1Univ Angers, Institut Agro, INRAE, IRHS, SFR QUASAV, 49000 Angers, France
2Research Centre for Cereal and Industrial Crops (CREA-CI), Council for Agricultural Research and Agricultural Economics Analysis, Via di Corticella 133, 40128 Bologna, Italy
3EA2106 Biomolécules et Biotechnologies Végétales, UFR Sciences Pharmaceutiques, Université François Rabelais, 31 avenue Monge, 37200 Tours, France
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