Skip to main content
PLOS ONE logoLink to PLOS ONE
. 2022 Jul 21;17(7):e0271504. doi: 10.1371/journal.pone.0271504

How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?: A simulation analysis in urban Namibia

Dana R Thomson 1,2,*, Douglas R Leasure 3,¤a, Tomas Bird 3,¤b, Nikos Tzavidis 2, Andrew J Tatem 3
Editor: Krishna Prasad Vadrevu4
PMCID: PMC9302737  PMID: 35862480

Abstract

Disaggregated population counts are needed to calculate health, economic, and development indicators in Low- and Middle-Income Countries (LMICs), especially in settings of rapid urbanisation. Censuses are often outdated and inaccurate in LMIC settings, and rarely disaggregated at fine geographic scale. Modelled gridded population datasets derived from census data have become widely used by development researchers and practitioners; however, accuracy in these datasets are evaluated at the spatial scale of model input data which is generally courser than the neighbourhood or cell-level scale of many applications. We simulate a realistic synthetic 2016 population in Khomas, Namibia, a majority urban region, and introduce several realistic levels of outdatedness (over 15 years) and inaccuracy in slum, non-slum, and rural areas. We aggregate the synthetic populations by census and administrative boundaries (to mimic census data), resulting in 32 gridded population datasets that are typical of LMIC settings using the WorldPop-Global-Unconstrained gridded population approach. We evaluate the cell-level accuracy of these gridded population datasets using the original synthetic population as a reference. In our simulation, we found large cell-level errors, particularly in slum cells. These were driven by the averaging of population densities in large areal units before model training. Age, accuracy, and aggregation of the input data also played a role in these errors. We suggest incorporating finer-scale training data into gridded population models generally, and WorldPop-Global-Unconstrained in particular (e.g., from routine household surveys or slum community population counts), and use of new building footprint datasets as a covariate to improve cell-level accuracy (as done in some new WorldPop-Global-Constrained datasets). It is important to measure accuracy of gridded population datasets at spatial scales more consistent with how the data are being applied, especially if they are to be used for monitoring key development indicators at neighbourhood scales within cities.

Introduction

Small area population counts, especially in low- and middle-income countries (LMICs), provide essential denominators for health, economic, and development indicators [1]. For example, small area population counts are used to calculate vaccination coverage rates [2], understand health service utilisation [3], and estimate infection rates of malaria, COVID-19, and many other health conditions [4]. Spatially-detailed and time-sensitive population counts are also essential to monitor and understand the accelerated pace of urbanisation in LMICs compared to HICs. Ninety percent of global population growth in the next 30 years is expected to occur in African and Asia cities alone [5], which means it is vital to monitor population trends across diverse LMIC cities with respect to economic development, human impacts on biodiversity and environment, and the changing climate [6,7]. Authoritative population data are traditionally collected via a national census. Censuses are generally collected every ten years, though one in ten LMICs has not held a census in the last 15 years [8], and some national censuses have poor data quality due to negligence (e.g., [9,10]) or deliberate mis-counting of sub-populations for political purposes (e.g., [1113]). Due to increasing rates of mobility and urbanisation worldwide, the urban poorest–especially in LMIC cities–are increasingly difficult to count as more people take-up residence in informal settlements or atypical housing locations (e.g., shops) [14].

In the absence of updated, fine-scale census data, many policy-makers, urban planners, researchers, and service providers have turned to gridded population estimates as a source of population counts in their work. Gridded population data are viewed by data producers and users as meeting a global development challenge to “leave no one off the map” and thus leave no one behind [15]. However, performing accuracy assessments of gridded population datasets at the scale at which they are applied (e.g., neighbourhood, grid cell) poses a conundrum; reliable fine-scale population counts are generally not available where they are needed most [16], and users often turn to gridded population estimates when census counts are excessively outdated or untrustworthy [14]. Despite these challenges, it is imperative to understand if, and how, census inaccuracies propagate through gridded population datasets, especially with respect to vulnerable populations.

Briefly, gridded population data provide estimates of the total population in small grid cells, and are derived with geo-statistical methods using population counts and spatial datasets [16]. “Top-down” gridded population estimates have been available for roughly 15 years and disaggregate census or other complete population counts from areal units (e.g., 3rd-, 4th-, or 5th-level administrative units) to grid cells (e.g., 30x30m, 100x100m, 1x1km) [14]. The simplest models assume a uniform distribution of population within areal units (i.e., GPW [17,18], GHS-POP [19,20], HRSL [21]), while the most complex models use spatial covariates to inform spatial disaggregation from the areal unit to grid cells (i.e., WorldPop [22,23], LandScan [24,25], WPE [26]). To estimate gridded population figures beyond the year of the last census; birth, migration, and death rates are used to project new population totals by areal unit [27]. “Bottom-up” gridded population estimates are derived from micro-census population counts in a sample of areas, or from assumptions about the average household size, and have only recently been developed [28,29]. Read papers by Leyk and colleagues (2019) and Thomson and colleagues (2020) for detailed descriptions and comparisons of gridded population datasets [14,16].

The accuracy of “top-down” gridded population data is generally calculated at the scale of the input population areal units because these are the finest-scale population counts available to the data producers. A number of factors contribute to gridded population model accuracy including: (1) the modelling algorithm itself, (2) inaccuracy of the input population data, (3) the geographic scale of the input population data (e.g., census tracts versus districts), (4) the age, accuracy, completeness, and type of ancillary data, (5) the nature of the relationship between ancillary data and population density, and (6) the geographic scale of the output grid. Of these, the two strongest predictors of accuracy (at the scale of areal units) in top-down gridded population models are the resolution and age of the input population data [30]. Among top-down gridded population datasets, the WorldPop-Global-Unconstrained Random Forest model was among the best documented and most accurate gridded population models available at the time of this analysis in 2017–2019 [22,31]. Specifically, the model code [32] and pre-processed model covariates [33,34] were publicly available enabling reproducibility and evaluation. WorldPop-Global-Unconstrained and its preceding data products (AfriPop, AsiaPop, and AmeriPop) result in estimates for all land areas; however, a new WorldPop-Global-Constrained dataset was published in 2020 limiting population estimates to cells with buildings or built-up features [35].

To evaluate cell-level accuracy of gridded population data, actual population counts are needed for each grid cell or in finer-scale units such as household point locations. Few censuses in LMICs collect household latitude-longitude coordinates, and where these censuses exist, the data are extremely sensitive and difficult to obtain. Furthermore, even the best census data might be problematic because vulnerable sub-populations including homeless and nomadic populations are supposed to be counted separately in special enumerations. Unfortunately, though, under-resourced statistical offices are often not able to perform these counts [36], and some censuses do not include certain refugee or internally displaced populations [37]. To ensure that this analysis of cell-level accuracy did not exclude the urban poorest and other hidden populations, we chose to simulate a realistic population in a LMIC setting. It was important that the synthetic population was located in a real-world location so that actual covariate datasets–with their own imperfections–could be used to generate realistic gridded population datasets. We adapted methods outlined by Thomson and colleagues (2018) for simulating a geo-located realistic household population, and added classification of urban households by slum/non-slum area in a final step to focus this analysis on dynamic, complex LMICs cities where inaccuracies in gridded data are likely to propagate [38].

This paper describes how we evaluated the cell-level accuracy of 32 simulated 100x100m WorldPop-Global-Unconstrained gridded population datasets which reflect realistic levels of census (1) outdatedness (0-, 5-, 10-, and 15-years outdated), (2) inaccuracy (none, low, middle, and high missing population counts), and (3) two administrative-level aggregations of the population in an urban LMIC setting. This is among the first assessments of cell-level accuracy of a gridded population dataset in a LMIC setting. While the methods and approach outlined here to evaluate cell-level accuracy (developing a realistic synthetic population, and from this, deriving several realistic versions of census data) were applied to just one gridded population dataset, they could be applied to other gridded population data products used for development monitoring and decision-making.

Methods

Setting

We chose to simulate a population in Khomas, Namibia–in which the vast majority of residents reside in Windhoek, the capital–because the government has produced numerous high-quality population datasets [39], and Windhoek’s population is incredibly dynamic (Fig 1). Namibia, like some other countries that inherited colonial boundaries, placed restrictions on freedom of movement until independence in 1990 [40]. After independence, vast numbers of people migrated to Windhoek, exaggerating rural-to-urban migration patterns observed globally during this time period [41,42]. Windhoek is also a destination for immigrants from neighbouring countries including financially unstable Zimbabwe [42,43]. The population of the Windhoek metropolitan area grew by a staggering 37% between the 2001 and 2011 censuses [39], with much of that growth in informal settlements [40].

Fig 1. Location of Khomas region in Namibia, and of constituencies in Windhoek area.

Fig 1

Source: Constituency boundaries publically available from https://gadm.org/.

Simulation overview

To simulate realistic gridded population datasets for Khomas, Namibia, we (a) simulated a “true” synthetic 2016 population geo-located to realistic manually-generated household point locations; (b) introduced realistic outdatedness by removing households in 2011, 2006, and 2001; (c) introduced realistic inaccuracies among urban-slum, urban-non-slum, and rural sub-populations; and (d) aggregated these 16 simulated population scenarios into two geographic areal units (census EA and constituency) to generate 32 realistic census datasets. These 32 realistic census datasets were consequently used to model 32 realistic WorldPop-Global-Unconstrained 100x100m gridded population datasets. This workflow is summarised in Fig 2 and detailed below.

Fig 2. Summary of the population and gridded population simulation workflow.

Fig 2

(1) Simulate a realistic population geo-located to realistic building point locations, (2) simulate three periods of outdatedness by removing households at point locations not present on satellite imagery in earlier years, (3) simulate low/middle/high census inaccuracy by removing points at random from rural, urban-slum, and urban-non-slum household types, (4) aggregate to 922 census enumeration areas (EAs) and 10 constituencies (admin-2), (5) generate 100x100m gridded population datasets in raster grid format using WorldPop-Global-Unconstrained approach and WorldPop-Global spatial covariates.

Simulating a “true” synthetic 2016 population geo-located to household latitude-longitude points

To simulate a realistic population in Khomas, Namibia, we used all of the same population inputs and spatial auxiliary datasets as Thomson and colleagues (2018) [38]. Broadly, this involved the creation of three datasets—modelled surfaces of household types, manually digitised building point locations, and synthetic (simulated) households—then linked synthetic households to point locations based on the household type probability surfaces.

  1. Modelled surfaces of household types. Household types were defined from Namibia 2013 Demographic and Health Survey (DHS) data using k-means analysis with variables that were also present in the Namibia 2011 census (e.g., improved sanitation facilities, gender of head of household). Next, probability surfaces of these household types were created using a Random Forrest model and spatial covariates to interpolate the likelihood of a given household type across Namibia between DHS survey locations [38]. The probability surfaces of “urban poor” and “urban non-poor” household types were manually adjusted due to high misclassification. These adjustments were made by manually assigning the proportion of households in each census enumeration area (EA) that appeared to be located in areas of small disorganised buildings based on visual inspection of 30m Quickbird satellite imagery.

  2. Synthetic households. Separately, we modelled a synthetic population of individuals nested within households across Khomas from Namibia 2011 census microdata using an iterative proportional fitting model and conditional annealing [44].

  3. Building locations. A third set of data, building point locations, were manually digitised from 2014–2016 30cm Quickbird imagery in ArcGIS 10.

To link synthetic households with building locations, we calculated the most likely household type of each synthetic household using k-means analysis scores. Next, we iteratively assigned synthetic households (2) to building point locations (3) based on the probability of each household type at a given building point (1). Finally, using the manually classified EAs (with our estimated portion of urban poor households), we classified all urban households as being located in either a slum or non-slum area. All of these steps are detailed in Supplement 1 and the paper by Thomson and colleagues (2018) [38]. This simulated population is meant to represent a realistic “true” synthetic reference population for 2016.

Simulating realistic outdatedness of Khomas census population

To simulate population outdatedness in Khomas, we imported the above 2016 synthetic population household point locations into Google Earth, and used the software’s historical Maxar and SPOT imagery (40cm) to flag all buildings that were not present in 2011, 2006, and 2001 imagery. The oldest imagery available at 40cm resolution in Google Earth was from 2004, so we used some judgement to flag buildings that looked recently built in 2004 (e.g., bare fresh soil) and assumed they were not present in 2001. During this exercise, we ensured that the number of household coordinates in each constituency matched the number of households reported in the 2001 and 2011 Population and Housing Census final reports to ensure that both patterns and degree of outdatedness were realistic [39] (Fig 3). The synthetic population is provided in Supplement 2 and is comparable to the Oshikoto, Namibia 2016 synthetic population created by Thomson and colleagues [38].

Fig 3. Household point locations in Khomas, Namibia by presence in 2016, 2011, 2006, and 2001.

Fig 3

Sources: Constituency boundaries publically available from https://gadm.org/. Synthetic population latitude-longitude coordinates available in Supplement 2.

Simulating realistic levels of under-count inaccuracy in censuses

To identify realistic levels of under-counts among urban-slum, urban-non-slum, and rural populations in LMIC censuses, we reviewed the scientific and grey literature. The review included census post enumeration surveys (PESs) in 108 LMICs listed by the UN Statistical Division Census Programme website [8], and a systematic search in PubMed and Scopus of articles published between January 1, 1990 and February 28, 2017 using the following search criteria: “census AND (listing OR enumerat* OR count OR coverage OR miss*) AND (nomad* OR pastoral* OR refugee OR displaced OR migrant OR slum OR poorest OR unregistered OR homeless OR [street] sleeper OR pavement [dweller] OR floating)”. The first wave of the literature search resulted in 459 unique articles, of which co-author DRT screened all titles and abstracts. Of 72 potentially eligible articles from LMICs, DRT reviewed the full-text, and kept five which reported a census under-count. In a second wave, we used Google Scholar to identify the top 20 “cited by” and top 20 “related” articles for each of the five articles identified in the first wave. The second wave resulted in 334 unique articles, of which 49 had potentially relevant titles or abstracts. After a full-text review of these articles, we found eight additional reported census under-counts. Together, census under-counts in LMICs were collated from 10 PESs [4554], and 13 articles [10,5566] (Fig 4). The average census under-counts were 46% in urban-slum populations, 6% in urban-non-slum populations, and 7% in rural populations (Table 2, see Supplement 3 for details).

Fig 4. Search terms and process used in the census under-count literature review.

Fig 4

Table 2. Number of households simulated in the "true" synthetic population and 15 realistic scenarios of census outdatedness and inaccuracy, by household type.
Year No inaccuracy Low inaccuracy Medium inaccuracy High inaccuracy
2016 (current)
    Urban slum
    Urban non-slum
    Rural

35,001
57,843
4,823

31,500
56,677
4,735

24,500
54,942
4,590

14,000
52,073
4,326
2011 (5 years old)
    Urban slum
    Urban non-slum
    Rural

28,583
55,680
5,175

25,724
54,566
5,071

20,008
52,895
4,917

11,433
50,122
4,647
2006 (10 years old)
    Urban slum
    Urban non-slum
    Rural

18,018
49,742
4,146

16,216
48,747
4,063

12,612
47,258
3,935

7,207
44,769
3,730
2001 (15 years old)
    Urban slum
    Urban non-slum
    Rural

13,149
41,700
3,731

11,834
40,866
3,656

9,204
39,612
3,547

5,259
37,514
3,373

Low inaccuracy: missing 2% rural and urban-non-slum households, and 10% urban-slum households. Medium inaccuracy: missing 5% rural and urban-non-slum households, and 30% urban-slum households. High inaccuracy: missing 10% rural and urban-non-slum households, and 60% urban-slum households.

Based on these findings, we simulated three levels of census inaccuracy: low inaccuracy was considered to be missing 2% of rural and urban-non-slum households, and 10% of urban-slum households; medium inaccuracy was considered to be missing 5% of rural and urban-non-slum households, and 30% of urban-slum households; and finally, high inaccuracy was classified as missing 10% of rural and urban-non-slum households, and 60% of urban-slum households (Table 1). We applied the inaccuracy rates at random within rural, urban-slum, and urban-non-slum households such that there was no spatial pattern inherent to the simulated under-counts. This exercise resulted in one “true” and 15 outdated-inaccurate simulated populations which we used to generate realistic gridded population datasets that reflect typical gridded population estimates currently available across LMICs (Table 2).

Table 1. Range and average percent of population missing from LMIC censuses based on literature review.
Location Literature review findings Simulated inaccuracy
Minimum Average Maximum Low Medium High
Urban-slum 5% 46% 100% 10% 30% 60%
Urban-non-slum 2% 6% 15% 2% 5% 10%
Rural 2% 7% 13% 2% 5% 10%

Simulating realistic gridded population datasets

To simulate realistic gridded population datasets, we aggregated each of the simulated household populations to EA or constituency (second-level administrative unit) boundaries, and applied the WorldPop-Global-Unconstrained modelling technique (for a total of 32 datasets). We applied the WorldPop-Global-Unconstrained model in three phases as described in WorldPop’s method publication [22] (Fig 5, Table 3).

Fig 5. Overview of WorldPop-Global random forest modelling workflow.

Fig 5

(A) Each decision tree in the ensemble is built upon a random bootstrap sample of the log-transformed population and ancillary data by administrative unit. (B) Population density prediction for each cell ycell(x) is based on an average of the individual trees. (C) Predicted cell densities are normalized by administrative unit and used to dasymetrically disaggregate log-transformed administrative unit population, then transformed to predict population per cell.

Table 3. Covariate data sources for Random Forest gridded population estimates.
Name Description (Year) Original scale Original source
cov_road Distance to OSM major roads (2016) Vector, <30 m OpenStreetMap [68]
cov_intsec Distance to OSM major road intersections (2016) Vector, <30 m OpenStreetMap [68]
cov_waterw Distance to OSM major waterways (2016) Vector, <30 m OpenStreetMap [68]
cov_wdpa Distance to IUCN nature reserve (2000–17) 30” (~900 m) UNEP-WCMS & IUCN [69]
cov_viirs Resampled VIIRS night-time lights (2012–2016) 30” (~900 m) NOAA [70]
cov_dmsp Resampled DMSP-OLS night-time lights (2011) 30” (~900 m) NOAA & Zhang, et al. [71,72]
cov_tt50k Resampled travel time to cities of 50,000+ (2000) 30” (~900 m) Weiss, et al. [73]
cov_001 Distance to cultivated areas (2015) 9” (~300 m) ESA CCI–LC [74]
cov_040 Distance to woody areas (2015) 9” (~300 m) ESA CCI–LC [74]
cov_130 Distance to cultivated areas (2015) 9” (~300 m) ESA CCI–LC [74]
cov_140 Distance to herbaceous areas (2015) 9” (~300 m) ESA CCI–LC [74]
cov_150 Distance to sparse vegetation areas (2015) 9” (~300 m) ESA CCI–LC [74]
cov_160 Distance to aquatic vegetation areas (2015) 9” (~300 m) ESA CCI–LC [74]
cov_190 Distance to urban areas (2015) 9” (~300 m) ESA CCI–LC [74]
cov_200 Distance to bare areas (2015) 9” (~300 m) ESA CCI–LC [74]
cov_cciwat Distance to ESA-CCI-LC inland waterbodies (2000–12) 4.5” (~150 m) ESA CCI [75]
cov_slope SRTM-based slope (2000) 3” (~90 m) de Ferranti [76,77]
cov_topo SRTM-based elevation (2000) 3” (~90 m) de Ferranti [76,77]
cov_coast Distance to open-water coastline (2000–20) 3” (~90 m) CIESIN [78]
cov_ghsl Distance to urban area (2012) 1.26” (~38 m) Pesaresi, et al. [79]
cov_guf Distance to settlement built-up areas (2012) 2.8” (~84 m) DLR EOC [80]
cov_bsgme Distance to built settlement expansion (2016) 3” (~90 m) Nieves, et al. [81]
cov_prec Average total annual precipitation (1970–2000) 30” (~900 m) Fick and Hijmans [82]
cov_temp Average annual temperature (1970–2000) 30” (~900 m) Fick and Hijmans [82]

OSM: OpenStreetMap; VIIRS: Visible Infrared Imaging Radiometer Suite; DMSP-OLS: Defence Meteorological Satellite Program Operational Linescan System; ESA-CCI-LC: European Space Agency Climate Change Initiative Land Cover; UNEP-WSMS: UN Environment Programme World Conservation Monitoring Centre; IUCN: International Union for Conservation of Nature; NOAA: US National Oceanic and Atmospheric Administration; CIESIN: Center for International Earth Science Information Network; DLR EOC: German Aerospace Center Earth Observation Center.

  1. In the first phase (A), a non-parametric Random Forest ensemble machine-learning algorithm grows a “forest” of decision trees for each input unit (EA or constituency) [67]. Each Random Forest tree is a model of the potential relationships between multiple auxiliary covariates and census population counts. In the Random Forest modelling workflow, this is where model uncertainty is calculated–at the scale of the input population areal unit.

  2. In the second phase (B), all of the covariates are prepared in 100x100m cells. In this phase, the split values of each classification tree developed in phase A are used to parameterize corresponding regression models to predict population density within 100x100m cells [22]. For each cell, the predicted population values from all regression models are averaged to make a single population estimate, though these population estimates are not pycnophylactic, meaning that estimates in cells do not necessarily sum to the original areal unit population.

  3. Thus the WorldPop-Global-Unconstrained workflow involves a third phase (C) outside of the Random Forest model to normalize cell-level predicted population densities to preserve census input population counts [22].

Analysing cell-level accuracy

To empirically measure cell-level accuracy of the 32 gridded population datasets, we compared each cell-level estimate against the “true” synthetic point-level 2016 population count in that cell, then calculated root mean square error (RMSE), a measure of error magnitude that penalises large errors. This was performed on 100x100m cells, and then estimated cell population counts were aggregated and assessed for accuracy at 200x200m, 300x300m, and so on up to 1x1km. This was to test a common assumption that large model errors at fine geographic scale are “smoothed out” and become less severe as population estimates are aggregated across larger zones. To compare RMSE across cells of different geographic sizes, we normalised the statistic by average population (Eq 1) and by area (Eq 2). The former represents RMSE of population counts expressed as a portion of the population [83], while the latter represents RMSE of population density per hectare (100x100m unit) [84]. We evaluated RMSE in urban-slum, urban-non-slum, and rural cells separately. In the calculation of RMSE, yi is the “true” synthetic population count in cell i, y^i is the gridded population estimate in cell i, Di is the “true” synthetic population density per hectare, D^i is the estimated population density per hectare, and n is the number of grid cells.

Pop-adj RMSE = i = 1ny^i-yi2n ÷ i = 1n(yi)n 1
Area-adj RMSE = i = 1nD^i-Di2n 2

To better understand the mechanics of the WorldPop-Global-Unconstrained model and workflow, we calculated bias, a measure of error direction and magnitude. This metric was especially useful for the two gridded population datasets derived from “true” synthetic population counts because any inaccuracies would be related to the model and covariate datasets alone; and not inaccuracies in the input population counts. Bias (Eq 3) reveals to what extent cell-level estimates are systematically under- or over-estimated, and reflects over/under-counts in cells of different sizes that a user might encounter in the field. Relative bias (Eq 4) refers to bias normalised by the average synthetic population which enables comparisons across grid scales. As above, bias and relative bias were assessed in 100x100m cells as well as cell sizes that ranged up to 1x1km, and separately in urban versus rural areas.

Bias = i = 1n(y^i-yi)n 3
Relative bias = i = 1n(y^i-yi)n / i = 1n(yi)n 4

To assess the degree to which non-zero population estimates in the WorldPop-Global-Unconstrained dataset resulted in misallocation of population, a third statistic was calculated counting the entire modelled population in Khomas that was misallocated to cells which were unsettled according to the “true” synthetic population. For all statistics, we excluded gridded population cell-level estimates of less than 1 person to avoid millions of near-zero cell-level estimates in unsettled areas of Khomas (located outside of Windhoek) from dominating the accuracy assessments.

Results

Neither measure of RMSE differed substantially across the simulated outdated-inaccurate census scenarios (Figs 6 and 7). Furthermore, errors only slightly decreased when the input data were aggregated to EA (finer) rather than constituency (coarser) (Figs 6 and 7). The major driver of RMSE in cells was urban versus rural location, with further difference between urban-slum and urban-non-slum. In urban cells, population-adjusted RMSE was substantially smaller than rural cells (Fig 6), but much larger per hectare due to larger population numbers (Fig 7). In urban areas, RMSE per hectare was lowest in 100x100m cells (slum range: 32–72, non-slum range: 21–33), while in rural areas, RMSE per hectare was lowest in cells 300x300m to 500x500m (rural range: 2–54) (Fig 7). Results for select scenarios are presented in Fig 6 ranging from the synthetic “true” 2016 population to the most outdated (2001) and inaccurate (missing 10% to 60%) population, though tables of all results are provided in Supplement 4.

Fig 6. Population-adjusted root mean square error (RMSE) according to input population aggregation, a selection of scenarios, and cell size.

Fig 6

Fig 7. Population density root mean square error (RMSE) per hectare according to input population aggregation, a selection of scenarios, and cell size.

Fig 7

Assessment of bias in the two gridded population datasets that were derived from synthetic “true” 2016 population counts revealed systematic and substantial under-estimates of populations in urban-slum and urban-non-slum cells due to the aggregation-level of the input population data and modelling approach, and not inaccuracies in the input data (Tables 4 and 5). For example, the average 300x300m urban-slum cell under-estimated the population by more than 350 people (EA-level input) up to 500 people per cell (constituency-level input). For comparison, the average 300x300m non-slum cell was under-estimated by 165 people (constituency-level input) to 187 people (EA-level input), while the average rural cell of the same size was over-estimated by 3 people (constituency-level input) to 14 people (EA-level input) (Table 4). When adjusted for population, the results indicate that for every person estimated in an urban non-slum cell, 0.5 to 1 person is omitted; and for every person estimated in an urban slum cell, 0.75 to 1.5 people are omitted (Table 5).

Table 4. Bias in gridded population estimates derived from “true” synthetic population counts, by output grid cell size and urban/rural location (in cells > = 1 estimated person).

Cell size EA-level input Constituency-level input
Non-slum Slum Rural Non-slum Slum Rural
100 0 0 20 -4 -34 7
200 -71 -135 18 -64 -212 6
300 -187 -353 14 -165 -498 3
400 -346 -678 8 -303 -929 -1
500 -549 -1029 3 -483 -1401 -8
600 -769 -1480 -22 -672 -2080 -34
700 -1094 -2114 -33 -981 -2747 -51
800 -1410 -2692 -72 -1247 -3359 -90
900 -1728 -3215 -126 -1576 -4437 -152
1000 -1928 -4453 -126 -1770 -5834 -167

Table 5. Population-adjusted bias in gridded population estimates derived from “true” synthetic population counts, by output grid cell size and urban/rural location (in cells > = 1 estimated person).

Cell size EA-level input Constituency-level input
Non-slum Slum Rural Non-slum Slum Rural
100 0.00 0.00 3.36 -0.10 -0.64 1.28
200 -0.58 -0.74 1.90 -0.52 -1.16 0.67
300 -0.76 -0.94 1.07 -0.67 -1.32 0.23
400 -0.85 -1.04 0.52 -0.74 -1.43 -0.08
500 -0.92 -1.07 0.15 -0.81 -1.46 -0.40
600 -0.96 -1.09 -0.77 -0.84 -1.53 -1.21
700 -0.99 -1.15 -1.02 -0.89 -1.50 -1.59
800 -1.03 -1.21 -1.70 -0.91 -1.51 -2.12
900 -1.00 -1.14 -2.41 -0.91 -1.58 -2.92
1000 -1.05 -1.09 -2.45 -0.96 -1.43 -3.25

Table 6 summarises the percent of the estimated population misallocated to “truly” unsettled cells according to the synthetic population. For this analysis, no cells in the estimated population were excluded. Roughly 20% (EA-level input) or 10% (constituency-level input) of the population was misallocated to unsettled 100x100m cells (Table 6). However, as cells were aggregated, the percent of misallocated population dropped precipitously. For example, at 300x300m, approximately 2% (EA-level input) or 1% (constituency-level input) of Khomas’s population was misallocated to unsettled cells. This indicates that most of the population was disaggregated to unsettled cells within, or near to, settlements. The rates of misallocation were similar when cells with less than one person were excluded (not reported).

Table 6. Percent of the overall population that is misallocated to unsettled cells (no exclusion), by aggregation level of the input data and output grid cell size.

Grid cell size (m2) EA-Level Input Constituency-Level Input
100 20.8% 12.5%
200 5.0% 2.6%
300 2.2% 1.0%
400 1.3% 0.5%
500 0.8% 0.3%
600 0.6% 0.2%
700 0.4% 0.1%
800 0.3% 0.1%
900 0.3% 0.1%
1000 0.2% 0.1%

Discussion

This is among the first accuracy assessments of a top-down gridded population model at the grid-cell level, and the first that we know of in a LMIC setting. By developing a simulated realistic population and several scenarios of the population with realistic levels of outdatedness and inaccuracy, we were able to evaluate the accuracy of a gridded population model, as well as assess the impact of outdated-inaccurate census inputs on estimates. In this paper, we evaluated just one of several gridded population models–WorldPop-Global-Unconstrained. We also only analysed one simulated population and focused on the particular setting of Khomas, Namibia, so the results do no necessarily generalize to other cities or datasets. In this specific analysis, cell-level inaccuracies between urban versus rural areas dominated the results.

In practical terms, this massive difference between urban versus rural accuracy means that urban development indicators calculated with a WorldPop-Global-Unconstrained dataset at fine scale (e.g., neighbourhood) would likely be incorrect, and could lead to poorly informed decisions. For example, an underestimate of the number of people living in a neighbourhood could overestimate both vaccination coverage and disease infection rates. Contrary to what some might assume, there was limited evidence in this study that outdated or inaccurate census data played a major role in cell-level inaccuracy of gridded population estimates. Instead, we address three other potential sources of the cell-level inaccuracies observed.

The first issue is specific to the WorldPop-Global-Unconstrained modelling approach. In this approach, input administrative units with zero population are excluded and the remaining population counts are log-transformed before inclusion in a Random Forest model. While this procedure ensures that population counts are normally distributed during modelling, it also means that unpopulated cells are assigned a very small fraction of a person [22]. A possible concern is that non-zero population estimates across millions of unsettled cells could result in a sizable portion of the population being misallocated. Our analysis of misallocation, however, indicates that this phenomenon played only a minor role in cell-level inaccuracies, if at all. Table 6 demonstrates that even in this context of vast unsettled areas, only a small portion of Khomas’ population was misallocated to cells far from actual settlements. Nearly all of the population was estimated to be in cells within 200 to 300 metres of the “true” synthetic population.

Most global gridded population producers constrain estimates to settled cells as defined with a settlement layer (e.g. LandScan [24,85], GHP-POP [19,20], HRSL [21], GRID3 [28,86], WPE [26]). Until recently, these settlement layers tended to be relatively coarse (e.g. GHS-BUILT 1x1km [87]) and/or had a tendency to omit areas with few sparse buildings (e.g. GUF [80]) which could under-estimate the population in rural areas and over-estimate the population in urban areas. However, new free very high resolution Sentinel-2 imagery, and major leaps in computing power for extracting building footprints and other features from imagery, have enabled development of several new detailed settlement layers in the last few years (e.g., GHS-BUILT-S2 [88], Maxar/Ecopia [89]). Recently, WorldPop-Global produced a constrained global gridded population estimate for 2020 that uses the same input population and covariate datasets as its unconstrained model plus several building footprint metrics (in Africa), and then masks all 100x100m cells without building footprints (in Africa) or built settlement (rest of the world) [35], eliminating the issue of non-zero population estimates in unsettled cells.

The second potential source of inaccuracy relates to covariate resolution and the relationship of covariates with population density. This issue seems to have contributed more substantially to errors in this analysis, particularly within the city of Windhoek. A number of the Random Forest model covariates, such a land cover type and night-time lights, had an original resolution substantially coarser than 100x100m which could have resulted in a “halo” effect around settlements, causing populations to be disaggregated to cells near a settlement, but not directly over it. Table 5 provides evidence of this; the accuracy of the estimated population distribution, and correct allocation of population to settled cells, both performed well when the estimated population was aggregated to 300x300m or larger. Other covariates, such as distance to roads and intersection locations were available at very fine spatial resolution and thus were precise at the 100x100m scale. Although they are good indicators of a settlement, they are not necessarily good indicators of higher or lower population density within a settlement. The lack of fine-scale covariates associated with population density within cities and towns likely explains a portion of the cell-level error observed in Khomas’s urban population. Other issues that might further decrease local spatial accuracy are temporal miss-match of covariates [16] and covariate spatial autocorrelation [90]. With the recent release of several building footprint datasets (e.g., Maxar/Ecopia in most of Africa [89], Bing in Tanzania and Uganda [91]), several new covariate layers have been created by the WorldPop team including number of buildings and total area of buildings in 100x100m cells [92]. Building footprints are likely associated with population density within settlements and have a finer spatial resolution than 100x100m, making it a potentially powerful covariate to differentiate lower and higher population density within urban areas in any gridded population model. The WorldPop team, among other gridded population producers, is currently working to test and incorporate building footprint datasets into gridded population models.

The third potential source of cell-level inaccuracies is use of average population densities from large administrative units to estimate population densities in much smaller grid cells. This is known as the ecological fallacy [93], and probably played the largest role in cell-level inaccuracies, especially within urban areas. Population densities are used by the Random Forest model to establish relationships between covariates and population density (total population divided by total area), not population totals. Even with perfect covariates and exclusion of unsettled areas, this would mean that cells with high “true” synthetic population counts are likely to be severely underestimated because the geographic size of input population units are larger (and population densities are smaller) than the output grid cells. When this happens, population counts that are not allocated to the densest cells will instead be allocated to other less dense cells in the same input areal unit. Tables 4 and 5 provide strong evidence of this issue with the population in urban cells, especially urban-slum cells, systematically underestimated regardless of cell size.

Although these results apply only to the WorldPop-Global-Unconstrained model, we can speculate about how these results might apply to other gridded population datasets. Most top-down gridded population datasets use average population densities from large input areal units in some way to populate smaller grid cells, and are thus likely subject to similar errors linked with the ecological fallacy. The High Resolution Settlement Layer (HRSL), for example, uses uniform areal disaggregation of the population from input units (e.g., EA) to 30x30m grid cells which contain a building footprint [21], and the Global Human Settlement GHS-POP dataset takes a similar approach disaggregating input populations into 250x250m cells that are classified as settled [19,20]. Gridded Population of the World (GPWv4) is likely even less accurate at the cell-level because the population from each input unit (e.g., EA) are smoothed across all cells in that unit, including unsettled cells [17]. Gridded population datasets based on complex models with variable disaggregation from units to grid cells, such as LandScan [24] and World Population Estimates (WPE) [26], are instead subject to the second limitation described above because, like WorldPop-Global-Unconstrained, they lack high-resolution model covariates (e.g., building density) to accurately differentiate population density within settled cells.

This analysis reinforces findings of other studies which find that currently available gridded population products tend to underestimate populations in urban areas [9496], especially in higher-density poorer neighbourhoods [97]. For example, Tuholske and colleagues (2021) compared five gridded population products to estimate the proportion of population affected by natural disasters (SDG 11.5) in three regions where disasters had occurred, and found that 1x1km population estimates varied widely among data products, and reflected anywhere from 20% to 80% of the total UN estimated population in each region. Furthermore, they found that WorldPop-Global-Unconstrained generally performed better than un-modelled products (e.g., GPW), but not as well as products that constrained estimates to settled cells (e.g., GHS-POP) [94]. In a separate comparison of nine gridded population estimates of Kenyan and Nigerian slum populations (SDG 11.1.1) where field counts were available for reference, the estimated population in each slum varied widely and WorldPop-Global-Unconstrained estimates reflected just 11% of the overall slum population while the best performing data product (HRSL) estimated just 34% of all slum dwellers [97]. A key take-away from gridded population comparison studies is that fine-scale accuracy across data products varies substantially depending on location, potentially leading to different conclusions and decisions (e.g., about the humanitarian need or health care burden) depending on the gridded population dataset used for analysis. Furthermore, these studies underscore the need to understand fine-scale accuracy across gridded population datasets and locations to inform improvements to the underlying modelling methods and inputs.

Our analysis of a simulated population offers a methodological approach that can be replicated in other settings to evaluate the accuracy of any gridded population dataset at the cell-level. This analysis also points toward two solutions–use of building footprint covariates and finer-scale training data–that stand to improve cell-level accuracy of gridded population datasets derived from complex models, including all WorldPop-Global datasets as well as LandScan [24,25], WPE [26], and GRID3 [28,86]. Other techniques would be needed to improve the accuracy of gridded population datasets that do not vary (weight) population densities within areal units based on auxiliary information (e.g., HRSL [21], GHS-POP [19,20], GPW [17,18]).

Our first suggestion to improve WorldPop-Global datasets is to incorporate finer-scale training data into models to overcome the problem of larger areal-unit average values being used in smaller grid cells. In cases where the input areal units are geographically large, WorldPop-Global-Unconstrained (and Constrained) models incorporate training data from a neighbouring country that has finer-scale input population counts [22]. Our analysis showed, however, that even when relatively small geographic units (census EAs) were used as the input population area unit, urban slum and non-slum cell-level errors were still substantial, and cell-level accuracy with EA-level input was only marginally improved compared to constituency-level input (Fig 7). This suggests that finer-scale training data (e.g., closer to 100x100m) should be incorporated during the model training phase, particularly from high-density urban areas, to ensure that the WorldPop Random Forest model contains sufficiently large population density values to assign to urban cells. Fine-scale training datasets might come from existing household survey enumerations (e.g., World Bank Living Standards Measurement Surveys), or slum community profiles such as those published on the Know Your City Campaign website [98]. Even if fine-scale densities are only available for a small sample of locations, they would provide the model with more accurate maximum population values at the scale of 100x100m during model training.

The second potential solution is to incorporate more spatially detailed datasets into models which correlate with variations in population density. This analysis of WorldPop-Global-Unconstrained data raises broader questions about the cell-level accuracy of all gridded population estimates in urban areas, especially the densest parts of cities such as in slums, informal settlements, and neighbourhoods with high-rise apartment buildings [99101]. New datasets derived from very high resolution satellite imagery, in particular building footprints, are a promising new covariate to reduce the “halo” effect of populations misallocated nearby, but not directly over, the highest density cells. More work will be needed to improve building footprint datasets by distinguishing residential and non-residential buildings to avoid population being misallocated to business districts, factories, universities, airports, and other non-residential cells [102,103].

Conclusions

Global gridded population data initiatives aim to fill a gap in available disaggregated and current population counts to ensure that everyone is counted and that all needs are met in development initiatives. However, many gridded population datasets are not evaluated for accuracy at fine spatial scale. This analysis of one simulated population in one setting revealed substantial and systematic under-estimation of population in slums. Further analyses of other gridded population datasets are needed across diverse settings. However, if severe under-estimates in slums and other high-density urban areas are widespread, this means that gridded population datasets might unintentionally reinforce marginalisation of the urban poorest by omitting them from maps and population counts. We offer two suggestions to address this challenge: inclusion of finer-scale training data from household survey listings or “slum” enumerations, and the addition of new building footprints data as model covariates. Given the increased use of gridded population datasets for monitoring health and development outcomes in small areas, it is imperative that gridded population datasets area assessed for cell-level accuracy and are improved where possible.

Supporting information

S1 Table. Percent of population missing from LMIC censuses by source.

(DOCX)

S2 Table. Root Mean Square Error (RMSE) statistics for all scenarios.

(DOCX)

S1 File. Simulating a population in Khomas, Namibia.

(PDF)

S2 File. Simulated population in Khomas, Namibia.

(CSV)

Acknowledgments

We would like to thank Drs. Angela Luna Hernandez and Ryan Engstrom for their feedback on an earlier version of this work.

Data Availability

All relevant data are within the paper and its Supporting Information files.

Funding Statement

Dana R. Thomson was funded by the Economic and Social Research Council (ESRC) grant number ES/5500161/1 (more information at https://esrc.ukri.org/). ESRC played no role in the design, analysis, decision to publish, or preparation of this manuscript.

References

  • 1.UN Human Settlements Programme (UN-Habitat). World cities report 2020: the value of sustainable urbanization. Nairobi: UN-Habitat; 2020. 377 p. [Google Scholar]
  • 2.Utazi CE, Wagai J, Pannell O, Cutts FT, Rhoda DA, Ferrari MJ, et al. Geospatial variation in measles vaccine coverage through routine and campaign strategies in Nigeria: analysis of recent household surveys. Vaccine. 2020;38(14):3062–71. doi: 10.1016/j.vaccine.2020.02.070 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 3.Ruktanonchai CW, Ruktanonchai NW, Nove A, Lopes S, Pezzulo C, Bosco C, et al. Equality in maternal and newborn health: modelling geographic disparities in utilisation of care in five East African countries. PLoS One. 2016;11(8):e0162006. doi: 10.1371/journal.pone.0162006 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 4.Cutts FT, Ferrari MJ, Krause LK, Tatem AJ, Mosser JF. Vaccination strategies for measles control and elimination: time to strengthen local initiatives. BMC Med. 2021;19(1):1–8. doi: 10.1186/s12916-020-01843-z [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 5.Turok I, McGranahan G. Urbanization and economic growth: the arguments and evidence for Africa and Asia. Environ Urban. 2013;25(2):465–82. doi: 10.1177/0956247813490908 [DOI] [Google Scholar]
  • 6.Chen M, Zhang H, Liu W, Zhang W. The global pattern of urbanization and economic growth: Evidence from the last three decades. PLoS One. 2014;9(8):e103799. doi: 10.1371/journal.pone.0103799 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 7.United Nations Statistics Division (UNSD). 2020 world population and housing census programme [Internet]. Census dates for all countries. 2021 [cited 2021 Sep 29]. Available from: https://unstats.un.org/unsd/demographic-social/census/censusdates/.
  • 8.Bekele S. The accuracy of demographic data in the Ethiopian censuses. East Afr Soc Sci Res Rev. 2017;33(1):15–38. doi: 10.1353/eas.2017.0001 [DOI] [Google Scholar]
  • 9.Carr-Hill R. Missing millions and measuring development progress. World Dev. 2013;46:30–44. doi: 10.1016/j.worlddev.2012.12.017 [DOI] [Google Scholar]
  • 10.Ahonsi BA. Deliberate falsification and census-data in Nigeria. Afr Aff (Lond). 1988. Oct;87(349):553–62. [Google Scholar]
  • 11.Okolo A. The Nigerian census: problems and prospects. Am Stat. 1999;53(4):321–5. doi: 10.2307/2686050 [DOI] [Google Scholar]
  • 12.Yin S. Objections surface over Nigerian census results [Internet]. Population Reference Bureau. 2007. [cited 2021 Sep 29]. p. 1–3. Available from: www.prb.org/resources/objections-surface-over-nigerian-census-results/. [Google Scholar]
  • 13.United Nations Department of Economic and Social Affairs (UN-DESA). World Urbanization Prospects: The 2018 Revision [Internet]. 2018 [cited 2021 Sep 29]. Available from: https://population.un.org/wup/DataQuery/.
  • 14.Thomson DR, Rhoda DA, Tatem AJ, Castro MC. Gridded population survey sampling: a systematic scoping review of the field and strategic research agenda. Int J Health Geogr. 2020;19:34. doi: 10.1186/s12942-020-00230-4 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 15.POPGRID Data Collaborative. Leaving no one off the map: a guide for gridded population data for sustainable development [Internet]. New York NY USA; 2020. Available from: www.popgrid.org/sites/default/files/documents/Leaving_no_one_off_the_map.pdf.
  • 16.Leyk S, Gaughan AE, Adamo SB, de Sherbinin A, Balk D, Freire S, et al. Allocating people to pixels: a review of large-scale gridded population data products and their fitness for use. Earth Syst Sci Data Discuss. 2019;11:1385–409. doi: 10.5194/essd-11-1385-2019 [DOI] [Google Scholar]
  • 17.Doxsey-Whitfield E, MacManus K, Adamo SB, Pistolesi L, Squires J, Borkovska O, et al. Taking advantage of the improved availability of census data: a first look at the Gridded Population of the World, version 4. Pap Appl Geogr. 2015. Jul 3;1(3):226–34. doi: 10.1080/23754931.2015.1014272 [DOI] [Google Scholar]
  • 18.Center for International Earth Science Information Network (CIESIN), Columbia University. Gridded Population of the World v4 [Internet]. 2016 [cited 2021 Sep 29]. Available from: http://sedac.ciesin.columbia.edu/data/collection/gpw-v4/sets/browse.
  • 19.Pesaresi M, Ehrlich D, Florczyk AJ, Freire S, Julea A, Kemper T, et al. Operating procedure for the production of the Global Human Settlement Layer from Landsat data of the epochs 1975, 1990, 2000, and 2014 [Internet]. Ispra Italy: European Commission Joint Research Centre; 2016. 67 p. Available from: http://publications.jrc.ec.europa.eu/repository/handle/JRC97705. [Google Scholar]
  • 20.European Commission Joint Research Centre (EC-JRC). Global human settlement population model (GHS-POP) [Internet]. 2020 [cited 2021 Sep 29]. Available from: https://ghsl.jrc.ec.europa.eu/data.php.
  • 21.Facebook Connectivity Lab, CIESIN—Columbia University. High Resolution Settlement Layer (HRSL) [Internet]. 2016 [cited 2021 Sep 29]. Available from: https://data.humdata.org/dataset/highresolutionpopulationdensitymaps.
  • 22.Stevens FR, Gaughan AE, Linard C, Tatem AJ. Disaggregating census data for population mapping using random forests with remotely-sensed and ancillary data. PLoS One. 2015;10(2):e0107042. doi: 10.1371/journal.pone.0107042 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 23.WorldPop. Population Counts 2000–2020 UN-Adjusted Unconstrained 100m [Internet]. 2020 [cited 2021 Sep 29]. Available from: www.worldpop.org/doi/10.5258/SOTON/WP00660.
  • 24.Dobson JE, Bright EA, Coleman PR, Worley BA, Bright EA, Coleman PR, et al. LandScan: a global population database for estimating populations at risk. Photogramm Eng Remote Sensing. 2000. Jul;66(7):849–57. [Google Scholar]
  • 25.Oak Ridge National Laboratories. LandScan Data Availability [Internet]. 2017 [cited 2021 Sep 29]. Available from: www.ornl.gov.
  • 26.Frye C, Nordstrand E, Wright DJ, Terborgh C, Foust J. Using classified and unclassified land cover data to estimate the footprint of human settlement. Data Sci J. 2018;17:1–12. doi: 10.5334/dsj-2018-020 [DOI] [Google Scholar]
  • 27.Long JF, McMillen DB. A survey of census bureau population projection methods. Clim Change. 1987;11:141–77. doi: 10.1007/BF00138799 [DOI] [PubMed] [Google Scholar]
  • 28.Leasure DR, Jochem WC, Weber EM, Seaman V, Tatem AJ. National population mapping from sparse survey data: a hierarchical Bayesian modeling framework to account for uncertainty. Proc Natl Acad Sci U S A. 2020;117(39):24173–9. doi: 10.1073/pnas.1913050117 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 29.Leasure DR, Dooley CA, Bondarenko M, Tatem AJ. peanutButter: an R package to produce rapid-response gridded population estimates from building footprints, version 0.3.0 [Internet]. 2020. [cited 2021 Sep 29]. Available from: https://apps.worldpop.org/peanutButter/. [Google Scholar]
  • 30.Hay S, Noor A, Nelson A, Tatem A. The accuracy of human population maps for public health application. Trop Med Int Heal. 2005;10:1073–86. doi: 10.1111/j.1365-3156.2005.01487.x [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 31.Gaughan AE, Stevens FR, Linard C, Jia P, Tatem AJ. High resolution population distribution maps for Southeast Asia in 2010 and 2015. PLoS One. 2013;8(2):e55882. doi: 10.1371/journal.pone.0055882 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 32.Bondarenko M, Nieves JJ, Stevens FR, Gaughan AE, Tatem A, Sorichetta A. wpgpRFPMS: random forests population modelling R scripts, version 0.1.0 [Internet]. Southampton UK; 2020. Available from: doi: 10.5258/SOTON/WP00665 [DOI] [Google Scholar]
  • 33.Lloyd CT, Chamberlain H, Kerr D, Yetman G, Pistolesi L, Stevens FR, et al. Global spatio-temporally harmonised datasets for producing high-resolution gridded population distribution datasets. Big Earth Data. 2019;3(2):108–39. doi: 10.1080/20964471.2019.1625151 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 34.WorldPop. WorldPop-Global covariates [Internet]. 2020 [cited 2021 Sep 29]. Available from: https://www.worldpop.org/project/categories?id=14.
  • 35.WorldPop. Top-down estimation modelling: constrained vs unconstrained [Internet]. 2020 [cited 2021 Sep 29]. Available from: www.worldpop.org/methods/top_down_constrained_vs_unconstrained.
  • 36.United Nations Statistics Division (UNSD). Report on the results of a survey on census methods used by countries in the 2010 census round [Internet]. New York NY USA; 2010. (Working paper). Report No.: UNSD/DSSB/1. Available from: http://unstats.un.org/unsd/census2010.htm.
  • 37.Cobham A. Uncounted: power, inequalities and the post-2015 data revolution. Development. 2014;57(3–4):320–37. doi: 10.1057/dev.2015.28 [DOI] [Google Scholar]
  • 38.Thomson DR, Kools L, Jochem WC. Linking synthetic populations to household geolocations: a demonstration in Namibia. Data. 2018;3(3):30. doi: 10.3390/data3030030 [DOI] [Google Scholar]
  • 39.Namibia Statistics Agency (NSA). Namibia 2011 Population and Housing Census main report [Internet]. Windhoek Namibia; 2011. Available from: https://cms.my.na/assets/documents/p19dmn58guram30ttun89rdrp1.pdf.
  • 40.Newaya TP. Rapid urbanization and its influence on the growth of informal settlements in Windhoek, Namibia. MSc Thesis, Cape Peninsula University of Technology. 2010. Available from: http://etd.cput.ac.za/handle/20.500.11838/1451.
  • 41.Lai S, Erbach-Schoenberg E zu, Pezzulo C, Ruktanonchai NW, Sorichetta A, Steele J, et al. Exploring the use of mobile phone data for national migration statistics. Palgrave Commun. 2019;5(1):34. doi: 10.1057/s41599-019-0242-9 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 42.Olivier M. Migration in Namibia: a country profile 2015. Geneva: International Organization for Migration (IOM); 2015. 174 p. [Google Scholar]
  • 43.WorldPop. Africa 1km internal migration flows [Internet]. 2016 [cited 2021 Sep 29]. Available from: www.worldpop.org/geodata/summary?id=1281.
  • 44.Alfons A, Kraft S, Templ M, Filzmoser P. Simulation of close-to-reality population data for household surveys with application to EU-SILC. Stat Methods Appl. 2011;20(3):383–407. doi: 10.1007/s10260-011-0163-2 [DOI] [Google Scholar]
  • 45.Oliveira LC de S, Freitas MPS de, Dias MRML, Nascimento CMF, Mattos E da S, Junior JJAR. Censo Demográfico 2000—pesquisa de avaliação da cobertura da coleta [Internet]. Rio de Janeiro; 2003. Available from: https://biblioteca.ibge.gov.br/biblioteca-catalogo.html?id=21402&view=detalhes.
  • 46.Korale RBM. Post Enumeration Survey 2001 [Nepal Population Census] Draft Report [Internet]. Kathmandu; 2002 [cited 2019 Jan 20]. Available from: https://nepal.unfpa.org/sites/default/files/pub-pdf/PopulationMonograph2014Volume1.pdf.
  • 47.Maro R. Post enumeration survey Tanzania experience [Internet]. Workshop on the 2010 World programme on population and housing censuses: census evaluation and post enumeration surveys, for English-speaking African countries. 2009 [cited 2021 Sep 29]. p. 12. Available from: https://unstats.un.org/unsd/demographic/meetings/wshops/Ethiopia_14_Sept_09/Country_Presentations/Tanzania.ppt.
  • 48.Uganda Bureau of Statistics (UBS). Post enumeration survey: 2002 Uganda population and housing census [Internet]. Entebbe Uganda; 2005 [cited 2021 Sep 29]. Available from: www.ubos.org/wp-content/uploads/publications/03_20182002_CensusPopnSizeGrowthAnalyticalReport.pdf.
  • 49.Ghana Statistical Service (GSS). 2010 Population and Housing Census Post Enumeration Survey Report [Internet]. Accra Ghana; 2012 [cited 2021 Sep 29]. Available from: www2.statsghana.gov.gh/docfiles/2010phc/2010_PHC_PES_Report.pdf.
  • 50.Central Statistical Office (CSO). [Zambia] 2010 Census of Population and Housing Post Enumeration Survey (PES) [Internet]. Lusaka Zambia; 2013 [cited 2021 Sep 29]. Available from: https://web.archive.org/web/20151113170741/ http://www.zamstats.gov.zm/report/Census/2010/National/2010%20Census%20Post%20Enumeration%20Report.pdf.
  • 51.Bangladesh Institute of Development Studies (BIDS). Report of the post enumeration check (PEC) of the [Bangladesh] Population and Housing Census, 2011 [Internet]. Dhaka Bangladesh; 2012 [cited 2021 Sep 29]. Available from: http://203.112.218.65:8008/WebTestApplication/userfiles/Image/LatestReports/PEC%20Report%202011.pdf.
  • 52.National Statistical Commission (NSC). Census of India 2011: Report on post enumeration survey [Internet]. New Delhi India; 2014 [cited 2021 Sep 29]. Available from: https://censusindia.gov.in/nada/index.php/catalog/1366.
  • 53.Statistics South Africa (SSA). Census 2011 post-enumeration survey [Internet]. Pretoria South Africa; 2012 [cited 2021 Sep 29]. Available from: www.datafirst.uct.ac.za/dataportal/index.php/catalog/485/download/8289.
  • 54.National Institute of Statistics of Rwanda (NISR). Post enumeration survey report: fourth Population and Housing Census, Rwanda, 2012 [Internet]. Kigali Rwanda; 2010 [cited 2021 Sep 29]. Available from: www.statistics.gov.rw/publication/rphc4-post-enumeration-survey.
  • 55.Agarwal S. The state of urban health in India: comparing the poorest quartile to the rest of the urban population in selected states and cities. Environ Urban. 2011;23(1):13–28. doi: 10.1177/0956247811398589 [DOI] [Google Scholar]
  • 56.Carr-Hill R. Improving population and poverty estimates with citizen surveys: evidence from East Africa. World Dev. 2017;93:249–59. doi: 10.1016/j.worlddev.2016.12.017 [DOI] [Google Scholar]
  • 57.Ebenstein A, Zhao Y. Tracking rural-to-urban migration in China: lessons from the 2005 inter-census population survey. Popul Stud (NY). 2015;69(3):337–53. doi: 10.1080/00324728.2015.1065342 [DOI] [PubMed] [Google Scholar]
  • 58.Gidado SO, Nguku PJ, Ndadilnasiya Waziri M, Ohuabunwo C, Etsano A, Mahmud MZ, et al. Polio field census and vaccination of underserved populations Northern Nigeria, 2012–2013. Morb Mortal Wkly Rep. 2013;62(33):663–5. [PMC free article] [PubMed] [Google Scholar]
  • 59.Gurgel RQ, Da Fonseca JDC, Neyra-Castañeda D, Gill G V., Cuevas LE. Capture-recapture to estimate the number of street children in a city in Brazil. Arch Dis Child. 2004;89:222–4. doi: 10.1136/adc.2002.023481 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 60.Jiang Q, Li X, Sánchez-Barricarte JJ. Data uncertainties in China’s population. Asian Soc Sci. 2015;11(13):200–5. doi: 10.5539/ass.v11n13p200 [DOI] [Google Scholar]
  • 61.Karanja I. An enumeration and mapping of informal settlements in Kisumu, Kenya, implemented by their inhabitants. Environ Urban. 2010;22(1):217–39. doi: 10.1177/0956247809362642 [DOI] [Google Scholar]
  • 62.Kronenfeld DA. Afghan refugees in Pakistan: not all refugees, not always in Pakistan, not necessarily Afghan? J Refug Stud. 2008;21(1):43–63. doi: 10.1093/jrs/fem048 [DOI] [Google Scholar]
  • 63.Lucci P, Bhatkal T, Khan A. Are we underestimating urban poverty? World Dev. 2018;103:297–310. doi: 10.1016/j.worlddev.2017.10.022 [DOI] [Google Scholar]
  • 64.Sabry S. How poverty is underestimated in Greater Cairo, Egypt. Environ Urban. 2010;22(2):523–41. doi: 10.1177/0956247810379823 [DOI] [Google Scholar]
  • 65.Stark L, Rubenstein BL, Pak K, Taing R, Yu G, Kosal S, et al. Estimating the size of the homeless adolescent population across seven cities in Cambodia. BMC Med Res Methodol. 2017;17:1–8. doi: 10.1186/s12874-017-0293-9 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 66.Treiman DJ, Mason WM, Lu Y, Pan Y, Qi Y, Song S. Observations on the design and implementation of sample surveys in China [Internet]. Los Angeles; 2005. Report No.: CCPR-006-05. Available from: http://papers.ccpr.ucla.edu/index.php/pwp/article/download/PWP-CCPR-2005-006/405.
  • 67.Breiman L. Random forests. Mach Learn. 2001;45:5–32. [Google Scholar]
  • 68.OpenStreetMap contributors. OpenStreetMap base data [Internet]. 2000 [cited 2021 Sep 29]. Available from: www.openstreetmap.org.
  • 69.United Nations Environment Programme-World Conservation Monitoring Centre (UNEP-WCMS), International Union for Conservation of Nature (IUCN). World database on protected areas & Global database on protected areas management effectiveness [Internet]. UNEP-WCMS & IUCN. 2016 [cited 2021 Sep 29]. Available from: www.protectedplanet.net.
  • 70.[USA] National Oceanic and Atmospheric Administration (NOAA). VIIRS nighttime lights [Internet]. 2012 [cited 2021 Sep 29]. Available from: www.ncei.noaa.gov/maps/VIIRS_DNB_nighttime_imagery.
  • 71.[USA]National Oceanic and Atmospheric Administration (NOAA). Version 4 DMSP-OLS Nighttime Lights Time Series [Internet]. 2017 [cited 2021 Sep 29]. Available from: www.ngdc.noaa.gov/eog/dmsp/downloadV4composites.html.
  • 72.Zhang Q, Pandey B, Seto KC. A robust method to generate a consistent time series from DMSP / OLS nighttime light data. IEEE Trans Geosci Remote Sens. 2016;54(10):5821–31. doi: 10.1109/AUTEST.2006.283598 [DOI] [Google Scholar]
  • 73.Weiss D, Nelson A, Gibson H, Temperley W, Peedell S, Lieber A, et al. A global map of travel time to cities to assess inequalities in accessibility in 2015. Nature. 2018;553(7688):333–6. doi: 10.1038/nature25181 [DOI] [PubMed] [Google Scholar]
  • 74.European Space Agency—Climate Change Initiative (ESA-CCI). Land Cover CCI Product—Annual LC maps from 2000 to 2015 (v2.0.7) [Internet]. 2017 [cited 2021 Sep 29]. Available from: http://maps.elie.ucl.ac.be/CCI/viewer/.
  • 75.European Space Agency—Climate Change Initiative (ESA-CCI). Land cover CCI product—MERIS Waterbody product v4.0 (150 m) [Internet]. 2017 [cited 2021 Sep 29]. Available from: http://maps.elie.ucl.ac.be/CCI/viewer/.
  • 76.de Ferranti J. Digital elevation data—Viewfinder panoramas [Internet]. 2017 [cited 2021 Sep 29]. Available from: www.viewfinderpanoramas.org/dem3.html.
  • 77.de Ferranti J. Digital elevation data: SRTM void fill—Viewfinder panoramas [Internet]. 2017 [cited 2021 Sep 29]. Available from: www.viewfinderPanoramas.org/voidfill.html.
  • 78.Center for International Earth Science Information Network—CIESIN—Columbia University. Gridded Population of the World, Version 4.11 (GPWv4.11) [Internet]. 2018 [cited 2021 Sep 29]. Available from: 10.7927/H4F47M65. [DOI]
  • 79.European Commission. Global human settlement city model (GHS-SMOD) [Internet]. 2017 [cited 2021 Sep 29]. Available from: https://ghsl.jrc.ec.europa.eu/download.php.
  • 80.DLR Earth Observation Center. Global Urban Footprint (GUF) [Internet]. 2017 [cited 2021 Sep 29]. Available from: www.dlr.de/eoc/en/desktopdefault.aspx/tabid-11725/20508_read-47944/.
  • 81.Nieves JJ, Sorichetta A, Linard C, Bondarenko M, Steele JE, Stevens FR, et al. Annually modelling built-settlements between remotely-sensed observations using relative changes in subnational populations and lights at night. Comput Environ Urban Syst. 2020;80:101444. doi: 10.1016/j.compenvurbsys.2019.101444 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 82.Fick SE, Hijmans RJ. WorldClim 2: new 1-km spatial resolution climate surfaces for global land areas. Int J Climatol. 2017;37(12):4302–15. doi: 10.1002/joc.5086 [DOI] [Google Scholar]
  • 83.Gregory IN. An evaluation of the accuracy of the areal interpolation of data for the analysis of long-term change in England and Wales. In: GeoComputation [Internet]. Greenwich UK; 2000. Available from: www.geocomputation.org/2000/GC045/Gc045.htm.
  • 84.Bozheva AM, Petrov AN, Sugumaran R. The effect of spatial resolution of remotely sensed data in dasymetric mapping of residential areas. GIScience Remote Sens. 2005;42(2):113–30. doi: 10.2747/1548-1603.42.2.113 [DOI] [Google Scholar]
  • 85.Oak Ridge National Laboratories (ORNL). LandScan documentation [Internet]. 2017 [cited 2021 Sep 29]. Available from: https://landscan.ornl.gov/about.
  • 86.CIESIN, UNFPA, WorldPop, Flowminder. Geo-Referenced Infrastructure and Demographic Data for Development (GRID3) [Internet]. 2018 [cited 2021 Sep 29]. Available from: www.grid3.org.
  • 87.European Commission Joint Research Centre. GHS-BUILT [Internet]. 2019 [cited 2021 Sep 29]. Available from: https://ghsl.jrc.ec.europa.eu/ghs_bu2019.php.
  • 88.Corbane C, Sabo F, Politis P, Syrris V. HS-BUILT-S2 R2020A - GHS built-up grid, derived from Sentinel-2 global image composite for reference year 2018 using Convolutional Neural Networks (GHS-S2Net). European Commission, Joint Research Centre (JRC); 2020. [Google Scholar]
  • 89.Maxar. Satellite Imagery [Internet]. 2019 [cited 2021 Sep 29]. Available from: www.maxar.com/products/satellite-imagery.
  • 90.Sinha P, Gaughan AE, Stevens FR, Nieves JJ, Sorichetta A, Tatem AJ. Assessing the spatial sensitivity of a random forest model: application in gridded population modeling. Comput Environ Urban Syst. 2019;75:132–45. doi: 10.1016/j.compenvurbsys.2019.01.006 [DOI] [Google Scholar]
  • 91.Microsoft. Building Footprints [Internet]. AI for Humanitarian Action program. 2020 [cited 2021 Sep 29]. Available from: www.microsoft.com/en-us/maps/building-footprints.
  • 92.Dooley CA, Leasure DR, Boo G, Tatem AJ. Gridded maps of building patterns throughout sub-Saharan Africa, version 2.0 [Internet]. WorldPop. 2021. [cited 2021 Sep 29]. Available from: https://wopr.worldpop.org/?/Buildings. [Google Scholar]
  • 93.Selvin HC. Durkheim’s suicide and problems of empirical research. Am J Sociol. 1958;63(6):607–19. [Google Scholar]
  • 94.Tuholske C, Gaughan AE, Sorichetta A, de Sherbinin A, Bucherie A, Hultquist C, et al. Implications for tracking SDG indicator metrics with gridded population data. Sustain. 2021;13(13). doi: 10.3390/su13137329 [DOI] [Google Scholar]
  • 95.Yin X, Li P, Feng Z, Yang Y, You Z, Xiao C. Which gridded population data product is better? Evidences from mainland southeast Asia (MSEA). ISPRS Int J Geo-Information. 2021;10(10). doi: 10.3390/ijgi10100681 [DOI] [Google Scholar]
  • 96.Archila Bustos MF, Hall O, Niedomysl T, Ernstson U. A pixel level evaluation of five multitemporal global gridded population datasets: a case study in Sweden, 1990–2015. Popul Environ. 2020;42(2):255–77. doi: 10.1007/s11111-020-00360-8 [DOI] [Google Scholar]
  • 97.Thomson DR, Gaughan AE, Stevens FR, Yetman G, Elias P, Chen R. Evaluating the accuracy of gridded population estimates in slums: a case study in Nigeria and Kenya. Urban Sci. 2021;5(2):48. doi: 10.3390/urbansci5020048 [DOI] [Google Scholar]
  • 98.Slum/Shack Dwellers International (SDI). Know Your City [Internet]. 2016 [cited 2021 Sep 29]. Available from: https://sdinet.org/explore-our-data/.
  • 99.Nuissl H, Heinrichs D. Slums: perspectives on the definition, the appraisal and the management of an urban phenomenon. J Geogr Soc Berlin. 2013;144(2):105–16. doi: 10.12854/erde-144-8 [DOI] [Google Scholar]
  • 100.Ezeh A, Oyebode O, Satterthwaite D, Chen Y, Ndugwa R, Sartori J, et al. The history, geography, and sociology of slums and the health problems of people who live in slums. Lancet. 2017;389:547–58. doi: 10.1016/S0140-6736(16)31650-6 [DOI] [PubMed] [Google Scholar]
  • 101.Mahabir R, Croitoru A, Crooks A, Agouris P, Stefanidis A. A critical review of high and very high-resolution remote sensing approaches for detecting and mapping slums: trends, challenges and emerging opportunities. Urban Sci. 2018;2:8. doi: 10.3390/urbansci2010008 [DOI] [Google Scholar]
  • 102.Sturrock HJW, Woolheater K, Bennett AF, Andrade-Pacheco R, Midekisa A. Predicting residential structures from open source remotely enumerated data using machine learning. PLoS One. 2018;13(9):e0204399. doi: 10.1371/journal.pone.0204399 [DOI] [PMC free article] [PubMed] [Google Scholar]
  • 103.Lloyd CT, Sturrock HJW, Leasure DR, Jochem WC, Lázár AN, Tatem AJ. Using GIS and machine learning to classify residential status of urban buildings in low and middle income settings. Remote Sens. 2020;12(23):3847. doi: 10.3390/rs12233847 [DOI] [Google Scholar]

Decision Letter 0

Krishna Prasad Vadrevu

28 Jul 2021

PONE-D-21-16343

How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?: A simulation analysis in urban Namibia

PLOS ONE

Dear Dr. Thomson,

Thank you for submitting your manuscript to PLOS ONE. After careful consideration, we feel that it has merit but does not fully meet PLOS ONE’s publication criteria as it currently stands. Therefore, we invite you to submit a revised version of the manuscript that addresses the points raised during the review process.

Please submit your revised manuscript by Sep 10 2021 11:59PM. If you will need more time than this to complete your revisions, please reply to this message or contact the journal office at plosone@plos.org. When you're ready to submit your revision, log on to https://www.editorialmanager.com/pone/ and select the 'Submissions Needing Revision' folder to locate your manuscript file.

Please include the following items when submitting your revised manuscript:

  • A rebuttal letter that responds to each point raised by the academic editor and reviewer(s). You should upload this letter as a separate file labeled 'Response to Reviewers'.

  • A marked-up copy of your manuscript that highlights changes made to the original version. You should upload this as a separate file labeled 'Revised Manuscript with Track Changes'.

  • An unmarked version of your revised paper without tracked changes. You should upload this as a separate file labeled 'Manuscript'.

If you would like to make changes to your financial disclosure, please include your updated statement in your cover letter. Guidelines for resubmitting your figure files are available below the reviewer comments at the end of this letter.

If applicable, we recommend that you deposit your laboratory protocols in protocols.io to enhance the reproducibility of your results. Protocols.io assigns your protocol its own identifier (DOI) so that it can be cited independently in the future. For instructions see: http://journals.plos.org/plosone/s/submission-guidelines#loc-laboratory-protocols. Additionally, PLOS ONE offers an option for publishing peer-reviewed Lab Protocol articles, which describe protocols hosted on protocols.io. Read more information on sharing protocols at https://plos.org/protocols?utm_medium=editorial-email&utm_source=authorletters&utm_campaign=protocols.

We look forward to receiving your revised manuscript.

Kind regards,

Krishna Prasad Vadrevu, Ph.D

Academic Editor

PLOS ONE

Journal requirements:

When submitting your revision, we need you to address these additional requirements.

1. Please ensure that your manuscript meets PLOS ONE's style requirements, including those for file naming. The PLOS ONE style templates can be found at

https://journals.plos.org/plosone/s/file?id=wjVg/PLOSOne_formatting_sample_main_body.pdf and

https://journals.plos.org/plosone/s/file?id=ba62/PLOSOne_formatting_sample_title_authors_affiliations.pdf

2.Please note that in order to use the direct billing option the corresponding author must be affiliated with the chosen institute. Please either amend your manuscript to change the affiliation or corresponding author, or email us at plosone@plos.org with a request to remove this option.

3. We note that Figures 1 &3  in your submission contain [map/satellite] images which may be copyrighted. All PLOS content is published under the Creative Commons Attribution License (CC BY 4.0), which means that the manuscript, images, and Supporting Information files will be freely available online, and any third party is permitted to access, download, copy, distribute, and use these materials in any way, even commercially, with proper attribution. For these reasons, we cannot publish previously copyrighted maps or satellite images created using proprietary data, such as Google software (Google Maps, Street View, and Earth). For more information, see our copyright guidelines: http://journals.plos.org/plosone/s/licenses-and-copyright.

We require you to either (1) present written permission from the copyright holder to publish these figures specifically under the CC BY 4.0 license, or (2) remove the figures from your submission:

a. You may seek permission from the original copyright holder of Figures 1 & 3 to publish the content specifically under the CC BY 4.0 license. 

We recommend that you contact the original copyright holder with the Content Permission Form (http://journals.plos.org/plosone/s/file?id=7c09/content-permission-form.pdf) and the following text:

“I request permission for the open-access journal PLOS ONE to publish XXX under the Creative Commons Attribution License (CCAL) CC BY 4.0 (http://creativecommons.org/licenses/by/4.0/). Please be aware that this license allows unrestricted use and distribution, even commercially, by third parties. Please reply and provide explicit written permission to publish XXX under a CC BY license and complete the attached form.”

Please upload the completed Content Permission Form or other proof of granted permissions as an "Other" file with your submission.

In the figure caption of the copyrighted figure, please include the following text: “Reprinted from [ref] under a CC BY license, with permission from [name of publisher], original copyright [original copyright year].”

b. If you are unable to obtain permission from the original copyright holder to publish these figures under the CC BY 4.0 license or if the copyright holder’s requirements are incompatible with the CC BY 4.0 license, please either i) remove the figure or ii) supply a replacement figure that complies with the CC BY 4.0 license. Please check copyright information on all replacement figures and update the figure caption with source information. If applicable, please specify in the figure caption text when a figure is similar but not identical to the original image and is therefore for illustrative purposes only.

The following resources for replacing copyrighted map figures may be helpful:

USGS National Map Viewer (public domain): http://viewer.nationalmap.gov/viewer/

The Gateway to Astronaut Photography of Earth (public domain): http://eol.jsc.nasa.gov/sseop/clickmap/

Maps at the CIA (public domain): https://www.cia.gov/library/publications/the-world-factbook/index.html and https://www.cia.gov/library/publications/cia-maps-publications/index.html

NASA Earth Observatory (public domain): http://earthobservatory.nasa.gov/

Landsat: http://landsat.visibleearth.nasa.gov/

USGS EROS (Earth Resources Observatory and Science (EROS) Center) (public domain): http://eros.usgs.gov/#

Natural Earth (public domain): http://www.naturalearthdata.com/

Additional Editor Comments (if provided):

Dear Authors,

First, thank you for submitting the manuscript to PLOS ONE. We have received reviews from two different experts with one major and another minor revision. Based on the suggestions, we ask you to submit a revised manuscript. Specifically, please see that the revised version includes clarification on the a). possibilities of extrapolating the results to other regions; b). the usefulness of the research for applications; c). Use of RMSE versus MAE while evaluating the gridded population datasets; d). Figures improvement, etc.

We look forward to a revised version.

Best,

Krishn

[Note: HTML markup is below. Please do not edit.]

Reviewers' comments:

Reviewer's Responses to Questions

Comments to the Author

1. Is the manuscript technically sound, and do the data support the conclusions?

The manuscript must describe a technically sound piece of scientific research with data that supports the conclusions. Experiments must have been conducted rigorously, with appropriate controls, replication, and sample sizes. The conclusions must be drawn appropriately based on the data presented.

Reviewer #1: Yes

Reviewer #2: Yes

**********

2. Has the statistical analysis been performed appropriately and rigorously?

Reviewer #1: Yes

Reviewer #2: Yes

**********

3. Have the authors made all data underlying the findings in their manuscript fully available?

The PLOS Data policy requires authors to make all data underlying the findings described in their manuscript fully available without restriction, with rare exception (please refer to the Data Availability Statement in the manuscript PDF file). The data should be provided as part of the manuscript or its supporting information, or deposited to a public repository. For example, in addition to summary statistics, the data points behind means, medians and variance measures should be available. If there are restrictions on publicly sharing data—e.g. participant privacy or use of data from a third party—those must be specified.

Reviewer #1: No

Reviewer #2: Yes

**********

4. Is the manuscript presented in an intelligible fashion and written in standard English?

PLOS ONE does not copyedit accepted manuscripts, so the language in submitted articles must be clear, correct, and unambiguous. Any typographical or grammatical errors should be corrected at revision, so please note any specific errors here.

Reviewer #1: No

Reviewer #2: Yes

**********

5. Review Comments to the Author

Please use the space provided to explain your answers to the questions above. You may also include additional comments for the author, including concerns about dual publication, research ethics, or publication ethics. (Please upload your review as an attachment if it exceeds 20,000 characters)

Reviewer #1: I find the topic interesting and the authors show to be experts in the topic. However, I think that the paper is too technical for most potential readers. I provide some simple comments hoping they can help in this regard.

1.The abstract is extremely and unnecessarily long.

2. I find the intro too technical. I recommend leaving technical thing for other sections and devoting more in the intro to tell the reader i) why is the topic relevant for policy debates, ii) what are the main contribution of the current paper?

3. I like the idea of a short and synthetic paper. However, even in a paper of this style, I think framing the paper in the relevant literature is essential. There are several papers on urbanisation, urban density, suburbanisation, etc. worldwide that the paper should cite and relate to. I recommend recent papers in the Journal of Economic Geography, Journal of Development Studies and in the Journal of Urban Economics.

4. Could the text be easier to read, leaving some technicalities for an appendix?

5. Sorry but I find really puzzling the use of “simulated” and “true” in the same sentences over and over in the paper to refer to the same numbers. How can be, at the same time, “simulate” and “true”?!

6. How could we extrapolate the findings for Namibia to other wold regions?

7. Finally, I miss a connection with applied research. For user of data sources like Gridded Population of the World, what does all mean? What are the implications? Alternatives? Etc. I think the authors should make discuss all this, leaving technicalities aside, in the conclusions

Reviewer #2: Review of the manuscript “How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?:A simulation analysis in urban Namibia”

As the authors point out, the paper presents a method of evaluating the cell-level accuracy of 32 simulated 100x100m WorldPop-Global-Unconstrained gridded population datasets which reflect realistic scenarios of census (1) outdatedness, (2) inaccuracy, and (3) aggregation in an urban LMIC setting. This topic is very interesting and timely, but the purpose of the article should be described more clearly.

A thorough overview of the literature is included in Introduction, and the quoted items exhaust the proposed topic. In line 123 authors state that they evaluate 32 simulated 100x100m WorldPop-Global-Unconstrained gridded population datasets. The authors should explain why they chose the 32 grid. What was the reason for choosing such a set of gridded population datasets?

The section Methods is well presented and illustrated with figures. Yet, it should be explained why the Root Mean Square Error (RMSE) was selected to evaluate the gridded population dataset. The literature provides ample evidence on the effectiveness and usefulness of the Mean Absolute Error (MAE).

The quality of figures and charts is quite unsatisfactory, and they need to be presented in adequate resolution.

In Figure 1, the black background interferes with map reading. What do white boundaries on the right of the map mean?

The Discussion section is presented in a clear way. Will the studies be continued?

Is the proposed method universal? Can it be used for other research areas?

The Conclusions section reinstates the main findings in an adequate way.

The article meets high scientific quality standards and fits the scope of PLOS ONE. It contributes to the existing knowledge, presenting the topic in an interesting and up-to-date way.

**********

6. PLOS authors have the option to publish the peer review history of their article (what does this mean?). If published, this will include your full peer review and any attached files.

If you choose “no”, your identity will remain anonymous but your review may still be made public.

Do you want your identity to be public for this peer review? For information about this choice, including consent withdrawal, please see our Privacy Policy.

Reviewer #1: No

Reviewer #2: No

[NOTE: If reviewer comments were submitted as an attachment file, they will be attached to this email and accessible via the submission site. Please log into your account, locate the manuscript record, and check for the action link "View Attachments". If this link does not appear, there are no attachment files.]

While revising your submission, please upload your figure files to the Preflight Analysis and Conversion Engine (PACE) digital diagnostic tool, https://pacev2.apexcovantage.com/. PACE helps ensure that figures meet PLOS requirements. To use PACE, you must first register as a user. Registration is free. Then, login and navigate to the UPLOAD tab, where you will find detailed instructions on how to use the tool. If you encounter any issues or have any questions when using PACE, please email PLOS at figures@plos.org. Please note that Supporting Information files do not need this step.

Decision Letter 1

Krishna Prasad Vadrevu

10 Nov 2021

PONE-D-21-16343R1How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?: A simulation analysis in urban NamibiaPLOS ONE

Dear Dr. Thomson,

Thank you for submitting your manuscript to PLOS ONE. After careful consideration, we feel that it has merit but does not fully meet PLOS ONE’s publication criteria as it currently stands. Therefore, we invite you to submit a revised version of the manuscript that addresses the points raised during the review process.

Please elaborate the discussion to include Application users in mind. Also, please see the suggestions on additional literature suggested by one of the reviewers - please refer and cite them as needed.

Please submit your revised manuscript by Dec 25 2021 11:59PM. If you will need more time than this to complete your revisions, please reply to this message or contact the journal office at plosone@plos.org. When you're ready to submit your revision, log on to https://www.editorialmanager.com/pone/ and select the 'Submissions Needing Revision' folder to locate your manuscript file.

Please include the following items when submitting your revised manuscript:

  • A rebuttal letter that responds to each point raised by the academic editor and reviewer(s). You should upload this letter as a separate file labeled 'Response to Reviewers'.

  • A marked-up copy of your manuscript that highlights changes made to the original version. You should upload this as a separate file labeled 'Revised Manuscript with Track Changes'.

  • An unmarked version of your revised paper without tracked changes. You should upload this as a separate file labeled 'Manuscript'.

If you would like to make changes to your financial disclosure, please include your updated statement in your cover letter. Guidelines for resubmitting your figure files are available below the reviewer comments at the end of this letter.

If applicable, we recommend that you deposit your laboratory protocols in protocols.io to enhance the reproducibility of your results. Protocols.io assigns your protocol its own identifier (DOI) so that it can be cited independently in the future. For instructions see: https://journals.plos.org/plosone/s/submission-guidelines#loc-laboratory-protocols. Additionally, PLOS ONE offers an option for publishing peer-reviewed Lab Protocol articles, which describe protocols hosted on protocols.io. Read more information on sharing protocols at https://plos.org/protocols?utm_medium=editorial-email&utm_source=authorletters&utm_campaign=protocols.

We look forward to receiving your revised manuscript.

Kind regards,

Krishna Prasad Vadrevu, Ph.D

Academic Editor

PLOS ONE

Journal Requirements:

Please review your reference list to ensure that it is complete and correct. If you have cited papers that have been retracted, please include the rationale for doing so in the manuscript text, or remove these references and replace them with relevant current references. Any changes to the reference list should be mentioned in the rebuttal letter that accompanies your revised manuscript. If you need to cite a retracted article, indicate the article’s retracted status in the References list and also include a citation and full reference for the retraction notice.

[Note: HTML markup is below. Please do not edit.]

Reviewers' comments:

Reviewer's Responses to Questions

Comments to the Author

1. If the authors have adequately addressed your comments raised in a previous round of review and you feel that this manuscript is now acceptable for publication, you may indicate that here to bypass the “Comments to the Author” section, enter your conflict of interest statement in the “Confidential to Editor” section, and submit your "Accept" recommendation.

Reviewer #1: (No Response)

Reviewer #2: All comments have been addressed

**********

2. Is the manuscript technically sound, and do the data support the conclusions?

The manuscript must describe a technically sound piece of scientific research with data that supports the conclusions. Experiments must have been conducted rigorously, with appropriate controls, replication, and sample sizes. The conclusions must be drawn appropriately based on the data presented.

Reviewer #1: Yes

Reviewer #2: Yes

**********

3. Has the statistical analysis been performed appropriately and rigorously?

Reviewer #1: Yes

Reviewer #2: Yes

**********

4. Have the authors made all data underlying the findings in their manuscript fully available?

The PLOS Data policy requires authors to make all data underlying the findings described in their manuscript fully available without restriction, with rare exception (please refer to the Data Availability Statement in the manuscript PDF file). The data should be provided as part of the manuscript or its supporting information, or deposited to a public repository. For example, in addition to summary statistics, the data points behind means, medians and variance measures should be available. If there are restrictions on publicly sharing data—e.g. participant privacy or use of data from a third party—those must be specified.

Reviewer #1: No

Reviewer #2: Yes

**********

5. Is the manuscript presented in an intelligible fashion and written in standard English?

PLOS ONE does not copyedit accepted manuscripts, so the language in submitted articles must be clear, correct, and unambiguous. Any typographical or grammatical errors should be corrected at revision, so please note any specific errors here.

Reviewer #1: Yes

Reviewer #2: Yes

**********

6. Review Comments to the Author

Please use the space provided to explain your answers to the questions above. You may also include additional comments for the author, including concerns about dual publication, research ethics, or publication ethics. (Please upload your review as an attachment if it exceeds 20,000 characters)

Reviewer #1: I acknowledge the work done in revising the manuscript. I think that the paper has improved significantly.

The Introduction is now much easier to read and it better motivates the paper.

I still find the paper very technical, but I understand that this is the contribution. My concern is that, given its focus and style, the reach of the paper will be limited (see my next comment). In this line, the new intro helps. The discussion had not changed much and could try to be broader in scope.

Related to the above, the literature continuous to be deficient. Most references are technical. For applied “users” of gridded data (rather than researchers “creating” or “adjusting” the data), one wants to relate to applied work using this data to study several outcomes that you mention in the intro (i.e., development, environmental outcomes, etc.). Aside some papers about vaccination and health outcomes, there are hardly any reference to this type of papers. Think that many of your potential readers will be authors in journal like the JouEcoGeo JouUrbEco, JourDevStud, etc. I recommended trying to relate to recent work in these journals. I see no reference.

Minor:

Try to shorten (or break) sentences were possible.

Reviewer #2: The authors adequately addressed the comments of reviewers. I believe that this manuscript is now acceptable for publication.

**********

7. PLOS authors have the option to publish the peer review history of their article (what does this mean?). If published, this will include your full peer review and any attached files.

If you choose “no”, your identity will remain anonymous but your review may still be made public.

Do you want your identity to be public for this peer review? For information about this choice, including consent withdrawal, please see our Privacy Policy.

Reviewer #1: No

Reviewer #2: No

[NOTE: If reviewer comments were submitted as an attachment file, they will be attached to this email and accessible via the submission site. Please log into your account, locate the manuscript record, and check for the action link "View Attachments". If this link does not appear, there are no attachment files.]

While revising your submission, please upload your figure files to the Preflight Analysis and Conversion Engine (PACE) digital diagnostic tool, https://pacev2.apexcovantage.com/. PACE helps ensure that figures meet PLOS requirements. To use PACE, you must first register as a user. Registration is free. Then, login and navigate to the UPLOAD tab, where you will find detailed instructions on how to use the tool. If you encounter any issues or have any questions when using PACE, please email PLOS at figures@plos.org. Please note that Supporting Information files do not need this step.

PLoS One. 2022 Jul 21;17(7):e0271504. doi: 10.1371/journal.pone.0271504.r004

Author response to Decision Letter 1


16 Mar 2022

31 January 2022

Dear Dr. Vadrevu,

Thank you for this opportunity to provide minor revisions to our manuscript, “How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?: A simulation analysis in urban Namibia”. We have responded to comments below in italics, and made corresponding revisions to the manuscript in track changes.

Reviewer #1

1. Authors have not made all data underlying the findings in their manuscript fully available.

It is unclear why the reviewer believes that the underlying findings are not fully available with our manuscript. All of the datasets that we used to simulate populations are publicly available and linked in the cited publication by Thomson et al. 2018 and in Table 3. The simulated outdated censuses are based on actual historical satellite imagery, which is publicly available and cited. Our parameters to define inaccurate censuses are based on a systematic literature search, which is described and cited. Finally, our simulated “true” population and all 32 versions of our simulated censuses are provided in Supplement 2. If we have missed any datasets, please let us know which ones and we will make them available or provide the corresponding links.

2. I still find the paper very technical, but I understand that this is the contribution. My concern is that, given its focus and style, the reach of the paper will be limited (see my next comment). In this line, the new intro helps. The discussion had not changed much and could try to be broader in scope.

Related to the above, the literature continuous to be deficient. Most references are technical. For applied “users” of gridded data (rather than researchers “creating” or “adjusting” the data), one wants to relate to applied work using this data to study several outcomes that you mention in the intro (i.e., development, environmental outcomes, etc.). Aside some papers about vaccination and health outcomes, there are hardly any reference to this type of papers. Think that many of your potential readers will be authors in journal like the JouEcoGeo JouUrbEco, JourDevStud, etc. I recommended trying to relate to recent work in these journals. I see no reference.

We appreciate the push to keep data users in mind because we, ultimately, hope this paper can impact how gridded population modellers measure and report accuracy, and thus improve the accuracy and usability of gridded datasets for users.

However, the stated focus of this paper is on a creative approach to measure fine-scale accuracy of a gridded population dataset (see last paragraph of introduction). Broadly, the implications are the same for all indicators and sectors, so we have added the following sentence to the first paragraph of the discussion, “In practical terms, this means that urban development indicators calculated with a WorldPop-Global-Unconstrained dataset at fine scale (e.g., neighbourhood) would likely be incorrect, and could lead to confusing results. For example, an underestimate of the number of people living in a neighbourhood could both make vaccination coverage rates as well as disease infection rates appear incorrectly high in that neighbourhood.”

However, we do not intend to broaden the scope of the discussion further to specific use cases of one or more gridded population datasets because the results do not support this.

Note that in our last revision, we added discussion of how our findings about the WorldPop-Global-Unconstrained model might translate to other gridded population models if assessed for accuracy in the same way. We also cited additional urban development journals, including Environ Urban, in the opening paragraph while listing potential uses cases for fine-scale gridded population estimates.

We hope these explanations and edits are acceptable to the reviewer.

3. Try to shorten (or break) sentences were possible.

We split longer sentences in several places throughout the paper (e.g. lines 94, 104, 147, 183).

Reviewer #2: “The authors adequately addressed the comments of reviewers. I believe that this manuscript is now acceptable for publication.”

We thank both reviewers for their time and constructive feedback which has helped to strengthen the paper. Please do not hesitate to contact us with any questions or concerns.

Most sincerely,

Dana R. Thomson (with Douglas R. Leasure, Tomas Bird, Nikos Tzavidis, and Andrew J. Tatem)

Attachment

Submitted filename: cell_accuracy_response2.docx

Decision Letter 2

Krishna Prasad Vadrevu

3 May 2022

PONE-D-21-16343R2How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?: A simulation analysis in urban NamibiaPLOS ONE

Dear Dr. Thomson,

Thank you for submitting your manuscript to PLOS ONE. After careful consideration, we feel that it has merit but does not fully meet PLOS ONE’s publication criteria as it currently stands. Therefore, we invite you to submit a revised version of the manuscript that addresses the points raised during the review process.

Please revise manuscript to reflect application potential of the topic with relevant references.

Please submit your revised manuscript by Jun 17 2022 11:59PM. If you will need more time than this to complete your revisions, please reply to this message or contact the journal office at plosone@plos.org. When you're ready to submit your revision, log on to https://www.editorialmanager.com/pone/ and select the 'Submissions Needing Revision' folder to locate your manuscript file.

Please include the following items when submitting your revised manuscript:

  • A rebuttal letter that responds to each point raised by the academic editor and reviewer(s). You should upload this letter as a separate file labeled 'Response to Reviewers'.

  • A marked-up copy of your manuscript that highlights changes made to the original version. You should upload this as a separate file labeled 'Revised Manuscript with Track Changes'.

  • An unmarked version of your revised paper without tracked changes. You should upload this as a separate file labeled 'Manuscript'.

If you would like to make changes to your financial disclosure, please include your updated statement in your cover letter. Guidelines for resubmitting your figure files are available below the reviewer comments at the end of this letter.

If applicable, we recommend that you deposit your laboratory protocols in protocols.io to enhance the reproducibility of your results. Protocols.io assigns your protocol its own identifier (DOI) so that it can be cited independently in the future. For instructions see: https://journals.plos.org/plosone/s/submission-guidelines#loc-laboratory-protocols. Additionally, PLOS ONE offers an option for publishing peer-reviewed Lab Protocol articles, which describe protocols hosted on protocols.io. Read more information on sharing protocols at https://plos.org/protocols?utm_medium=editorial-email&utm_source=authorletters&utm_campaign=protocols.

We look forward to receiving your revised manuscript.

Kind regards,

Krishna Prasad Vadrevu, Ph.D

Academic Editor

PLOS ONE

Journal Requirements:

Please review your reference list to ensure that it is complete and correct. If you have cited papers that have been retracted, please include the rationale for doing so in the manuscript text, or remove these references and replace them with relevant current references. Any changes to the reference list should be mentioned in the rebuttal letter that accompanies your revised manuscript. If you need to cite a retracted article, indicate the article’s retracted status in the References list and also include a citation and full reference for the retraction notice.

[Note: HTML markup is below. Please do not edit.]

Reviewers' comments:

[NOTE: If reviewer comments were submitted as an attachment file, they will be attached to this email and accessible via the submission site. Please log into your account, locate the manuscript record, and check for the action link "View Attachments". If this link does not appear, there are no attachment files.]

While revising your submission, please upload your figure files to the Preflight Analysis and Conversion Engine (PACE) digital diagnostic tool, https://pacev2.apexcovantage.com/. PACE helps ensure that figures meet PLOS requirements. To use PACE, you must first register as a user. Registration is free. Then, login and navigate to the UPLOAD tab, where you will find detailed instructions on how to use the tool. If you encounter any issues or have any questions when using PACE, please email PLOS at figures@plos.org. Please note that Supporting Information files do not need this step.

PLoS One. 2022 Jul 21;17(7):e0271504. doi: 10.1371/journal.pone.0271504.r006

Author response to Decision Letter 2


3 Jul 2022

2 July 2022

Dear Dr. Vadrevu,

Thank you for this opportunity to provide minor revisions to our manuscript, “How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?: A simulation analysis in urban Namibia”. Please find our responses below in italics and track changes in the manuscript.

Editor:

1. Please revise manuscript to reflect application potential of the topic with relevant references.

We have added the following paragraph to the discussion:

This analysis reinforces findings of other studies which find that currently available gridded population products tend to underestimate populations in urban areas [94–96], especially in higher-density poorer neighbourhoods [97]. For example, Tuholske and colleagues (2021) compared five gridded population products to estimate the proportion of population affected by natural disasters (SDG 11.5) in three regions where disasters had occurred, and found that 1x1 km population estimates varied widely among data products, and reflected anywhere from 20% to 80% of the total UN estimated population in each region. Furthermore, they found that WorldPop-Global-Unconstrained generally performed better than un-modelled products (e.g., GPW), but not as well as products that constrained estimates to settled cells (e.g., GHS-POP) [94]. In a separate comparison of nine gridded population estimates in Kenyan and Nigerian slum populations (SDG 11.1) where field counts were available for reference, the estimated population in each slum varied widely and WorldPop-Global-Unconstrained estimates reflected just 11% of the overall slum population while the best performing data product (HRSL) estimated just 34% of all slum dwellers [97]. A key take-away from gridded population comparison studies is that fine-scale accuracy across data products varies substantially depending on location, potentially leading to different conclusions and decisions (e.g., about the humanitarian need or health care burden) depending on the gridded population dataset used for analysis. Furthermore, these studies underscore the need to understand fine-scale accuracy across gridded population datasets and locations to inform improvements to the underlying modelling methods and inputs.

2. Please review your reference list to ensure that it is complete and correct.

We checked the references to ensure they are complete and correct, including URL links.

Review #1:

3. I still find the paper very technical, but I understand that this is the contribution. My concern is that, given its focus and style, the reach of the paper will be limited (see my next comment). In this line, the new intro helps. The discussion had not changed much and could try to be broader in scope.

Related to the above, the literature continuous to be deficient. Most references are technical. For applied “users” of gridded data (rather than researchers “creating” or “adjusting” the data), one wants to relate to applied work using this data to study several outcomes that you mention in the intro (i.e., development, environmental outcomes, etc.). Aside some papers about vaccination and health outcomes, there are hardly any reference to this type of papers. Think that many of your potential readers will be authors in journal like the JouEcoGeo JouUrbEco, JourDevStud, etc. I recommended trying to relate to recent work in these journals. I see no reference.

We did not find any studies that applied or evaluated gridded population datasets in the Journal of Economic Geography, Journal of Urban Ecology, and Journal of Development Studies. We still stand by our previous response to this comment – that we do not intent to discuss “specific use cases of one or more gridded population datasets because the results do not support this.” However, as detailed above, we did add a paragraph to the discussion about other gridded population accuracy assessments and comparison studies in the contexts of disaster response (SDG 11.5) and estimating slum populations (SDG 11.1).

4. Try to shorten (or break) sentences were possible.

We split a few additional sentences throughout the paper to improve readability (e.g., lines 52, 392) and made a few additional minor edits to improve readability.

Please do not hesitate to contact us with any questions or concerns.

Most sincerely,

Dana R. Thomson (with Douglas R. Leasure, Tomas Bird, Nikos Tzavidis, and Andrew J. Tatem)

Attachment

Submitted filename: cell_accuracy_response3.docx

Decision Letter 3

Krishna Prasad Vadrevu

5 Jul 2022

How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?: A simulation analysis in urban Namibia

PONE-D-21-16343R3

Dear Dr. Thomson,

We’re pleased to inform you that your manuscript has been judged scientifically suitable for publication and will be formally accepted for publication once it meets all outstanding technical requirements.

Within one week, you’ll receive an e-mail detailing the required amendments. When these have been addressed, you’ll receive a formal acceptance letter and your manuscript will be scheduled for publication.

An invoice for payment will follow shortly after the formal acceptance. To ensure an efficient process, please log into Editorial Manager at http://www.editorialmanager.com/pone/, click the 'Update My Information' link at the top of the page, and double check that your user information is up-to-date. If you have any billing related questions, please contact our Author Billing department directly at authorbilling@plos.org.

If your institution or institutions have a press office, please notify them about your upcoming paper to help maximize its impact. If they’ll be preparing press materials, please inform our press team as soon as possible -- no later than 48 hours after receiving the formal acceptance. Your manuscript will remain under strict press embargo until 2 pm Eastern Time on the date of publication. For more information, please contact onepress@plos.org.

Kind regards,

Krishna Prasad Vadrevu, Ph.D

Academic Editor

PLOS ONE

Additional Editor Comments (optional):

Reviewers' comments:

Acceptance letter

Krishna Prasad Vadrevu

13 Jul 2022

PONE-D-21-16343R3

How accurate are WorldPop-Global-Unconstrained gridded population data at the cell-level?: A simulation analysis in urban Namibia

Dear Dr. Thomson:

I'm pleased to inform you that your manuscript has been deemed suitable for publication in PLOS ONE. Congratulations! Your manuscript is now with our production department.

If your institution or institutions have a press office, please let them know about your upcoming paper now to help maximize its impact. If they'll be preparing press materials, please inform our press team within the next 48 hours. Your manuscript will remain under strict press embargo until 2 pm Eastern Time on the date of publication. For more information please contact onepress@plos.org.

If we can help with anything else, please email us at plosone@plos.org.

Thank you for submitting your work to PLOS ONE and supporting open access.

Kind regards,

PLOS ONE Editorial Office Staff

on behalf of

Dr Krishna Prasad Vadrevu

Academic Editor

PLOS ONE

Associated Data

    This section collects any data citations, data availability statements, or supplementary materials included in this article.

    Supplementary Materials

    S1 Table. Percent of population missing from LMIC censuses by source.

    (DOCX)

    S2 Table. Root Mean Square Error (RMSE) statistics for all scenarios.

    (DOCX)

    S1 File. Simulating a population in Khomas, Namibia.

    (PDF)

    S2 File. Simulated population in Khomas, Namibia.

    (CSV)

    Attachment

    Submitted filename: cell_accuracy_response1.docx

    Attachment

    Submitted filename: cell_accuracy_response2.docx

    Attachment

    Submitted filename: cell_accuracy_response3.docx

    Data Availability Statement

    All relevant data are within the paper and its Supporting Information files.


    Articles from PLoS ONE are provided here courtesy of PLOS

    RESOURCES