2022 TAXONOMIC UPDATE OF PHYLUM Negarnaviricota (Riboviria: Orthornavirae), INCLUDING THE LARGE ORDERS Bunyavirales AND Mononegavirales

Abstract

In March 2022, following the annual International Committee on Taxonomy of Viruses (ICTV) ratification vote on newly proposed taxa, the phylum Negarnaviricota was amended and emended. The phylum was expanded by two new families (bunyaviral Discoviridae and Tulasviridae), 41 new genera, and 98 new species. Three hundred forty-nine species were renamed and/or moved. The accidentally misspelled names of seven species were corrected. This article presents the updated taxonomy of Negarnaviricota as now accepted by the ICTV.

INTRODUCTION

The phylum Negarnaviricota was established in 2019 by the International Committee on Taxonomy of Viruses (ICTV) for negative-sense RNA viruses that can be connected evolutionarily through the possession of virally encoded RNA-directed RNA polymerases (RdRps). The phylum includes two subphyla, Haploviricotina and Polyploviricotina, for negative-sense RNA viruses that encode RdRps with or without mRNA capping activity, respectively. The two subphyla include four classes (Chunqiuviricetes, Milneviricetes, Monjiviricetes, and Yunchangviricetes) and two classes (Ellioviricetes and Insthoviricetes), respectively [33, 59, 74]. The phylum was last amended/emended in 2021 [34]. Here we present the changes that were proposed to the phylum via official taxonomic proposals (TaxoProps) in 2021 and accepted by the ICTV in March and May 2022 [23, 70]. These changes are now part of the official ICTV taxonomy [24].

TAXONOMIC CHANGES IN SUBPHYLUM Haploviricotina

Chunqiuviricetes

The names of all species in the class were changed to binomials as required by the recently amended International Code of Virus Classification and Nomenclature (ICVCN) [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename).

Milneviricetes

The names of all species in the class were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.014P.R.Aspiviridae_rename).

Monjiviricetes

1. Family Aliusviridae

The species name Olluvirus shayangense was corrected to Ollusvirus shayangense (TaxoProp 2021.042M.R.Corrections_Riboviria_Duplodnaviria).

2. Family Artoviridae

The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.009M.R.Artoviridae_sprename).

Genus Peropuvirus was expanded by one new species, Peropuvirus melongenae, for Solanum melongena rhabdo-like virus (SmRLV) identified in a metagenomic study of eggplant (Solanum melongena L.) leaf tissue sampled in Zhènjiāng (镇江市), Jiāngsū Province (江苏省), China [78] (TaxoProp 2021.029M.R.Peropuvirus_1nsp).

3. Family Bornaviridae

The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (2021.010M.R.Bornaviridae_sprename).

Genus Orthobornavirus was expanded by one new species, Orthobornavirus caenophidiae, for Caribbean watersnake bornavirus (CWBV) and Mexican black-tailed rattlesnake bornavirus (MRBV) identified in archived raw transcriptomic read data of a Caribbean watersnake (colubrid Tretanorhinus variabilis A.M.C. Duméril, Bibron & A.H.A. Duméril, 1854) and a Mexican black-tailed rattlesnake (viperid Crotalus molossus nigrescens Gloyd, 1936), respectively [49] (TaxoProp 2021.021M.R.Orthobornavirus_1nsp).

4. Family Filoviridae

Family Filoviridae was expanded by two genera:

• genus Oblavirus was established to include one new species, Oblavirus percae, for Oberland virus (OBLV) discovered by high-throughput sequencing (HTS) in European perch (percid Perca fluviatilis Linnaeus, 1758) sampled in Switzerland [20] (TaxoProp 2021.011M.R.Filoviridae_1ngen_3nsp); and

• genus Tapjovirus was established to include one new species, Tapjovirus bothropis, for Tapajós virus (TAPV) discovered in sequence read archive (SRA) data obtained from a common lancehead (viperid Bothrops atrox (Linnaeus, 1758)) sampled in Brazil [21] (TaxoProp 2021.005M.R.Filoviridae_1ngen_1nsp).

Genus Thamnovirus was expanded by two new species, Thamnovirus percae for Fiwi virus (FIWIV) and Thamnovirus kanderense for Kander virus (KNDV), both discovered by HTS in European perch (percid Perca fluviatilis Linnaeus, 1758) sampled in Switzerland [20] (TaxoProp 2021.011M.R.Filoviridae_1ngen_3nsp).

5. Family Lispiviridae

Family Lispiviridae was thoroughly revised (TaxoProp 2021.016M.R.Lispiviridae_16ngen_13nsp):

• in genus Arlivirus, species Lishi arlivirus was renamed Arlivirus arachnae. Species Gerrid arlivirus, Hubei arlivirus, Odonate arlivirus, Tacheng arlivirus, and Wuchang arlivirus were renamed Sanstrivirus gerridis, Leocovirus coleopteris, Damravirus dentatis, Ganiavirus tachengense, and Nematovirus wuchangense, respectively, and moved into the newly created genera Sanstrivirus, Leocovirus, Damravirus, Ganiavirus, and Nematovirus, respectively;

• genus Anicalvirus was created to include the new species Anicalvirus hangzhouense for Anisopteromalus calandrae negative-strand RNA virus 2 (AcNSRV-2) discovered by HTS in a parasitoid wasp (pteromalid Anisopteromalus calandrae (Howard, 1881)) sampled in Hángzhōu (杭州市), Zhèjiāng Province (浙江省), China [72];

• genus Anidravirus was created to include the new species Anidravirus hangzhouense for Anisopteromalus calandrae negative-strand RNA virus 1 (AcNSRV-1) discovered by HTS in parasitoid wasps (pteromalid Anisopteromalus calandrae (Howard, 1881)) sampled in Hángzhōu (杭州市), Zhèjiāng Province (浙江省), China [72];

• genus Copasivirus was created to include two new species:

  • Copasivirus manlyvaleense for Jimsystermes virus (JIMV) discovered by HTS in termites (termitid Occasitermes sp.) sampled in Manly Vale, New South Wales, Australia [35]; and
  • Copasivirus ivindoense for isopteran arli-related virus OKIAV103 (IARV-103) discovered by HTS in termites (rhinotermitid Coptotermes sp.) sampled in Ivindo National Park (Parc national d'Ivindo), Ogooué-Ivindo Province, Gabon [27];

• genus Cybitervirus was created to include new species Cybitervirus niederense for coleopteran arli-related virus OKIAV107 (CARV-107) discovered by HTS in beetles (dytiscid Cybister lateralimarginalis (De Geer, 1774)) sampled in Lüchow-Dannenberg, Lower Saxony (Niedersachsen), Germany [27];

• genus Phelinovirus was created to include new species Phelinovirus aphidis for hymenopteran arli-related virus OKIAV99 (HARV-99) discovered by HTS in parasitoid wasps (aphelinid Aphelinus abdominalis (Dalman, 1820)) in laboratory culture samples of unknown geographical origin [27];

• genus Rivapovirus was created to include new species Rivapovirus aleyrodidae for hemipteran arli-related virus OKIAV94 (HARV-94) discovered by HTS in greenhouse whiteflies (aleyrodid Trialeurodes vaporariorum Westwood, 1856) of commercial laboratory cultures sampled in Brandenburg, Germany [27];

• genus Stylovirus was created to include new species Stylovirus niederense for strepsipteran arli-related virus OKIAV104 (SARV-104) discovered by HTS in endoparasitic insects (stylopid Stylops melittae Kirby, 1802) sampled in Osnabrück, Lower Saxony (Niedersachsen), Germany [27];

• genus Supelovirus was created to include the new species Supelovirus thailandense for blattodean arli-related virus OKIAV102 (BARV-102) discovered by HTS in brown-banded cockroaches (ectobiid Supella longipalpa Fabricius, 1798) of laboratory culture samples originating from Thailand [27];

• genus Synelinevirus was created to include two new species:

  • Synelinevirus paranaense for Linepithema humile rhabdo-like virus 1 (LhuRLV-1) 1 discovered by HTS in Argentine ants (dolichoderine Linepithema humile (Mayr, 1868)) sampled in Catalonia (Cataluña), Spain [68]; and
  • Synelinevirus bonnense for hymenopteran arli-related virus OKIAV98 (HARV-98) discovered by HTS in gall wasps (cynipid Synergus umbraculus Walker, 1874) sampled in Bonn, North Rhine-Westphalia (Nordrhein-Westfalen), Germany [27];

• genus Usmuvirus was created to include the new species Usmuvirus newyorkense for Amsterdam virus (AMSV) discovered by HTS in house mice (murid Mus musculus Linnaeus, 1758) sampled in New York, New York, USA [73]; and

• genus Xenophyvirus was created to include the new species Xenophyvirus mathesonense for hemipteran arli-related virus OKIAV95 (HARV-95) discovered by HTS in moss bugs (peloridiid Xenophyes metoponcus Burckhardt, 2011) sampled in Lake Matheson, South Island Westland District, New Zealand [27].

6. Family Mymonaviridae

Genus Auricularimonavirus was expanded by one new species, Auricularimonavirus bondarzewiae, for Bondarzewia berkeleyi negative-strand RNA virus 1 (BbNSRV-1) discovered by HTS in stump blossoms (bondarzewiaceaen Bondarzewia berkeleyi (Fr.) Bondartsev & Singer (1941)) [65] (TaxoProp 2021.041M.R.Mymonaviridae_5nsp).

Genus Botrytimonavirus was expanded by one new species, Botrytimonavirus alphabotrytidis, for Botrytis cinerea negative-stranded RNA virus 5 (BcNSRV-5) and Sclerotinia sclerotiorum negative-stranded RNA virus 10 (SsNSRV-10) discovered in metatranscriptomes of sclerotiniaceaen fungi (Botrytis cinerea Pers. (1794) and Sclerotinia sclerotiorum (Lib.) de Bary (1884) sampled in Italy/Spain and China, respectively) [26, 53] (TaxoProp 2021.041M.R.Mymonaviridae_5nsp). Sclerotinia sclerotiorum negative-stranded RNA virus 11 (SsNSRV-11) discovered in a Sclerotinia sclerotiorum (Lib.) de Bary (1884) metatranscriptome data set [26] was assigned to the already established species Botrytimonavirus botrytidis (TaxoProp 2021.041M.R.Mymonaviridae_5nsp).

Genus Sclerotimonavirus was expanded by three new species (TaxoProp 2021.041M.R.Mymonaviridae_5nsp):

• Sclerotimonavirus alphabotrytidis for Botrytis cinerea negative-stranded RNA virus 3 (BcNSRV-3) and Sclerotinia sclerotiorum negative-stranded RNA virus 9 (SsNSRV-9) discovered in metatranscriptomes of sclerotiniaceaen fungi (Botrytis cinerea Pers. (1794) and Sclerotinia sclerotiorum (Lib.) de Bary (1884) sampled in Italy/Spain and China, respectively) [26, 53];

• Sclerotimonavirus betabotrytidis for Botrytis cinerea negative-stranded RNA virus 4 (BcNSRV-4), discovered in the metatranscriptome of a sclerotiniaceaen fungus (Botrytis cinerea Pers. (1794)) sampled in Italy/Spain [53]; and

• Sclerotimonavirus cryphonectriae for Cryphonectria parasitica sclerotimonavirus 1 (CpSV-1), discovered by HTS in a cryphonectriaceaen fungus (Cryphonectria parasitica (Murrill) M. E. Barr (1978)) sampled in Azerbaijan [18].

7. Family Nyamiviridae

The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.020M.R.Nyamiviridae_sprename).

Genus Formivirus was expanded by one new species, Formivirus solenopsi, for Solenopsis invicta virus 15 (SoINV-15) identified in the transcriptome of red fire ants (myrmicine Solenopsis invicta Buren, 1972) sampled in the US and Taiwan [77] (TaxoProp 2021.019M.R.Nyamiviridae_3nsp).

Genus Nyavirus was expanded by two new species (TaxoProp 2021.019M.R.Nyamiviridae_3nsp):

• Nyavirus argatis, for Sekira virus (SEKRV) identified in the RNA virome of a pool of ticks (argasid Argas japonicus Yamaguti, Clifford and Tipton, 1968) sampled in Mishima (三島市), Shizuoka Prefecture (静岡県), Japan [28]; and

• Nyavirus somateriae for Jeremy Point nyavirus (JPNV) identified in the spleen of a common eider (anatid Somateria mollissima (Linnaeus, 1758)) collected at Jeremy Point, Massachusetts, USA, in 2013 (unpublished; GenBank: MN045233).

8. Family Paramyxoviridae

Genus Jeilongvirus was expanded by eight new species (TaxoProp: 2021.014M.R.Jeilongvirus_8nsp):

• Jeilongvirus anhuiense for Miniopterus schreibersii paramyxovirus (MisPV) discovered by HTS in Schreibers’s long-fingered bats (miniopterid Miniopterus schreibersii (Kuhl, 1817)) sampled in China [75];

• Jeilongvirus apodemi for rodent paramyxovirus (RoPV) discovered by HTS in a Korean field mouse (murid Apodemus peninsulae (Thomas, 1907)) sampled in China [76];

• Jeilongvirus comorosense for bat paramyxovirus 16797 (BatPV-1) identified in a Comoran long-fingered bat (miniopterid Miniopterus griveaudi Harrison, 1959) sampled in Comoros (unpublished; GenBank: MG203877);

• Jeilongvirus erinacei for belerina virus (BeV) discovered by HTS in West European hedgehogs (erinaceid Erinaceus europaeus Linnaeus, 1758) sampled in Belgium [66];

• Jeilongvirus felis for feline paramyxovirus (FPaV) identified in domestic cats (feline Felis catus Linnaeus, 1758) sampled in Germany [55, 61];

• Jeilongvirus madagascarense for bat paramyxovirus 17770 (BatPV-2) identified in a Comoran long-fingered bat (miniopterid Miniopterus griveaudi Harrison, 1959) sampled in Madagascar (unpublished; GenBank: MG203878);

• Jeilongvirus murinae for bat paramyxovirus (BatPV-3) identified in a greater tube-nosed bat (vespertilionid Murina leucogaster (Milne-Edwards, 1872) sampled in China [75]; and

• Jeilongvirus rungweense for ruloma virus (RulV) detected by HTS in a Machangu’s brush-furred rat (murid Lophuromys machangui Verheyen, Hulselmans, Dierckx, Mulungu, Leirs, Corti & Verheyen, 2007) sampled in Rungwe District, Mbeya Region, Tanzania [67].

9. Family Rhabdoviridae

The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProps 2021.036M.R.Rhabdoviridae_sprename, 2021.042M.R.Corrections_Riboviria_Duplodnaviria, and 2021.043M.R.Corrections_Mononegavirales_Bunyavirales).

Five new genera were created and included in the subfamily Alpharhabdovirinae:

• genus Amplylivirus, including one novel species, Amplylivirus cinereus, for frog lyssa-like virus 1 (FLLV-1) detected in an American green tree frog (hylid Dryophytes cinereus (Schneider, 1799)) (unpublished; GenBank: MK473367) (2021.035M.R.Rhabdoviridae_2ngen_2nsp);

• genus Cetarhavirus for two new species (2021.003M.R.Alpharhabdovirinae_3ngen_7nsp):

  • Cetarhavirus laganorhynchus for dolphin rhabdovirus (DRV) isolated from a white-beaked dolphin (delphinid Lagenorhynchus albirostris (Gray, 1846)) sampled in Schiermonnikoog, Friesland, Holland [45]; and
  • Cetarhavirus phocoena for harbour porpoise rhabdovirus (HPRV) isolated from a harbour porpoise (phocoenid Phocoena phocoena (Linnaeus, 1758)) stranded off the coast of Alaska [15];

• genus Replylivirus, including one novel species, Replylivirus allogus, for anole lyssa-like virus 1 (ALLV-1) identified in RNAseq data from a Spanish flag anole (dactyloid Anolis allogus Barbour & Ramsden, 1919) [22] (2021.035M.R.Rhabdoviridae_2ngen_2nsp);

• genus Scophrhavirus for two new species (2021.003M.R.Alpharhabdovirinae_3ngen_7nsp):

  • Scophrhavirus chanodychthys for Wuhan redfin culter dimarhabdovirus (WhRFCRV) detected by high-throughput sequencing in redfin culter (cyprinid Chanodichthys erythropterus (Basilewsky, 1855)) collected in Húběi Province (湖北省), China [58];
  • Scophrhavirus maximus for Scophthalmus maximus rhabdovirus (SMRV) isolated from turbot fish (scophthalmid Scophthalmus maximus (Linnaeus, 1758)) sampled in Shāndōng Province (山东省), China [82];
• genus Siniperhavirus for two new species (2021.003M.R.Alpharhabdovirinae_3ngen_7nsp):

  • Siniperhavirus chuatsi for Siniperca chuatsi rhabdovirus (SRCV) isolated from mandarin fish (sinipercid Siniperca chuatsi (Basilewsky, 1855)) collected in Guǎngdōng Province (广东省), China [86, 87]; Chinese rice-field eel rhabdovirus (CrERV) isolated from Chinese ricefield eels (synbranchid Monopterus albus (Zuiew, 1793)) sampled in Qiánjiāng (潜江市), Húběi Province (湖北省), China [38]; hybrid snakehead rhabdovirus (HSHRV) isolated from hybrid snakehead (channid Channa maculata × Channa argus) sampled in Fóshān (佛山市), Guǎngdōng Province (广东省), China [80]; and Micropterus salmoides rhabdovirus (MSRV) isolated from largemouth bass (centrarchid Micropterus salmoides (Lacépède, 1802)) sampled in Yúháng (余杭区), Zhèjiāng Province (浙江省), China [40]; and
  • Siniperhavirus zoarces for harbour eelpout rhabdovirus (EPRV) detected by HTS in eelpout (zoarcid Zoarces viviparous (Linnaeus, 1758)) near Stockholm, Sweden [2];

Genus Alphanucleorhabdovirus was expanded by one new species, Alphanucleorhabdovirus joa, for joa yellow blotch associated virus (JYBaV) identified in joá (solanaceaen Solanum aculeatissimum L.) plants sampled in Manaus, State of Amazonas (Estado de Amazonas), Brazil [11] (TaxoProp 2021.001M.A.v1.Betarhabdovirinae_7nsp).

Genus Betanucleorhabdovirus was expanded by one new species, Betanucleorhabdovirus bacopae, for Bacopa monnieri virus 2 (BmV-2) identified in transcriptomic data of water hyssop (plantaginaceaen Bacopa monnieri L.) tissues sampled in Lucknow, Uttar Pradesh, India [60] (TaxoProp 2021.001M.A.v1.Betarhabdovirinae_7nsp).

Genus Cytorhabdovirus was expanded by five new species (TaxoProp 2021.001M.A.v1.Betarhabdovirinae_7nsp):

• Cytorhabdovirus bacopae for Bacopa monnieri virus 1 (BmV-1) identified in transcriptomic data of water hyssop (plantaginaceaen Bacopa monnieri L.) tissues sampled in Lucknow, Uttar Pradesh, India [60];

• Cytorhabdovirus chrysanthemi for chrysanthemum yellow dwarf associated virus (CYDaV) identified in grey-leaved euryops (asteraceaen Euryops pectinatus (L.) Cass.) plant sampled in Chóngqìng (重庆市), China [37];

• Cytorhabdovirus cucurbitae for cucurbit cytorhabdovirus 1 (CuCV-1) identified in zucchini (cucurbitaceaen Cucurbita pepo L.) plants sampled in Greece [44];

• Cytorhabdovirus broussonetiae for paper mulberry mosaic associated virus (PMuMaV) identified in paper mulberry (moraceaen Broussonetia papyrifera L.) plants sampled in Chóngqìng (重庆市), China [51]; and

• Cytorhabdovirus rosae for rose virus R (RVR), identified in a rose ‘Hugh Dickson’ plant from Maryland, USA [5].

Genus Hapavirus was expanded by two new species (TaxoProp 2021.004M.R.Alpharhabdovirinae_13nsp):

• Hapavirus bangoran for Bangoran virus (BGNV) found in mosquitoes (culicid Culex perfuscus Edwards, 1914) sampled in Bangoran, Bamingui-Bangoran Prefecture, Central African Republic [6, 14]; and

• Hapavirus porton for Porton virus (PORV) found in mosquitoes (culicid Mansonia uniformis (Theobald, 1901)) sampled in Sarawak state, Malaysia [6].

Genus Ledantevirus was expanded by one new species, Ledantevirus taiyi, for Taiyi bat virus (TYBV) found in Chinese rufous horseshoe bats (rhinolophid Rhinolophus sinicus K. Andersen, 1905) sampled in Xiánníng (咸宁市), Húběi Province (湖北省), China [39] (TaxoProp 2021.004M.R.Alpharhabdovirinae_13nsp).

Genus Perhabdovirus was expanded by one new species, Perhabdovirus leman, for Leman virus (LeRV) isolated from perch fry sampled from Lake Geneva (Lac Léman), France [47] (TaxoProp 2021.003M.R.Alpharhabdovirinae_3ngen_7nsp).

Genus Sunrhavirus was expanded by eight new species (TaxoProp 2021.004M.R.Alpharhabdovirinae_13nsp):

• Sunrhavirus sandjimba for Sandjimba virus (SJAV) found in sedge warblers (acrocephalid Acrocephalus schoenobaenus (Linnaeus, 1758)) collected at the Landjia river, Ombella-M'Poko Prefecture, Central African Republic [6, 14];

• Sunrhavirus nasoule for Nasoule virus (NASV) found in little greenbuls (pycnonotid Eurillas virens (Cassin, 1857)) sampled in Nasoule, Mambéré-Kadéi Prefecture, Central African Republic [6, 14];

• Sunrhavirus boteke for Boteke virus (BOTV) found in mosquitoes (culicid Coquillettidia maculipennis (Theobald, 1911)) sampled in Boteke, Ombella-M'Poko Prefecture, Central African Republic [6, 14];

• Sunrhavirus bimbo for Bimbo virus (BBOV) found in yellow-crowned bishop (ploceid Euplectes afer (Gmelin, 1789)) sampled in Kolongo, Ombella-M'Poko Prefecture, Central African Republic [6, 14];

• Sunrhavirus kolongo for Kolongo virus (KOLV) found in yellow-crowned bishop (ploceid Euplectes afer (Gmelin, 1789)) sampled in Bangui, Central African Republic [6, 14];

• Sunrhavirus ouango for Ouango virus (OUAV) found in black-headed weaves (ploceid Ploceus melanocephalus (Linnaeus, 1758)) collected at the Landjia river, Ombella-M'Poko Prefecture, Central African Republic [6, 14];

• Sunrhavirus alexandria for Burg el Arab virus (BEAV) found in lesser whitethroats (sylviid Curruca curruca (Linnaeus, 1758)) sampled in Borg El Arab/Burj al-'Arab (قسم برج العرب), Alexandria Governorate (محافظة الإسكندرية), Egypt [6]; and

• Sunrhavirus matariya for Matariya virus (MTYV) found in lesser whitethroats (sylviid Curruca curruca (Linnaeus, 1758)) sampled in Port Saïd/Būrsaʿīd (بورسعيد), Port Saïd Governorate (محافظة بورسعيد), Egypt [6].

Genus Vesiculovirus was expanded by two new species (TaxoProp 2021.004M.R.Alpharhabdovirinae_13nsp):

• Vesiculovirus mediterranean for Mediterranean bat virus (MBV) found in bats sampled in Algeria, Morocco, and Spain (unpublished; GenBank: MW557331); and

• Vesiculovirus yinshui for Yinshui bat virus (YSBV) found in Chinese rufous horseshoe bats (rhinolophid Rhinolophus sinicus K. Andersen, 1905) sampled in Xiánníng (咸宁市), Húběi Province (湖北省), China [39].

10. Family Sunviridae

The name of the only species in the family was changed to a binomial as required by the recently amended ICVCN [25, 69, 81] (2021.037M.R.Sunviridae_sprename).

11. Family Xinmoviridae

Family Xinmoviridae was thoroughly revised (2021.039M.R.Xinmoviridae_11ngen_8nsp):

• in the genus Anphevirus, the species Shuangao anphevirus was abolished, and the species Xincheng anphevirus was renamed Anphevirus xinchengense. Species Dipteran anphevirus, Odonate anphevirus, Drosophilid anphevirus, Bolahun anphevirus, and Orthopteran anphevirus were renamed Alasvirus muscae, Draselvirus dentati, Drunivirus chambonense, Gambievirus bolahunense, and Hoptevirus orthopteris, respectively, and moved into the newly created genera Alasvirus, Draselvirus, Drunivirus, Gambievirus, and Hoptevirus, respectively;

• genus Doupovirus was created to include the new species Doupovirus australiaense for Culex mononega-like virus 2 (CMLV-2) discovered by HTS in mosquitoes (culicid Culex australicus Dobrotworsky & Drummond, 1953) sampled in Point Douro, Western Australia, Australia [57];

• genus Gambievirus was created to include the new species Gambievirus senegalense for Gambie virus (GAMV) discovered by HTS in mosquitoes (culicid Anopheles gambiae Giles, 1902) sampled in Senegal [17];

• genus Gylbovirus was created to include new species Gylbovirus aagae for Aedes anphevirus (AeAV) discovered by HTS in yellow fever mosquitoes (culicid Aedes aegypti (Linnaeus in Hasselquist, 1762)) and the Aag2 Aedes albopictus cell line [10, 48];

• genus Madalivirus was created to include two new species:

  • Madalivirus amapaense for Anopheles marajoara virus (AnMV) discovered by HTS in mosquitoes (culicid Anopheles marajoara Galvao & Damesceno 1942) sampled in Macapá, Amapá State, Brazil [56]; and
  • Madalivirus amazonaense for Anopheles darlingi virus (AnDV) discovered by HTS in American malaria mosquitoes (culicid Anopheles darlingi Root, 1926) sampled in Macapá, Amapá State, and Manaus, Amazonas State, Brazil [56];

• genus Pelmivirus was created to include new species Pelmivirus eymattense for hymenopteran anphe-related virus OKIAV71 (HARV-71) discovered by HTS in parasitoid wasps (ichneumonid Heteropelma amictum (Fabricius, 1775)) sampled in Eymatt, Bern, Switzerland [27];

• genus Triniovirus was created to include the new species Triniovirus yonagoense for Culex tritaeniorhynchus anphevirus (CtAV) discovered by HTS in mosquitoes (culicid Culex tritaeniorhynchus Giles, 1901) sampled in Yonago (米子市), Tottori Prefecture (鳥取県), Japan [16]; and

• genus Ulegvirus was created for new species Ulegvirus freckenfeldense for odonatan anphe-related virus OKIAV59 (OARV-59) discovered by HTS in golden-ringed dragonflies (cordulegastrid Cordulegaster boltonii (Donovan, 1807)) sampled in Freckenfeld, Rhineland-Palatinate (Rheinland-Pfalz), Germany [27].

Yunchangviricetes

The names of all species in the class were changed to binomials as required by the recently amended International Code of Virus Classification and Nomenclature (ICVCN) [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename).

TAXONOMIC CHANGES IN SUBPHYLUM Polyploviricotina

Taxonomic changes within order Articulavirales (Polyploviricotina: Insthoviricetes)

Family Amnoonviridae

The name of the only species in the family was changed to a binomial as required by the recently amended International Code of Virus Classification and Nomenclature (ICVCN) [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename).

Family Orthomyxoviridae

The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.024M.R.Orthomyxoviridae_sprename).

One new genus, Mykissvirus, was established to include one new species, Mykissvirus tructae, for rainbow trout orthomyxovirus (RbtOV) and steelhead trout orthomyxovirus (SttOV-1) first isolated from commercially reared rainbow trout (salmonid Oncorhynchus mykiss (Walbaum, 1792)) in Hagerman Valley, Idaho, USA [4] (TaxoProp 2021.022M.R.Orthomyxoviridae_1ngen_1nsp_Mykiss).

One new genus, Sardinovirus, was established to include one new species, Sardinovirus pilchardi, for pilchard orthomyxovirus (POMV) first isolated from blue pilchards (clupeid Sardinops sagax (Jenyns, 1842)) sampled off the coast of South Australia and subsequently Tasmania [19, 41] (TaxoProp 2021.023M.R.Orthomyxoviridae_1ngen_1nsp_Sardino).

Genus Quaranjavirus was expanded by four new species (TaxoProp 2021.034M.R.Quaranjavirus_4nsp):

• Quaranjavirus araguariense for Araguari virus (ARAV) isolated from a gray four-eyes opossum (didelphid Philander opossum (Linnaeus, 1758)) sampled in Serra do Navio, Amapá State, Brazil [9, 79];

• Quaranjavirus chadense for Lake Chad virus (LKCV) isolated from a vitelline masked weaver (ploceid Ploceus vitellinus (Lichtenstein, 1823)), sampled at Lake Chad, Nigeria [50];

• Quaranjavirus tyulekense for Tyulek virus (TLKV) isolated from ticks (argasid Argas vulgaris Filippova, 1961) sampled near Tûlëk/Tölök (Тюлёк/Төлөк), Naryn Region (Нарынская область), Kirghiz Soviet Socialist Republic, USSR [85]; and

• Quaranjavirus wellfleetense for Wellfleet Bay virus (WFBV) isolated from common eiders (anatid Somateria mollissima (Linnaeus, 1758)) sampled at Cape Cod, Massachusetts, USA [1].

Genus Thogotovirus was expanded by six new species (TaxoProp 2021.038M.R.Thogotovirus_6nsp):

• Thogotovirus bourbonense for Bourbon virus (BRBV) isolated from a human sampled in Bourbon County, Kansas, USA [30];

• Thogotovirus josense for Jos virus (JOSV) isolated from a zebu (bovid Bos indicus Linnaeus, 1758) and ticks (ixodid Amblyomma variegatum (Fabricius, 1794) and Rhipicephalus decoloratus Koch, 1844)) sampled in Jos, Plateau State, Nigeria [36];

• Thogotovirus ozense for Oz virus (OZV) isolated from ticks (ixodid Amblyomma testudinarium Koch, 1844) sampled in Ehime Prefecture (愛媛県), Japan [13];

• Thogotovirus sinuense for Sinu virus (SINUV) isolated from a pool of mosquitoes collected in San Bernardo del Viento, Córdoba Department, Colombia [8];

• Thogotovirus thailandense for Thailand tick thogotovirus (TT-THOV) detected by HTS in ticks (ixodid Boophilus sp.) sampled in Tha Wang Pha District (ท่าวังผา), Nan Province (น่าน), Thailand [63]; and

• Thogotovirus upoluense for Upolu virus (UPOV) isolated from ticks (ixodid Ornithodoros capensis Neumann, 1901) sampled on Upolu Cay, Great Barrier Reef, Australia [12].

Taxonomic changes within order Bunyavirales (Polyploviricotina: Ellioviricetes)

The order Bunyavirales was expanded by two new families (2021.040M.R.Bunyavirales_2nfam_2ngen_6nsp):

• Discoviridae: genus Orthodiscovirus was created for five species:

  • Orthodiscovirus coniellae for Coniothyrium diplodiella negative-stranded RNA virus 1 (CdNSRV-1) and Plasmopara viticola lesion associated mycobunyavirales-like virus 9, discovered by HTS in an isolate of a fungus (melanconidaceaen Coniella diplodiella (Speg.) Petr. & Syd., (1927)) and in plant lesions caused by grapevine downy mildew (peronosporaceaen Plasmopara viticola (Berk. & M.A. Curtis) Berl. & De Toni, (1888)), respectively [42];
  • Orthodiscovirus hispaniae for Plasmopara viticola lesion associated mycobunyavirales-like virus 8 (PvLAM-LV8) discovered by HTS in plant lesions caused by grapevine downy mildew (peronosporaceaen Plasmopara viticola (Berk. & M.A. Curtis) Berl. & De Toni, (1888)) [7];
  • Orthodiscovirus iberiae for Plasmopara viticola lesion associated mycobunyavirales-like virus 4 (PvLAM-LV4) discovered by HTS in plant lesions caused by grapevine downy mildew (peronosporaceaen Plasmopara viticola (Berk. & M.A. Curtis) Berl. & De Toni, (1888)) [7];
  • Orthodiscovirus missouriense for Penicillium discovirus (PDV) discovered by HTS in a fungus (trichocomaceaen Penicillium atramentosum Thom, C. 1910) [31]; and
  • Orthodiscovirus penicillii for Penicillium roseopurpureum negative ssRNA virus 1 (PrNssV-1) discovered by HTS in a fungus (trichocomaceaen Penicillium roseopurpureum Dierckx, R.P. 1901) [42];

• Tulasviridae: genus Orthotulasvirus was created to include one species, Orthotulasvirus tulasnellae, for Tulasnella bunyavirales-like virus 1 (TB-LV) discovered by HTS in an orchid mycorrhizal virome and confirmed to be present in an isolate of an effused fungus (tulasnellaceaen Tulasnella sp.) by full length Sanger sequencing of its cDNA [62].

Family Arenaviridae

Genus Mammarenavirus was expanded by two new species:

• Mammarenavirus bituense for Bitu virus (BITV) detected by HTS in Namaqua micaelamys (Micaelamys namaquensis (A. Smith, 1834)) sampled in Angola (TaxoProp 2021.006M.R.Mammarenavirus_1nsp_Bitu) [64]; and

• Mammarenavirus kwanzaense, for Kwanza virus (KWAV) detected by HTS in gray-bellied mice (Mus triton (Thomas, 1909)) sampled in Angola (TaxoProp 2021.007M.R.Mammarenavirus_1nsp_Kwanza) [64].

Family Fimoviridae

The names of all species in the family were changed to binomials as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.013P.R.Fimoviridae_rename).

Genus Emaravirus was expanded by three new species:

• Emaravirus aceris for maple mottle-associated emaravirus (MaMaV) discovered by HTS in maple (sapindaceaen Acer sp.) sampled in Germany (TaxoProp 2021.010P.R.Emaravirus_1ns) [54];

• Emaravirus chrysanthemi for chrysanthemum mosaic-associated emaravirus (ChMaV) discovered by HTS in chrysanthemum (asteraceaen Chrysanthemum morifolium L.) sampled in Japan (TaxoProps 2021.011P.R.Emaravirus_1ns and 2021.043M.R.Corrections_Mononegavirales_Bunyavirales) [32]; and

• Emaravirus quercus for common oak ringspot-associated emaravirus (CORaV) discovered by HTS in oak (fagaceaen Quercus robur L.) sampled in Germany and Scandinavia (TaxoProp 2021.012P.R.Emaravirus_1ns) [52].

Family Hantaviridae

The species name Seewis orhtohantavirus was corrected to Seewis orthohantavirus (TaxoProp 2021.042M.R.Corrections_Riboviria_Duplodnaviria).

The species name Thottopalayam thottimvirus was corrected to Thottapalayam thottimvirus (TaxoProp 2021.042M.R.Corrections_Riboviria_Duplodnaviria).

Family Leishbuviridae

The name of the only species in the family was changed to a binomial as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename).

Family Mypoviridae

The name of the only species in the family was changed to a binomial as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename).

Family Peribunyaviridae

Family Peribunyaviridae was expanded by three new genera (TaxoProp 2021.027M.R.Peribunyaviridae_3ngen_3nsp):

• Khurdivirus, including one new species, Khurdivirus volgaense, for Khurdun virus (KHURV) isolated from a Eurasian coot (rallid Fulica atra Linnaeus, 1758) sampled in Astrakhan Region (Астраханская область), Russia [83, 84];

• Lakivirus, including one new species, Lakivirus lakamhaense, for Lakamha virus (LAKV) discovered by HTS in a mosquito (culicid Wyeomyia complosa (Dyar, 1928)) sampled in Palenque, Chiapas, Mexico [29]; and

• Lambavirus, including one new species, Lambavirus wisconsinense, for largemouth bass bunyavirus (LBBV) discovered by HTS in largemouth bass (centrarchid Micropterus salmoides (Lacépède, 1802)) sampled in the upper Mississippi River, Wisconsin, USA [71].

The species name Lumbo orthbunyavirus was corrected to Lumbo orthobunyavirus (TaxoProp 2021.042M.R.Corrections_Riboviria_Duplodnaviria).

Family Phasmaviridae

The family was expanded by one new genus, Cicadellivirus, to include one new species, Cicadellivirus scaphoidei, for Scaphoideus titanus bunya-like virus 1 (StHV) (TaxoProp 2021.030M.R.Phasmaviridae_1ngen_5nsp) [46].

Genus Orthophasmavirus was expanded by four new species (TaxoProps 2021.030M.R.Phasmaviridae_1ngen_5nsp and 2021.043M.R.Corrections_Mononegavirales_Bunyavirales):

• Orthophasmavirus coredoense for Coredo virus (CORV) discovered by HTS in mosquitoes (culicid Mansonia titillans (Walker, 1848)) sampled in Colombia (unpublished; GenBank: MN661021–3);

• Orthophasmavirus barstukasense for Barstukas virus (BARV) discovered by HTS in mosquitoes (culicid Aedes sp.) sampled in California, USA [3];

• Orthophasmavirus miglotasense for Miglotas virus (MIGV) discovered by HTS in mosquitoes (culicid Aedes sp. and Culex sp.) sampled in California, USA [3]; and

• Orthophasmavirus flenense for Flen virus (FLNV) discovered by HTS in mosquitoes (culicid Aedes cantans (Meigen, 1818)) sampled in Flen, Södermanland County (Södermanlands län), Sweden [43].

Family Wupedeviridae

The name of the only species in the family was changed to a binomial as required by the recently amended ICVCN [25, 69, 81] (TaxoProp 2021.018M.R.Negarnaviricota_sprename).

Table 1.

ICTV-accepted taxonomy of the order Muvirales (Negarnaviricota: Haploviricotina: Chunqiuviricetes) as of March 2022.

Genus	Species¶	Virus (Abbreviation)&
Family Qinviridae
Yingvirus	Yingvirus beihaiense	Běihǎi sesarmid crab virus 4 (BhSCV-4)
	Yingvirus charybdis	Wēnzhōu qínvirus-like virus 2 (WzQLV-2)
	Yingvirus hubeiense	Húběi qínvirus-like virus 1 (HbQLV-1)
	Yingvirus sanxiaense	Sānxiá qínvirus-like virus 1 (SxQLV-1)
	Yingvirus shaheense	Shāhé qínvirus-like virus 1 (ShQLV-1)
	Yingvirus wenzhouense	Wēnzhōu qínvirus-like virus 1 (WzQLV-1)
	Yingvirus wuhanense	Wǔhàn insect virus 15 (WhIV-15)
	Yingvirus xinzhouense	Xīnzhōu nematode virus 3 (XzNV-3)

Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, families, and orders are taxa.

^¶Taxon names are always italicized and always begin with a capital letter.

^&Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization.

Table 2.

ICTV-accepted taxonomy of the order Serpentovirales (Negarnaviricota: Haploviricotina: Milneviricetes) as of March 2022.

Genus	Species¶	Virus (Abbreviation)&
Family Aspiviridae
Ophiovirus	Ophiovirus citri	citrus psorosis virus (CPsV)
	Ophiovirus freesiae	freesia sneak virus (FreSV)
	Ophiovirus lactucae	lettuce ring necrosis virus (LRNV)
	Ophiovirus mirafioriense	Mirafiori lettuce big-vein virus (MLBVV)
	Ophiovirus ranunculi	ranunculus white mottle virus (RWMV)
	Ophiovirus tulipae	tulip mild mottle mosaic virus (TMMMV)
	Ophiovirus vaccinii	blueberry mosaic associated virus (BlMaV)

Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, families, and orders are taxa.

^¶Taxon names are always italicized and always begin with a capital letter.

^&Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization.

Table 3.

ICTV-accepted taxonomy of the order Jingchuvirales (Negarnaviricota: Haploviricotina: Monjiviricetes) as of March 2022.

Genus	Species¶	Virus (Abbreviation)&
Family Aliusviridae
Obscuruvirus	Obscuruvirus quintum	Atrato chu-like virus 5 (AClV-5)
Ollusvirus	Ollusvirus coleopteri	Húběi coleoptera virus 3 (HbCV-3)
	Ollusvirus culvertonense	Culverton virus (CvV)
	Ollusvirus hanchengense	Hánchéng leafhopper mivirus (HLMV)
	Ollusvirus hymenopteri	hymenopteran chu-related virus 123 (HCrV-123)
	Ollusvirus insectii	hymenopteran chu-related virus 126 (HCrV-126)
	Ollusvirus scaldisense	Scaldis River bee virus (SRBV)
	Ollusvirus shayangense	Shāyáng fly virus 1 (SyFV-1)
	Ollusvirus taiyuanense	Tàiyuán leafhopper virus (TYLeV)
Family Chuviridae
Boscovirus	Boscovirus hippoboscidae	Wǔhàn louse fly virus 7 (WhLFV-7)
	Boscovirus hypoboscidae	Wǔhàn louse fly virus 6 (WhLFV-6)
Chuvivirus	Chuvivirus brunnichi	Wēnlǐng crustacean virus 14 (WlCV-14)
	Chuvivirus canceris	Wēnzhōu crab virus 2 (WzCV-2)
Culicidavirus	Culicidavirus culicidae	Wǔhàn mosquito virus 8 (WhMV-8)
	Culicidavirus culicis	Culex mosquito virus 5 (ClMV-5)
	Culicidavirus imjinense	Imjin River virus 1 (IjRV-1)
	Culicidavirus quitotaense	Culex mosquito virus 4 (ClMV-4)
Demapteravirus	Demapteravirus dermapteri	dermapteran chu-related virus 142 (DCrV-142)
Doliuvirus	Doliuvirus culisetae	Mos8Chu0 chuvirus (MoCV)
Mivirus	Mivirus amblyommae	lone star tick chuvirus 1 (LSTCV-1)
	Mivirus boleense	Bólè tick virus 3 (BTV-3)
	Mivirus changpingense	Chāngpíng tick virus 2 (CpTV-2)
	Mivirus dermacentoris	Chāngpíng tick virus 3 (CpTV-3)
		Tǎchéng tick virus 5 (TcTV-5)
	Mivirus genovaense	Genoa virus (GeV)
	Mivirus karukeraense	Karukera tick virus (KtV)
	Mivirus rhipicephali	brown dog tick mivirus 1 (BDTMV-1)
	Mivirus suffolkense	Suffolk virus (SFKV)
	Mivirus wuhanense	Wǔhàn tick virus 2 (WhTV-2)
Morsusvirus	Morsusvirus argatis	Tǎchéng tick virus 4 (TcTV-4)
Nigecruvirus	Nigecruvirus ixodes	blacklegged tick chuvirus-2 (BlTC-2)
Odonatavirus	Odonatavirus draconis	odonatan chu-related virus 137 (OCrV-137)
	Odonatavirus fabricii	Húběi odonate virus 11 (HbOV-11)
	Odonatavirus odontis	odonatan chu-related virus 136 (OCrV-136)
Pediavirus	Pediavirus cirripedis	Běihǎi barnacle virus 9 (BhBV-9)
Piscichuvirus	Piscichuvirus franki	Herr Frank virus 1 (HFrV-1)
	Piscichuvirus lycodontis	Guǎngdōng red-banded snake chuvirus-like virus (GRSCV)
	Piscichuvirus sanxiaense	Sānxiá atyid shrimp virus 4 (SxASC-4)
	Piscichuvirus wenlingense	Wēnlǐng fish chu-like virus (WFClV)
Pterovirus	Pterovirus chulinense	hymenopteran chu-related virus OKIAV147 (HCrV-147)
Scarabeuvirus	Scarabeuvirus blattae	Wǔchāng cockroach virus 3 (WcLFV-3)
	Scarabeuvirus dentati	Húběi chuvirus-like virus 3 (HbCLV-3)
	Scarabeuvirus hubeiense	Húběi chuvirus-like virus 1 (HbCLV-1)
	Scarabeuvirus lampyris	Lampyris noctiluca chuvirus-like virus 1 (LNClV-1)
	Scarabeuvirus lishiense	Lĭshì spider virus 1 (LsSV-1)
Taceavirus	Taceavirus wenlingense	Wēnlǐng crustacean virus 15 (WlCV-15)
Family Crepuscuviridae
Aqualaruvirus	Aqualaruvirus sialis	megalopteran chu-related virus 119 (MCrV-119)
Family Myriaviridae
Myriavirus	Myriavirus myriapedis	Húběi myriapoda virus 8 (HbMV-8)
Family Natareviridae
Charybdivirus	Charybdivirus charybdis	Wēnzhōu crab virus 3 (WzCV-3)

Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, families, and orders are taxa.

^¶Taxon names are always italicized and always begin with a capital letter.

^&Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization. Lists of viruses within a given species are provisional at this point and will likely be amended in the near future.

Table 4.

ICTV-accepted taxonomy of the order Mononegavirales (Negarnaviricota: Haploviricotina: Monjiviricetes) as of March 2022.

Genus	Species¶	Virus (Abbreviation)&
Family Artoviridae
Hexartovirus	Hexartovirus cirripedis	Běihǎi barnacle virus 8 (BhBV-8)
	Hexartovirus lepeophtheiri	Lepeophtheirus salmonis negative-stranded RNA virus 1 (LsNSRV-1)
Peropuvirus	Peropuvirus beihaiense	Běihǎi rhabdo-like virus 1 (BhRLV-1)
	Peropuvirus dentati	Húběi rhabdo-like virus 8 (HbRLV-8)
	Peropuvirus hubeiense	Húběi rhabdo-like virus 6 (HbRLV-6)
	Peropuvirus juli	Húběi rhabdo-like virus 5 (HbRLV-5)
	Peropuvirus lignarii	Běihǎi rhabdo-like virus 2 (BhRLV-2)
	Peropuvirus melongenae	Solanum melongena rhabdo-like virus (SmRLV)
	Peropuvirus pteromali	Pteromalus puparum negative-strand RNA virus 1 (PpNSRV-1)
Family Bornaviridae
Carbovirus	Carbovirus queenslandense	jungle carpet python virus (JCPV)
	Carbovirus tapeti	southwest carpet python virus (SWCPV)
Cultervirus	Cultervirus hemicultri	Wǔhàn sharpbelly bornavirus (WhSBV)
Orthobornavirus	Orthobornavirus alphapsittaciforme	parrot bornavirus 1 (PaBV-1)
		parrot bornavirus 2 (PaBV-2)
		parrot bornavirus 3 (PaBV-3)
		parrot bornavirus 4 (PaBV-4)
		parrot bornavirus 7 (PaBV-7)
	Orthobornavirus avisaquaticae	aquatic bird bornavirus 1 (ABBV-1)
		aquatic bird bornavirus 2 (ABBV-2)
	Orthobornavirus betapsittaciforme	parrot bornavirus 5 (PaBV-5)
	Orthobornavirus bornaense	Borna disease virus 1 (BoDV-1)
		Borna disease virus 2 (BoDV-2)
	Orthobornavirus caenophidiae	Caribbean watersnake bornavirus (CWBV)
		Mexican black-tailed rattlesnake bornavirus (MRBV)
	Orthobornavirus elapsoideae	Loveridge’s garter snake virus 1 (LGSV-1)
	Orthobornavirus estrildidae	estrildid finch bornavirus 1 (EsBV-1)
	Orthobornavirus sciuri	variegated squirrel bornavirus 1 (VSBV-1)
	Orthobornavirus serini	canary bornavirus 1 (CnBV-1)
		canary bornavirus 2 (CnBV-2)
		canary bornavirus 3 (CnBV-3)
Family Filoviridae
Cuevavirus	Lloviu cuevavirus	Lloviu virus (LLOV)
Dianlovirus	Mengla dianlovirus	Měnglà virus (MLAV)
Ebolavirus	Bombali ebolavirus	Bombali virus (BOMV)
	Bundibugyo ebolavirus	Bundibugyo virus (BDBV)
	Reston ebolavirus	Reston virus (RESTV)
	Sudan ebolavirus	Sudan virus (SUDV)
	Tai Forest ebolavirus	Taï Forest virus (TAFV)
	Zaire ebolavirus	Ebola virus (EBOV)
Marburgvirus	Marburg marburgvirus	Marburg virus (MARV)
		Ravn virus (RAVV)
Oblavirus	Oblavirus percae	Oberland virus (OBLV)
Striavirus	Xilang striavirus	Xīlǎng virus (XILV)
Tapjovirus	Tapjovirus bothropis	Tapajós virus (TAPV)
Thamnovirus	Huangjiao thamnovirus	Huángjiāo virus (HUJV)
	Thamnovirus kanderense	Kander virus (KNDV)
	Thamnovirus percae	Fiwi virus (FIWIV)
Family Lispiviridae
Anicalvirus	Anicalvirus hangzhouense	Anisopteromalus calandrae negative-strand RNA virus 2 (AcNSRV-2)
Anidravirus	Anidravirus hangzhouense	Anisopteromalus calandrae negative-strand RNA virus 1 (AcNSRV-1)
Arlivirus	Arlivirus arachnae	Lĭshì spider virus 2 (LsSV-2)
Copasivirus	Copasivirus ivindoense	isopteran arli-related virus OKIAV103 (IARV-103)
	Copasivirus manlyvaleense	Jimsystermes virus (JIMV)
Cybitervirus	Cybitervirus niederense	coleopteran arli-related virus OKIAV107 (CARV-107)
Damravirus	Damravirus dentatis	Húběi odonate virus 10 (HbOV-10)
Ganiavirus	Ganiavirus tachengense	Tǎchéng tick virus 6 (TcTV-6)
Leocovirus	Leocovirus coleopteris	Húběi rhabdo-like virus 3 (HbRLV-3)
Nematovirus	Nematovirus wuchangense	Wǔchāng romanomermis nematode virus 2 (WcRNV-2)
Phelinovirus	Phelinovirus aphidis	hymenopteran arli-related virus OKIAV99 (HARV-99)
Rivapovirus	Rivapovirus aleyrodidae	hemipteran arli-related virus OKIAV94 (HARV-94)
Sanstrivirus	Sanstrivirus gerridis	Sānxiá water strider virus 4 (SxWSV-4)
Stylovirus	Stylovirus niederense	strepsipteran arli-related virus OKIAV104 (SARV-104)
Supelovirus	Supelovirus thailandense	blattodean arli-related virus OKIAV102 (BARV-102)
Synelinevirus	Synelinevirus bonnense	hymenopteran arli-related virus OKIAV98 (HARV-98)
	Synelinevirus paranaense	Linepithema humile rhabdo-like virus 1 (LhuRLV-1)
Usmuvirus	Usmuvirus newyorkense	Amsterdam virus (AMSV)
Xenophyvirus	Xenophyvirus mathesonense	hemipteran arli-related virus OKIAV95 (HARV-95)
Family Mymonaviridae
Auricularimonavirus	Auricularimonavirus auriculariae	Auricularia heimuer negative-stranded RNA virus 1 (AhNRSV-1)
	Auricularimonavirus bondarzewiae	Bondarzewia berkeleyi negative-strand RNA virus 1 (BbNSRV-1)
Botrytimonavirus	Botrytimonavirus alphabotrytidis	Botrytis cinerea negative-stranded RNA virus 5 (BcNSRV-5)
		Sclerotinia sclerotiorum negative-stranded RNA virus 10 (SsNSRV-10)
	Botrytimonavirus botrytidis	Botrytis cinerea negative-stranded RNA virus 7 (BcNSRV-7)
		Sclerotinia sclerotiorum negative-stranded RNA virus 11 (SsNSRV-11)
	Botrytimonavirus glycinis	soybean leaf-associated negative-stranded RNA virus 3 (SLaNSRV-3)
	Botrytimonavirus sclerotiniae	Sclerotinia sclerotiorum negative-stranded RNA virus 2 (SsNSRV-2)
		Sclerotinia sclerotiorum negative-stranded RNA virus 2-A (SsNSRV-2-A)
		Sclerotinia sclerotiorum negative-stranded RNA virus 4 (SsNSRV-4)
		Sclerotinia sclerotiorum negative-stranded RNA virus 4-A (SsNSRV-4-A)
Hubramonavirus	Hubramonavirus hubeiense	Húběi rhabdo-like virus 4 (HbRLV-4)
	Hubramonavirus terrae	H2BulkLitter1223 virus
Lentimonavirus	Lentimonavirus lentinulae	Lentinula edodes negative-strand RNA virus 1 (LeNSRV-1)
Penicillimonavirus	Penicillimonavirus alphapenicillii	Penicillium adametzioides negative-stranded RNA virus 1 (PaNsRV-1)
	Penicillimonavirus alphaplasmoparae	Plasmopara viticola lesion associated mononegaambi virus 1 (PvaMV-1)
	Penicillimonavirus betapenicillii	Penicillium glabrum negative-stranded RNA virus 1 (PgRlV-1)
	Penicillimonavirus betaplasmoparae	Plasmopara viticola lesion associated mononegaambi virus 2 (PvaMV-2)
		Plasmopara viticola lesion associated mononegaambi virus 4 (PvaMV-4)
	Penicillimonavirus deltaplasmoparae	Plasmopara viticola lesion associated mononegaambi virus 5 (PvaMV-5)
	Penicillimonavirus epsilonplasmoparae	Plasmopara viticola lesion associated mononegaambi virus 6 (PvaMV-6)
	Penicillimonavirus etaplasmoparae	Plasmopara viticola lesion associated mononegaambi virus 9 (PvaMV-9)
	Penicillimonavirus gammaplasmopara	Plasmopara viticola lesion associated mononegaambi virus 3 (PvaMV-3)
	Penicillimonavirus kilnbarnense	Kiln Barn virus (KBV)
	Penicillimonavirus zetaplasmoparae	Plasmopara viticola lesion associated mononegaambi virus 7 (PvaMV-7)
Plasmopamonavirus	Plasmopamonavirus plasmoparae	Plasmopara viticola lesion associated mononegaambi virus 8 (PvaMV-8)
Phyllomonavirus	Phyllomonavirus gysingense	Gysinge virus (GYSV)
	Phyllomonavirus phyllospherae	soybean leaf-associated negative-stranded RNA virus 4 (SLaNSRV-4)
Rhizomonavirus	Rhizomonavirus mali	apple virus B (APPv-B)
Sclerotimonavirus	Sclerotimonavirus alphabotrytidis	Botrytis cinerea negative-stranded RNA virus 3 (BcNSRV-3)
		Sclerotinia sclerotiorum negative-stranded RNA virus 9 (SsNSRV-9)
	Sclerotimonavirus alphaclarireediae	Sclerotinia homoeocarpa TSA contig 1 (ShTSA-1)
		Sclerotinia homoeocarpa TSA contig 2 (ShTSA-2)
	Sclerotimonavirus alphaplasmoparae	Plasmopara viticola lesion associated mymonavirus 1 (PvaMV-1)
	Sclerotimonavirus alternariae	Alternaria tenuissima negative-stranded RNA virus 1 (AtNsRV-1)
	Sclerotimonavirus betabotrytidis	Botrytis cinerea negative-stranded RNA virus 4 (BcNSRV-4)
	Sclerotimonavirus betaclarireediae	Sclerotinia homoeocarpa TSA contig 3 (ShTSA-3)
	Sclerotimonavirus betaplasmoparae	Plasmopara viticola lesion associated mononega virus 2 (PvaMV-2)
	Sclerotimonavirus botrytidis	Botrytis cinerea mymonavirus 1 (BcMyV-1)
		Sclerotinia sclerotiorum negative-stranded RNA virus 7 (SsNSRV-7)
	Sclerotimonavirus cryphonectriae	Cryphonectria parasitica sclerotimonavirus 1 (CpSV-1)
	Sclerotimonavirus fusarii	Fusarium graminearum negative-stranded RNA virus 1 (FgNSRV-1)
		soybean leaf-associated negative-stranded RNA virus 1 (SLaNSRV-1)
	Sclerotimonavirus illinoisense	soybean leaf-associated negative-stranded RNA virus 2 (SLaNSRV-2)
	Sclerotimonavirus penicillii	Penicillium cairnsense negative-stranded RNA virus 1 (PcNsRV-1)
	Sclerotimonavirus sclerotiniae	Sclerotinia sclerotiorum negative-stranded RNA virus 1 (SsNSRV-1)
		Sclerotinia sclerotiorum negative-stranded RNA virus 1-A (SsNSRV-1-A)
		Sclerotinia sclerotiorum negative-stranded RNA virus 3 (SsNSRV-3)
		Sclerotinia sclerotiorum negative-stranded RNA virus 3-A (SsNSRV-3-A)
	Sclerotimonavirus terrae	H4BulkLitter234 virus
Family Nyamiviridae
Berhavirus	Berhavirus beihaiense	Běihǎi rhabdo-like virus 4 (BhRLV-4)
	Berhavirus radialis	Běihǎi rhabdo-like virus 5 (BhRLV-5)
	Berhavirus sipunculi	Běihǎi rhabdo-like virus 3 (BhRLV-3)
Crustavirus	Crustavirus beihaiense	Běihǎi rhabdo-like virus 6 (BhRLV-6)
	Crustavirus wenlingense	Wēnlǐng crustacean virus 12 (WlCV-12)
	Crustavirus wenzhouense	Wēnzhōu crab virus 1 (WzCV-1)
Formivirus	Formivirus angliae	Formica fusca virus 1 (FfusV-1)
	Formivirus chalybii	hymenopteran orino-related virus OKIAV87 (HORV-87)
	Formivirus finnoniae	Formica exsecta virus 4 (FeV-4)
	Formivirus gorytis	hymenopteran orino-related virus OKIAV85 (HORV-85)
	Formivirus solenopsi	Solenopsis invicta virus 15 (SoINV-15)
Nyavirus	Nyavirus argatis	Sekira virus (SEKRV)
	Nyavirus midwayense	Midway virus (MIDWV)
	Nyavirus nyamaniniense	Nyamanini virus (NYMV)
	Nyavirus sanjacintoense	San Jacinto virus (SJCV)
	Nyavirus sierranevadaense	Sierra Nevada virus (SNVV)
	Nyavirus somateriae	Jeremy Point nyavirus (JPNV)
Orinovirus	Orinovirus pasiphilae	Orinoco virus (ONCV)
Socyvirus	Socyvirus heteroderae	soybean cyst nematode virus 1 (SbCNV-1)
Tapwovirus	Tapwovirus cesti	Wēnzhōu tapeworm virus 1 (WzTWV-1)
Family Paramyxoviridae
Subfamily Avulavirinae
Metaavulavirus	Avian metaavulavirus 2	avian paramyxovirus 2 (APMV-2)
	Avian metaavulavirus 5	avian paramyxovirus 5 (APMV-5)
	Avian metaavulavirus 6	avian paramyxovirus 6 (APMV-6)
	Avian metaavulavirus 7	avian paramyxovirus 7 (APMV-7)
	Avian metaavulavirus 8	avian paramyxovirus 8 (APMV-8)
	Avian metaavulavirus 10	avian paramyxovirus 10 (APMV-10)
	Avian metaavulavirus 11	avian paramyxovirus 11 (APMV-11)
	Avian metaavulavirus 14	avian paramyxovirus 14 (APMV-14)
	Avian metaavulavirus 15	avian paramyxovirus 15 (APMV-15)
	Avian metaavulavirus 20	avian paramyxovirus 20 (APMV-20)
	Avian metaavulavirus 22	avian paramyxovirus 22 (APMV-22)
Orthoavulavirus	Avian orthoavulavirus 1	avian paramyxovirus 1 (APMV-1)1
	Avian orthoavulavirus 9	avian paramyxovirus 9 (APMV-9)
	Avian orthoavulavirus 12	avian paramyxovirus 12 (APMV-12)
	Avian orthoavulavirus 13	avian paramyxovirus 13 (APMV-13)
	Avian orthoavulavirus 16	avian paramyxovirus 16 (APMV-16)
	Avian orthoavulavirus 17	Antarctic penguin virus A (APV-A)
	Avian orthoavulavirus 18	Antarctic penguin virus B (APV-B)
	Avian orthoavulavirus 19	Antarctic penguin virus C (APV-C)
	Avian orthoavulavirus 21	avian paramyxovirus 21 (APMV-21)
Paraavulavirus	Avian paraavulavirus 3	avian paramyxovirus 3 (APMV-3)
	Avian paraavulavirus 4	avian paramyxovirus 4 (APMV-4)
Subfamily Metaparamyxovirinae
Synodonvirus	Synodus synodonvirus	Wēnlǐng triplecross lizardfish paramyxovirus (WTLPV)
Subfamily Orthoparamyxovirinae
Aquaparamyxovirus	Oncorhynchus aquaparamyxovirus	Pacific salmon paramyxovirus (PSPV)
	Salmo aquaparamyxovirus	Atlantic salmon paramyxovirus (AsaPV)
Ferlavirus	Reptilian ferlavirus	fer-de-lance virus (FDLV)
Henipavirus	Cedar henipavirus	Cedar virus (CedV)
	Ghanaian bat henipavirus	Ghana virus (GhV)
	Hendra henipavirus	Hendra virus (HeV)
	Mojiang henipavirus	Mòjiāng virus (MojV)
	Nipah henipavirus	Nipah virus (NiV)
Jeilongvirus	Beilong jeilongvirus	Beilong virus (BeiV)
	Jeilongvirus anhuiense	Miniopterus schreibersii paramyxovirus (MisPV)
	Jeilongvirus apodemi	rodent paramyxovirus (RoPV)
	Jeilongvirus comorosense	bat paramyxovirus 16797 (BatPV-1)
	Jeilongvirus erinacei	belerina virus (BeV)
	Jeilongvirus felis	feline paramyxovirus (FPaV)
	Jeilongvirus madagascarense	bat paramyxovirus 17770 (BatPV-2)
	Jeilongvirus murinae	bat paramyxovirus (BatPV-3)
	Jeilongvirus rungweense	ruloma virus (RulV)
	Jun jeilongvirus	J virus (JV)
	Lophuromys jeilongvirus 1	Mount Mabu Lophuromys virus 1 (MMLV-1)
	Lophuromys jeilongvirus 2	Mount Mabu Lophuromys virus 2 (MMLV-2)
	Miniopteran jeilongvirus	Shaan virus (ShaV)
	Myodes jeilongvirus	Pohorje Myodes paramyxovirus 1 (PMPV-1)
	Tailam jeilongvirus	Tailam virus (TaiV)
Morbillivirus	Canine morbillivirus	canine distemper virus (CDV)
	Cetacean morbillivirus	cetacean morbillivirus (CeMV)
	Feline morbillivirus	feline morbillivirus (FeMV)
	Measles morbillivirus	measles virus (MeV)
	Phocine morbillivirus	phocine distemper virus (PDV)
	Rinderpest morbillivirus	rinderpest virus (RPV)
	Small ruminant morbillivirus	peste-des-petits-ruminants virus (PPRV)
Narmovirus	Mossman narmovirus	Mossman virus (MossV)
	Myodes narmovirus	bank vole virus 1 (BaV-1)
	Nariva narmovirus	Nariva virus (NarV)
	Tupaia narmovirus	Tupaia paramyxovirus (TupV)
Respirovirus	Bovine respirovirus 3	bovine parainfluenza virus 3 (BPIV-3)
	Caprine respirovirus 3	caprine parainfluenzavirus 3 (CPIV-3)
	Human respirovirus 1	human parainfluenza virus 1 (HPIV-1)
	Human respirovirus 3	human parainfluenza virus 3 (HPIV-3)
	Murine respirovirus	Sendai virus (SeV)
	Porcine respirovirus 1	porcine parainfluenza virus 1 (PPIV-1)
	Squirrel respirovirus	giant squirrel virus (GSqV)
Salemvirus	Salem salemvirus	Salem virus (SalV)
Subfamily Rubulavirinae
Orthorubulavirus	Human orthorubulavirus 2	human parainfluenza virus 2 (HPIV-2)
	Human orthorubulavirus 4	human parainfluenza virus 4a (HPIV-4a)
		human parainfluenza virus 4b (HPIV-4b)
	Mammalian orthorubulavirus 5	parainfluenza virus 5 (PIV-5)
	Mammalian orthorubulavirus 6	Alston virus (AlsV)
	Mapuera orthorubulavirus	Mapuera virus (MapV)
	Mumps orthorubulavirus	mumps virus (MuV)
	Porcine orthorubulavirus	La Piedad Michoacán Mexico virus (LPMV)
	Simian orthorubulavirus	simian virus 41 (SV-41)
Pararubulavirus	Achimota pararubulavirus 1	Achimota virus 1 (AchPV-1)
	Achimota pararubulavirus 2	Achimota virus 2 (AchPV-2)
	Hervey pararubulavirus	Hervey virus (HerV)
	Menangle pararubulavirus	Menangle virus (MenPV)
	Sosuga pararubulavirus	Sosuga virus (SOSV)
	Teviot pararubulavirus	Teviot virus (TevPV)
	Tioman pararubulavirus	Tioman virus (TioPV)
	Tuhoko pararubulavirus 1	Tuhoko virus 1 (ThkPV-1)
	Tuhoko pararubulavirus 2	Tuhoko virus 2 (ThkPV-2)
	Tuhoko pararubulavirus 3	Tuhoko virus 3 (ThkPV-3)
Unassigned (to subfamilies)
Cynoglossusvirus	Cynoglossus cynoglossusvirus	Wēnlǐng tonguesole paramyxovirus (WTSPV)
Hoplichthysvirus	Hoplichthys hoplichthysvirus	Wēnlǐng hoplichthys paramyxovirus (WHPV)
Scoliodonvirus	Scoliodon scoliodonvirus	Wēnzhōu pacific spadenose shark paramyxovirus (WPSSPV)
Family Pneumoviridae
Metapneumovirus	Avian metapneumovirus	avian metapneumovirus (AMPV)
	Human metapneumovirus	human metapneumovirus (HMPV)
Orthopneumovirus	Bovine orthopneumovirus	bovine respiratory syncytial virus (BRSV)
	Human orthopneumovirus	human respiratory syncytial virus (HRSV)
	Murine orthopneumovirus	murine pneumonia virus (MPV)
Family Rhabdoviridae
Subfamily Alpharhabdovirinae
Almendravirus	Almendravirus almendras	Puerto Almendras virus (PTAMV)
	Almendravirus arboretum	Arboretum virus (ABTV)
	Almendravirus balsa	Balsa virus (BALV)
	Almendravirus chico	Rio Chico virus (RCHV)
	Almendravirus cootbay	Coot Bay virus (CBV)
	Almendravirus menghai	Menghai virus (MRV)
Alphanemrhavirus	Alphanemrhavirus xingshan	Xingshan nematode virus 4 (XsNV-4)
	Alphanemrhavirus xinzhou	Xinzhou nematode virus 4 (XzNV-4)
Alphapaprhavirus	Alphapaprhavirus hubei	Hubei lepidoptera virus 2 (HbLV-2)
	Alphapaprhavirus pararge	Pararge aegeria rhabdovirus (PAeRV)
Alpharicinrhavirus	Alpharicinrhavirus blanchseco	Blanchseco virus (BCOV)
	Alpharicinrhavirus bole	Bole tick virus 2 (BlTV-2)
	Alpharicinrhavirus wuhan	Wuhan tick virus 1 (WhTV-1)
Amplylivirus	Amplylivirus cinereus	frog lyssa-like virus 1 (FLLV-1)
Arurhavirus	Arurhavirus aruac	Aruac virus (ARUV)
	Arurhavirus inhangapi	Inhangapi virus (INHV)
	Arurhavirus santabarbara	Santa Barbara virus (SBAV)
	Arurhavirus xiburema	Xiburema virus (XIBV)
Barhavirus	Barhavirus bahia	Bahia Grande virus (BGV)
		Harlingen virus (HARV)
	Barhavirus muir	Muir Springs virus (MSV)
Caligrhavirus	Caligrhavirus caligus	Caligus rogercresseyi rhabdovirus (CRogRV)
	Caligrhavirus lepeophtheirus	Lepeophtheirus salmonis rhabdovirus 127 (LSalRV-127)
	Caligrhavirus salmonlouse	Lepeophtheirus salmonis rhabdovirus 9 (LSalRV-9)
Cetarhavirus	Cetarhavirus laganorhynchus	dolphin rhabdovirus (DRV)
	Cetarhavirus phocoena	harbour porpoise rhabdovirus (HPRV)
Curiovirus	Curiovirus curionopolis	Curionopolis virus (CURV)
	Curiovirus iriri	Iriri virus (IRIRV)
	Curiovirus itacaiunas	Itacaiunas virus (ITAV)
	Curiovirus rochambeau	Rochambeau virus (RBUV)
Ephemerovirus	Ephemerovirus adelaide	Adelaide River virus (ARV)
	Ephemerovirus berrimah	Berrimah virus (BRMV)
	Ephemerovirus febris	bovine ephemeral fever virus (BEFV)
	Ephemerovirus hayes	Hayes Yard virus (HYV)
	Ephemerovirus kent	New Kent County virus (NKCV)
	Ephemerovirus kimberley	Kimberley virus (KIMV)
		Malakal virus (MALV)
	Ephemerovirus koolpinyah	Koolpinyah virus (KOOLV)
	Ephemerovirus kotonkan	kotonkan virus (KOTV)
	Ephemerovirus obodhiang	Obodhiang virus (OBOV)
	Ephemerovirus puchong	Puchong virus (PUCV)
	Ephemerovirus yata	Yata virus (YATV)
Hapavirus	Hapavirus bangoran	Bangoran virus (BGNV)
	Hapavirus flanders	Flanders virus (FLAV)
	Hapavirus graylodge	Gray Lodge virus (GLOV)
	Hapavirus hartpark	Hart Park virus (HPV)
	Hapavirus holmes	Holmes Jungle virus (HOJV)
	Hapavirus joinjakaka	Joinjakaka virus (JOIV)
	Hapavirus kamese	Kamese virus (KAMV)
	Hapavirus lajoya	La Joya virus (LJV)
	Hapavirus landjia	Landjia virus (LANV = LJAV)
	Hapavirus manitoba	Manitoba virus (MANV = MNTBV)
	Hapavirus marco	Marco virus (MCOV)
	Hapavirus mosqueiro	Mosqueiro virus (MQOV)
	Hapavirus mossuril	Mossuril virus (MOSV)
	Hapavirus ngaingan	Ngaingan virus (NGAV)
	Hapavirus ord	Ord River virus (ORV)
	Hapavirus parry	Parry Creek virus (PCV)
	Hapavirus porton	Porton virus (PORV)
	Hapavirus wongabel	Wongabel virus (WONV)
Ledantevirus	Ledantevirus barur	Barur virus (BARV)
	Ledantevirus bughendera	Bughendera virus (BUGV)
	Ledantevirus elgon	Mount Elgon bat virus (MEBV)
	Ledantevirus fikirini	Fikirini virus (FKRV)
	Ledantevirus fukuoka	Fukuoka virus (FUKV)
	Ledantevirus kanyawara	Kanyawara virus (KYAV)
	Ledantevirus kern	Kern Canyon virus (KCV)
	Ledantevirus keuraliba	Keuraliba virus (KEUV)
	Ledantevirus kolente	Kolente virus (KOLEV)
	Ledantevirus kumasi	Kumasi rhabdovirus (KRV)
	Ledantevirus ledantec	Le Dantec virus (LDV)
	Ledantevirus nishimuro	Nishimuro virus (NISV)
	Ledantevirus nkolbisson	Nkolbisson virus (NKOV)
	Ledantevirus oita	Oita virus (OITAV)
	Ledantevirus taiyi	Taiyi bat virus (TYBV)
	Ledantevirus vaprio	Vaprio virus (VAPV)
	Ledantevirus wuhan	Wuhan louse fly virus 5 (WLFV-5)
	Ledantevirus yongjia	Yongjia tick virus 2 (YTV-2)
Lostrhavirus	Lostrhavirus hyalomma	Xinjiang tick rhabdovirus (XjTRV)
	Lostrhavirus lonestar	lone star tick rhabdovirus (LITRV)
Lyssavirus	Lyssavirus aravan	Aravan virus (ARAV)
	Lyssavirus australis	Australian bat lyssavirus (ABLV)
	Lyssavirus bokeloh	Bokeloh bat lyssavirus (BBLV)
	Lyssavirus caucasicus	West Caucasian bat virus (WCBV)
	Lyssavirus duvenhage	Duvenhage virus (DUVV)
	Lyssavirus formosa	Taiwan bat lyssavirus (TWBLV)
	Lyssavirus gannoruwa	Gannoruwa bat lyssavirus (GBLV)
	Lyssavirus hamburg	European bat lyssavirus 1 (EBLV-1)
	Lyssavirus helsinki	European bat lyssavirus 2 (EBLV-2)
	Lyssavirus ikoma	Ikoma lyssavirus (IKOV)
	Lyssavirus irkut	Irkut virus (IRKV)
	Lyssavirus khujand	Khujand virus (KHUV)
	Lyssavirus lagos	Lagos bat virus (LBV)
	Lyssavirus lleida	Lleida bat lyssavirus (LLEBV)
	Lyssavirus mokola	Mokola virus (MOKV)
	Lyssavirus rabies	rabies virus (RABV)
	Lyssavirus shimoni	Shimoni bat virus (SHIBV)
Merhavirus	Merhavirus merida	Merida virus (MERDV)
	Merhavirus tritaeniorhynchus	Culex tritaeniorhynchus rhabdovirus (CTRV)
Mousrhavirus	Mousrhavirus moussa	Moussa virus (MOUV)
Ohlsrhavirus	Ohlsrhavirus angeles	Culex rhabdo-like virus Los Angeles (CRLVLA)
	Ohlsrhavirus culex	Culex rhabdo-like virus (CRLV)
	Ohlsrhavirus lobeira	Lobeira virus (LOBV)
	Ohlsrhavirus northcreek	North Creek virus (NORCV)
	Ohlsrhavirus ohlsdorf	Ohlsdorf virus (OHLDV)
	Ohlsrhavirus pseudovishnui	Culex pseudovishnui rhabdo-like virus (CpRLV)
	Ohlsrhavirus riverside	Riverside virus (RISV)
	Ohlsrhavirus tongilchon	Tongilchon virus 1 (TCHV-1)
Perhabdovirus	Perhabdovirus anguilla	eel virus American (EVA)
		eel virus European X (EVEX)
	Perhabdovirus leman	Leman virus (LeRV)
	Perhabdovirus perca	perch rhabdovirus (PRV)
	Perhabdovirus trutta	lake trout rhabdovirus (LTRV)
Replylivirus	Replylivirus allogus	anole lyssa-like virus 1 (ALLV-1)
Sawgrhavirus	Sawgrhavirus connecticut	Connecticut virus (CNTV)
	Sawgrhavirus longisland	Long Island tick rhabdovirus (LITRV)
	Sawgrhavirus minto	New Minto virus (NMV)
	Sawgrhavirus sawgrass	Sawgrass virus (SAWV)
Scophrhavirus	Scophrhavirus chanodychthys	Wuhan redfin culter dimarhabdovirus (WhRFCRV)
	Scophrhavirus maximus	Scophthalmus maximus rhabdovirus (SMRV)
Sigmavirus	Sigmavirus affinis	Drosophila affinis sigmavirus (DAffSV)
	Sigmavirus ananassae	Drosophila ananassae sigmavirus (DAnaSV)
	Sigmavirus capitata	Ceratitis capitata sigmavirus (CCapSV)
	Sigmavirus domestica	Wuhan fly virus 2 (WhFV-2)
	Sigmavirus hippoboscid	Wuhan louse fly virus 9 (WhLFV-9)
	Sigmavirus hubei	Hubei diptera virus 9 (HbDV-9)
	Sigmavirus immigrans	Drosophila immigrans sigmavirus (DImmSV)
	Sigmavirus lousefly	Wuhan louse fly virus 10 (WhLFV-10)
	Sigmavirus melanogaster	Drosophila melanogaster sigmavirus (DMelSV)
	Sigmavirus muscina	Muscina stabulans sigmavirus (MStaSV)
	Sigmavirus myga	Hubei diptera virus 10 (HbDV-10)
	Sigmavirus obscura	Drosophila obscura sigmavirus (DObsSV)
	Sigmavirus shayang	Shayang fly virus 2 (SyFV-1)
	Sigmavirus sturtevanti	Drosophila sturtevanti sigmavirus (DStuSV)
	Sigmavirus tristis	Drosophila tristis sigmavirus (DTriSV)
	Sigmavirus wuhan	Wuhan house fly virus 1 (WhHFV-1)
	Sigmavirus ying	Hubei dimarhabdovirus 1 (HbDRV-1)
Siniperhavirus	Siniperhavirus chuatsi	Chinese rice-field eel rhabdovirus (CrERV)
		hybrid snakehead rhabdovirus (HSHRV)
		Siniperca chuatsi rhabdovirus (SCRV)
		Micropterus salmoides rhabdovirus (MSRV)
	Siniperhavirus zoarces	eelpout rhabdovirus (EPRV)
Sprivivirus	Sprivivirus cyprinus	spring viremia of carp virus (SVCV)
	Sprivivirus esox	grass carp rhabdovirus (GrCRV)
		pike fry rhabdovirus (PFRV)
		tench rhabdovirus (TenRV)
Sripuvirus	Sripuvirus almpiwar	Almpiwar virus (ALMV)
	Sripuvirus chaco	Chaco virus (CHOV)
	Sripuvirus charleville	Charleville virus (CHVV)
	Sripuvirus cuiaba	Cuiaba virus (CUIV)
	Sripuvirus hainan	Hainan black-spectacled toad rhabdovirus (HnBSTRV)
	Sripuvirus madureira	Sena Madureira virus (SMV)
	Sripuvirus niakha	Niakha virus (NIAV)
	Sripuvirus sripur	Sripur virus (SRIV)
Sunrhavirus	Sunrhavirus alexandria	Burg el Arab virus (BEAV)
	Sunrhavirus bimbo	Bimbo virus (BBOV)
	Sunrhavirus boteke	Boteke virus (BOTV)
	Sunrhavirus dillard	Dillard’s Draw virus (DDRV)
	Sunrhavirus garba	Garba virus (GARV)
	Sunrhavirus harrison	Harrison Dam virus (HARDV)
	Sunrhavirus kolongo	Kolongo virus (KOLV)
	Sunrhavirus kwatta	Kwatta virus (KWAV)
	Sunrhavirus matariya	Matariya virus (MTYV)
	Sunrhavirus nasoule	Nasoule virus (NASV)
	Sunrhavirus oakvale	Oak Vale virus (OVV)
	Sunrhavirus ouango	Ouango virus (OUAV)
	Sunrhavirus sandjimba	Sandjimba virus (SJAV)
	Sunrhavirus sunguru	Sunguru virus (SUNV)
	Sunrhavirus walkabout	Walkabout Creek virus (WACV)
Tibrovirus	Tibrovirus alphaekpoma	Ekpoma virus 1 (EKV-1)
	Tibrovirus beatrice	Beatrice Hill virus (BHV)
	Tibrovirus betaekpoma	Ekpoma virus 2 (EKV-2)
	Tibrovirus coastal	Coastal Plains virus (CPV)
	Tibrovirus congo	Bas-Congo virus (BASV)
	Tibrovirus sweetwater	Sweetwater Branch virus (SWBV)
	Tibrovirus tibrogargan	Bivens Arm virus (BAV)
		Tibrogargan virus (TIBV)
Tupavirus	Tupavirus durham	Durham virus (DURV)
	Tupavirus klamath	Klamath virus (KLAV)
	Tupavirus tupaia	tupaia rhabdovirus (TUPV)
Vesiculovirus	Vesiculovirus alagoas	vesicular stomatitis Alagoas virus (VSAV)
	Vesiculovirus bogdanovac	Yug Bogdanovac virus (YBV)
	Vesiculovirus carajas	Carajás virus (CJSV)
	Vesiculovirus chandipura	Chandipura virus (CHPV)
	Vesiculovirus cocal	Cocal virus (COCV)
	Vesiculovirus eptesicus	American bat vesiculovirus (ABVV)
	Vesiculovirus indiana	vesicular stomatitis Indiana virus (VSIV)
	Vesiculovirus isfahan	Isfahan virus (ISFV)
	Vesiculovirus jurona	Jurona virus (JURV)
	Vesiculovirus malpais	Malpais Spring virus (MSPV)
	Vesiculovirus maraba	Maraba virus (MARAV)
	Vesiculovirus mediterranean	Mediterranean bat virus (MBV)
	Vesiculovirus morreton	Morreton virus (MORV)
	Vesiculovirus newjersey	vesicular stomatitis New Jersey virus (VSNJV)
	Vesiculovirus perinet	Perinet virus (PERV)
	Vesiculovirus piry	Piry virus (PIRYV)
	Vesiculovirus radi	Radi virus (RADV)
	Vesiculovirus rhinolophus	Jinghong bat virus (JhBV)
		Qiongzhong bat virus (QZBV)
	Vesiculovirus yinshui	Yinshui bat virus (YSBV)
Zarhavirus	Zarhavirus zahedan	Zahedan rhabdovirus (ZARV)
Subfamily Betarhabdovirinae
Alphanucleorhabdovirus	Alphanucleorhabdovirus colocasiae	taro vein chlorosis virus (TaVCV)
	Alphanucleorhabdovirus constrictae	constricta yellow dwarf virus (CYDV)
	Alphanucleorhabdovirus joa	joa yellow blotch associated virus (JYBaV)
	Alphanucleorhabdovirus maydis	maize mosaic virus (MMV)
	Alphanucleorhabdovirus melongenae	eggplant mottled dwarf virus (EMDV)
	Alphanucleorhabdovirus morogoromaydis	Morogoro maize-associated virus (MMaV)
	Alphanucleorhabdovirus oryzae	rice transitory yellowing virus (RTYV)
		rice yellow stunt virus (RYSV)
	Alphanucleorhabdovirus physostegiae	Physostegia chlorotic mottle virus (PhCMoV)
	Alphanucleorhabdovirus pruni	peach virus 1 (PeV1)
	Alphanucleorhabdovirus tritici	wheat yellow striate virus (WYSV)
	Alphanucleorhabdovirus tuberosum	potato yellow dwarf virus (PYDV)
	Alphanucleorhabdovirus zeairanense	maize Iranian mosaic virus (MIMV)
Betanucleorhabdovirus	Betanucleorhabdovirus bacopae	Bacopa monnieri virus 2 (BmV-2)
	Betanucleorhabdovirus cardamomi	cardamom vein clearing virus (CdVCV)
	Betanucleorhabdovirus daturae	datura yellow vein virus (DYVV)
	Betanucleorhabdovirus loti	birds-foot trefoil-associated virus (BFTV)
	Betanucleorhabdovirus mali	apple rootstock virus A (ApRVA)
	Betanucleorhabdovirus medicagonis	alfalfa-associated nucleorhabdovirus (AaNV)
	Betanucleorhabdovirus retesonchi	Sonchus yellow net virus (SYNV)
	Betanucleorhabdovirus ribes	blackcurrant-associated rhabdovirus (BCaRV)
	Betanucleorhabdovirus venasonchi	sowthistle yellow vein virus (SYVV)
	Betanucleorhabdovirus zanthoxyli	Zhuye pepper nucleorhabdovirus (ZPNRV)
Cytorhabdovirus	Cytorhabdovirus alphatrifolii	Trifolium pratense virus A (TpVA)
	Cytorhabdovirus alphawuhaninsectum	Wuhan insect virus 4 (WuIV-4)
	Cytorhabdovirus bacopae	Bacopa monnieri virus 1 (BmV-1)
	Cytorhabdovirus betatrifolii	Trifolium pratense virus B (TpVB)
	Cytorhabdovirus betawuhaninsectum	Wuhan insect virus 5 (WuIV-5)
	Cytorhabdovirus brassicae	broccoli necrotic yellows virus (BNYV)
	Cytorhabdovirus brassicicolae	cabbage cytorhabdovirus 1 (CCyV-1)
	Cytorhabdovirus broussonetiae	paper mulberry mosaic associated virus (PMuMaV)
	Cytorhabdovirus caricae	papaya virus E (PpVE)
	Cytorhabdovirus chrysanthemi	chrysanthemum yellow dwarf associated virus (CYDaV)
	Cytorhabdovirus colocasiae	Colocasia bobone disease-associated virus (CBDaV)
	Cytorhabdovirus cucurbitae	cucurbit cytorhabdovirus 1 (CuCV-1)
	Cytorhabdovirus festucae	Festuca leaf streak virus (FLSV)
	Cytorhabdovirus flaviyerbamate	yerba mate chlorosis-associated virus (YmCaV)
	Cytorhabdovirus fragariae	strawberry-associated virus 1 (SaV-1)
		strawberry virus 1 (StrV-1)
	Cytorhabdovirus fragariarugosus	strawberry crinkle virus (SCV)
	Cytorhabdovirus gammawuhaninsectum	Wuhan insect virus 6 (WuIV-6)
	Cytorhabdovirus gramineae	northern cereal mosaic virus (NCMV)
	Cytorhabdovirus hordei	barley yellow striate mosaic virus (BYSMV)
	Cytorhabdovirus lactucamaculante	lettuce yellow mottle virus (LYMoV)
	Cytorhabdovirus lactucanecante	lettuce necrotic yellows virus (LNYV)
	Cytorhabdovirus lycopersici	tomato yellow mottle-associated virus (TYMaV)
	Cytorhabdovirus maydis	maize-associated cytorhabdovirus (MaCV)
	Cytorhabdovirus maysflavostriatis	maize yellow striate virus (MYSV)
	Cytorhabdovirus medicagonis	alfalfa dwarf virus (ADV)
	Cytorhabdovirus oryzae	rice stripe mosaic virus (RSMV)
	Cytorhabdovirus persimmon	persimmon virus A (PeVA)
	Cytorhabdovirus rosae	rose virus R (RVR)
	Cytorhabdovirus rubus	raspberry vein chlorosis virus (RVCV)
	Cytorhabdovirus sonchi	Sonchus virus (SonV)
	Cytorhabdovirus trichosanthei	Trichosanthes associated rhabdovirus 1 (TrARV1)
	Cytorhabdovirus tritici	wheat American striate mosaic virus (WASMV)
	Cytorhabdovirus yerbamate	yerba mate virus A (YmVA)
Dichorhavirus	Dichorhavirus citri	citrus chlorotic spot virus (CiCSV)
	Dichorhavirus clerodendri	clerodendrum chlorotic spot virus (ClCSV)
	Dichorhavirus coffeae	coffee ringspot virus (CoRSV)
	Dichorhavirus leprosis	citrus leprosis virus N (CiLV-N)
	Dichorhavirus orchidaceae	orchid fleck virus (OFV)
Gammanucleorhabdovirus	Gammanucleorhabdovirus maydis	maize fine streak virus (MFSV)
Varicosavirus	Varicosavirus alopecuri	Alopecurus myosuroides varicosavirus 1 (AMVV-1)
	Varicosavirus lactucae	lettuce big-vein associated virus (LBVaV)
	Varicosavirus trifolii	red clover associated varicosavirus (RCaVV)
Subfamily Gammarhabdovirinae
Novirhabdovirus	Novirhabdovirus hirame	hirame rhabdovirus (HIRRV = HIRV)
	Novirhabdovirus piscine	viral hemorrhagic septicemia virus (VHSV)
	Novirhabdovirus salmonid	infectious hematopoietic necrosis virus (IHNV)
	Novirhabdovirus snakehead	snakehead rhabdovirus (SHRV)
Unassigned (to subfamilies)
Alphacrustrhavirus	Alphacrustrhavirus wenling	Wenling crustacean virus 10 (WlCV-10)
	Alphacrustrhavirus zhejiang	Wenling crustacean virus 11 (WlCV-11)
Alphadrosrhavirus	Alphadrosrhavirus hubei	Wuhan house fly virus 2 (WhHFV-2)
	Alphadrosrhavirus shayang	Shayang fly virus 3 (SyFV-3)
Alphahymrhavirus	Alphahymrhavirus cinereus	hymenopteran rhabdo-related virus 38 (HyRRV-38)
	Alphahymrhavirus hirtum	hymenopteran rhabdo-related virus 109 (HyRRV-109)
	Alphahymrhavirus neglectus	Lasius neglectus virus 2 (LnegV-2)
	Alphahymrhavirus radians	hymenopteran rhabdo-related virus 46 (HyRRV-46)
Betahymrhavirus	Betahymrhavirus austriaca	hymenopteran rhabdo-related virus 23 (HyRRV-23)
	Betahymrhavirus heterodontonyx	hymenopteran rhabdo-related virus 24 (HyRRV-24)
Betanemrhavirus	Betanemrhavirus hubei	Hubei rhabdo-like virus 9 (HbRLV-9)
	Betanemrhavirus shayang	Shayang ascaridia galli virus 2 (SyAGV-2)
Betapaprhavirus	Betapaprhavirus frugiperda	Spodoptera frugiperda rhabdovirus (SfruRV)
	Betapaprhavirus sylvina	lepidopteran rhabdo-related virus 34 (LeRRV-34)
Betaricinrhavirus	Betaricinrhavirus chimay	Chimay rhabdovirus (CRV)
	Betaricinrhavirus scapularis	blacklegged tick rhabdovirus 1 (BLTRV-1)
Family Sunviridae
Sunshinevirus	Sunshinevirus reptilis	Sunshine Coast virus (SunCV)
Family Xinmoviridae
Alasvirus	Alasvirus muscae	Húběi diptera virus 11 (HbDV-11)
Anphevirus	Anphevirus xinchengense	Xīnchéng mosquito virus (XcMV)
Doupovirus	Doupovirus australiaense	Culex mononega-like virus 2 (CMLV-2)
Draselvirus	Draselvirus dentati	Húběi rhabdo-like virus 7 (HbRLV-7)
Drunivirus	Drunivirus chambonense	Drosophila unispina virus 1 (DuniV-1)
Gambievirus	Gambievirus bolahunense	Bolahun virus (BLHV)
	Gambievirus senegalense	Gambie virus (GAMV)
Gylbovirus	Gylbovirus aagae	Aedes anphevirus (AeAV)
Hoptevirus	Hoptevirus orthopteris	Húběi orthoptera virus 5 (HbOV-5)
Madalivirus	Madalivirus amapaense	Anopheles marajoara virus (AnMV)
	Madalivirus amazonaense	Anopheles darlingi virus (AnDV)
Pelmivirus	Pelmivirus eymattense	hymenopteran anphe-related virus OKIAV71 (HARV-71)
Triniovirus	Triniovirus yonagoense	Culex tritaeniorhynchus anphevirus (CtAV)
Ulegvirus	Ulegvirus freckenfeldense	odonatan anphe-related virus OKIAV59 (OARV-59)

Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, subfamilies, families, and orders are taxa.

¹Includes: Newcastle disease virus (NDV).

^¶Taxon names are always italicized and always begin with a capital letter.

^&Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization. Lists of viruses within a given species are provisional at this point and will likely be amended in the near future.

Table 5.

ICTV-accepted taxonomy of the order Goujianvirales (Negarnaviricota: Haploviricotina: Yunchangviricetes) as of March 2022.

Genus	Species¶	Virus (Abbreviation)&
Family Yueviridae
Yuyuevirus	Yuyuevirus beihaiense	Běihǎi sesarmid crab virus 3 (BhSCV-3)
	Yuyuevirus shaheense	Shāhé yuèvirus-like virus 1 (ShYLV-1)

Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, families, and orders are taxa.

^¶Taxon names are always italicized and always begin with a capital letter.

^&Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization.

Table 6.

ICTV-accepted taxonomy of the order Bunyavirales (Negarnaviricota: Polyploviricotina: Ellioviricetes) as of March 2022.

Genus	Species¶	Virus (Abbreviation)&
Family Arenaviridae
Antennavirus	Hairy antennavirus	Wēnlǐng frogfish arenavirus 2 (WlFAV-2)
	Salmon antennavirus	salmon pescarenavirus 1 (SPAV-1)
		salmon pescarenavirus 2 (SPAV-2)
	Striated antennavirus	Wēnlǐng frogfish arenavirus 1 (WlFAV-1)
Hartmanivirus	Haartman hartmanivirus	Haartman Institute snake virus 1 (HISV-1)
		Haartman Institute snake virus 2 (HISV-2)
	Heimat hartmanivirus	andere Heimat virus 1 (aHeV-1)
	Muikkunen hartmanivirus	Dante Muikkunen virus 1 (DaMV-1)
	Schoolhouse hartmanivirus	old schoolhouse virus 1 (OScV-1)
		old schoolhouse virus 2 (OScV-2)
	Setpatvet hartmanivirus	SetPatVet virus 1 (SPVV-1)
	Zurich hartmanivirus	veterinary pathology Zurich virus 1 (VPZV-1)
		veterinary pathology Zurich virus 2 (VPZV-2)
Mammarenavirus	Allpahuayo mammarenavirus	Allpahuayo virus (ALLV)
	Alxa mammarenavirus	Alxa virus (ALXV)
	Argentinian mammarenavirus	Junín virus (JUNV)
	Bear Canyon mammarenavirus	Bear Canyon virus (BCNV)
	Brazilian mammarenavirus	Sabiá virus (SBAV)
	Cali mammarenavirus	Pichindé virus (PICHV)
	Chapare mammarenavirus	Chapare virus (CHAPV)
	Chevrier mammarenavirus	Lìjiāng virus (LIJV)
	Cupixi mammarenavirus	Cupixi virus (CUPXV)
	Flexal mammarenavirus	Flexal virus (FLEV)
	Gairo mammarenavirus	Gairo virus (GAIV)
	Guanarito mammarenavirus	Guanarito virus (GTOV)
	Ippy mammarenavirus	Ippy virus (IPPYV)
	Kitale mammarenavirus	Kitale virus (KTLV)
	Lassa mammarenavirus	Lassa virus (LASV)
	Latino mammarenavirus	Latino virus (LATV)
	Loei River mammarenavirus	Loei River virus (LORV)
	Lujo mammarenavirus	Lujo virus (LUJV)
	Luna mammarenavirus	Luli virus (LULV)
		Luna virus (LUAV)
	Lunk mammarenavirus	Lunk virus (LNKV)
	Lymphocytic choriomeningitis mammarenavirus	Dandenong virus (DANV)
		lymphocytic choriomeningitis virus (LCMV)
	Machupo mammarenavirus	Machupo virus (MACV)
	Mammarenavirus bituense	Bitu virus (BITV)
	Mammarenavirus kwanzaense	Kwanza virus (KWAV)
	Mariental mammarenavirus	Mariental virus (MRLV)
	Merino Walk mammarenavirus	Merino Walk virus (MRWV)
	Mobala mammarenavirus	mobala virus (MOBV)
	Mopeia mammarenavirus	Mopeia virus (MPOV)
		Morogoro virus (MORV)
	Okahandja mammarenavirus	Okahandja virus (OKAV)
	Oliveros mammarenavirus	Oliveros virus (OLVV)
	Paraguayan mammarenavirus	Paraná virus (PRAV)
	Planalto mammarenavirus	Aporé virus (APOV)
	Pirital mammarenavirus	Pirital virus (PIRV)
	Ryukyu mammarenavirus	Ryukyu virus (RYKV)
	Serra do Navio mammarenavirus	Amaparí virus (AMAV)
	Solwezi mammarenavirus	Solwezi virus (SOLV)
	Souris mammarenavirus	souris virus (SOUV)
	Tacaribe mammarenavirus	Tacaribe virus (TCRV)
	Tamiami mammarenavirus	Tamiami virus (TMMV)
	Wenzhou mammarenavirus	Wēnzhōu virus (WENV)
	Whitewater Arroyo mammarenavirus	Big Brushy Tank virus (BBRTV)
		Catarina virus (CTNV)
		Skinner Tank virus (SKTV)
		Tonto Creek virus (TTCV)
		Whitewater Arroyo virus (WWAV)
	Xapuri mammarenavirus	Xapuri virus (XAPV)
Reptarenavirus	California reptarenavirus	CAS virus (CASV)
	Giessen reptarenavirus	University of Giessen virus 1 (UGV-1)
		University of Giessen virus 2 (UGV-2)
		University of Giessen virus 3 (UGV-3)
	Golden reptarenavirus	Golden Gate virus (GOGV)
	Ordinary reptarenavirus	tavallinen suomalainen mies virus 2 (TSMV-2)
	Rotterdam reptarenavirus	ROUT virus (ROUTV)
		University of Helsinki virus 1 (UHV-1)
Family Cruliviridae
Lincruvirus	Lincruvirus europense	European shore crab virus 1 (EscV-1)
	Lincruvirus sinense	Chinese mitten crab virus 1 (CmcV-1)
	Lincruvirus wenlingense	Wēnlǐng crustacean virus 9 (WlCV-9)
Family Discoviridae
Orthodiscovirus	Orthodiscovirus coniellae	Coniothyrium diplodiella negative-stranded RNA virus 1 (CdNSRV-1)
		Plasmopara viticola lesion associated mycobunyavirales-like virus 9 (PvLAM-LV-9)
	Orthodiscovirus hispaniae	Plasmopara viticola lesion associated mycobunyavirales-like virus 8 (PvLAM-LV8)
	Orthodiscovirus iberiae	Plasmopara viticola lesion associated mycobunyavirales-like virus 4 (PvLAM-LV4)
	Orthodiscovirus missouriense	Penicillium discovirus (PDV)
	Orthodiscovirus penicillii	Penicillium roseopurpureum negative ssRNA virus 1 (PrNssV-1)
Family Fimoviridae
Emaravirus	Emaravirus aceris	maple mottle-associated virus (MaMaV)
	Emaravirus actinidiae	Actinidia chlorotic ringspot-associated virus (AcCRaV)
	Emaravirus cajani	pigeonpea sterility mosaic virus 1 (PPSMV-1)
	Emaravirus camelliae	Camellia japonica-associated virus 1 (CjaV-1)
	Emaravirus cercidis	redbud yellow ringspot-associated virus (RYRaV)
	Emaravirus chrysanthemi	chrysanthemum mosaic-associated virus (ChMaV)
	Emaravirus cordylinae	ti ringspot-associated virus (TiRSaV)
	Emaravirus fici	fig mosaic virus (FMV)
	Emaravirus idaeobati	raspberry leaf blotch virus (RLBV)
	Emaravirus kiwii	Actinidia virus 2 (AcV-2)
	Emaravirus parkinsoniae	palo verde broom virus (PVBV)
	Emaravirus perillae	perilla mosaic virus (PerMV)
	Emaravirus pistaciae	Pistacia virus B (PiVB)
	Emaravirus populi	aspen mosaic-associated virus (AsMaV)
	Emaravirus pyri	pear chlorotic leaf spot-associated virus (PCLSaV)
	Emaravirus quercus	common oak ringspot-associated virus (CORaV)
	Emaravirus rosae	rose rosette virus (RRV)
	Emaravirus rubi	blackberry leaf mottle-associated virus (BLMaV)
	Emaravirus sorbi	European mountain ash ringspot-associated virus (EMARaV)
	Emaravirus syringae	lilac chlorotic ringspot-associated virus (LiCRaV)
	Emaravirus toordali	pigeonpea sterility mosaic virus 2 (PPSMV-2)
	Emaravirus tritici	High Plains wheat mosaic virus (HPWMoV)
	Emaravirus verbanni	Camellia japonica-associated virus 2 (CjaV-2)
	Emaravirus ziziphi	jujube yellow mottle-associated virus (JYMaV)
Family Hantaviridae
Subfamily Actantavirinae
Actinovirus	Batfish actinovirus	Wēnlǐng minipizza batfish virus (WEMBV)
	Goosefish actinovirus	Wēnlǐng yellow goosefish virus (WEYGV)
	Perch actinovirus	Bern perch virus (BRPV)
	Spikefish actinovirus	Wēnlǐng red spikefish virus (WERSV)
Subfamily Agantavirinae
Agnathovirus	Hagfish agnathovirus	Wēnlǐng hagfish virus (WEHV)
Subfamily Mammantavirinae
Loanvirus	Brno loanvirus	Brno virus (BRNV)
	Longquan loanvirus	Lóngquán virus (LQUV)
Mobatvirus	Laibin mobatvirus	Láibīn virus (LAIV)
	Lena mobatvirus	Lena virus (LENV)
	Nova mobatvirus	Nova virus (NVAV)
	Quezon mobatvirus	Quezon virus (QZNV)
	Xuan Son mobatvirus	Xuân Sơn virus (XSV)
Orthohantavirus	Andes orthohantavirus	Andes virus (ANDV)
		Castelo dos Sonhos virus (CASV)
		Lechiguanas virus (LECV = LECHV)
		Orán virus (ORNV)
	Asama orthohantavirus	Asama virus (ASAV)
	Asikkala orthohantavirus	Asikkala virus (ASIV)
	Bayou orthohantavirus	bayou virus (BAYV)
		Catacamas virus (CATV)
	Black Creek Canal orthohantavirus	Black Creek Canal virus (BCCV)
	Bowe orthohantavirus	Bowé virus (BOWV)
	Bruges orthohantavirus	Bruges virus (BRGV)
	Cano Delgadito orthohantavirus	Caño Delgadito virus (CADV)
	Cao Bang orthohantavirus	Cao Bằng virus (CBNV)
		Liánghé virus (LHEV)
	Choclo orthohantavirus	Choclo virus (CHOV)
	Dabieshan orthohantavirus	Dàbiéshān virus (DBSV)
	Dobrava-Belgrade orthohantavirus	Dobrava virus (DOBV)
		Kurkino virus (KURV)
		Saaremaa virus (SAAV)
		Sochi virus (SOCV)
	El Moro Canyon orthohantavirus	Carrizal virus (CARV)
		El Moro Canyon virus (ELMCV)
		Huitzilac virus (HUIV)
	Fugong orthohantavirus	Fúgòng virus (FUGV)
	Fusong orthohantavirus	Fǔsōng virus (FUSV)
	Hantaan orthohantavirus	Amur virus (AMRV)
		Hantaan virus (HTNV)
		Soochong virus (SOOV)
	Jeju orthohantavirus	Jeju virus (JJUV)
	Kenkeme orthohantavirus	Kenkeme virus (KKMV)
	Khabarovsk orthohantavirus	Khabarovsk virus (KHAV)
		Topografov virus (TOPV)
	Laguna Negra orthohantavirus	Laguna Negra virus (LANV)
		Maripa virus (MARV)
		Rio Mamoré virus (RIOMV)
	Luxi orthohantavirus	Lúxī virus (LUXV)
	Maporal orthohantavirus	Maporal virus (MAPV)
	Montano orthohantavirus	Montaño virus (MTNV)
	Necocli orthohantavirus	Necoclí virus (NECV)
	Oxbow orthohantavirus	Oxbow virus (OXBV)
	Prospect Hill orthohantavirus	Prospect Hill virus (PHV)
	Puumala orthohantavirus	Hokkaido virus (HOKV)
		Muju virus (MUJV)
		Puumala virus (PUUV)
	Robina orthohantavirus	Robina virus (ROBV)
	Rockport orthohantavirus	Rockport virus (RKPV)
	Sangassou orthohantavirus	Sangassou virus (SANGV)
	Seewis orthohantavirus	Seewis virus (SWSV)
	Seoul orthohantavirus	gōu virus (GOUV)
		Seoul virus (SEOV)
	Sin Nombre orthohantavirus	New York virus (NYV)
		Sin Nombre virus (SNV)
	Tatenale orthohantavirus	Tatenale virus (TATV)
	Thailand orthohantavirus	Anjozorobe virus (ANJZV)
		Serang virus (SERV)
		Thailand virus (THAIV)
	Tigray orthohantavirus	Tigray virus (TIGV)
	Tula orthohantavirus	Adler virus (ADLV)
		Tula virus (TULV)
	Yakeshi orthohantavirus	Yákèshí virus (YKSV)
Thottimvirus	Imjin thottimvirus	Imjin virus (MJNV)
	Thottapalayam thottimvirus	Thottapalayam virus (TPMV)
Subfamily Repantavirinae
Reptillovirus	Gecko reptillovirus	Hǎinán oriental leaf-toed gecko virus (HOLGV)
Family Leishbuviridae
Shilevirus	Shilevirus leptomonadis	Leptomonas moramango virus (LEPMV)
Family Mypoviridae
Hubavirus	Hubavirus myriapedis	Húběi myriapoda virus 5 (HbMV-5)
Family Nairoviridae
Norwavirus	Grotenhout norwavirus	Grotenhout virus (GRHV)
Ocetevirus	Blattodean ocetevirus	red goblin roach virus 1 (RGRV-1)
Orthonairovirus	Abu Hammad orthonairovirus	Abū Ḥammād virus (AHV)
	Abu Mina orthonairovirus	Abū Mīnā virus (AMV)
	Artashat orthonairovirus	Artashat virus (ARTSV)
	Avalon orthonairovirus	Avalon virus (AVAV)
	Chim orthonairovirus	Chim virus (CHIMV)
	Bandia orthonairovirus	Bandia virus (BDAV)
	Burana orthonairovirus	Burana virus (BURV)
	Congoid orthonairovirus	Aigai virus (AIGV)
	Crimean-Congo hemorrhagic fever orthonairovirus	Crimean-Congo hemorrhagic fever virus (CCHFV)
	Dera Ghazi Khan orthonairovirus	Dera Ghazi Khan virus (DGKV)
	Dugbe orthonairovirus	Dugbe virus (DUGV)
	Erve orthonairovirus	Erve virus (ERVEV)
	Estero Real orthonairovirus	Estero Real virus (ERV)
	Gossas orthonairovirus	Gossas virus (GOSV)
	Hazara orthonairovirus	Hazara virus (HAZV)
	Huangpi orthonairovirus	Huángpí tick virus 1 (HpTV-1)
	Hughes orthonairovirus	Caspiy virus (CASV)
		Farallon virus (FARV)
		Great Saltee virus (GRSV)
		Hughes virus (HUGV)
		Raza virus (RAZAV)
	Issyk-kul orthonairovirus	Issyk-kul virus (ISKV)
	Kasokero orthonairovirus	Kasokero virus (KASV = KASOV)
	Keterah orthonairovirus	Keterah virus (KTRV)
		Uzun-Agach virus (UZAV)
	Kupe orthonairovirus	kupe virus (KUPEV)
	Leopards Hill orthonairovirus	Leopards Hill virus (LPHV)
	Meram orthonairovirus	Meram virus (MEMV)
	Nairobi sheep disease orthonairovirus	Nairobi sheep disease virus (NSDV)
	Pacific Coast orthonairovirus	Pacific Coast tick nairovirus (PCTNV)
	Punta orthonairovirus	Punta Salinas virus (PSV)
	Qalyub orthonairovirus	Geran virus (GERV)
		Qalyub virus (QYBV)
	Sakhalin orthonairovirus	Sakhalin virus (SAKV)
		Tillamook virus (TILLV)
	Sapphire orthonairovirus	Sapphire II virus (SAPV)
	Scot orthonairovirus	Clo Mor virus (CMV = CLMV)
	Soldado orthonairovirus	Soldado virus (SOLV)
	Tacheng orthonairovirus	Tǎchéng tick virus 1 (TcTV-1)
	Taggert orthonairovirus	Taggert virus (TAGV)
	Tamdy orthonairovirus	Tamdy virus (TAMV)
	Thiafora orthonairovirus	Thiafora virus (TFAV)
	Tofla orthonairovirus	tofla virus (TFLV)
	Tunis orthonairovirus	Tunis virus (TUNV)
	Vinegar Hill orthonairovirus	Vinegar Hill virus (VINHV)
	Wenzhou orthonairovirus	Wēnzhōu tick virus (WzTV)
	Yogue orthonairovirus	Yogue virus (YOGV)
	Zirqa orthonairovirus	Zirqa virus (ZIRV)
Sabavirus	South Bay sabavirus	South Bay virus (SBV)
Shaspivirus	Spider shaspivirus	Shāyáng spider virus 1 (SySV-1)
Striwavirus	Strider striwavirus	Sānxiá water strider virus 1 (SxWSV-1)
Xinspivirus	Xinzhou xinspivirus	Xīnzhōu spider virus (XSV)
Family Peribunyaviridae
Herbevirus	Herbert herbevirus	Herbert virus (HEBV)
	Kibale herbevirus	Kibale virus (KIBV)
	Tai herbevirus	Taï virus (TAIV)
Khurdivirus	Khurdivirus volgaense	Khurdun virus (KURV)
Lakivirus	Lakivirus lakamhaense	Lakamha virus (LAKV)
Lambavirus	Lambavirus wisconsinense	largemouth bass bunyavirus (LBBV)
Orthobunyavirus	Abras orthobunyavirus	Abras virus (ABRV)
	Acara orthobunyavirus	Acará virus (ACAV)
		Moriche virus (MORV)
	Aino orthobunyavirus	Aino virus (AINOV)
	Akabane orthobunyavirus	Akabane virus (AKAV)
		Tinaroo virus (TINV)
		Yaba-7 virus (Y7V)
	Alajuela orthobunyavirus	Alajuela virus (ALJV)
		San Juan virus (SJV)
	Anadyr orthobunyavirus	Anadyr virus (ANADV)
	Ananindeua orthobunyavirus	Ananindeua virus (ANUV)
	Anhembi orthobunyavirus	Anhembi virus (AMBV)
	Anopheles A orthobunyavirus	Anopheles A virus (ANAV)
		Arumateua virus (ARTV = ARMTV)
		Caraipé virus (CPEV = CRPV)
		Las Maloyas virus (LMV)
		Lukuni virus (LUKV)
		Trombetas virus (TRMV)
		Tucuruí virus (TUCV = TUCRV)
	Anopheles B orthobunyavirus	Anopheles B virus (ANBV)
		Boracéia virus (BORV)
	Apeu orthobunyavirus	Apeú virus (APEUV)
	Bakau orthobunyavirus	Bakau virus (BAKV)
		Ketapang virus (KETV)
		Nola virus (NOLAV)
		Tanjong Rabok virus (TRV)
		Telok Forest virus (TFV)
	Batai orthobunyavirus	Batai virus (BATV)
	Batama orthobunyavirus	Batama virus (BMAV)
	Bellavista orthobunyavirus	Bellavista virus (BELLV)
	Benevides orthobunyavirus	Benevides virus (BVSV = BENV)
	Bertioga orthobunyavirus	Bertioga virus (BERV)
		Cananéia virus (CNAV)
		Guaratuba virus (GTBV)
		Itimirim virus (ITIV)
		Mirim virus (MIRV)
	Bimiti orthobunyavirus	bimiti virus (BIMV)
	Birao orthobunyavirus	Birao virus (BIRV)
	Botambi orthobunyavirus	Botambi virus (BOTV)
	Bozo orthobunyavirus	Bozo virus (BOZOV)
	Brazoran orthobunyavirus	brazoran virus (BRAZV)
	Bruconha orthobunyavirus	Bruconha virus (BRUV)
	Buffalo Creek orthobunyavirus	Buffalo Creek virus (BUCV)
	Bunyamwera orthobunyavirus	Bunyamwera virus (BUNV)
		Germiston virus (GERV)
		Lokern virus (LOKV)
		Mboké virus (MBOV)
		Ngari virus (NRIV)
		Northway virus (NORV)
		Santa Rosa virus (SARV)
		Shokwe virus (SHOV)
		Stanfield virus (STAV)
		Xingu virus (XINV)
	Bushbush orthobunyavirus	Benfica virus (BENV = BNFV)
		Bushbush virus (BSBV)
		Juan Díaz virus (JDV)
	Buttonwillow orthobunyavirus	Buttonwillow virus (BUTV)
	Bwamba orthobunyavirus	Bwamba virus (BWAV)
		Pongola virus (PGAV)
	Cache Valley orthobunyavirus	Cache Valley virus (CVV)
		Cholul virus (CHLV)
		Tlacotalpan virus (TLAV)
	Cachoeira Porteira orthobunyavirus	Cachoeira Porteira virus (CPOV)
	California encephalitis orthobunyavirus	California encephalitis virus (CEV)
		Morro Bay virus (MBV)
	Capim orthobunyavirus	Capim virus (CAPV)
	Caraparu orthobunyavirus	Caraparú virus (CARV)
		El Huayo virus (EHUV)
		Itaya virus (ITYV)
		Ossa virus (OSSAV)
		Vinces virus (VINV)
	Cat Que orthobunyavirus	Cát Quế virus (CQV)
		Oya virus (OYAV)
	Catu orthobunyavirus	Catú virus (CATUV)
	Enseada orthobunyavirus	Enseada virus (ENSV)
	Faceys paddock orthobunyavirus	Facey’s paddock virus (FPV)
	Fort Sherman orthobunyavirus	Fort Sherman virus (FSV)
	Gamboa orthobunyavirus	Brus Laguna virus (BLAV)
		Calchaquí virus (CQIV)
		Gamboa virus (GAMV)
		Pueblo Viejo virus (PVV)
		Soberanía virus (SOBV)
	Gan Gan orthobunyavirus	Gan Gan virus (GGV)
	Guajara orthobunyavirus	Guajará virus (GJAV)
	Guama orthobunyavirus	Guamá virus (GMAV)
	Guaroa orthobunyavirus	Guaroa virus (GROV)
	Iaco orthobunyavirus	Iaco virus (IACOV)
	Ilesha orthobunyavirus	Ilesha virus (ILEV)
	Ingwavuma orthobunyavirus	Ingwavuma virus (INGV)
	Jamestown Canyon orthobunyavirus	Inkoo virus (INKV)
		Jamestown Canyon virus (JCV)
		Jerry Slough virus (JSV)
		South River virus (SORV)
	Jatobal orthobunyavirus	Jatobal virus (JATV)
	Kaeng Khoi orthobunyavirus	Kaeng Khoi virus (KKV)
	Kairi orthobunyavirus	kairi virus (KRIV)
	Keystone orthobunyavirus	Keystone virus (KEYV)
	Koongol orthobunyavirus	koongol virus (KOOV)
		wongal virus (WONV)
	La Crosse orthobunyavirus	La Crosse virus (LACV)
	Leanyer orthobunyavirus	Leanyer virus (LEAV)
	Lumbo orthobunyavirus	Lumbo virus (LUMV)
	Macaua orthobunyavirus	Macauã virus (MCAV)
	Madrid orthobunyavirus	Madrid virus (MADV)
	Maguari orthobunyavirus	Maguari virus (MAGV)
		Playas virus (PLAV)
	Mahogany Hammock orthobunyavirus	Mahogany Hammock virus (MHV)
	Main Drain orthobunyavirus	Main Drain virus (MDV)
	Manzanilla orthobunyavirus	Manzanilla virus (MANV)
		Inini virus (INIV)
	Mapputta orthobunyavirus	Mapputta virus (MAPV)
	Maprik orthobunyavirus	Maprik virus (MPKV)
	Marituba orthobunyavirus	Gumbo Limbo virus (GLV)
		Marituba virus (MTBV)
		Murutucú virus (MURV)
		Nepuyo virus (NEPV)
		Restan virus (RESV)
		Zungarococha virus (ZUNV)
	Matruh orthobunyavirus	Matruh virus (MTRV)
	Melao orthobunyavirus	Melao virus (MELV)
	Moju orthobunyavirus	Moju virus (MOJUV)
	Mermet orthobunyavirus	Mermet virus (MERV)
	Minatitlan orthobunyavirus	Minatitlán virus (MNTV)
		Palestina virus (PLSV)
	MPoko orthobunyavirus	M'Poko virus (MPOV)
		Yaba-1 virus (Y1V)
	Nyando orthobunyavirus	Eretmapodites virus (ERETV)
		Mojuí dos Campos virus (MDCV)
		Nyando virus (NDV)
	Olifantsvlei orthobunyavirus	Bobia virus (BIAV)
		Dabakala virus (DABV)
		Olifantsvlei virus (OLIV)
		Oubi virus (OUBIV)
	Oriboca orthobunyavirus	Itaquí virus (ITQV)
		Oriboca virus (ORIV)
	Oropouche orthobunyavirus	Iquitos virus (IQTV)
		Madre de Dios virus (MDDV)
		Oropouche virus (OROV)
		Perdões virus (PDEV)
		Pintupo virus (PINTV)
	Oyo orthobunyavirus	Oyo virus (OYOV)
	Patois orthobunyavirus	Babahoya virus (BABV)
		Pahayokee virus (PAHV)
		Patois virus (PATV)
	Peaton orthobunyavirus	peaton virus (PEAV)
	Potosi orthobunyavirus	Potosi virus (POTV)
	Sabo orthobunyavirus	Sabo virus (SABOV)
	San Angelo orthobunyavirus	San Angelo virus (SAV)
	Sango orthobunyavirus	Sango virus (SANV)
	Schmallenberg orthobunyavirus	Douglas virus (DOUV)
		Sathuperi virus (SATV)
		Schmallenberg virus (SBV)
		Shamonda virus (SHAV)
	Sedlec orthobunyavirus	Sedlec virus (SEDV)
	Serra do Navio orthobunyavirus	Serra do Navio virus (SDNV)
	Shark River orthobunyavirus	Shark River virus (SRV)
	Shuni orthobunyavirus	Kaikalur virus (KAIV)
		Shuni virus (SHUV)
	Simbu orthobunyavirus	Para virus (PARAV)
		Simbu virus (SIMV)
	Snowshoe hare orthobunyavirus	Khatanga virus (KHATV)
		snowshoe hare virus (SSHV)
	Sororoca orthobunyavirus	Sororoca virus (SORV)
	Tacaiuma orthobunyavirus	CoAr 1071 virus (CA1071V)
		CoAr 3627 virus (CA3626V)
		Tacaiuma virus (TCMV)
		Virgin River virus (VRV)
	Tahyna orthobunyavirus	Ťahyňa virus (TAHV)
	Tataguine orthobunyavirus	Tataguine virus (TATV)
	Tensaw orthobunyavirus	Tensaw virus (TENV)
	Tete orthobunyavirus	Bahig virus (BAHV)
		Tete virus (TETEV)
		Tsuruse virus (TSUV)
		Weldona virus (WELV)
	Thimiri orthobunyavirus	Thimiri virus (THIV)
	Timboteua orthobunyavirus	Timboteua virus (TBTV)
	Triniti orthobunyavirus	Triniti virus (TNTV)
	Trivittatus orthobunyavirus	Achiote virus (ACHOV)
		Trivittatus virus (TVTV)
	Turlock orthobunyavirus	Lednice virus (LEDV)
		Turlock virus (TURV)
		Umbre virus (UMBV)
	Utinga orthobunyavirus	Utinga virus (UTIV)
	Witwatersrand orthobunyavirus	Witwatersrand virus (WITV)
	Wolkberg orthobunyavirus	Wolkberg virus (WBV)
	Wyeomyia orthobunyavirus	Rio Pracupi virus
		Taiassui virus (TAIAV)
		Tucunduba virus (TUCV)
		Wyeomyia virus (WYOV)
Pacuvirus	Caimito pacuvirus	Caimito virus (CAIV)
	Chilibre pacuvirus	Chilibre virus (CHIV)
	Pacui pacuvirus	Pacui virus (PACV)
	Rio Preto da Eva pacuvirus	Rio Preto da Eva virus (RPEV)
	Tapirape pacuvirus	Tapirapé virus (TAPV)
Shangavirus	Insect shangavirus	Shuāngào insect virus 1 (SgIV-1)
Family Phasmaviridae
Feravirus	Ferak feravirus	Ferak virus (FRKV)
	Hemipteran feravirus	hemipteran phasma-related virus OKIAV247 (HeFV)
	Neuropteran feravirus	neuropteran phasma-related virus OKIAV248 (NeFV)
Hymovirus	Hymenopteran hymovirus 1	hymenopteran phasma-related virus OKIAV252 (HyHV-1)
	Hymenopteran hymovirus 2	hymenopteran phasma-related virus OKIAV250 (HyHV-2)
Jonvirus	Jonchet jonvirus	jonchet virus (JONV)
Orthophasmavirus	Anopheles orthophasmavirus	Anopheles triannulatus orthophasmavirus (AtOPV)
	Coleopteran orthophasmavirus	coleopteran phasma-related virus OKIAV235 (CPRV)
	Culex orthophasmavirus	Culex phasma-like virus (CPLV)
	Ganda orthophasmavirus	Ganda bee virus (GBEEV)
	Hymenopteran orthophasmavirus 1	hymenopteran phasma-related virus OKIAV228 (HyOV-1)
	Hymenopteran orthophasmavirus 2	hymenopteran phasma-related virus OKIAV227 (HyOV-2)
	Kigluaik phantom orthophasmavirus	Kigluaik phantom virus (KIGV)
	Niukluk phantom orthophasmavirus	Niukluk phantom virus (NUKV)
	Odonate orthophasmavirus	Húběi odonate virus 8 (HbOV-8)
	Orthophasmavirus barstukasense	Barstukas virus (BARV)
	Orthophasmavirus coredoense	Coredo virus (CORV)
	Orthophasmavirus flenense	Flen virus (FLNV)
	Orthophasmavirus miglotasense	Miglotas virus (MIGV)
	Qingling orthophasmavirus	Húběi odonate virus 9 (HbOV-9)
	Wuchang cockroach orthophasmavirus 1	Wǔchāng cockroach virus 1 (WcCV-1)
	Wuhan mosquito orthophasmavirus 1	Wǔhàn mosquito virus 1 (WhMV-1)
	Wuhan mosquito orthophasmavirus 2	Wǔhàn mosquito virus 2 (WhMV-2)
Sawastrivirus	Sanxia sawastrivirus	Sānxiá water strider virus 2 (SxWSV-2)
Wuhivirus	Insect wuhivirus	Wǔhàn insect virus 2 (WhIV-2)
Family Phenuiviridae
Bandavirus	Bhanja bandavirus	Bhanja virus (BHAV)
	Dabie bandavirus	severe fever with thrombocytopenia syndrome virus (SFTSV)
	Guertu bandavirus	Gùěrtú virus (GTV)
	Heartland bandavirus	Heartland virus (HRTV)
	Hunter Island bandavirus	Albatross Island virus (ABIV)
	Kismaayo bandavirus	Kismaayo virus (KISV)
	Lone star bandavirus	lone star virus (LSV)
	Razdan bandavirus	Razdan virus (RAZV)
Beidivirus	Dipteran beidivirus	Húběi diptera virus 3 (HbDV-3)
Coguvirus	Citrus coguvirus	citrus concave gum-associated virus (CCGaV)
	Coguvirus eburi	citrus virus A (CiV-A)
	Grapevine coguvirus	grapevine associated cogu-like virus 1 (GaCLV-1)
Entovirus	Entoleuca entovirus	Entoleuca phenui-like virus 1 (EnPLV-1)
Goukovirus	Cumuto goukovirus	Cumuto virus (CUMV)
	Gouleako goukovirus	Gouléako virus (GOLV)
	Yichang insect goukovirus	Yíchāng insect virus (YcIV)
Horwuvirus	Horsefly horwuvirus	Wǔhàn horsefly virus (WhHV)
	Kimberley horwuvirus	Fitzroy Crossing tenui-like virus 1 (FCTenV1)
Hudivirus	Dipteran hudivirus	Húběi diptera virus 4 (HbDV-4)
Hudovirus	Lepidopteran hudovirus	Húběi lepidoptera virus 1 (HbLV-1)
Ixovirus	Blackleg ixovirus	blacklegged tick virus 1 (BLTV-1)
	Norway ixovirus	Fairhair virus (FHAV)
	Scapularis ixovirus	blacklegged tick virus 3 (BLTV-3)
Laulavirus	Grapevine laulavirus 2	grapevine associated cogu-like virus 2 (GaCLV-2)
	Grapevine laulavirus 3	grapevine associated cogu-like virus 3 (GaCLV-3)
	Grapevine laulavirus 4	grapevine associated cogu-like virus 4 (GaCLV-4)
	Laurel Lake laulavirus	Laurel Lake virus (LLV)
Lentinuvirus	Lentinula lentinuvirus	Lentinula edodes negative-strand RNA virus 2 (LeNSRV-2)
Mobuvirus	Mothra mobuvirus	Mothra virus (MTHV)
	Narangue mobuvirus	Ñarangue virus (NRGV)
Phasivirus	Badu phasivirus	Badu virus (BADUV)
	Dipteran phasivirus	Húběi diptera virus 5 (HbDV-5)
	Fly phasivirus	Wǔhàn fly virus 1 (WhFV-1)
	Guadeloupe phasivirus	Guadeloupe mosquito phasivirus (GMPV)
	Kimberley phasivirus	Parry’s Creek phasivirus 1 (PCPhasV1)
	Phasi Charoen-like phasivirus	Phasi Charoen-like virus (PCLV)
	Wutai mosquito phasivirus	Wǔtái mosquito virus (WtMV)
Phlebovirus	Adana phlebovirus	Adana virus (ADAV)
	Aguacate phlebovirus	Aguacate virus (AGUV)
	Alcube phlebovirus	Alcube virus (ACBV)
	Alenquer phlebovirus	Alenquer virus (ALEV)
	Ambe phlebovirus	Ambé virus (ABEV)
	Anhanga phlebovirus	Anhangá virus (ANHV)
	Arumowot phlebovirus	Arumowot virus (AMTV)
	Buenaventura phlebovirus	Buenaventura virus (BUEV)
	Bogoria phlebovirus	Bogoria virus (BGRV)
	Bujaru phlebovirus	Bujaru virus (BUJV)
	Cacao phlebovirus	Cacao virus (CACV)
	Campana phlebovirus	Campana virus (CMAV)
	Candiru phlebovirus	Ariquemes virus (ARQV)
		Candirú virus (CDUV)
		Jacundá virus (JCNV)
		Morumbi virus (MR(M)BV)
		Mucura virus (MCRV/MRAV)
		Serra Norte virus (SRNV)
	Chagres phlebovirus	Chagres virus (CHGV)
	Cocle phlebovirus	Coclé virus (CCLV)
	Corfou phlebovirus	Corfou virus (CFUV)
	Dashli phlebovirus	Dāshlī virus (DASV)
	Durania phlebovirus	Durania virus (DRNV)
	Echarate phlebovirus	Echarate virus (ECHV)
	Embossos phlebovirus	Embossos virus (EMBV)
	Gabek phlebovirus	Gabek forest virus (GFV)
	Gordil phlebovirus	Gordil virus (GORV)
	Icoaraci phlebovirus	Icoaraci virus (ICOV)
	Itaituba phlebovirus	Itaituba virus (ITAV)
	Itaporanga phlebovirus	Itaporanga virus (ITPV)
	Ixcanal phlebovirus	Ixcanal virus (IXCV)
	Karimabad phlebovirus	Karimabad virus (KARV)
	Kiborgoch phlebovirus	Kiborgoch virus (KBGV)
	La Gloria phlebovirus	La Gloria virus (LAGV)
	Lara phlebovirus	Rio Claro virus (RICV)
	Leticia phlebovirus	Leticia virus (LTCV)
	Maldonado phlebovirus	Maldonado virus (MLOV)
	Massilia phlebovirus	Massilia virus (MASV)
	Medjerda phlebovirus	Medjerda Valley virus (MVV)
	Mona Grita phlebovirus	Mona Grita virus (MOGV)
	Mukawa phlebovirus	Mukawa virus (MKWV)
	Munguba phlebovirus	Munguba virus (MUNV)
	Naples phlebovirus	Arrábida virus (ARRV)
		Balkan virus (BALKV)
		Fermo virus (FERV)
		Granada virus (GRV = GRAV)
		Saddaguia virus (SADV)
		sandfly fever Naples virus (SFNV)
	Nique phlebovirus	Nique virus (NIQV)
	Ntepes phlebovirus	Ntepes virus (NTPV)
	Odrenisrou phlebovirus	Odrénisrou virus (ODRV)
	Oriximina phlebovirus	Oriximiná virus (ORXV)
	Pena Blanca phlebovirus	Peña Blanca virus (PEBV)
	Penshurt phlebovirus	Penshurt virus (PEHV)
	Perkerra phlebovirus	Perkerra virus (PKEV)
	Punique phlebovirus	Punique virus (PUNV)
	Punta Toro phlebovirus	Capira virus (CAPIV)
		Punta Toro virus (PTV)
	Rift Valley fever phlebovirus	Rift Valley fever virus (RVFV)
	Rio Grande phlebovirus	Rio Grande virus (RGV)
	Saint Floris phlebovirus	Saint-Floris virus (SAFV)
	Salanga phlebovirus	Salanga virus (SLGV)
	Salehabad phlebovirus	Adria virus (ADRV)
		Arbia virus (ARBV)
		Bregalaka virus (BREV)
		Olbia virus (OLBV)
		Salehabad virus (SALV)
		Zaba virus (ZABAV)
	Salobo phlebovirus	Salobo virus (SLBOV)
	Sicilian phlebovirus	sandfly fever Sicilian virus (SFSV)
	Tapara phlebovirus	Tapará virus (TPRV)
	Tehran phlebovirus	Tehran virus (THEV)
	Tico phlebovirus	Tico virus (TICV)
	Toros phlebovirus	Toros virus (TORV)
	Toscana phlebovirus	Toscana virus (TOSV)
	Tres Almendras phlebovirus	Tres Almendras virus (TRAV)
	Turuna phlebovirus	Turuna virus (TUAV)
	Uriurana phlebovirus	Uriurana virus (URIV)
	Urucuri phlebovirus	Urucuri virus (URUV)
	Viola phlebovirus	viola virus (VIOV)
	Zerdali phlebovirus	Zerdali virus (ZERV)
Pidchovirus	Coleopteran pidchovirus	coleopteran phenui-related virus 308 (CoPrV-308)
	Pidgey pidchovirus	Pidgey virus (PGYV)
Rubodvirus	Apple rubodvirus 1	apple rubbery wood virus 1 (ARWV-1)
	Apple rubodvirus 2	apple rubbery wood virus 2 (ARWV-2)
	Grapevine rubodvirus 1	grapevine Garan dmak virus (GGDV)
	Grapevine rubodvirus 2	grapevine Muscat rose virus (GMRV)
Tanzavirus	Human tanzavirus	Dar es Salaam virus (DeSV)
Tenuivirus	Echinochloa hoja blanca tenuivirus	Echinochloa hoja blanca virus (EHBV)
	European wheat striate mosaic tenuivirus	European wheat striate mosaic virus (EWSMV)
	Iranian wheat stripe tenuivirus	Iranian wheat stripe virus (IWSV)
	Maize stripe tenuivirus	maize stripe virus (MStV = MSpV)
	Melon tenuivirus	melon chlorotic spot virus (MeCSV)
	Rice grassy stunt tenuivirus	rice grassy stunt virus (RGSV)
	Rice hoja blanca tenuivirus	rice hoja blanca virus (RHBV)
	Rice stripe tenuivirus	rice stripe virus (RSV = RStV)
	Urochloa hoja blanca tenuivirus	Urochloa hoja blanca virus (UHBV)
Uukuvirus	American dog uukuvirus	American dog tick virus (ADAV)
	Dabieshan uukuvirus	Dàbiéshān tick virus (DBSH)
	Grand Arbaud uukuvirus	Grand Arbaud virus (GAV)
	Huangpi uukuvirus	Huángpí tick virus 2 (HpTV-2)
	Kabuto mountain uukuvirus	Kabuto mountain virus (KAMV)
	Kaisodi uukuvirus	Kaisodi virus (KASDV)
	Lihan uukuvirus	Lǐhán tick virus (LITV)
	Murre uukuvirus	murre virus (MURV)
	Pacific coast uukuvirus	Pacific coast tick virus (PACV)
	Precarious Point uukuvirus	Precarious Point virus (PPV)
	Rukutama uukuvirus	Rukutama virus (RUKV)
	Schmid uukuvirus	Nile Warbler virus (NIWV)
	Silverwater uukuvirus	Silverwater virus (SILV)
	Tacheng uukuvirus	Tǎchéng tick virus 2 (TCGV)
	Uukuniemi uukuvirus	Chizé virus (CHZV)
		Fin V 707 virus (FINV)
		Oceanside virus (OCV = OCEV)
		Pontevès virus (PTVV)
		St. Abbs Head virus (SAHV)
		Uukuniemi virus (UUKV)
	Yongjia uukuvirus	Yǒngjiā tick virus (YONV)
	Zaliv Terpeniya uukuvirus	Zaliv Terpeniya virus (ZTV)
Wenrivirus	Shrimp wenrivirus	Mourilyan virus (MoV)
Family Tospoviridae
Orthotospovirus	Alstroemeria necrotic streak orthotospovirus	Alstroemeria necrotic streak virus (ANSV)
	Alstroemeria yellow spot orthotospovirus	Alstroemeria yellow spot virus (AYSV)
	Bean necrotic mosaic orthotospovirus	bean necrotic mosaic virus (BeNMV)
	Calla lily chlorotic spot orthotospovirus	calla lily chlorotic spot virus (CCSV)
	Capsicum chlorosis orthotospovirus	Capsicum chlorosis virus (CaCV)
	Chrysanthemum stem necrosis orthotospovirus	Chrysanthemum stem necrosis virus (CSNV)
	Groundnut bud necrosis orthotospovirus	groundnut bud necrosis virus (GBNV)
	Groundnut chlorotic fan spot orthotospovirus	groundnut chlorotic fan-spot virus (GCFSV)
	Groundnut ringspot orthotospovirus	groundnut ringspot virus (GRSV)
	Groundnut yellow spot orthotospovirus	groundnut yellow spot virus (GYSV)
	Hippeastrum chlorotic ringspot orthotospovirus	Hippeastrum chlorotic spot virus (HCRV)
	Impatiens necrotic spot orthotospovirus	impatiens necrotic spot virus (INSV)
	Iris yellow spot orthotospovirus	iris yellow spot virus (IYSV)
	Melon severe mosaic orthotospovirus	melon severe mosaic virus (MSMV)
	Melon yellow spot orthotospovirus	melon yellow spot virus (MYSV)
	Mulberry vein banding associated orthotospovirus	mulberry vein banding-associated virus (MVBaV)
	Pepper chlorotic spot orthotospovirus	pepper chlorotic spot virus (PCSV)
	Polygonum ringspot orthotospovirus	Polygonum ringspot virus (PolRSV)
	Soybean vein necrosis orthotospovirus	soybean vein necrosis virus (SVNV)
	Tomato chlorotic spot orthotospovirus	tomato chlorotic spot virus (TCSV)
	Tomato spotted wilt orthotospovirus	tomato spotted wilt virus (TSWV)
	Tomato yellow ring orthotospovirus	tomato yellow ring virus (TYRV)
	Tomato zonate spot orthotospovirus	tomato zonate spot virus (TZSV)
	Watermelon bud necrosis orthotospovirus	watermelon bud necrosis virus (WBNV)
	Watermelon silver mottle orthotospovirus	watermelon silver mottle virus (WSMoV)
	Zucchini lethal chlorosis orthotospovirus	zucchini lethal chlorosis virus (ZLCV)
Family Tulasviridae
Orthotulasvirus	Orthotulasvirus tulasnellae	Tulasnella bunyavirales-like virus 1 (TB-LV)
Family Wupedeviridae
Wumivirus	Wumivirus millepedae	Wǔhàn millipede virus 2 (WhMV-2)

Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, subfamilies, families, and orders are taxa.

^¶Taxon names are always italicized and always begin with a capital letter.

^&Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization. Lists of viruses within a given species are provisional at this point and will likely be amended in the near future.

Table 7.

ICTV-accepted taxonomy of the order Articulavirales (Negarnaviricota: Polyploviricotina: Insthoviricetes) as of March 2022.

Genus	Species¶	Virus (Abbreviation)&
Family Amnoonviridae
Tilapinevirus	Tilapinevirus tilapiae	tilapia lake virus (TiLV)
Family Orthomyxoviridae
Alphainfluenzavirus	Alphainfluenzavirus influenzae	influenza A virus (FLUAV)
Betainfluenzavirus	Betainfluenzavirus influenzae	influenza B virus (FLUBV)
Deltainfluenzavirus	Deltainfluenzavirus influenzae	influenza D virus (FLUDV)
Gammainfluenzavirus	Gammainfluenzavirus influenzae	influenza C virus (FLUCV)
Isavirus	Isavirus salaris	infectious salmon anemia virus (ISAV)
Mykissvirus	Mykissvirus tructae	rainbow trout orthomyxovirus (RbtOV)
		steelhead trout orthomyxovirus (SttOV-1)
Quaranjavirus	Quaranjavirus araguariense	Araguari virus (ARAV)
	Quaranjavirus chadense	Lake Chad virus (LKCV)
	Quaranjavirus johnstonense	Johnston Atoll virus (JAV)
	Quaranjavirus quaranfilense	Quaranfil virus (QRFV)
	Quaranjavirus tyulekense	Tyulek (Tjuloc) virus (TLKV)
	Quaranjavirus wellfleetense	Wellfleet Bay virus (WFBV)
Sardinovirus	Sardinovirus pilchardi	pilchard orthomyxovirus (POMV)
Thogotovirus	Thogotovirus bourbonense	Bourbon virus (BRBV)
	Thogotovirus dhoriense	Dhori virus (DHOV)
	Thogotovirus josense	Jos virus (JOSV)
	Thogotovirus ozense	Oz virus (OZV)
	Thogotovirus sinuense	Sinu virus (SINUV)
	Thogotovirus thailandense	Thailand tick thogotovirus (TT-THOV)
	Thogotovirus thogotoense	Thogoto virus (THOV)
	Thogotovirus upoluense	Upolu virus (UPOV)

Note that viruses are real objects that are assigned to concepts that are called taxa. Species, genera, families, and orders are taxa.

^¶Taxon names are always italicized and always begin with a capital letter.

^&Virus names are not italicized and are not capitalized, except if the name or a name component is a proper noun. This column lists the virus names with their correct (lack of) capitalization.