in Pimeliinae: Nycteliini. Note: unjustified emendation of Cerostena Solier, 1836, not in prevailing usage; junior homonym of Stenocera Brullé, 1834 [Coleoptera: Carabidae].

Stenochia W. Kirby, 1819a: 423 [F]. Type species: Stenochiarufipes W. Kirby, 1819, by subsequent designation (Hope 1841: 133). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Latreille (1829b: 683).

Stenochidus J.L. LeConte, 1862: 244 [M]. Type species: Stenochiagracilis J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 608). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Stenochinus Motschulsky, 1860a: 102 [M]. Type species: Stenochinusreticulatus Motschulsky, 1860, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stenodesia Reitter, 1916a: 4 [F]. Type species: Stenocaraglobulum Haag-Rutenberg, 1875, by monotypy. Status: valid genus in Pimeliinae: Adesmiini.

Stenogena Fairmaire, 1895a: 33 [F]. Type species: Stenogenamadecassa Fairmaire, 1895, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: placed in the subfamily Alleculinae by Chatanay (1915a: 526).

Stenogenomorpha Pic, 1919b: 5 [F]. Type species: Stenogenomorphaimpressa Pic, 1919, by monotypy. Status: valid genus in Alleculinae: incertae sedis.

Stenogonopus Gebien, 1938b: 91 [M]. Type species: Tenebrioplumosus Thunberg, 1787, by original designation. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Stenohelops Reitter, 1922a: 22 [M]. Type species: Isopedusplicatulus Kraatz, 1880, by subsequent designation (Gebien 1943: 423). Status: valid genus and subgenus in Tenebrioninae: Helopini: Helopina. Note: the First Revisers (Stenohelops Reitter, 1922 versus Gunarellus Reitter, 1922) are Nabozhenko et al. (2020b: 297).

Stenolagria Merkl, 1987: 124, 157 [F]. Type species: Stenolagriamatthewsi Merkl, 1987, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Stenolamus Gebien, 1920: 107 [M]. Type species: Stenolamussulciceps Gebien, 1920, by subsequent designation (Gebien 1938a: 397). Status: valid genus in Blaptinae: incertae sedis. Note: this taxon is placed in “Blaptinae incertae sedis” based on comments in Kamiński et al. (2021b: 150).

†Stenolassus Nabozhenko, Chigray & Bukejs, 2020a: 519 [M]. Type species: Nalassusklebsi Nabozhenko, Perkovsky & Chernei, 2016, by original designation. Status: valid subgenus of Stenohelops Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Note: described from Eocene Baltic amber.

Stenomacidius Seidlitz, 1895: 791 [M]. Type species: Hedyphanesacutangulus Seidlitz, 1895, by subsequent designation (G.S. Medvedev 1990: 242). Status: junior synonym of Cylindrinotus Faldermann, 1837 in Tenebrioninae: Helopini: Cylindrinotina. Synonymy: Nabozhenko (2006: 799).

Stenomaleis Español, 1957b: 21, 34 [M]. Type species: Stenohelopsardoini Español, 1957 (= Helopsprotensulus Seidlitz, 1896), by original designation. Status: junior synonym of Helopelius Reitter, 1922 in Tenebrioninae: Helopini: Helopina. Synonymy: Nabozhenko et al. (2020b: 293).

Stenomax Allard, 1876a: 4 [M]. Type species: Tenebriolanipes Linnaeus, 1771 (= Tenebrioaeneus Scopoli, 1763), by subsequent designation (Español and Comas 1989: 166, 170). Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina.

Stenomorpha Solier, 1836: 407, 487 [F]. Type species: Stenomorphablapsoides Solier, 1836, by subsequent designation (Desmarest 1860: 150). Status: valid genus and subgenus in Pimeliinae: Asidini.

Stenopalorus Blair, 1930: 135 [M]. Type species: Palorushypophloeoides Blair, 1930, by monotypy. Status: junior synonym of Palorus Mulsant, 1854 in Tenebrioninae: Palorini. Synonymy: Halstead (1967a: 114, 115).

Stenophanes Solsky, 1876: 294 [M]. Type species: Hedyphanesmesostenus Solsky, 1871, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Stenophloeus Blair, 1921: 1 [M]. Type species: Hypophlaeusfilum Fairmaire, 1893, by subsequent designation (Löbl et al. 2008b: 312). Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Stenopsis Rafinesque, 1815: 113 [F]. Type species [automatic]: Pimeliareflexa Fabricius, 1775, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Akis Herbst, 1799 in Pimeliinae: Akidini. Note: unnecessary replacement name for Akis Herbst, 1799.

Stenoscapha Bates, 1873b: 237 [F]. Type species: Stenoscaphatibialis Bates, 1873, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Stenoscapha Fairmaire, 1885b: 234 [F]. Type species: Stenoscaphaspissicornis Fairmaire, 1885, by monotypy. Status: senior synonym of Brachypophlaeus Fairmaire, 1897 in Tenebrioninae: Ulomini. Note: junior homonym of Stenoscapha Bates, 1873 [Coleoptera: Tenebrionidae: Diaperinae: Diaperini: Diaperina].

Stenosethas Kaszab, 1975a: 11 [M]. Type species: Stenosethascarinipennis Kaszab, 1975, by original designation. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Stenosida Solier, 1835b: 253, 281 [F]. Type species: Stenosidatenuicollis Solier, 1835 (= Tageniastriatopunctata Wiedemann, 1821), by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Stenosides Solier, 1836: 406, 484 [M]. Type species: Stenosidesgraciliformis Solier, 1836, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Stenosidops Koch, 1940b: 733 [M]. Type species: Tageniasabulosa Guérin-Méneville, 1849, by monotypy. Status: valid subgenus of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina.

Stenosis Herbst, 1799: 160 [F]. Type species: fixed herein (ICZN 1999, Article 70.3) as Tageniaintermedia Solier, 1838, misidentified as Pimeliaangustata Fabricius, 1775 in the subsequent designation in Lacordaire (1859a). Status: valid genus and subgenus in Pimeliinae: Stenosini: Stenosina. Note: the type species “Pimeliaangustata Fabricius” (as angustata Herbst) was selected by Lacordaire (1859a: 102) as the type species; Baudi de Selve (1875: 67) first noted that Pimeliaangustata Fabricius of Herbst (1799) was misidentified and corresponded to Tageniaintermedia Solier, 1838; we follow currently accepted concepts (e.g., Löbl et al. 2008b: 179) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Pimeliaangustata Fabricius, 1775 is a valid species in the genus Mesostena Eschscholtz, 1831 [Coleoptera: Tenebrionidae].

Stenothesilea Kulzer, 1951a: 131, 134 [F]. Type species: Stenothesileacylindriformis Kulzer, 1951, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stenotrichus J.L. LeConte, 1862: 239 [M]. Type species: Amphidorarufipes J.L. LeConte, 1851, by original designation. Status: junior synonym of Helops Fabricius, 1775 in Tenebrioninae: Helopini: Helopina. Synonymy: Aalbu et al. (2002: 496).

Steptochora Koch, 1952b: 10 [F]. Type species: Eurychorasetosula Fairmaire, 1887, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Sterces Champion, 1891: 640 [M]. Type species: Stercesviolaceipennis Champion, 1891, by present designation. Status: junior synonym of Alcyonotus Pascoe, 1882 in Stenochiinae: Cnodalonini. Synonymy: Champion (1894b: lxiii).

Steriphanides Casey, 1907: 515 [M]. Type species: Emmenastusstolidus Champion, 1892, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Steriphanus Casey, 1907: 289 [M]. Type species: Emmenastusconicicollis Casey, 1890, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Sternocnera Skopin, 1964: 394 [F]. Type species: Sternocneralindti Skopin, 1964, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Sternodes Fischer von Waldheim, 1837: 10 [M]. Type species: Sternodeskarelini Fischer von Waldheim, 1837 (= Tenebriocaspicus Pallas, 1781), by monotypy. Status: valid genus in Pimeliinae: Pimeliini.

Sternomaia Kulzer, 1952: 731 [F]. Type species: Sternomaiacoeruleovirens Kulzer, 1952, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Sternoplax Frivaldszky, 1890: 207 [F]. Type species: Trigonoscelisszechenyii Frivaldszky, 1890, by monotypy. Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Sternotrigon Skopin, 1973: 109, 110 [M]. Type species: Trigonoscelissetosa Bates, 1879, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Stethasida Casey, 1912: 78, 203 [F]. Type species: Pelecyphorusmuricatulus J.L. LeConte, 1851, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Stethotrypes Gebien, 1914c: 26 [M]. Type species: Stethotrypesbicornutus Gebien, 1914, by subsequent designation (Gebien 1940: 431). Status: valid genus in Diaperinae: Leiochrinini.

Sthenoboea Champion, 1885: 112 [F]. Type species: Sthenoboeaapicalis Champion, 1885, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stibia Horn, 1870: 258, 260 [F]. Type species: Stibiapuncticollis Horn, 1870, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Stictodera Casey, 1907: 289, 352 [F]. Type species: Emmenastuspinguis J.L. LeConte, 1866, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Stictodere Gebien, 1928: 101 [F]. Type species: Stictoderesubseriata Gebien, 1928, by subsequent designation (Gebien 1937a: 571). Status: valid genus in Pimeliinae: Epitragini.

Stictoderia Gebien, 1937a: 571 [F]. Type species [automatic]: Stictoderesubseriata Gebien, 1928, by subsequent designation (Gebien 1937a: 571). Status: junior synonym of Stictodere Gebien, 1928 in Pimeliinae: Epitragini. Note: unnecessary replacement name for Stictodere Gebien, 1928, which is not a homonym of Stictodera Casey, 1907 [Coleoptera: Tenebrionidae: Pimeliinae: Edrotini].

Stierlinius Forel, 1893: 167 [M]. Type species [automatic]: Dolichoderusacuminatus Klug, 1833, by monotypy. Status: junior synonym of Macellocerus Solier, 1848 in Tenebrioninae: Toxicini: Nycteropina. Note: replacement name for Dolichoderus Klug, 1833.

Stigmatoma J.L. LeConte, 1862: 244 [F]. Type species: Cistelafraterna Say, 1824, by monotypy. Status: junior synonym of Ernocharis C.G. Thomson, 1859 in Alleculinae: Alleculini: Mycetocharina. Synonymy: Seidlitz (1896: 147).

Stilbocistela Borchmann, 1932a: 319 [F]. Type species: Stilbocistelaluzonica Borchmann, 1932, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: genus described as new again by Borchmann (1932b: 125).

Stips Koch, 1950a: 66 [F]. Type species [automatic]: Steiracostata Westwood, 1837, by monotypy. Status: valid genus in Pimeliinae: Adelostomini. Note: replacement name for Steira Westwood, 1837.

Stipsostoma Koch, 1952b: 34, 105 [N]. Type species: Steirasculpta Gebien, 1920, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Stira Agassiz, 1846b: 350, 353 [F]. Type species [automatic]: Steiracostata Westwood, 1837, by monotypy. Status: senior synonym of Stips Koch, 1950 in Pimeliinae: Adelostomini. Note: unjustified emendation of Steira Westwood, 1837 (and Steira Eschscholtz, 1825 in Mollusca), not in prevailing usage; we act as First Revisers and treat Stira Agassiz, 1846 [Coleoptera] as a junior homonym of Stira Agassiz, 1846 [Mollusca].

Stizopus Erichson, 1843: 245 [M]. Type species: Stizopuslaticollis Erichson, 1843, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Stomion G.R. Waterhouse, 1845a: 27 [N]. Type species: Stomiongalapagoense G.R. Waterhouse, 1845, by subsequent designation (Gebien 1937a: 590). Status: valid genus in Pimeliinae: Edrotini.

Stomium Agassiz, 1846b: 354 [N]. Type species [automatic]: Stomiongalapagoense G.R. Waterhouse, 1845, by subsequent designation (Gebien 1937a: 590). Status: junior synonym of Stomion G.R. Waterhouse, 1845 in Pimeliinae: Edrotini. Note: unjustified emendation of Stomion G.R. Waterhouse, 1845, not in prevailing usage.

Stomylus Fåhraeus, 1870: 300 [M]. Type species: Stomylusbicolor Fåhraeus, 1870, by subsequent designation (R. Lucas 1920: 616). Status: valid genus in Diaperinae: Diaperini: Diaperina.

Stonavus Reitter, 1904: 161 [M]. Type species: Penthicusalaiensis Reitter, 1896, by subsequent designation (Iwan and Löbl 2007: 734). Status: junior synonym of Penthicus Faldermann, 1836 in Blaptinae: Opatrini: Opatrina. Synonymy: Iwan and Löbl (2007: 734).

Storthephora Mäklin, 1875: 658 [F]. Type species: Storthephoradenticollis Mäklin, 1875, by subsequent designation (Bousquet and Bouchard 2014: 26). Status: junior synonym of Paratenetus Spinola, 1845 in Lagriinae: Goniaderini. Synonymy: Champion (1893b: 47).

Storthocnemis Karsch, 1881: 47 [F]. Type species: Storthocnemissteckeri Karsch, 1881, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Stratodemus Gebien, 1921b: 98 [M]. Type species: Stratodemusheraldicus Gebien, 1921, by monotypy. Status: valid genus in Lagriinae: Pycnocerini.

Strepsius Fairmaire, 1896b: 351 [M]. Type species: Strepsiusspretulus Fairmaire, 1896, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Stridigula Koch, 1954a: 60 [F]. Type species: Stridigulaarguta Koch, 1954, by original designation. Status: valid genus in Blaptinae: Platynotini: Eurynotina.

Stridulomus Koch, 1955a: 37 [M]. Type species: Psammodessulcicollis Péringuey, 1885, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Strongyallecula Pic, 1955: 186 [F]. Type species: Strongyalleculabasilewskyi Pic, 1955, by original designation. Status: valid genus in Alleculinae: incertae sedis.

Strongylacanthus Brèthes, 1925: 13 [M]. Type species: Strongylacanthusperuvianus Brèthes, 1925, by monotypy. Status: valid genus in Stenochiinae: Stenochiini.

Strongylagria Pic, 1915b: 4 [F]. Type species: Strongylagriametallica Pic, 1915, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Strongyliastrum Fairmaire, 1894a: 39 [N]. Type species: Strongyliastrumbraetii Fairmaire, 1894 (= Strongyliumrufipenne Kollar & Redtenbacher, 1844), by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Strongylium W. Kirby, 1819a: 417 [N]. Type species: Strongyliumchalconatum W. Kirby, 1819, by monotypy. Status: valid genus and subgenus in Stenochiinae: Stenochiini. Note: nomen protectum (see Bouchard et al. 2005: 501); the First Reviser (Strongylium W. Kirby, 1819 versus Stenochia W. Kirby, 1819) is Latreille (1829b: 683); we recognize two valid subgenera in the diverse genus Strongylium W. Kirby, 1819, i.e., the recently described Afrostrongylium Robiche, 2019 and the nominotypical subgenus; however, much taxonomic research is needed to evaluate the possible validity of the many synonyms of the subgenusStrongylium W. Kirby, 1819.

Strophia Robiche, 2004b: 130 [F]. Type species: Oncosomaertli Gebien, 1910, by original designation. Status: senior synonym of Strophiamixa Robiche, 2005 in Blaptinae: Pedinini: Helopinina. Note: junior homonym of Strophia Meigen, 1832 [Lepidoptera] and Strophia Albers, 1850 [Mollusca].

Strophiamixa Robiche, 2005: 358 [F]. Type species [automatic]: Oncosomaertli Gebien, 1910, by original designation. Status: valid subgenus of Amatodes Dejean, 1834 in Blaptinae: Pedinini: Helopinina. Note: replacement name for Strophia Robiche, 2004.

Stygohelops Leo & Liberto, 2003: 299 [M]. Type species: Probaticuskalavriticus Schawaller, 2001, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Styphacus Fairmaire, 1901a: 71 [M]. Type species: Styphacusdecorsii Fairmaire, 1901, by subsequent designation (R. Lucas 1920: 619). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Styphloeus Kaszab, 1941a: 4, 36 [M]. Type species: Styphloeusindicus Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Styrax Westwood, 1875: 227 [M]. Type species: Styraxtricondyloides Westwood, 1875, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Styrus Bates, 1873e: 348 [M]. Type species: Styruselongatulus Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Cyphaleina.

Suarezius Fairmaire, 1895a: 22 [M]. Type species: Suareziusgibbosulus Fairmaire, 1895, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Subalphasida Escalera, 1928: 136 [F]. Type species: Asidaluctuosa Boisduval, 1835, by original designation. Status: junior synonym of Betasida Reitter, 1917 in Pimeliinae: Asidini. Synonymy: Viñolas and Cartagena (2005: 282).

Subpterocoma Bouchard & Bousquet, new replacement name [F]. Type species [automatic]: Pterocomatuberculata Motschulsky, 1845, by subsequent designation (Skopin 1974b: 159). Status: valid subgenus of Pterocoma Dejean, 1834 in Pimeliinae: Pimeliini. Note: replacement name for Pseudopimelia Motschulsky, 1860.

Sulcipectus Louw, 1979: 99, 109 [N]. Type species: Sulcipectusleve Louw, 1979, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Sulcolagria Pic, 1955: 181 [F]. Type species: Sulcolagriasemiopaca Pic, 1955, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Sulcosis Penrith, 1977: 19, 220 [F]. Type species: Zophosisangolensis Erichson, 1843, by original designation. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Sulpius Fairmaire, 1906: 273 [M]. Type species: Sulpiuspunctostriatus Fairmaire, 1906, by monotypy. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Sulpiusoma Ferrer, 2006c: 81 [N]. Type species: Sulpiusomamedquisti Ferrer, 2006, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: we act as First Revisers and reject the alternative original spelling Sulpiosoma, used by Ferrer (2006c: 79).

Sumbawia Gebien, 1925b: 441 [F]. Type species: Sumbawiatetrops Gebien, 1925, by monotypy. Status: valid genus in Tenebrioninae: Bolitophagini.

Sundon Pic, 1923d: 18 [N]. Type species: Sundonatricorne Pic, 1923, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Syachis Bates, 1879b: 467 [M]. Type species: Syachishimalaicus Bates, 1879, by subsequent designation (Gebien 1937a: 594). Status: valid genus in Pimeliinae: Tentyriini. Note: redescribed as new by Bates (1890: 55).

Sycophantes Kirsch, 1866: 198 [M]. Type species: Sycophantesdentipes Kirsch, 1866, by subsequent designation (Gebien 1928: 203). Status: valid genus in Stenochiinae: Cnodalonini.

Sycophantomorphus Pic, 1924b: 13 [M]. Type species: Sycophantomorphusater Pic, 1924, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Syggona Fåhraeus, 1870: 330 [F]. Type species: Syggonaconcinna Fåhraeus, 1870, by monotypy. Status: junior synonym of Luprops Hope, 1833 in Lagriinae: Lupropini. Synonymy: Ferrer (1995b: 62).

Sylvanoplonyx Bremer, 2010: 156, 157 [M]. Type species: Sylvanoplonyxfemoralis Bremer, 2010, by original designation. Status: valid genus in Tenebrioninae: Amarygmini.

Sympetes Pascoe, 1866a: 464 [M]. Type species: Sympetesmacleayi Pascoe, 1866 (= Emcephalustricostellus White, 1841), by subsequent designation (Gebien 1940: 1075). Status: valid genus in Tenebrioninae: Heleini: Heleina.

Symphochora Koch, 1952b: 12 [F]. Type species: Eurychorahumerifera Gebien, 1937, by original designation. Status: valid genus in Pimeliinae: Adelostomini.

Symphoxycara Koch, 1943a: 577 [N]. Type species: Oxycarabreviusculum Fairmaire, 1892, by monotypy. Status: valid subgenus of Oxycara Solier, 1835 in Pimeliinae: Tentyriini.

Sympiezocnemis Solsky, 1876: 290 [F]. Type species: Sympiezocnemiskessleri Solsky, 1876, by subsequent designation (Löbl et al. 2008b: 164). Status: junior synonym of Pisterotarsa Motschulsky, 1860 in Pimeliinae: Pimeliini. Synonymy: Heyden (1882: 140).

Synallecula Kolbe, 1883: 25 [F]. Type species: Alleculalivida Sahlberg, 1823, by subsequent designation (R. Lucas 1920: 621). Status: valid genus in Alleculinae: Alleculini: Alleculina. Note: raised from a subgenus of Alogista Fåhraeus, 1870 to valid genus by Novák (2017b: 130).

Synatractus W.J. MacLeay, 1887: 312 [M]. Type species: Synatractusvariabilis W.J. MacLeay, 1887, by monotypy. Status: junior synonym of Casnonidea Fairmaire, 1882 in Lagriinae: Lagriini: Statirina. Synonymy: Carter (1920b: 199), Merkl (1986: 188).

Syncolydium Kolbe, 1897a: 110 [N]. Type species: Syncolydiumglabratum Kolbe, 1897, by monotypy. Status: junior synonym of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini. Synonymy: Bremer (1995: 3). Note: genus originally described in Tenebrionoidea: Colydiidae.

Syngona Rye, 1873: 293 [F]. Type species [automatic]: Syggonaconcinna Fåhraeus, 1870, by monotypy. Status: junior synonym of Luprops Hope, 1833 in Lagriinae: Lupropini. Note: unjustified emendation of Syggona Fåhraeus, 1870, not in prevailing usage.

Synhimba Koch, 1952d: 216 [N]. Type species: Psammodescordiformis Haag-Rutenberg, 1871, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Oxurina.

Synquadrideres Iwan, 2003a: 182 [M]. Type species: Synquadrideresnaivashaensis Iwan, 2003, by original designation. Status: junior synonym of Glyptopteryx Gebien, 1910 in Blaptinae: Platynotini: Platynotina. Synonymy: Kamiński (2015a: 91).

Syntyphlus Koch, 1953e: 243 [M]. Type species: Syntyphlussubterraneus Koch, 1953, by original designation. Status: valid genus in Blaptinae: Opatrini: Stizopodina.

Szentivanya Kaszab, 1958: 1 [F]. Type species: Szentivanyametasternalis Kaszab, 1958, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Tabarus Gebien, 1921a: 317 [M]. Type species: Tabarusinfernalis Gebien, 1921, by subsequent designation (Gebien 1940: 1089). Status: valid genus in Stenochiinae: Cnodalonini.

Taclamacanius Ferrer & Yvinec, 2005: 121 [M]. Type species [automatic]: Taklamakanialepetzi Ferrer & Yvinec, 2004, by monotypy. Status: junior synonym of Paranemia Heyden, 1892 in Diaperinae: Phaleriini. Synonymy: G.S. Medvedev (2006: 557). Note: replacement name for Taklamakania Ferrer & Yvinec, 2004.

Tactoderus Fairmaire, 1892a: 112 [M]. Type species: Tactoderussubopacus Fairmaire, 1892 (= Praogenagagatina Mäklin, 1863), by monotypy. Status: junior synonym of Praeugena Laporte, 1840 in Tenebrioninae: Praeugenini. Synonymy: Gebien (1948: 544).

Tadzhikistania Bogatchev, 1960b: 35 [F]. Type species: Tadzhikistaniamystacea Bogatchev, 1960, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Taenobates Motschulsky, 1872: 25 [M]. Type species: Tenebriosaperdoides G.-A. Olivier, 1795, by original designation. Status: junior synonym of Xylopinus J.L. LeConte, 1862 in Stenochiinae: Cnodalonini. Synonymy: C.O. Waterhouse (1876: 288). Note: we act as First Revisers and reject the alternative original spelling Taeniobates, used by Motschulsky (1872: 32).

Tagalinus Kaszab, 1977a: 301, 333 [M]. Type species: Ulomalifuanum Montrouzier, 1860, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Tagalopsis Kaszab, 1955a: 471, 475 [F]. Type species: Tagalopsisszekessyi Kaszab, 1955, by original designation. Status: valid genus in Phrenapatinae: Penetini.

Tagalus Gebien, 1914b: 388 [M]. Type species: Tagalusimpressicollis Gebien, 1914, by subsequent designation (Gebien 1940: 756). Status: junior synonym of Dioedus J.L. LeConte, 1862 in Phrenapatinae: Penetini. Synonymy: Kaszab (1977a: 314).

Tagenesthes Koch, 1941: 276, 282 [F]. Type species: Asphaltesthesafrogermanica Koch, 1941, by monotypy. Status: valid subgenus of Asphaltesthes Kraatz, 1865 in Pimeliinae: Tentyriini.

Tagenia Latreille, 1802: 170 [F]. Type species: Tenebriofiliformis Fabricius, 1792, by monotypy. Status: junior synonym of Stenosis Herbst, 1799 in Pimeliinae: Stenosini: Stenosina. Synonymy: G.-A. Olivier (1803: 355, through synonymy of the type species).

Tagenodes Haag-Rutenberg, 1876: 87 [M]. Type species: Tagenodesmoufleti Haag-Rutenberg, 1876, by monotypy. Status: junior synonym of Rhammatodes Haag-Rutenberg, 1876 in Pimeliinae: Tentyriini. Synonymy: Koch (1952a: 133).

†Tagenopsis Heer, 1864: 377 [F]. Type species: Tagenopsisbrevicornis Heer, 1864, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6); described from Middle Miocene deposits (Germany).

Tagenostola Reitter, 1916d: 138, 151 [F]. Type species: Stenosisturkestanica Reitter, 1886, by subsequent designation (Gebien 1937a: 684). Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Tagona Fischer, 1820: pl. 16 [F]. Type species: Tagonaacuminata Fischer, 1820, by subsequent designation (Hope 1841: 124). Status: valid genus in Blaptinae: Blaptini: Prosodina.

Tagonoides Fairmaire, 1886d: 343 [F]. Type species: Tagonoidesdelavayi Fairmaire, 1886, by subsequent designation (Blair 1923a: 283). Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Taichius Ando, 1996: 196, 197 [M]. Type species: Platycrepishemiceroides Blair, 1929, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Taiwanocryphaeus Masumoto, 1996b: 67 [M]. Type species: Taiwanocryphaeusrhinoceros Masumoto, 1996, by original designation. Status: valid genus in Tenebrioninae: Toxicini: Toxicina.

Taiwanolagria Masumoto, 1988b: 41 [F]. Type species: Taiwanolagriamerkli Masumoto, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Taiwanomenephilus Masumoto, 1986b: 61 [M]. Type species: Taiwanomenephiluschui Masumoto, 1986, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Taiwanomenimus Masumoto, Akita & Lee, 2019: 161 [M]. Type species: Taiwanomenimusnakasatoi Masumoto, Akita & Lee, 2019, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Gnathidiina.

Taiwanotagalus Masumoto, 1982b: 143 [M]. Type species: Taiwanotagalusklapperichi Masumoto, 1982, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Taiwanotrachyscelis Masumoto, Akita & Lee, 2012: 29 [F]. Type species: Taiwanotrachyscelischengi Masumoto, Akita & Lee, 2012, by original designation. Status: valid genus in Diaperinae: Trachyscelini.

Taklamakania Ferrer & Yvinec, 2004: 42 [F]. Type species: Taklamakanialepetzi Ferrer & Yvinec, 2004, by monotypy. Status: junior synonym of Paranemia Heyden, 1892 in Diaperinae: Phaleriini. Synonymy: G.S. Medvedev (2006: 557). Note: junior homonym of Taklamakania Zhang, 1979 [Trilobita].

Talanus Jacquelin du Val, 1857: 156 [M]. Type species: Talanuscribrarius Jacquelin du Val, 1857, by monotypy. Status: valid genus in Stenochiinae: Talanini.

Tamatasida Koch, 1962a: 90, 145 [F]. Type species: Scotinesthestuberculosa Fairmaire, 1895, by original designation. Status: valid genus in Pimeliinae: Asidini.

Tamdaous Pic, 1923b: 11 [M]. Type species: Tamdaousimpressus Pic, 1923, by monotypy. Status: valid subgenus of Oedemutes Pascoe, 1860 in Stenochiinae: Cnodalonini.

Tamena Reitter, 1900c: 90, 143 [F]. Type species: Psammocryptusrugiceps Reitter, 1897, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Tanchirus Fairmaire, 1897e: 224 [M]. Type species: Tanchiruscompactus Fairmaire, 1897, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tangiprosodes G.S. Medvedev, 2005b: 84 [M]. Type species: Prosodespygmaea Kraatz, 1882, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina. Note: originally described as a section within a subgenus.

Tanychilus Newman, 1838: 487 [M]. Type species: Tanychilusstriatus Newman, 1838, by subsequent designation (Cazurro Ruiz 1897a: 213). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Tanylypa Pascoe, 1869: 152 [F]. Type species: Tanylypamorio Pascoe, 1869, by monotypy. Status: valid genus in Zolodininae.

Tapenopsis Solier, 1843: 5 [F]. Type species: Tapenopsiscostata Solier, 1843, by original designation. Status: valid genus in Pimeliinae: Leptodini.

Taphrosoma Kirsch, 1866: 195 [N]. Type species: Taphrosomadohrnii Kirsch, 1866, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tapinocomus Gebien, 1928: 102 [M]. Type species: Tapinocomussubnudus Gebien, 1928, by monotypy. Status: valid genus in Pimeliinae: Epitragini.

Tapinopsis Agassiz, 1846b: 361 [F]. Type species [automatic]: Tapenopsiscostata Solier, 1843, by original designation. Status: junior synonym of Tapenopsis Solier, 1843 in Pimeliinae: Leptodini. Note: unjustified emendation of Tapenopsis Solier, 1843, not in prevailing usage.

Taraxides C.O. Waterhouse, 1876: 289 [M]. Type species: Helopssinuatus Fabricius, 1801 (= Tenebriolaevigatus Fabricius, 1781), by subsequent designation (Gebien 1941: 340). Status: valid genus in Stenochiinae: Cnodalonini. Note: the original combination of the accepted name of the type species, Tenebriolaevigatus Fabricius, 1781, is a junior primary homonym of Tenebriolaevigatus Linnaeus, 1767.

Tarpela Bates, 1870: 272 [F]. Type species: Tarpelabrownii Bates, 1870, by subsequent designation (Gebien 1943: 407). Status: valid genus in Tenebrioninae: Helopini: Helopina.

Tarphiophasis Wollaston, 1877: 227 [F]. Type species: Tarphiophasistuberculata Wollaston, 1877, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Tarsocnodes Gebien, 1920: 82 [M]. Type species: Psammodesmolossus Haag-Rutenberg, 1871, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Tarsosis Gebien, 1920: 33, 34 [F]. Type species: Stenocaradamarense Péringuey, 1886, by monotypy. Status: valid subgenus of Zophosis Latreille, 1802 in Pimeliinae: Zophosini.

Tauroceras Hope, 1841: 130 [N]. Type species: Tenebriocornutus Fabricius, 1775, by original designation. Status: valid genus in Tenebrioninae: Centronopini.

Tauroceropedus Pic, 1913b: 4 [M]. Type species: Tauroceropedusdifformipes Pic, 1913, by subsequent designation (Gebien 1941: 344). Status: junior synonym of Tauroceras Hope, 1841 in Stenochiinae: Cnodalonini. Synonymy: Ferrer et al. (2005: 272).

Taurohelops Keskin & Nabozhenko, 2015: 84 [M]. Type species: Stenomaxincultus Allard, 1877, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Taxes Champion, 1895a: 226 [M]. Type species: Taxesdepressus Champion, 1895, by subsequent designation (R. Lucas 1920: 628). Status: valid genus in Alleculinae: Alleculini: Alleculina.

Tearchus Kraatz, 1880b: 110 [M]. Type species: Tearchusannulipes Kraatz, 1880, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tedinus Casey, 1891: 153 [M]. Type species: Tedinusangustus Casey, 1891, by monotypy. Status: junior synonym of Isomira Mulsant, 1856 in Alleculinae: Alleculini: Gonoderina. Synonymy: Bousquet and Campbell (1991: 259).

Telabis Casey, 1890b: 331 [M]. Type species: Eurymetoponlongipenne Casey, 1890, by subsequent designation (Casey 1907: 288). Status: valid genus in Pimeliinae: Edrotini. Note: see Alonso-Zarazaga in Bousquet et al.(2018: 116) for comments on the gender of this genus-group name.

Telacis Poey, 1854: 322 [M]. Type species [automatic]: Chrysomelasulphurea Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Cteniopus Solier, 1835 in Alleculinae: Cteniopodini. Note: replacement name for Cistela Fabricius, 1775, a senior synonym of Cteniopus Solier, 1835.

Telaponium Blaisdell, 1923: 209 [N]. Type species: Telaponiumcastaneum Blaisdell, 1923, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Telchis Champion, 1886: 142 [M]. Type species: Telchisclavicornis Champion, 1886, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Teles Mulsant & Godart, 1876: 163 [M]. Type species: Teleseutymi Mulsant & Godart, 1876, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: the alternative original spelling Tales, used by Mulsant and Godart (1876: 164), was rejected by Rye (1879: 62) who acted as the First Reviser.

Telesicles Champion, 1888: 450 [M]. Type species: Telesiclescordatus Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Telethrus Pascoe, 1882: 29 [M]. Type species: Telethrusebeninus Pascoe, 1882, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Telleus Fairmaire, 1904b: 465 [M]. Type species: Telleuscrenatus Fairmaire, 1904, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Temnes Champion, 1888: 410 [M]. Type species: Temnescaeruleus Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Temnoaphelus Ferrer, 1988: 377 [M]. Type species: Temnoaphelushispidus Ferrer, 1988, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Temnophthalmus Gebien, 1921b: 62, 75 [M]. Type species: Temnophthalmusscalaris Gebien, 1921, by subsequent designation (Gebien 1941: 341). Status: valid genus in Stenochiinae: Cnodalonini.

Teneatopus Reitter, 1920a: 23 [M]. Type species: Tenebrioatronitens Fairmaire, 1891, by monotypy. Status: junior synonym of Ariarathus Fairmaire, 1891 in Tenebrioninae: Tenebrionini. Synonymy: Ardoin (1969b: 126).

Tenebrio Linnaeus, 1758: 417 [M]. Type species: Tenebriomolitor Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: valid genus and subgenus in Tenebrioninae: Tenebrionini.

Tenebriocamaria Pic, 1919b: 3 [F]. Type species: Tenebriocamariaatra Pic, 1919, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tenebriocephalon Pic, 1925b: 10 [N]. Type species: Tenebriocephalonpiceum Pic, 1925, by monotypy. Status: valid genus in Pimeliinae: Ceratanisini.

Tenebrioloma Gebien, 1910c: 386 [N]. Type species: Tenebriolomasemicostatum Gebien, 1910, by monotypy. Status: junior synonym of Tribolium W.S. MacLeay, 1825 in Tenebrioninae: Triboliini. Synonymy: Halstead (1967b: 270).

Tenebriomimus Kolbe, 1901: 342 [M]. Type species: Tenebriomimusadansoniarum Kolbe, 1901 (= Palembusocularis Casey, 1891), by monotypy. Status: junior synonym of Ulomoides Blackburn, 1888 in Diaperinae: Diaperini: Diaperina. Synonymy: Gebien (1922b: 268, with Martianus Fairmaire, 1893, a junior synonym of Ulomoides Blackburn, 1888).

Tenebrionellus Crotch, 1874: 105 [M]. Type species [automatic]: Tenebriomolitor Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Tenebrio Linnaeus, 1758 in Tenebrioninae: Tenebrionini. Note: unnecessary replacement name for Tenebrio Linnaeus, 1758.

†Tenebrionites Cockerell, 1920: 67 [M]. Type species: Tenebrionitesanglicus Cockerell, 1920, by monotypy. Status: valid genus in Tenebrionidae: incertae sedis. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6); described from Middle Eocene deposits (United Kingdom).

Tenebriopsis Gebien, 1928: 170, 186 [F]. Type species: Tenebriopsissubtilicostis Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tenesis Duvivier, 1892: 163 [M]. Type species: Tenesisfemoratus Duvivier, 1892, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tentyria Latreille, 1802: 170 [F]. Type species: Tentyrialigurica Solier, 1835, by plenary powers (ICZN 2010a, Opinion 2244). Status: valid genus in Pimeliinae: Tentyriini. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2010a, Opinion 2244).

Tentyriina Peyerimhoff, 1907: 31 [F]. Type species [automatic]: Pimeliaorbiculata Fabricius, 1775, by subsequent designation (Gebien 1937a: 624; see ICZN 2010a, Opinion 2244). Status: junior synonym of Tentyrina Reitter, 1900 in Pimeliinae: Tentyriini. Note: unjustified emendation of Tentyrina Reitter, 1900, not in prevailing usage.

Tentyrina Reitter, 1900c: 92, 166 [F]. Type species: Pimeliaorbiculata Fabricius, 1775, by subsequent designation (Gebien 1937a: 624; see ICZN 2010a, Opinion 2244). Status: valid genus in Pimeliinae: Tentyriini. Note: name placed on the Official List of Generic Names in Zoology (ICZN 2010a, Opinion 2244).

Tentyriomorpha Peyerimhoff, 1927: 52 [F]. Type species [automatic]: Tentyromorphatelueti Escalera, 1913, by monotypy. Status: junior synonym of Tentyromorpha Escalera, 1913 in Pimeliinae: Tentyriini. Note: unjustified emendation of Tentyromorpha Escalera, 1913, not in prevailing usage.

Tentyriopsis Gebien, 1928: 168, 174 [F]. Type species: fixed herein (ICZN 1999, Article 70.3) as Tentyriopsispertyi Gebien, 1940, misidentified as Tentyriastriipuncta Perty, 1830 in the original designation by monotypy in Gebien (1928). Status: valid genus in Stenochiinae: Cnodalonini. Note: the type species Tentyriastriipuncta Perty was first established by monotypy; Gebien (1940: 1091) reported that Tentyriastriipuncta Perty of Gebien (1928) was misidentified and proposed the name Tentyriopsispertyi for it; we follow currently accepted concepts (e.g., Blackwelder 1945: 533) and fix the type species according to the requirements of Article 70.3.2 (ICZN 1999); the nominal species Tentyriastriipuncta Perty, 1830 is also a valid species in the genus Tentyriopsis Gebien, 1928.

Tentyrodera Koch, 1943a: 572 [F]. Type species: Microderamarginata Baudi di Selve, 1874, by monotypy. Status: valid subgenus of Microdera Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Tentyromorpha Escalera, 1913: 38 [F]. Type species: Tentyromorphatelueti Escalera, 1913, by monotypy. Status: valid subgenus of Pachychila Eschscholtz, 1831 in Pimeliinae: Tentyriini.

Tentyronota Reitter, 1900c: 93, 188 [F]. Type species: Micipsarotundicollis Kraatz, 1865, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Terametus Motschulsky, 1869: 193 [M]. Type species: Terametuscapicola Motschulsky, 1869, by monotypy. Status: valid genus in Lagriinae: Lupropini.

Teremenes Carter, 1914b: 54 [M]. Type species: Tenebriolongipennis Hope, 1843, by present designation. Status: junior synonym of Zophophilus Fairmaire, 1881 in Stenochiinae: Cnodalonini. Synonymy: Carter (1930: 547).

Termitonebria Wasmann in Wasmann and Brauns, 1925: 105 [F]. Type species: Termitonebriabraunsi Wasmann, 1925 (= Asyleptusfumosus Péringuey, 1896), by monotypy. Status: junior synonym of Asyleptus Péringuey, 1896 in Tenebrioninae: Amarygmini. Synonymy: Schawaller and Bremer (2013: 81).

Tessaromma Boheman, 1858: 91 [N]. Type species: Tessarommalugubre Boheman, 1858, by subsequent designation (Aalbu and Triplehorn 1991: 170). Status: junior synonym of Blapstinus Dejean, 1821 in Blaptinae: Opatrini: Blapstinina. Synonymy: Gemminger in Gemminger and Harold (1870: 1923, with Pedonoeces G.R. Waterhouse, 1845, a junior synonym of Blapstinus Dejean, 1821), Aalbu and Triplehorn (1991: 170). Note: junior homonym of Tessaromma Newman, 1840 [Coleoptera: Cerambycidae].

Tetragonomecus Rye, 1880: 87 [M]. Type species [automatic]: Tetragonomenessemiviridis Chevrolat, 1878, by monotypy. Status: junior synonym of Tetragonomenes Chevrolat, 1878 in Stenochiinae: Cnodalonini. Note: unjustified emendation for Tetragonomenes Chevrolat, 1878, not in prevailing usage.

Tetragonomenes Chevrolat, 1878b: clii [M]. Type species: Tetragonomenessemiviridis Chevrolat, 1878, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Tetranillus Wasmann, 1899a: 167 [M]. Type species: Tetranilluscostatus Wasmann, 1899, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Tetranosis G.S. Medvedev, 1995a: 858 [F]. Type species: Tetranosisclypeoloba Koch, 1940, by original designation. Status: valid subgenus of Microtelopsis Koch, 1940 in Pimeliinae: Stenosini: Stenosina. Note: as pointed out by Aalbu et al. (2017: 326) this name was first introduced by Koch (1940b: 740); however, Koch did not designate a type species and therefore his name is unavailable; Löbl and Merkl (2003: 251) designated a type species for this genus but did not describe the genus-group taxon as new (ICZN 1999, Article 16.1); G.S. Medvedev (1995a: 858) was the first to provide a description of Tetranosis as well as fixing a type species for the genus, thereby making the name nomenclaturally available for the first time; we hereby act as First Revisers and select Microtelopsis Koch, 1940 as the valid name of the genus based on the Principle of Priority (ICZN 1999, Article 23.3.5).

Tetraphyllus Laporte & Brullé, 1831: 332, 404 [M]. Type species: Tetraphylluslatreillei Laporte & Brullé, 1831, by subsequent designation (Gebien 1935: 64). Status: valid genus in Stenochiinae: Cnodalonini. Note: nomenclatural stability in this genus is threatened by the discovery of an older type species designation (Tetraphyllusformosus Laporte & Brullé, 1831, by subsequent designation by Bates (1879a: 293), which is the type species of Damatris Laporte, 1840); we recommend that an application be submitted to the International Commission on Zoological Nomenclature to maintain the type species designation proposed by Gebien (1935: 64).

Tetrethas Koch, 1962a: 23, 145 [M]. Type species: Anethasxylophilus Koch, 1962, by original designation. Status: valid subgenus of Anethas Jakobson, 1924 in Pimeliinae: Stenosini: Stenosina.

Tetromma Dejean, 1834: 183 [N]. Type species: Upisunicolor Herbst, 1797, by subsequent designation (Löbl et al. 2008a: 43). Status: junior synonym of Hyperops Eschscholtz, 1831 in Pimeliinae: Tentyriini. Synonymy: Dejean (1836: 203).

Texaponium Thomas, 1984: 658 [N]. Type species: Cryptadiustriplehorni Berry, 1974, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Thaioblaps Masumoto, 1989: 187 [F]. Type species: Thaioblapspunneeae Masumoto, 1989, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Thalpobia Fairmaire, 1871a: 385 [F]. Type species: Thalpobialaevipennis Fairmaire, 1871, by monotypy. Status: valid genus in Pimeliinae: Tentyriini.

Thalpophila Solier, 1835b: 253, 370 [F]. Type species: Akisabbreviata Fabricius, 1801, by subsequent designation (Chevrolat 1849: 544). Status: senior synonym of Thalpophilodes Strand, 1942 in Pimeliinae: Tentyriini. Note: junior homonym of Thalpophila Hübner, 1820 [Lepidoptera].

Thalpophilodes Strand, 1942: 391 [M]. Type species [automatic]: Akisabbreviata Fabricius, 1801, by subsequent designation (Chevrolat 1849: 544). Status: valid genus in Pimeliinae: Tentyriini. Note: replacement name for Thalpophila Solier, 1835.

Tharsus J.L. LeConte, 1862: 233 [M]. Type species: Tharsusseditiosus J.L. LeConte, 1862, by monotypy. Status: junior synonym of Metaclisa Jacquelin du Val, 1861 in Tenebrioninae: Metaclisini. Synonymy: Steiner (2016: 538).

Thaumatoblaps Kaszab & G.S. Medvedev, 1984: 79 [F]. Type species: Thaumatoblapsmarikovskiji Kaszab & G.S. Medvedev, 1984, by original designation. Status: valid genus in Blaptinae: Blaptini: Blaptina.

Theatetes Champion, 1888: 420 [M]. Type species: Theatetesbasicornis Champion, 1888, by monotypy. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Thecacerus Lacordaire, 1859b: 420 [M]. Type species: Cnodalonnodosum Gray, 1831, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Theresea Pic, 1917d: 12 [F]. Type species: Thereseadiversipennis Pic, 1917, by monotypy. Status: valid genus in Stenochiinae: Stenochiini. Note: Masumoto (1999b: 130) transferred this genus from Cnodalonini.

Thesilea Haag-Rutenberg, 1878: 103 [F]. Type species: Thesileaversicolor Haag-Rutenberg, 1878, by subsequent designation (R. Lucas 1920: 638). Status: valid genus in Stenochiinae: Cnodalonini.

Thettea Bates, 1879a: 290 [F]. Type species: Thetteatenuitarsis Bates, 1879, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Thinobatis Eschscholtz, 1831: 5, 8 [F]. Type species: Thinobatisferruginea Eschscholtz, 1831, by monotypy. Status: valid genus in Pimeliinae: Thinobatini.

Thoracon Gistel, 1848a: xi [N]. Type species [automatic]: Silphasabulosa Linnaeus, 1758, by subsequent designation (Latreille 1810: 429). Status: junior synonym of Opatrum Fabricius, 1775 in Blaptinae: Opatrini: Opatrina. Note: unnecessary replacement name for Opatrum Fabricius, 1775.

Thoracophorus Hope, 1841: 188 [M]. Type species: Thoracophoruswalckenaerii Hope, 1841, by original designation. Status: senior synonym of Cardiothorax Motschulsky, 1860 in Lagriinae: Adeliini. Synonymy: Motschulsky (1860a: 67). Note: junior homonym of Thoracophorus Motschulsky, 1837 [Coleoptera: Staphylinidae].

Thoracostira Borchmann, 1936: 240, 508 [F]. Type species: Statirasculpta Kirsch, 1873, by original designation. Status: valid genus in Lagriinae: Lagriini: Statirina.

Thorictophasis Koch, 1950c: 80 [F]. Type species: Caenocrypticusdeserticus Koch, 1950, by original designation. Status: junior synonym of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini. Synonymy: Endrödy-Younga (1996: 13).

Thorictosoma Lea, 1919: 257 [N]. Type species: Thorictosomaectatommae Lea, 1919, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Thorictosomatina.

Thornella Novák, 2019e: 83 [F]. Type species: Alleculaholomelaena Fairmaire, 1894, by original designation. Status: valid subgenus of Upinella Mulsant, 1856 in Alleculinae: Alleculini: Alleculina.

Thoseus Pic, 1925b: 9 [M]. Type species: Thoseusrufus Pic, 1925, by monotypy. Status: valid genus in Lagriinae: Belopini.

Thraustocolus Kraatz, 1866: 414 [M]. Type species [automatic]: Calobamonleptoderus Kraatz, 1865, by monotypy. Status: valid genus & subgenus in Pimeliinae: Tentyriini. Note: replacement name for Calobamon Kraatz, 1865.

Threnus Motschulsky, 1870: 404 [M]. Type species: Threnusniger Motschulsky, 1870, by original designation. Status: senior synonym of Argoporis Horn, 1870 in Tenebrioninae: Cerenopini. Synonymy: Aalbu et al. (1995: 483). Note: see the entry for Argoporis Horn, 1870 regarding the priority of these two names.

Thriptera Solier, 1836: 9, 48 [F]. Type species: Thripteramaillei Solier, 1836 (= Pimeliacrinita Klug, 1830), by subsequent designation (Hope 1841: 118). Status: valid genus in Pimeliinae: Pimeliini.

Thurea Ferrer, 1998b: 151 [F]. Type species: Thureapalmi Ferrer, 1998, by original designation. Status: junior synonym of Platycotylus Olliff, 1883 in Tenebrioninae: Palorini. Synonymy: Schawaller (2014: 51).

Thydemorphus Pic, 1918b: 19 [M]. Type species: Thydemorphuspilitarsis Pic, 1918, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Thydemus Lewis, 1894: 475 [M]. Type species: Scotaeuspurpurivittatus Marseul, 1876, by original designation. Status: junior synonym of Pseudonautes Fairmaire, 1892 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1914a: 58).

Thylacoderes Solier, 1843: 44, 123 [M]. Type species: Thylacodereseumolpoides Solier, 1843, by original designation. Status: valid genus in Pimeliinae: Praociini.

Tibinella Novák, 2019e: 91 [F]. Type species: Upinellapahangica Novák, 2019, by original designation. Status: valid subgenus of Upinella Mulsant, 1856 in Alleculinae: Alleculini: Alleculina.

​Tibiocnodes ​Gearner & Kamiński in Gearner et al., 2021: 8 [M]. Type species: Psammodeslucidus ​Fåhraeus, 1870, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Tidiguinia Español, 1959: 249 [F]. Type species: Tidiguiniabolivari Español, 1959, by original designation. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Timeneca Carter, 1914b: 78 [F]. Type species [automatic]: Ctimenebreweri Bates, 1873, by monotypy. Status: junior synonym of Mitrothorax Carter, 1914 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 456). Note: replacement name for Ctimene Bates, 1873, a senior synonym of Mitrothorax Carter, 1914.

Timogebienus Ardoin, 1963b: 308, 331 [M]. Type species: Hoplonyxcollaris Gebien, 1910, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Timosmithus Ardoin, 1974b: 457 [M]. Type species: Timosmithusbasilewskyi Ardoin, 1974, by monotypy. Status: valid genus in Pimeliinae: Stenosini: Stenosina.

Tinophyllus Fairmaire, 1869b: 234 [M]. Type species: Tinophyllusgracilicornis Fairmaire, 1869, by monotypy. Status: junior synonym of Camariodes Fairmaire, 1869 in Stenochiinae: Cnodalonini. Synonymy: Fairmaire (1886c: 75).

Tisamenes Champion, 1884: 64 [M]. Type species: Tisamenestruquii Champion, 1884, by monotypy. Status: valid subgenus of Philolithus Lacordaire, 1858 in Pimeliinae: Asidini.

Titaena Erichson, 1842a: 178 [F]. Type species: Titaenacolumbina Erichson, 1842, by subsequent designation (Chevrolat 1849: 594). Status: valid genus in Tenebrioninae: Titaenini.

Tithassa Pascoe, 1860b: 125 [F]. Type species: Tithassacorynomelas Pascoe, 1860, by monotypy. Status: valid genus in Lagriinae: Goniaderini.

Tjikoraia Pic, 1921d: 18 [F]. Type species: Tjikoraiajavana Pic, 1921, by subsequent designation (Riley 1923: 130). Status: junior synonym of Menimus Sharp, 1876 in Diaperinae: Gnathidiini: Gnathidiina. Synonymy: Kaszab (1979a: 81).

Tlascalinus Casey, 1907: 370 [M]. Type species: Trimytisflohri Champion, 1892, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Toktokkus Kamiński & Gearner in Kamiński et al., 2020: 10 [M]. Type species: Toktokkustschinkeli Kamiński & Gearner, 2020, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Tomogria Merkl, 1988a: 135 [F]. Type species: Tomogriaperlata Merkl, 1988, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Tonibiastes Casey, 1895: 617 [M]. Type species: Notibiuscostipennis Horn, 1894, by original designation. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Tonibius Casey, 1895: 617 [M]. Type species: Notibiussulcatus J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 644). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Tonkinius Fairmaire, 1903a: 13 [M]. Type species: Tonkiniussculptilis Fairmaire, 1903, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Toreuma Carter, 1913a: 84 [N]. Type species: Toreumacupreum Carter, 1913, by monotypy. Status: senior synonym of Atoreuma Gebien, 1941 in Tenebrioninae: Heleini: Cyphaleina. Note: junior homonym of Toreuma Haeckel, 1880 [Cnidaria].

Toxicum Latreille, 1802: 174 [N]. Type species: Toxicumrichesianum Latreille, 1802, by monotypy. Status: valid genus and subgenus in Tenebrioninae: Toxicini: Toxicina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6).

Toxocnema Fåhraeus, 1870: 304 [F]. Type species: Toxocnemarufitarsis Fåhraeus, 1870, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini. Note: transferred from Tenebrioninae: Ulomini by Schawaller (2009b: 363).

Trachasida Reitter, 1917a: 41, 62 [F]. Type species: Asidaruficornis Solier, 1836, by subsequent designation (F. Soldati 2008: 33). Status: junior synonym of Gracilasida Escalera, 1905 in Pimeliinae: Asidini. Synonymy: Wilke (1922: 259, with Planasida Escalera, 1907, a junior synonym of Gracilasida Escalera, 1905).

Tracheloeum Hope, 1841: 116 [N]. Type species: Tracheloeumlaticolle Hope, 1841, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6).

Trachelostenus Solier, 1851: 255 [M]. Type species: Trachelostenusinaequalis Solier, 1851, by monotypy. Status: valid genus in Tenebrioninae: Trachelostenini. Note: the family-group name based on this genus was downgraded from the family Trachelostenidae (within the superfamily Tenebrionoidea) to the tribe Trachelostenini (within the tenebrionid subfamily Tenebrioninae) by Matthews and Lawrence (2015: 290).

Trachyderes Koch, 1955a: 112 [M]. Type species: Trachynotusbipunctatus Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Trachyderma Latreille, 1828: 576 [N]. Type species: Pimeliahispida Fabricius, 1775 (= Tenebriohispidus Forskål, 1775), by subsequent designation (P.H. Lucas 1839: 50). Status: valid genus and subgenus in Pimeliinae: Pimeliini. Note: gender of genus corrected from feminine to neuter by Löbl and Smetana (2011: 33).

Trachydermum Gistel, 1848a: xi [N]. Type species [automatic]: Pimeliahispida Fabricius, 1775 (= Tenebriohispidus Forskål, 1775), by subsequent designation (P.H. Lucas 1839: 50). Status: junior synonym of Trachyderma Latreille, 1828 in Pimeliinae: Pimeliini. Note: unnecessary replacement name for Trachyderma Latreille, 1828.

Trachymetus Reitter, 1904: 51, 76 [M]. Type species: Pachypteruselongatus Mulsant & Rey, 1859, by monotypy. Status: junior synonym of Amblysphagus Fairmaire, 1896 in Blaptinae: Opatrini: Neopachypterina. Synonymy: Kaszab (1975a: 27).

Trachynotidus Péringuey, 1899: 296 [M]. Type species: Psammodesthoreyi Haag-Rutenberg, 1871, by subsequent designation (Kamiński et al. 2019b: 18). Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Trachynotus Latreille, 1828: 579 [M]. Type species: Sepidium vittatum Fabricius, 1781, by subsequent designation (Kamiński et al. 2019b: 93). Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Trachyscelis Latreille, 1809: 379 [F]. Type species: Trachyscelisaphodioides Latreille, 1809, by monotypy. Status: valid genus in Diaperinae: Trachyscelini.

Tragardhus Koch, 1956a: 369 [M]. Type species: Tragardhusgrandipleurum Koch, 1956, by original designation. Status: valid genus and subgenus in Blaptinae: Dendarini: Melambiina.

Trestonia Rafinesque, 1815: 113 [F]. Type species [automatic]: Toxicumrichesianum Latreille, 1802, by monotypy. Status: junior synonym of Toxicum Latreille, 1802 in Tenebrioninae: Toxicini: Toxicina. Note: unnecessary replacement name for Toxicum Latreille, 1802.

Triangulipenna Louw, 1979: 100, 114 [F]. Type species: Triangulipennalacuna Louw, 1979, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Tribolium W.S. MacLeay, 1825: 47 [N]. Type species: Colydiumcastaneum Herbst, 1797, by monotypy (see ICZN 1988, Opinion 1495). Status: valid genus and subgenus in Tenebrioninae: Triboliini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1988, Opinion 1495).

Trichamarygmus Carter, 1913b: 46 [M]. Type species: Trichamarygmuspilosus Carter, 1913, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Trichanemia Ardoin, 1971: 361, 406 [F]. Type species: Anemiaschmitzi Ardoin, 1971, by monotypy. Status: valid subgenus of Cheirodes Gené, 1839 in Tenebrioninae: Melanimonini.

Tricheleodes Blaisdell, 1909: 138 [M]. Type species: Eleodeshirsuta J.L. LeConte, 1861, by subsequent designation (Johnston 2016: 666). Status: valid subgenus of Eleodes Eschscholtz, 1829 in Blaptinae: Amphidorini.

Trichethmus Gebien, 1937b: 45 [M]. Type species: Trichethmuslobicollis Gebien, 1937, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Trachynotina.

Trichiasida Casey, 1912: 77, 172 [F]. Type species: Pelecyphorushirsutus J.L. LeConte, 1851, by original designation. Status: valid subgenus of Stenomorpha Solier, 1836 in Pimeliinae: Asidini.

Trichiotes Casey, 1907: 432, 443 [M]. Type species: Trichiotesseriatus Casey, 1907, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Trichochianalus Kaszab, 1940a: 149, 201 [M]. Type species: Platynoscelismonticola Kaszab, 1940, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini. Note: the spellings Trichochianalis (p. 201 – also used in the directory of new taxa published in volume 30 of the journal Mitteilungen der Münchner Entomologischen Gesellschaft (p. xi)) and Trichochianalus (pp 125, 149, 201) were used in the original publication; Kaszab (1940b: 989) used the spelling Trichochianalus only and therefore acted as the First Reviser (ICZN 1999, Article 24.2.4).

Trichodamatris Chatanay, 1915a: 506, 524 [M]. Type species: Porphyrhybaraffrayi Fairmaire, 1884, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Trichoderulus Blaisdell, 1923: 281 [M]. Type species: Trichoderuluslongipilosus Blaisdell, 1923 (= Eleodestribulus Thomas, 2005), by original designation. Status: junior synonym of Pseudeleodes Blaisdell, 1909 in Blaptinae: Amphidorini. Synonymy: Johnston (2016: 672).

Tricholeipopleura Kaszab, 1940a: 152, 223 [F]. Type species: Platynoscelislucidicollis Kraatz, 1882, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini. Note: the spellings Tricholeipopleura (pp 131, 223, 225–228, also used in the directory of new taxa published in volume 30 of the journal Mitteilungen der Münchner Entomologischen Gesellschaft (p. xii)) and Tricholeipoleura (pp. 152) were used in the original publication; Kaszab (1940b: 989) used the spelling Tricholeipopleura only and therefore acted as the First Reviser (ICZN 1999, Article 24.2.4).

Trichomyatis Schuster in Reinig, 1931: 893 [F]. Type species: Trichomyatisconradti Schuster, 1931, by original designation. Status: valid genus in Blaptinae: Platyscelidini.

Trichoodescelis Kaszab, 1940b: 940, 954 [F]. Type species: Platyscelisacutangula Kraatz, 1884, by original designation. Status: junior synonym of Longuloodescelis Kaszab, 1940 in Blaptinae: Platyscelidini. Synonymy: Egorov (2004: 595).

Trichoplatynoscelis Kaszab, 1940b: 896 [F]. Type species: Trichoplatynoscelispamirensis Kaszab, 1940, by original designation. Status: junior synonym of Trichomyatis Schuster, 1931 in Blaptinae: Platyscelidini. Synonymy: Egorov (2004: 603). Note: this name was also used in several instances by Kaszab (1940a: 126, 132, 142, 143, 146); however, it is not available from that publication since there is no type species designated.

Trichoplatyscelis Reinig, 1931: 895 [F]. Type species: Trichoplatyscelispamirensis Reinig, 1931, by original designation. Status: valid subgenus of Bioramix Bates, 1879 in Blaptinae: Platyscelidini.

Trichopodus Mulsant & Rey, 1859c: 58, 59 [M]. Type species: Trichopodusvalidus Mulsant & Rey, 1859, by monotypy. Status: senior synonym of Trichosternum Wollaston, 1861 in Blaptinae: Opatrini: Opatrina. Synonymy: Bates (1872a: 98). Note: junior homonym of Trichopodus Lacepède, 1802 [Pisces].

Trichosaragus Blackburn, 1890: 1269 [M]. Type species: Trichosaraguspilosellus Blackburn, 1890, by monotypy. Status: valid genus in Tenebrioninae: Heleini: Heleina.

Trichosphaena Reitter, 1916c: 140, 145 [F]. Type species: Sphenariaolgae Semenov, 1889, by subsequent designation (Gebien 1937a: 577). Status: valid genus in Pimeliinae: Tentyriini.

Trichosternum Wollaston, 1861: 205 [N]. Type species: Trichosternumstriatum Wollaston, 1861, by monotypy. Status: valid genus in Blaptinae: Opatrini: Opatrina.

Trichostethe Koch, 1950b: 285 [F]. Type species: Cyphostethepilosa Koch, 1950, by original designation. Status: valid subgenus of Cyphostethe Marseul, 1866 in Pimeliinae: Tentyriini.

Trichotenebrio Ardoin, 1962a: 64 [M]. Type species: Trichotenebrioatronitidus Ardoin, 1962, by monotypy. Status: valid genus in Tenebrioninae: Tenebrionini.

Trichotoides Marcuzzi, 1954: 23 [M]. Type species: Scapteshintoni Kaszab, 1949, by monotypy. Status: junior synonym of Ammodonus Mulsant, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Ferrer and Moraguès (2001: 499).

Trichoton Hope, 1841: 111 [N]. Type species: Trichotoncayennense Hope, 1841, by original designation. Status: valid genus in Blaptinae: Opatrini: Blapstinina. Note: combined description of a new genus and a single new species (ICZN 1999, Article 12.2.6).

Trichotrachys Koch, 1955a: 201 [M]. Type species: Trachynotussordidus Gerstaecker, 1854, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Trichotrichus Koch, 1955a: 108 [M]. Type species: Trachynotuscrinitus Haag-Rutenberg, 1873, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Trichotum Agassiz, 1846b: 376 [N]. Type species [automatic]: Trichotoncayennense Hope, 1841, by original designation. Status: junior synonym of Trichoton Hope, 1841 in Blaptinae: Opatrini: Blapstinina. Note: unjustified emendation of Trichoton Hope, 1841, not in prevailing usage.

Trichulodes Carter, 1914a: 223 [M]. Type species: Trichulodespunctatus Carter, 1914, by monotypy. Status: junior synonym of Pseudolyprops Fairmaire, 1882 in Lagriinae: Goniaderini. Synonymy: Doyen et al. (1990: 231).

Trientoma Solier, 1835b: 253, 256 [F]. Type species: Trientomavarvasi Solier, 1835, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Trigonocnera Reitter, 1893: 202, 213 [F]. Type species: Trigonoscelispseudopimelia Reitter, 1889, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Trigonopachys Skopin, 1968b: 300 [M]. Type species: Trigonopachysmichailovi Skopin, 1968, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Trigonopilus Fairmaire, 1893b: 22 [M]. Type species: Trigonopiluslaticeps Fairmaire, 1893, by monotypy. Status: valid genus in Blaptinae: Opatrini: incertae sedis. Note: placed in Opatrini incertae sedis by Kamiński et al. (2021b: 151).

Trigonopoda Gebien, 1914a: 2 [F]. Type species: Trigonopodacrassipes Gebien, 1914, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Trigonopus Mulsant & Rey, 1853b: 105 [M]. Type species: Trigonopuscapicola Mulsant & Rey, 1853, by subsequent designation (Cazurro Ruiz 1897b: 539). Status: valid genus in Blaptinae: Platynotini: Platynotina.

Trigonoscelis Dejean, 1834: 179 [F]. Type species: Pimelianodosa Fischer, 1820, by subsequent designation (Hope 1841: 118). Status: valid genus and subgenus in Pimeliinae: Pimeliini.

Trigonotarsus Hope, 1843: 357 [M]. Type species: Trigonotarsusaustralis Hope, 1843, by original designation. Status: senior synonym of Sobas Pascoe, 1863 in Blaptinae: Opatrini: Opatrina. Note: junior homonym of Trigonotarsus Guérin-Méneville, 1838 [Coleoptera: Curculionidae].

Trilobocara Solier, 1851: 129 [N]. Type species: Trilobocaraciliatum Solier, 1851, by original designation. Status: valid genus in Pimeliinae: Trilobocarini.

Trimytantron Ardoin, 1977: 381 [N]. Type species: Trimytantrondecui Ardoin, 1977, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Trimytis J.L. LeConte, 1851: 141 [F]. Type species: Trimytispruinosa J.L. LeConte, 1851, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Triorophus J.L. LeConte, 1851: 141 [M]. Type species: Triorophuslaevis J.L. LeConte, 1851, by subsequent designation (Casey 1907: 432). Status: valid genus in Pimeliinae: Edrotini.

Triphalopsis Blaisdell, 1923: 232 [F]. Type species: Triphalopsispartida Blaisdell, 1923, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Triphalopsoides Doyen, 1990: 222 [M]. Type species: Triphalopsoideslasiodorsa Doyen, 1990, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Triphalus J.L. LeConte, 1866b: 104 [M]. Type species: Triphalusperforatus J.L. LeConte, 1866, by monotypy. Status: valid genus in Pimeliinae: Edrotini.

Triplehornia Matthews & Lawrence, 2005: 543 [F]. Type species: Triplehorniametallica Matthews & Lawrence, 2005, by original designation. Status: valid genus in Diaperinae: incertae sedis. Note: Matthews and Lawrence (2019: 651) mentioned that this genus belongs to an unnamed tribe within the subfamily Diaperinae.

Tripolicryptus Strand, 1929: 24 [M]. Type species [automatic]: Brachycryptustripolitanus Quedenfeldt, 1891, by monotypy. Status: valid genus in Alleculinae: Cteniopodini. Note: replacement name for Brachycryptus Quedenfeldt, 1891.

Trisilus Haag-Rutenberg, 1878: 101 [M]. Type species: Trisilusfemoralis Haag-Rutenberg, 1878, by monotypy. Status: junior synonym of Cyphaleus Westwood, 1841 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Matthews (1992: 490).

Trogloderus J.L. LeConte, 1879: 2 [M]. Type species: Trogloderuscostatus J.L. LeConte, 1879, by monotypy. Status: valid genus in Blaptinae: Amphidorini.

Troglogeneion Aalbu, 1985: 541 [N]. Type species: Troglogeneionzapoteca Aalbu, 1985, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Tromosternus Harold, 1876: 130 [M]. Type species: Tromosternushaagi Harold, 1876 (= Gnesishelopioides Pascoe, 1866), by monotypy. Status: junior synonym of Gnesis Pascoe, 1866 in Stenochiinae: Cnodalonini. Synonymy: Lewis (1894: 476).

Tropidopterus Cazurro Ruiz, 1897b: 637 [M]. Type species: Tropidopteruscarinatus Cazurro Ruiz, 1897, by original designation. Status: junior synonym of Adelium W. Kirby, 1819 in Lagriinae: Adeliini. Synonymy: Doyen et al. (1990: 230). Note: Tropidoptère was first used by Blanchard (1845: 35) but it was not formed as a scientific name and is therefore unavailable (ICZN 1999, Article 1.1); the latinized form of this name, Tropidopterus, was used subsequently (e.g., Lacordaire, 1859b: 438) but was not made available to our knowledge until Cazurro Ruiz (1897b: 637) published a combined description of a new genus and single new species (ICZN 1999, Article 12.2.6); since we believe that the combined description by Cazurro Ruiz (1897b : 637) was based on the same concept as the unavailable name “Tropidoptère Blanchard” we follow the intentions of authors in recent literature (e.g., Doyen et al. 1990: 230; Matthews 1998: 777; Matthews and Lawrence 2019: 615) and treat Tropidopterus Cazurro Ruiz, 1897 as a junior synonym of Adelium W. Kirby, 1819.

Tropitrachys Koch, 1955a: 229 [M]. Type species: Trachynotusperegrinator Koch, 1953, by original designation. Status: valid subgenus of Somaticus Hope, 1841 in Pimeliinae: Sepidiini: Trachynotina.

Truncatocamaria Pic, 1922b: 27 [F]. Type species: Camariaspinipes Pic, 1917 (as “Camariaspinifer”), by monotypy. Status: junior synonym of Camaria Lepeletier & Audinet-Serville, 1828 in Stenochiinae: Cnodalonini. Synonymy: Gebien (1942a: 320), Masumoto (1993a: 147).

Truncatoodescelis Kaszab, 1940b: 942, 962 [F]. Type species: Platyscelislongicollis Kraatz, 1884, by original designation. Status: valid subgenus of Oodescelis Motschulsky, 1845 in Blaptinae: Platyscelidini.

Tuberocnodes ​Gearner & Kamiński in Gearner et al., 2021: 8 [M]. Type species: Psammodeshumeralis ​Haag-Rutenberg, 1871, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Molurina.

Tucumana Gebien, 1911b: 604 [F]. Type species [automatic]: Eusteniatenuimembris Fairmaire, 1905, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina. Note: replacement name for Eustenia Fairmaire, 1905; placed in Alleculinae by Chatanay (1915a: 526).

Turcmenicola Bogatchev, 1952: 44 [M]. Type species: Turcmenicolajachontovi Bogatchev, 1952, by monotypy. Status: valid subgenus of Colposcelis Dejean, 1834 in Pimeliinae: Tentyriini.

Turkmenohelops G.S. Medvedev, 1987: 98, 102 [M]. Type species: Zophohelopsbalchanicus G.S. Medvedev & Nepesova, 1985, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Turkonalassus Keskin, Nabozhenko & Alpagut-Keskin, 2017: 727 [M]. Type species: Helopsadimonius Allard, 1876, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Tydeolus Champion, 1884: 37 [M]. Type species: Tydeolusatratus Champion, 1884, by subsequent designation (W.F. Kirby 1885b: 80). Status: valid genus in Pimeliinae: Epitragini.

Tylophloeus Bremer, 1998: 10, 13 [M]. Type species: Hypophlaeusflavipennis Motschulsky, 1860, by original designation. Status: valid subgenus of Corticeus Piller & Mitterpacher, 1783 in Diaperinae: Hypophlaeini.

Tyndarisus Pascoe, 1869: 294 [M]. Type species: Tyndarisuslongitarsus Pascoe, 1869 (= Lepispilusstygianus Pascoe, 1869), by monotypy. Status: junior synonym of Pachycoelia Boisduval, 1835 in Tenebrioninae: Heleini: Cyphaleina. Synonymy: Carter (1926: 119, with Lepispilus Westwood, 1841, a junior synonym of Pachycoelia Boisduval, 1835).

Tynteria Reitter, 1897a: 301 [F]. Type species: Pachychilahumerosa Fairmaire, 1875, by monotypy. Status: junior synonym of Oterophloeus Desbrochers des Loges, 1881 in Pimeliinae: Tentyriini. Synonymy: Reitter (1900c: 89, 143).

Tynthlobia Fairmaire, 1888d: 261 [F]. Type species: Tynthlobiaquadricostata Fairmaire, 1888, by monotypy. Status: junior synonym of Ethmus Haag-Rutenberg, 1873 in Pimeliinae: Sepidiini: Trachynotina. Synonymy: Fairmaire (1891b: 250).

Typhlophloeus Jeannel & Paulian, 1945: 51 [M]. Type species: Typhlophloeuschappuisi Jeannel & Paulian, 1945, by original designation. Status: valid genus in Diaperinae: Hypophlaeini.

Typhluloma Lea, 1912: 475 [N]. Type species: Typhlulomainops Lea, 1912, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Typhlusechus Linell, 1897: 154 [M]. Type species: Typhlusechussingularis Linell, 1897, by original designation. Status: valid genus in Pimeliinae: Stenosini: Typhlusechina.

Typhobia Pascoe, 1869: 279 [F]. Type species: Typhobiafuliginea Pascoe, 1869, by monotypy. Status: junior synonym of Platydema Laporte & Brullé, 1831 in Diaperinae: Diaperini: Diaperina. Synonymy: Champion (1886: 181).

Tyrtaeus Champion, 1913: 76 [M]. Type species: Tyrtaeusrufus Champion, 1913, by original designation. Status: valid genus in Diaperinae: Gnathidiini: Anopidiina.

Ubangia Gebien, 1914e: 54 [F]. Type species: Ubangialatifrons Gebien, 1914, by monotypy. Status: junior synonym of Crypsinous Fairmaire, 1891 in Tenebrioninae: Amarygmini. Synonymy: Ardoin (1964a: 839).

Ucalegon Champion, 1884: 65 [M]. Type species: Ucalegonpulchellus Champion, 1884, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini.

Udebra Reitter, 1896a: 236 [F]. Type species: Udebrahauseri Reitter, 1896 (= Erodiusfimbriatus Ménétriés, 1849), by monotypy. Status: junior synonym of Adavius Mulsant & Rey, 1859 in Blaptinae: Opatrini: Ammobiina. Synonymy: Kaszab (1942: 28).

Uenomisolampidius Masumoto, 1996a: 36 [M]. Type species: Ueonomisolampidiusshunichii Masumoto, 1996, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Uenostrongylium Masumoto, 1999a: 123 [N]. Type species: Cryptobateslaosensis Pic, 1928, by original designation. Status: valid genus in Stenochiinae: Stenochiini.

Uleda Laporte, 1840: 220 [F]. Type species: Ulomadiaperoides Laporte, 1840, by monotypy. Status: valid genus in Tenebrioninae: Ulomini.

Uloma Dejean, 1821: 67 [N]. Type species: Tenebrioculinaris Linnaeus, 1758, by plenary powers (ICZN 1975, Opinion 1039). Status: valid genus and subgenus in Tenebrioninae: Ulomini. Note: placed on the Official List of Generic Names in Zoology (ICZN 1975, Opinion 1039). Note: the name Uloma is a Greek noun οὔλωμα (meaning cicatrisation) and the gender is neuter instead of the previously accepted feminine gender (e.g., ICZN 1975, Opinion 1039; see ICZN 2021, Official Correction 134).

Ulomimus Bates, 1873a: 201 [M]. Type species: Ulomimusindicus Bates, 1873, by monotypy. Status: valid genus in Tenebrioninae: Ulomini. Note: unjustified emendation of the original spelling Ulomimimus, introduced by Rye (1875: 288), in prevailing usage and treated as a justified emendation (ICZN 1999, Article 33.2.3.1).

Ulomina Baudi di Selve, 1876a: 112 [F]. Type species: Ulominacarinata Baudi di Selve, 1876, by monotypy. Status: valid genus in Tenebrioninae: Palorini. Note: redescribed as new by Baudi di Selve (1876b: 235).

Ulomoides Blackburn, 1888: 274 [M]. Type species: Ulomoideshumeralis Blackburn, 1888, by monotypy. Status: valid genus in Diaperinae: Diaperini: Diaperina.

Ulomoides Escalera, 1927: 501 [M]. Type species [automatic]: Crypticusviaticus Fairmaire, 1851, by original designation. Status: senior synonym of Platycrypticus Español, 1952 in Diaperinae: Crypticini. Note: junior homonym of Ulomoides Blackburn, 1888 [Coleoptera: Tenebrionidae: Diaperinae: Diaperini: Diaperina].

Ulomotypus Broun, 1886: 841 [M]. Type species: Ulomotypuslaevigatus Broun, 1886, by monotypy. Status: valid genus in Tenebrioninae: Palorini.

Ulosonia Laporte, 1840: 220 [F]. Type species: Ulomatricorne Laporte, 1840 (= Phaleriatricornis Dalman, 1823), by subsequent designation (Gebien 1940: 786). Status: junior synonym of Hypogena Dejean, 1834 in Tenebrioninae: Triboliini. Synonymy: Jacquelin du Val (1857: 148).

Ulus Horn, 1870: 349, 358 [M]. Type species: Blapstinuscrassus J.L. LeConte, 1851, by subsequent designation (R. Lucas 1920: 665). Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Umslatus Péringuey, 1899: 312 [M]. Type species: Umslatusagilis Péringuey, 1899, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Uniungulum Koch, 1962b: 113 [N]. Type species: Uniungulumhoeschi Koch, 1962, by original designation. Status: valid genus in Pimeliinae: Sepidiini: Hypomelina.

Upembarus Koch, 1956a: 220 [M]. Type species: Upembarussaegeri Koch, 1956, by original designation. Status: valid genus and subgenus in Blaptinae: Platynotini: Platynotina.

Upinella Mulsant, 1857: 17 [F]. Type species: Alleculaaterrima Rosenhauer, 1847, by monotypy. Status: valid genus and subgenus in Alleculinae: Alleculini: Alleculina.

Upis Fabricius, 1792: 515 [F]. Type species: Attelabusceramboides Linnaeus, 1758, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Uptona G.S. Medvedev & Lawrence, 1986: 582 [F]. Type species: Uptonapallida G.S. Medvedev & Lawrence, 1986, by original designation. Status: valid genus in Diaperinae: Hyociini: Uptonina.

Uria Gistel, 1848a: x [F]. Type species [automatic]: Tenebriocadaverinus Fabricius, 1792, by subsequent designation (Westwood 1838: 32). Status: junior synonym of Phaleria Latreille, 1802 in Diaperinae: Phaleriini. Note: unnecessary replacement name for Phaleria Latreille, 1802.

Uriela Reitter, 1887b: 518 [F]. Type species: Podhomalafausti Kraatz, 1881, by monotypy. Status: junior synonym of Podhomala Solier, 1836 in Pimeliinae: Pimeliini. Synonymy: Sénac (1888: lv).

Urielina Reitter, 1888: 331 [F]. Type species: Podhomalanitida Baudi di Selve, 1876, by monotypy. Status: valid subgenus of Podhomala Solier, 1836 in Pimeliinae: Pimeliini.

Uroblaps Motschulsky, 1860c: 530 [F]. Type species: Blaps producta Brullé, 1832 (= Blaps lusitanica Herbst, 1799), by subsequent designation (Nabozhenko 2008: 36). Status: junior synonym of Blaps Fabricius, 1775 in Blaptinae: Blaptini: Blaptina. Synonymy: Gemminger in Gemminger and Harold (1870: 1860)

Uroplatopsis Champion, 1889: 53 [F]. Type species: Uroplatopsisimitator Champion, 1889, by subsequent designation (R. Lucas 1920: 666). Status: valid genus in Lagriinae: Lagriini: Statirina.

Uroprosodes Reitter, 1909a: 119 [M]. Type species: Prosodescostifera Kraatz, 1886, by original designation. Status: valid subgenus of Prosodes Eschscholtz, 1829 in Blaptinae: Blaptini: Prosodina.

Uyttenboogaartia Koch, 1943a: 592, 595 [F]. Type species: Hegetercribricollis Brullé, 1839, by original designation. Status: valid genus in Pimeliinae: Tentyriini.

Uzagaria Ancey, 1881: 509 [F]. Type species: Uzagariapubens Ancey, 1881, by monotypy. Status: junior synonym of Emmalus Erichson, 1843 in Blaptinae: Opatrini: Ammobiina. Synonymy: Gebien (1910b: 306, as Emmallus).

†Vabole Alekseev & Nabozhenko, 2015: 128 [F]. Type species: Vaboletriplehorni Alekseev & Nabozhenko, 2015, by original designation. Status: valid genus in Tenebrioninae: Palorini. Note: described from Eocene Baltic amber.

Vacronus Casey, 1907: 501, 508 [M]. Type species: Vacronustenuicornis Casey, 1907, by original designation. Status: junior synonym of Alaephus Horn, 1870 in Pimeliinae: Vacronini. Synonymy: Doyen and Lawrence (1979: 350).

Vadalus Mulsant & Rey, 1853b: 150 [M]. Type species: Pedinuspunctulatus Mulsant & Rey, 1853, by monotypy. Status: junior synonym of Pedinus Latreille, 1797 in Blaptinae: Pedinini: Pedinina. Synonymy: Gemminger in Gemminger and Harold (1870: 1918), Kamiński and Iwan (2017: 599).

Valdivium Matthews, 1998: 709, 721 [N]. Type species: Adeliumsulcatulum Fairmaire & Germain, 1860, by original designation. Status: valid genus in Lagriinae: Adeliini.

Vaniosus Kulzer, 1956b: 896 [M]. Type species: Vaniosusparadoxus Kulzer, 1956, by original designation. Status: valid genus in Pimeliinae: Evaniosomini.

Vansonium Koch, 1950b: 354 [N]. Type species: Vansoniumbushmanicum Koch, 1950, by original designation. Status: valid genus in Pimeliinae: Cryptochilini: Vansoniina.

Varogeton Bremer, 2014a: 37, 80 [M]. Type species: Dietysussubannulipes Pic, 1923, by original designation. Status: valid subgenus of Amarygmus Dalman, 1823 in Tenebrioninae: Amarygmini.

Vernayella Koch, 1958: 129 [F]. Type species: Vernayellanoctivaga Koch, 1958, by original designation. Status: valid subgenus of Caenocrypticus Gebien, 1920 in Pimeliinae: Caenocrypticini.

Vieta Laporte, 1840: 196 [F]. Type species: Sepidium vestitum Guérin-Méneville, 1831, by subsequent designation (Hope 1841: 116). Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Vietnalia Novák, 2021: 440 [F]. Type species: Vietnaliacatcatica Novák, 2021, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Vietomorpha Fairmaire, 1887a: 186 [F]. Type species: Vietomorphafoveipennis Fairmaire, 1887, by monotypy. Status: valid genus in Pimeliinae: Sepidiini: Sepidiina.

Viettagona G.S. Medvedev & Merkl, 2003: 317, 328 [F]. Type species: Viettagonavietnamensis G.S. Medvedev & Merkl, 2003, by original designation. Status: valid genus in Blaptinae: Blaptini: Gnaptorinina.

Villiersia Gridelli, 1951: 219, 227 [F]. Type species: Tenebrioclypealis Gebien, 1920, by original designation. Status: senior synonym of Gridellia Kammerer, 2006 in Tenebrioninae: Tenebrionini. Note: junior homonym of Villiersia d’Orbigny, 1837 [Mollusca].

Viriathus Fairmaire, 1902b: 339 [M]. Type species: Viriathusstrigipennis Fairmaire, 1902, by monotypy. Status: valid genus in Alleculinae: Alleculini: Gonoderina.

Vizcainyx Aalbu & Smith, 2020: 198 [M]. Type species: Vizcainyxandrewsi Aalbu & Smith, 2020, by original designation. Status: valid genus in Pimeliinae: Edrotini.

Vutsimus Péringuey, 1899: 308 [M]. Type species: Vutsimuspraetorius Péringuey, 1899, by subsequent designation (Gebien 1943: 922). Status: valid genus in Tenebrioninae: Amarygmini.

Wahlbergylium Ferrer, 2011: 141 [N]. Type species: Wahlbergyliumviklundi Ferrer, 2011, by original designation. Status: junior synonym of Argutiolana Robiche, 2001 in Stenochiinae: Cnodalonini. Synonymy: Robiche (2019a: 97).

Wallardilagria Pic, 1910: 74 [F]. Type species: Heterogriapallidicolor Pic, 1910, by monotypy. Status: junior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Synonymy: Merkl (2004: 285).

Warchalowskiellus Iwan, 1998a: 60 [M]. Type species: Trigonopuslongulus Mulsant & Rey, 1853, by original designation. Status: junior synonym of Schelodontes Koch, 1956 in Blaptinae: Platynotini: Platynotina. Synonymy: Iwan and Kamiński (2014: 171).

Waterhousia Skopin, 1973: 109, 110 [F]. Type species: Trigonoscelislongipes C.O. Waterhouse, 1889, by original designation. Status: valid genus in Pimeliinae: Pimeliini.

Wattadelium Emberson, 2000: 24 [N]. Type species [automatic]: Edalusopacus Broun, 1893, by subsequent designation (R. Lucas 1920: 255). Status: valid genus in Lagriinae: Adeliini. Note: replacement name for Edalus Broun, 1893.

Wattiana Matthews & Lawrence, 2005: 537 [F]. Type species: Wattianagreensladei Matthews & Lawrence, 2005, by original designation. Status: valid genus in Pimeliinae: Cnemeplatiini: Thorictosomatina.

Wattius Kaszab, 1982b: 50 [M]. Type species: Calymmuscucullatus Pascoe, 1871, by original designation. Status: valid genus in Tenebrioninae: Toxicini: Dysantina.

Weisea Semenov, 1891: 370 [F]. Type species: Weiseasabulicola Semenov, 1891, by monotypy. Status: valid genus in Blaptinae: Opatrini: Ammobiina.

Wolladrus Iwan & Kamiński, 2016: 483 [M]. Type species [automatic]: Hadrusalpinus Wollaston, 1854, by subsequent designation (R. Lucas 1920: 313). Status: valid genus in Blaptinae: Opatrini: Opatrina. Note: replacement name for Hadrus Wollaston, 1854.

Xanthalia Fairmaire, 1894f: 395 [F]. Type species [automatic]: Xanthiacurticollis Fairmaire, 1893, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina. Note: replacement name for Xanthia Fairmaire, 1893; new placement [OM], previously included in Lagriinae: Lagriini: Lagriina.

Xanthia Fairmaire, 1893b: 31 [F]. Type species: Xanthiacurticollis Fairmaire, 1893, by monotypy. Status: senior synonym of Xanthalia Fairmaire, 1894 in Lagriinae: Lagriini: Statirina. Note: junior homonym of Xanthia Hübner, 1813 [Lepidoptera].

Xanthicles Champion, 1886: 231 [M]. Type species: Xanthiclescaraboides Champion, 1886, by subsequent designation (Gebien 1941: 815). Status: valid genus in Lagriinae: Goniaderini.

Xanthobates Gebien, 1928: 170, 185 [M]. Type species: Xanthobatesflavus Gebien, 1928, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Xanthohelops Nabozhenko, 2006: 822 [M]. Type species: Xanthohelopskarakumicus Nabozhenko & G.S. Medvedev, 2006, by original designation. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Xanthomus Mulsant, 1854: 302 [M]. Type species: Helopspallidus Curtis, 1830, by monotypy. Status: valid genus in Tenebrioninae: Helopini: Cylindrinotina.

Xanthothopeia Mäklin, 1867: 223 [F]. Type species: Xanthothopeiarufipennis Mäklin, 1867, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1948: 519).

Xanthothopia Gemminger in Gemminger and Harold, 1870: 2001 [F]. Type species [automatic]: Xanthothopeiarufipennis Mäklin, 1867, by monotypy. Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Note: unjustified emendation of Xanthothopeia Mäklin, 1867, not in prevailing usage.

Xantusiella Kaszab, 1941a: 4, 18 [F]. Type species: Xantusiellacrenulata Kaszab, 1941, by original designation. Status: valid genus in Stenochiinae: Cnodalonini.

Xenius Champion, 1886: 224 [M]. Type species: Xeniusscabripennis Champion, 1886, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Xenocera Borchmann, 1936: 18, 116 [F]. Type species: Lagriocerafeai (as “feae”) Borchmann, 1911, by original designation. Status: senior synonym of Xenocerogria Merkl, 2007 in Lagriinae: Lagriini: Lagriina. Note: junior homonym of Xenocera Broun, 1881 [Coleoptera: Ptinidae].

Xenocerogria Merkl, 2007: 269 [F]. Type species [automatic]: Lagriocerafeai (as “feae”) Borchmann, 1911, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina. Note: replacement name for Xenocera Borchmann, 1936.

Xenogena Borchmann, 1936: 22, 211 [F]. Type species: Adynatacrinita Borchmann, 1915, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Xenogloeus Wollaston, 1861: 251 [M]. Type species: Xenogloeuspolitus Wollaston, 1861, by monotypy. Status: valid genus in Tenebrioninae: Triboliini.

Xenolagria Merkl, 1987: 124, 126 [F]. Type species: Lagriatincta Blackburn, 1889, by original designation. Status: valid genus in Lagriinae: Lagriini: Lagriina.

Xenostethus Bates, 1868: 321 [M]. Type species: Xenostethuslacordairii Bates, 1868, by monotypy. Status: valid genus in Lagriinae: Lagriini: Statirina.

Xenostira Borchmann, 1921: 217, 221 [F]. Type species: Xenostiragiraffa Borchmann, 1921, by original designation. Status: valid subgenus of Statira Lepeletier & Audinet-Serville, 1828 in Lagriinae: Lagriini: Statirina.

Xenotermes Wasmann, 1896: 616 [M]. Type species: Xenotermesfeai (as “feae”) Wasmann, 1896, by monotypy. Status: valid genus in Tenebrioninae: Rhysopaussini.

Xenus Péringuey, 1899: 255 [M]. Type species: Xenustricorniger Péringuey, 1899, by monotypy. Status: senior synonym of Aphrotus Péringuey, 1904 in Pimeliinae: Epitragini. Note: junior homonym of Xenus Kaup, 1829 [Aves].

Xerolinus Ivie & Hart, 2016: 470 [M]. Type species: Diastolinussallei Mulsant & Rey, 1859, by original designation. Status: valid genus in Blaptinae: Opatrini: Blapstinina.

Xyloborus Motschulsky, 1858a: 64 [M]. Type species: Xyloboruscrenipennis Motschulsky, 1858, by monotypy. Status: senior synonym of Rhipidandrus J.L. LeConte, 1862 in Tenebrioninae: Bolitophagini. Synonymy: Merkl and Kompantzeva (1996: 91). Note: although Schwarz and Barber (1914: 175) pointed out that the name Xyloborus was made available for the first time by Motschulsky (1858: 64) there are few occurrences of this name in the literature; reversal of precedence cannot be used to conserve usage of the broadly used name Rhipidandrus J.L. LeConte, 1862 since Xyloborus was used as valid after 1899 (e.g., Swezey 1942: 167); an application to the ICZN is necessary to conserve usage Rhipidandrus J.L. LeConte, 1862; the older names Xyloborus Kirby & Spence, 1828 and Xyloborus Dejean, 1834 [Coleoptera] are unavailable because they were published before 1931 without a description, a definition or an indication (ICZN 1999, Article 12.1).

Xylochus Broun, 1880: 396 [M]. Type species: Xylochustibialis Broun, 1880, by subsequent designation (Watt 1992: 24). Status: valid genus in Alleculinae: Alleculini: incertae sedis.

Xylopinus J.L. LeConte, 1862: 230 [M]. Type species: Tenebrioanthracinus Knoch, 1801 (= Tenebriosaperdoides G.-A. Olivier, 1795), by subsequent designation (Gebien 1941: 336). Status: valid genus in Stenochiinae: Cnodalonini.

Xysta Eschscholtz, 1829: 9 [F]. Type species: Eleodesgravida Eschscholtz, 1829, by subsequent designation (Hope 1841: 124). Status: senior synonym of Steneleodes Blaisdell, 1909 in Blaptinae: Amphidorini. Synonymy: Smith and Johnston in Bousquet et al. (2018: 166). Note: junior homonym of Xysta Meigen, 1824 [Diptera].

Xystronia Solier, 1835a: 238 [F]. Type species: Xystroniacoerulea Solier, 1835, by monotypy. Status: valid subgenus of Lystronychus Latreille, 1829 in Alleculinae: Alleculini: Xystropodina.

Xystropus Solier, 1835a: 241 [M]. Type species: Xystropuspilosus Solier, 1835, by monotypy. Status: valid genus in Alleculinae: Alleculini: Xystropodina. Note: see Bousquet et al. (2015: 139) for comments regarding the species originally included in this genus.

Yamatotakeru Ando, 2015: 385 [M]. Type species: Ischnodactylusloripes Lewis, 1894, by original designation. Status: valid genus in Diaperinae: Diaperini: Diaperina.

†Yantaroxenos Nabozhenko, Kirejtshuk & Merkl, 2016c: 564 [M]. Type species: Yantaroxenoscolydioides Nabozhenko, Kirejtshuk & Merkl, 2016, by original designation. Status: valid genus in Lagriinae: Belopini. Note: described from Eocene Baltic amber.

Yarranum Matthews, 1998: 708, 734 [N]. Type species: Seirotranacrenicollis Pascoe, 1869, by original designation. Status: valid genus in Lagriinae: Adeliini.

Zabroideus Fairmaire, 1894c: 219 [M]. Type species: Zabroideuspinguis Fairmaire, 1894, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Zadenos Laporte, 1840: 210 [M]. Type species: Opatrumlongipalpe Wiedemann, 1823, by monotypy. Status: valid subgenus of Selenepistoma Dejean, 1834 in Blaptinae: Dendarini: Melambiina. Note: this name was recently treated as a valid genus (Kamiński 2015: 531); however, the older available genus name Selenepistoma Dejean, 1834 has priority and therefore Zadenos is downgraded to a valid subgenus of Dejean’s name; new status.

Zaleucus Champion, 1892: 491 [M]. Type species [automatic]: Zamolxisdilatatus Champion, 1884, by monotypy. Status: valid subgenus of Pelecyphorus Solier, 1836 in Pimeliinae: Asidini. Note: replacement name for Zamolxis Champion, 1884.

Zambesmia Bouchard & Bousquet, new subgenus [F]. Type species: Macropodachiyakensis Kuntzen, 1916, by present designation. Status: valid subgenus of Adesmia Fischer, 1822 in Pimeliinae: Adesmiini. Note: Koch (1944b: 149) introduced the new subgenus name Zambesmia for three nominal species, but unfortunately did not designate a type species; the subgenusZambesmia, which has been treated as valid since 1944, is therefore unavailable (ICZN 1999, Article 13.3); we hereby make the name available by selecting Macropodachiyakensis Kuntzen, 1916 as type species and referring to Koch (1944b: 149) for the character states that characterise and differentiate Zambesmia.

Zamolxis Champion, 1884: 70 [M]. Type species: Zamolxisdilatatus Champion, 1884, by monotypy. Status: senior synonym of Zaleucus Champion, 1892 in Pimeliinae: Asidini. Note: junior homonym of Zamolxis Stål, 1865 [Hemiptera].

Zarudnionymus Semenov-Tjan-Shansky & Bogatchev, 1947: 175 [M]. Type species: Zarudnionymuspersis Semenov-Tjan-Shansky & Bogatchev, 1947 (= Adelostomagrande Haag-Rutenberg, 1879), by monotypy. Status: valid subgenus of Adelostoma Duponchel, 1827 in Pimeliinae: Adelostomini.

Zeadelium Watt, 1992: 32 [N]. Type species: Adeliumlentum Broun, 1880, by original designation. Status: valid genus in Lagriinae: Adeliini.

Ziaelas Fairmaire, 1892e: cx [M]. Type species: Ziaelasinsolitus Fairmaire, 1892, by monotypy. Status: valid genus in Tenebrioninae: Amarygmini.

Zidalus Mulsant & Rey, 1853b: 71 [M]. Type species: Opatrinuscorvinus Mulsant & Rey, 1853, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Zizu Novák, 2019b: 186 [M]. Type species: Zizukejvali Novák, 2019, by original designation. Status: valid genus in Alleculinae: Alleculini: Alleculina.

Zodinus Mulsant & Rey, 1853b: 90 [M]. Type species: Opatrinusservus Mulsant & Rey, 1853, by subsequent designation (Koch 1956a: 93). Status: junior synonym of Zidalus Mulsant & Rey, 1853 in Blaptinae: Platynotini: Platynotina. Synonymy and First Reviser action (Zodinus Mulsant & Rey, 1853 versus Zidalus Mulsant & Rey, 1853) is Iwan (1995b: 362).

Zolodinus Blanchard, 1853: 159 [M]. Type species: Zolodinuszelandicus Blanchard, 1853, by monotypy. Status: valid genus in Zolodininae.

Zomedes Watt, 1992: 25 [M]. Type species: Zomedesborealis Watt, 1992, by original designation. Status: valid genus in Alleculinae: Alleculini: incertae sedis.

Zophius Dejean, 1834: 189 [M]. Type species: Helopsrufopictus Wiedemann, 1823, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Zophobas Dejean, 1834: 204 [M]. Type species: Helopsmorio Fabricius, 1777 (= Tenebrioatratus Fabricius, 1775), by subsequent designation (Motschulsky 1872: 26). Status: valid genus and subgenus in Tenebrioninae: Tenebrionini. Note: according to Matthews and Lawrence (2019: 628) more research is needed to establish the correct placement of this genus within the subfamily Tenebrioninae.

Zophodes Fåhraeus, 1870: 298 [M]. Type species: Zophodestristis Fåhraeus, 1870, by monotypy. Status: valid genus in Blaptinae: Platynotini: Platynotina.

Zophohelops Reitter, 1902a: 221 [M]. Type species [automatic]: Euryhelopstiro Reitter, 1902, by subsequent designation (Gebien 1943: 424). Status: valid genus and subgenus in Tenebrioninae: Helopini: Cylindrinotina. Note: replacement name for Euryhelops Reitter, 1902.

Zophondrus Nabozhenko, 2014: 240 [M]. Type species: Zophohelopsiranensis Nabozhenko, 2014, by original designation. Status: valid subgenus of Zophohelops Reitter, 1902 in Tenebrioninae: Helopini: Cylindrinotina.

Zophophilus Fairmaire, 1881b: 359 [M]. Type species: Zophophiluscurticornis Fairmaire, 1881, by monotypy. Status: valid genus in Stenochiinae: Cnodalonini.

Zophoserodius Reitter, 1914a: 58, 60 [M]. Type species: Erodiuszophosoides Allard, 1865, by subsequent designation (Löbl et al. 2008a: 43). Status: valid subgenus of Erodius Fabricius, 1775 in Pimeliinae: Erodiini.

Zophosis Latreille, 1802: 167 [F]. Type species: Erodiustestudinarius Fabricius, 1781, by monotypy. Status: valid genus and subgenus in Pimeliinae: Zophosini.

Zophosodactylus Koch, 1962b: 146 [M]. Type species: Protodactylussanctaemariae Koch, 1962, by original designation. Status: junior synonym of Protodactylus Koch, 1952 in Pimeliinae: Zophosini. Synonymy: Penrith (1981c: 129).

Zoutpansbergia Koch, 1956a: 388 [F]. Type species: Zoutpansbergiaserricostata Koch, 1956, by monotypy. Status: valid genus in Blaptinae: Dendarini: Melambiina. Note: combined description of new genus-group taxon and new species (ICZN 1999, Article 13.4).

Zuercheria Reitter, 1908: 134 [F]. Type species: Zuercheriamatthiesseni Reitter, 1908, by subsequent designation (Löbl et al. 2008a: 43). Status: junior synonym of Strongylium W. Kirby, 1819 in Stenochiinae: Stenochiini. Synonymy: Gebien (1911b: 590).

Zygas Pascoe, 1866a: 487 [M]. Type species: Eurychoracimicoides Quensel, 1806, by original designation. Status: junior synonym of Lycanthropa J. Thomson, 1860 in Pimeliinae: Adelostomini. Synonymy: Haag-Rutenberg (1875b: 61–64).

Zypoetes Champion, 1893a: 532 [M]. Type species: Zypoetesepieroides Champion, 1893, by monotypy. Status: valid genus in Phrenapatinae: Penetini.

Appendix 1

List of unavailable genus-group names in Tenebrionidae Latreille, 1802. Each entry includes the name, its usage in the literature (or one example when the name was used more than once), and the reason for treating it as unavailable. See the Methods section for additional information. The incorrect subsequent spellings below are not in prevailing usage.

Acanthodactylus; Escalera, 1914: 347; incorrect subsequent spelling for Acontodactylus Desbrochers des Loges, 1894.

Acanthopus; Latreille, 1829a: 38; incorrect subsequent spelling for Accanthopus Dejean, 1821.

Achaemenus; G.S. Medvedev and Iwan, 2006: 618; incorrect subsequent spelling for Achaemenes Bogatchev, 1949.

Acilagria; Borchmann, 1916a: 90; published before 1931 without a description, a definition, or an indication.

Acmaeus; Gebien, 1910b: 304; incorrect subsequent spelling for Acmoeus Fåhraeus, 1870.

Acrothymus; Gemminger in Gemminger and Harold, 1870: 2009; incorrect subsequent spelling for Arcothymus Pascoe, 1866.

Acthosus; Carter, 1906: 251; incorrect subsequent spelling for Achthosus Pascoe, 1863.

Acysba; Villa and Villa, 1833: 18; published before 1931 without a description, a definition, or an indication.

Adosagria; Pic, 1955: 179, 180; incorrect subsequent spelling for Adosogria Borchmann, 1936.

Aediotorix; Gebien, 1905: 258; incorrect subsequent spelling for Aediatorix Bates, 1868.

Aemyone; Ferrer and Delatour, 2007: 275, 276; incorrect subsequent spelling for Aemymone Bates, 1868.

Afghanopachya; Anonymous in Staff of BIOSIS, 1981: 283; incorrect subsequent spelling for Afghanopachys Kwieton, 1978.

Afghanprosodes; Schuster, 1936: 197; published after 1930 without a type species designation.

Agapetus; Dejean, 1834: 212; published before 1931 without a description, a definition, or an indication.

Agroecus; Chatanay, 1915: 508; incorrect subsequent spelling for Agraecus Fairmaire, 1900.

Agymnonix; Kaszab, 1980b: 320; incorrect subsequent spelling for Agymnonyx Gebien, 1921.

Akixa; Guérin-Méneville, 1834: 13; incorrect subsequent spelling for Akis Herbst, 1799.

Alcione; Reitter, 1897a: 300; incorrect subsequent spelling for Alcinoe Ménétriés, 1849.

Alcmaeonis; Blackburn, 1893b: 133; incorrect subsequent spelling for Alcmeonis Bates, 1868.

Alcyone; Kraatz, 1865: 80; incorrect subsequent spelling for Alcinoe Ménétriés, 1849.

Aletha; Poole and Gentili, 1996: 425; incorrect subsequent spelling for Alethia Champion, 1888.

Alienolonyx; Bremer, 2019: 60; alternative original spelling of Alienoplonyx Bremer, 2019, herein rejected.

Allegoria; Gemminger in Gemminger and Harold, 1870: 1959; incorrect subsequent spelling for Alegoria Laporte, 1840.

Allolagria; Sharp, 1922: 144; incorrect subsequent spelling for Allogria Borchmann, 1916.

Alphitoplogus; Motschulsky, 1873: 474; incorrect subsequent spelling for Alphitophagus Stephens, 1832.

Amacarus; Dejean, 1834: 212; published before 1931 without a description, a definition, or an indication.

Amarigmus; Hope, 1841: 75; incorrect subsequent spelling for Amarygmus Dalman, 1823.

Amathodes; Erichson, 1845a: 114; incorrect subsequent spelling for Amatodes Dejean, 1834.

Amblychara; Reitter, 1900c: 94; incorrect subsequent spelling for Amblycara Fairmaire, 1893.

Amblycyphrus; Aalbu et al. 1995: 481; incorrect subsequent spelling for Amblycyphus Motschulsky, 1870.

Ammophthorus; Jacquelin du Val, 1861: 288; incorrect subsequent spelling for Ammophtorus Lacordaire, 1859.

Amophorus; Dejean, 1836: 203; incorrect subsequent spelling for Ammophorus Guérin-Méneville, 1831.

Amozoum; Reitter, 1914a: 45, 50; incorrect subsequent spelling for Ammozoum Semenov, 1891.

Amphitrix; Koch, 1956a: 49; incorrect subsequent spelling for Amphithrix Español, 1953.

Amphydora; Dejean, 1834: 210; incorrect subsequent spelling for Amphidora Eschscholtz, 1829.

Amphysus; Dejean, 1834: 189; published before 1931 without a description, a definition, or an indication.

Amplypteraca; Mas-Peinado et al., 2018: 531, 543; alternative original spelling of Amblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 2018, herein rejected.

Amyone; Ferrer and Delatour, 2007: 279; incorrect subsequent spelling for Aemymone Bates, 1868.

Anadesis; Laporte, 1840: 185; incorrect subsequent spelling for Anodesis Solier, 1834.

Anaemia; Horn, 1870: 377; incorrect subsequent spelling for Anemia Laporte, 1840.

Anarmastodera; Schawaller, 2010: 285; incorrect subsequent spelling for Anarmostodera Fairmaire, 1897.

Anchophtalmus; Lesne, 1922: 703; incorrect subsequent spelling for Anchophthalmus Gerstaecker, 1854.

Aniosis; Ferreira, 1967: 741; incorrect subsequent spelling for Anisosis Deyrolle, 1867.

Anisochara; Löbl et al. 2008b: 309; incorrect subsequent spelling for Anisocara Gebien, 1925.

Anisocheira; Dejean, 1834: 197; published before 1931 without a description, a definition, or an indication.

Anisocrepis; Dejean, 1834: 198; published before 1931 without a description, a definition, or an indication.

Anodesia; Agassiz, 1846a: 11; incorrect subsequent spelling for Anodesis Solier, 1834.

Anoedus; Blanchard, 1845: 35; incorrect subsequent spelling for Anaedus Blanchard, 1842.

Anomaearthrum; Aalbu et al., 2002: 501; incorrect subsequent spelling for Anomoearthrum Mäklin, 1867.

Anomalipes; Guérin-Méneville, 1831b: pl. 29; incorrect original spelling, emended to Anomalipus Guérin-Méneville, 1831, under prevailing usage.

Anthcarohelops; Nabozhenko, 2019: 7; incorrect subsequent spelling for Anthracohelops Haupt, 1950.

Antocera; Ragusa, 1898: 121; incorrect subsequent spelling for Autocera Wollaston, 1857.

Apatocerus; Schawaller, 2013: 68; incorrect subsequent spelling for Apistocerus Fairmaire, 1899.

Apaxo; Fairmaire, 1877b: 167; incorrect subsequent spelling for Anaxo Bates, 1868.

Aphthora; Champion, 1895b: 148; incorrect subsequent spelling for Aphtora Bates, 1872.

Aplocheirus; Macquart, 1850: 191; published before 1931 without a description, a definition or an indication.

Archeoglenes; Doyen and Lawrence, 1979: 333; incorrect subsequent spelling for Archaeoglenes Broun, 1893.

Arcolymus; Gebien, 1942a: 745; incorrect subsequent spelling for Arcothymus Pascoe, 1866.

Argasidius; Gebien, 1937a: 674; incorrect subsequent spelling for Argasidus Péringuey, 1899.

Argenis; Burmeister, 1875: 463; incorrect subsequent spelling for Aryenis Bates, 1868.

Arrhenoplitis; Gemminger in Gemminger and Harold, 1870: 1949; incorrect subsequent spelling for Arrhenoplita W. Kirby, 1837.

Arthrocomus; Gebien, 1911b: 614; incorrect subsequent spelling for Arthroconus Solier, 1851.

Arthrodactyla; Laporte, 1840: 211; incorrect subsequent spelling for Athrodactyla Klug, 1833.

Ascalabos; Fairmaire, 1894f: 395; incorrect subsequent spelling for Ascalabus Fairmaire, 1893.

Asdidius; Rye, 1870: 268; incorrect subsequent spelling for Asididius Fairmaire, 1869.

Asidax; Gistel, 1848b: 149; incorrect subsequent spelling for Asida Latreille, 1802.

Aspicephalus; Motschulsky, 1839: 63; incorrect original spelling, emended to Aspidocephalus Motschulsky, 1839, under prevailing usage.

Aspidus; Gebien, 1910b: 298; incorrect subsequent spelling for Aspidius Mulsant & Rey, 1859.

Astenochirus; R. Lucas, 1920: 123; incorrect subsequent spelling for Asthenochirus Fairmaire, 1885

Astenorhinus; Fairmaire, 1894g: 664; incorrect subsequent spelling for Asthenochirus Fairmaire, 1885.

Asthenorhinus; Kolbe, 1897b: 611; incorrect subsequent spelling for Asthenochirus Fairmaire, 1885.

Asyleplus; Gebien, 1943: 922; incorrect subsequent spelling for Asyleptus Péringuey, 1896.

Batulinus; Papp, 1961: 105; incorrect subsequent spelling for Batulius J.L. LeConte, 1851.

Beplegenes; Carter, 1926: 128; incorrect subsequent spelling for Blepegenes Pascoe, 1868.

Biophanes; Marseul, 1857: 117; incorrect subsequent spelling for Bioplanes Mulsant, 1854.

Blapicauda; Ren et al., 2000: 48; published after 1930 without a type species designation.

Blapiplanula; Ren et al., 2000: 49; published after 1930 without a type species designation.

Blapsibreva; Ren et al., 2000: 49; published after 1930 without a type species designation.

Blaptinus; Latreille, 1829a: 21; incorrect subsequent spelling for Blapstinus Dejean, 1821.

Blaptyscelis; Koch, 1965: 127; published after 1930 without a type species designation.

Blaptyscellis; Koch in Pierre, 1962: 213; published after 1930 without a type species designation.

Brachicula; Fairmaire, 1906: 278; alternative original spelling of Brachycula, rejected by Bousquet et al. (2015: 143).

Brachydium; Kaszab, 1939b: 187; incorrect subsequent spelling for Brachyidium Fairmaire, 1883.

Brachypophloeus; Alluaud, 1899: 342; incorrect subsequent spelling for Brachypophlaeus Fairmaire, 1897.

Bradytes; Dejean, 1834: 182; published before 1931 without a description, a definition, or an indication.

Bucerus; Dejean, 1834: 203; published before 1931 without a description, a definition, or an indication.

Bysacnus; Fairmaire, 1897f: 124; incorrect subsequent spelling for Byzacnus Pascoe, 1866.

Cacloplesia; Pic, 1935: 97; incorrect subsequent spelling for Cacoplesia Fairmaire, 1898.

Calliosis; Deyrolle, 1867: 81; original spelling of Calosis, corrected in the same work.

Calobomon; Reitter, 1900c: 93; incorrect subsequent spelling for Calobamon Kraatz, 1865.

Calobosca; Champion, 1895b: 170; incorrect subsequent spelling for Calabosca Fairmaire, 1894.

Calymmaphorus; Solier, 1841a: 209, 245; incorrect original spelling for Calymmophorus Solier, 1841.

Calyptosis; Maseul, 1857: 111; incorrect original spelling for Calyptopsis Solier, 1835.

Camariocropteron; Gebien, 1942a: 332; incorrect subsequent spelling for Camariocropterum Pic, 1920.

Camarioides; Seidlitz, 1900: 277; incorrect original spelling for Camariodes Fairmaire, 1869.

Cameria; Laporte, 1840: 231; incorrect subsequent spelling for Camaria Lepeletier & Audinet-Serville, 1828.

Campanotiphilus; Carter, 1926: 148; incorrect subsequent spelling for Camponotiphilus Lea, 1914.

Caragonia; Wollaston, 1861: 206; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Caribanoisis; Nabozhenko et al., 2016d: 569 [header]; as pointed out by Bousquet et al. (2018: 136), this is an error by the publisher for Caribanosis Nabozhenko, Kirejtshuk, Merkl, Varela, Aalbu & Smith, 2016.

Cataphronetes; Imhoff, 1856: 239; incorrect subsequent spelling for Cataphronetis P.H. Lucas, 1846.

Catonus; Kaszab, 1968: 395; incorrect subsequent spelling for Catomus Allard, 1876.

Centronipus; Macquart, 1850: 188; incorrect subsequent spelling for Centronopus Solier, 1848.

Ceracostira; Pic, 1955: 185; incorrect subsequent spelling for Coracostira Fairmaire, 1899.

Chardiochianalus; Kaszab, 1940a: 202; alternative original spelling of Cardiochianalus Kaszab, 1940, rejected by Kaszab (1975a: 4).

Charinotus; Dejean, 1834: 206; published before 1931 without a description, a definition or an indication.

Charistela; Kolbe, 1897b: 620; incorrect subsequent spelling for Caristela Fairmaire, 1894.

Charitotheca; Agassiz, 1846: 78; proposed as an unjustified emendation for the unavailable name Chariotheca Dejean, 1834; published before 1931 without a description, a definition, or an indication.

Chatanayas; Anonymous in Commonwealth Institute of Entomology, 1960: 393; incorrect subsequent spelling for Chatanayus Ardoin, 1957.

Chelenodus; Berthold, 1827: 369; published before 1931 without a description, a definition, or an indication.

Chemolamus; Fairmaire, 1895b: 447, 448; incorrect subsequent spelling for Chemolanus Bates, 1879.

Chilenodus: Agassiz, 1846a: 35; proposed as a replacement name for the unavailable name Chelenodus Berthold, 1827; published before 1931 without a description, a definition, or an indication.

Chiron; Villa and Villa, 1833: 18; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Cholopus Agassiz, 1846b: 83; proposed as a replacement name for the unavailable name Cholipus Dejean, 1834; published before 1931 without a description, a definition, or an indication.

Chorodes; Cazurro Ruiz, 1895: 730; incorrect subsequent spelling for Chaerodes White, 1846.

Chremolamus; Fairmaire, 1884d: 74; incorrect subsequent spelling for Chemolanus Bates, 1879.

Chromomoea; Carter, 1915a: 59; incorrect subsequent spelling for Chromomaea Pascoe, 1866.

Cichillus; Bertkau, 1890: 257; incorrect subsequent spelling for Dichillus Jacquelin du Val, 1860.

Cilibe; Dejean, 1836: 208; incorrect subsequent spelling for Celibe Boisduval, 1835.

Clinocramon; Péringuey, 1904: 297; incorrect subsequent spelling for Clinocranion Solier, 1843.

Clinocranium; Agassiz, 1846a: 39; incorrect subsequent spelling for Clinocranion Solier, 1843.

Cnecozochara; Ogloblin and Znojko, 1950: 57; incorrect subsequent spelling for Cnecosochara Reitter, 1913.

Cnodalium; Gray, 1831: pl. 80; incorrect subsequent spelling for Cnodalon Latreille, 1797.

Cnodulon; Fabricius, 1801a: xxi; incorrect subsequent spelling for Cnodalon Latreille, 1797.

Cnudalon; Latreille, 1816a: 306; incorrect subsequent spelling for Cnodalon Latreille, 1797.

Coedius; Blanchard, 1845: 13; incorrect original spelling of Caedius Blanchard, 1845.

Coelecnemis; Imhoff, 1856: 238; incorrect subsequent spelling for Coelocnemis Mannerheim, 1843.

Colposcytis; Reitter, 1897a: 298; incorrect subsequent spelling for Colposcythis Reitter, 1889.

Colposphaena; Koch, 1950b: 283; incorrect subsequent spelling for Colposphena Semenov, 1889.

Comphonota; Marseul, 1857: 112; incorrect subsequent spelling for Camphonota Solier, 1836.

Coriogeton; Pic, 1915e: 20; incorrect subsequent spelling for Cyriogeton Pascoe, 1871.

Coronus; Dejean, 1834: 192; published before 1931 without a description, a definition or an indication.

Corypera; Fauvel, 1904: 190; incorrect subsequent spelling for Coripera Pascoe, 1866.

Coscinopter; Allard, 1876a: 15; original spelling corrected to Coscinoptilix in the same work.

Coscinoptilax; Blackwelder, 1945: 542; incorrect subsequent spelling for Coscinoptilix Allard, 1876.

Cossiphus; Laporte, 1833b: 34; incorrect subsequent spelling for Cossyphus G.-A. Olivier, 1791.

Costatostira; Pic, 1954: 232; incorrect subsequent spelling for Costatosora Pic, 1934.

Crirasida; Gebien, 1937a: 726; incorrect subsequent spelling for Cribrasida Reitter, 1917.

Cripticopsis; Español, 1955: 19; incorrect subsequent spelling for Crypticopsis Antoine, 1945.

Cripticus; Español, 1955: 9; incorrect subsequent spelling for Crypticus Latreille, 1816.

Crypsinon: Fairmaire, 1894f: 395; incorrect subsequent spelling for Crypsinous Fairmaire, 1891.

Cryptochyle; Latreille, 1829a: 7; incorrect subsequent spelling for Cryptochile Latreille, 1828.

Ctenoposomus; Reitter, 1906b: 135; alternative original spelling of Ctenioposomus Reitter, 1906, rejected by Bousquet et al. (2015: 140).

Curismosphena; Gebien, 1921b: 3; incorrect subsequent spelling for Curimosphena Gebien, 1920.

Cymathotes; Blanchard, 1845: 33; incorrect subsequent spelling for Cymatothes Dejean, 1834.

Cyphagenia; Laporte, 1840: 191; incorrect subsequent spelling for Cyphogenia Solier, 1837.

Cyphoglossa; Silbermann, 1838: 32; incorrect subsequent spelling for Cryptoglossa Solier, 1837.

Cypobiestes; Gebien, 1942a: 756; incorrect subsequent spelling for Cybopiestes Reitter, 1917.

Deilognatha; Sahlberg, 1903: 26; incorrect subsequent spelling for Dailognatha Steven, 1828.

Delanurops; Iablokoff-Khnzorian, 1964: 309; incorrect subsequent spelling for Delonurops Reitter, 1922.

Dendronomus; Dejean, 1834: 205; published before 1931 without a description, a definition or an indication.

Derilis; Motschulsky, 1872: 27; alternative original spelling of Deriles Motschulsky, 1872, herein rejected.

Diaphanidius; Semenov-Tjan-Shansky and Bogatchev, 1940: 208; incorrect subsequent spelling for Diaphanidus Reitter, 1900.

Diatopsis; Laporte, 1840: 242; incorrect subsequent spelling for Dietopsis Solier, 1835.

Dichroma; Gemminger in Gemminger and Harold, 1870: 1916; incorrect subsequent spelling for Dichromma Mulsant & Rey, 1855.

Dichtrethus; Péringuey, 1892a: 52; incorrect subsequent spelling for Distretus Haag-Rutenberg, 1871.

Dimonus; Reitter, 1914c: 389; incorrect subsequent spelling for Dymonus Solier, 1843.

Diodoeus; Kaszab, 1955: 477; incorrect subsequent spelling for Dioedus J.L. LeConte, 1862.

Diodontus; Laporte, 1840: 185; incorrect subsequent spelling for Diodontes Solier, 1834.

Diphyrhynchus; Fairmaire, 1849: 445; incorrect original spelling for Diphyrrhynchus Fairmaire, 1849.

Diplocyrthus; Escalera, 1914: 355; incorrect original spelling for Diplocyrtus Quedenfeldt, 1887.

Dirosus; Dallas, 1866: 470; incorrect original spelling for Dirosis Miller, 1858.

Dissonus; Marseul, 1863: 181; incorrect original spelling for Dissonomus Jacquelin du Val, 1861.

Dolichopterus; Gemminger in Gemminger and Harold, 1870: 2028; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Doryacus; Gebien, 1938a: 73; incorrect subsequent spelling for Doryagus Pascoe, 1887.

Dromeus; Reitter, 1900c: 186; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Dyla; Marseul, 1857: 116; incorrect subsequent spelling for Dila Fischer von Waldheim, 1844.

Dzungaropterocoma; Löbl et al., 2008b: 166; incorrect subsequent spelling for Dzhungaropterocoma Skopin, 1974.

Ebertia; Kaszab, 1970d: 429; incorrect subsequent spelling for Ebertius Jedlička, 1965.

Ecoptostira; Borchmann, 1936: 236; alternative original spelling of Eccoptostira Borchmann, 1936, herein rejected.

†Edromus; Haupt, 1950: 120; alternative original spelling of †Eodromus Haupt, 1950, herein rejected.

Edrotoporus; Champion, 1895b: 93; incorrect subsequent spelling for Edrotopus Haag-Rutenberg, 1877.

Elanophorus; Dahl, 1823: 41; incorrect subsequent spelling for Elenophorus Dejean, 1821; published in a work suppressed for the purposes of zoological nomenclature (ICZN 1964, Opinion 710).

Eleutheris; Dejean, 1834: 206; published before 1931 without a description, a definition or an indication.

Ellidonius; Gebien, 1937a: 743; incorrect subsequent spelling for Ellidoneus Wilke, 1922.

Elodona; Erichson, 1845b: 256; incorrect subsequent spelling for Eledona Latreille, 1797.

Emallodera; Burmeister, 1875: 467; incorrect subsequent spelling for Emalodera Blanchard, 1842.

Emalodera; Blanchard, 1842: pl. 13; incorrect original spelling, emended to Emmallodera Blanchard, 1842, under prevailing usage.

Emmalodera; Champion, 1895b: 46; incorrect subsequent spelling for Emalodera Blanchard, 1842.

Emsipara; Reitter, 1916b: 10; incorrect subsequent spelling for Emypsara Pascoe, 1866.

Encyrtus; Fairmaire, 18cix; incorrect subsequent spelling for Eucyrtus Lacordaire, 1859.

Endomoderes; Silbermann, 1838: 28; incorrect subsequent spelling for Entomoderes Solier, 1836.

Endothyna; Gebien, 1939: 470; incorrect subsequent spelling for Endothina Carter, 1924.

Entelogonus; Iablokoff-Khnzorian, 1964: 304; incorrect subsequent spelling for Eutelogonus Reitter, 1922.

Enthriptera; Reitter, 1893: 203; alternative original spelling of Euthriptera Reitter, 1893, herein rejected.

Entomoderon; Silbermann, 1838: 30; incorrect subsequent spelling for Entomoderes Solier, 1836.

Eocoptostira; Neave, 1950: 86; incorrect subsequent spelling for Eccoptostira Borchmann, 1936.

Epaerops; Champion, 1895b: 92; incorrect subsequent spelling for Epairops Fåhraeus, 1870.

Epeirops; Bertkau, 1876: 78; incorrect subsequent spelling for Epairops Fåhraeus, 1870.

Ephicerus; Macquart, 1850: 186; published before 1931 without a description, a definition or an indication.

Epicamptus; Dejean, 1834: 198; published before 1931 without a description, a definition or an indication.

Epipedonata; Solier, 1836: 342; alternative original spelling of Epipedonota Solier, 1836, rejected by Solier (1851: 157).

Erganna; Oustalet, 1898: 972; incorrect subsequent spelling for Ergenna Fairmaire, 1897.

Eriodontes; Gebien, 1937a: 546; incorrect subsequent spelling for Erodiontes Reitter, 1914

Eschaptoporis; Seidlitz, 1908: 336; incorrect subsequent spelling for Eschatoporis Blaisdell, 1906.

Etaceta; Ferrer and Delatour, 2007: 275; incorrect subsequent spelling for Etazeta Fairmaire, 1889.

Eucirtus; Schoenfeldt, 1897: 126; incorrect subsequent spelling for Eucyrtus Lacordaire, 1859.

Eudustomus Kolbe, 1889: 127; incorrect subsequent spelling for Endustomus Brême, 1842.

Euganodia; Seidlitz, 1900: 277; incorrect subsequent spelling for Enganodia Fairmaire, 1898.

Eulimenes; Lacordaire, 1859: 467; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Eunotus; Dejean, 1834: 198; published before 1931 without a description, a definition or an indication.

Euoplus; Bertkau, 1879: 502; incorrect subsequent spelling for Evoplus J.L. LeConte, 1866.

Euplopus; Ragusa, 1898: 127; incorrect subsequent spelling for Enoplopus Solier, 1848.

Eurhyzodina; Bremer and Lillig, 2014: 145; incorrect subsequent spelling for Eurhysodina Wasmann, 1921.

Eurichora; G.-A. Olivier, 1795: 2; incorrect subsequent spelling for Eurychora Thunberg, 1789.

Euripimelia; Chernei, 2005: 104; incorrect subsequent spelling for Eurypimelia Reitter, 1915.

Eurygonus; Lacordaire, 1859a: 217; incorrect subsequent spelling for Eurygona Laporte, 1840.

Eusemostene; Gebien, 1940: 764; published after 1930 without a type species designation (see Bousquet et al. 2018: 224).

Eutychus Péringuey, 1908: 405; incorrect subsequent spelling of Eutichus Haag-Rutenberg, 1875.

Eutypodera; Pic, 1907b: 148; incorrect subsequent spelling for Entypodera Gerstaecker, 1871.

Evaniosamus; Laporte, 1840: 192; incorrect subsequent spelling for Evaniosomus Guérin-Méneville, 1834.

Evrychora; Thunberg, 1789: 9; incorrect original spelling for Eurychora Thunberg, 1789.

Evtelus; Fåhraeus, 1870: 304; incorrect original spelling for Eutelus Solier, 1843.

Exangelutus; Kulzer, 1964: 274; incorrect subsequent spelling for Exangeltus Blackburn, 1897.

Excavatoprostenus; Pic, 1944: 8; published after 1930 without a type species designation (see Bousquet et al. 2015: 139).

Falsostenochirus; Gebien, 1943: 924; incorrect subsequent spelling for Falsastenochirus Pic, 1938.

Fatuellus; Gistel, 1850: 409; proposed as a replacement name for the unavailable name Lateranus Sturm, 1843; published before 1931 without a description, a definition, or an indication.

Galymmaphorus; Solier, 1841a: 247; alternative original spelling of Calymmophorus Solier, 1841, herein rejected.

Gampsia; Gistel, 1848a: 126; incorrect subsequent spelling for Campsia Lepeletier & Audinet-Serville, 1828.

Gasthrhaema; Reitter, 1906b: 118; incorrect subsequent spelling for Gastrhaema Jacquelin du Val, 1863.

Gentianidis; Gemminger in Gemminger and Harold, 1870: 2028; incorrect subsequent spelling for Gentinadis Laporte, 1840.

Glyptophrinus; Lesne, 1922: 688; incorrect subsequent spelling for Glyptophrynus Fairmaire, 1899.

Glyptoteryx; Koch, 1956a: 216; incorrect subsequent spelling for Glyptopteryx Gebien, 1910.

Gnata; Dahl, 1823: 41; published in a work suppressed for the purposes of zoological nomenclature (ICZN 1964, Opinion 710).

Gonyodera; Perty, 1832: 63; alternative original spelling of Goniadera, rejected by Perty (1833: 14).

Gymnogathus; Kaszab, 1978: 52; incorrect subsequent spelling for Gymnognathus Solier, 1851.

Hadrobates; Seidlitz, 1905: 275; incorrect subsequent spelling for Habrobates Semenov, 1903.

Halamobia; Reitter, 1911: 329; incorrect subsequent spelling for Halammobia Semenov, 1901.

Harotoma; Kulzer 1951b: 563; incorrect subsequent spelling for Horatoma Solier, 1841.

Harpiscius; Reitter, 1914b: 370; incorrect subsequent spelling for Herpiscius Solier, 1838.

Heiliofugus; Laporte, 1840: 200; incorrect subsequent spelling for Heliofugus Guérin-Méneville, 1831.

Helaeus; Kirby, 1819b: 467; incorrect subsequent spelling for Helea Latreille, 1804.

Heleus; Latreille, 1817: 261; incorrect subsequent spelling for Helea Latreille, 1804.

Heliopathes; Mulsant, 1854: 157; incorrect subsequent spelling for Heliopates Dejean, 1834.

Heliophugus; Lacordaire, 1859b: 744; incorrect subsequent spelling for Heliofugus Guérin-Méneville, 1831.

Heliosthraema; Reitter, 1890a: 34; incorrect original spelling, emended to Heliostrhaema Reitter, 1890 under prevailing usage.

Helisteres Gemminger in Gemminger and Harold, 1870: 2008; incorrect subsequent spelling for Heliosteres Hope, 1841.

Hemicyclys; Imhoff, 1856: 243; incorrect subsequent spelling for Hemicyclus Westwood, 1841.

Hemisphaeresmia; Koch, 1944b: 157; published after 1930 without a type species designation.

Hemitrichesthes; Gebien, 1911b: 620; incorrect subsequent spelling for Hemitrichestes Reitter, 1904.

Hemosodes; Seidlitz, 1909: 298; incorrect subsequent spelling for Hemasodes Casey, 1907.

Hepadrinus; Luederwaldt, 1929: 48; incorrect subsequent spelling for Opatrinus Dejean, 1821.

Heteroblaps; Kolbe, 1928: 202; published before 1931 without a description, a definition, or an indication.

Hexagonocheilus; Solier, 1851: 168; incorrect original spelling for Hexagonochilus Solier, 1851.

Heydyphanes; Kraatz in Heyden and Kraatz, 1882: 332; incorrect subsequent spelling for Hedyphanes Fischer von Waldheim, 1820.

Hipponomus; Iablokoff-Khnzorian, 1964: 303; incorrect subsequent spelling for Hipponome Laporte, 1840.

Hipsosoma; Marseul, 1887: 307; incorrect subsequent spelling for Hypsosoma Ménétriés, 1854.

Holania; Fauvel, 1904: 186; incorrect subsequent spelling for Holaniara Fairmaire, 1871.

Hologlyptus; J.L. LeConte, 1866a: 59; incorrect original spelling for Ologlyptus Lacordaire, 1858.

Homalopus; Solier, 1836: 78; name not used as valid when proposed.

Homoeotrysis; Haag-Rutenberg, 1879b: 136; incorrect subsequent spelling for Homotrysis Pascoe, 1866.

Hovamarygmus; Gebien, 1943: 925; incorrect subsequent spelling for Hovarygmus Fairmaire, 1898.

Hybraenia; Pascoe 1866c: 497; incorrect subsequent spelling for Hybrenia Pascoe, 1866.

Hylobates; Dejean, 1834: 204; published before 1931 without a description, a definition, or an indication.

Hylocurus; Dejean, 1834: 203; published before 1931 without a description, a definition, or an indication.

Hymenophorus; Mulsant, 1852: 68; original spelling of Hymenorus Mulsant, 1852, corrected in the same work.

Hyonthis; Reitter, 1897a: 297; incorrect subsequent spelling for Hionthis Miller, 1861.

Hypophloeus; Panzer, 1795: 339; incorrect subsequent spelling for Hypophlaeus Fabricius, 1790.

Hypsoderes; Dejean, 1834: 195; published before 1931 without a description, a definition, or an indication.

Iophon; Champion, 1895a: 224; incorrect original spelling for Jophon Champion, 1895.

Iphicerus; Dejean, 1834: 203; published before 1931 without a description, a definition, or an indication.

Iphtinus; Reiche, 1854: 16; incorrect original spelling for Iphthinus Dejean, 1834.

Ipitragus; Macquart, 1850: 186; incorrect original spelling for Epitragus Latreille, 1802.

Isomera; Fairmaire and Coquerel, 1866: 46; incorrect subsequent spelling for Isomira Mulsant, 1856.

Isophon; Doyen et al., 1990: 235; incorrect subsequent spelling for Jophon Champion, 1895.

Jintainum; Iwan and Löbl, 2008: 267; incorrect subsequent spelling for Jintaium Ren, 1999.

Lachnoderus; Gebien, 1910b: 301; incorrect subsequent spelling for Lachnoderes Mulsant & Rey, 1859.

Lachnogyia; Reitter, 1904: 34; incorrect subsequent spelling for Lachnogya Ménétriés, 1849.

Lachriomus; Solsky, 1881: 50; incorrect subsequent spelling for Lechriomus Morawitz, 1865.

Lasiotata; Laporte, 1840: 181; incorrect subsequent spelling for Lasiostola Dejean, 1834.

Lathelicus; Heyden et al., 1883: 133; incorrect subsequent spelling for Latheticus C.O. Waterhouse, 1880.

Lateranus; Sturm, 1843: 147; published before 1931 without a description, a definition, or an indication.

Leaena; Dahl, 1823: 41; incorrect subsequent spelling for Laena Dejean, 1821; published in a work suppressed for the purposes of zoological nomenclature (ICZN 1964, Opinion 710).

Leichrodes; Pic, 1934d: 84; incorrect subsequent spelling for Leiochrodes Westwood, 1883.

Leiochbrota; Kirby, 1885a: 90; incorrect subsequent spelling for Leiochrota Westwood, 1883.

Leiochritina; Neervoort van de Poll, 1886: 34; incorrect subsequent spelling for Leiochrotina Westwood, 1883.

Leiochrodomorphus; Kaszab, 1946a: 31; incorrect subsequent spelling for Leichrodomorphus Pic, 1921.

Leporina; Reitter, 1920a: 18; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Leptynoderus; Curtis, 1844: 221; incorrect subsequent spelling for Leptynoderes Solier, 1838.

Leucolephus; Medvedev, 1973: 644; incorrect subsequent spelling for Leucolaephus P.H. Lucas, 1859.

Leucoloephus; P.H. Lucas, 1859: xxii; incorrect original spelling of Leucolaephus P.H. Lucas, 1859.

Liochrinus; Fairmaire, 1893b: 25; incorrect subsequent spelling for Leiochrinus Westwood, 1883.

Listronychus; Laporte, 1840: 244; incorrect subsequent spelling for Lystronychus Latreille, 1829.

Loboderus; Gemminger, 1870: 1925; incorrect subsequent spelling for Lobodera Mulsant & Rey, 1859.

Lobophilus; Pic, 1911a: 183; incorrect subsequent spelling for Lophophyllus Fairmaire, 1887.

Longicerenopus; Berry, 1975: 929; published after 1930 without a type species designation (see Bousquet et al. 2018: 181).

Lutelus; Solier, 1843: 4; alternative original spelling of Eutelus Solier, 1843, herein rejected.

Lyprocaulus; Pic, 1934d: 84; incorrect subsequent spelling for Leprocaulus Fairmaire, 1896.

Lyprochelida; Gebien, 1904b: 20; incorrect subsequent spelling for Lyprochelyda Fairmaire, 1899.

Lyproehelyda; Gebien, 1921b: 238; incorrect subsequent spelling for Lyprochelyda Fairmaire, 1899.

Lyprops; Hope, 1841: 133; incorrect subsequent spelling for Luprops Hope, 1833.

Lystronichus; Latreille, 1829a: 41; incorrect original spelling for Lystronychus Latreille, 1829.

Machlopis; Reitter, 1900c: 191; incorrect subsequent spelling for Machlopsis Pomel, 1871.

Macrophthalmus; Lacordaire, 1859b: 732; incorrect subsequent spelling for Macrophtalmus Montrouzier, 1855.

Macrotis; Dejean, 1834: 186; published before 1931 without a description, a definition, or an indication.

Marcuzzichoton; Ferrer and Moraguès, 2001: 500, 517; published after 1930 without a type species designation (see Kamiński et al. 2019c: 361).

Medaris; Kolbe, 1898: 598; incorrect subsequent spelling for Mederis Motschulsky, 1872.

Megagrius; Laporte, 1840: 183; incorrect subsequent spelling for Megagenius Solier, 1835.

Megliphus; Motschulsky, 1872: 41; alternative original spelling of Meglyphus Motschulsky, 1872, herein rejected.

Meladiesa; Gebien, 1937a: 807; incorrect subsequent spelling for Meladiesia Reitter, 1909.

Melanchrus; Dejean, 1834: 185; published before 1931 without a description, a definition or an indication.

Melanchrus; Lillig and Pavlíček, 2003: 62; incorrect subsequent spelling for Melancrus Reiche & Saulcy, 1857.

Metopocerus; Dejean, 1834: 190; published before 1931 without a description, a definition, or an indication.

Metriopa; Hope, 1841: 118; incorrect subsequent spelling for Metriopus Solier, 1835.

Micispa; Motschulsky, 1858b: 189; incorrect subsequent spelling for Micipsa P.H. Lucas, 1855.

Micrantherus; Bertkau, 1876: 78; incorrect subsequent spelling for Micrantereus Solier, 1848.

Micretyche; Carter, 1926: 134; incorrect subsequent spelling for Micrectyche Bates, 1873.

Micreuphloeus; Gebien, 1911a: 526; incorrect subsequent spelling for Micreuphlaeus Fairmaire, 1897.

Microblattelus; Bremer, 2014: 181; incorrect subsequent spelling for Microblattellus Ferrer, 2006.

Microperithrix; Pierre, 1972: 951; name published after 1930 without a description, a definition or a bibliographic reference to such a published statement.

Microperitrix; Reymond, 1954: 45; name published after 1930 without a description, a definition or a bibliographic reference to such a published statement.

Microps; Dahl, 1823: 46; published in a work suppressed for the purposes of zoological nomenclature (ICZN 1964, Opinion 710).

Microsomira; Anonymous, 1931: 204; incorrect subsequent spelling for Micrisomira Pic, 1930.

Miladion; Reitter, 1887a: 385; alternative original spelling of Myladion Reitter, 1887, rejected by Reitter (1896b: 166).

Milaris; Motschulsky, 1872: 23; incorrect subsequent spelling for Mylaris Pallas, 1781.

Mimoborchmannia; Borchmann, 1936: 490; incorrect subsequent spelling for Mimoborchmania Pic, 1934.

Minantereus; Desmarest, 1860: 168; incorrect subsequent spelling for Micrantereus Solier, 1848.

Mitrogenius; Solier, 1836: 330; alternative original spelling of Mitragenius Solier, 1836, rejected by Solier (1851: 152).

Monogolesthes; Seidlitz, 1906: 268; incorrect subsequent spelling for Mongolesthes Reitter, 1904.

Morocaula; Sharp, 1899: 144; incorrect subsequent spelling for Morocaulus Fairmaire, 1899.

Mosostena; Marseul, 1863: 170; incorrect subsequent spelling for Mesostena Eschscholtz, 1831.

Myaladion; Kaszab, 1960a: 161; incorrect subsequent spelling for Myladion Reitter, 1887.

Myaldion; Kaszab, 1960a: 161; incorrect subsequent spelling for Myladion Reitter, 1887.

Mycetocharus; Stephens, 1832b: 27; incorrect subsequent spelling for Mycetochares Latreille, 1829.

Mycetophylla; Sturm, 1843: 166; incorrect subsequent spelling for Mycetophila Gyllenhal, 1810.

Myria; Gemminger in Gemminger and Harold, 1870: 1963; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Myrmecixenus; Erichson, 1844: 287; incorrect subsequent spelling for Myrmechixenus Chevrolat, 1835.

Neoandrosus; Pic, 1921: 12; alternative original spelling of Neandrosus Pic, 1921, herein rejected.

Nesiotaurus; Fairmaire, 1897b: 385; incorrect subsequent spelling for Nesotaurus Fairmaire, 1896.

Nilion; Latreille, 1802: 179; incorrect original spelling of Nilio Latreille, 1802.

Nocturna; Voet, 1806: 81; published in a work that did not include consistent application of binominal nomenclature.

Nodosogilium; Pic, 1951: 12; alternative original spelling of Nodosogylium Pic, 1951, herein rejected.

Nonpenicillus; Ren, Wang and Yu, 2000: 50; published after 1930 without a type species designation.

Notha; Dejean, 1834: 182; published before 1931 without a description, a definition or an indication.

Notiolosthus; Motschulsky, 1872: 34; alternative original spelling of Notiolesthus, herein rejected.

Notoprateus; Gebien, 1941: 821; incorrect subsequent spelling for Notoprataeus Carter, 1924.

Nyctipathes; Marseul, 1857: 116; incorrect subsequent spelling for Nyctipates Gebler, 1841.

Nystagmus; Gistel, 1848a: 125; proposed as a replacement name for the unavailable name Bucerus Dejean, 1834; published before 1931 without a description, a definition or an indication.

Odontomoplus; Ragusa, 1898: 200; incorrect subsequent spelling for Odontomophlus Seidlitz, 1896.

Oedemetus; Bertkau, 1876: 79; incorrect subsequent spelling for Oedemutes Pascoe, 1860.

Ogcoosoma; Westwood, 1844: 227; incorrect subsequent spelling for Oncosoma Westwood, 1843.

Ogcosoma; Westwood, 1843: 121; incorrect original spelling, emended to Oncosoma Westwood, 1843 under prevailing usage.

Ohionthis; Reitter, 1900c: 89, 139; incorrect subsequent spelling for Ohyonthis Reitter, 1898.

Oleroscelis; Gemminger in Gemminger and Harold, 1870: 1812; incorrect subsequent spelling for Oteroscelis Solier, 1835.

Olisthoena; Fairmaire, 1849: 451; incorrect subsequent spelling for Olisthaena Erichson, 1842.

Omalois; Allard, 1876a: 4; incorrect original spelling of Omaleis Allard, 1876.

Omalus; Allard, 1876a: 4; original spelling of Omalois Allard, 1876, corrected in the same work.

Omocrates; Mulsant, 1854: 150; original spelling of Olocrates, corrected in the same work.

Onymachris; Champion, 1895b: 9; incorrect subsequent spelling for Onymacris Allard, 1885.

Opatrinops; Reitter, 1904: 73; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Ophtalmosis; Deyrolle, 1867: 229; original spelling of Ophthalmosis, corrected in the same work.

Opiestus; Chevrolat, 1833b: 30; alternative original spelling of Oopiestus, rejected by Chevrolat (1847a: 118).

Opisidus; Masters, 1872: 182; incorrect subsequent spelling for Ospidus Pascoe, 1866.

Oplochirus; Bertkau, 1876: 78; incorrect subsequent spelling for Oplocheirus Klug, 1835.

Oplomerus; Dejean, 1834: 206; published before 1931 without a description, a definition, or an indication.

Opsidus; Masters, 1887: 328; incorrect subsequent spelling for Ospidus Pascoe, 1866.

Orchrotus; Cazurro Ruiz, 1894a: 63; incorrect subsequent spelling for Oochrotus Lucas, 1852.

Orientacara; Penrith, 1986: 294; incorrect subsequent spelling for Orientocara Koch, 1952.

Orthogonoderus; Germain, 1855: 402, 403; incorrect subsequent spelling for Orthogonoderes Solier, 1841.

Oxipistoma; Koch, 1940a: 259; alternative original spelling of Oxypistoma, also proposed after 1930 without type species designation.

Pachynoscelis; Heyden, 1882: 144; published without a description, a definition, or an indication.

Paecilesthus; Dejean, 1834: 207; incorrect original spelling of Poecilesthus Dejean, 1834.

Paleosclerum; Nabozhenko, 2019: 6; incorrect subsequent spelling for Palaeosclerum Nabozhenko & Kirejtshuk, 2017.

Paracupezus; Riley, 1923: 129; incorrect subsequent spelling for Pareupezus Kolbe, 1889.

Parapiophorus; Nabozhenko, 2019: 8; incorrect subsequent spelling for Paropiophorus Haupt, 1950.

Parenicmosoma; Ardoin, 1959a: 61; published after 1930 without a type species designation.

Pectenepitragus; Gebien, 1937a: 570; incorrect subsequent spelling for Pectinepitragus Pic, 1927.

Pectophegoneus; Anonymous in Staff of the Zoological Society of London, 1972: 507; incorrect subsequent spelling for Pectphegoneus Freude, 1968.

Pedirus; Doyen et al., 1990: 237; incorrect subsequent spelling for Pediris Motschulsky, 1872.

Pelecipalpus; Kaszab, 1982b: 48; incorrect subsequent spelling for Pelecypalpus Hinton, 1947.

Penthapyllus; Ragusa, 1898: 123; incorrect subsequent spelling for Pentaphyllus Dejean, 1821.

Penticinus; Reitter, 1896b: 171; alternative original spelling of Penthicinus Reitter, 1896, rejected by Reitter (1904: 135, 170).

Penticus; Medvedev and Iwan, 2007: 613; incorrect subsequent spelling for Penthicus Faldermann, 1836.

Periclytus; Bertkau, 1876: 78; incorrect subsequent spelling for Proselytus Fåhraeus, 1870.

Periseptus; Fairmaire, 1887a: 176; incorrect subsequent spelling for Peristeptus Haag-Rutenberg, 1875.

Petrobius; Brullé, 1832: 202; incorrect subsequent spelling for Gnaptor Brullé, 1831.

Phalera; Dejean, 1821: 68; incorrect subsequent spelling for Phaleria Latreille, 1802.

Phanerentoma; Lacordaire, 1859a: 195; incorrect subsequent spelling for Phanerotoma Solier, 1843.

Philammus; Escalera 1914: 339; incorrect subsequent spelling for Philhammus Fairmaire, 1871.

Philax; Dejean, 1834: 192; incorrect subsequent spelling for Phylax Brullé, 1832.

Philethus; Macquart, 1850: 182; incorrect subsequent spelling for Phyletes Redtenbacher, 1845.

Philoscotus; Dejean, 1834: 186; published before 1931 without a description, a definition or an indication.

Phlaegmatus; Dejean, 1834: 208; published before 1931 without a description, a definition or an indication.

Phloeotribon; Gerstaecker, 1866: 427; incorrect subsequent spelling for Phaeotribon Kraatz, 1865.

Phrepates; Solier, 1834: 488; incorrect subsequent spelling for Phrenapates Gray, 1831.

Phylacoprosodes; Schuster, 1934: 75; replacement name for Aulonoscelis Reitter, 1909, which is not an available name.

Phylas; Berthold, 1827: 368; incorrect subsequent spelling for Phylan Sturm, 1826.

Phylethus; Chevrolat, 1847b: 57; incorrect subsequent spelling for Phyletes Redtenbacher, 1845.

Phyletus; Marseul, 1863: 184; incorrect subsequent spelling for Phyletes Redtenbacher, 1845.

Phylhammus; Español, 1963b: 188; incorrect subsequent spelling for Philhammus Fairmaire, 1871.

Phymatesthes; Gebien, 1928: 189, 191; incorrect subsequent spelling for Phymatestes Pascoe, 1866.

Phymathodes; Blanchard, 1845: 39; incorrect subsequent spelling for Phymatodes Dejean, 1834.

Phymatisoma; Laporte and Brullé, 1831: 332, 408; incorrect original spelling, emended to Phymatosoma Laporte & Brullé, 1831 under prevailing usage.

Phythora; Marquet, 1897: 162; incorrect subsequent spelling for Phtora Germar, 1836.

Phytophilus; Guérin-Méneville, 1838: 99; incorrect subsequent spelling for Phitophilus Guérin-Méneville, 1831.

Phyxelius; Philippi, 1864: 348; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Piazomera; Gebien, 1910a: 154; incorrect subsequent spelling for Piesomera Solier, 1843.

Piezomera; Gemminger in Gemminger and Harold, 1870: 1898; incorrect subsequent spelling for Piesomera Solier, 1843.

Piliobola; Hope, 1841: 112; published before 1931 without a description, a definition, or an indication.

Pimelionotus; Ardoin, 1962b: 969; published after 1930 without a type species designation.

Pineta; Desmarest, 1860: 159; incorrect subsequent spelling for Peneta Lacordaire, 1859.

Plaesia; Fairmaire, 1875: 43; incorrect subsequent spelling for Plesia Klug, 1833.

Platomodes; Dallas, 1866: 473; incorrect subsequent spelling for Platamodes Ménétriés, 1849.

Platysum; Mulsant and Rey, 1859a: 73, 81; original spelling of Platynosum, corrected in the same work.

Plectrascelis; Silbermann, 1838: 28; incorrect subsequent spelling for Psectrascelis Solier, 1836.

Podhamala; Motschulsky, 1860c: 535; incorrect subsequent spelling for Podhomala Solier, 1836.

Polpagenia; Laporte, 1840: 183; incorrect subsequent spelling for Polpogenia Solier, 1836.

Polycoelagastridion; Kaszab, 1942: 14; incorrect subsequent spelling for Polycoelogastridion Reichardt, 1936.

Polycoelogastridium; Gebien, 1938a: 428; incorrect subsequent spelling for Polycoelogastridion Reichardt, 1936.

Porphyrrhyba; Champion, 1895b: 195; incorrect subsequent spelling for Porphyrhyba Fairmaire, 1877.

Porphyruba; Bertkau, 1890: 260; incorrect subsequent spelling for Porphyrhyba Fairmaire, 1877.

Porphyryba; Fairmaire, 1894b: 144; incorrect subsequent spelling for Porphyrhyba Fairmaire, 1877.

Prachoma; Laporte, 1840: 195; incorrect subsequent spelling for Prochoma Solier, 1835.

Praosis; Laporte, 1840: 186; incorrect subsequent spelling for Praocis Eschscholtz, 1829.

Praygena; Bertkau, 1876: 78; incorrect subsequent spelling for Praeugena Laporte, 1840.

Priopus; Hope, 1841: 73; alternative original spelling of Prioscelis, rejected by Westwood (1844: 211).

Procris; Curtis, 1844: 221; incorrect subsequent spelling for Praocis Eschscholtz, 1829.

Prodhomala; Marseul, 1857: 112; incorrect subsequent spelling for Podhomala Solier, 1836.

Proplatamodes; Kaszab, 1960a: 16; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Prorythinota; Ferrer et al., 2016: 160; incorrect subsequent spelling for Prorhytinota Koch, 1943, an unavailable name.

Prothraustocolus; Kaszab, 1979b: 278; incorrect subsequent spelling for Prothraustocola Kaszab, 1957.

Psammeticus; Guérin-Méneville, 1831b: pl. 28bis; incorrect subsequent spelling for Psammetichus Latreille, 1828.

Psammodius; Berthold, 1827: 366; incorrect subsequent spelling for Psammodes W. Kirby, 1819.

Psaryphys; Cazurro Ruiz, 1895: 498; incorrect subsequent spelling for Psaryphis Erichson, 1843.

Pseudammobius; Franz, 1996: 71; new nominal genus published in a combined description with a single included new nominal species not marked by “gen. nov., sp. nov.” or an equivalent expression (Löbl et al. 2008a: 45).

Pseudelops; Fauvel, 1904: 190; incorrect subsequent spelling for Pseudhelops Guérin-Méneville, 1841.

Pseudemallus; Edwards and Vevers, 1975: 281; incorrect subsequent spelling for Pseudemmallus Koch, 1956.

Pseudoammobius; Franz, 1996: 131; new nominal genus published in a combined description with a single included new nominal species not marked by “gen. nov., sp. nov.” or an equivalent expression (see Löbl et al. 2008a: 45).

Pseudobax; Gebien, 1942a: 308; incorrect subsequent spelling for Pseudabax Kraatz, 1880.

Pseudocylibe; Kaszab, 1982b: 291; alternative original spelling of Pseudocylibe Kaszab, 1982, herein rejected.

Pseudocyrtus; Pic 1916: 14; alternative original spelling of Pseudeucyrtus Pic, 1916, herein rejected.

Pseudoderilis; Gebien, 1941: 343; incorrect subsequent spelling for Pseudoderiles Gebien, 1928.

Pseudohelops; Cazurro Ruiz, 1895: 506; incorrect subsequent spelling for Pseudhelops Guérin-Méneville, 1841.

Pseudonantes; Pic, 1925b: 7; incorrect subsequent spelling for Pseudonautes Fairmaire, 1892.

Pseudostena; Gemminger in Gemminger and Harold, 1870: 1958; incorrect subsequent spelling for Pseudostene Wollaston, 1861.

Pseudotrichoplatycelis; Kaszab, 1960a: 83; alternative original spelling of Pseudotrichoplatyscelis Kaszab, 1960, herein rejected.

Pseudotrichoplatynoscelis; Kaszab, 1960a: 82; alternative original spelling of Pseudotrichoplatyscelis Kaszab, 1960, herein rejected.

Psilioloba; Mulsant and Rey, 1853b: 20; incorrect subsequent spelling of Pilioloba Erichson in Agassiz, 1846.

Pyganisia; Hope, 1841: 133; incorrect subsequent spelling for Pyanisia Laporte, 1840.

Raibosceles; Allard, 1876a: 5; incorrect original spelling for Raiboscelis Allard, 1876.

Reichardtia; Kaszab, 1982c: 79; incorrect subsequent spelling for Reichardtiella Kaszab, 1942.

Reiterella; Seidlitz, 1905: 276; incorrect subsequent spelling for Reitterella Semenov, 1891.

Rhamnatodes Champion, 1895b: 20; incorrect subsequent spelling for Rhammatodes Haag-Rutenberg, 1876.

Rhicnodes; Gebien, 1941: 817; incorrect subsequent spelling for Rhicnodus Fairmaire, 1892.

Rhytimorpha; Koch, 1943: 888; alternative original spelling of Rhydimorpha Koch, 1943, herein rejected.

Rophobas; Motschulsky, 1872: 36; alternative original spelling of Rhophobas Motschulsky, 1872, herein rejected.

Rythinota; Ferrer et al., 2016: 159; incorrect subsequent spelling for Rhytinota Eschscholtz, 1831.

Rytinota; Eschscholtz, 1831: 5, 7; incorrect original spelling, emended to Rhytinota Eschscholtz, 1831 under prevailing usage.

Saragdonius; Bertkau, 1876: 78; incorrect subsequent spelling for Saragodinus Bates, 1872.

Sarathropus; Heyden, 1895: 111; incorrect subsequent spelling for Sarothropus Kraatz, 1865.

Saurothropus; Dallas, 1866: 471; incorrect subsequent spelling for Sarothropus Kraatz, 1865.

Scelidopsecta; Kulzer, 1954b: 204; incorrect original spelling for Scelidospecta Kulzer, 1954.

Sceloblaps; Kolbe, 1928: 203; published before 1931 without a description, a definition or an indication.

Schizomena; Sharp, 1922: 142; incorrect subsequent spelling for Schizomma Gebien, 1921.

Scytodonta; Reitter, 1900c: 166; incorrect subsequent spelling for Scythodonta Reitter, 1894.

Selenepistomus; Mulsant and Rey, 1853b: 20; incorrect subsequent spelling for Selenepistoma Dejean, 1834.

Selenoma; Peña, 1966: 408; incorrect subsequent spelling for Selenomma Dejean, 1836.

Sepidiostemis; Reitter, 1914c: 382; incorrect subsequent spelling for Sepidiostenus Fairmaire, 1884.

Sepidostenus; Fairmaire, 1887a: 184; incorrect subsequent spelling for Sepidiostenus Fairmaire, 1884.

Sericeus Ragusa, 1898: 114; incorrect subsequent spelling for Seriscius Motschulsky, 1845.

Solenopistoma; Mulsant and Rey, 1854: 29; incorrect subsequent spelling for Selenepistoma Dejean, 1834.

Solskia; Semenov, 1891: 263; incorrect subsequent spelling for Solskyia Solsky, 1881.

Solskya; Semenov-Tjan-Shansky, 1908: 265; incorrect subsequent spelling for Solskyia Solsky, 1881.

Somaticum; Hope, 1841: 117; incorrect original spelling for Somaticus Hope, 1841.

Spaerotidius; Kaszab, 1941a: 39, 40; alternative original spelling of Sphaerotidius Kaszab, 1941, herein rejected.

Sphaenariopsis; Blair, 1935a: 103; incorrect original spelling for Sphenariopsis Kraatz, 1865.

Sphingidophorus; Pic, 1917: 1; incorrect original spelling for Sphragidophorus Champion, 1889.

Sphoerotus; Brême, 1842a: 106; incorrect original spelling for Sphaerotus W. Kirby, 1819.

Spinanaedus; Gebien, 1941: 826; incorrect original spelling for Spinadaenus Pic, 1921.

Spinolystronychus; Pic, 1944: 5; published after 1930 without a type species designation (see Bousquet et al. 2015: 139).

Statyra; Lacordaire, 1830b: 154; incorrect original spelling for Statira Lepeletier & Audinet-Serville, 1828.

Stegatopsis; Allard, 1884: 23; incorrect subsequent spelling for Stegastopsis Kraatz, 1865.

Stenochara; Hope, 1841: 70; incorrect subsequent spelling for Stenocara Solier, 1835.

Stenogenius; Solier, 1834: 520; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Sternodus; Agassiz, 1846b: 353; incorrect subsequent spelling for Sternodes Fischer von Waldheim, 1837.

Subtentyrina; Koch, 1939: 260; published after 1930 without a type species designation.

Subtentyrina; Löbl and Merkl, 2003: 251; published after 1999 and not explicitly indicated as intentionally new.

Sulpiosoma; Ferrer, 2006c: 79; alternative original spelling of Sulpiusoma Ferrer, 2006, herein rejected.

Symnetasida; Ragusa, 1921: 100; incorrect subsequent spelling for Gymnetasida Reitter, 1917

Syntractus; Borchmann, 1909b: 2; incorrect subsequent spelling for Synatractus W.J. Macleay, 1887.

Taeniobates; Motschulsky, 1872: 32; alternative original spelling of Taenobates Motschulsky, 1872, herein rejected.

Taganoides; Kaszab, 1965: 109; incorrect subsequent spelling for Tagonoides Fairmaire, 1886.

Tagaulus; Hayashi, 1980: 19; incorrect subsequent spelling for Tagalus Gebien, 1914.

Talpophila; Macquart, 1850: 174; incorrect subsequent spelling for Thalpophila Solier, 1835.

Tanchyrus; Kaszab, 1941a: 2, 42; incorrect subsequent spelling for Tanchirus Fairmaire, 1897.

Tanuria; Lacordaire, 1859: 454; first published as a synonym and not treated before 1961 as an available name and adopted as the name of a taxon or treated as a senior homonym.

Targesius; Gistel, 1848: xi; proposed as a replacement name for the unavailable name Melancrus Dejean, 1834; published before 1931 without a description, a definition, or an indication.

Tarsoconodes; Sharp, 1922: 142; incorrect subsequent spelling for Tarsocnodes Gebien, 1920.

Taurocerus; Agassiz, 1846b: 362; incorrect subsequent spelling for Tauroceras Hope, 1841.

Taxonema; Bertkau, 1876: 78; incorrect subsequent spelling for Toxocnema Fåhraeus, 1870.

Telleas; Gebien, 1911a: 447; incorrect subsequent spelling for Telleus Fairmaire, 1904.

Teneobrionites; Bousquet et al. 2018: 381; incorrect subsequent spelling for TenebrionitesCockerell 1920.

Tentiria; Dahl, 1823: 41; incorrect subsequent spelling for Tentyria Latreille, 1802; published in a work suppressed for the purposes of zoological nomenclature (ICZN 1964, Opinion 710).

Tentyrionota; Gridelli, 1937: 38; incorrect subsequent spelling for Tentyronota Reitter, 1900.

Thecocerus; Agassiz, 1846b: 368; proposed as an unjustified emendation for the unavailable name Thecacerus Dejean, 1834; published before 1931 without a description, a definition, or an indication.

Thoracophora; Imhoff, 1856: 242; incorrect subsequent spelling for Thoracophorus Hope, 1841.

Thraucostolus; Gebien, 1910a: 74; incorrect subsequent spelling for Thraustocolus Kraatz, 1866.

Thryptera; Gaubil, 1849: 213; incorrect subsequent spelling for Thriptera Solier, 1836.

Thylacoderus; Burmeister, 1875: 489; incorrect subsequent spelling for Thylacoderes Solier, 1843.

Trachelaeum; Agassiz, 1846b: 373; incorrect subsequent spelling for Tracheloeum Hope, 1841.

Tracheloblaps; Kolbe, 1928: 203; published before 1931 without a description, a definition or an indication.

Trachinotus; Gistel, 1848a: xi; incorrect subsequent spelling for Trachynotus Latreille, 1828.

Treintoma; Wolcott, 1950: 326; incorrect subsequent spelling for Trientoma Solier, 1835.

Trelolosodis; Cazurro Ruiz, 1897b: 427; incorrect subsequent spelling for Scelosodis Solier, 1835.

Tribolocara; Lacordaire, 1859a: 69, 72; incorrect subsequent spelling for Trilobocara Solier, 1851.

Trichochianalis; Kaszab, 1940a: 201; alternative original spelling of Trichochianalus, rejected by Ksazab (1940b: 989).

Tricholeipoleura; Kaszab, 1940a: 152; alternative original spelling of Tricholeipopleura, rejected by Kaszab (1940b: 989).

Trichopodum; Gebien, 1905: 253; incorrect subsequent spelling for Trichopodus Mulsant & Rey, 1859.

Trichosphena; Gebien, 1937a: 577; incorrect subsequent spelling for Trichosphaena Reitter, 1916.

Trigonoides; Fairmaire, 1901c: 267; incorrect subsequent spelling for Tagonoides Fairmaire, 1886.

Trinobatis; Laporte, 1840: 195; incorrect subsequent spelling for Thinobatis Eschscholtz, 1831.

Trisilius; Bertkau, 1879: 505; incorrect subsequent spelling for Trisilus Haag-Rutenberg, 1878.

Trogloderes; Gebien, 1938a: 64; incorrect subsequent spelling for Trogloderus J.L. LeConte, 1879.

Ulomimimus; Bates, 1873a: 201; incorrect original spelling, emended to Ulomimus Bates, 1873 under prevailing usage.

Ulona; Dahl, 1823: 43; incorrect subsequent spelling for Uloma Dejean, 1821; published in a work suppressed for the purposes of zoological nomenclature (ICZN 1964, Opinion 710).

Umbraticus; Voet, 1806: 81; published in a work that did not include consistent application of binominal nomenclature.

Urosis; Deyrolle, 1867: 81; alternative original spelling of Anisosis Deyrolle, 1867, rejected by Dallas (1868: 265–266).

Usagaria; Péringuey, 1892a: 56; incorrect subsequent spelling for Uzagaria Ancey, 1881.

Usoma; Berthold, 1827: 369; incorrect subsequent spelling for Uloma Dejean, 1821.

Valeron; Cazurro Ruiz, 1897b: 97; incorrect subsequent spelling for Scleron Hope, 1841.

Verticiphloeus; Bremer and Lillig, 2017b: 201; published after 1930 without a description, a definition, or a bibliographic reference to such a published statement.

Victa; Allard, 1870: 50; incorrect subsequent spelling for Vieta Laporte, 1840.

Xanthomerus; Escalera, 1914: 353; incorrect subsequent spelling for Xanthomus Mulsant, 1854.

Xanthotopeia; Fairmaire, 1894g: 671, 672; incorrect subsequent spelling for Xanthothopeia Mäklin, 1867.

Xanthotopia; Kolbe, 1897b: 619; incorrect subsequent spelling for Xanthothopeia Mäklin, 1867.

Zaleucos; Gebien, 1937a: 715; incorrect subsequent spelling for Zaleucus Champion, 1892.

Zaphobas; Papp, 1961: 128; incorrect subsequent spelling for Zophobas Dejean, 1834.

Zarudnyonymus; Kaszab 1981b: 367; incorrect subsequent spelling for Zarudnionymus Semenov-Tjan-Shansky & Bogatchev, 1947.

Zioelas; Gebien, 1943: 914; incorrect subsequent spelling for Ziaelas Fairmaire, 1892.

Zophabas; Motschulsky, 1872: 35; incorrect subsequent spelling for Zophobas Dejean, 1834.

Zophelops; Gebien, 1911a: 534; incorrect subsequent spelling for Zophohelops Reitter, 1902.

Zophobius; Dejean, 1834: 180; published before 1931 without a description, a definition, or an indication.

Zypoetus; Kaszab, 1977: 303; incorrect subsequent spelling for Zypoetes Champion, 1893.

Appendix 2

Supporting references for conservation of Cyphaleus Westwood, 1841 over Chrysobalus Boisduval, 1835 through reversal of precedence (ICZN 1999, Article 23.9.2). To our knowledge Chrysobalus Boisduval, 1835 has not been used as a valid name after 1899.

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4. Hangay G, Zborowski P (2010) A guide to the beetles of Australia. CSIRO Publishing, Collingwood. x + 238 pp.

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18. Matthews EG, Scupola A (2003) Entomological investigations in Australia by the Natural History Museum of Turin: Coleoptera, Tenebrionidae. Museo Regionale di Scienze Naturali, Monografie 35: 281–302.

19. Moore BP (1980) A guide to the beetles of southeastern Australia. Australian Entomological Press, Greenwich, Australia, 226 pp.

20. Naumann I (1993) CSIRO handbook of Australian insect names: common and scientific names for insects and allied organisms of economic and environmental importance. Sixth Edition. CSIRO Publishing, East Melbourne, Australia, 200 pp.

21. Steiner Jr WE (2009) Tenebrionidae of Australia. Descriptions of tribes. Keys to genera. Catalogue of species by E.G. Matthews and P. Bouchard. Australian Biological Resources Study, GPO Box 787, Canberra ACT 2601, Australia, 2008, pp. 398. Systematic Entomology 34: 198.

22. Stone c, Goodyer G, Sims K, Penman T, Carnegie A (2010) Beetle assemblages captured using static panel traps within New South Wales pine plantations. Australian Journal of Entomology 49: 304–316.

23. Williams G (2002) A taxonomic and biogeographic review of the invertebrates of the Central Eastern Rainforest Reserves of Australia (CERRA) World Heritage Area, and adjacent regions. Technical Reports of the Australian Museum 16: 1–208.

24. Williams G (2020) The invertebrate world of Australia’s subtropical rainforests. CSIRO Publishing, Clayton South, 385 pp.

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Citation

Bouchard P, Bousquet Y, Aalbu RL, Alonso-Zarazaga MA, Merkl O, Davies AE (2021) Review of genus-group names in the family Tenebrionidae (Insecta, Coleoptera). ZooKeys 1050: 1–633. https://doi.org/10.3897/zookeys.1050.64217