1UMR ENTROPIE (UR‑IRD‑CNRS‑IFREMER‑UNC), LabEx « Corail », Université de La Nouvelle-Calédonie, BP R4, 98851 Nouméa Cedex, New Caledonia
2CRIOBE, PSL Research University, USR 3278 EPHE-CNRS-UPVD, LabEx « Corail », Université de Perpignan, Avenue Paul Alduy, 66860 Perpignan Cedex, France
3Mediterranean Institute of Oceanography, UM 110 (AMU-UTV-CNRS-IRD), Campus de Luminy, Case 901, 13288 Marseille Cedex 9, France
4GEOMAR Helmholtz Centre for Ocean Research, Research Division Marine Ecology, Düsternbrooker Weg 20, 24105 Kiel, Germany
5UMR LIENSs 7266 (CNRS-ULR), Institut du Littoral Et de L’environnement, 2 rue Olympe de Gouges, 17000 La Rochelle, France
6Laboratoire BOREA, Muséum National d’Histoire Naturelle, CNRS 7208, IRD 207-SU-UCN-UA, Muséum National d’Histoire Naturelle, 61 rue Buffon, 5 CP 53, 75231 Paris Cedex, France
7Newcastle University, School of Natural and Environmental Sciences, Newcastle‑upon‑Tyne, NE1 7RU UK
8UMR EIO (UPF-IRD-ILM-IFREMER), Université de La Polynésie Française, LabEx « Corail », BP 6570, 98702 Faa’a, Tahiti French Polynesia
© The Author(s) 2021
Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.