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Resistance landscape of surface polar relevant BSLA variants towards DMSO. Residual activity, activity, variants, and the empty vector are denoted as R, A, V, and EV, respectively. The OS resistance was measured in the absence or presence of 60 % (v/v) DMSO cosolvents after 2 h incubation with crude culture supernatant. The beneficial, unchanged, decreased, and inactive variants are colored green, yellow, red, and purple, respectively.
