Abstract
The type specimens of the genus Andrena Fabricius, 1775, described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko deposited in the Zoological Museum of the Moscow State University and in the Zoological Institute, Russian Academy of Sciences, St. Petersburg (Russia), are critically reviewed. Precise information with illustrations of types for 52 taxa is provided; of these 39 species are valid and thirteen are invalid (ten synonyms and three homonyms). Lectotypes are here designated for the following 24 nominal taxa: Andrenaacutilabris Morawitz, 1876, A.aulica Morawitz, 1876, A.bicarinata Morawitz, 1876, A.combusta Morawitz, 1876, A.comparata Morawitz, 1876, A.corallina Morawitz, 1876, A.discophora Morawitz, 1876, A.fedtschenkoi Morawitz, 1876, A.initialis Morawitz, 1876, A.laeviventris Morawitz, 1876, A.leucorhina Morawitz, 1876, A.mucorea Morawitz, 1876, A.nitidicollis Morawitz, 1876, A.oralis Morawitz, 1876, A.planirostris Morawitz, 1876, A.ravicollis Morawitz, 1876, A.rufilabris Morawitz, 1876, A.sarta Morawitz, 1876, A.smaragdina Morawitz, 1876, A.sogdiana Morawitz, 1876, A.subaenescens Morawitz, 1876, A.tuberculiventris Morawitz, 1876, A.turkestanica Morawitz, 1876, and A.virescens Morawitz, 1876.
Keywords: Anthophila, Apiformes, lectotypes, Palaearctic Region, taxonomy
Introduction
The present paper is the second part of a series of works dealing with the bee taxa described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko deposited in the Zoological Museum of the Moscow State University, Moscow (ZMMU) and in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg (ZISP). In the first part, the family Halictidae was considered (Astafurova and Proshchalykin 2020). The main goal of this work is to make Fedtschenko’s collection of bees more accessible and useful to scientists.
More than 140 years ago (1876), the second part of Ferdinand Morawitz’s critical study on the bees collected by Aleksey Fedtschenko 1869–1871 Expeditions in “Turkestan” was published. In the first volume, “Apidae genuinae” (Morawitz 1875), Morawitz treated a total of 255 species of numerous genera, of which many species were described as new. In this second part, “Andrenidae” (Morawitz 1876), the remaining bees were dealt with, including the species of the difficult genera Andrena Fabricius, Halictus Latreille, and Hylaeus Fabricius, totalling 183 species (Pesenko and Astafurova 2003). The species treatments are of a high professional standard, the localities are precisely documented (A. Fedtschenko 1871; O. Fedtschenko 1874; Baker 2004; Kuhlmann 2005; Dathe and Proshchalykin 2017) and the type series have been carefully conserved over a long period, generally in the collections of the ZMMU and ZISP. To this day, these remain some of the most important manuscripts on bees of Central Asia.
Andrena Fabricius, 1775 is the one of the largest bee genera, numbering about 1550 species worldwide, of which about 950 are known from the Palaearctic region (Gusenleitner and Schwarz 2002; Ascher and Pickering 2022). The genus is distributed throughout the Holarctic region, south in the Western Hemisphere to Panama, in Africa through the East African highlands and south to the Cape of Good Hope, and in Asia to the mountains of southern India and of Malaysia (Michener 2007). Only three species of Andrena are distributed in both the Palearctic and the Holarctic regions, whereas the others occur in only one of these zoogeographic realms. Many species appear to be widespread, and many are bivoltine, which has led to separate descriptions in different regions, especially in the Palearctic. Thus, about 3000 species descriptions were recognised at the beginning of 21st century (Gusenleitner and Schwarz 2002).
The genus Andrena is represented in Morawitz’s 1876 publication by 68 species (Nos. 258–326). Only 16 species were previously known, while the remaining 52 were newly described (Table 1). Few of these taxa have been mentioned in subsequent publications, remaining enigmatic for decades.
Table 1.
The nominal taxa of Fedtschenko’s Andrena, described by F. Morawitz, and their current status.
| N | Species name | Sex | Current status | Depositaries of types |
|---|---|---|---|---|
| 1. | Andrenaacutilabris | ♀, ♂ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 2. | Andrenaahenea | ♀, ♂ | Valid | LT, PLT (ZMMU) |
| 3. | Andrenaamoena | ♀ | Valid | LT, PLT (ZMMU) |
| 4. | Andrenaarenaria | ♂ | Synonym | HT (ZMMU) |
| 5. | Andrenaaulica | ♀, ♂ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 6. | Andrenabairacumensis | ♀ | Valid | HT (ZMMU) |
| 7. | Andrenabicarinata | ♀ | Synonym | LT (ZISP); PLT (ZISP/ZMMU) |
| 8. | Andrenacapillosa | ♀ | Valid | LT (ZMMU); PLT (ZISP) |
| 9. | Andrenacarinifrons | ♀ | Valid | LT (ZISP) |
| 10. | Andrenacombusta | ♀, ♂ | Valid | LT, PLT (ZMMU) |
| 11. | Andrenacomparata | ♂ | Synonym | LT (ZISP); PLT (ZISP/ZMMU) |
| 12. | Andrenacorallina | ♀ | Synonym | LT (ZMMU) |
| 13. | Andrenadiscophora | ♀, ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 14. | Andrenafedtschenkoi | ♀, ♂ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 15. | Andrenaferghanica | ♀ | Valid | HT (ZISP) |
| 16. | Andrenaflavitarsis | ♀, ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 17. | Andrenafuscicollis | ♀, ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 18. | Andrenahieroglyphica | ♀ | Valid | HT (ZMMU) |
| 19. | Andrenainfirma | ♀, ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 20. | Andrenainitialis | ♀, ♂ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 21. | Andrenalaeviventris | ♀ | Valid | LT, PLT (ZMMU) |
| 22. | Andrenalateralis | ♀, ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 23. | Andrenaleucorhina | ♂ | Synonym | LT (ZMMU); PLT (ZISP/ZMMU) |
| 24. | Andrenalucidicollis | ♂ | Valid | ST (ZMMU) |
| 25. | Andrenamaculipes | ♀, ♂ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 26. | Andrenamajalis | ♀ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 27. | Andrenamordax | ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 28. | Andrenamucorea | ♀, ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 29. | Andrenanigrita | ♂ | Homonym | HT (ZMMU) |
| 30. | Andrenanitidicollis | ♀, ♂ | Valid | LT, PLT (ZMMU) |
| 31. | Andrenanupta | ♀, ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 32. | Andrenaoralis | ♀ | Valid | LT (ZISP); PLT (ZMMU) |
| 33. | Andrenapannosa | ♀ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 34. | Andrenaplanirostris | ♀, ♂ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 35. | Andrenapunctifrons | ♀ | Valid | HT (ZMMU) |
| 36. | Andrenapunctiventris | ♀ | Valid | HT (ZMMU) |
| 37. | Andrenaquadrifasciata | ♀ | Valid | HT (ZMMU) |
| 38. | Andrenaravicollis | ♀ | Synonym | LT (ZISP); PLT (ZMMU) |
| 39. | Andrenarufilabris | ♀ | Synonym | LT, PLT (ZMMU) |
| 40. | Andrenarufina | ♀, ♂ | Valid | LT, PLT (ZMMU) |
| 41. | Andrenasarta | ♀ | Valid | LT (ZMMU) |
| 42. | Andrenasemiaenea | ♀ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 43. | Andrenasmaragdina | ♀, ♂ | Valid | LT (ZISP); PLT (ZMMU) |
| 44. | Andrenasogdiana | ♂ | Synonym | LT (ZISP) |
| 45. | Andrenasordida | ♀, ♂ | Homonym | LT (ZMMU); PLT (ZISP/ZMMU) |
| 46. | Andrenasubaenescens | ♀, ♂ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 47. | Andrenatemporalis | ♂ | Homonym | LT (ZMMU); PLT (ZISP/ZMMU) |
| 48. | Andrenatuberculiventris | ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
| 49. | Andrenaturkestanica | ♀, ♂ | Synonym | LT (ZISP); PLT (ZMMU) |
| 50. | Andrenaurmitana | ♂ | Synonym | HT (ZMMU) |
| 51. | Andrenavirescens | ♀, ♂ | Valid | LT (ZISP); PLT (ZISP/ZMMU) |
| 52. | Andrenaviridigastra | ♀, ♂ | Valid | LT (ZMMU); PLT (ZISP/ZMMU) |
LT – lectotype, PLT – paralectotype/s, ST – syntype.
Since the 1940s, V. Popov [ZISP] based his studies on the taxonomy and ecology of the Central Asian Andrena fauna using these collections (Popov 1940, 1947, 1949, 1952, 1958, 1960).
Type material of Fedtschenko’s Andrena was studied by K. Warncke, a teacher in Dachau (Germany), who visited the ZMMU from 26.03.1975 to 01.04.1975 (Dathe and Proshchalykin 2017). He worked his way through the drawers containing the Fedtschenko material and labelled specimens of all nominal taxa found there as “Lectotypus.” Thereby, certain specimens from some groups of bees (Hylaeus, Andrena, Anthidium, Halictus, etc.) received a red label with the inscription “Lectotypus Warncke 1975.” All other type specimens were left without nomenclatural status labels. However, unlike other groups of bees (Warncke 1981, 1982), he did not publish information about the designation of Andrena lectotypes.
In 1980 A.Z. Osytshnjuk continued the study of Andrena in the Fedtschenko’s collection of the ZISP and ZMMU. The lectotype and paralectotype designations by A. Osytshnjuk were largely unpublished, because of her tragic death in 1998. Some of these designations have been posthumously validated in subsequent publications (Osytshnjuk et al. 2005, 2008).
The present paper is the first complete, critically reviewed, illustrated summary of all species of the genus Andrena described by F. Morawitz from the collection of A. Fedtschenko, an invaluable reference for researchers across this region who otherwise could not easily assign names to these difficult bees.
Materials and methods
All of the material listed below was examined for this study. In the following list, the taxa are treated in alphabetical order of the names used in the original descriptions. Each entry includes the name of the taxon in its original combination, the complete reference to the original description of the species (including the original combination and spelling of the name and the author, year, and page of the description) and a list of type specimens present in the collections of the Zoological Museum of the Moscow State University, Moscow (ZMMU) and Zoological Institute of the Russian Academy of Sciences, St. Petersburg (ZISP). The data from each label are separated by two slashes (//). Square brackets are used for English translations and when information is added to specimen label information (e.g., geographical coordinates) or published data (e.g., current name of an old place name; affiliation to a present-day country).
Photographs were made using a combination of a stereomicroscope Olympus SZX10 and a digital camera (Olympus OM-D). Illustrations were obtained by montaging from an image series that covers different focal planes into a single in-focus image with the program Helicon Focus 7. The final illustrations were post-processed for contrast and brightness using Adobe Photoshop software.
The classification and current species status mostly follow Gusenleitner and Schwarz (2002) and Pisanty et al. (2022), distribution follows Osytshnjuk et al. (2005, 2008), Proshchalykin et al. (2017), Wood and Monfared (2022), and Ascher and Pickering (2022). In the dating the publications of Morawitz’s paper, we follow Kerzhner (1984) and Ebmer (2021).
Taxonomy
List of species
Genus Andrena Fabricius, 1775
1. Andrena acutilabris
Morawitz, 1876
9B7E5D93-5C81-551C-BEC4-7012E1FA0BC7
Figure 1.
Andrenaacutilabris Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T4, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena acutilabris Morawitz, 1876: 165 (key), 175, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent, Urmitan.
Lectotype (designated here).
♀, Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // acutilabris [handwritten by F. Morawitz] // Paralectotypus Andr.acutilabris Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenaacutilabris Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000300 [ZISP].
Paralectotypes
(3 ♀, 10 ♂). 2 ♀, 4 ♂, 2, 3, 8, 11[IV.1871] // Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // acutilabris Mor. [handwritten by F. Morawitz] // Paralectotypus Andrenaacutilabris Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 6 ♂, 2., 5.[IV.1871] // Ташкентъ; 1 ♀, 11.[IV.1871] // Ташкентъ // Lectotypus Warncke 1975 // Paralectotypus Andrenaacutilabris Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Nobandrena) acutilabris Morawitz, 1876.
Distribution.
Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan.
2. Andrena ahenea
Morawitz, 1876
4EB0DE5A-2A36-5D91-BFE3-B0E72FF30969
Figure 2.
Andrenaahenea Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena ahenea Morawitz, 1876: 164, 166 (key), 210, ♀, ♂.
Type locality.
Samarkand (Uzbekistan).
Published (original) locality.
Uzbekistan: Samarkand.
Lectotype.
♀, designated by Osychnjuk et al. 2008: 51, 16.[III.1869] // Самаркандъ [Uzbekistan, Samarkand, 39°39'N, 66°57'E] // Andrenaahenea Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenaahenea Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotype.
1 ♂, 16.[III.1876] // Самаркандъ // Paralectotypus Andrenaahenea Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu.Astafurova> [ZMMU].
Current status.
Andrena (Euandrena) ahenea Morawitz, 1876.
Distribution.
Uzbekistan.
3. Andrena amoena
Morawitz, 1876
C47DE2A4-D7F6-5EA9-AF77-05732ADA4358
Figure 3.
Andrenaamoena Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena amoena Morawitz, 1876: 164 (key), 211, ♀.
Type locality.
Chardara (Kazakhstan).
Published (original) locality.
Kazakhstan: Syr Darja River near Chardara.
Lectotype.
♀, designated by Osytshnjuk et al. 2005: 67, 25.[IV.1871] // Чардара [Kazakhstan, Chardara, 41°18'N, 67°57'E] // Andrenaamoena Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenaamoena Morawitz, 1876, design. Osytshnjuk et al., 2005 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(3 ♀). 3 ♀, 25.[IV.1871] // Чардара [Chardara] // Paralectotypus Andrenaamoena Morawitz, 1876, design. Osytshnjuk et al., 2005 <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Aciandrena) amoena Morawitz, 1876 (according to Osytshnjuk et al. 2005: 67).
Remarks.
Description of male: Gusenleitner and Schwarz 2001: 99, fig. 1.
Distribution.
Turkmenistan, Uzbekistan, Tajikistan.
4. Andrena arenaria
Morawitz, 1876
2F76774E-CF43-5C40-B7A3-D8059933E633
Figure 4.
Andrenaarenaria Morawitz, 1876, holotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena arenaria Morawitz, 1876: 201, ♂.
Type locality.
Central Asia.
Published (original) locality.
“Turkestan”.
Holotype.
♂, 487. // Туркест.[анский] кр.[ай] [Turkestan] // Andrenaarenaria Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Holotypus Andrenaarenaria Morawitz, 1876 <red label, labelled by Yu. Astafurova> // Andrenalateralis, D.A. Sidorov det. 2022 [ZMMU].
Current status.
Andrena (incertae sedis) lateralis Morawitz, 1876 (synonymised by Gusenleitner and Schwarz 2001: 135).
Distribution.
Europe, Russia (to East Siberia), Caucasus, Turkey, Israel, Iran, Afghanistan, Central Asia.
5. Andrena aulica
Morawitz, 1876
FBBADC3D-7CF1-5500-B7F1-E508B1E2BC1B
Figure 5.
Andrenaaulica Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T4, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena aulica Morawitz, 1876: 162, 166 (key), 187, ♀, ♂.
Type locality.
Panjakent (Tajikistan).
Published (original) locality.
Uzbekistan: Tashkent, Samarkand, Oalyk Gorge; Tajikistan: Panjakent, Fan [River], Varzaminor [= Ayni], Iori Gorge.
Lectotype (designated here).
♀, Пянджикентъ [Tajikistan, Panjakent, 39°30'N, 67°36'E] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenaaulica Morawitz, ♀ [handwritten by F. Morawitz] // Paralectotypus Andr.aulica F. Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenaaulica Morawitz, 1876, design. Astafurova et al., 2022 // Zoological Institute St. Petersburg INS_HYM_0000299 [ZISP].
Paralectotypes
(21 ♀, 3 ♂). 1 ♂, <golden circle> // Варзаминоръ // aulica Mor. Typ. ♂ [handwritten by F. Morawitz] // Andr.ferghanica F. Mor, ♂, Popov det, 1936; 1 ♀, Заравш. дол. // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // aulica Mor. [handwritten by F. Morawitz]; Paralectotypus Andrenaaulica Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 4 ♀, 8.[IV.1871] // Ташкентъ; 9 ♀, 20., 21., 23., 26., 30.[III], 9.[V.1869] // Самаркандъ; 3 ♀, 28., 29.[IV.1869] // Пянджикентъ; 1 ♀, 12.[VI.1870] // Фанъ; 2 ♀, 7.[VI.1870] // Варзаминоръ; 1 ♀, 2 ♂, 25.[III], 2., 7.[V.1869] // Заравш. дол. // Paralectotypus Andrenaaulica Morawitz, 1876, design. Astafurova et. al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
There is one female specimen labelled by Warcke as “Lectotype” [31. // Заравш. дол. // Andrenaaulica Mor. Typ. [handwritten by F. Morawitz] // Lectotypus Warncke 1975; ZMMU]. However, the label date (31.[II.1869]) does not correspond any date mentioned for type series by Morawitz (1876).
Current status.
Andrena (Plastandrena) aulica Morawitz, 1876.
Remarks.
According to Warncke (1967: 179) and Gusenleitner and Schwarz (2002: 130) A.aulica Morawitz, 1876 is a junior synonym of A.bimaculata (Kirby, 1802). However, Popov (1949), Osytshnjuk et al. (1978) and Astafurova et al. (2021) regarded A.aulica as a valid species. Taxonomic status of A.bimaculata sensu lato is problematic and requires a revision.
Distribution.
Russia (European part), Caucasus, Iran, Central Asia, India.
6. Andrena bairacumensis
Morawitz, 1876
C7F2DBE7-1D67-52CA-B700-AD8ACAE36A69
Figure 6.
Andrenabairacumensis Morawitz, 1876, holotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena bairacumensis Morawitz, 1876: 164 (key), 170, ♀.
Type locality.
Bairkum (Kazakhstan).
Published (original) locality.
Kazakhstan: Bayrakum [= Bairkum].
Holotype.
♀, 4.[V.1871] // Байракумъ [Kazakhstan, Syr-Darya River, Bairkum, 42°05'N, 68°10'E] // Andrenabairacumensis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Holotypus Andrenabairacumensis Mor., 1876 <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Leimelissa) bairacumensis Morawitz, 1876.
Remarks.
Description of male: Osytshnjuk 1984: 21, fig. 1.
Distribution.
Kazakhstan.
7. Andrena bicarinata
Morawitz, 1876
90CFDA06-597E-5519-A15B-62EBD3F0EC04
Figure 7.
Andrenabicarinata Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T5, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena bicarinata Morawitz, 1876: 164 (key), 197, ♀.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent, Katty-Kurgan, Urmitan [near Katty-Kurgan]; Tajikistan: Panjakent.
Lectotype (designated here).
♀, 11.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenabicarinata Mor. [handwritten by F. Morawitz] // Paralectotypus Andrenabicarinata Mor., design. Osychnjuk, 1980 <red label> // Andrenatuberculiventris Mor. A. Osytshnjuk det. // Lectotypus Andrenabicarinata Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000298 [ZISP].
Paralectotypes
(5 ♀, 1 ♂). 1 ♀, 2.[IV.1871] // Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenabicarinata Mor. [handwritten by F. Morawitz] // Paralectotypus Andrenabicarinata Mor., design. Osychnjuk, 1980 < red label > // Andrenatuberculiventris, D.A. Sidorov det.; 1 ♀, 5.[IV.1871] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenabicarinata F.Mor, Cotype!, F. Morawitz det. // Andrenatuberculiventris, D.A. Sidorov det. [ZISP]; 1 ♀, 5.[V.1871] // Урмитанъ [Urmitan] // Lectotypus Warncke 1975 <red label> // A.tuberculiventris [det. A.Osytshnjuk]; 1 ♂, 5.[IV.1871] // Ташкентъ [Tashkent] // Andrenatuberculiventris, D.A. Sidorov det.; 1 ♀, 29.[VI.1869] // Katty-Kurgan // Andrenatuberculiventris, D.A. Sidorov det.; 1 ♀, 2.[V.1871] // Урмитанъ [Urmitan] // Andrenatuberculiventris, D.A. Sidorov det. // Paralectotypus Andrenabicarinata Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Parandrenella) tuberculiventris Morawitz, 1876 (synonymised by Popov 1958: 118).
Distribution.
Uzbekistan, Tajikistan.
8. Andrena capillosa
Morawitz, 1876
76D969E1-5112-5B2B-92E5-C70764C11795
Figure 8.
Andrenacapillosa Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena capillosa Morawitz, 1876: 163 (key), 205, ♀.
Type locality.
Samarkand (Uzbekistan).
Published (original) locality.
Uzbekistan: Samarkand.
Lectotype.
♀, designated by Osytshnjuk et al. 2008: 49, ♀, 23.[II.1869] // Самаркандъ [Uzbekistan, Samarkand, 39°39'N, 66°57'E] // Andrenacapillosa Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 // Lectotypus Andrenacapillosa Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(5 ♀). 1 ♀, <golden circle>, 27.[II.1869] // Самаркандъ // capillosa Mor. Typ. [handwritten by F. Morawitz]; 4 ♀, 27.[II.1869], 3., 4.[III.1869] // Самаркандъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andrenacapillosa Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP].
Current status.
Andrena (Euandrena) capillosa Morawitz, 1876.
Remarks.
Description of male: Osytshnjuk et al. 2008: 50.
Distribution.
Uzbekistan, Tajikistan, Kazakhstan. The record from Russia (Western Siberia) by Krainov (2013) is doubtful.
9. Andrena carinifrons
Morawitz, 1876
71B337F1-1FD1-5B2B-A154-0B57EB5FE4C0
Figure 9.
Andrenacarinifrons Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1-T4, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena carinifrons Morawitz, 1876: 164 (key), 198, ♀.
Type locality.
Turkistan Province (Kazakhstan).
Published (original) locality.
Kazakhstan: between the Karak Mts and Bairkum.
Lectotype.
♀, designated by Osytshnjuk et al. 2005: 161, Туркест.[анский] кр.[ай] [Kazakhstan, Turkistan Province, ≈ 42°46'N, 67°24'E] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenacarinifrons Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenacarinifrons Morawitz, 1876 design. by Osytchjuk et al., 2005 <red label, labelled by Yu. Astafurova> // Zoological Institute St. Petersburg INS_HYM_0000200.
Current status.
Andrena (Carinandrena) carinifrons Morawitz, 1876.
Remarks.
Description of male: Osytshnjuk 1993: 18, figs 2, 3.
Distribution.
Turkmenistan, Uzbekistan, Kazakhstan.
10. Andrena combusta
Morawitz, 1876
6B61927A-F633-5629-AF93-B179FBF73B45
Figure 10.
Andrenacombusta Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena combusta Morawitz, 1876: 163, 165 (key), 189, ♀, ♂.
Type locality.
Kattakurgan (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent and near Kattakurgan.
Lectotype (designated here).
♀, 29. [IV.1869] // Верхн.[ий] Заравш.[ан] [Uzbekistan, Upper Zaravshan, near Kattakurgan, 39°53'N, 66°15'E] // Andrenacombusta Mor. [handwritten by F. Morawitz] // Lectotypus Warncke, 1975 <red label> // Lectotypus Andrenacombusta Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Paralectotypes
(1 ♀, 1 ♂). 1 ♀, 29. [IV.1869] // Верхн. Заравш. [Upper Zaravshan]; 1 ♂, 11.[IV.1871] // Ташкентъ // Paralectotypus Andrenacombusta Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Truncandrena) combusta Morawitz, 1876.
Distribution.
Azerbaijan, Turkey, Syria, Iran, Afghanistan, Uzbekistan, Tajikistan.
11. Andrena comparata
Morawitz, 1876
172B32CF-0E9E-56B0-BEA5-69B9EF64D435
Figure 11.
Andrenacomparata Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena comparata Morawitz, 1876: 166 (key), 188, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent and Samarkand.
Lectotype (designated here).
♂, 9.[III.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenacomparata Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andr.aulica F. Mor, ♂, Popov 1936 det. // Paralectotypus Andr.comparata Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenacomparata Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000297 [ZISP].
Paralectotypes
(9 ♂). 1 ♂, <golden circle>, 21.[III.1869] // Самаркандъ [Samarkand] // Andrenacomparata Mor. Typ. [handwritten by F. Morawitz] // Andr.aulica F.Mor, ♂, Popov det. 1936 // Paralectotypus Andrenacomparata Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 1 ♂, 21.[III.1869] // Самаркандъ // Andrenacomparata Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Andrenaaulica, D.A. Sidorov det., 2022; 7 ♂, 21., 26., 30.[III.1869] // Самаркандъ // Paralectotypus Andrenacomparata Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Plastandrena) aulica Morawitz, 1876 (synonymised by Popov 1949: 392).
Remarks.
Listed as Andrena (Plastandrena) bimaculata (Kirby, 1802) by Gusenleitner and Schwarz 2002: 130 (see remarks on Andrenaaulica, above).
Distribution.
Russia (European part), Caucasus, Iran, Central Asia, India.
12. Andrena corallina
Morawitz, 1876
50D305FD-02E6-5309-B6E0-03759C1DE0F6
Figure 12.
Andrenacorallina Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena corallina Morawitz, 1876: 162 (key), 203, ♀.
Type locality.
Urmetan (Tajikistan).
Published (original) locality.
Tajikistan: Urmitan.
Lectotype (designated here).
♀, 2.[V.1869] // Урмитанъ [Tajikistan, Urmetan, 39°26'N, 68°15'E] // Andrenacorallina Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 // Lectotypus Andrenacorallina Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Current status.
Andrena (Trachandrena) discophora Morawitz, 1876 (synonymised by Popov 1958: 159).
Distribution.
Uzbekistan, Tajikistan, Kazakhstan.
13. Andrena discophora
Morawitz, 1876
2A1A6A11-E2A5-5BE0-ACE5-8E5EECBF9249
Figure 13.
Andrenadiscophora Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena discophora Morawitz, 1876: 162, 166 (key), 202, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent.
Lectotype (designated here).
♀, 8.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenadiscophora Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenadiscophora Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Paralectotypes
(21 ♀, 9 ♂). 1 ♂, <golden circle> // 28.[II.1871] // Ташкентъ // discophora Mor. ♂, Typ. [handwritten by F. Morawitz]; 1 ♀, 4 ♂, Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenadiscophora Mor. [handwritten by F. Morawitz]; 1 ♀, <golden circle> // 11.[IV.1871] // Ташкентъ // discophora Mor. ♀, Typ. [handwritten by F. Morawitz]; 2 ♀, 8.[IV.1871] and 27.[III.1987] // Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♀, 11.[IV.1981] // Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andr.discophora F. Mor., Popov det. // Paralectotypus Andrena Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 16 ♀, 4 ♂, [28.II–11.IV.1871] // Ташкентъ // Paralectotypus Andrenadiscophora Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Trachandrena) discophora Morawitz, 1876.
Distribution.
Uzbekistan, Tajikistan, Kazakhstan.
14. Andrena fedtschenkoi
Morawitz, 1876
507C9637-6F8D-5654-8D5F-0E328CC6C9BC
Figure 14.
Andrenafedtschenkoi Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena Fedtschenkoi Morawitz, 1876: 162, 165 (key), 184, ♀, ♂.
Type locality.
Chardara (Kazakhstan).
Published (original) locality.
Kazakhstan: Kysyl-Kum [desert] near draw-well Chakany, Karak steppe, Chardara; Uzbekistan: Zeravshan River valley.
Lectotype (designated here).
♀, Чардара [Kazakhstan, Chardara, 41°18'N, 67°57'E] // AndrenaFedtschenkoi Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andrenafedtschenkoi Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenafedtschenkoi Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000296.
Paralectotypes
(7 ♀, 11 ♂). 1 ♀, Кизилъкум [Kyzylkum] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // fedtschenkoi F. Morawitz, ♀ [handwritten by F. Morawitz]; 3 ♂, Чардара // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // AndrenaFedtschenkoi Mor. [handwritten by F. Morawitz]; 1 ♂, 25.[IV.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley] // fedtschenkoi F. Morawitz, ♂ Typ. [handwritten by F. Morawitz] // Paralectotypus Andrenafedtschenkoi Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 1 ♀, 28.[IV.1871] // Кизилъкум // AndrenaFedtschenkoi Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label>; 4 ♀, 4 ♂, 25.[IV.1869], 12., 13., 18., 23.[V.1869] // Заравш. дол.; 3 ♂, 25.[IV.1871] // Чардара; 1 ♀, 5.[V.1871] // Карак.[ская] степь [Karak steppe] // Paralectotypus Andrenafedtschenkoi Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Ulandrena) fedtschenkoi Morawitz, 1876.
Distribution.
Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan.
15. Andrena ferghanica
Morawitz, 1876
09C1F838-36F7-50E1-B769-D69BF110A5D9
Figure 15.
Andrenaferghanica Morawitz, 1876, holotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T4, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena ferghanica Morawitz, 1876: 163 (key), 189, ♀.
Type locality.
Alai Mts, Kavuk Pass (Kyrgyzstan).
Published (original) locality.
Kyrgyzstan: Kavuk Pass.
Holotype.
♀, 24.[VI.1871] // Алай [Kyrgyzstan, Alay Mts, Kavuk Pass, 39°40'N, 72°15'E] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenaferghanica Morawitz [handwritten by F. Morawitz] // Lectotypus Andrenaferghanica Mor., design. Osychnjuk, 1980 <red label> // Holotypus Andrenaferghanica Mor., 1876 <red label, labelled by Yu. Astafurova> // Zoological Institute St. Petersburg INS_HYM_0000261 [ZISP].
Current status.
Andrena (Plastandrena) ferghanica Morawitz, 1876.
Remarks.
Description of male: Morawitz 1876: 187 as A.aulica, in partim (according to Popov 1949: 391).
Distribution.
Iran, Pakistan, Central Asia.
16. Andrena flavitarsis
Morawitz, 1876
BC8174E3-1DF8-5368-A470-59979B36B891
Figure 16.
Andrenaflavitarsis Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena flavitarsis Morawitz, 1876: 163, 166 (key), 204, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent.
Lectotype.
♀, designated by Osytshnjuk et al. 2008: 47, 3.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenaflavitarsis Mor. ♀ [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenaflavitarsis Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(1 ♀, 1 ♂). 1 ♀, 5.[IV.1871] // Ташкентъ [Tashkent] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenaflavitarsis Mor. [handwritten by F. Morawitz] // Andrenaflavitarsis Mor., design. Osychnjuk, 1980 < red label > [ZISP]; 1 ♂, 16.[III.1871] // Ташкентъ [Tashkent] // Andrenaflavitarsis Mor. [handwritten by F. Morawitz] // Paralectotypus Andrenaflavitarsis Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Euandrena) flavitarsis Morawitz, 1876.
Distribution.
Uzbekistan.
17. Andrena fuscicollis
Morawitz, 1876
7919437C-157C-5B68-91C2-DB45AB668844
Figure 17.
Andrenafuscicollis Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena fuscicollis Morawitz, 1876: 164, 165 (key), 208, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent.
Lectotype.
♂, designated by Osytshnjuk et al. 2008: 73, 11.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenafuscicollis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenafuscicollis Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(5 ♀, 5 ♂). 1 ♂, <golden circle>, 1.[IV.1871] // Ташкентъ // fuscicollis Mor. ♂ [handwritten by F. Morawitz]; 4 ♂, 5., 10.[IV.1871] // Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♀, <golden circle>, 10.[IV.1871] // Ташкентъ // fuscicollis Mor. Typ., ♀ [handwritten by F. Morawitz] // Paralectotypus Andrenafuscicollis Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 4 ♀, 1.,10., 11.[IV.1871] Ташкентъ // Paralectotypus Andrenafuscicollis Morawitz, 1876, design. Osytshnjuk et al., 2008 <identical red labels on each paralectotype specimen, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Fuscandrena) fuscicollis Morawitz, 1876.
Remarks.
The collection date in the original publication is “from 11 February to 11 March”. However, the lectotype and paralectotype specimens designated by A. Osytshnjuk are labelled as collected in April [green label]. Probably “March” was mistakenly mentioned in the Morawitz’ publication instead of April.
Distribution.
Turkmenistan, Uzbekistan, Tajikistan.
18. Andrena hieroglyphica
Morawitz, 1876
C32A70DA-D344-5CB0-8FDB-BF59457419E3
Figure 18.
Andrenahieroglyphica Morawitz, 1876, holotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena hieroglyphica Morawitz, 1876: 163 (key), 192, ♀.
Type locality.
Sokh Enclave (Uzbekistan).
Published (original) locality.
Uzbekistan: near Sokh.
Holotype.
♀, 28.[VI.1871] // Сохъ [Uzbekistan, Sokh Enclave, 39°57'N, 71°07'E] // Andrenahieroglyphica Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Holotypus Andrenahieroglyphica Mor., 1876 <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (incertae sedis) hieroglyphica Morawitz, 1876.
Remarks.
Description of male: Morawitz 1876: 204, as Andrenatemporalis Morawitz, 1876 (synonymised by Osytshnjuk 1984: 3).
The lectotype designation by Osytshnjuk et al. (2005: 151) is unnecessary as the species was described from a single male that was directly written about by Morawitz (1876: 192).
Distribution.
Iran, Turkmenistan, Uzbekistan, Tajikistan, Pakistan.
19. Andrena infirma
Morawitz, 1876
19934B0E-CBF8-514A-9109-668CCF477E38
Figure 19.
Andrenainfirma Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena infirma Morawitz, 1876: 164, 166 (key), 195, ♀, ♂.
Type locality.
Ayni (Tajikistan).
Published (original) locality.
Tajikistan: Varzaminor, near Iskander-Kul Lake; Kyrgyzstan/Tajikistan: Khodzha-Chiburgan River.
Lectotype.
♂, designated by by Osytshnjuk et al. 2008: 199, 8.[VI.1869] // Варзаминоръ [Tajikistan, Varzaminor (= Ayni), 39°23'N, 68°32'E] // Andrenainfirma Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenainfirma Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(1 ♀, 4 ♂). 1 ♂, 8.[VI.1870] // Варзаминоръ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♂, <golden circle> // 21.[VI.1870] // Искандеръ [Iskander] // infirma Mor. Typ. [handwritten by F. Morawitz]; 1 ♂, 21.[VI.1870] // Искандеръ // Andrenainfirma Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andrenainfirma Mor., design. Osytshnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 1 ♀ [without head], 9.[VI.1869] // Варзаминоръ // Федченко [Fedtschenko leg.]; 1 ♂, 26.[VI.1871] // Чибурганъ [Khodzha-Chiburgan River] // Paralectotypus Andrenainfirma Morawitz, 1876., design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Melandrena) infirma Morawitz, 1876.
Remarks.
Osytshnjuk et al. (2008: 199) mistakenly mentioned sex of the lectotype as a female instead of male.
Distribution.
Afghanistan, Central Asia.
20. Andrena initialis
Morawitz, 1876
FC99A7E8-DD11-5754-AFDC-B0A7621D887E
Figure 20.
Andrenainitialis Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena initialis Morawitz, 1876: 164, 166 (key), 199, ♀, ♂.
Type locality.
Shardara District (Kazakhstan).
Published (original) locality.
Kazakhstan: “between Keles [River] and Kosaral [Lake]”.
Lectotype (designated here).
♀, 24.[IV.1871] // Косаралъ [Kazakhstan, “Kosaral Lake” (now Shardara Reservoir), ≈ 41°10'N, 68°06'E] // Andrenainitialis Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andrenainitialis Mor., design. Osychnjuk, 1980 // Lectotypus Andrenainitialis Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000295.
Paralectotypes
(3 ♀, 1 ♂). 1 ♀, Косаралъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andrenainitialis Mor., design. Osychnjuk, 1980 [ZISP]; 1 ♀, 24.[IV.1871] // Косаралъ // Andrenainitialis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975; 1 ♀, 24.[IV.1871] // Косаралъ; 1 ♂, 24.[IV.1871] // Косаралъ // Andrenalateralis, D.A. Sidorov det., 2022 // Paralectotypus Andrenainitialis Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (incertae sedis) initialis Morawitz, 1876.
Remarks.
Male unknown. The single male from the type series belongs to Andrena (incertae sedis) lateralis Morawitz, 1876, as was mentioned by Gusenleitner and Schwarz (2001: 133).
Distribution.
Uzbekistan, Kazakhstan.
21. Andrena laeviventris
Morawitz, 1876
D63503E2-A06A-541E-8961-220548D83CFA
Figure 21.
Andrenalaeviventris Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena laeviventris Morawitz, 1876: 163 (key), 182, ♀.
Type locality.
Obburdon (Tajikistan).
Published (original) locality.
Uzbekistan: Gus [near Urgut]; Tajikistan: Pyandzhikent, Obburden [Obburdon].
Lectotype (designated here).
♀, 4.[VI.1870] // Оббурденъ [Tajikistan, Obburdon, 40°25'N, 69°18'E] // Andrenalaeviventris Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenalaeviventris Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Paralectotype.
1 ♀, 24.[V.1869] // Заравш.[анская] дол.[ина] [Zaravshan River Valley] // Paralectotypus Andrenalaeviventris Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Current status.
Andrena (Aciandrena) laeviventris Morawitz, 1876.
Remarks.
Male unknown (according to Gusenleitner and Schwarz 2002: 410).
Distribution.
Uzbekistan, Tajikistan.
22. Andrena lateralis
Morawitz, 1876
5E95C847-D890-53BE-AF42-B943B500DC75
Figure 22.
Andrenalateralis Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena lateralis Morawitz, 1876: 163, 166 (key), 200, ♀, ♂.
Type locality.
Shardara District (Kazakhstan).
Published (original) locality.
Kazakhstan: “between Keles [River] and Kosaral [Lake]”, Kyzyl-Kum [desert]; Uzbekistan: Urmitan [near Katty-Kurgan]; Tajikistan: Varzaminor [Ayni], near Obburden [Obburdon].
Lectotype.
♀, designated by by Osytshnjuk et al. 2005: 144, 24.[VI.1871] // Косаралъ [Kazakhstan, “Kosaral Lake” (now Shardara Reservoir), ≈ 41°10'N, 68°06'E] // Andrenalateralis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenalateralis Morawitz, 1876., design. Osytshnjuk et al., 2005 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(16 ♀, 12 ♂). 1 ♀, 24.[VI.1871] // Косаралъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenalateralis F.Mor. Popov 1935 det.; 1 ♀, 1 ♂, <golden circle>, 24.[VI.1871] // Косаралъ // lateralis Mor. Typ. [handwritten by F. Morawitz]; 2 ♂, 25.[IV.1871] // Чардара [Shardara] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenalateralis Mor. [handwritten by F. Morawitz] // Paralectotypus Andrenalateralis Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 4 ♀, 24.[IV.1871] // Косараль; 1 ♀, 30.[IV.1871] // Кизилкумъ [Kyzyl-Kum desert]; 9 ♀, Оббурденъ [Obburden]; 2 ♀, Варзаминоръ [Varzaminor]; 9 ♂, 25.[IV.1871] // Чардара // Lectotypus Andrenalateralis Morawitz, 1876, design. Osytshnjuk et al., 2005 <identical red labels on each paralectotype specimen, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (incertae sedis) lateralis Morawitz, 1876.
Remarks.
Osytshnjuk et al. (2005: 144) mistakenly mentioned sex of the lectotype as a male instead of female.
Distribution.
Europe, Russia (to East Siberia), Caucasus, Turkey, Israel, Iran, Afghanistan, Central Asia.
23. Andrena leucorhina
Morawitz, 1876
6D8B14C6-E3AC-5BD3-A0AC-A334A76E8EEE
Figure 23.
Andrenaleucorhina Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena leucorhina Morawitz, 1876: 165 (key), 169, ♂.
Type locality.
Shardara (Kazakhstan).
Published (original) locality.
Kazakhstan: Syr-Darja River, near Chardara.
Lectotype (designated here).
♂, 25.[IV.1871] // Чардара [Kazakhstan, Shardara, 41°18'N, 67°57'E] // Andrenaleucorhina Mor. [handwritten by F. Morawitz] // Lectotypus Warmcke 1975 <red label> // Lectotypus Andrenaleucorhina Morawitz, 1876 design. Astafurova et al., 2022 <red label> [ZMMU].
Paralectotypes
(2 ♂). 2 ♂, 25.[IV.1871] // Чардара [Chardara] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenaleucorhina Morawitz [handwritten by F. Morawitz] // Paralectotypus Andrenaleucorhina Morawitz, 1876 design. Astafurova et al., 2022 <red label> [1 ♂ – ZISP, 1 ♂ – ZMMU].
Current status.
Andrena (Ulandrena) abbreviata Dours, 1873 (according to Gusenleitner and Schwarz 2002).
Remarks.
Description of female: Morawitz 1876: 185 as Andrenaravicollis Morawitz, 1876 (synonymised by Osytshnjuk 1982: 28).
Distribution.
Southern Balkans, Cyprus, Turkey, Levant, Caucasus, Ukraine, Russia, Central Asia.
24. Andrena lucidicollis
Morawitz, 1876
341C135E-6C05-5F90-BACD-83421509E43E
Figure 24.
Andrenalucidicollis Morawitz, 1876, syntype, male A genitalia, dorsal view B habitus, lateral view and labels C head, frontal view D labrum, dorsal view E metasoma, dorsal view F mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena lucidicollis Morawitz, 1876: 165 (key), 181, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent.
Syntype.
♂, 5.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenalucidicollis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke, 1975 <red label> // Syntypus Andrenalucidicollis Mor., 1876 <red label> [ZMMU].
Current status.
Andrena (Poecilandrena) lucidicollis Morawitz, 1876
Remarks.
Female unknown (according to Gusenleitner and Schwarz 2002: 443).
Gusenleitner and Schwarz (2001: 140) refer to a “lectotype” deposited at the ZISP. However, we have found a single male specimen labelled “10.[IV.1871] // Ташкентъ // Andrenalucidicollis Mor. var. [handwritten by F. Morawitz] // Paralectotypus Andrenalucidicollis Mor., design. Osychnjuk, 1980”. This specimen does not belong to the type series formally, as it is labelled “10.[10 April 1871]” (in the original description Morawitz mentioned only specimens collected on 5 April). The syntype (Fig. 24) deposited at the ZMMU and labelled by Warncke as a “Lectotypus” on our opinion is in too poor a condition to be designated as a lectotype. However, both exemplars belong to one species and could be considered to be syntypes.
Distribution.
Uzbekistan.
25. Andrena maculipes
Morawitz, 1876
0AEAA303-FCD4-5360-A601-5590013F9B1D
Figure 25.
Andrenamaculipes Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena maculipes Morawitz, 1876: 162, 166 (key), 178, ♀, ♂.
Type locality.
Samarkand (Uzbekistan).
Published (original) locality.
Uzbekistan: near Samarkand.
Lectotype.
♂, designated by Osytshnjuk et al. 2005: 184, <golden circle>, 16.[III.1869] // Самаркандъ [Uzbekistan, Samarkand, 39°39'N, 66°57'E] // maculipes Mor. Typ. [handwritten by F. Morawitz] // Lectotypus Andrenamaculipes Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenamaculipes Morawitz, 1876, design. Osytshnjuk et al., 2005 <red label, labelled by Yu. Astafurova> // Zoological Institute St. Petersburg INS_HYM_0000187 [ZISP].
Paralectotypes
(12 ♀, 2 ♂). 2 ♀, 2 ♂, Самаркандъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenamaculipes F. Moraw. [handwritten by F. Morawitz]; 1 ♀, <golden circle>, 16.[III.1869] // Самаркандъ // maculipes Mor. Typ. [handwritten by F. Morawitz] // Paralectotypus Andr.maculipes Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 7 ♀, 7, 16.[III.1869] // Самаркандъ; 2 ♀, 4.[III.1869] // Заравш.[анская] дол.[ина] [Zaravshan River Valley] // Paralectotypus Andrenamaculipes Morawitz, 1876, design. Osytshnjuk et al., 2005 <identical red labels on each paralectotype specimen, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Chrysandrena) maculipes Morawitz, 1876.
Remarks.
Osytshnjuk et al. (2005: 184) mistakenly mentioned the lectotype depository as the ZMMU instead of the ZISP.
Distribution.
Central Asia.
26. Andrena majalis
Morawitz, 1876
AFACB171-0C0A-54F1-AC08-46299ECAF5A7
Figure 26.
Andrenamajalis Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena majalis Morawitz, 1876: 163 (key), 182, ♀.
Type locality.
Bairkum (Kazakhstan).
Published (original) locality.
Kazakhstan: Bairakum, Karak steppe; Uzbekistan: Zeravshan River valley, Dzham Gorge, Oalyk Gorge between Oalyk and Aksay.
Lectotype.
♀, designated by Osytshnjuk et al. 2008: 48, 4.[V.1871] // Байракумъ [Kazakhstan, Bairkum, Syr-Darya River, 42°05'N, 68°10'E] // Andrenamajalis Mor. [handwritten by F. Morawitz] // Lectotypus Warmcke, 1975 <red label> // Lectotypus Andrenamajalis Morawitz, 1876., design. Osytshnjuk et al., 2005 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(14 ♀). 3 ♀, the same labels as in the lectotype; 3 ♀, 5.[V.1871] // Каракск.[ая] степь [Karak steppe] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]// Paralectotypus Andr.majalis Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 4 ♀, 4.[V.1871] // Байракумъ; 1 ♀, 5.[V.1871] // Каракск.[ая] степь; 3 ♀, 16.,17.[V.1869] // Заравш.[анская] дол.[ина] [Zaravshan River Valley] // Paralectotypus Andrenamajalis Morawitz, 1876 design. Osychnjuk et al., 2008 <identical red labels on each paralectotype specimen, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Euandrena) majalis Morawitz, 1876.
Remarks.
Description of male: Shebl and Tadauchi 2009: 49, figs 23, 24.
Distribution.
Central Asia.
27. Andrena mordax
Morawitz, 1876
055E3059-8E4C-54CB-BF56-36294DB2D4EE
Figure 27.
Andrenamordax Morawitz, 1876, lectotype, male A genitalia, dorsal view B habitus, lateral view and labels C head, frontal view D labrum, dorsal view E T1–T3, dorsal view F mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena mordax Morawitz, 1876: 165 (key), 196, ♂.
Type locality.
Ayni (Tajikistan).
Published (original) locality.
Tajikistan: Varzaminor.
Lectotype.
♂, designated by Osytshnjuk et al. 2008: 122, 8.[VI.1870] // Варзаминоръ [Tajikistan, Varzaminor (= Ayni), 39°23'N, 68°32'E] // Andrenamordax Mor. [handwritten by F. Morawitz] // Lectotypus Warmcke 1975 <red label> // Lectotypus Andrenamordax Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(3 ♂). 2 ♂, 8.[VI.1870] // Варзаминоръ // Andrenamordax Mor. [handwritten by F. Morawitz] // Paralectotypus Andrenamordax Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 1 ♂, 9.[VI.1870] // Варзаминоръ // Paralectotypus Andrenamordax Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Hoplandrena) mordax Morawitz, 1876.
Remarks.
Description of female: Morawitz 1880: 362.
Distribution.
Uzbekistan, Tajikistan, Kyrgyzstan, Kazakhstan, China.
28. Andrena mucorea
Morawitz, 1876
5A85590B-42AF-594E-9EFF-0A4FAA607D96
Figure 28.
Andrenamucorea Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena mucorea Morawitz, 1876: 164, 165 (key), 212, ♀, ♂.
Type locality.
Between Tashkent and Keless River (Uzbekistan).
Published (original) locality.
Uzbekistan: between Tashkent and Keless River, Urmitan [near Katty-Kurgan]; Kazakhstan: “between Keles [River] and Kosaral [Lake]”, Kyzyl Kum [desert].
Lectotype (designated here).
♀, 23.[IV.1871] // Келесъ [River] [Uzbekistan, between Tashkent and Keless River, ≈ 41°15'N, 69°10'E] // Andrenamucorea Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andr.mucorea Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenamucorea Morawitz, 1876 design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000294 [ZMMU].
Paralectotypes
(12 ♀, 1 ♂). 1 ♀, 1 ♂, 22., 23.[IV.1871] // Келесъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenamucorea Mor. [handwritten by F. Morawitz]; 1 ♀, 24.IV.1871 // Косаралъ [Kosaral] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♀, Косаралъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // mucorea F. Mor., ♀ [handwritten by F. Morawitz] // Paralectotypus Andr.mucorea Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen > [ZISP]; 1 ♀, 10.[V.1869] // Урмитанъ [Urmitan] // Andrenamucorea Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label>; 7 ♀, 22., 23.[IV.1871] // Келесъ; 2♀, 9., 10.[V.1869] // Урмитанъ // Paralectotypus Andrenamucorea Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Poecilandrena) mucorea Morawitz, 1876.
Distribution.
Uzbekistan, Tajikistan, Kazakhstan.
29. Andrena nigrita
Morawitz, 1876
91DEE2B8-6FED-5FB1-9E67-2823415B3F1F
Figure 29.
Andrenanigrita Morawitz, 1876, holotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena nigrita Morawitz, 1876: 166 (key), 196, ♂.
Type locality.
Iskanderkul Lake (Tajikistan).
Published (original) locality.
Tajikistan: Iskanderkul Lake.
Holotype.
♂, 18.[VI.1870] // Искандеръ [Tajikistan, Iskanderkul Lake, Hissar Ridge, 39°04'N, 68°22'E] // Andrenanigrita Mor. [handwritten by F. Morawitz] // Lectotypus Warmcke 1975 <red label> // Holotypus Andrenanigrita Mor., 1876 <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Euandrena) nigritula Cockerell, 1906 [replacement name by Cockerell 1906: 74 (nec Andrenanigrita Fabricius, 1775; nec Nomadanigrita Panzer, 1800)].
Remarks.
Description of female: Morawitz 1893: 69.
The lectotype designation by Osytshnjuk et al. (2008: 36) is unnecessary as the species was described from a single male that was directly written about by Morawitz (1876: 196).
Distribution.
Uzbekistan, Tajikistan, Kyrgyzstan, Kazakhstan.
30. Andrena nitidicollis
Morawitz, 1876
6C0500DB-E21A-586A-B461-FBC4277C16E2
Figure 30.
Andrenanitidicollis Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena nitidicollis Morawitz, 1876: 162, 166 (key), 180, ♀, ♂.
Type locality.
Kyzylkum Desert (Kazakhstan).
Published (original) locality.
Kazakhstan: Kyzyl-kum Desert; Karak mountains [western part of Kysyl-kum, NW Bairkum].
Lectotype (designated here).
♀, 29.[IV.1871] // Кизилъкумъ [Kazakhstan, Kyzylkum Desert, NW of Shardara, ≈ 41°58'N, 67°03'E] // Andrenanitidicollis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenanitidicollis Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Paralectotype.
1 ♂, 6.[V.1871] // Кизилъкумъ // Paralectotypus Andrenanitidicollis Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Current status.
Andrena (incertae sedis) nitidicollis Morawitz, 1876.
Distribution.
Kazakhstan.
31. Andrena nupta
Morawitz, 1876
1D4DCA7C-1F1C-5485-AD49-3D7F45537F8A
Figure 31.
Andrenanupta Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena nupta Morawitz, 1876: 163, 166 (key), 191, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent and Samarkand.
Lectotype.
♀, designated by Osytshnjuk et al. 2008: 46, 8.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenanupta Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenanupta Morawitz, 1876, design. Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(5 ♀, 4 ♂). 1 ♀, the same label as in the lectotype; 2 ♀, 2 ♂, Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // nupta F. Mor. [handwritten by F. Morawitz] // Paralectotypus Andr.nupta Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 2 ♀, 1 ♂, 27.[III.1871], 3., 8.[IV.1871] // Ташкентъ; 1 ♂, 5.[IV.1869] // Самаркандъ [Samarkand] // Paralectotypus Andrenanupta Morawitz, 1876, design. Osytshnjuk et al., 2008 <identical red labels on each paralectotype specimen, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Euandrena) nupta Morawitz, 1876.
Distribution.
Uzbekistan, Kazakhstan.
32. Andrena oralis
Morawitz, 1876
9B29B2C9-3AE3-56A6-B52E-07F70DB32B87
Figure 32.
Andrenaoralis Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena oralis Morawitz, 1876: 162 (key), 177, ♀.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: near Tashkent.
Lectotype (designated here).
♀, <golden circle> // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // oralis Mor. Typ. [handwritten by F. Morawitz] // Lectotypus Andrenaoralis Mor., design. Osychnjuk, 1980 <red label> // Lectotypus, Andrenaoralis Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000201 [ZISP].
Paraectotype.
1 ♀, 5.[IV.1871] // Ташкент // Andrenaoralis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Paralectotypus Andrenaoralis Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Current status.
Andrena (Orandrena) oralis Morawitz, 1876.
Remarks.
Description of male: Morawitz 1876: 177, as Andrenasogdiana Morawitz, 1876 (according to Gusenleitner and Schwarz 2001: 150).
Distribution.
Europe, Russia (to the Urals), Turkey, Turkmenistan, Uzbekistan, Kazakhstan.
33. Andrena pannosa
Morawitz, 1876
9532633F-6932-54FE-9D00-E3F908AAA849
Figure 33.
Andrenapannosa Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena pannosa Morawitz, 1876: 162 (key), 197, ♀.
Type locality.
Khodzha-Chiburgan Gorge (Kyrgyzstan).
Published (original) locality.
Kyrgyzstan: Khodzha-Chiburgan Gorge.
Lectotype.
♀, <golden circle>, 21.[VI.1871] // Чибурганъ [Kyrgyzstan, Khodzha-Chiburgan Gorge, S Varuch, ≈ 39°48'N, 70°41'E] // pannosa Mor. Typ. [handwritten by F. Morawitz] // Lectotypus Andrenapannosa Mor., design. Osychnjuk, 1980 <red label> // Lectotypus, Andrenapannosa Morawitz, 1876 design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000287 [ZISP].
Paralectotypes
(2 ♀). 1 ♀, 21.[VI.1871] // Чибурганъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenapannosa Mor. [handwritten by F. Morawitz] // Paralectotypus Andr.pannosa Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 1 ♀, 21.[VI.1871] // Чибурганъ // Paralectotypus Andrenapannosa Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Euandrena) pannosa Morawitz, 1876.
Remarks.
Description of male: Gusenleitner and Schwarz 2001: 153. The lectotype was mistakenly designated by Osytshnjuk et al. (2008: 54) in ZMMU. This specimen is a female labelled “21.[06.1971] // Soch [Uzbek enclave] Andrenapannosa Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975.” However, according to the original description (Morawitz, 1876), the type location is “Khodzha-Chiburgan Gorge” [Kyrgyzstan, S Varuch]; thus, the lectotype published by Osytshnjuk et al. (2008) is not valid. There are two females in ZISP from “Chiburgan”, which correspond to the original description of Morawitz. One of these females labelled by A. Osytshnjuk as “lectotype” is designated here as a lectotype of Andrenapannosa.
Distribution.
Uzbekistan, Kyrgyzstan, Tajikistan, Kazakhstan.
34. Andrena planirostris
Morawitz, 1876
B23B5039-ED84-5D62-A6C3-AB2075AB4F9B
Figure 34.
Andrenaplanirostris Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena planirostris Morawitz, 1876: 163, 165 (key), 174, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: near Tashkent.
Lectotype (designated here).
♀, 23.[III.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenaplanirostris Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andr.planirostris Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenaplanirostris Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000256 [ZISP].
Paralectotypes
(20 ♀, 2 ♂). 1 ♀, <golden circle> // 25.[III.1871] // Ташкентъ // planirostris Mor. Typ. ♀ [handwritten by F. Morawitz]; 9 ♀, 23., 25.[III.1871] // Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♂, Ташкентъ // planirostris Mor. [handwritten by F. Morawitz] // Paralectotypus Andr.planirostris Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 10 ♀, 1 ♂, 11., 22., 23., 24., 25., 26. Ташкентъ // Paralectotypus Andrenaplanirostris Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Remarks.
There is a female specimen labelled by Warcke as a “Lectotypus” [<golden circle> // 5.[IV.1871] // Ташкент // Andrenaplanirostris Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975; ZMMU]. However, the label date does not correspond to any date [11–26.III.1871) mentioned for the type series by Morawitz (1876: 174).
Current status.
Andrena (Planiandrena) planirostris Morawitz, 1876.
Distribution.
Uzbekistan, Kazakhstan.
35. Andrena punctifrons
Morawitz, 1876
42502708-0CC4-593B-85A3-AEC547D2C9AD
Figure 35.
Andrenapunctifrons Morawitz, 1876, holotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena punctifrons Morawitz, 1876: 164 (key), 202, ♀.
Type locality.
Khodzha-Chiburgan River (Kyrgyzstan).
Published (original) locality.
Kyrgyzstan: “Khodzha-Chiburgan River”.
Holotype.
♀, Чибурганъ [Kyrgyzstan: Khodzha-Chiburgan River/Gorge, S Varuch, ≈ 39°48'N, 70°41'E] // Andrenapunctifrons Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Holotypus Andrenapunctifrons Mor., 1876 <red label> [ZMMU].
Current status.
Andrena (Micrandrena) punctifrons Morawitz, 1876.
Remarks.
Listed as Graecandrena by Osytshnjuk et al. (2008: 93). Male unknown (according to Osytshnjuk et al. 2008: 93).
Distribution.
Kyrgyzstan.
36. Andrena punctiventris
Morawitz, 1876
9252B04B-2569-5D89-8F09-732E73B8D1B8
Figure 36.
Andrenapunctiventris Morawitz, 1876, holotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena punctiventris Morawitz, 1876: 162 (key), 186, ♀.
Type locality.
Iskanderkul Lake (Tajikistan).
Published (original) locality.
Tajikistan: near Iskander-Kul Lake.
Holotype.
♀, 15.[VI.1870] // Искандеръ [Tajikistan, Iskanderkul Lake, Hissar Ridge, 39°04'N, 68°22'E] // Andrenapunctiventris Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Holotypus Andrenapunctiventris Mor., 1876 <red label> [ZMMU].
Current status.
Andrena (Lepidandrena) punctiventris Morawitz, 1876.
Remarks.
Listed as Poliandrena by Osytshnjuk (1994: 30). Description of male: Osytshnjuk 1994: 30, as Andrenapunctiventrisssp.basagiensis Osytshnjuk, 1994.
Distribution.
Uzbekistan, Tajikistan, Kazakhstan.
37. Andrena quadrifasciata
Morawitz, 1876
3DB49B1D-B095-50BA-878D-2362AF9DC8B7
Figure 37.
Andrenaquadrifasciata Morawitz, 1876, holotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena quadrifasciata Morawitz, 1876: 163 (key), 168, ♀.
Type locality.
Shakhimardan (Uzbekistan).
Published (original) locality.
Uzbekistan: near Shakhimardan.
Holotype.
♀, 7.[VII.1871] // Шагимарданъ [Shakhimardan in the Uzbek enclave in the territory of Kyrgyzstan, Alai Ridge, 39°58'N, 71°47'E] // Andrenaquadrifasciata Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Holotypus Andrenaquadrifasciata Mor., 1876 <red label> [ZMMU].
Current status.
Andrena (Simandrena) quadrifasciata Morawitz, 1876.
Remarks.
Description of male: Gusenleitner and Schwarz 2001: 157.
Distribution.
Tajikistan, Kazakhstan.
38. Andrena ravicollis
Morawitz, 1876
98834552-FE5D-5218-B35F-00F3F44B8B6B
Figure 38.
Andrenaravicollis Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena ravicollis Morawitz, 1876: 163 (key), 185, ♀.
Type locality.
Shardara (Kazakhstan).
Published (original) locality.
Kazakhstan: Chardara.
Lectotype (designated here).
♀, <golden circle> // 25.[IV.1871] // Чардара [Kazakhstan, Shardara, 41°18'N, 67°57'E] // ravicollis Mor. Typ. [handwritten by F. Morawitz] // Paralectotypus Andrenaravicollis Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenaravicollis Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Andrenaleucorhina, D.A. Sidorov det., 2022 // Zoological Institute St. Petersburg INS_HYM_0000286 [ZISP].
Paralectotype.
1 ♀, 25.[IV.1871] // Чардара // Andrenaleucorhina, D.A. Sidorov det., 2022 // Paralectotypus Andrenaravicollis Morawitz, 1876, design. Astafurova et al., 2022 < red label> [ZMMU].
Remarks.
There is a female specimen (in bad condition) labelled by Warcke as a “Lectotypus” [27.[IV.1871] // Чардара // Andrenaravicollis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975; ZMMU]. However, the label date does not correspond the date [25.III.1871] mentioned for the type series by Morawitz (1876).
Current status.
Andrena (Ulandrena) abbreviata Dours, 1873 (according to Gusenleitner and Schwarz 2002).
Distribution.
Southern Balkans, Cyprus, Turkey, Levant, Caucasus, Ukraine, Russia, Central Asia.
39. Andrena rufilabris
Morawitz, 1876
29FB8B08-629A-5DB2-A68A-1EDC2E1D675D
Figure 39.
Andrenarufilabris Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena rufilabris Morawitz, 1876: 162 (key), 180, ♀.
Type locality.
NW of Bairkum (Kazakhstan).
Published (original) locality.
Uzbekistan: Katty-Kurgan; Kazakhstan: Karak steppe.
Lectotype (designated here).
♀, 5.[V.1871] // Каракск.[ая] степь [Kazakhstan, Karak steppe, NW of Bairkum, ≈ 42°46'N, 67°24'E] // Andrenarufilabris Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenarufilabris Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Paralectotypes
(2 ♀). 2 ♀, 29.[VI.1869] and 6.[V.1869] // Катты Курганъ // Andrenasatra, D.A. Sidorov det., 2022 // Paralectotypus, Andrenarufilabris Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Current status.
Andrena (Simandrena) sarta Morawitz, 1876 (according to Gusenleitner and Schwarz 2002: 660).
Distribution.
Uzbekistan, Kazakhstan.
40. Andrena rufina
Morawitz, 1876
2698CE2C-0CD4-5A2A-89ED-33ADE9DA7DCA
Figure 40.
Andrenarufina Morawitz, 1876, lectotype, male A genitalia, dorsal view B habitus, lateral view and labels C head, frontal view D labrum, latero-dorsal view E metasoma, dorsal view F mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena rufina Morawitz, 1876: 162, 165 (key), 167, ♀, ♂.
Type locality.
Anzob (Tajikistan).
Published (original) locality.
Tajikistan: Anzob.
Lectotype.
♂, designated by Osytshnjuk et al. 2005: 204, 20.VI.1870 // Ягнобъ [Tajikistan, Yagnob River near Anzob, 39°04'N, 68°52'E] // Andrenarufina Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 // Lectotypus Andrenarufina Morawitz, 1876, design. Osytshnjuk et al., 2005 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotype.
♀, 75. // Туркест.[анский] кр.[ай] [Turkestan] // Paralectotypus, Andrenarufina Morawitz, 1876, design. Osytshnjuk et al., 2005 <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Cnemidandrena) rufina Morawitz, 1876.
Distribution.
Uzbekistan, Tajikistan, Kazakhstan.
41. Andrena sarta
Morawitz, 1876
4801D484-8040-534D-9125-A0496776FEA9
Figure 41.
Andrenasarta Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena sarta Morawitz, 1876: 164 (key), 171, ♀.
Type locality.
30 km SE of Kozhatogai, Turkistan Province (Kazakhstan).
Published (original) locality.
Kazakhstan: steppe between Tashkent and Syrdarya River.
Lectotype (designated here).
♀, 20.[V.1871] // Степь м.[ежду] С.[ыр] [-] д.[арьей] и Т.[ашкентом] [Kazakhstan, Turkistan Province, steppe between Syrdarya River and Tashkent, 30 km SE of Kozhatogai, 41°47'N, 68°23'E] // satra Mor. [handwritten by Osytshnjuk] // Lectotypus Warncke 1975 // Lectotypus Andrenasarta Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Current status.
Andrena (Simandrena) sarta Morawitz, 1876.
Remarks.
Description of male: Gusenleitner and Schwarz 2001: 163.
Distribution.
Uzbekistan, Kazakhstan.
42. Andrena semiaenea
Morawitz, 1876
7EC68AB0-DB8B-50C4-B89A-0CF6054B5A10
Figure 42.
Andrenasemiaenea Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena semiaenea Morawitz, 1876: 164 (key), 213, ♀.
Type locality.
Keles River, Turkistan Province (Kazakhstan).
Published (original) locality.
Kazakhstan: between Keles River and dry Keles; near Kosaral Lake; near Chardara.
Lectotype.
♀, designated by Osytshnjuk et al. 2008: 155, 23.[IV.1871] // Келесъ [Kazakhstan, Turkistan Province, Keles River] // Andrenasemiaenea Mor. [handwritten by F. Morawitz] // Lectotypus Warncke, 1975 // Lectotypus Andrenasemiaenea Morawitz, 1876, Osytshnjuk et al., 2008 <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(4 ♀). 3 ♀, 25.IV.1871 // Чардара [Chardara] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andr.semiaenea Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 1 ♀, 24.[IV.1871] // Косараль [Kosaral] // Paralectotypus Andenasemiaenea Mor., design. Osytshnjuk et al., 2008 <identical red labels on each paralectotype specimen, labelled Yu. Astafurova> [ZMMU].
Current status.
Andrena (Poecilandrena) semiaenea Morawitz, 1876.
Remarks.
Listed as Lepidandrena by Osytshnjuk et al. (2008: 155). Description of male: Gusenleitner and Schwarz 2001: 167.
Distribution.
Afghanistan, Uzbekistan, Kazakhstan, Mongolia.
43. Andrena smaragdina
Morawitz, 1876
1DE20C98-571A-51BA-9C30-4D65FD27CD34
Figure 43.
Andrenasmaragdina Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena smaragdina Morawitz, 1876: 164, 165 (key), 211, ♀, ♂.
Type locality.
Samarkand (Uzbekistan).
Published (original) locality.
Uzbekistan: Samarkand.
Lectotype (designated here).
♀, Самаркандъ [Uzbekistan, Samarkand, 39°39'N, 66°57'E] // Andrenasmaragdina Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andrenasmaragdina Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenasmaragdina Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000293 [ZISP].
Paralectotypes
(2 ♀, 1 ♂). 1 ♀, 28.[II.1869] // Самаркандъ // Andrenasmaragdina Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label>; 1 ♀, 1 ♂, 10., 28.[II.1869] // Самаркандъ // Paralectotypus Andrenasmaragdina Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Current status.
Andrena (Notandrena) smaragdina Morawitz, 1876.
Remarks.
Listed as Poecilandrena by Osytshnjuk et al. (2005: 28).
Distribution.
Uzbekistan, Tajikistan.
44. Andrena sogdiana
Morawitz, 1876
3BEFB11C-F0B3-50CF-BB78-B4EBFD4EF6E0
Figure 44.
Andrenasogdiana Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, latero-dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena sogdiana Morawitz, 1876: 165 (key), 177, ♂.
Type locality.
Samarkand (Uzbekistan).
Published (original) locality.
Uzbekistan: Samarkand.
Lectotype (designated here).
♂, 11.[IV.1869] // Самаркандъ [Uzbekistan, Samarkand, 39°39'N, 66°57'E] // Andrenasogdiana Mor. [handwritten by F. Morawitz] // Paralectotypus Andrenasogdiana Mor., design. Osychnjuk, 1980 <red label> // Andrenaoralis, ♂, D.A. Sidorov det. 2022 // Lectotypus Andrenasogdiana Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000292 [ZISP].
Current status.
Andrena (Orandrena) oralis Morawitz, 1876 (according to Gusenleitner and Schwarz 2001: 150).
Remarks.
There is a male specimen labelled by Warncke as a “Lectotypus” (3.[IV.1869] // Самарканд // Andrenasogdiana Mor.[handwritten by F. Morawitz] // Lectotypus Warncke 1975; ZMMU). However, the label date does not correspond the date [11.IV.1869] mentioned for the type series by Morawitz (1876).
Distribution.
Europe, Russia (to the Urals), Turkey, Turkmenistan, Uzbekistan, Kazakhstan.
45. Andrena sordida
Morawitz, 1876
1462616D-C19B-5A54-94EC-7FD24029765F
Figure 45.
Andrenasordida Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena sordida Morawitz, 1876: 163, 166 (key), 173, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: near Tashkent.
Lectotype.
♀, designated by Osytshnjuk et al. 2008: 161, 5.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenasordida Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenasordida Morawitz, 1876, design. Osytshnjuk et al., 2008 // <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(83 ♀, 2 ♂). 1 ♀, 1 ♂, the same label as in the lectotype; 4 ♀, Ташкентъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // sordida [handwritten by F. Morawitz] // Paralectotypus Andr.sordida Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 78 ♀, 1 ♂, 1., 3., 5., 8.[IV.1871], 28.[III.1871] // Ташкентъ // Paralectotypus Andrenasordida Mor. design. Osytshnjuk et al., 2008 // <red label, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Leucandrena) sordidella Viereck, 1918, replacement name for A.sordida Morawitz, 1876 (nec Apissordida Scopoli, 1763; nec Apissordida Gmelin, 1790).
Distribution.
Uzbekistan.
46. Andrena subaenescens
Morawitz, 1876
D0F5D5E8-812B-5682-9441-5276A1375D7D
Figure 46.
Andrenasubaenescens Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena subaenescens Morawitz, 1876: 164, 166 (key) 207, ♀, ♂.
Type locality.
Samarkand (Uzbekistan).
Published (original) locality.
Uzbekistan: Samarkand.
Lectotype (designated here).
♀, <golden circle> // 27.[II.1869] // Самаркандъ [Uzbekistan, Samarkand, 39°39'N, 66°57'E] // subaenescens Mor. Typ. [handwritten by F. Morawitz] // Paralectotypus Andrenasubaenescens Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenasubaenescens Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000255 [ZISP].
Paralectotypes
(8 ♀, 1 ♂). 3 ♀, 27.[II.1869] // Самаркандъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andr.sordida Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 1 ♀, 27.[II.1869] // Самаркандъ // Andrenasubaenescens Mor. [handwritten by F. Morawitz] // Lectotypus Warncke, 1975 <red label>; 4 ♀, 1 ♂. 27., 28.[II–18.III.1869] // Самаркандъ // Paralectotypus Andrenasubaenescens Morawitz, 1876, design. Astafurova et. al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Poecilandrena) subaenescens Morawitz, 1876.
Distribution.
Uzbekistan.
47. Andrena temporalis
Morawitz, 1876
6C6D8114-958B-53F9-A23C-DC08B526346F
Figure 47.
Andrenatemporalis Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena temporalis Morawitz, 1876: 165 (key), 204, ♂.
Type locality.
Samarkand (Uzbekistan).
Published (original) locality.
Uzbekistan: Samarkand.
Lectotype.
♂, designated by Osytshnjuk et al. 2005: 151, 20.[III.1869] // Самаркандъ [Uzbekistan, Samarkand, 39°39'N, 66°57'E] // Andrenatemporalis Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus, Andrenatemporalis Morawitz, 1876, design. Osytshnjuk et al., 2008 // <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(7 ♂). 1 ♂, <golden circle> // 20.[III.1869] // Самаркандъ // temporalis Mor. Typ. [handwritten by F. Morawitz]; 1 ♂, Самаркандъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Andrenatemporalis F. Morawitz, ♂ [handwritten by F. Morawitz] // Paralectotypus Andr.temporalis Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 5 ♂, 16., 20.[III.1869] // Самаркандъ // Paralectotypus, Andrenatemporalis Morawitz, 1876, design. Osytshnjuk et al., 2005 et al. <identical red labels on each paralectotype specimen, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Notandrena) hieroglyphica Morawitz, 1876 (synonymised by Osytshnjuk 1984: 3).
Distribution.
Iran, Turkmenistan, Uzbekistan, Tajikistan, Pakistan.
48. Andrena tuberculiventris
Morawitz, 1876
24AED2F6-5BDF-5506-83FF-44B1EA1B7476
Figure 48.
Andrenatuberculiventris Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena tuberculiventris Morawitz, 1876: 166 (key), 184, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent.
Lectotype (designated here).
♂, 10.[IV.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenatuberculiventris Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus Andrenatuberculiventris Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Paralectotypes
(3 ♂). 1 ♂, <golden circle> // 10.[IV.1871] // Ташкентъ // tuberculiventris Mor. Typ. [handwritten by F. Morawitz] // Paralectotypus Andr.tuberculiventris Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 2 ♂, 10.[IV.1871] // Ташкентъ // Paralectotypus Andrenatuberculiventris Morawitz, 1876, design. Astafurova et al., 2022 <red label> [ZMMU].
Current status.
Andrena (Parandrenella) tuberculiventris Morawitz, 1876 (according to Scheuchl et al. 2011: 225).
Remarks.
Listed as Andrena (Parandrenella) bicarinata Morawitz, 1876 by Gusenleitner and Schwarz (2002: 123).
Distribution.
Uzbekistan, Tajikistan.
49. Andrena turkestanica
Morawitz, 1876
D1059395-B44F-5680-B205-0F4EA578CEE0
Figure 49.
Andrenaturkestanica Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena turkestanica Morawitz, 1876: 162, 166 (key), 192, ♀, ♂.
Type locality.
Urgut, Zeravshan River valley (Uzbekistan).
Published (original) locality.
Uzbekistan: Zeravshan River valley; steppe between Ulus and Dzham.
Lectotype (designated here).
♀, Верхн.[ий] Заравш.[ан] [Uzbekistan, Zeravshan River valley, near Urgut, 39°24'N, 67°13'E] // turkestanica F. Mor. ♀, [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andr.turkestanica Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenaturkestanica Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000285 [ZISP].
Paralectotypes
(3 ♀). 2 ♀, <golden circle> // Верхн. Заравш. // turkestanica Mor. Typ. ♀ [handwritten by F. Morawitz] // Paralectotypus Andr.turkestanica Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 1 ♀, 12.[V.1869] // Верхн. Заравш. // Andrenaturkestanica Mor. // Lectotypus Warncke 1975 <red label> // Paralectotypus Andrenaturkestanica Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Melanapis) fuscosa Erichson, 1835 (synonymised by Warncke 1967: 315).
Remarks.
Listed as “turcestanica” by Gusenleitner and Schwarz (2002: 303).
Distribution.
Europe, North Africa, Russia (Dagestan Rep.), Caucasus, Turkey, Syria, Lebanon, Israel, Jordan, Iran, Afghanistan, Pakistan, Central Asia, India.
50. Andrena urmitana
Morawitz, 1876
EFE42609-8123-5324-8A1B-58479B7E0AD7
Figure 50.
Andrenaurmitana Morawitz, 1876, lectotype, male A genitalia, dorsal view B habitus, lateral view and labels C head, frontal view D labrum, dorsal view E T1–T3, dorsal view F mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena urmitana Morawitz, 1876: 165 (key), 175, ♂.
Type locality.
Urmetan (Tajikistan).
Published (original) locality.
Tajikistan: near Urmitan [Urmetan].
Holotype.
♂, 2.[V.1869] // Урмитанъ [Tajikistan, Urmetan, 39°26'N, 68°15'E] // Andrenaurmitana Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Holotypus Andrenaurmitana Mor., 1876 <red label> [ZMMU].
Current status.
Andrena (Nobandrena) acutilabris Morawitz, 1876 (according to Gusenleitner and Schwarz 2002: 410).
Distribution.
Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan.
51. Andrena virescens
Morawitz, 1876
3B63BDF3-4103-57A6-84AF-18ABE16ECA87
Figure 51.
Andrenavirescens Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena virescens Morawitz, 1876: 164, 165 (key), 209, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: near Samarkand, near Tashkent, between Tashkent and Keless River.
Lectotype (designated here).
♀, Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // virescens Mor., ♀ [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andr.virescens Mor., design. Osychnjuk, 1980 <red label> // Lectotypus Andrenavirescens Morawitz, 1876, design. Astafurova et al., 2022 <red label> // Zoological Institute St. Petersburg INS_HYM_0000254 [ZISP].
Paralectotypes
(7 ♀, 8 ♂). 4 ♂, 27., 28.[II.1869] // Самаркандъ // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Andr.virescens Mor., design. Osychnjuk, 1980 <red label> [ZISP]; 1 ♂, 27.[II.1869] // Самаркандъ // Andrenavirescens Mor. [handwritten by F. Morawitz] // Lectotypus Warncke, 1975 <red label>; 5 ♀, 1 ♂, 10., 23., 24.[III.1871], 8.[IV.1871] // Ташкентъ; 1 ♀, 2 ♂, 27.[II.1869] // Самаркандъ ; 1 ♀, 22.[IV.1871] // Келесъ // // Paralectotypus, Andrenavirescens Morawitz, 1876, design. Astafurova et al., 2022 <identical red labels on each paralectotype specimen> [ZMMU].
Current status.
Andrena (Poecilandrena) virescens Morawitz, 1876.
Distribution.
Uzbekistan, Tajikistan.
52. Andrena viridigastra
Morawitz, 1876
ACA99AF2-42E2-566A-AEAA-5EDD94580840
Figure 52.
Andrenaviridigastra Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C labrum, dorsal view D T1–T3, dorsal view E mesosoma, dorsal view. Scale bar: 1.0 mm.
Andrena viridigastra Morawitz, 1876: 164, 165 (key), 206, ♀, ♂.
Type locality.
Tashkent (Uzbekistan).
Published (original) locality.
Uzbekistan: Tashkent, Samarkand.
Lectotype.
♀, designated by Osytshnjuk et al. 2008: 220, 28.[II.1871] // Ташкентъ [Uzbekistan, Tashkent, 41°18'N, 69°16'E] // Andrenaviridigastra Mor. [handwritten by F. Morawitz] // Lectotypus Warncke 1975 <red label> // Lectotypus, Andrenaviridigastra Morawitz, 1876, design. Osytshnjuk et al., 2008 // <red label, labelled by Yu. Astafurova> [ZMMU].
Paralectotypes
(29 ♀, 32 ♂). 1 ♀, <golden circle>, 16.[III.1869] // Самаркандъ // viridigastra Mor., ♀, Typ. [handwritten by F. Morawitz]; 2 ♀, 27.[II.1871] // Ташкентъ [Tashkent] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // viridigastra Mor. [handwritten by F. Morawitz]; 1 ♂, <golden circle> // 16.[III.1871] // Ташкентъ // viridigastra Mor. ♂, Typ. [handwritten by F. Morawitz] // Paralectotypus Andr.viridigastra Mor., design. Osychnjuk, 1980 <identical red labels on each paralectotype specimen> [ZISP]; 20 ♀, 14 ♂, 28.[II.1871], 16., 23., 24., 25., 27.[III.1871], 5., 8., 10., 11.[IV.1871] // Ташкентъ; 6 ♀, 17 ♂, 16., 18., 21., 25[III.1869] // Самаркандъ // Paralectotypus, Andrenaviridigastra Morawitz, 1876, design. Osytshnjuk et al., 2005 <identical red labels on each paralectotype specimen, labelled by Yu. Astafurova> [ZMMU].
Current status.
Andrena (Melandrena) viridigastra Morawitz, 1876.
Distribution.
Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan.
Supplementary Material
Acknowledgements
We are grateful to A.V. Antropov (ZMMU) for assisting during the work in the ZMMU. We also thank A.S. Lelej (Vladivostok, Russia) and T.J. Wood (Mons, Belgium) for careful review of the manuscript and comments that improved the text. This investigation was supported by the Russian Funds for Basic Research (grant number 20–54–44014), and the state research projects 122031100272-3 and 121031000151-3.
Citation
Astafurova YuV, Proshchalykin MYu, Sidorov DA (2022) The bees of the genus Andrena Fabricius, 1775 (Hymenoptera, Andrenidae) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko. ZooKeys 1120: 105–176. https://doi.org/10.3897/zookeys.1120.90206
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