An updated checklist of liverworts and hornworts of Malaysia

Abstract

An updated checklist of the liverworts and hornworts of Malaysia accepts 773 species and 31 infraspecific taxa of liverworts, in 120 genera and 40 families, and 7 species of hornworts (6 genera, 3 families). The largest family is Lejeuneaceae with 312 species in 30 genera, accounting for 40% of the total number of species. The largest genera are Cololejeunea, Bazzania and Frullania with 90, 61 and 55 species, respectively. The greatest number of species has been recorded from Sabah with 568 species, followed by Pahang and Sarawak with 338 and 265 species, respectively.

Introduction

The country of Malaysia consists of two separate regions, Peninsular Malaysia in the West, being part of the Asian mainland, and the states of Sabah and Sarawak on the island of Borneo in the East. Physiogeographically, the two regions are characterized by hills and mountain ranges. On Peninsular Malaysia, mountain ranges cover about one-third of the total land surface, running parallel to the long axis of the peninsula. They include Titiwangsa Range (the main range), Bintang Range, Nakawan Range, Kedah-Singgora Range, Keledang Range, Benom Range, Tahan Range and Pantai Timur Range. Many of them rise to 2000 m and about 23% of the land lies above 300 m (Wyatt-Smith, 1963; Mohamed and Tan 1988). Mount Tahan (2189 m) in Titiwangsa Range, at the border with Thailand, is the highest peak on the peninsula. The rest of the area consists of coastal plains and alluvial terraces, usually extending 15 to 65 km inland from the coast, as well as riverine flood plains.

Sabah’s highlands include Crocker Range, Trus Madi Range, Witti Range, Maitland Range and Brassey Range. Among these, Crocker Range is the longest and highest mountain range, rising to 4095 m on Mt Kinabalu, being the highest mountain of Malaysia and of Southeast Asia. Those of Sarawak are Tama Abu Range, Iran Mountains, Hose Mountains, Kapuas Hulu Range and Kelinkang Range. The physiographies of these two states are somewhat different. Only a small proportion of the land of Sabah consists of lowland, including coastal plains and river valleys. Sarawak, in contrast, has more low-lying, flat lands and swampy coastal areas. Both states, however, have steep undulating hills and mountains in the interior parts, acting as a barrier against inland penetration.

Climate

Malaysia is located near the Equator and experiences a humid tropical climate with constant temperatures, high humidity and copious rainfall. Rainfall distribution is usually seasonal dependent on wind flow patterns and physiogeography (Malaysian Meteorological 2021). Generally, the East Coast states of Peninsular Malaysia receive high rainfall almost throughout the year with June and July being the driest months. The remaining states (except for the southwest coastal region) have two maximum rainfall periods (October-November and April-May) that are separated by drier periods (Malaysian Meteorological 2021). The country is subjected to four monsoonal seasons each year: northeast monsoon, southwest monsoon and two shorter inter-monsoon periods. The northeast monsoon normally occurs from November to March and the southwest monsoon during May or early June to September.

Vegetation and flora

The vegetation of Malaysia can be classified into different types based on climate and soil. Climatic vegetation types include evergreen and semi-evergreen lowland rainforest, lower montane forest and the upper montane forest, while the edaphic types include heath vegetation, limestone forest, beach vegetation, mangrove forest, brackish water vegetation, peat swamp forest, freshwater swamp forest and riparian vegetation (Symington 2004). In lowland and hill areas, dipterocarp forests are the most prominent forest type (Ashton 1995; Saw 2010). Malaysia has a very rich flora that is made up of Sundaic and mainland Asiatic floristic elements in Peninsular Malaysia and of Sundaic elements with many local endemics on the island of Borneo (Saw and Chung 2015). The Malaysian rainforests stand out by their very high species diversity (e.g., Austin et al. 1973; Baillie and Ashton 1983; Baillie et al. 1987; Saw and Chung 2015), as well as by intricate stratification and regeneration dynamics patterns that have been the subject of numerous major studies by specialists from all over the world (e.g., Burgess 1961, 1975; Denslow 1987; Ashton and Hall 1992; Manokaran et al. 1992; Saiful and Latiff 2017).

Malaysian liverwort and hornwort flora

The first checklist of liverworts and hornworts of Malaysia, recognizing 727 species and 37 infraspecific taxa in 127 genera and 39 families, was published by Chuah-Petiot (2011). Lee et al. (2018) presented a review of the history of liverwort exploration in Malaysia and the more recent papers on Malaysian liverworts, and further additions to the liverwort flora were published by Pócs et al. (2020), Pesiu et al. (2021) and Sarimi et al. (2021). A manual of the genera of liverworts and hornworts of Malaysia has been prepared by Lee and Gradstein (2021). The present paper provides an updated checklist of liverworts and hornworts of Malaysia based on a compilation of the published literature and a critical revision of the nomenclature of the taxa and new synonymy, resulting from taxonomic revisions. The updated checklist accepts 773 species and 31 infraspecific taxa in 120 genera in 40 families of liverworts, and 7 species (3 families, 6 genera) of hornworts. The largest family is Lejeuneaceae with 312 species in 30 genera, accounting for 40% of the total number of liverwort species. The largest genera are Cololejeunea, Bazzania and Frullania with 90, 61 and 55 species, respectively. Genera with more than ten species include Lejeunea (40), Plagiochila (35), Radula (34), Colura (27), Drepanolejeunea (26), Cheilolejeunea (24), Riccardia (19), Lepidozia (18), Acromastigum (16), Leptolejeunea (16), Thysananthus (16), Solenostoma (15), Lopholejeunea (12), Heteroscyphus (13) and Anastrophyllum (10). Forty eight genera have fewer than 10 species (e.g., Acrolejeunea, Andrewsianthus, Cephalozia, Ceratolejeunea, Diplasiolejeunea, Kurzia, Marchantia, Microlejeunea, Metzgeria, Odontoschisma, Schistochila, Syzygiella, Trichocolea and Tricholepidozia) and 54 are represented only by a single species, such as Acroscyphella, Balantiopsis, Conoscyphus, Eotrichocolea, Haplomitrium, Jubula, Kymatocalyx, Mastigopelma, Metalejeunea, Mesoptygia, Nardia, Pictolejeunea, Soella, Southbya, Treubia, Triandrophyllum, Tritomaria and the five genera of hornworts. Sabah has the largest number of species with 568 recorded species, followed by Pahang and Sarawak with 338, 265, respectively (Table 1).

Table 1.

The number of recorded species of liverworts and hornworts per state in Malaysia.

State	Number of species
	Liverworts	Hornworts	Total
Sabah	565	3	568
Pahang	334	4	338
Sarawak	263	2	265
Penang	95	–	95
Perak	82	1	83
Johor	81	–	81
Terengganu	62	–	62
Kedah including Langkawi	59	1	60
Selangor	48	–	48
Negeri Sembilan	44	–	44
Kelantan	35	–	35
Malacca	11	–	11
Perlis	5	–	5

An updated checklist of liverworts and hornworts of Malaysia

The nomenclature follows Söderström et al. (2016) with updates. For explanation of some of the new nomenclature, see Lee and Gradstein (2021). In taxon authorities, only the name of the validating author is cited and “ex” authors are omitted for brevity and because their citation is not obligatory. Reports from Malaysia that are only based on secondary information (= papers dealing with liverworts of other regions (not Malaysia) and citing ‘Malaya’, ‘Malaysia’, ‘Malay Peninsular’, ‘Malacca’ etc. in the section on species distribution) and are not supported by direct evidence, are omitted. Similarly, records based on doubtful identifications (“cf.”, “aff.”) are not taken into account. Furthermore, we have omitted all reports citing only “Borneo” unless there is evidence that the record was from Malaysia (Sarawak, Sabah). For example, species reported by Stephani from Borneo based on material collected by Everett (e.g., Bazzaniafleischeri (Steph.) Abeyw., B.inaequitexta Steph., Plagiochilaborneensis Steph., P.kuhliana Sande Lac.) are included in the checklist because these Everett collections are presumably from Sarawak (Menzel 1988).

Liverworts (Marchantiophyta)

Acrobolbusciliatus (Mitt.) Schiffn. – Kodama and Narita (1974). Sabah.

Acrobolbussaccatus (Hook.) Briscoe – Kitagawa and Kodama (1974 as Tylimanthussaccatus). Sabah.

Acrolejeuneaarcuata (Nees) Grolle et Gradst. – Kodama and Kitagawa (1974 as Ptychocoleuscoroniformis), Gradstein (1975), Kürschner (1990), Lee et al. (2013), Renner (2013), Wang et al. (2014b), Cheah (2017). Pahang, Sabah.

Acrolejeuneacrassicaulis (Steph.) J.Wang bis et Gradst. – Mizutani (1989 as Trocholejeuneacrassicaulis), Gradstein et al. (2002 as T.crassicaulis), Wang et al. (2014a). Sabah.

Acrolejeuneafertilis (Reinw., Blume et Nees) Schiffn. – Verdoorn (1933 as Ptychocoleuswichurae, 1934a as P.fertilis), Mizutani (1969 as P.fertilis), Gradstein (1975), Lee et al. (2013). Penang, Perak, Pahang, Sabah.

Acrolejeuneapycnoclada (Taylor) Schiffn. – Schiffner (1898), Verdoorn (1933, 1934a both as Ptychocoleuspycnocladus), Mizutani (1969 as P.pycnocladus), Kitagawa (1971 as P.pycnocladus), Kamimura (1974 as P.pycnocladus), Gradstein (1975), Lee et al. (2013), Wang et al. (2016). Penang, Perak, Malacca, Sarawak, Sabah.

Acrolejeuneatjibodensis (Verd.) Grolle et Gradst. – Kamimura (1974 as Ptychocoleustjibodensis), Gradstein (1975), Lee et al. (2013). Pahang, Sabah.

Acromastigumaurescens A.Evans – Evans (1934). Johor.

Acromastigumbancanum (Sande Lac.) A.Evans – De Notaris (1874 as Mastigobryumbancanum), Schiffner (1898 as Bazzaniabancana), Evans (1934, also as A.bidenticulatum), Herzog (1950), Kitagawa and Kodama (1973), Tixier (1974 as A.bidenticulatum), Kürschner (1990), Pócs et al. (2020). Kedah, Johor, Sarawak, Sabah.

Acromastigumbifidum (Steph.) A.Evans – Evans (1934 as A.denticulatum), Herzog (1950), Grolle (1978). Sarawak.

Acromastigumbrotheri (Steph.) A.Evans – Stephani (1909 as Mastigobryumbrotheri), Evans (1934), Kitagawa and Kodama (1973). Sarawak, Sabah.

Acromastigumcurtilobum A.Evans – Kitagawa and Kodama (1973), Kürschner (1990). Sabah.

Acromastigumdivaricatum (Nees) Reimers – Evans (1934), Kitagawa and Kodama (1973). Johor, Sabah.

Acromastigumechinatiforme (De Not.) A.Evans – De Notaris (1874 as Mastigobryumechinatiforme), Schiffner (1898 as Bazzaniaechinatiformis), Evans (1934), Herzog (1950), Kitagawa and Kodama (1973), Grolle (1978), Kürschner (1990), Cheah (2017). Pahang, Sarawak, Sabah.

Acromastigumechinatum (Gottsche) A.Evans – Evans (1934), Herzog (1950), Kitagawa and Kodama (1973), Tixier (1974). Kedah, Johor, Sarawak, Sabah.

Acromastigum fimbriatum (Steph.) A.Evans – Stephani (1909 as Mastigobryumfimbriatum), Evans (1934), Herzog (1950). Sarawak.

Acromastigumherzogii Grolle – Grolle (1964), Kürschner (1990). Sarawak, Sabah.

Acromastiguminaequilaterum (Lehm. et Lindenb.) A.Evans – De Notaris (1874 as Mastigobryumelegantulum), Schiffner (1898 as Bazzanianotarisii), Stephani (1909 as M.notarisii), Evans (1934), Herzog (1950), Kitagawa (1969c), Kitagawa and Kodama (1973), Kürschner (1990), Wolseley et al. (1996). Penang, Negeri Sembilan, Johor, Sarawak, Sabah.

Acromastigumlaevigatum A.Evans – Evans (1934), Tixier (1974). Kedah, Sarawak.

Acromastigumleptophyllum Herzog – Herzog (1950). Sarawak.

Acromastigumlinganum (De Not.) A.Evans – De Notaris (1874 as Mastigobryumlinganum), Schiffner (1898 as Bazzanialingana), Stephani (1909 as M.linganum), Evans (1934). Sarawak.

Acromastigumlobuliferum A.Evans – Grolle (1978). Sabah.

Acromastigumobliquatum (Mitt.) A.Evans – Kitagawa and Kodama (1973 as A.gemmiferum), Grolle (1978). Sabah.

Acroscyphellatjiwideiensis (Sande Lac.) N.Kitag. et Grolle – Cheah (2017). Pahang.

Allorgellasemperiana (Steph.) Bechteler, G.E.Lee, Schäf.-Verw. et Heinrichs – Tixier (1980b as A.changiana), Grolle (1985 as Otolejeuneasemperiana), Zhu and So (2001 as O.semperiana), Schäfer-Verwimp (2006 as O.semperiana). Kedah, Perak, Pahang.

Anastrophyllumassimile (Mitt.) Steph. – Kitagawa (1970), Gradstein and Váňa (1987), Váňa and Piippo (1989), Váňa (1991c). Sabah.

Anastrophyllumauritum (Lehm.) Steph. – Kitagawa (1970 as A.appendiculatum), Váňa and Piippo (1989), Váňa (1991c). Sabah.

Anastrophyllumbidens (Reinw., Blume et Nees) Steph. – Inoue (1967a, 1968b), Kitagawa (1970), Kitagawa and Kodama (1973), Kürschner (1990), Váňa (1991c). Pahang, Sarawak, Sabah.

Anastrophyllumbidens var. aristatum N.Kitag. – Herzog (1950), Kitagawa (1970), Kitagawa and Kodama (1973), Kürschner (1990), Váňa (1991c). Sarawak, Sabah.

Anastrophyllumborneense Herzog – Herzog (1950), Inoue (1967a,b, 1968b all as A.malayense), Kitagawa (1970), Váňa (1991c). Pahang, Sarawak, Sabah.

Anastrophyllumfissum Steph. – Herzog (1950 as A.divergens), Váňa (1991c as A.divergens). Sarawak, Sabah.

Anastrophyllumobtusum Herzog – Herzog (1950), Váňa (1991c). Sabah.

Anastrophyllumpiligerum (Nees) Steph. – De Notaris (1874 as Jungermanniapiligera [‘pilifera’]), Schiffner (1898, also as A.imbricatum), Herzog (1950, also as A.imbricatum), Inoue (1967a), Kitagawa (1969c, 1970), Kitagawa and Kodama (1973), Kürschner (1990), Váňa (1991c), Álvaro-A (2011). Penang, Pahang, Johor (“Mt. Ophir” (Gunung Ledang) was mistakenly listed as in Malacca), Sarawak, Sabah.

Anastrophyllumprionophyllum S.Hatt. – Kitagawa (1970), Váňa (1991c). Sabah.

Anastrophyllumrevolutum Steph. – Kitagawa (1970), Váňa (1991c), Cheah (2017). Pahang, Sabah.

Anastrophyllumsquarrosum Herzog – Kitagawa (1970), Váňa (1991c), Cheah (2017). Pahang, Sabah.

Andrewsianthusbidens (Steph.) R.M.Schust. – Cheah (2017). Pahang.

Andrewsianthuschimbuensis R.M.Schust. – Kürschner (1990), Váňa (1991c). Sabah.

Andrewsianthuskinabaluensis N.Kitag. – Kitagawa (1970), Váňa (1974, 1991c). Sabah.

Andrewsianthusmizutanii N.Kitag. – Amakawa (1969a as Jamesoniellaminutissima), Kitagawa (1969d), Váňa (1974, 1991c). Sabah.

Andrewsianthuspapillosus N.Kitag. – Kitagawa (1970), Kitagawa and Kodama (1973), Váňa (1974, 1991c). Sabah.

Andrewsianthuspuniceus (Nees) Grolle – Kitagawa (1970), Pocock et al. (1984), Váňa (1991c). Pahang, Sabah.

Andrewsianthusrecurvifolius (Nees) R.M.Schust. – Váňa (1991c). Sabah.

Aneuramaxima (Schiffn.) Steph. – Kitagawa and Kodama (1974). Sabah.

Aneurapinguis (L.) Dumort. – Johnson (1972 as Riccardiapinguis). Pahang.

Aphanotropissaxicola Herzog – Herzog (1952a). Sarawak. (synonym of Colura?, see Gradstein and Benitez 2014, Lee and Gradstein 2021).

Asterellalimbata D.G.Long et Grolle – Long and Grolle (1994), Long (2006). Sabah.

Balantiopsisciliaris S.Hatt. – Hattori (1966), Kitagawa and Kodama (1973). Sabah.

Bazzaniaalbifolia Horik. – Cheah and Yong (2016). Pahang.

Bazzaniaangustistipula N.Kitag. – Cheah and Yong (2016). Pahang.

Bazzaniaasymmetrica (Steph.) N.Kitag. – Cheah and Yong (2016). Pahang.

Bazzaniabicrenata N.Kitag. – Cheah and Yong (2016). Pahang.

Bazzaniabidentula (Steph.) Yasuda – Cheah and Yong (2016). Pahang.

Bazzaniabilobata N.Kitag. – Pócs et al. (2014). Pahang.

Bazzaniaborneensis N.Kitag. – Stephani (1908 as as Mastigobryumborneense), Kitagawa (1973). Sarawak.

Bazzaniacalcarata (Sande Lac.) Schiffn. – Stephani (1908 as Mastigobryumcalcaratum), Herzog (1950 as B.richardsii), Meijer (1960), Tixier (1971). Pahang, Sarawak.

Bazzaniacaudata (Steph.) Herzog – Stephani (1908 as Mastigobryumcaudatum), Herzog (1950), Tixier (1971), Kitagawa and Kodama (1973). Penang, Pahang, Sarawak, Sabah.

Bazzaniaceylanica (Mitt.) Steph. – Kamimura (1975). Sabah.

Bazzaniacincinnata (De Not.) Trevis. – De Notaris (1874 as Mastigobryumcincinnatum), Schiffner (1898), Stephani (1908 as M.cincinnatum), Kamimura (1975). Sarawak, Sabah.

Bazzaniacommutata (Lindenb. et Gottsche) Schiffn. – Tixier (1971). Pahang.

Bazzaniaconfertifolia (Steph.) Herzog – Herzog (1950). Sarawak.

Bazzaniaconophylla (Sande Lac.) Schiffn. – Kitagawa and Kodama (1973), Kitagawa (1973). Sabah.

Bazzaniadensa (Sande Lac.) Schiffn. – Kitagawa (1971), Kitagawa and Kodama (1973), Kürschner (1990). Penang, Sabah.

Bazzaniadiminuta Herzog – Herzog (1950). Sarawak.

Bazzaniadistans (Nees) Trevis. – Johnson (1972). Pahang.

Bazzaniadrepanophylla Herzog – Herzog (1950). Sarawak.

Bazzaniadulitensis Herzog – Herzog (1950), Kitagawa and Kodama (1973), Kürschner (1990). Sarawak, Sabah.

Bazzaniaerosa (Reinw., Blume et Nees) Trevis. – De Notaris (1874 as Mastigobryumvagum), Schiffner (1898), Stephani (1908 as M.vagum), Herzog (1950 as B.vaga and B.serrulata), Kitagawa and Kodama (1973), Kitagawa (1977), Cheah and Yong (2016). Penang, Pahang, Sarawak, Sabah.

Bazzaniaerosa var. pulopenangensis (Lindenb. et Gottsche) Schiffn. – Lindenberg and Gottsche (1851 as Mastigobryumerosum δ pulopenangense), Schiffner (1898). Penang.

Bazzaniafallax (Sande Lac.) Schiffn. – De Notaris (1874 as Mastigobryumborneense), Schiffner (1898), Herzog (1950 as B.fallaxf.fissa), Meijer (1960). Sarawak.

Bazzania fleischeri (Steph.) Abeyw. – Stephani (1908, 1909 both as Mastigobryumfleischeri). Sarawak.

Bazzaniafriabilis N.Kitag. et T.Kodama – Kitagawa and Kodama (1973, 1975), Kürschner (1990), Cheah and Yong (2016). Pahang, Sabah.

Bazzaniagrandiretis (Steph.) Herzog – Herzog (1950), Kitagawa and Kodama (1973). Sarawak, Sabah.

Bazzaniaharpago (De Not.) Schiffn. – De Notaris (1874 as Mastigobryumferox and M.harpago), Schiffner (1898), Stephani (1908 as M.harpago), Herzog (1950, 1952b), Kitagawa (1972), Kitagawa and Kodama (1973), Kamimura (1975, also as B.ferox), Kürschner (1990), Ludwiczuk and Asakawa (2010), Ng et al. (2021). Sarawak, Sabah.

Bazzaniaherzogiana Meijer – Herzog (1950 as B.remotifolia). Sarawak.

Bazzaniahorridula Schiffn. – Herzog (1932 as Mastigobryummuricatulum, 1950), Kitagawa and Kodama (1973), Kürschner (1990), Cheah and Yong (2016). Pahang, Sarawak, Sabah.

Bazzaniainaequitexta Steph. – Stephani (1908, 1909 both as Mastigobryuminaequitextum). Sarawak.

Bazzaniaindica (Gottsche et Lindenb.) Trevis. – Kamimura (1975), Kürschner (1990). Sabah.

Bazzaniainsignis (De Not.) Trevis. – De Notaris (1874 as Mastigobryuminsigne), Schiffner (1898), Evans (1932), Kamimura (1975). Sarawak, Sabah.

Bazzaniaintermedia (Gottsche et Lindenb.) Trevis. – De Notaris (1874 as Mastigobryumconcinnum), Schiffner (1898 as B.concinna), Stephani (1908 as M.concinnum), Herzog (1950 as B.concinna), Meijer (1960), Kitagawa (1969c), Tixier (1971), Kitagawa and Kodama (1973), Kamimura (1975), Grolle (1987). Penang, Pahang, Sarawak, Sabah.

Bazzaniaintermedia var. sarawakiana (De Not.) Schiffn. – De Notaris (1874 as Mastigobryumintermediumvar.sarawakianum), Schiffner (1898). Sarawak.

Bazzaniainvolutiformis (De Not.) Trevis. – De Notaris (1874 as Mastigobryumduplex and M.involutiforme), Schiffner (1898), Herzog (1950, 1952b), Kitagawa (1972), Kitagawa and Kodama (1973), Kamimura (1975), Kürschner (1990). Sarawak, Sabah.

Bazzaniakokawana N.Kitag. et T.Kodama – Kitagawa and Kodama (1973). Sabah.

Bazzanialinearis Herzog – Herzog (1950). Sarawak.

Bazzanialinguiformis (Sande Lac.) Trevis. – Stephani (1908 as Mastigobryumlinguiforme [‘linguaeforme’]). Penang.

Bazzanialongicaulis (Sande Lac.) Schiffn. – Herzog (1950), Johnson (1972), Kitagawa and Kodama (1973), Kitagawa (1977), Kürschner (1990). Pahang, Sarawak, Sabah.

Bazzanialoricata (Reinw., Blume et Nees) Trevis. – Stephani (1908 as Mastigobryumloricatum), Herzog (1950), Inoue (1967a, 1968b), Pocock et al. (1984). Pahang, Sarawak.

Bazzanialowii (Steph.) Schiffn. – Stephani (1886, 1908 both as Mastigobryumlowii), Schiffner (1898), Evans (1932 as M.lowii). Sabah.

Bazzaniamalaccensis (Steph.) Tixier – Stephani (1908 as Mastigobryummalaccense), Tixier (1971, 1974). Kedah, Perak, Pahang.

Bazzaniamanillana (Steph.) Steph. – Stephani (1917 as Mastigobryumcrenatistipulum), Kamimura (1975), Mizutani (1967a). Penang, Sabah.

Bazzaniamarginella (Herzog) N.Kitag. et T.Kodama – Kitagawa and Kodama (1973). Sabah.

Bazzaniamenzelii E.D.Cooper – Herzog (1950 as Acromastigumemarginatum). Sarawak.

Bazzaniaparadoxa (Sande Lac.) Steph. – Stephani (1908 as Mastigobryumnatunense), Meijer (1960), Kitagawa and Kodama (1973). Pahang, Sarawak, Sabah.

Bazzaniaparvitexta Steph. – Kitagawa (1979a). Sabah.

Bazzaniapatentistipa (Sande Lac.) Schiffn. – Kitagawa and Kodama (1973 [‘patentistipula’]). Sabah.

Bazzaniapectinata (Lindenb. et Gottsche) Schiffn. – Herzog (1950, 1952b), Tixier (1971), Kitagawa and Kodama (1973), Pocock et al (1984). Pahang, Sarawak, Sabah.

Bazzaniapraerupta (Reinw., Blume et Nees) Trevis. – Kitagawa and Kodama (1973), Kürschner (1990), Ludwiczuk and Asakawa (2010). Sabah.

Bazzaniapseudovittata N.Kitag. et T.Kodama – Kitagawa and Kodama (1973, 1975), Kürschner (1990), Cheah and Yong (2016). Pahang, Sabah.

Bazzaniarecurva (Mont.) Trevis. – Montagne (1843 as Herpetiumrecurvum, 1856 as Mastigobryumrecurvum), Gottsche et al. (1845 as M.recurvum), Lindenberg and Gottsche (1851 as M.recurvum), De Notaris (1874 as M.recurvumvar.pallens), Schiffner (1898 as B.recurvavar.pallens), Stephani (1908 as M.recurvum), Inoue (1968b). Penang, Pahang, Sarawak.

Bazzaniarevoluta (Steph.) N.Kitag. – Kamimura (1975 as B.recurvolimbata). Sabah.

Bazzaniaserpentina (Nees) Trevis. – Cheah and Yong (2016). Pahang.

Bazzaniasikkimensis Schiffn. – Kamimura (1975). Sabah.

Bazzaniaspiralis (Reinw., Blume et Nees) Meijer – Meijer (1960), Kitagawa and Kodama (1973), Tixier (1971, 1974), Kamimura (1975), Kürschner (1990), Ludwiczuk and Asakawa (2010). Kedah, Pahang, Sarawak, Sabah.

Bazzaniasubtilis (Sande Lac.) Trevis. – De Notaris (1874 as Mastigobryumsubtile), Stephani (1917 as M.repandistipulum), Meijer (1960), Kitagawa and Kodama (1973), Kitagawa (1977, 1979 as B.palmatifida), Kürschner (1990), Wolseley et al. (1996). Negeri Sembilan, Sarawak, Sabah.

Bazzaniasumbavensis (Steph.) Steph. – Kamimura (1975). Sabah.

Bazzaniatridens (Reinw., Blume et Nees) Trevis. – Schiffner (1898 as B.australis), Herzog (1950, also as B.coreana), Kitagawa (1969c, 1972, also as B.australis), Tixier (1971), Kitagawa and Kodama (1973, also as B.australis), Kamimura (1975, also as B.tridensf.minutissima), Kürschner (1990). Penang, Pahang, Sarawak, Sabah.

Bazzaniatridens var. assamica (Steph.) Pócs – Kamimura (1975), Pócs et al. (2014 as B.assamica). Pahang, Sabah.

Bazzaniauncigera (Reinw., Blume et Nees) Trevis. – Herzog (1950), Kitagawa and Kodama (1973), Kürschner (1990), Cheah and Yong (2016). Pahang, Sarawak, Sabah.

Bazzaniauncigera var. brevifolia Herzog – Herzog (1950). Sarawak.

Bazzaniaundulata Herzog – Herzog (1950). Sarawak.

Bazzaniavittata (Gottsche) Trevis. – Herzog (1950), Inoue (1967a), Kitagawa and Kodama (1973), Kürschner (1990). Pahang, Sarawak, Sabah.

Bazzaniavittata var. luxurians (De Not.) Schiffn. – De Notaris (1874 as Mastigobryumvittatumvar.luxurians), Schiffner (1898). Sarawak.

Bazzaniawiltensii (Steph.) Schiffn. – Pocock et al. (1984). Pahang.

Bazzaniazollingeri (Lindenb.) Trevis. – De Notaris (1874 as Mastigobryumzollingeri). Sarawak.

Blepharostomatrichophyllum (L.) Dumort. – Akiyama et al. (2001). Sabah.

Calypogeiaapiculata (Steph.) Steph. – Kitagawa and Kodama (1973). Sabah.

Calypogeiaarguta Nees et Mont – Wolseley et al. (1996), Bisang et al. (2003). Pahang, Negeri Sembilan.

Caudalejeuneacristiloba (Steph.) Gradst. – Verdoorn (1934a, 1937 both as C.circinata), Mizutani (1966, 1969 both as C.circinata, 1988), Kitagawa and Kodama (1974 as C.circinata). Johor, Sarawak, Sabah.

Caudalejeuneareniloba (Gottsche) Steph. – Stephani (1912 as C.serrata), Verdoorn (1934a), Herzog (1952b), Mizutani (1966, 1969, also as C.stephanii, 1988, also as C.recurvistipula), Tixier (1971 as C.stephanii, 1980a), Kitagawa and Kodama (1974), Thiers (1985), Pócs et al. (2020), Pesiu et al. (2021, in press). Penang, Perak, Selangor, Pahang, Malacca, Terengganu, Johor, Sarawak, Sabah.

Cephalolejeuneaparvilobula Mizut. – Mizutani (1979b). Sabah.

Cephaloziaacuminata (Herzog) Váňa – Herzog (1950 as Metahygrobiellaacuminata), Váňa (1993 as M.acuminata). Sarawak.

Cephaloziaacutiloba (Inoue) Váňa – Inoue (1967a,b, both as Metahygrobiellaacutiloba). Pahang.

Cephaloziahamatiloba Steph. – Müller and Schäfer-Verwimp (1999), Cheah (2017). Pahang.

Cephaloziamollusca (De Not.) Váňa – De Notaris (1874 as Jungermanniamollusca), Schiffner (1898 as Hygrobiellamollusca), Stephani (1908 as H.mollusca), Herzog (1950 as Metahygrobiellamollusca), Kitagawa and Kodama (1974 as M.mollusca), Váňa (1993 as M.mollusca). Sarawak, Sabah.

Cephaloziastolonacea (Herzog) Váňa – Herzog (1950 as Metahygrobiellastolonacea and Hygrobiellamolluscaf.subintegerrima), Grolle (1963 as M.stolonacea), Váňa (1993 as M.stolonacea). Sarawak.

Cephaloziellacapillaris (Steph.) Douin – Inoue (1967a), Inoue and Miller (1967). Pahang.

Cephaloziellakiaeri (Austin) Douin – Kitagawa and Kodama (1974 as C.willisana), Váňa (1992). Sabah.

Cephaloziellamicrophylla (Steph.) Douin – Kitagawa (1969c). Penang.

Ceratolejeuneaaliena Herzog – Herzog (1952b), Kitagawa and Kodama (1974), Mizutani (1981a). Sarawak, Sabah.

Ceratolejeuneabelangeriana (Gottsche) Steph. – Kitagawa and Kodama (1974, also as C.ryukyuensis), Mizutani (1981a as C.oceanica), Zhu et al. (2005), Cheah (2017). Pahang, Johor, Sabah.

Ceratolejeuneacornuta (Lindenb.) Steph. – Inoue (1967a as C.tahitensis). Pahang.

Ceratolejeuneaminor Mizut. – Mizutani (1981a), Zhu et al. (2005), Sarimi et al. (2021). Terengganu, Sabah.

Ceratolejeuneamoniliata Herzog – Herzog (1952b), Mizutani (1970, 1981a), Pesiu et al. (in press). Terengganu, Johor, Sarawak, Sabah.

Ceratolejeuneasingapurensis (Lindenb.) Steph. – Herzog (1952b), Cheah (2017), Sarimi et al. (2021). Pahang, Terengganu, Sarawak.

Ceratolejeuneaspinistipula (Herzog) R.L.Zhu, L.Shu, Xiong He et Y.M.Wei – Herzog (1948 as Drepanolejeuneaspinistipula), Pócs et al. (1995 as Acantholejeuneaspinistipula), Zhu et al. (2018a as D.spinistipula). Johor, Sarawak, Sabah.

Cheilolejeuneaadnata (Lehm.) Grolle var. autoica Gradst. et Ilk.-Borg. – Shu et al. (2015 as C.larsenii). Penang.

Cheilolejeuneaceylanica (Gottsche) R.M.Schust. et Kachroo – Schiffner (1898 as Pycnolejeuneaceylanica), Mizutani (1966, 1970, 1980), Tixier (1972, 1974 both as P.arietina, 1980a as Xenolejeuneaceylanica), Kitagawa and Kodama (1974), Grolle (1979), Kürschner (1990), Wolseley et al. (1996), Zhu and So (2001), Pócs et al. (2020), Pesiu et al. (2021). Kedah, Penang, Pahang, Negeri Sembilan, Malacca, Terengganu, Sabah.

Cheilolejeuneadecursiva (Sande Lac.) R.M.Schust. – Mizutani (1970 as C.spathulata), Grolle (1977), Zhu and Lai (2005), Shu et al. (2014), Cheah (2017). Pahang, Sabah.

Cheilolejeuneafalsinervis (Sande Lac.) R.M.Schust. et Kachroo – Mizutani (1966, 1970), Kitagawa and Kodama (1974), Tixier (1974 as Xenolejeuneafalsinervis), Thiers (1986), Kürschner (1990), Wolseley et al. (1996). Kedah, Negeri Sembilan, Sabah.

Cheilolejeuneagigantea (Steph.) R.M.Schust. et Kachroo – Herzog (1950 as Pycnolejeuneagigantea). Sarawak.

Cheilolejeuneaincisa (Gottsche) R.M.Schust. et Kachroo – Hoffmann (1935 as Pycnolejeuneaexcisula), Mizutani (1970 as C.excisula), Tixier (1971 as P.excisula), Kamimura (1974 as C.excisula), Kitagawa and Kodama (1974 as C.excisula), Grolle and Piippo (1990 as C.excisula). Pahang, Johor, Sabah.

Cheilolejeuneainsignis Jovet-Ast et Tixier – Kitagawa and Kodama (1974). Sabah.

Cheilolejeuneaintertexta (Lindenb.) Steph. – Mizutani (1967b, 1970), Kitagawa (1971), Grolle (1979), Kürschner (1990), Pócs et al. (2020). Penang, Sarawak, Sabah.

Cheilolejeuneakrakakammae (Lindenb.) R.M.Schust. – Cheah (2017). Pahang.

Cheilolejeunealindenbergii (Gottsche) Mizut. – Herzog (1950 as Hygrolejeunealindenbergii and Euosmolejeunealuerssenii), Inoue (1967 as C.luerssenii [‘luersonii’]), Kitagawa (1969a as C.luerssenii, 1971), Mizutani (1970), Kitagawa and Kodama (1974), Kürschner (1990), Cheah (2017). Penang, Pahang, Sarawak, Sabah.

Cheilolejeunealoriana (Steph.) W.Ye et R.L.Zhu – Mizutani (1970 as Leucolejeunealoriana). Sabah.

Cheilolejeuneamalaccensis (G.Hoffm.) Xiao L.He – Hoffmann (1935 as Pycnolejeuneamalaccensis), Thiers (1986 as P.malaccensis), He (1996). Penang.

Cheilolejeuneamicholitzii (Steph.) R.M.Schust. et Kachroo – Hoffmann (1935 as Pycnolejeuneamicholitzii). Johor.

Cheilolejeuneamizutanii W.Ye et R.L.Zhu – Mizutani (1976 as Leucolejeuneadecurrens), Grolle and Piippo (1990 as L.decurrens). Sabah.

Cheilolejeuneanipponica (S.Hatt.) S.Hatt. – Pócs and Lee (2016). Selangor.

Cheilolejeuneaocclusa (Herzog) T.Kodama et N.Kitag. – Herzog (1950 as Strepsilejeuneaocclusa), Kitagawa and Kodama (1974), Mizutani (1980), Wolseley et al. (1996), Pócs et al. (2020). Negeri Sembilan, Sarawak, Sabah.

Cheilolejeuneaorientalis (Gottsche) Mizut. – Kitagawa and Kodama (1974). Sabah.

Cheilolejeuneaserpentina (Mitt.) Mizut. – Hashimoto et al. (1993), Cheah (2017). Langkawi, Pahang.

Cheilolejeuneastreimannii Pócs et Ninh – Pócs and Lee (2016). Pahang.

Cheilolejeuneatrapezia (Nees) R.M.Schust. et Kachroo – Stephani (1890 as Lejeuneaimbricata), Hoffmann (1935 as Pycnolejeuneaimbricata, P.meyenianaand itsvar.ligulata, C.longiloba), Herzog (1948 as Placolejeuneasubarrhyncha), Mizutani (1966, 1970 both as C.imbricata, 1980 as C.longiloba), Inoue (1967a as C.imbricata), Kitagawa (1971 as C.imbricata), Tixier (1971, 1974, 1980a as Xenolejeunealongiloba and X.meyeniana), Kitagawa and Kodama (1974 as C.imbricata and C.meyeniana), Grolle (1979), Thiers (1986 as C.meyeniana), Kürschner (1990 as C.imbricata), Cheah (2017 as C.imbricata), Pócs et al. (2020, also as C.meyeniana), Pesiu et al. (2021). Kedah, Penang, Pahang, Terengganu, Johor, Sarawak, Sabah.

Cheilolejeuneatrifaria (Reinw., Blume et Nees) Mizut. – Inoue (1967a as C.viridula), Mizutani (1970, 1982), Kamimura (1974), Kitagawa and Kodama (1974), Hashimoto et al. (1993), Lee et al. (2013), Pócs et al. (2020), Pesiu et al. (2021). Langkawi, Perak, Pahang, Terengganu, Sabah.

Cheilolejeuneaventricosa (P.Syd.) Xiao L.He – Hoffmann (1935 as Pycnolejeuneaventricosa), Mizutani (1970 as P.ventricosa), Hattori and Kamimura (1971 as P.ventricosa), Zhu and Lai (2005), Pócs et al. (2020). Penang, Sabah.

Cheilolejeuneaverrucosa Steph. – Tixier (1974), Mizutani (1979b). Kedah, Sabah.

Cheilolejeuneavittata (G.Hoffm.) R.M.Schust. et Kachroo – Mizutani (1970, 1980), Kürschner (1990), Wolseley et al. (1996), Pócs et al. (2020). Negeri Sembilan, Sabah.

Cheilolejeuneaxanthocarpa (Lehm. et Lindenb.) Malombe – Verdoorn (1934a as Leucolejeuneaxanthocarpa), Inoue (1967a as L.xanthocarpa), Tixier (1971 as L.xanthocarpa), Kamimura (1975 as L.xanthocarpa). Pahang, Sabah.

Chiastocaulonbraunianum (Nees) S.D.F.Patzak, M.A.M Renner, Schäf.-Verw. et Heinrichs – Kitagawa and Kodama (1974 as Plagiochilionbraunianum), Akiyama et al. (2001 as P.braunianum). Sabah.

Chiastocaulondendroides (Nees) Carl – De Notaris (1874 as Plagiochiladendroides), Schiffner (1898 as P.dendroides), Herzog (1950 as P.dendroides), Kitagawa and Kodama (1974 as P.dendroides), Inoue (1967a, 1968b, 1984 as P.dendroides), Kürschner (1990 as P.dendroides), So (2001 as P.dendroides), Akiyama et al. (2001 as P.dendroides). Pahang, Sarawak, Sabah.

Chiastocaulonoppositum (Reinw., Blume et Nees) S.D.F.Patzak, M.A.M Renner, Schäf.-Verw. et Heinrichs – Stephani (1904 as Plagiochilaopposita), Hattori (1942a as Plagiochilaopposita, 1950), Inoue (1964, 1967a both as Plagiochilionoppositum), Kitagawa and Kodama (1974 as Plagiochilionoppositum), Kamimura (1975 as Plagiochilionoppositum), Kürschner (1990 as Plagiochilionoppositum). Perak, Pahang, Sarawak, Sabah.

Chiastocaulonpachycephalum (De Not.) Herzog – De Notaris (1874 as Plagiochilapachycephala), Schiffner (1898 as Plagiochilapachycephala), Herzog (1950 as Plagiochilionpachycephalum), Inoue (1964 as Plagiochilionpachycephalum), So and Grolle (2000 as Plagiochilionpachycephalum), So (2001 as Plagiochilapachycephala). Sarawak.

Chiastocaulontheriotianum (Steph.) S.D.F.Patzak, M.A.M Renner, Schäf.-Verw. et Heinrichs – Inoue (1964 as Plagiochiliontheriotianum). Sarawak.

Cololejeuneaabnormis Mizut. – Mizutani (1970), Tixier (1985). Penang, Sabah.

Cololejeuneaacuminata Mizut. – Mizutani (1970), Wolseley et al. (1996), Pócs et al. (2020). Negeri Sembilan, Sabah.

Cololejeuneaaequabilis (Sande Lac.) Schiffn. – Benedix (1953 as C.yulensis), Mizutani (1966 as C.yulensis), Kitagawa and Kodama (1974 as C.yulensis), Pesiu et al. (2021). Terengganu, Johor, Sarawak, Sabah.

Cololejeuneaamphibola B.M.Thiers – Eggers et al. (1998), Pócs and Lee (2016). Selangor, Kelantan, Pahang, Sarawak.

Cololejeuneaangulata (Steph.) Mizut. – Kitagawa (1969a, 1969c), Wolseley et al. (1996), Mizutani (1966). Penang, Negeri Sembilan, Sabah.

Cololejeuneaangustiflora (Steph.) Mizut. – Mizutani (1966, 1970 both as C.javanica and C.crenulata), Kitagawa and Kodama (1974 as C.crenulata), Tixier (1985 as C.javanica), Zhu and So (2000a as C.mackeeana, 2002). Penang, Sarawak, Sabah.

Cololejeuneaappressa (A.Evans) Benedix – Mizutani (1966, 1970), Sarimi et al. (2021). Terengganu, Sabah.

Cololejeuneabidentula (Steph.) E.W.Jones – Pócs and Lee (2016). Selangor.

Cololejeuneaceatocarpa (Ångstr.) Steph – Pócs and Lee (2016). Kelantan, Pahang.

Cololejeuneaceratilobula (P.C.Chen) R.M.Schust. – Mizutani (1966, 1970 both as C.formosana), Pesiu et al. (in press). Terengganu, Sabah.

Cololejeuneachamlongiana Tixier – Tixier (1985). Perak.

Cololejeuneachinii Tixier – Tixier (1973b), Kitagawa (1981). Selangor, Kelantan.

Cololejeuneacordiflora Steph. – Tixier (1971 as C.karstenii, 1985). Perak, Penang, Pahang.

Cololejeuneadecliviloba Steph. – Tixier (1980a). Pahang.

Cololejeuneadesciscens Steph. – Eggers et al. (1998). Sarawak.

Cololejeuneadiaphana A.Evans – Eggers et al. (1998 as Aphanolejeuneaproboscoidea), Pócs and Lee (2016). Kelantan, Sarawak.

Cololejeuneadilatata (Steph.) Mizut. – Mizutani (1966, 1970). Sabah.

Cololejeuneadozyana (Sande Lac.) Schiffn. – Benedix (1953), Tixier (1980a), Pócs et al. (2020). Pahang, Sabah.

Cololejeuneaensifera Tixier – Pócs et al. (2020), Sarimi et al. (2021). Sabah.

Cololejeuneaequialbi Tixier – Pócs et al. (2020), Sarimi et al. (2021), Pesiu et al (in press). Terengganu, Sabah.

Cololejeuneafalcata (Horik.) Benedix – Mizutani (1966, 1970 both as C.falcatoides), Tixier (1971, 1974 both as C.falcatoides, 1978, 1980a), Kitagawa and Kodama (1974 as C.falcatoides), Pesiu et al. (2021, in press). Kedah, Perak, Penang, Pahang, Terengganu, Sabah.

Cololejeuneafilidens Benedix – Tixier (1985). Perak, Selangor.

Cololejeuneafissilobula Herzog – Herzog (1950). Sarawak.

Cololejeuneafloccosa (Lehm. et Lindenb.) Schiffn. – Benedix (1953), Kitagawa (1969c), Mizutani (1966, 1970, 1984a), Tixier (1971, 1974, 1978), Kitagawa and Kodama (1974), Wolseley et al. (1996), Pesiu et al. (2021, in press). Kedah, Penang, Perak, Pahang, Negeri Sembilan, Terengganu, Sarawak, Sabah.

Cololejeuneafloccosa var. amoena (Benedix) Pócs – Benedix (1953 as C.amoena), Mizutani (1966, 1970, both as C.amoena). Johor, Sabah.

Cololejeuneafloccosa var. aurita Benedix – Benedix (1953), Tixier (1978). Kedah, Penang, Perak, Johor.

Cololejeuneafloccosa var. trivittata Tixier – Eggers et al. (1998 as C.floccosavar.vittata), Crosby and Engel (2006). Pahang, Sarawak.

Cololejeuneafructumarginata Tixier – Tixier (1985). Penang.

Cololejeuneageorgiana Tixier – Tixier (1985). Penang.

Cololejeuneagottschei (Steph.) Pandé, K.P.Srivast. et Ahmad – Mizutani (1966 as C.paroica), Pócs et al. (2020), Sarimi et al. (2021), Pesiu et al. (in press). Kelantan, Terengganu, Sabah.

Cololejeuneagradsteinii M.J.Lai et R.L.Zhu – Tixier (1971 as C.pusilla). Pahang.

Cololejeuneagynophthalma Benedix – Tixier (1971, 1978), Mizutani (1966, 1970). Penang, Pahang, Sabah.

Cololejeuneahaskarliana (Lehm.) Schiffn. – Benedix (1953), Mizutani (1966, 1970, 1986a), Kitagawa (1971), Kitagawa and Kodama (1974), Tixier (1980a, 1985), Wolseley et al. (1996), Pócs et al. (2020). Penang, Perak, Pahang, Negeri Sembilan, Johor, Sarawak, Sabah.

Cololejeuneahattoriana Mizut. et Pócs – Mizutani (1966 as Campylolejeuneapusilla). Sabah.

Cololejeuneahildebrandii (Austin) Steph. – Tixier (1980a as C.filicaulis, 1985), Pócs et al. (2014, 2020). Penang, Perak, Selangor, Pahang, Sabah.

Cololejeuneahirta Steph. – Tixier (1985). Perak, Pahang.

Cololejeuneaindica Pandé et R.N.Misra – Tixier (1985 as C.planiflora). Penang.

Cololejeuneainflata Steph. – Benedix (1953 as C.oshimensis), Mizutani (1966, 1970 both as C.oshimensis), Tixier (1971, 1974 both as C.oshimensis, 1978, 1980a), Kitagawa and Kodama (1974 as C.oshimensis), Pesiu et al. (2021). Kedah, Pahang, Terengganu, Johor, Sabah.

Cololejeuneainflectens (Mitt.) Benedix – Benedix (1953 as C.ciliatilobula and C.peculiaris), Mizutani (1970 as Campylolejeuneapeculiaris), Tixier (1971 as C.ciliatilobula, 1974 as Campylolejeuneapeculiaris, 1980a as C.inflectidens), Pesiu et al. (2021). Kedah, Pahang, Terengganu, Johor, Sarawak, Sabah.

Cololejeuneajohannis-winkleri (Herzog) R.L.Zhu – Zhu et al. (2004). Sabah.

Cololejeuneakulenensis Tixier – Tixier (1985). Johor.

Cololejeunealacinulata Benedix – Benedix (1953). Sabah.

Cololejeunealanciloba Steph. – Mizutani (1966, 1984b), Kitagawa (1971), Tixier (1971 as C.pahangiana, 1985 as C.bolombensis), Kitagawa and Kodama (1974), Pócs et al. (2020), Pesiu et al. (2021, in press). Penang, Pahang, Terengganu, Johor, Sabah.

Cololejeunealatilobula (Herzog) Tixier – Pócs and Lee (2016). Kelantan.

Cololejeunealongifolia (Mitt.) Mizut. – Tixier (1984, 1985), Pócs et al. (2020). Penang, Johor, Sabah.

Cololejeuneamacounii (Spruce) A.Evans – Mizutani (1966, 1970 both as C.kinabalensis), Pócs et al. (2020). Sabah.

Cololejeuneamadothecoides (Steph.) Benedix. – Benedix (1953). Sarawak.

Cololejeuneamagnilobula (Horik.) S.Hatt. – Pócs et al. (2020). Sabah.

Cololejeuneamagnilobula var. falcidentata Pócs et G.E.Lee – Pócs and Lee (2016). Kelantan.

Cololejeuneamalaccensis Tixier – Tixier (1985). Johor.

Cololejeuneamalayana Tixier – Tixier (1980a, 1985), Eggers (2006). Perak, Pahang.

Cololejeuneamaritima Tixier – Tixier (1979, 1985). Perak, Johor.

Cololejeuneametzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilk.-Borg. et Heinrichs – Mizutani (1966 as Metzgeriopsispusilla), Kitagawa (1969b as M.pusilla), Kitagawa and Kodama (1974 as M.pusilla), Tixier (1980a as M.pusilla), Akiyama et al. (2001 as M.pusilla), Gradstein et al. (2006), Pócs and Piippo (2011), Pócs et al. (2020), Pesiu et al. (2021). Perak, Pahang, Terengganu, Sabah.

Cololejeuneamutabilis Benedix – Benedix (1953 as C.mutabilisf.borneensis), Tixier (1978 as C.mutabilisf.borneensis,f.floccodoidesandf.subfalcata), Mizutani (1966), Pócs et al. (2020). Perak, Sabah.

Cololejeuneaobliqua (Nees et Mont.) Schiffn. – Benedix (1953 as C.androgyna, C.nymannii), Mizutani (1966 as C.jelinekii and C.nymannii, 1970 as C.nymannii), Tixier (1971 as C.jelinekii, 1980a, 1985 both as C.scabriflora), Kitagawa and Kodama (1974 as C.nymannii), Asthana and Srivastava (2003 as C.jelinekii), Pócs et al. (2020), Pesiu et al. (2021). Penang, Perak, Pahang, Terengganu, Sarawak, Sabah.

Cololejeuneaocellata (Horik.) Benedix – Pócs and Lee (2016). Pahang.

Cololejeuneaocelloides (Horik.) Mizut. – Benedix (1953 as C.leonidensand itsvar.saccata), Mizutani (1966, 1970 both as C.leonidens, 1984a), Inoue (1967a as C.leonidens), Tixier (1969 as C.meijeri, 1971, 1978, 1980a all as C.leonidens), Kitagawa and Kodama (1974 as C.leonidens), Pesiu et al. (2021). Perak, Pahang, Terengganu, Sabah.

Cololejeuneapacifica Pócs – Pócs and Lee (2016). Kelantan.

Cololejeuneapapillosa (K.I.Goebel) Mizut. – Herzog (1950 as Aphanolejeuneamicroscopicavar.borneensis), Mizutani (1966), Pócs and Piippo (1999 as A.borneensis), Pócs and Bernecker (2009), Pócs and Lee (2016), Pócs et al. (2020). Pahang, Sarawak, Sabah.

Cololejeuneapeponiformis Mizut. – Mizutani (1970), Pócs et al. (2020). Sabah.

Cololejeuneaperaffinis (Schiffn.) Schiffn. – Herzog (1952b as Taeniolejeuneaperaffinis), Benedix (1953), Mizutani (1966, 1970, 1984a), Kitagawa (1969a), Kitagawa and Kodama (1974), Tixier (1978 as C.peraffinisvar.elegans, 1980a). Penang, Pahang, Johor, Sarawak, Sabah.

Cololejeuneaperakensis Tixier – Tixier (1985). Perak.

Cololejeuneaplanissima (Mitt.) Abeyw. – Mizutani (1970, 1984b), Kitagawa (1971), Kitagawa and Kodama (1974), Tixier (1985), Asthana and Srivastava (2003), Pócs et al. (2020), Pesiu et al. (2021). Penang, Perak, Terengganu, Johor, Sabah.

Cololejeuneaplatyneura (Spruce) A.Evans – Mizutani (1966 as C.astyla), Zhu and So (1998b, 2001), Cheah (2017). Pahang, Sabah

Cololejeuneapretiosa Benedix – Kitagawa and Kodama (1974). Sabah.

Cololejeunea pseudofloccosa (Horik.) Benedix – Benedix (1953), Mizutani (1966, 1970), Kitagawa and Kodama (1974), Tixier (1980a). Pahang, Sabah.

Cololejeuneapseudoschmidtii Tixier – Tixier (1985). Penang.

Cololejeuneapseudostephanii Tixier – Pócs and Lee (2016). Pahang.

Cololejeuneapseudostipulata P.Syd. – Mizutani (1970). Sabah.

Cololejeunearaduliloba Steph. – Mizutani (1966), Pócs et al. (2020). Sabah.

Cololejeuneareineckeana Steph. – Tixier (1985). Perak.

Cololejeunearosellata Mizut. – Mizutani (1966), Kitagawa and Kodama (1974), Tixier (1985). Sabah.

Cololejeuneaschmidtii Steph. – Mizutani (1966, 1970 as C.benedixii), Kitagawa and Kodama (1974), Cheah (2017), Sarimi et al. (2021), Pesiu et al. (in press). Pahang, Terengganu, Sabah.

Cololejeuneaselangorensis Tixier – Tixier (1985). Selangor.

Cololejeuneaserrulata Steph. – Kitagawa (1971). Penang.

Cololejeuneasetosa Mizut. – Mizutani (1966), Eggers et al. (1998). Sarawak, Sabah.

Cololejeuneashimizui N.Kitag. – Kitagawa (1969c, 1981). Kelantan.

Cololejeuneasiamensis Steph. – Benedix (1953 as C.pluriplicata, orth. var. of C.pluripunctata), Mizutani (1966, 1970 both as C.pluripunctata), Tixier (1978), Wolseley et al. (1996). Penang, Negeri Sembilan, Johor, Sabah.

Cololejeuneasigmoidea Jovet-Ast et Tixier – Mizutani (1966), Kitagawa (1969c), Tixier (1971, 1980a), Kitagawa and Kodama (1974), Bernecker-Lücking and Morales (1999), Pesiu et al. (2021). Penang, Pahang, Terengganu, Sabah.

Cololejeuneasigmoidea var. dubia Tixier – Tixier (1985). Perak.

Cololejeuneasmitinandii Tixier – Pócs and Lee (2016). Kelantan.

Cololejeuneaspathulifolia (Steph.) H.A.Mill. – Pócs and Lee (2016). Kelantan.

Cololejeuneasphaerodonta Mizut. – Mizutani (1966), Zhu (1995), Zhu and So (2001), Pócs et al. (2020). Sabah.

Cololejeuneastephanii Benedix – Benedix (1953), Mizutani (1966, 1970), Kitagawa and Kodama (1974), Tixier (1974, 1978, 1980a), Zhu (1995), Eggers et al. (1998), Pócs et al. (2020), Pesiu et al. (2021, in press). Kedah, Perak, Pahang, Terengganu, Johor, Sarawak, Sabah.

Cololejeuneastoniana Tixier – Tixier (1976, 1985), Eggers et al. (1998). Johor, Sarawak.

Cololejeuneastylosa Steph. – Tixier (1985), Pócs and Lee (2016), Pócs et al. (2020). Penang, Kelantan, Sabah.

Cololejeuneatenella Benedix – Benedix (1953), Tixier (1974, 1978, 1985 the latter two as C.tenellavar.vittata), Wolseley et al. (1996), Pócs et al. (2020). Kedah, Penang, Perak, Negeri Sembilan, Johor, Sabah.

Cololejeuneathailandensis Tixier – Tixier (1985). Penang.

Cololejeuneatriapiculata (Herzog) Tixier – Tixier (1971, 1980a, 1985). Pahang.

Cololejeuneatrichomanis (Gottsche) Besch. – Mizutani (1966, 1970 both as C.goebelii), Kitagawa (1971 as C.goebelii), Kitagawa and Kodama (1974 as C.goebelii), Tixier (1974, 1985 as C.balansae and C.goebelii), Asthana and Srivastava (2003). Kedah, Penang, Perak, Sabah.

Cololejeuneatridentata Tixier – Tixier (1985), Eggers et al. (1998). Johor, Sarawak.

Cololejeuneauchimae Amakawa – Tixier (1985 as C.pakseana). Selangor.

Cololejeuneaveillonii Tixier – Pócs and Lee (2016). Pahang.

Cololejeuneaverrucosa Steph. – Benedix (1953), Pesiu et al. (2021, in press). Terengganu, Johor.

Cololejeuneaverrucosa var. rectispina (Herzog) Benedix – Benedix (1953), Kitagawa and Kodama (1974). Sarawak, Sabah.

Cololejeuneawightii Steph. – Stephani (1895), Kitagawa (1971, 1972), Tixier (1971, 1985). Penang, Pahang.

Coluraacroloba (Prantl) Jovet-Ast – Jovet-Ast (1954, 1967, 1976), Mizutani (1966, 1970), Kitagawa (1969a), Tixier (1971, 1980a), Kitagawa and Kodama (1974), Wolseley et al. (1996), Pócs et al. (2020), Pesiu et al. (2021, in press). Kedah, Selangor, Pahang, Negeri Sembilan, Terengganu, Johor, Sarawak, Sabah.

Coluraari (Steph.) Steph. – Herzog (1952b as C.javanica), Jovet-Ast (1976), Mizutani (1966, 1970), Sarimi et al. (2021), Pesiu et al. (in press). Perak, Terengganu, Sarawak, Sabah.

Colurabisvoluta Herzog et Jovet-Ast – Jovet-Ast (1954). Malacca.

Colurabrevistyla Herzog – Pócs and Lee (2016). Kelantan.

Coluraclementis Grolle – Jovet-Ast (1967). Sabah.

Coluraconica (Sande Lac.) K.I.Goebel – Jovet-Ast (1958, 1967), Mizutani (1966, 1970 both as C.acutifolia), Kitagawa (1969a as C.acutifolia), Kitagawa and Kodama (1974), Wolseley et al. (1996), Pócs et al. (2020), Pesiu et al. (2021, in press). Penang, Negeri Sembilan, Terengganu, Sarawak, Sabah.

Coluracorynophora (Nees, Lindenb. et Gottsche) Trevis. – Herzog (1952b as C.trialata), Mizutani (1966, 1970), Jovet-Ast (1954, 1958, 1967, 1976), Kitagawa (1969c), Wolseley et al. (1996), Pócs et al. (2020), Pesiu et al. (2021, in press). Penang, Perak, Negeri Sembilan, Terengganu, Johor, Sarawak, Sabah.

Coluracrenulata Grolle – Jovet-Ast (1967). Sabah.

Coluracristata Jovet-Ast – Jovet-Ast (1976), Sangrattanaprasert et al. (2019), Pócs et al. (2020). Penang, Sabah.

Coluracymbalifera Herzog et Jovet-Ast – Jovet-Ast (1954). Peninsular Malaysia (type locality).

Coluragaleata Jovet-Ast – Jovet-Ast (1967). Sabah.

Coluraimperfecta Steph. – Jovet-Ast (1954), Eggers et al. (1998). Pahang, Sarawak.

Colurainflata K.I.Goebel – Müller and Schäfer-Verwimp (1999). Perak.

Colurainuii Horik. – Cheah (2017), Sarimi et al. (2021), Pesiu et al. (in press). Terengganu, Pahang.

Colurakarstenii K.I.Goebel – Jovet-Ast (1954), Lee et al. (2013), Sangrattanaprasert et al. (2018). Kedah, Pahang, Johor, Sabah.

Coluraleratii (Steph.) Steph. – Jovet-Ast (1967 as C.apiculata). Sabah.

Coluramaxima Jovet-Ast – Pócs et al. (2020). Sabah.

Coluramosenii Steph. – Pócs and Lee (2016). Kelantan.

Coluraornata K.I.Goebel – Schiffner (1893 as Lejeuneaornata,1898 as Colurolejeuneaornata), Jovet-Ast (1954, 1967), Mizutani (1966), Kitagawa and Kodama (1974), Wolseley et al. (1996), Pócs et al. (2020). Penang, Pahang, Negeri Sembilan, Sarawak, Sabah.

Colurapluridentata Jovet-Ast – Mizutani (1970). Sabah.

Colurasigmoidea Sangratt., Chantanaorr. et R.L.Zhu – Sangrattanaprasert et al. (2019). Pahang.

Coluraspeciosa Jovet-Ast – Pócs and Lee (2016). Kelantan.

Colurastrophiolata Jovet-Ast – Jovet-Ast (1976). Kedah.

Colurasuperba (Mont.) Steph. – Jovet-Ast (1967), Pócs et al. (2020). Sabah.

Coluratenuicornis (A.Evans) Steph. – Jovet-Ast (1954, 1976), Mizutani (1970), Tixier (1980a), Pócs et al. (2020). Pahang, Johor, Sabah.

Coluravalida Jovet-Ast – Sangrattanaprasert et al. (2019). Pahang.

Coluraverdoornii Herzog et Jovet-Ast – Jovet-Ast (1954, 1976), Tixier (1980a), Sangrattanaprasert et al. (2019), Pócs et al. (2020). Kedah, Pahang, Johor, Sabah.

Conoscyphustrapezioides (Sande Lac.) Schiffn. – De Notaris (1874 as Diploscyphusborneensis), Herzog (1950 as C.inflexifolius), Inoue (1967a), Kitagawa and Kodama (1974), Piippo (1989), Piippo et al. (2014). Pahang, Malacca, Sarawak, Sabah.

Cryptolophocoleaciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler et Váňa – Kitagawa and Kodama (1974 as Lophocoleaciliolata), Piippo (1989 as Chiloscyphusciliolatus), Inoue (1967a, 1968b both as L.ciliolata), Cheah (2017 as C.ciliolatus). Pahang, Sabah.

Cryptolophocoleacostata (Nees) L.Söderstr. – Herzog (1950 as Lophocoleacostata), Inoue (1967a as L.costata), Kitagawa and Kodama (1974 as L.costata), Piippo (1989 as Chiloscyphuscostatus), Akiyama et al. (2001 as L.costata). Pahang, Sarawak, Sabah.

Cryptolophocolealevieri (Schiffn.) L.Söderstr. – Piippo (1989 as Chiloscyphusparoicus). Sabah.

Cryptolophocoleamassalongoana (Schiffn.) L.Söderstr. – Pocock et al. (1984 as Lophocoleamassalongoana). Pahang.

Cyathodiumfoetidissimum Schiffn. – Lang (1905). Perak.

Cyathodiumcavernarum Kunze – Lee and Gradstein (2021). Kelantan.

Cylindrocoleatagawae (N.Kitag.) R.M.Schust. – Kitagawa (1971 as Cephaloziellatagawae). Penang.

Dactylophorellamuricata (Gottsche) R.M.Schust. – Mizutani (1970 as Drepanolejeuneamuricata), Yong and Chan (2017). Perak, Sabah.

Denotarisialinguifolia (De Not.) Grolle – De Notaris (1874 as Plagiochilalinguifolia), Inoue (1968b as Jamesoniellaovifolia), Amakawa (1969a as J.flexicaulisf.affinis), Kitagawa (1970 as J.pulchra), Grolle (1971), Kitagawa and Kodama (1973), Kürschner (1990), Váňa (1991c), Akiyama et al. (2001). Pahang, Sarawak, Sabah.

Dinckleriasingularis (Schiffn.) M.A.M.Renner, Schäf.-Verw. et Heinrichs – Inoue (1989 as Plagiochilasingularis), Cheah (2017 as P.singularis). Pahang, Sabah.

Diplasiolejeuneacavifolia Steph. – Tixier (1971 as D.brachyclada, 1980a as D.javanica), Mizutani (1966 as D.javanica, 1970 as D.brachyclada), Kitagawa and Kodama (1974), Dong et al. (2012), Pócs et al. (2020). Perak, Pahang, Sabah.

Diplasiolejeuneacobrensis Steph. – Mizutani (1970 as D.incurvata). Sabah.

Diplasiolejeuneajovet-astiae Grolle – Grolle (1966), Kitagawa and Kodama (1974), Tixier (1980a), Dong et al. (2012), Pócs et al. (2020). Pahang, Sarawak, Sabah.

Diplasiolejeunealongilobula Herzog – Herzog (1950). Sarawak.

Diplasiolejeuneapatelligera Herzog – Mizutani (1970 as D.neobrachyclada), Tixier (1980a, also as D.neobrachyclada), Dong et al. (2012). Pahang, Sabah.

Diplasiolejeuneaunidentata (Lehm. et Lindenb.) Schiffn. – Pócs and Lee (2016 as D.rudolphiana). Kelantan.

Diplophyllumkinabaluense Furuki et Suleiman – Furuki and Suleiman (2016). Sabah.

Diplophyllumnanum Herzog – Müller and Schäfer-Verwimp (1999). Pahang.

Drepanolejeuneaangustifolia (Mitt.) Grolle – Herzog (1950 as D.tenuis), Tixier (1971 as D.tenuis), Kamimura (1974 as D.tenuis). Pahang, Sarawak, Sabah.

Drepanolejeuneabischlerae (Grolle) Grolle et R.L.Zhu – Grolle and Zhu (2000). Sabah.

Drepanolejeuneaciliata Mizut. – Mizutani (1970), Kamimura (1974). Sabah.

Drepanolejeuneacyclops (Sande Lac.) Grolle et R.L.Zhu – Herzog (1943 as Rhaphidolejeuneacyclops), Mizutani (1966, 1970 both as R.cyclops), Bischler (1968 as R.cyclops). Selangor, Sarawak, Sabah.

Drepanolejeuneadactylophora (Nees, Lindenb. et Gottsche) J.B.Jack et Steph. – Herzog (1934, 1950), Mizutani (1966, 1970), Kamimura (1974), Kitagawa and Kodama (1974), Pócs et al. (2020), Pesiu et al. (2021). Terengganu, Johor, Sarawak, Sabah.

Drepanolejeuneaelegans Herzog – Herzog (1936). Sabah.

Drepanolejeuneafissicornua Steph. – Mizutani (1990), Pócs et al. (2014, 2020), Pócs and Lee (2016). Kelantan, Pahang, Selangor, Sabah.

Drepanolejeuneafoliicola Horik. – Eggers (2006). Pahang.

Drepanolejeuneaintermedia Zwickel – Grolle (1976). Sabah.

Drepanolejeunealevicornua Steph. – Tixier (1971 [‘laevicornua’]), Mizutani (1990), Eggers et al. (1998), Pócs et al. (2014), Pesiu et al. (2021, in press). Selangor, Pahang, Terengganu, Sarawak.

Drepanolejeunealongicornua (Herzog) Mizut. – Mizutani (1970 as D.levicornuavar.longicornua, 1990), Kitagawa and Kodama (1974 as D.levicornuavar.longicornua), Pócs et al. (2020), Pesiu et al. (2021). Penang, Terengganu, Sarawak, Sabah.

Drepanolejeunealongicruris (Steph.) Grolle et R.L.Zhu – Herzog (1943 as Rhaphidolejeunealongicruris), Bischler (1968 as R.longicruris), Kitagawa and Kodama (1974 as R.longicruris), Grolle and Zhu (2000). Sabah.

Drepanolejeuneanymanii Steph. – Mizutani (1990). Sabah.

Drepanolejeuneaobliqua Steph. – Herzog (1939), Mizutani (1970). Johor, Sabah.

Drepanolejeuneaobtriangulata T.Kodama – Kodama (1976), Pócs et al. (2014). Selangor, Sabah.

Drepanolejeuneapentadactyla (Mont.) Steph. – Herzog (1934 as D.micholitziiand itsvar.angustissima), Tixier (1974 as D.micholitzii, 1980a as D.micholitziivar.brevifolia), Mizutani (1966, 1970 both as D.micholitzii), Kitagawa and Kodama (1974 as D.micholitzii), Pócs et al. (2014, 2020), Pesiu et al. (2021, in press). Kedah, Selangor, Pahang, Terengganu, Johor, Sarawak, Sabah.

Drepanolejeuneapentadactyla var. dactylophoroides (Herzog) Pócs – Herzog (1934 as D.micholitziivar.dactylophoroides). Sabah.

Drepanolejeuneapleiodictya Herzog – Tixier (1980a). Pahang.

Drepanolejeuneaserricalyx Herzog – Pócs et al. (2020). Sabah.

Drepanolejeuneaspicata (Steph.) Grolle et R.L.Zhu – Mizutani (1966 as Rhaphidolejeuneaspicata), Bischler (1968 as R.spicata), Grolle and Zhu (2000), Pócs et al. (2014), Pesiu et al. (2021, in press). Pahang, Terengganu, Sarawak, Sabah.

Drepanolejeuneaspinosocornuta Steph. – Tixier (1980a). Pahang.

Drepanolejeuneatenera K.I.Goebel – Herzog (1934 as D.teneravar.genuinaandf.goebelii, 1952b), Mizutani (1966, 1970), Tixier (1971, 1974, 1980a as D.teneraf.goebelii). Kitagawa and Kodama (1974), Pócs et al. (2020). Kedah, Pahang, Sarawak, Sabah.

Drepanolejeuneaternatensis (Gottsche) Schiffn. – Herzog (1950, also as D.ternatensisvar.lancispina), Mizutani (1966, 1970), Kamimura (1974), Cheah (2017), Pócs et al. (2020), Pesiu et al. (2021, in press). Pahang, Terengganu, Sarawak, Sabah.

Drepanolejeuneateysmannii (Gottsche) Steph. – Herzog (1950), Mizutani (1966, 1970), Inoue (1968b), Kamimura (1974, 1975), Kitagawa and Kodama (1974), Kürschner (1990), Pócs et al. (2020). Pahang, Sarawak, Sabah.

Drepanolejeuneathwaitesiana (Mitt.) Steph. – Goebel (1930 [‘Thwaitesii’]), Mizutani (1966, 1970, 1990), Tixier (1971, 1974, 1980a), Kitagawa and Kodama (1974), Pócs et al. (2020), Pesiu et al. (2021, in press). Kedah, Perak, Selangor, Pahang, Terengganu, Sabah.

Drepanolejeuneathwaitesiana var. zhengii R.L.Zhu – Zhu and So (2001). Pahang.

Drepanolejeuneatricornua Herzog – Mizutani (1970, 1990), Pócs et al. (2014, 2020). Selangor, Pahang, Sarawak, Sabah.

Drepanolejeuneavesiculosa (Mitt.) Steph. – Herzog (1950 as D.vesiculosaf.robusta), Inoue (1967a, 1968b), Mizutani (1966, 1970), Kamimura (1974), Kitagawa and Kodama (1974), Kürschner (1990), Lee et al. (2013), Pócs et al. (2020). Perak, Pahang, Sarawak, Sabah.

Dumortierahirsuta (Sw.) Nees – Johnson (1958), Akiyama et al. (2001), Ludwiczuk and Asakawa (2010). Pahang, Sabah.

Dumortierahirsuta subsp. nepalensis (Taylor) R.M.Schust. – Campbell (1915 as D.trichocephala, 1918 as D.calcicola), Evans (1919 as D.nepalensis), Kitagawa and Kodama (1974 as D.nepalensis). Perak, Selangor, Sarawak, Sabah.

Eotrichocoleafurukii T.Katag. – Katagiri et al. (2012). Sabah.

Frullaniaalstonii Verd. – Verdoorn (1932b), Hattori (1976). Johor, Sabah.

Frullaniaapiculata (Reinw., Blume et Nees) Nees – Verdoorn (1932a), Herzog (1950), Inoue (1967a, 1968b), Tixier (1974, 1980a), Kamimura (1974, 1975), Kitagawa and Kodama (1974), Hattori (1975c, 1976), Kodama (1976), Pócs et al. (2020), Pesiu et al. (2021). Kedah, Terengganu, Pahang, Selangor, Sarawak, Sabah.

Frullaniaapiculata var. goebelii Schiffn. – Verdoorn (1934b), Herzog (1950), Tixier (1971). Pahang, Sarawak, Sabah.

Frullaniaarmatifolia Verd. – Hattori (1976 as F.altemammillata). Sabah.

Frullaniaarmitiana Steph. – Kamimura (1974). Sabah.

Frullaniabenjaminiana Inoue – Inoue (1967a,b), Hattori (1975a). Pahang.

Frullaniaberthoumieui Steph. – Hattori (1976). Sabah.

Frullaniabrotheri Steph. – Stephani (1894), Schiffner (1898), Bonner (1965), Hattori (1975a,c, 1976, 1980), Kürschner (1990), Sukkharak (2018). Perlis, Perak, Selangor, Pahang, Malacca, Johor, Sabah.

Frullaniaclaviloba Steph. – Verdoorn (1930, 1932b), Herzog (1950), Kamimura (1974), Kitagawa and Kodama (1974), Hattori (1974b, 1975b,c, 1976). Selangor, Johor, Sarawak, Sabah.

Frullaniaclemensiana Verd. – Verdoorn (1932a), Hattori and Kamimura (1971 as Steereamastigophoroides), Hattori (1976 as S.clemensiana). Sabah.

Frullaniacordistipula (Reinw., Blume et Nees) Nees – Kamimura (1974, 1975). Sabah.

Frullaniaericoides (Nees) Mont. – Verdoorn (1929 as F.squarrosa), Hattori (1975a, 1976). Penang, Pahang, Sabah.

Frullaniafallax Gottsche – Verdoorn (1930). Pahang.

Frullaniagaudichaudii (Nees et Mont.) Nees et Mont. – Verdoorn (1930, 1932a), Herzog (1950), Kitagawa and Kodama (1974), Hattori (1975c, 1976), Cheah (2017). Selangor, Pahang, Sarawak, Sabah.

Frullaniagracilis (Reinw., Blume et Nees) Nees – Verdoorn (1932a as F.minor), Herzog (1950), Kamimura (1974, 1975), Kitagawa and Kodama (1974), Hattori (1975a, 1975c as F.minor, 1976), Kürschner (1990). Selangor, Pahang, Malacca, Johor, Sarawak, Sabah.

Frullaniagrandistipula Lindenb. – Verdoorn (1932a), Tixier (1971), Hattori (1976). Pahang, Sabah.

Frullaniahasskarliana Lindenb. – Verdoorn (1932a), Hattori (1973, 1975a, 1976 as F.hasskarlianavar.integribracteata), Kitagawa and Kodama (1974). Pahang, Sabah.

Frullaniahottana S.Hatt. – Hattori (1976, 1986). Sarawak.

Frullaniahypoleuca Nees – Hattori (1976), Kamimura (1974). Sarawak, Sabah.

Frullaniaintegristipula (Nees) Nees – Verdoorn (1930, 1932a), Herzog (1950), Kamimura (1974), Kitagawa and Kodama (1974), Tixier (1974, 1980a), Hattori (1976, 1980). Kedah, Pahang, Johor, Sarawak, Sabah.

Frullaniaintermedia (Reinw., Blume et Nees) Nees – Verdoorn (1930, 1934b), Herzog (1950), Kitagawa and Kodama (1974), Hattori (1975c, 1976 as F.intermediavar.submorokensis and F.intermediaf.billardieriana, 1980), Wolseley et al. (1996). Penang, Perak, Kelantan, Negeri Sembilan, Johor, Sarawak, Sabah.

Frullaniajunghuhniana Gottsche – Verdoorn (1932a, 1934b), Hattori (1974c, 1976), Kürschner (1990). Sabah.

Frullaniajunghuhniana var. bisexualis S.Hatt. – Hattori (1976), Kodama (1976). Sabah.

Frullaniajunghuhniana var. tenella (Sande Lac.) Grolle et S.Hatt. – Kamimura (1974 as F.perversa), Tixier (1974 as Neohattoriaperversa), Hattori (1975a, 1976 as F.junghuhnianavar.minutissima and F.perminuta). Kedah, Pahang, Sabah.

Frullaniameijeri S.Hatt. – Hattori (1974c, 1976). Sabah.

Frullaniameyeniana Lindenb. – Verdoorn (1934b), Tixier (1971), Hattori (1975c). Pahang, Sabah.

Frullaniamizutanii Kamim. et S.Hatt. – Hattori and Kamimura (1973), Kamimura (1974), Hattori (1976, 1978), Kodama (1976). Sabah.

Frullaniamoniliata (Reinw., Blume et Nees) Mont. – Verdoorn (1932a as F.moniliatasubsp.breviramea), Hattori (1972, 1976 both as F.tamariscivar.breviramea). Sabah.

Frullaniamonocera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees – Hattori (1976 as F.hampeana). Sabah.

Frullanianepalensis (Spreng.) Lehm. et Lindenb. – Verdoorn (1932b). Pahang.

Frullanianigricaulis (Reinw., Blume et Nees) Nees – Verdoorn (1930, 1932a), Hattori (1975a). Pahang, Sabah.

Frullanianodulosa (Reinw., Blume et Nees) Nees – Verdoorn (1930, 1932a, 1934b), Herzog (1950), Inoue (1968b), Kitagawa and Kodama (1974), Hattori (1975c, 1980). Perak, Selangor, Pahang, Johor, Sarawak, Sabah.

Frullanianotarisii Steph. – Verdoorn (1934b), Hattori (1974c, 1976), Kitagawa and Kodama (1974), Tixier (1974). Kedah, Sarawak, Sabah.

Frullaniaobscura (Sw.) Mont. – Hattori (1975a as F.wallichiana). Pahang.

Frullaniaocellata S.Hatt. et Kamim. – Hattori and Kamimura (1973), Kamimura (1974). Sabah.

Frullaniaorientalis Sande Lac. – Verdoorn (1937), Hattori (1976). Pahang, Sabah.

Frullaniaornithocephala (Reinw., Blume et Nees) Nees – Verdoorn (1934b), Hattori (1975a, 1976 as F.ornithocephalaf.magnilobulaandf.retusa, 1980). Pahang, Sabah.

Frullaniapapillata Steph. – Kamimura (1974, 1975). Sabah.

Frullaniapapillilobula S.Hatt. – Hattori (1975b, 1976). Sabah.

Frullaniapolyptera Taylor – Hattori (1975a). Pahang.

Frullaniapullei Verd. – Tixier (1974). Kedah.

Frullaniaramuligera (Nees) Mont. – Verdoorn (1932a, 1934b), Hattori (1972), Pócs et al. (2020). Sabah.

Frullaniarecurvistipula S.Hatt. – Hattori (1975a, 1986). Pahang.

Frullaniareflexistipula Sande Lac. – Verdoorn (1932a), Hattori (1975a as F.philippinensis, 1976). Pahang, Sabah.

Frullaniarepandistipula Sande Lac. subsp. spinibractea S.Hatt. – Hattori (1975c, 1976 as F.repandistipulasubsp.spinifera), Pesiu et al. (in press). Terengganu, Sabah.

Frullaniasabahana S.Hatt. – Hattori (1976, 1980). Sarawak, Sabah.

Frullaniasarawakensis S.Hatt. – Hattori (1976, 1980). Sarawak.

Frullaniaserrata Gottsche – Verdoorn (1932b, 1934b), Herzog (1950), Kamimura (1974, 1975), Kitagawa and Kodama (1974), Hattori (1975a, 1976 as F.serrataf.crispulodentata), Ludwiczuk and Asakawa (2010). Pahang, Johor, Sarawak, Sabah.

Frullaniasinuata Sande Lac. – Verdoorn (1932b, 1934b), Hattori (1975a, 1980). Johor, Pahang.

Frullaniasublignosa Steph. – Stephani (1910 as F.borneensis), Verdoorn (1930), Hattori (1976). Sarawak.

Frullaniasubnigricaulis S.Hatt. – Hattori (1974b, 1975a, 1976). Pahang, Sabah.

Frullaniasubocellata S.Hatt. – Pócs et al. (2014). Pahang.

Frullaniaternatensis Gottsche – Verdoorn (1930, 1932a, 1934b), Herzog (1950), Tixier (1971, 1974), Kamimura (1974, 1975), Kitagawa and Kodama (1974), Hattori (1975c). Kedah, Pahang, Sarawak, Sabah.

Frullaniaternatensis var. nonappendiculata S.Hatt. – Hattori (1976). Sarawak, Sabah.

Frullaniatogashiana S.Hatt. – Hattori (1975a). Pahang.

Frullaniatricarinata Sande Lac. – Kamimura (1974). Sabah.

Frullaniatrichodes Mitt. – Verdoorn (1930, 1934b as F.picta, F.tenuicaulis and F.vethii), Herzog (1950 as F.picta, F.tenuicaulis and F.vethii), Kitagawa (1969c as F.picta), Kamimura (1974, 1975 both as F.tenuicaulis), Kitagawa and Kodama (1974 as F.tenuicaulis), Tixier (1974 as F.picta), Hattori (1976 as F.vethiif.acutiloba), Pócs et al. (2014), Cheah (2017). Kedah, Penang, Pahang, Johor, Sarawak, Sabah.

Frullaniavaginata (Sw.) Nees – Hattori (1975a). Pahang.

Frullaniavenusta S.Hatt. – Hattori (1974b, 1976). Sabah.

Gottscheliaschizopleura (Spruce) Grolle – Inoue (1967a as Jamesoniellamicrophylla), Amakawa (1969a as J.microphylla), Váňa (1991c). Pahang, Sabah.

Gymnomitrionincompletum (Gottsche) Váňa – Kitagawa (1973 as G.laceratumvar.borneense), Kitagawa and Kodama (1973 as G.laceratumvar.borneense), Váňa (1991a). Sabah.

Gymnomitrionrevolutum (Nees) H.Philib. – Kitagawa (1967 as Marsupellarevoluta), Kitagawa and Kodama (1973 as M.revoluta), Váňa (1991a as M.revoluta), Akiyama et al. (2001 as M.revoluta). Sabah.

Gymnomitrionsubintegrum (S.W.Arnell) Váňa – Kitagawa (1967 as Marsupellaintegra), Váňa (1991aM.subintegra). Sabah.

Haplomitriumblumei (Nees) R.M.Schust. – Bartholomew-Began (1991). Pahang.

Herbertusaduncus (Dicks.) Gray – Kürschner (1990 as H.dicranus [‘dicrananus’]), Juslén (2006 as H.dicranus and H.longispinus), Cheah (2017 as H.dicranus). Pahang, Sabah.

Herbertusceylanicus (Steph.) Abeyw. – Juslén (2006). Pahang.

Herbertuspilifer (Steph.) H.A.Mill. – Juslén (2006). Sarawak.

Herbertusramosus (Steph.) H.A.Mill. – Kamimura (1975 [‘Herbertaramosa’]). Sabah.

Herbertussendtneri (Nees) Lindb. – Kürschner (1990 as H.armitanus and H.circinatus), Akiyama et al. (2001 as H.armitanus), Juslén (2006 as H.circinatus), Cheah (2017 as H.armitanus). Pahang, Sabah.

Heteroscyphusacutangulus (Schiffn.) Schiffn. – Inoue (1968b as Chiloscyphusacutangulus). Pahang.

Heteroscyphusargutus (Reinw., Blume et Nees) Schiffn. – Herzog (1950 as Chiloscyphusargutus), Kitagawa (1969c), Tixier (1971 as C.argutus), Kitagawa and Kodama (1974 as C.argutus), Inoue (1981), Piippo (1989), Wolseley et al. (1996). Penang, Pahang, Negeri Sembilan, Sarawak, Sabah.

Heteroscyphusaselliformis (Reinw., Blume et Nees) Schiffn. – Mitten and Wright (1894 as Chiloscyphusaselliformis), Herzog (1950 as C.aselliformisvar.neesii), Inoue (1967a as C.aselliformis), Kitagawa and Kodama (1974), Piippo (1989), Kürschner (1990 as C.aselliformis), Ludwiczuk and Asakawa (2010). Pahang, Sarawak, Sabah.

Heteroscyphusbalnetii (Herzog) Grolle – Piippo (1989). Sabah.

Heteroscyphuscoalitus (Hook.) Schiffn. – Herzog (1950 as Chiloscyphuscommunis), Inoue (1967a, 1968b both as C.communis), Kitagawa (1971 as H.communis), Tixier (1971 as C.communis), Kitagawa and Kodama (1974 as C.communis), Kamimura (1975 as H.bescherellei), Piippo (1989). Penang, Pahang, Sarawak, Sabah.

Heteroscyphusdiestianus (Sande Lac.) Piippo – Kitagawa and Kodama (1974 as Chiloscyphusdiestianus), Piippo (1989), Akiyama et al. (2001). Sarawak, Sabah.

Heteroscyphusintegerrimus (Schiffn.) Gradst. et G.E.Lee – Pócs et al. (2014 as Chiloscyphusintegerrimus). Pahang.

Heteroscyphusiwatsukii (S.Hatt.) Piippo – Hattori (1964 as Saccogynidiumiwatsukii), Piippo (1989). Sabah.

Heteroscyphusparvulus (Schiffn.) Schiffn. – Kitagawa and Kodama (1974 as Chiloscyphusparvulus). Sabah.

Heteroscyphussplendens (Lehm. et Lindenb.) Grolle – De Notaris (1874 as Chiloscyphusdecurrens and C.densifolius), Herzog (1950, 1952b both as C.decurrens), Inoue (1967a, 1968b, both as C.decurrens), Kitagawa (1969c as C.decurrens), Kitagawa and Kodama (1974 as C.decurrens), Tixier (1974 as C.decurrens), Piippo (1989), Kürschner (1990), Akiyama et al. (2001). Kedah, Penang, Pahang, Sarawak, Sabah.

Heteroscyphussucculentus (Gottsche) Schiffn. – De Notaris (1874 as Chiloscyphusconcinnus), Stephani (1907 as C.succulentus), Herzog (1950 as C.succulentus), Kitagawa (1973 as C.succulentus), Kitagawa and Kodama (1974 as C.succulentus), Piippo (1989). Penang, Sarawak, Sabah.

Heteroscyphustridentatus (Sande Lac.) Grolle – Cheah (2017). Pahang.

Heteroscyphuswettsteinii (Schiffn.) Schiffn. – Kitagawa and Kodama (1974 as Chiloscyphuswettsteinii), Müller and Schäfer-Verwimp (1999), Cheah (2017). Perak, Pahang, Sabah.

Hygrolembidiumboschianum (Sande Lac.) R.M.Schust. – Grolle (1967). Malacca.

Isotachisjaponica Steph. – Inoue (1967a). Pahang.

Jackiellaangustifolia Herzog – Herzog (1950, 1952a), Váňa (1992). Sarawak, Sabah.

Jackiellajavanica Schiffn. – Herzog (1950), Inoue (1967a), Kitagawa (1971), Váňa (1992). Penang, Pahang, Sarawak, Sabah.

Jackiellasingapurensis Schiffn. – Kitagawa (1969c, 1981). Penang.

Jenseniadecipiens (Mitt.) Grolle – Herzog (1950 as Makednothallusobtusidens), Kitagawa and Kodama (1974 as J.zollingeri), Grolle and Piippo (1986), Akiyama et al. (2001), Forrest et al. (2005). Sarawak, Sabah.

Jubulajavanica Steph. – Herzog (1950 as J.hutchinsiaesubsp.javanica), Kitagawa and Kodama (1974 as J.hutchinsiaesubsp.javanica), Váňa and Piippo (1989b as J.hutchinsiaesubsp.javanica), Pätsch et al. (2010 as J.hutchinsiaesubsp.javanica). Pahang, Sarawak, Sabah.

Kurziaabbreviata Mizut. – Mizutani (1974), Hürlimann (1985), Cheah (2017). Pahang, Sarawak, Sabah.

Kurziaabietinella (Herzog) Grolle – Herzog (1950 as Lepidoziaabietinella), Kitagawa and Kodama (1973), Mizutani (1974), Kürschner (1990), Cheah (2017). Pahang, Sarawak, Sabah.

Kurziaborneensis Mizut. – Mizutani (1974), Kamimura (1975), Akiyama et al. (2001), Cheah (2017). Pahang, Sabah.

Kurziageniculata Mizut. – Mizutani (1974), Cheah (2017). Pahang, Sabah.

Kurziagonyotricha (Sande Lac.) Grolle – Herzog (1952b as Lepidoziagonyotricha), Kitagawa (1969c), Mizutani (1974), Akiyama et al. (2001). Penang, Sarawak, Sabah.

Kurzialineariloba Mizut. – Mizutani (1974), Kürschner (1990), Cheah (2017). Pahang, Sabah.

Kymatocalyxrhizomaticus (Herzog) Gradst. et Váňa – Herzog (1950, 1952a both as Stenorrhipisrhizomatica), Váňa (1991c as S.rhizomatica). Sarawak.

Leiomitramerrillana (Steph.) T.Katag. – Kitagawa (1973 as Trichocoleamerrillana), Akiyama et al. (2001 as T.merrillana), Katagiri and Deguchi (2012). Pahang, Sabah.

Lejeuneaadpressa Nees – Inoue (1968b as L.boninensis), Kitagawa (1971 as L.borneensis), Mizutani (1978 as L.borneensis), Wolseley et al. (1996 as L.caespitosa), Zhu and So (2001 as L.anisophylla), Lee (2013 as L.anisophylla), Pesiu et al. (2021, in press). Perlis, Langkawi, Perak, Penang, Selangor, Kuala Lumpur, Pahang, Negeri Sembilan, Terengganu, Johor, Sarawak, Sabah.

Lejeuneaalata Gottsche – Mizutani (1970 as L.mitracalyx), Kitagawa and Kodama (1974 as L.mitracalyx), Lee (2013). Pahang, Sabah.

Lejeuneaalbescens (Steph.) Mizut. – Eifrig (1937 as Taxilejeuneaalbescens), Inoue (1967a as T.albescens), Mizutani (1970), Hattori and Kamimura (1971), Lee (2013). Pahang, Sabah.

Lejeuneaapiculata Sande Lac. – Mizutani (1966 as Prionolejeuneaungulata, 1970 as Stenolejeuneaapiculata), Tixier (1971 as S.apiculata), Kitagawa and Kodama (1974 as P.ungulata), Lee (2013), Cheah (2017). Pahang, Sabah.

Lejeuneacocoes Mitt. – Mizutani (1963), Kamimura (1974), Lee (2013), Pesiu et al. (in press). Perlis, Pahang, Terengganu, Sarawak, Sabah.

Lejeuneacompacta (Steph.) Steph. – Lee (2013). Sabah.

Lejeuneacompressiuscula (Steph.) G.E.Lee et Heinrichs – Eifrig (1937 as Taxilejeuneacompressiuscula), Tixier (1971 as T.compressiuscula). Pahang, Johor.

Lejeuneacontracta Mizut. – Mizutani (1970), Kitagawa and Kodama (1974), Lee (2013). Sabah.

Lejeuneaconvexiloba M.L.So et R.L.Zhu – Sarimi et al. (2021). Sabah.

Lejeuneadimorpha Kodama – Kodama (1976), Lee (2013), Pócs et al. (2014). Selangor, Kelantan, Pahang, Sabah.

Lejeuneadipterota (Eifrig) G.E.Lee – Lee (2013). Sabah.

Lejeuneadiscreta Lindenb. – Mizutani (1970), Kitagawa and Kodama (1974), Tixier (1980a as Xylolejeunealongiloba), Lee et al. (2011b), Lee (2013). Perlis, Perak, Pahang, Sabah.

Lejeuneaeifrigii Mizut. – Mizutani (1970), Lee et al. (2011b), Lee (2013). Pahang, Sabah.

Lejeuneaexilis (Reinw., Blume et Nees) Grolle – Mizutani (1970), Pócs et al. (2020). Sabah.

Lejeuneaexilis var. abnormis (Herzog) G.E.Lee – Lee (2013). Kelantan, Pahang.

Lejeuneaflava (Sw.) Nees – Inoue (1967a), Mizutani (1970), Kitagawa and Kodama (1974), Lee (2013), Pócs et al. (2020). Kedah, Penang, Perak, Selangor, Kelantan, Pahang, Sabah.

Lejeunea flavida Mitt. – Kamimura (1974 as Pycnolejeuneaflavida), Inoue (1967a as P.flavida). Pahang, Sabah.

Lejeuneafleischeri (Steph.) Mizut. – Lee (2013). Perak, Pahang.

Lejeuneagradsteinii G.E.Lee, Damanhuri et Latiff – Lee et al. (2011a), Lee (2013), Pócs and Lee (2016). Pahang, Sabah.

Lejeuneakinabalensis Mizut. – Mizutani (1970), Lee (2013). Sabah.

Lejeunealeratii (Steph.) Mizut. – Eifrig (1937 as Taxilejeuneapatersonii). Sabah.

Lejeunealumbricoides (Nees) Nees – Eifrig (1937 as Taxilejeunealumbricoides), Mizutani (1970), Kitagawa and Kodama (1974), Tixier (1980a), Lee (2013). Pahang, Sabah.

Lejeuneamalaysiana G.E.Lee et Pócs - Lee et al. (in press). Pahang.

Lejeuneamicholitzii Mizut. – Mizutani (1970), Lee (2013), Pócs et al. (2014, 2020), Pesiu et al. (2021). Langkawi, Selangor, Pahang, Negeri Sembilan, Kelantan, Terengganu, Sarawak, Sabah.

Lejeuneamicroloba Taylor – Mizutani (1970 as L.chalmersii), Lee (2013). Sabah.

Lejeuneamimula Hürl. – Tixier (1971 as Taxilejeunealuteola), Mizutani (1970 as L.luteola), Kitagawa and Kodama (1974 as L.luteola), Lee (2013). Pahang, Sabah.

Lejeuneamizutanii Grolle – Mizutani (1970 as L.zollingeri), Lee (2013). Sabah.

Lejeuneapapilionacea Prantl – Mizutani (1966 as L.infestans and L.pterota, 1970 as L.herzogii and L.infestans, 1972b as L.diversitexta), Kitagawa and Kodama (1974 as L.diversitexta), Zhu and Grolle (2001), Lee (2013), Pócs et al. (2020). Kedah, Langkawi, Selangor, Kelantan, Pahang, Sarawak, Sabah.

Lejeuneapatersonii (Steph.) Steph. – Inoue (1967a), Lee (2013). Kuala Lumpur, Kelantan, Pahang, Johor.

Lejeuneapatriciae Schäf.-Verw. – Tixier (1971 as L.pilifera), Schäfer-Verwimp (2001, 2006), Lee (2013). Penang, Perak, Kelantan, Pahang.

Lejeuneapectinella Mizut. – Mizutani (1970), Lee (2013). Sabah.

Lejeuneapulchriflora (Pearson) G.E.Lee, Bechteler, Pócs, Schäf.-Verw. et Heinrichs – Lee (2013 as L.tamaspocsii), Lee and Gradstein (2013 as L.tamaspocsii), Lee et al. (2016). Kelantan, Pahang, Sabah.

Lejeuneasordida (Nees) Nees – Eifrig (1937 as Taxilejeuneasordida), Mizutani (1970), Lee et al. (2011a, 2011b), Lee (2013). Perlis, Langkawi, Kedah, Penang, Perak, Selangor, Kelantan, Pahang, Johor, Sarawak, Sabah.

Lejeuneastenodentata M.A.M.Renner et Pócs – Herzog (1950 as Crossotolejeuneaborneensis). Mizutani (1966, 1970 both as C.borneensis, 1972a as Drepanolejeuneadentata). Sarawak, Sabah.

Lejeuneastephaniana Mizut. – Mizutani (1966, 1970), Lee (2013). Sabah.

Lejeuneathallophora (Eifrig) Gradst. – Zhu et al. (2018b). Sabah.

Lejeuneatrinitensis Lindenb. – Reiner-Drehwald and Grolle (2012). Johor.

Lejeuneatuberculosa Steph. – Lee (2013). Kedah, Penang, Perak, Selangor, Pahang, Sabah.

Lejeuneaumbilicata (Nees) Nees – Herzog (1950 as Taxilejeuneaumbilicata), Mizutani (1970), Kitagawa and Kodama (1974), Tixier (1980a), Grolle (1988 as L.cuculliflora), Lee (2013). Perak, Selangor, Kelantan, Pahang, Johor, Sarawak, Sabah.

Lejeuneautriculata (Steph.) Mizut. – Mizutani (1966 as Pycnolejeuneautriculata, 1970), Lee (2013). Sabah.

Lejeuneawightii Steph. – Mizutani (1966, 1970). Sabah.

Lepicolearara (Steph.) Grolle – Inoue (1967a, 1968b both as L.loriana), Kitagawa (1978 as L.loriana), Pocock et al. (1984 as L.loriana), Kürschner (1990), Cheah (2017). Pahang, Sabah.

Lepicoleayakusimensis (S.Hatt.) S.Hatt. – Inoue (1967a), Kitagawa and Kodama (1973), Akiyama et al. (2001). Pahang, Sabah.

Lepidolejeuneabidentula (Steph.) R.M.Schust – Hoffmann (1935 as Pycnolejeuneacorticola and P.decurvifolia), Herzog (1950 as P.badiaf.parvistipa, 1952b as P.corticola and P.nicobarica), Mizutani (1966, 1970 as P.bidentula, P.punctata and P.nicobarica), Inoue (1967a as P.bidentula), Kitagawa (1969c as P.nicobarica), Kitagawa and Kodama (1974 as P.bidentula and P.nicobarica), Piippo (1986), Wolseley et al. (1996), Pócs et al. (2020), Pesiu et al. (2021). Penang, Selangor, Pahang, Negeri Sembilan, Terengganu, Johor, Sarawak, Sabah.

Lepidolejeuneaintegristipula (J.B.Jack et Steph.) R.M.Schust. – Mizutani (1970, 1972b both as Pycnolejeuneaintegristipula), Kitagawa and Kodama (1974 as P.integristipula), Piippo (1986), Cheah (2017). Pahang, Sabah.

Lepidoziaambigua De Not. – De Notaris (1874), Schiffner (1898), Stephani (1909). Sarawak.

Lepidoziaborneensis Steph. – Herzog (1950), Kitagawa and Kodama (1973), Mizutani (1974), Kamimura (1975), Ludwiczuk and Asakawa (2010). Sarawak, Sabah.

Lepidoziabrotheri Steph. – Kürschner (1990). Sabah.

Lepidoziacladorhiza (Reinw., Blume et Nees) Nees – Mitten and Wright (1894), Herzog (1950 as L.cladorhizavar.macgregorii), Kitagawa and Kodama (1973), Mizutani (1974), Kürschner (1990), Akiyama et al. (2001). Sarawak, Sabah.

Lepidoziaeverettii Steph. – Stephani (1909), Herzog (1950), Mizutani (1968). Sarawak.

Lepidoziafauriana Steph. – Mizutani (1974), Ludwiczuk and Asakawa (2010). Sabah.

Lepidoziaferdinandi-muelleri Steph. – Kitagawa and Kodama (1973), Mizutani (1974). Sabah.

Lepidoziahaskarliana (Gottsche, Lindenb. et Nees) Steph. – Mizutani (1974). Sabah.

Lepidoziaholorhiza (Reinw., Blume et Nees) Nees – Mitten and Wright (1894), Inoue (1968b). Pahang, Sabah.

Lepidoziakinabaluensis Mizut. – Mizutani (1974). Sabah.

Lepidozialacerifolia Steph. – Herzog (1950). Sarawak.

Lepidoziamiqueliana Sande Lac. – De Notaris (1874), Schiffner (1898), Herzog (1950). Sarawak.

Lepidoziareptans (L.) Dumort. – Mizutani (1974). Sabah.

Lepidoziarichardsii Herzog – Herzog (1950). Sarawak.

Lepidoziasquamifolia Steph. – Mizutani (1968, 1974). Sarawak, Sabah.

Lepidoziasubintegra Lindenb. – Mitten and Wright (1894), Herzog (1950). Sarawak, Sabah.

Lepidoziasupradecomposita Lindenb. – Kitagawa and Kodama (1973), Mizutani (1974), Akiyama et al. (2001). Sabah.

Lepidoziatrichodes (Reinw., Blume et Nees) Nees – Mitten and Wright (1894), Herzog (1950), Inoue (1967a), Kitagawa (1969c), Kitagawa and Kodama (1973), Mizutani (1974), Kamimura (1975), Pocock et al. (1984), Kürschner (1990). Penang, Pahang, Sarawak, Sabah.

Leptolejeuneaamphiophthalma Zwickel – Mizutani (1966 as L.picta), Kitagawa and Kodama (1974 as L.picta), Tixier (1974 as L.picta), Grolle (1988), Pócs et al. (2020), Pesiu et al. (2021). Kedah, Terengganu, Sarawak, Sabah.

Leptolejeuneaapiculata (Horik.) S.Hatt. – Pócs and Lee (2016). Pahang.

Leptolejeuneabalansae Steph. – Dürhammer and Schäfer-Verwimp (1995). Perak.

Leptolejeuneaborneensis Herzog – Herzog (1942), Mizutani (1966). Sabah.

Leptolejeuneadentistipula Steph. – Herzog (1942). Sarawak.

Leptolejeuneaepiphylla (Mitt.) Steph. – Herzog (1942), Mizutani (1966), Kitagawa (1969c), Kitagawa and Kodama (1974), Tixier (1974), Thiers (1986), Wolseley et al. (1996), Pócs et al. (2020), Pesiu et al. (2021, in press). Kedah, Penang, Selangor, Pahang, Negeri Sembilan, Terengganu, Sabah.

Leptolejeuneafoliicola Steph. – Herzog (1942), Tixier (1971, 1974). Kedah, Pahang, Johor.

Leptolejeunealigulata Herzog – Herzog (1942), Pócs et al. (2020). Pahang, Sabah.

Leptolejeuneamaculata (Mitt.) Schiffn. – Herzog (1942 as L.schiffnerif.latifolia,f.angustifoliaandvar.subintegerrima), Tixier (1974, 1980a both as L.radiata), Kitagawa and Kodama (1974 as L.schiffneri), Mizutani (1966, 1970 both as L.schiffneri, 1975), Dürhammer and Schäfer-Verwimp (1995), Wolseley et al. (1996 as L.schiffneri), Pócs et al. (2020), Pesiu et al. (2021, in press). Kedah, Perak, Pahang, Negeri Sembilan, Terengganu, Johor, Sarawak, Sabah.

Leptolejeuneamassartiana Herzog – Tixier (1971). Pahang.

Leptolejeunearenneri Herzog – Shu et al. (2021). Kelantan.

Leptolejeuneasubacuta A.Evans – Herzog (1942), Mizutani (1966, also as L.elliptica), Tixier (1971), Kitagawa and Kodama (1974 as L.elliptica), Wolseley et al. (1996 as L.ellipticasubsp.subacuta), Pócs et al. (2020 as L.elliptica), Pesiu et al. (2021, in press). Perak, Pahang, Negeri Sembilan, Terengganu, Johor, Sabah.

Leptolejeuneasubdentata Herzog – Herzog (1942), Mizutani (1966), Pócs et al. (2020). Johor, Sarawak, Sabah.

Leptolejeuneasubrotundifolia Herzog – Shu et al. (2021). Kelantan, Pahang.

Leptolejeuneatripuncta (Mitt.) Steph. – Herzog (1942 as L.serrulata), Inoue (1967a as L.serrulata), Kitagawa (1969c as L.serrulata), Thiers (1986 as L.serrulata), Pócs et al. (2020), Shu et al. (2021). Penang, Selangor, Pahang, Sabah.

Leptolejeuneavitrea (Nees) Schiffn. – Herzog (1942), Mizutani (1966), Kitagawa (1969c), Tixier (1971, 1974, 1980a), Kitagawa and Kodama (1974), Wolseley et al. (1996), Pócs et al. (2020), Pesiu et al. (2021, in press). Kedah, Penang, Perak, Pahang, Negeri Sembilan, Terengganu, Johor, Sabah.

Lobatiriccardiacoronopus (De Not.) Furuki – Herzog (1950 as Riccardialobataandf.gigantea), Furuki (1996 as L.lobata). Pahang, Sarawak, Sabah.

Lophocoleabidentata (L.) Dumort. – Piippo (1989 as Chiloscyphuslatifolius), Cheah (2017 as C.coadunatus and C.cuspidatus). Pahang, Sabah.

Lophocoleakurzii Sande Lac. – Kitagawa and Kodama (1974), Piippo (1989 as Chiloscyphuskurzii). Sabah.

Lophocoleamollis (Nees) Nees – Inoue (1967a). Pahang.

Lophocoleamuricata (Lehm.) Nees – Kitagawa and Kodama (1974), Piippo (1989 as Chiloscyphusmuricatus), Müller and Schäfer-Verwimp (1999), Cheah (2017 as C.muricatus). Pahang, Sabah.

Lophocoleasikkimensis (Steph.) Herzog et Grolle – Piippo (1989 as Chiloscyphussikkimensis). Sabah.

Lophocoleasteetziae De Not. – De Notaris (1874), Schiffner (1898), Piippo (1989 as Chiloscyphussteetziae).

Lopholejeuneaapplanata (Reinw., Blume et Nees) Schiffn. – Verdoorn (1934a), Mizutani (1969), Kitagawa and Kodama (1974), Zhu and Gradstein (2005). Perak, Sabah.

Lopholejeuneaborneensis (Steph.) Verd. – Verdoorn (1934a), Zhu and Gradstein (2005). Sarawak, Sabah.

Lopholejeuneaceylanica Steph. – Mizutani (1985), Zhu and Gradstein (2005), Cheah (2017), Pesiu et al. (in press). Pahang, Terengganu, Sabah.

Lopholejeuneaeulopha (Taylor) Schiffn. – Stephani (1912 as L.cranstonii), Verdoorn (1934a), Kitagawa (1969a), Mizutani (1969, 1979b), Kitagawa and Kodama (1974), Zhu and Gradstein (2005), Lee et al. (2013), Cheah (2017 as L.nicobarica), Pócs et al. (2020), Pesiu et al. (in press). Penang, Perak, Pahang, Terengganu, Johor, Sarawak, Sabah.

Lopholejeuneaherzogiana Verd. – Verdoorn (1934a), Mizutani (1969, 1985), Kitagawa and Kodama (1974), Tixier (1974), Thiers (1983), Zhu and Gradstein (2005). Kedah, Pahang, Sabah.

Lopholejeuneahorticola Schiffn. – Mizutani (1985). Sabah.

Lopholejeuneamagna Mizut. – Mizutani (1969), Kitagawa and Kodama (1974), Zhu and Gradstein (2005). Sabah.

Lopholejeuneanigricans (Lindenb.) Schiffn. – Mizutani (1969, 1985 as L.javanica), Kitagawa and Kodama (1974), Zhu and Gradstein (2005), Lee et al. (2013), Cheah (2017), Pócs et al. (2020), Pesiu et al. (in press). Perak, Pahang, Terengganu, Sarawak, Sabah.

Lopholejeunearecurvata Mizut. – Zhu and Gradstein (2005). Pahang.

Lopholejeuneasubfusca (Nees) Schiffn. – Verdoorn (1934a), Herzog (1950), Mizutani (1966, 1969), Kitagawa (1969a), Tixier (1971, 1980a), Kamimura (1974), Kitagawa and Kodama (1974), Kürschner (1990), Wolseley et al. (1996), Zhu and Gradstein (2005), Lee et al. (2013), Pócs et al. (2020). Perak, Penang, Pahang, Negeri Sembilan, Sarawak, Sabah.

Lopholejeuneawiltensii Steph. – Mizutani (1979a), Akiyama et al. (2001), Zhu and Gradstein (2005). Pahang, Sabah.

Lopholejeuneazollingeri (Steph.) Schiffn. – Verdoorn (1934a), Mizutani (1969, also as L.nipponica, 1981b), Kitagawa and Kodama (1974), Zhu and Gradstein (2005), Cheah (2017 as L.latialata). Pahang, Sabah.

Marchantiaacaulis Steph. – Tixier (1971), Bischler-Causse (1989). Perak, Pahang, Sarawak, Sabah.

Marchantiaemarginata Reinw., Blume et Nees – Bischler-Causse (1989), Gepp (1914). Selangor, Sarawak, Sabah.

Marchantiageminata Reinw., Blume et Nees – Johnson (1958), Bischler-Causse (1989). Perak, Pahang, Selangor.

Marchantianovoguineensis Bischl. – Bischler-Causse (1989). Sabah.

Marchantiapolymorpha L. – Johnson (1958), Kitagawa and Kodama (1974), Bischler-Causse (1989). Perak, Sabah.

Marchantiastreimannii Bischl. – Bischler-Causse (1989). Sabah.

Marchantiasubgeminata Steph. – Bischler-Causse (1989). Sabah.

Marsupellaemarginata (Ehrh.) Dumort. – Kitagawa (1967), Gradstein and Váňa (1987), Váňa (1991a). Sabah.

Marsupellastoloniformis N.Kitag. – Kitagawa (1967), Váňa (1991a). Sabah.

Marsupellavermiformis (R.M.Schust.) Bakalin et Fedosov. – Bakalin et al. (2021). Sabah.

Mastigopelmapulvinulatum (De Not.) Grolle – De Notaris (1874 as Mastigobryumpulvinulatum), Schiffner (1898 as Bazzaniapulvinulata), Herzog (1950 as M.bilobum). Sarawak.

Mastigophoradiclados (F.Weber) Nees – De Notaris (1874 as Sendtneradicladosvar.borneensis), Verdoorn (1932c), Herzog (1950, 1952b), Johnson (1958), Inoue (1967a, 1968b), Tixier (1971), Kitagawa and Kodama (1973), Kamimura (1975), Kodama (1976), Kürschner (1990), Ludwiczuk and Asakawa (2010), Ng et al. (2017). Kedah, Pahang, Johor, Sarawak, Sabah.

Mastigophoradiclados var. borneensis (De Not.) Schiffn. – Schiffner (1898). Sarawak.

Mastigophoradiclados var. ramentifissa Herzog – Herzog (1950). Sarawak.

Mastigophoradiclados var. villosa Herzog – Herzog (1950). Sarawak.

Mesoptychiasubcrispa (Herzog) L.Söderstr. et Váňa – Cheah (2017 as Hattoriellasubcrispa). Pahang.

Metalejeuneacucullata (Reinw., Blume et Nees) Grolle – Herzog (1950 as Microlejeuneasundaica), Mizutani (1966, 1970 both as Lejeuneacucullata), Hattori and Kamimura (1971), Tixier (1971, 1974, 1980a as M.cucullata), Kitagawa and Kodama (1974 as L.cucullata), Kürschner (1990 as L.cucullata), Wolseley et al. (1996 as M.cucullata), Pócs et al. (2020), Pesiu et al. (2021). Kedah, Pahang, Negeri Sembilan, Terengganu, Sarawak, Sabah.

Metzgeriaciliata Raddi – Kuwahara (1965 as M.decipiens), Kürschner (1990 as M.decipiens), Cheah (2017). Pahang, Sabah.

Metzgeriafoliicola Schiffn. – Kuwahara (1965), Kitagawa and Kodama (1974). Sabah.

Metzgeriafrancana Steph. – Kuwahara (1965 as Austrometzgeriafrancana). Sabah.

Metzgeriafurcata (L.) Corda – Kuwahara (1965 as M.molokaiensis), Tixier (1971). Pahang, Sabah.

Metzgeriakinabaluensis (Kuwah.) Masuzaki – Kuwahara (1965 as Apometzgeriapubescensvar.kinabaluensis), Kitagawa and Kodama (1974 as A.pubescensvar.kinabaluensis), Kürschner (1990 as A.pubescensvar.kinabaluensis), Akiyama et al. (2001 as A.pubescensvar.kinabaluensis), Masuzaki (2011). Sabah.

Metzgerialeptoneura Spruce – Herzog (1950 as M.hamata), Kuwahara (1965 as M.borneensis, M.hamata and M.sandei, 1976 as M.thomeensis and M.papulosa, 1984 as M.sharpii), Johnson (1972 as M.hamata), Kitagawa and Kodama (1974 as M.hamata), So (2003a). Pahang, Sarawak, Sabah.

Metzgerialindbergii Schiffn. – Kuwahara (1965 as M.conjugatasubsp.japonica and M.pectinata), So (2003a), Cheah (2017). Pahang, Sabah.

Metzgeriascobina Mitt. – Mitten (1886), Schiffner (1898), Kitagawa (1969c), Thiers (1983), So (2003a). Penang, Selangor, Sarawak.

Microlejeuneaconstricta (Grolle) Grolle – Mizutani (1973 as Harpalejeuneaconstricta), Pócs et al. (2020). Sabah.

Microlejeuneafilicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong – Mizutani (1970 as Drepanolejeuneafilicuspis, 1973 as Harpalejeuneafilicuspis and H.riddleana), Pócs and Lee (2016), Cheah (2017 as H.filicuspis), Pócs et al. (2020). Kelantan, Pahang, Sabah.

Microlejeuneakinabaluensis (Mizut.) Grolle – Mizutani (1973 as Harpalejeuneakinabaluensis), Grolle and Reiner-Drehwald (1999). Sabah.

Microlejeunealunulatiloba Horik. – Bischler et al. (1962 as M.gracillima), Pócs et al. (2020). Malacca, Sabah.

Microlejeuneamammillosa (Mizut.) Grolle – Mizutani (1973 as Harpalejeuneamammillosa), Grolle and Reiner-Drehwald (1999). Sabah.

Microlejeuneaminutissima (Mizut.) Grolle – Mizutani (1973 as Harpalejeuneaminutissima), Grolle and Reiner-Drehwald (1999). Sabah.

Microlejeuneapunctiformis (Taylor) Steph. – Kamimura (1974 as Lejeuneapunctiformis), Pócs and Lee (2016), Pócs et al. (2020), Pesiu et al. (2021, in press). Kelantan, Terengganu, Sabah.

Microlejeuneaspinosa (Mizut.) Grolle – Mizutani (1973 as Harpalejeuneaspinosa), Grolle and Reiner-Drehwald (1999). Sabah.

Mizutaniariccardioides Furuki et Z.Iwats. – Furuki and Iwatsuki (1989), Masuzaki et al. (2010). Pahang, Johor, Sarawak, Sabah.

Mniolomafuscum (Lehm.) R.M.Schust. – Grolle (1965 as Calypogeiabaldwinii), Bisang et al. (2003 as C.fusca), Cheah (2017). Pahang, Sarawak.

Mohamediaborneensis (Steph.) R.L.Zhu et L.Shu – Mizutani (1972b as Pycnolejeuneaborneensis), Piippo (1986 as Lepidolejeuneaborneensis), Zhu et al. (2019). Sarawak, Sabah.

Mohamediabrunnea (Mizut.) R.L.Zhu et L.Shu – Mizutani (1970 as Drepanolejeuneabrunnea), Pócs and Lee (2016 as D.brunnea), Zhu et al. (2019). Pahang, Sabah.

Myriocoleopsisminutissima (Sm.) R.L.Zhu, Y.Yu et Pócs – Pócs et al. (2020). Sabah.

Nardiaassamica (Mitt.) Amakawa – Váňa (1991b). Sabah.

Neolepidozialongitudinalis (Herzog) E.D.Cooper – Herzog (1950 as Lepidozialongitudinalis), Engel and Merrill (2004 as Telaranealongitudinalis). Sarawak.

Neolepidoziamamillosa (Schiffn.) E.D.Cooper – Grolle (1967 as Lepidoziamamillosa), Mizutani (1974b as L.mamillosa). Sabah.

Neolepidoziaophiria (Gottsche) E.D.Cooper – Stephani (1909 as Lepidoziaophiria), Mizutani (1974b as L.ophiria), Pocock et al. (1984 as L.ophiria), Engel and Merrill (2004 as Telaraneaophiria). Pahang, Johor (“Mt. Ophir” (Gunung Ledang) was mistakenly listed as in Malacca), Sabah.

Neolepidoziapapulosa (Steph.) Fulford et J.Taylor – Kitagawa (1973 as Lepidoziapapulosa), Kitagawa and Kodama (1973 as L.papulosa). Sarawak, Sabah.

Neolepidoziawallichiana (Gottsche) Fulford et J.Taylor – De Notaris (1874 as Lepidoziawallichiana), Mitten and Wright (1894 as L.wallichiana), Herzog (1950 as L.wallichiana), Kitagawa and Kodama (1973 as L.wallichiana), Mizutani (1974 as L.wallichiana), Kamimura (1975 as L.wallichiana), Pocock et al. (1984 as L.wallichiana), Kürschner (1990 as L.wallichiana), Wolseley et al. (1996 as L.wallichiana), Bisang et al. (2003 as L.wallichiana). Penang, Pahang, Negeri Sembilan, Sarawak, Sabah.

Notoscyphuslutescens (Lehm. et Lindenb.) Mitt. – Herzog (1950 as N.paroicus), Inoue (1967a), Kitagawa (1969c as N.paroicus), Kitagawa and Kodama (1973 as N.paroicus). Penang, Pahang, Sarawak, Sabah.

Nowelliaborneensis (De Not.) Schiffn. – De Notaris (1874 as Jungermanniacurvifoliavar.borneensis), Schiffner (1898), Herzog (1950), Grolle (1968), Kitagawa and Kodama (1974), Akiyama et al. (2001), Cheah (2017). Pahang, Sarawak, Sabah.

Nowelliacurvifolia (Dicks.) Mitt. – Grolle (1968), Váňa (1993). Johor, Sabah.

Nowellialangii Pearson – Pearson (1922), Grolle (1968), Kitagawa and Kodama (1974), Váňa (1993). Perak, Johor, Sabah.

Odontoschismadenudatum (Mart.) Dumort. subsp. naviculare (Steph.) Gradst., S.C.Aranda et Vanderp. – Kitagawa and Kodama (1974 as O.denudatum), Gradstein and Váňa (1987 as O.denudatum), So (2004 as O.denudatum), Gradstein and Ilkiu-Borges (2015). Pahang, Johor, Sabah.

Odontoschismajishibae (Steph.) L.Söderstr. et Váňa – Kitagawa (1964 as Iwatsukiaexigua), Kürschner (1990 as I.exigua), Váňa (1993 as I.exigua), Gradstein and Ilkiu-Borges (2015). Sabah.

Odontoschismapurpuratum Herzog – Herzog (1950, 1952b), So (2004), Gradstein and Ilkiu-Borges (2015). Pahang, Sarawak, Sabah.

Pallaviciniaambigua (Mitt.) Steph. – Kitagawa and Kodama (1974 as P.fistulosa), Grolle and Piippo (1986). Sabah.

Pallaviciniaindica Schiffn. – Herzog (1950), Grolle (1969), Kitagawa (1971), Johnson (1972). Penang, Pahang, Sarawak, Sabah.

Pallavicinialevieri Schiffn. – Johnson (1972), Grolle and Piippo (1986). Pahang, Sabah.

Pallavicinialyellii (Hook.) Gray – Grolle and Piippo (1986). Sarawak.

Pedinophyllumautoicum (Steph.) Inoue – Kamimura (1975). Sabah.

Pictolejeuneamizutanii Grolle – Mizutani (1970 as Cheilolejeuneapicta), Kitagawa and Kodama (1974 as C.picta), Grolle (1977), Thiers (1986), Wolseley et al. (1996), Bi et al. (2019). Negeri Sembilan, Sarawak, Sabah.

Plagiochilaabietina (Nees) Mont. et Nees – Herzog (1950), Inoue (1984). Sarawak, Sabah.

Plagiochilaarbuscula (Lehm. et Lindenb.) Lindenb. – Kitagawa and Kodama (1974), Kamimura (1975), Inoue (1984, 1989), Kürschner (1990). Pahang, Terengganu, Sabah.

Plagiochilabantamensis (Reinw., Blume et Nees) Mont. – De Notaris (1874 as P.mutabilis), Herzog (1950 as P.mutabilis and P.richardsii), Inoue (1967a as P.lobulata, 1984, 1989), Kitagawa and Kodama (1974), Wolseley et al. (1996), So and Grolle (2000), So (2001), Pesiu et al. (in press). Perak, Pahang, Negeri Sembilan, Terengganu, Sarawak, Sabah.

Plagiochilabicornuta Steph. – Stephani (1903 as P.laxissima), Inoue (1989 as P.laxissima). Cheah (2017). Pahang, Sarawak, Sabah.

Plagiochilablepharophora (Nees) Lindenb. – Herzog (1950 as P.estipulata and P.elegantissima), Inoue (1984, 1989), So and Grolle (2000), So (2001), Cheah (2017). Pahang, Sarawak, Sabah.

Plagiochilaborneensis Steph. – Stephani (1918). Sarawak.

Plagiochilachauviniana Mont. – Herzog (1950 as P.vanikorensis). Sarawak.

Plagiochilaclavatosaccata Steph. – Inoue (1984), So and Grolle (2000 as P.longifolia), So (2001). Perak, Sabah.

Plagiochiladenigrata Inoue – Inoue (1989), So and Grolle (2000). Sabah.

Plagiochiladolichoblasta – Herzog (1950), So and Grolle (2000). Sarawak.

Plagiochilafrondescens (Nees) Lindenb. – De Notaris (1874), Herzog (1950), Kitagawa and Kodama (1974), Kamimura (1975), Inoue (1967a, 1984, 1989). Pahang, Sarawak, Sabah.

Plagiochilagymnoclada Sande Lac. – Inoue (1984, 1989), Kürschner (1990), So and Grolle (2000), So (2001). Sabah.

Plagiochilahampeana Gottsche – Inoue (1968b, 1969, 1984 all as P.cameronensis, 1989), Kürschner (1990). Pahang, Sabah.

Plagiochilaintegrilobula Schiffn. – Inoue (1989). Sabah.

Plagiochilajavanica (Sw.) Nees et Mont – Inoue (1989, also as P. infirma), Kamimura (1975), Cheah (2017). Pahang, Sabah.

Plagiochilajunghuhniana Sande Lac. – Kitagawa and Kodama (1974), Kamimura (1975), Cheah (2017). Pahang, Sabah.

Plagiochilakorthalsiana Molk. – Kitagawa and Kodama (1974 as P.accedens), Inoue (1989), So (2000). Sabah.

Plagiochilakuhliana Sande Lac. – Stephani (1903, 1906). Sarawak.

Plagiochilakurzii Steph. – Kitagawa and Kodama (1974 as P.sambusana), Inoue (1984), Wolseley et al. (1996), So and Grolle (2000), So (2001). Negeri Sembilan, Sarawak, Sabah.

Plagiochilamassalongoana Schiffn. – Kitagawa and Kodama (1974), Inoue (1984). Pahang, Sabah.

Plagiochilanitens Inoue – Inoue (1989), So and Grolle (2000), So (2001). Sabah.

Plagiochilanobilis Gottsche – Inoue (1989). Sabah.

Plagiochilaobtusa Lindenb. – Inoue (1968b). Pahang.

Plagiochilapeculiaris Schiffn. – Inoue (1984, 1989), Kürschner (1990), Renner (2018). Pahang, Sarawak, Sabah.

Plagiochilaperserrata Herzog – Inoue (1975, 1984 both as P.hottae), So and Grolle (2000), So (2001). Sabah.

Plagiochilaphilippinensis Steph. – Inoue (1989). Sabah.

Plagiochilapropinqua Sande Lac. – De Notaris (1874), Schiffner (1898), Stephani (1903), Herzog (1950), Inoue (1984, 1989). Perak, Sarawak, Sabah.

Plagiochilarenitens (Nees) Lindenb. – Inoue (1967a, 1984, 1989), Kürschner (1990). Pahang, Sabah.

Plagiochilasandei Dozy – De Notaris (1874), Schiffner (1898), Herzog (1950 as P.sandeif.remotidens), Kitagawa and Kodama (1974), Kamimura (1975), Inoue (1984, 1989), Müller and Schäfer-Verwimp (1999), Cheah (2017), Renner (2018). Pahang, Sarawak, Sabah.

Plagiochilasciophila Nees – Inoue (1984, 1989), So (2001), Cheah (2017). Pahang, Sarawak, Sabah.

Plagiochilaspathulifolia Mitt. – Kitagawa and Kodama (1974), Inoue (1984, 1989). Sarawak, Sabah.

Plagiochilasumatrana Schiffn. – Inoue (1989). Sabah.

Plagiochilateysmannii Sande Lac. – Kitagawa and Kodama (1974), Inoue (1984, 1989), Grolle and So (1999 as P.fraseri). Perak, Sabah.

Plagiochilatrapezoidea Lindenb. – Inoue (1989). Sabah.

Plagiochilaungarangana Sande Lac. – Stephani (1917 as P.tridens), Inoue (1975 as P.tsutomui, 1984), So and Grolle (2000). Perak, Sabah.

Pleuroziaacinosa (Mitt.) Trevis. – De Notaris (1874 as Physiotiummyriangium), Schiffner (1898), Inoue (1967a), Kitagawa (1979b), Thiers (1993). Selangor, Pahang, Sarawak, Sabah.

Pleuroziaarticulata (Lindb.) Lindb. et Lackström – Kürschner (1990). Sabah.

Pleuroziaconchifolia (Hook. et Arn.) Austin – Herzog (1950), Kitagawa and Kodama (1974), Kürschner (1990), Thiers (1993), Akiyama et al. (2001). Sarawak, Sabah.

Pleuroziagigantea (F.Weber) Lindb. – De Notaris (1874 as P.giganteavar.borneensis), Herzog (1950 as Eopleuroziasimplicissima), Inoue (1967a, 1968b), Kitagawa and Kodama (1974 also as E.simplicissima), Tixier (1974), Kürschner (1990), Thiers (1993), Akiyama et al. (2001), Ludwiczuk and Asakawa (2010). Kedah, Pahang, Malacca, Johor, Sarawak, Sabah.

Pleuroziajohannis-winkleri Herzog – Thiers (1993), Akiyama et al. (2001). Sabah.

Pleuroziasubinflata (Austin) Austin – Kamada et al. (2020). Sabah.

Plicanthushirtellus (F.Weber) R.M.Schust. – Herzog (1950 as Chandonanthushirtellus), Kitagawa (1970 as C.hirtellus), Kitagawa and Kodama (1973 as C.hirtellus), Kürschner (1990 as C.hirtellus), Akiyama et al. (2001 as C.hirtellus), Ludwiczuk and Asakawa (2010 as C.hirtellus), Cheah (2017 as C.hirtellus). Pahang, Sarawak, Sabah.

Podomitriummalaccense (Steph.) Campb. – Kitagawa (1969c), Kitagawa and Kodama (1974), Grolle and Piippo (1986), Campbell (1915), Wolseley et al. (1996). Penang, Negeri Sembilan, Johor, Sarawak, Sabah.

Porellaacutifolia (Lehm. et Lindenb.) Trevis. – Kitagawa and Kodama (1974 as P.acutifoliavar.elbertii), Hattori (1967 as P.johannis-winkleri, 1969 as P.acutifoliavar.johannis-winkleri). Sarawak, Sabah.

Porellacaespitans (Steph.) S.Hatt. subsp. latior (S.Hatt.) S.Hatt. – Hattori (1969 as P.acutifoliasubsp.latior). Pahang.

Porellajaponica (Sande Lac.) Mitt. – Hattori (1967). Sabah.

Porellajavanica (Gottsche) Inoue – Inoue (1967a). Pahang.

Pseudolepicoleaandoi (R.M.Schust.) Inoue – Hattori and Mizutani (1968 as P.trolliisubsp.andoi). Sabah.

Pseudolepicoleatrollii (Herzog) Grolle et Ando – Hattori (1966). Sabah.

Pseudomarsupidiumborneensis (Grolle) Váňa, L.Söderstr., A.Hagborg et von Konrat – Grolle (1972 as Adelanthusborneensis), Kitagawa and Kodama (1974 as A.borneensis). Sabah.

Psilocladaclandestina Mitt. – Herzog (1950), Kitagawa and Kodama (1973), Mizutani (1974), Pocock et al. (1984), Kürschner (1990), Akiyama et al. (2001), Cheah (2017). Pahang, Sarawak, Sabah.

Ptychanthusstriatus (Lehm. et Lindenb.) Nees – Verdoorn (1934a), Herzog (1950), Mizutani (1969), Tixier (1971), Kamimura (1974, 1975), Kitagawa and Kodama (1974). Pahang, Sarawak, Sabah.

Pycnolejeuneacavistipula (Steph.) Mizut. – Mizutani (1970 as P.molaris), Kitagawa and Kodama (1974), Kürschner (1990). Sabah.

Pycnolejeuneacontigua (Nees) Grolle – Mizutani (1970 as P.bancana), Kürschner (1990 as P.bancana), He (1999). Sabah.

Pycnolejeuneagrandiocellata Steph. – Kürschner (1990), Wolseley et al. (1996), He (1999), Schäfer-Verwimp (2006). Pahang, Negeri Sembilan, Sarawak.

Pycnolejeuneagrossiloba Steph. – Hoffmann (1935). Johor.

Pycnolejeuneasphaeroides (Sande Lac.) J.B.Jack et Steph. – Kodama (1976 as P.kinabaluensis), He (1999), Schäfer-Verwimp (2006). Pahang, Sabah.

Radulaacuminata Steph. – Yamada (1973, 1979, 1989), Wolseley et al. (1996), Pesiu et al. (2021, in press). Selangor, Negeri Sembilan, Terengganu, Sabah.

Radulaamentulosa Mitt. – Herzog (1950), Yamada (1979, 1989), Cheah (2017). Pahang, Sarawak, Sabah.

Radulaamoena Herzog – Yamada (1979), Yamada and Piippo (1989). Sarawak, Sabah.

Radulaanceps Sande Lac. – De Notaris (1874), Schiffner (1898), Herzog (1950 as R.laciniata), Castle (1961), Inoue and Miller (1965), Yamada (1973, also as R.apiculata, 1979, 1989), Kitagawa and Kodama (1974), Wolseley et al. (1996), Cheah (2017 as R.apiculata). Perak, Pahang, Negeri Sembilan, Sarawak, Sabah.

Radulaassamica Steph. – Pócs et al. (2014), Pesiu et al. (2021, in press). Selangor, Pahang, Terengganu.

Radulaborneensis Steph. – Inoue (1967a), Castle (1936), Yamada (1979, 1989). Pahang, Sarawak, Sabah.

Radulacaduca K.Yamada – Cheah (2017). Pahang.

Radulacampanigera Mont. – Castle (1936), Yamada (1973, 1979, 1989). Perak, Sabah.

Radulacavifolia Gottsche, Lindenb. et Nees – Yamada (1973, 1979, 1989). Sabah.

Radulaceylanica K.Yamada – Yamada (1979). Pahang.

Raduladensifolia Castle – Yamada (1973, 1979, 1989). Sabah.

Radulafalcata Steph. – Cheah (2017). Pahang.

Radulaformosa (Spreng.) Nees – Inoue (1967a as R.novae-guineae), Yamada (1973 as R.novae-guineae, 1979, 1989), Kitagawa and Kodama (1974), Tixier (1974). Kedah, Pahang, Sabah.

Radulagedena Gottsche – Yamada (1973, 1979, 1989). Sabah.

Radulagrandilobula Promma et Chantanaorr. – Sarimi et al. (2021), Zhang et al. (2021), Pesiu et al. (in press). Terengganu, Sarawak.

Radulaiwatsukii K.Yamada – Yamada (1979, 1989), Yamada and Piippo (1989). Sabah.

Radulajavanica Gottsche – De Notaris (1874), Schiffner (1898), Herzog (1950), Yamada (1973, 1979 also as R.yangii, 1989), Kitagawa and Kodama (1974), Wolseley et al. (1996), Pesiu et al. (in press). Selangor, Pahang, Negeri Sembilan, Terengganu, Sarawak, Sabah.

Radulakinabaluensis K.Yamada – Yamada (1979, 1989). Sabah.

Radulalacerata Steph. – Castle (1961), Yamada (1979, 1989), Wolseley et al. (1996). Negeri Sembilan, Sarawak, Sabah.

Radulalingulata Gottsche – Cheah (2017). Pahang.

Radulamadagascariensis Gottsche – Yamada (1979, 1989). Sabah.

Radulamizutanii K.Yamada – Yamada (1973, 1979, 1989), Yamada and Piippo (1989). Sabah.

Radulamorobeana K.Yamada et Piippo – Kürschner (1990). Sabah.

Radulamultiflora Schiffn. – Yamada (1979, 1989). Sabah.

Radulanymannii Steph. – Tixier (1971, 1980a), Yamada (1973, 1979, 1989), Kitagawa and Kodama (1974), Pesiu et al. (2021, in press). Pahang, Terengganu, Sabah.

Radulaobscura Mitt. – Yamada (1973, 1979, 1989). Sabah.

Radulaphilippinensis K.Yamada – Yamada (1979). Selangor.

Radulaprotensa Lindenb. – Herzog (1952b), Wolseley et al. (1996). Negeri Sembilan, Sarawak.

Radularetroflexa Taylor – Inoue (1968b as R.miqueliana), Tixier (1974 as R.miqueliana). Yamada (1979, also as R.retroflexavar.fauciloba, 1989). Kedah, Selangor, Pahang, Terengganu, Sabah.

Radulasumatrana Steph. – Kamimura (1975), Yamada (1979, 1989). Sarawak, Sabah.

Radulatabularis Steph. – Yamada (1973 as R.indica, 1979, 1989), Cheah (2017). Pahang, Sabah.

Radulatjibodensis K.I.Goebel – Kitagawa (1969c), Yamada (1973, 1979, 1989), Kamimura (1975), Kitagawa and Kodama (1974), Pócs et al. (2020), Pesiu et al. (2021, in press). Penang, Pahang, Terengganu, Sabah.

Radulaventricosa Steph. – Yamada (1973, 1979, 1989 all as R.subpallens), Kitagawa and Kodama (1974 as R.subpallens). Sarawak, Sabah.

Radulavrieseana Sande Lac. – Kitagawa and Kodama (1974), Yamada (1979, 1989), Wolseley et al. (1996). Negeri Sembilan, Sarawak, Sabah.

Rebouliahemisphaerica (L.) Raddi subsp. australis R.M.Schust. – Gepp (1914 as R.hemisphaericavar.javanica). Sabah.

Riccardiaalbomarginata (Steph.) Schiffn. – Furuki (1995), Cheah (2017). Pahang, Sabah.

Riccardiaaspera (Steph.) Grolle – Furuki (1998). Sabah.

Riccardiabaumannii Hürl. – Furuki (1994). Sabah.

Riccardiacrassa (Schwägr.) C.Massal. – Herzog (1950 as R.scabra), Johnson (1958 as R.scabra), Kitagawa and Kodama (1974 as R.scabra). Pahang, Sarawak, Sabah.

Riccardiacrassiretis Schiffn. – Furuki (2001). Pahang, Sabah.

Riccardiacrenulata Schiffn. – Furuki (2001). Sabah.

Riccardiadeguchii Furuki et K.T.Yong – Furuki et al. (2013). Pahang.

Riccardiaelata (Steph.) Schiffn. – Herzog (1950 also as R.ridleyi), Cheah (2017). Pahang, Sarawak.

Riccardiafruticosa (Steph.) Furuki – Furuki (1998), Cheah (2017). Pahang, Sabah.

Riccardiagrollei Furuki – Furuki (1999), Cheah (2017). Pahang, Sabah.

Riccardiagrossitexta (Steph.) Furuki – Furuki (1994). Johor, Sarawak, Sabah.

Riccardiahattorii Furuki – Furuki (1994). Sabah.

Riccardiaheteroclada Schiffn. – Herzog (1950), Furuki (1998), Cheah (2017). Pahang, Sarawak.

Riccardiainconspicua (Steph.) Reeb et Bardat – Furuki (1994 as R.tenuicostata). Johor, Sarawak, Sabah.

Riccardiajackii Schiffn. – Johnson (1958). Pahang.

Riccardianobilis (Steph.) Schiffn. – Stephani (1893, 1900), Schiffner (1898). Sarawak.

Riccardiaparvula Schiffn. – Johnson (1958, 1972), Furuki (2001 as Aneurasubcrenulata). Perak, Pahang, Sarawak, Sabah.

Riccardiapindensis Hewson – Furuki (1995). Sabah.

Riccardiaplaniflora (Steph.) S.Hatt. var. aequatorialis Furuki – Furuki (1997), Cheah (2017). Pahang, Sabah.

Riccardiasumatrana Schiffn. – Herzog (1950). Sarawak.

Ricciafluitans L. – Gepp (1914). Sabah.

Ricciocarposnatans (L.) Corda – Gepp (1914). Sabah.

Saccogynidiumgoebelii (Herzog) Grolle – Inoue (1967a, 1968b). Pahang.

Saccogynidiummuricellum (De Not.) Grolle – De Notaris (1874 as Chiloscyphusmuricellus), Schiffner (1898), Mitten and Wright (1894 as Saccogynamuricella), Schiffner (1898 as C.muricellus), Kitagawa and Kodama (1974), Piippo (1989). Sarawak, Sabah.

Saccogynidiumrigidulum (Nees) Grolle – Herzog (1950 as Saccogynarigidula), Grolle (1960, 1963 both as Saccgynidiumjugatum), Inoue (1967a). Perak, Pahang, Johor, Sarawak, Sabah.

Sandeothallusradiculosus (Schiffn.) R.M.Schust. – Campbell and Williams (1914), Pocock et al. (1984), Crandall-Stotler and Stotler (2007). Perak, Pahang, Sabah.

Scapaniajavanica Gottsche – Herzog (1950 as S.javanicaf.amplifolia), Schäfer-Verwimp (2009), Ludwiczuk and Asakawa (2010). Pahang, Sarawak, Sabah.

Scapanialepida Mitt. – Mitten and Wright (1894), Hattori (1964), Kitagawa and Kodama (1973), Akiyama et al. (2001). Sabah.

Scapaniaornithopodioides (With.) Waddell. – Kodama and Narita (1974). Sabah.

Schiffneriahyalina Steph. – Kitagawa (1973), Kitagawa and Kodama (1974), Grolle and Piippo (1984). Pahang, Sabah.

Schiffneriolejeuneacumingiana (Mont.) Gradst. – Lee et al. (2013). Perak.

Schiffneriolejeuneanymannii (Steph.) Gradst. et Terken. – Stephani (1912 as Ptychocoleuslongispicus), Verdoorn (1934a as P.longispicus), Gradstein and Terken (1981). Sarawak.

Schiffneriolejeuneaomphalanthoides Verd. – Gradstein (2015). Sabah.

Schiffneriolejeuneapulopenangensis (Gottsche) Gradst. – Gottsche et al. (1845 as Phragmicomapulopenangensis), Stephani (1890 as P.pulopenangensis, Phragmicomatumida as Ptychocoleuscranstonii and P.tridens), Schiffner (1898 as Acrolejeuneapulopenangensis), Verdoorn (1934a as Ptychocoleuspulopenangensis), Herzog (1950 as Ptychocoleuspulopenangensis), Kitagawa (1969a as P.pulopenangensis), Mizutani (1969 as Ptychocoleuspulopenangensis), Thiers and Gradstein (1989), Haerida et al. (2010), Lee et al. (2013). Penang, Pahang, Sarawak, Sabah.

Schiffneriolejeuneatumida (Nees) Gradst. – Nees (1838 as Ptychanthustumidus), Stephani (1890 as Phragmicomatumida, 1912 as Ptychocoleussarawakensis and P.squarrosifolius), Schiffner (1898 as Acrolejeuneatumida), Verdoorn (1933 as Phragmicomatumida, 1934a as, Ptychocoleussarawakensis), Inoue (1967a as Ptychocoleussarawakensis), Mizutani (1969 as Ptychocoleussarawakensis and P.haskarlianus), Kitagawa (1971 as Ptychocoleussarawakensis), Gradstein and Terken (1981), Lee et al. (2013), Gradstein (2015), Pócs et al. (2020). Penang, Perak, Pahang, Sarawak, Sabah.

Schistochilaacuminata Steph. – Herzog (1950), Kitagawa and Kodama (1973 as S.wrayana), So (2003b), Ng et al. (2016). Perak, Sarawak, Sabah.

Schistochilaaligera (Nees et Blume) J.B.Jack et Steph. – De Notaris (1874 as Gottscheaaligera, G.aligeriformis and G.philippinensis), Schiffner (1898 as G.gaudichaudii and S.aligeriformis), Herzog (1950 also as S.philippinensis), Kitagawa (1969c as S.philippinensis, 1973), Kitagawa and Kodama (1973, also as S.philippinensis), So (2003b). Penang, Selangor, Pahang, Johor, Sarawak, Sabah.

Schistochilabeccariana (De Not.) Trevis. – De Notaris (1874 as Gottscheabeccariana), Schiffner (1898), So (2003b). Sarawak, Sabah.

Schistochilablumei (Nees) Trevis. – Gepp (1914 as S.wallisii), Inoue (1967a), Kitagawa and Kodama (1973), So (2003b). Perak, Kelantan, Pahang, Sarawak, Sabah.

Schistochiladoriae (De Not) Trevis. – De Notaris (1874 as Gottscheadoriae), Schiffner (1898), So (2003b). Pahang, Sarawak, Sabah.

Schistochilareinwardtii (Nees) Schiffn. – So (2003b), Cheah (2017). Pahang, Sabah.

Schistochilarubriseta Steph. – Kürschner (1990). Sabah.

Schistochilasciurea (Nees) Schiffn. – De Notaris (1874 as Gottscheasciurea), Schiffner (1898), Herzog (1950), Inoue (1967a, 1968b), Kitagawa and Kodama (1973), Kürschner (1990), So (2003b). Perak, Pahang, Sarawak, Sabah.

Soellaspinistipula (Mizut.) R.L.Zhu et L.Shu – Mizutani (1970 as Pycnolejeuneaspinistipula), Bi et al. (2019 as Leptolejeuneaspinistipula), Shu et al. (2021). Sarawak, Sabah.

Solenostomaappressifolium (Mitt.) Váňa et D.G.Long – Amakawa (1969b as Jungermanniakinabalensis and J.decolyana), Váňa (1991b as J.appressifolia). Sabah.

Solenostomaariadne (Taylor) Váňa et D.G.Long – Inoue (1967a as Jungermanniaariadne), Amakawa (1968 as J.ariadne), Kitagawa and Kodama (1973 as J.ariadne), Váňa (1991b, 1972 both as J.ariadne). Penang, Perak, Pahang, Negeri Sembilan, Sabah.

Solenostomaborneense (Amakawa) Váňa, Hentschel et Heinrichs – Amakawa (1970 as Jungermanniaborneensis), Váňa (1991b as J.borneensis). Sabah.

Solenostomacomatum (Nees) C.Gao – Inoue (1967a as Jungermanniacomata). Pahang.

Solenostomahaskarlianum (Nees) Váňa et D.G.Long – Váňa (1972, 1975, 1991b all as J.haskarliana). Perak, Pahang, Sarawak, Sabah.

Solenostomajavanicum (Schiffn.) Steph. – Váňa (1991b as Jungermanniaherzogiana). Sabah.

Solenostomapseudocyclops (Inoue) Váňa et D.G.Long – Amakawa (1969b as Jungermanniapseudocyclops). Sabah.

Solenostomariclefii Váňa et D.G.Long – Amakawa (1969b as Jungermanniagrollei), Váňa (1991b as J.grollei). Sabah.

Solenostomasphaerocarpum (Hook.) Steph. – Gradstein and Váňa (1987 as Jungermanniasphaerocarpa), Váňa (1991b as J.sphaerocarpa). Sabah.

Solenostomastephanii (Schiffn.) Steph. – Amakawa (1969b as Jungermanniastephanii and J.mizutanii), Váňa (1991b as J.stephanii). Sabah.

Solenostomastrictum (Schiffn.) Váňa, Hentschel et Heinrichs – Amakawa (1969b as Jungermanniastricta), Váňa (1991b as J.stricta). Sabah.

Solenostomatetragonum (Lindenb.) Váňa et D.G.Long – Herzog (1950 as Plectocoleaparabolica), Váňa (1991b as Jungermanniatetragona). Sarawak, Sabah.

Solenostomatruncatum (Nees) Váňa et D.G.Long – Stephani (1901 as Jungermanniapolyrhiza), Horikawa (1943 as Aploziapolyrhiza), Herzog (1950 as Plectocoleatruncata), Amakawa (1972 as J.truncata), Váňa (1991 as J.truncata). Penang, Sarawak, Sabah.

Solenostomatuberculiferum (Herzog) Váňa, Hentschel et Heinrichs – Váňa (1991b as Jungermanniatuberculifera). Sabah.

Solenostomavirgatum (Mitt.) Váňa et D.G.Long – Váňa (1991b as Jungermanniavirgata). Sabah.

Southbyaorganensis Herzog – Kodama and Narita (1974 as S.grollei). Sabah.

Sphenolobopsispearsonii (Spruce) R.M.Schust. – Kitagawa (1970 as Cephaloziapearsonii), Kodama and Narita (1974), Kürschner (1990 as S.kitagawae), Váňa (1991c). Sabah.

Sphenolobusminutus (Schreb.) Berggr. – Kitagawa (1970 as Anastrophyllumminutum), Kitagawa and Kodama (1973 as A.minutum), Váňa (1991c as A.minutum). Sabah.

Spruceanthusplaniusculus (Mitt.) X.Q.Shi, Gradst. et R.L.Zhu – Verdoorn (1934a as Archilejeuneamariana and Spruceanthusmarianus), Mizutani (1969 as S.marianus), Kitagawa and Kodama (1974 as S.marianus), Lee et al. (2013 as A.planiuscula). Penang, Sarawak, Sabah.

Spruceanthuspolymorphus (Sande Lac.) Verd. – Verdoorn (1934a), Mizutani (1969), Kitagawa and Kodama (1974), Wolseley et al. (1996), Lee et al. (2013). Perak, Pahang, Negeri Sembilan, Sabah.

Spruceanthussemirepandus (Nees) Verd. – Verdoorn (1934a), Sun et al. (2018). Kedah, Sabah.

Spruceanthussulcatus (Nees) Gradst. – Verdoorn (1934a as Ptychanthussulcatus), Mizutani (1969 as P.sulcatus). Sabah.

Stictolejeuneabalfourii (Mitt.) E.W.Jones – Herzog (1950 as S.richardsii), Mizutani (1969 as S.richardsii), Gradstein (1985), Thiers and Gradstein (1989). Sarawak, Sabah.

Symphyogynopsisgottscheana (Mont. et Nees) Grolle – Grolle and Piippo (1986 as S.filicum), Cheah (2017). Perak, Pahang.

Syzygiellaautumnalis (DC.) K.Feldberg, Váňa, Hentschel et Heinrichs – Akiyama et al. (2001 as Jamesoniellaautumnalis). Sabah.

Syzygiellacontracta (Reinw., Blume et Nees) Gradst. et G.E.Lee – Kitagawa (1970 as Cuspidatulacontracta), Váňa (1991c as Jamesoniellacontracta). Sabah.

Syzygiellaflaccida (Steph.) Gradst. et G.E.Lee – Grolle (1971 as Anomacaulisflaccidus), Váňa (1991c as A.flaccidus). Sabah.

Syzygiellaflexicaulis (Nees) Gradst. et G.E.Lee – Inoue (1967a as Jamesoniella flexicaulis), Amakawa (1969a as J.flexicaulisvar.mizutanii), Grolle (1971 as J. flexicaulis), Kürschner (1990 as J.flexicaulis), Váňa (1991c as J. flexicaulis). Pahang, Sabah.

Syzygiellanipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs – Amakawa (1969a as Jamesoniellaperverrucosa), Grolle (1971 as J.nipponica), Kürschner (1990 as J.nipponica), Váňa (1991c as J.nipponica). Sabah.

Syzygiellaovalifolia Inoue – Váňa (1991c). Sabah.

Syzygiellasecurifolia (Nees) Inoue – Inoue (1966, 1967a, 1968b all as S.variegata), Kitagawa and Kodama (1974 as S.variegata). Pahang, Sabah.

Syzygiellasonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs – Amakawa (1969b as Jungermanniaiwatsukii), Kitagawa and Kodama (1973 as J.iwatsukii), Váňa (1991c as Cryptochilagrandiflora). Sabah.

Syzygiellasubintegerrima (Reinw., Blume et Nees) Spruce – Inoue (1966, 1967a, 1968a), Kitagawa and Kodama (1974), Váňa (1991c). Pahang, Sabah.

Telaraneamajor (Herzog) J.J.Engel et G.L.Merr – Herzog (1950 as Arachniopsismajor), Kitagawa and Kodama (1973 as A.major), Mizutani (1974b as A.major), Manuel (1981 as A.major), Engel and Merrill (2004). Pahang, Sarawak, Sabah.

Telaraneamonocera (R.M.Schust. et Grolle) J.J.Engel et G.L.Merr. – Piippo (1985 as Arachniopsismonocera), Engel and Merrill (2004). Johor [“Mt. Ophir” (Gunung Ledang)].

Telaraneapapulosa (Steph.) J.J.Engel et G.L.Merr. – Cheah (2017). Pahang.

Temnomasetigerum (Lindenb.) R.M.Schust. – Kitagawa and Kodama (1973). Sabah.

Tetralophoziafiliformis (Steph.) Urmi – Kitagawa (1973 as Chandonanthusfiliformis), Kitagawa and Kodama (1973 as C. filiformis), Kodama and Narita (1974 as C. filiformis). Sabah.

Tetralophoziapilifera (Steph.) R.M.Schust. – Kürschner (1990 as Chandonanthuspilifer [‘piliferus’]). Sabah.

Thiersianthussilamensis R.L.Zhu et L.Shu – Zhu et al. (2017). Sabah.

Thysananthusaculeatus Herzog – Verdoorn (1934a as T.richardsianus), Herzog (1950 as T.richardsianus), Inoue (1967a, 1968b), Mizutani (1969), Kamimura (1974), Thiers (1985), Sukkharak (2015). Pahang, Sarawak, Sabah.

Thysananthusciliaris (Sande Lac.) Sukkharak – Sukkharak (2015). Perak.

Thysananthuscomosus Lindenb. – Lehmann (1844), Stephani (1890), Schiffner (1898), Mizutani (1977), Sukkharak et al. (2011), Sukkharak (2015). Kedah, Penang, Perak, Selangor, Johor, Sarawak, Sabah.

Thysananthusconvolutus Lindenb. – Stephani (1912 as T.laceratus), Verdoorn (1934a), Herzog (1950), Kamimura (1974), Kitagawa and Kodama (1974), Sukkharak et al. (2011), Sukkharak (2015). Penang, Selangor, Kelantan, Pahang, Johor, Sarawak, Sabah.

Thysananthusconvolutus var. laceratus (Steph.) Sukkharak – Sukkharak (2015). Sabah.

Thysananthusfruticosus (Lindenb. et Gottsche) Schiffn. – Verdoorn (1934a), Herzog (1950 as T.fruticosusf.pendulus), Mizutani (1969, 1987), Kitagawa and Kodama (1974), Kürschner (1990), Wolseley et al. (1996), Sukkharak (2015). Pahang, Selangor, Negeri Sembilan, Johor, Sarawak, Sabah.

Thysananthusgottschei (J.B.Jack et Steph.) Steph. – Stephani (1912 as T.borneensis), Verdoorn (1934a), Kamimura (1974), Kitagawa and Kodama (1974), Thiers (1985), Sukkharak (2015). Penang, Pahang, Sarawak, Sabah.

Thysananthusgottschei var. continuus Sukkharak – Sukkharak (2015). Sarawak.

Thysananthusgradsteinii (Sukkharak) Sukkharak et Gradst. – Sukkharak and Gradstein (2014 as Mastigolejeuneagradsteinii). Kelantan, Pahang.

Thysananthushumilis (Gottsche) Sukkharak et Gradst. – Mizutani (1969 as Mastigolejeuneahumilis, 1986b as M.recurvifolia), Kitagawa (1971 as M.humilis), Kamimura (1974 as M.humilis), Sukkharak and Gradstein (2014 as M.humilis), Cheah (2017 as M.humilis). Penang, Pahang, Sarawak, Sabah.

Thysananthusindicus (Steph.) Sukkharak et Gradst. – Sukkharak and Gradstein (2014 as M.indica). Kedah.

Thysananthusligulatus (Lehm. et Lindenb.) Sukkharak et Gradst. – Stephani (1890 as Phragmicomaligulata), Schiffner (1898 as Mastigolejeunealigulata), Verdoorn (1934a as M.ligulata), Mizutani (1969 as M.ligulata), Kitagawa and Kodama (1974 as M.ligulata), Thiers and Gradstein (1989 as M.ligulata), Kürschner (1990), Gradstein et al. (2002 as M.ligulata), Lee et al. (2013 as M.ligulata), Sukkharak and Gradstein (2014 as M.ligulata). Penang, Pahang, Sarawak, Sabah.

Thysananthusreconditus (Steph.) Sukkharak et Gradst. – Mizutani (1969 as Mastigolejeunearecondita), Sukkharak and Gradstein (2014 as M.recondita), Sun et al. (2018). Pahang, Sarawak, Sabah.

Thysananthusrepletus (Taylor) Sukkharak et Gradst. – Mizutani (1969 as Mastigolejeuneaatypos, 1986b as M.repleta), Sukkharak and Gradstein (2014 as M.repleta). Kedah, Selangor, Sabah.

Thysananthusretusus (Reinw., Blume et Nees) B.M.Thiers et Gradst. – Mizutani (1969, 1987 both as T.planus), Tixier (1971). Pahang, Sabah.

Thysananthusspathulistipus (Reinw., Blume et Nees) Lindenb. – Verdoorn (1934a), Herzog (1950 as T.spathulistipusf.borneensis), Inoue (1967a, 1968b), Tixier (1971, 1974), Kamimura (1974), Kitagawa and Kodama (1974), Kodama (1976), Thiers (1985), Kürschner (1990), Sukkharak (2015), Cheah (2017 as T.minor). Kedah, Penang, Perak, Kelantan, Pahang, Negeri Sembilan, Malacca, Johor, Sarawak, Sabah.

Thysananthustruncatus (Mizut.) Sukkharak et Gradst. – Mizutani (1986b as Mastigolejeuneatruncata), Sukkharak and Gradstein (2014 as M.truncata). Sabah.

Thysananthusvirens Ångstr. – Mizutani (1986b as Mastigolejeuneavirens), Sukkharak and Gradstein (2014 as M.virens). Penang, Sabah.

Treubiainsignis K.I.Goebel – Akiyama et al. (2001). Sabah.

Triandrophyllumheterophyllum (Steph.) Grolle – Cheah (2017). Pahang.

Trichocoleamagna T.Katag. – Katagiri et al. (2013). Sarawak.

Trichocoleamollissima (Hook.f et Taylor) Gottsche – Katagiri et al. (2013). Sabah.

Trichocoleapluma (Reinw., Blume et Nees) Mont. – De Notaris (1874), Herzog (1950), Inoue (1965), Kitagawa (1971, 1973), Kitagawa and Kodama (1973), Tixier (1974), Pocock et al. (1984), Kürschner (1990), Akiyama et al. (2001), Ludwiczuk and Asakawa (2010), Katagiri et al. (2013). Kedah, Penang, Pahang, Sarawak, Sabah.

Trichocolearudimentaris Steph. – Katagiri et al. (2013). Pahang.

Trichocoleatomentella (Ehrh.) Dumort. – Gottsche et al. (1845 as T.tomentellaf.javanica), Johnson (1972), Katagiri et al. (2013). Penang, Pahang, Sabah.

Tricholepidozianeesii (Lindenb.) E.D.Cooper – De Notaris (1874), Herzog (1950 as Lepidozianeesii), Inoue (1967a, 1968b both as Telaraneaneesii), Kitagawa and Kodama (1973), Mizutani (1974 as Telaraneaneesii), Akiyama et al. (2001 as Telaraneaneesii), Engel and Merrill (2004 as Telaraneaneesii). Pahang, Sarawak, Sabah.

Tricholepidoziaoctoloba (Del Ros.) E.D.Cooper – Shi and Zhu (2008 as Telaraneaoctoloba). Johor.

Tricholepidoziaquadriseta (Steph.) E.D.Cooper – Pócs et al. (2014 as Telaraneaquadriseta). Pahang.

Tricholepidoziasemperiana (Steph.) E.D.Cooper – Mizutani (1974b as Telaraneasemperiana). Sabah.

Tricholepidoziatrichocoleoides (Herzog) E.D.Cooper – Herzog (1950 as Lepidoziatrichocoleoides), Kitagawa and Kodama (1973 as L.trichocoleoides), Engel and Merrill (2004 as Telaraneatrichocoleoides). Sarawak, Sabah.

Tritomariaexsecta (Schmidel) Loeske – Kitagawa (1970). Sabah.

Tuyamaellaangulistipa (Steph.) R.M.Schust. et Kachroo – Stephani (1896 as Pycnolejeuneaangulistipa), Schiffner (1898 as P.angulistipa), Mizutani (1966, 1970), Tixier (1971, 1973a), Zhu and So (2000b). Perak, Pahang, Johor, Sabah.

Tuyamaellaborneensis Tixier – Tixier (1973a). Sabah.

Tuyamaellamolischii (Schiffn.) S.Hatt. – Tixier (1973a), Pesiu et al. (2021). Kedah, Terengganu.

Tuyamaellaserratistipa S.Hatt. – Herzog (1951 as Pycnolejeuneaappendiculata), Mizutani (1966), Tixier (1973a), Kitagawa and Kodama (1974), Zhu and So (1998a, 2000c), Pócs et al. (2020). Sarawak, Sabah.

Wiesnerelladenudata (Mitt.) Steph. – Akiyama et al. (2001), Ludwiczuk and Asakawa (2010). Sabah.

Zoopsisliukiuensis Horik. – Herzog (1950 as Z.argentea), Kitagawa and Kodama (1973), Manuel (1981), Grolle and Piippo (1984), Pocock et al. (1984), Mizutani and Chang (1986), Bisang et al. (2003). Pahang, Sarawak, Sabah.

Zoopsissetigera K.I.Goebel – Kitagawa and Kodama (1973 as Z.rigida), Schuster (1999). Sarawak, Sabah.

Hornworts (Anthocerotophyta)

Anthocerosangustus Steph. – Mohamed and Yong (2005 as A.formosae), Mohamed et al. (2005 as A.formosae), Lee and Gradstein (2021). Langkawi.

Dendroceroscavernosus J.Haseg. – Hasegawa (1980). Sabah.

Dendrocerosdifficilis Steph. – Chantanaorrapint et al. (2014). Pahang.

Foliocerosglandulosus (Lehm. et Lindenb.) D.C.Bharadwaj – Meijer (1957 as Anthocerosgrandulosus), Tixier (1971 as A.grandulosus). Pahang, Sarawak.

Megaceros flagellaris (Mitt.) Steph. – Herzog (1950 as M.celebensis), Hasegawa (1983), Lee and Gradstein (2021). Pahang, Sarawak, Sabah.

Phaeoceroscarolinianus (Michx.) Prosk. – Johnson (1958 as Anthocerosvalidus). Perak, Pahang.

Phaeomegacerosfoveatus (J.Haseg.) J.C.Villarreal – Hasegawa (2001 as Phaeocerosfoveatus). Sabah.

Further records

The following records from Malaysia (Genting Highlands, Pahang; Cheah 2017) need verification.

Cololejeuneadenticulata (Horik.) S.Hatt.

Cololejeuneadinghuiana R.L.Zhu et Y.F.Wang

Cololejeuneadrepanolejeuneoides (Horik.) R.M.Schust.

Cololejeuneagrossepapillosa (Horik.) N.Kitag.

Cololejeuneaplagiophylla Benedix

Cololejeuneaspinosa (Horik.) Pandé et R.N.Misra

Plagiochilafusca Sande Lac.

Plagiochilagracilis Lindenb. et Gottsche

Plagiochilasalacensis Gottsche

Plagiochilastephanii Schiffn.

Riccardiadiminuta Schiffn.

Riccardiahattorii Furuki

Riccardiamultifida (L.) Gray

Riccardiamultifidoides Schiffn.

Riccardiapumila Furuki

Riccardiasingapurensis Schiffn.

Doubtful records

Anastrophyllumpiligerumf.denticalyx – Reported from Sarawak (Herzog 1950); status unclear.

Aspiromitusvesiculosusvar.latifrons Herzog – Reported from Sarawak (Herzog 1950), but its status is unclear. Aspiromitus is now a synonym of Anthoceros.

Bazzaniaeverettii (Steph.) Meijer nom. inval. (ICN Art. 41.5) – Reported from Sarawak (Stephani 1908 as Mastigobryumeverettii); status unclear.

Bazzaniakarcharias Herzog, nom. inval. (ICN 38.1) – Sarawak (Herzog 1950).

Bazzaniastonii Inoue nom. inval. (ICN Art. 38.1(a) – Reported from Pahang (Inoue 1968b); the record should be checked.

Bazzaniasubhyalina (Steph.) Chuah-Pet. nom. inval. (ICN Art. 41.5) – Reported from Sarawak (Stephani 1924 as Mastigobryumsubhyalinum); status unclear.

Cephaloziellastephanii Douin – Reported from Pahang (Cheah 2017); the record should be checked.

Chiloscyphusceylanicus Tixier nom. inval. (ICN Art. 38.1a) – Reported from Kedah (Tixier 1974); the record should be checked.

Chiloscyphusschiffneri J.J.Engel et R.M.Schust. nom illeg. – Synonym of Lophocoleajavanica Schiffn., its occurrence in Pahang (Cheah 2017) is doubtful.

Chiloscyphus flaccidens Steph. – Reported from Kedah (Tixier 1974); the record should be checked.

Drepanolejeuneablumeivar.angulistipa Herzog nom inval. (ICN Art. 39.1) – Reported from Sabah (Herzog 1936); the record should be checked.

Drepanolejeuneamalayana Tixier (“Grolle”) nom. inval. – Reported from Kedah (Tixier 1974); the record should be checked.

Drepanolejeuneavesiculosasubsp.propagulifera Herzog nom. inval. (ICN Art. 39.1) – Reported from Negeri Sembilan (Wolseley et al. 1996) and Pahang (Inoue 1967a); the record should be checked.

Frullaniaornithocephalaf.magnilobulaandf.retusa – Reported from Sabah (Hattori 1976), but their status is unclear.

Gottscheaintegerrima (Steph.) Grolle et Zijlstra – Accepted name: Schistochilaintegerrima Steph., only known from Vanuatu and New Caledonia (Thouvenot 2021). Its occurrence in Pahang (Cheah 2017) is doubtful and the record is most likely S.nuda Horik., known from Thailand and Southeast Asia.

Radulakurzii Steph. – Reported from Pahang (Cheah 2017); not known in tropical Southeast Asia and very similar to the widespread Asian R.javanica Gottsche.

Symphyogynasimilis Grolle – Reported from Pahang (Cheah 2017); the record should be checked, only known from the Huon Peninsula and Java (Grolle and Piippo 1986).

Telaraneawallichianavar.remotifolia Herzog – Reported from Sarawak (Herzog 1950), but its status is unclear.

Excluded and erroneous records (See Menzel 1988 for further excluded records)

Acrolejeuneaaulacophora (Mont.) Steph. – The record from Sabah (Mizutani 1969 as Ptychocoleusaulacophorus) is doubtful.

Acrolejeuneaborneensis Bonner, nom. inval. from Sarawak (Bonner 1962) is a Schiffneriolejeunea sp. fide Gradstein (1975).

Acrolejeuneasandvicensis (Gottsche) J.Wang et Gradst. – The record from Malaysia (Tixier (1975 as Brachiolejeuneasandvicensis) belongs to Thysananthusreconditus (Sun et al. 2018).

Bazzaniatricrenata (Wahlenb.) Lindb. – This is a Holarctic species, not known in tropical Southeast Asia, and its occurence in Malaysia (Pahang; Cheah 2017) is considered erroneous. Also reported from Sabah by Sande Lacoste (1864 as Mastigobryumdeflexum).

Cheilolejeuneaparvidens B.M.Thiers – The record from Sabah (Pócs et al. 2020) belongs to C.intertexta.

Chiloscyphusminor (Nees) J.J.Engel et R.M.Schust. – This is a Holarctic species, not known in tropical Southeast Asia, and its occurence in Malaysia (Pahang; Cheah 2017) is considered erroneous.

Frullaniaintegristipulavar.emarginata Verd. – Reported from Sabah by Chuah-Petiot (2011) based on Hattori (1982), but the latter author did not report this variety from Malaysia.

Frullaniapulogensis Steph. – Reported from Sabah by Verdoorn (1932a) but it was rejected by Hattori (1978) due to the scarcity of the specimen. No material of F.pulogensis but fragmental shoots of F.mizutanii are present.

Kurziamauiensis (H.A.Mill.) H.A.Mill. – Reported from Pahang (Cheah 2017). Only known from the Pacific Islands and its occurrence in Malaysia needs verification.

Lejeuneadentata Mitt. – Reported from Pahang (Cheah 2017). Synonym of Radulacuspidata Steph., only known in New Zealand.

Lejeuneaulicina (Taylor) Gottsche, Lindenb. et Nees – Accepted name: Microlejeuneaulicina (Taylor) Steph., a Holarctic species, not known in tropical Southeast Asia. Its occurrence in Malaysia (Pahang; Cheah 2017) is considered erroneous.

Metacalypogeiaalternifolia (Nees) Grolle – Reported from Sarawak by Herzog (1950 as Calypogeiaalternifolia). According to Grolle (1964), this record is erroneous.

Metzgeriaalbinea Spruce – This is a Neotropical species according to So (2002), and its occurrence in Malaysia (Pahang; Cheah 2017) needs verification.

Mniolomastamatotonum M.A.M.Renner et E.A.Br. – Reported from Pahang (Cheah 2017); an Australasian species, not known in tropical Southeast Asia.

Ricciatreubiana Steph. – Reported from Peninsular Malaysia by Chuah Petiot (2011) based on Meijer (1958) but the latter author reported this species from Singapore. Ricciatreubiana is a common species and often found in gardens, recreational parks and secondary forests. Its occurrence in Malaysia is to be expected.

Odontoschismadenudatum (Nees) Dumort. – Reported from Sabah by various authors (see Chuah-Petiot 2011), but the material belongs to O.denudatumsubsp.naviculare (see Gradstein and Ilkiu-Borges 2015).

Odontoschismasphagni (Dicks.) Dumort. – Reported from Sabah by Mitten and Wright (1894), but the record is considered erroneous (Váňa 1993). According to Gradstein and Ilkiu-Borges (2015)O.sphagni is a species from Europe and North America that does not occur in Asia.

Saccogynidiumirregularospinum C.Gao, T.Cao et M.J.Lai – Reported from Pahang (Cheah 2017); a temperate Asiatic species from Sikkim and China, not known in tropical Southeast Asia.

Schistochilanyamanii – Reported from Pahang (Cheah 2017), but the species name does not exist.

Solenostomaobliquifolium (Schiffn.) Váňa, Hentschel et Heinrichs – Reported from Sabah by Chuah-Petiot (2011) based on Váňa (1991b as Jungermanniaobliquifolia) but the latter author did not report this species in Sabah.

Telaraneagranulata J.J.Engel et G.L.Merr. – Reported from Pahang (Cheah 2017); endemic to New Zealand, not known in tropical Southeast Asia.

Telaraneapatentissima (Hook.f et Taylor) E.A.Hodgs. – Reported from Pahang (Cheah 2017); an Australasian species from Australia and New Zealand, not known in tropical Southeast Asia.

Thysananthusauriculatus (Wilson et Hook.) Sukkharak et Gradst. – Reported from Pahang by Lee et al. (2013 as Mastigolejeuneaauriculata). According to Sukkharak and Gradstein (2014, 2017) T.auriculatus is an Afro-American species; Asiatic records of this species belong to T.humilis.

Wettsteiniainversa (Sande Lac.) Schiffn. – Reported from Sabah by Frahm et al. (1990) and Chuah-Petiot (2011) based on Menzel (1988), but the latter author reported this species only from Kalimantan Selatan (leg. Korthals, paratype of Plagiochilascabra Sande Lac.). Wettsteiniainversa is a widespread Malesian species (see Piippo 1984) and its occurrence in Malaysia is to be expected.

Synonyms

New synonymy published after the appearance of World Checklist (Söderström et al. 2016) is provided with a reference.

Acantholejeuneaspinistipula (Herzog) R.M.Schust. ≡ Ceratolejeuneaspinistipula (Herzog) R.L.Zhu, L.Shu, Qiong He et Y.M.Wei

Acrolejeuneapulopenangensis (Gottsche) Schiffn. ≡ Schiffneriolejeuneapulopenangensis (Gottsche) Gradst.

Acrolejeuneatumida (Nees) Steph. ≡ Schiffneriolejeuneatumida (Nees) Gradst.

Acromastigumbidenticulatum A.Evans = A.bancanum (Sande Lac.) A.Evans

Acromastigumdenticulatum A.Evans = A.bifidum (Steph.) A.Evans

Acromastigumemarginatum Herzog ≡ Bazzaniamenzelii E.D.Cooper

Acromastigumgemmiferum N.Kitag. et T.Kodama = A.obliquatum (Mitt.) A.Evans

Adelanthusborneensis Grolle ≡ Pseudomarsupidiumborneensis (Grolle) Váňa, L.Söderstr., A.Hagborg et von Konrat

Allorgellachangiana Tixier = Otolejeuneasemperiana (Steph.) Grolle

Anastrophyllumappendiculatum N.Kitag = A.auritum (Lehm.) Steph.

Anastrophyllumdivergens Herzog = A.fissum Steph.

Anastrophyllumimbricatum (Gottsche, Lindenb. et Nees) Steph. = A.piligerum (Nees) Steph.

Anastrophyllumminutum (Schreb.) R.M.Schust. ≡ Sphenolobusminutus (Schreb.) Berggr.

Aneurasubcrenulata Herzog = Riccardiaparvula Schiffn.

Anomacaulisflaccidus (Steph.) Grolle ≡ Syzygiellaflaccida (Steph.) Gradst. et G.E.Lee (Lee and Gradstein 2021)

Anthocerosformosae Steph. = A.angustus Steph.

Anthocerosglandulosus Lehm. et Lindenb. ≡ Foliocerosglandulosus (Lehm. et Lindenb.) Bhardwaj

Anthocerosvalidus Steph. = Phaeoceroscarolinianus (Michx.) Prosk.

Aploziapolyrhiza (Hook.) Horik. = Solenostomatruncatum (Nees) Váňa et D.G.Long

Apometzgeriapubescensvar.kinabaluensis Kuwah. = Metzgeriakinabaluensis (Kuwah.) Masuzaki

Aphanolejeuneaborneensis (Herzog) Pócs = Cololejeuneapapillosa (K.I.Goebel) Mizut.

Aphanolejeuneamicroscopicavar.borneensis Herzog = Cololejeuneapapillosa (K.I.Goebel) Mizut.

Aphanolejeuneaproboscoidea (K.I.Goebel) R.M.Schust. = Cololejeuneadiaphana A.Evans

Arachniopsismajor Herzog ≡ Telaraneamajor (Herzog) J.J.Engel et G.L.Merr.

Arachniopsismonocera R.M.Schust. et Grolle ≡ Telaraneamonocera J.J.Engel et G.L.Merr.

Archilejeuneamariana auct. (non (Gottsche) Steph.) = Spruceanthusplaniusculus (Mitt.) X.Q.Shi, R.L.Zhu et Gradst. (Shi et al. 2015)

Archilejeuneaplaniuscula (Mitt.) Steph. ≡ Spruceanthusplaniusculus (Mitt.) X.Q.Shi, R.L.Zhu et Gradst. (Shi et al. 2015)

Austrometzgeriafrancana (Steph.) Kuwah. ≡ Metzgeriafrancana Steph.

Bazzaniaassamica (Steph.) S.Hatt. = B.tridensvar.assamica (Steph.) Pócs

Bazzaniaaustralis Gottsche, Lindenb. et Nees = B.tridens (Reinw., Blume et Nees) Trevis.

Bazzaniabancana (Sande Lac.) Trevis. ≡ Acromastigumbancanum (Sande Lac.) A.Evans

Bazzaniaconcinna (De Not.) Trevis. = B.intermedia (Gottsche et Lindenb.) Trevis.

Bazzaniacoreana Herzog = B.tridens (Reinw., Blume et Nees) Trevis.

Bazzaniaechinatiformis (De Not.) Trevis. ≡ Acromastigumechinatiforme (De Not.) A Evans

Bazzaniaferox (De Not.) Trevis. = B.harpago (De Not.) Schiffn.

Bazzaniaremotifolia Herzog nom. illeg. ≡ B.herzogiana Meijer

Bazzaniapalmatifida (Steph.) Grolle = B.subtilis (Sande Lac.) Trevis.

Bazzaniafallaxf.fissa Herzog = B.fallax (Sande Lac.) Schiffn.

Bazzanialingana (De Not.) Trevis. ≡ Acromastigumlinganum (De Not.) A.Evans

Bazzanianotarisii (Steph.) Schiffn. = Acromastiguminaequilaterum (Lehm. et Lindenb.) A.Evans

Bazzaniapulvinulata (De Not.) Schiffn. ≡ Mastigopelmapulvinulatum (De Not.) Grolle

Bazzaniarecurvavar.pallens (De Not.) Schiffn. = B.recurva (Mont.) Trevis.

Bazzaniarecurvolimbata (Steph.) N.Kitag. = B.revoluta (Steph.) N.Kitag.

Bazzaniarichardsii Herzog = B.calcarata (Sande Lac.) Schiffn.

Bazzaniaserrulata (Mitt.) Steph. = B.erosa (Reinw., Blume et Nees) Trevis.

Bazzaniatridensf.minutissima (Kamim.) Pócs = B.tridens (Reinw., Blume et Nees) Trevis.

Bazzaniavaga (De Not.) Trevis. = B.erosa (Reinw., Blume et Nees) Trevis.

Calypogeiabaldwinii Austin = Mniolomafuscum (Lehm.) R.M.Schust.

Calypogeiafusca (Lehm.) Steph. ≡ Mniolomafuscum (Lehm.) R.M.Schust.

Campylolejeuneapeculiaris (Herzog) Amakawa = Cololejeuneainflectens (Mitt.) Benedix

Campylolejeuneapusilla Mizut. ≡ Cololejeuneahattoriana Mizut. et Pócs

Caudalejeuneacircinata Steph. = C.cristiloba (Steph.) Gradst.

Caudalejeuneaserrata Steph. = C.reniloba (Gottsche) Steph.

Caudalejeunearecurvistipula (Gottsche) Schiffn. = C.reniloba (Gottsche) Steph.

Caudalejeuneastephanii Steph. nom. inval. = C.reniloba (Gottsche) Steph.

Cephaloziapearsonii (Spruce) Steph. ≡ Sphenolobopsispearsonii (Spruce) R.M.Schust.

Cephaloziellatagawae N.Kitag. = Cylindrocoleatagawae (N.Kitag.) R.M.Schust.

Cephaloziellawillisana (Steph.) N.Kitag. = Cylindrocoleakiaeri (Austin) Douin

Ceratolejeuneaoceanica (Mitt.) Steph. = C.belangeriana (Gottsche) Steph.

Ceratolejeunearyukyuensis Amakawa = C.belangeriana (Gottsche) Steph.

Ceratolejeuneatahitensis Steph. = C.cornuta (Lindenb.) Steph.

Chandonanthusfiliformis Steph. ≡ Tetralophoziafiliformis (Steph.) Urmi

Chandonanthushirtellus (F.Weber) Mitt. ≡ Plicanthushirtellus (F.Weber) R.M.Schust.

Chandonanthuspilifer Steph. ≡ Tetralophoziapilifera (Steph.) R.M.Schust.

Cheilolejeuneaexcisula (Steph.) Mizut. = C.incisa (Gottsche) R.M.Schust.

Cheilolejeuneaimbricata (Nees) S.Hatt. = C.trapezia (Nees) R.M.Schust. et Kachroo

Cheilolejeunealarsenii Mizut. = C.adnatavar.autoica Gradst. et Ilk.-Borg. (Bastos and Gradstein 2020).

Cheilolejeunealongiloba (G.Hoffm.) R.M.Schust. = C.trapezia (Nees) R.M.Schust. et Kachroo

Cheilolejeunealuerssenii (Steph.) Mizut. = C.lindenbergii (Gottsche) Mizut.

Cheilolejeuneameyeniana (Nees, Lindenb. et Gottsche) R.M.Schust. et Kachroo = C.trapezia (Nees) R.M.Schust. et Kachroo

Cheilolejeuneapicta Mizut. ≡ Pictolejeuneamizutanii Grolle

Cheilolejeuneaspathulata Mizut. = C.decursiva (Sande Lac.) R.M.Schust.

Cheilolejeuneaviridula (Herzog) Mizut. = C.trifaria (Reinw., Blume et Nees) Mizut.

Chiloscyphusacutangulus Schiffn. ≡ Heteroscyphusacutangulus (Schiffn.) Schiffn.

Chiloscyphusargutus (Reinw., Blume et Nees) Nees ≡ Heteroscyphusargutus (Reinw., Blume et Nees) Schiffn.

Chiloscyphusaselliformis (Reinw., Blume et Nees) Nees ≡ Heteroscyphusaselliformis (Reinw., Blume et Nees) Schiffn.

Chiloscyphusaselliformisvar.neesii Schiffn. = Heteroscyphusaselliformis (Reinw., Blume et Nees) Schiffn.

Chiloscyphusciliolatus (Nees) J.J.Engel et R.M.Schust. ≡ Cryptolophocoleaciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler et Váňa

Chiloscyphuscoadunatus (Sw.) J.J.Engel et R.M.Schust. = Lophocoleabidentata (L.) Dumort.

Chiloscyphuscommunis Steph. = Heteroscyphuscoalitus (Hook.) Schiffn.

Chiloscyphusconcinnus De Not. = Heteroscyphussucculentus (Gottsche) Schiffn.

Chiloscyphuscostatus (Nees) J.J.Engel et R.M.Schust. ≡ Cryptolophocoleacostata (Nees) L.Söderstr.

Chiloscyphuscuspidatus (Nees) J.J.Engel et R.M.Schust. = Lophocoleabidentata (L.) Dumort.

Chiloscyphusdecurrens (Reinw., Blume et Nees) Nees = Heteroscyphussplendens (Lehm. et Lindenb.) Grolle

Chiloscyphusdensifolius De Not. = Heteroscyphussplendens (Lehm. et Lindenb.) Grolle

Chiloscyphusdiestianus Sande Lac. ≡ Heteroscyphusdiestianus (Sande Lac.) Piippo

Chiloscyphusintegerrimus Schiffn. ≡ Heteroscyphusintegerrimus (Schiffn.) Gradst. et G.E.Lee (Lee and Gradstein 2021).

Chiloscyphuskurzii (Sande Lac.) J.J.Engel et R.M.Schust. ≡ Lophocoleakurzii Sande Lac.

Chiloscyphuslatifolius (Nees) J.J.Engel et R.M.Schust. = Lophocoleabidentata (L.) Dumort.

Chiloscyphusmuricatus (Lehm.) J.J.Engel et R.M.Schust. ≡ Lophocoleamuricata (Lehm.) Nees

Chiloscyphusmuricellus De Not. ≡ Saccogynidiummuricellum (De Not.) Grolle

Chiloscyphusparoicus J.J.Engel et R.M.Schust. ≡ Cryptolophocolealevieri (Schiffn.) L.Söderstr.

Chiloscyphusparvulus Schiffn. ≡ Heteroscyphusparvulus (Schiffn.) Schiffn.

Chiloscyphussikkimensis (Steph.) J.J.Engel et R.M.Schust. ≡ Lophocoleasikkimensis (Steph.) Herzog et Grolle

Chiloscyphussteetziae (De Not.) J.J.Engel et R.M.Schust. ≡ Lophocoleasteetziae De Not.

Chiloscyphussucculentus Gottsche ≡ Heteroscyphussucculentus (Gottsche) Schiffn.

Chiloscyphuswettsteinii Schiffn. ≡ Heteroscyphuswettsteinii (Schiffn.) Schiffn.

Chiloscyphuszollingerivar.borneensis Herzog = Heteroscyphuszollingeri (Gottsche) Schiffn.

Cololejeuneaandrogyna Benedix = C.obliqua (Nees et Mont.) Schiffn.

Cololejeuneaamoena Benedix ≡ C.floccosavar.amoena (Benedix) Pócs

Cololejeuneaastyla Mizut. = C.platyneura (Spruce) A.Evans

Cololejeuneabalansae (Steph.) Mizut. = C.trichomanis (Gottsche) Steph.

Cololejeuneabenedixii Tixier = C.schmidtii Steph.

Cololejeuneabolombensis (Steph.) Vanden Berghen = C.lanciloba Steph.

Cololejeuneaciliatilobula (Schiffn.) Schiffn. = C.inflectens (Mitt.) Benedix

Cololejeuneacrenulata (Pearson) H.A.Mill. = C.angustiflora (Steph.) Mizut.

Cololejeuneafalcatoides Benedix = C.falcata (Horik.) Benedix

Cololejeuneafilicaulis Steph. = C.hildebrandii (Austin) Steph.

Cololejeuneaformosana Mizut. = C.ceratilobula (P.C.Chen) R.M.Schust.

Cololejeuneagoebelii (Schiffn.) Schiffn. = C.trichomanis (Gottsche) Steph.

Cololejeuneainflectidens (Mitt.) Benedix nom. inval. ≡ C.inflectens (Mitt.) Benedix

Cololejeuneajavanica (Steph.) Mizut. = C.angustiflora (Steph.) Mizut.

Cololejeuneajelinekii Steph. = C.obliqua (Nees et Mont.) Schiffn.

Cololejeuneakarstenii K.I.Goebel = C.cordiflora Steph.

Cololejeuneakinabalensis Mizut. = C.macounii (Underw.) A.Evans

Cololejeunealeonidens Benedix = C.ocelloides (Horik.) Mizut.

Cololejeunealeonidensvar.saccata Benedix = C.ocelloides (Horik.) Mizut.

Cololejeuneamackeeana Tixier = C.angustiflora (Steph.) Mizut.

Cololejeuneameijeri Tixier = C.ocelloides (Horik.) Mizut.

Cololejeuneamutabilisf.borneensis Benedix = C.mutabilis Benedix

Cololejeuneamutabilisf.floccodoides Tixier = C.mutabilis Benedix

Cololejeuneamutabilisf.subfalcata Tixier = C.mutabilis Benedix

Cololejeuneanymannii (Steph.) Benedix = C.obliqua (Nees et Mont.) Schiffn.

Cololejeuneaoshimensis (Horik.) Benedix = C.inflata Steph.

Cololejeuneapahangiana Tixier = C.lanciloba Steph.

Cololejeuneapakseana Tixier = C.uchimae Amakawa

Cololejeuneaparoica Mizut. = C.gottschei (Steph.) Mizut.

Cololejeuneapeculiaris (Herzog) Benedix = C.inflectens (Mitt.) Benedix

Cololejeuneaperaffinisvar.elegans Benedix = C.peraffinis (Schiffn.) Schiffn.

Cololejeuneaplaniflora Benedix = Cololejeuneaindica Pandé et R.N.Misra

Cololejeuneapluripunctata Benedix = C.siamensis Steph.

Cololejeuneapusilla Steph. = C.gradsteinii M.J.Lai et R.L.Zhu

Cololejeuneascabriflora Steph = C.obliqua (Nees et Mont.) Schiffn.

Cololejeuneatenellavar.vittata Tixier = C.tenella Benedix

Cololejeuneatrichomanissubsp.cordiflora (Steph.) Pócs ≡ C.cordiflora Steph.

Cololejeuneayulensis (Steph.) Benedix = C.aequabilis (Sande Lac) Schiffn.

Coluraacutifolia Jovet-Ast = C.conica (Sande Lac.) K.I.Goebel

Coluraapiculata (Schiffn.) Steph. = C.leratii (Steph.) Steph.

Colurajavanica Steph. = C.ari (Steph.) Steph.

Coluratrialata (Steph.) Herzog et Zwickel = C.corynophora (Nees, Lindenb. et Gottsche) Trevis.

Colurolejeuneaornata A.Evans ≡ Coluraornata K.I.Goebel

Conoscyphusinflexifolius Mitt. = C.trapezioides (Sande Lac.) Schiffn.

Crossotolejeuneaborneensis Herzog = Lejeuneastenodentata M.A.M.Renner et Pócs

Cryptochilagrandiflora (Lindenb. et Gottsche) Grolle = Syzygiellasonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs

Cuspidatulacontracta (Reinw., Blume et Nees) Steph. ≡ Syzygiellacontracta (Reinw., Blume et Nees) Gradst. et G.E.Lee (Lee and Gradstein 2021)

Cylindrocoleakiaeri (Austin) Váňa ≡ Cephaloziellakiaeri (Austin) Douin (Lee and Gradstein 2021)

Diplasiolejeuneabrachyclada A.Evans = D.cavifolia Steph.

Diplasiolejeuneaincurvata Ast et Tixier = D.cobrensis Steph.

Diplasiolejeuneajavanica Steph. = D.cavifolia Steph.

Diplasiolejeuneaneobrachyclada S.Hatt = D.patelligera Herzog

Diplasiolejeunearudolphiana Steph. = D.unidentata (Lehm. et Lindenb.) Schiffn.

Diploscyphusborneensis De Not. = Conoscyphustrapezioides (Sande Lac.) Schiffn.

Drepanolejeuneabrunnea Mizut. ≡ Mohamediabrunnea (Mizut.) R.L.Zhu et L.Shu

Drepanolejeuneadentata Steph. ≡ Lejeuneastenodentata M.A.M.Renner et Pócs

Drepanolejeuneafilicuspis (Steph.) Mizut. ≡ Microlejeuneafilicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong

Drepanolejeunealevicornuavar.longicornua Herzog = D.longicornua (Herzog) Mizut.

Drepanolejeuneamicholitzii Steph. = D.pentadactyla (Mont.) Steph.

Drepanolejeuneamicholitziivar.angustissima Herzog = D.pentadactyla (Mont.) Steph.

Drepanolejeuneamicholitziivar.brevifolia Herzog = D.pentadactyla (Mont.) Steph.

Drepanolejeuneamicholitziivar.dactylophoroides Herzog ≡ D.pentadactylavar.dactylophoroides (Herzog) Pócs

Drepanolejeuneamuricata (Gottsche) Schiffn. ≡ Dactylophorellamuricata (Gottsche) R.M.Schust.

Drepanolejeuneaspinistipula Herzog ≡ Ceratolejeuneaspinistipula (Herzog) R.L.Zhu, L.Shu, Qiong He et Y.M.Wei

Drepanolejeuneateneraf.goebelii Herzog = D.tenera K.I.Goebel

Drepanolejeuneateneravar.genuina Herzog nom. illeg. = D.tenera K.I.Goebel

Drepanolejeuneatenuis (Nees) Schiffn. = D.angustifolia (Mitt.) Grolle

Drepanolejeuneaternatensisvar.lancispina Herzog = D.ternatensis (Gottsche) Schiffn.

Drepanolejeuneavesiculosaf.robusta Herzog = D.vesiculosa (Mitt.) Steph.

Dumortieracalcicola Campb. = D.hirsutasubsp.nepalensis (Taylor) R.M.Schust.

Dumortieranepalensis (Taylor) Nees ≡ D.hirsutasubsp.nepalensis (Taylor) R.M.Schust.

Dumortieratrichocephala (Hook.) Nees = D.hirsutasubsp.nepalensis (Taylor) R.M.Schust.

Eopleuroziasimplicissima (Herzog) R.M.Schust. = Pleuroziagigantea (F.Weber) Lindb.

Euosmolejeunealuerssenii Steph. = Cheilolejeunealindenbergii (Gottsche) Mizut.

Frullaniaaltemammillata S.Hatt = F.armatifolia Verd.

Frullaniaborneensis Steph. = F.sublignosa Steph.

Frullaniahampeana Nees = F.monocera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees

Frullaniahasskarlianavar.integribracteata (Verd.) S.Hatt. = F.hasskarliana Lindenb.

Frullaniaintermediaf.billardieriana (Nees et Mont.) Verd. = F.intermedia (Reinw., Blume et Nees) Nees

Frullaniaintermediavar.submorokensis S.Hatt = F.intermedia (Reinw., Blume et Nees) Nees

Frullaniajunghuhnianavar.minutissima Verd. = F.junghuhnianavar.tenella (Sande Lac.) Grolle et S.Hatt.

Frullaniaminor Sande Lac. = F.gracilis (Reinw., Blume et Nees) Nees

Frullaniamoniliatasubsp.breviramea (Steph.) Verd. = F.moniliata (Reinw., Blume et Nees) Mont.

Frullaniaornithocephalaf.magnilobula S.Hatt. = F.ornithocephala (Reinw., Blume et Nees) Nees

Frullaniaornithocephalaf.retusa S.Hatt. = F.ornithocephala (Reinw., Blume et Nees) Nees

Frullaniaperminuta S.Hatt. = F.junghuhnianavar.tenella (Sande Lac.) Grolle et S.Hatt.

Frullaniaperversa Steph. = F.junghuhnianavar.tenella (Sande Lac.) Grolle et S.Hatt.

Frullaniaphilippinensis Steph. = F.reflexistipula Sande Lac.

Frullaniapicta Steph. = F.trichodes Mitt.

Frullaniarepandistipulasubsp.spinifera S.Hatt. nom. inval. = F.repandistipulasubsp.spinibractea S.Hatt.

Frullaniaserrataf.crispulodentata Verd. = F.serrata Gottsche

Frullaniasquarrosa (Mont.) Nees = F.ericoides (Nees) Mont.

Frullaniatamariscivar.breviramea (Steph.) S.Hatt. = F.moniliata (Reinw., Blume et Nees) Mont.

Frullaniatenuicaulis Mitt. = F.trichodes Mitt.

Frullaniavethii Sande Lac. = F.trichodes Mitt.

Frullaniavethiif.acutiloba S.Hatt. = F.trichodes Mitt.

Frullaniawallichiana Mitt = F.obscura (Sw.) Mont.

Gottscheaaligera (Nees et Blume) Nees ≡ Schistochilaaligera (Nees et Blume) J.B.Jack et Steph.

Gottscheaaligeriformis De Not. = Schistochilaaligera (Nees et Blume) J.B.Jack et Steph.

Gottscheabeccariana De Not. ≡ Schistochilabeccariana (De Not.) Trevis.

Gottscheadoriae De Not. ≡ Schistochiladoriae De Not) Trevis.

Gottscheagaudichaudii Gottsche = Schistochilaaligera (Nees et Blume) J.B.Jack et Steph.

Gottscheaphilippinensis Mont. = Schistochilaaligera (Nees et Blume) J.B.Jack et Steph.

Gottscheasciurea (Nees) Sande Lac. ≡ Schistochilasciurea (Nees) Schiffn.

Gymnomitrionlaceratumvar.borneense N.Kitag. = G.incompletum (Gottsche) Váňa

Harpalejeuneaconstricta Grolle ≡ Microlejeuneaconstricta (Grolle) Grolle

Harpalejeuneafilicuspis (Steph.) Mizut. ≡ Microlejeuneafilicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong

Harpalejeuneakinabaluensis Mizut. ≡ Microlejeuneakinabaluensis (Mizut.) Grolle

Harpalejeuneamammillosa Mizut. ≡ Microlejeuneamammillosa (Mizut.) Grolle

Harpalejeuneaminutissima Mizut. ≡ Microlejeuneaminutissima (Mizut.) Grolle

Harpalejeuneariddleana (Steph.) Mizut. = Microlejeuneafilicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong

Harpalejeuneaspinosa Mizut. ≡ Microlejeuneaspinosa (Mizut.) Grolle

Hattoriellasubcrispa (Herzog) Bakalin ≡ Mesoptychiasubcrispa (Herzog) L.Söderstr. et Váňa

Herbertusarmitanus (Steph.) H.A.Mill. = H.sendtneri (Nees) Lindb.

Herbertuscircinatus (Steph.) H.A.Mill. = H.sendtneri (Nees) Lindb.

Herbertusdicranus (Taylor) Trevis. = H.aduncus (Dicks.) Gray

Herbertuslongispinus J.B.Jack et Steph. = H.aduncus (Dicks.) Gray

Herpetiumrecurvum Mont. ≡ Bazzaniarecurva (Mont.) Trevis.

Heteroscyphusbescherellei (Steph.) S.Hatt. = H.coalitus (Hook.) Schiffn.

Heteroscyphuscommunis (Steph.) Schiffn. = H.coalitus (Hook.) Schiffn.

Hygrobiellamollusca (De Not.) Steph. ≡ Cephaloziamollusca (De Not.) Váňa

Hygrobiellamolluscaf.subintegerrima Herzog = Cephaloziastolonacea (Herzog) R.M.Schust.

Hygrolejeunealindenbergii (Gottsche) Steph. ≡ Cheilolejeunealindenbergii (Gottsche) Mizut.

Iwatsukiaexigua N.Kitag. = Odontoschismajishibae (Steph.) L.Söderstr. et Váňa

Jamesoniellaautumnalis (DC.) Steph. ≡ Syzygiellaautumnalis (DC.) K.Feldberg, Váňa, Hentschel et Heinrichs

Jamesoniellacontracta (Reinw., Blume et Nees) N.Kitag. ≡ Syzygiellacontracta (Reinw., Blume et Nees) Gradst. et G.E.Lee (Lee and Gradstein 2021)

Jamesoniella flexicaulis (Nees) Schiffn. ≡ Syzygiellaflexicaulis (Nees) Gradst. et G.E.Lee (Lee and Gradstein 2021)

Jamesoniellaflexicaulisf.affinis (Schiffn.) Amakawa = Denotarisialinguifolia (De Not.) Grolle

Jamesoniellaflexicaulisvar.mizutanii Amakawa = Syzygiellaflexicaulis (Nees) Gradst. et G.E.Lee (Lee and Gradstein 2021)

Jamesoniellamicrophylla (Gottsche, Lindenb. et Nees) Schiffn. = Gottscheliaschizopleura (Spruce) Grolle

Jamesoniellaminutissima Amakawa = Andrewsianthusmizutanii N.Kitag.

Jamesoniellanipponica S.Hatt. ≡ Syzygiellanipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs

Jamesoniellaovifolia (Schiffn.) Schiffn. = Denotarisialinguifolia (De Not.) Grolle

Jamesoniellaperverrucosa Amakawa = Syzygiellanipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs

Jamesoniellapulchra N.Kitag. = Denotarisialinguifolia (De Not.) Grolle

Jenseniazollingeri (Gottsche) Grolle = J.decipiens (Mitt.) Grolle

Jubulahutchinsiaesubsp.javanica (Steph.) Verd. = J.javanica Steph.

Jungermanniaappressifolia Mitt. ≡ Solenostomaappressifolium (Mitt.) Váňa

Jungermanniaariadne Taylor ≡ Solenostomaariadne (Taylor) Váňa et D.G.Long

Jungermanniaborneensis Amakawa ≡ Solenostomaborneense (Amakawa) Váňa, Hentschel et Heinrichs

Jungermanniacomata Nees ≡ Solenostomacomatum (Nees) C.Gao

Jungermanniacurvifoliavar.borneensis De Not. ≡ Nowelliaborneensis (De Not.) Schiffn.

Jungermanniadecolyana Steph. = Solenostomaappressifolium (Mitt.) Váňa

Jungermanniagrollei Amakawa = Solenostomariclefii Váňa et D.G.Long

Jungermanniahaskarliana (Nees) Mitt. ≡ Solenostomahaskarlianum (Nees) Váňa et D.G.Long

Jungermanniaherzogiana Váňa ≡ Solenostomajavanicum (Schiffn.) Steph.

Jungermanniaiwatsukii Amakawa = Syzygiellasonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs

Jungermanniakinabalensis Amakawa = Solenostomaappressifolium (Mitt.) Váňa

Jungermanniamizutanii Amakawa = Solenostomastephanii (Schiffn.) Steph.

Jungermanniamollusca De Not. ≡ Cephaloziamollusca (De Not.) Váňa

Jungermanniapiligera Nees ≡ Anastrophyllumpiligerum (Nees) Steph.

Jungermanniapolyrhiza Lehm. et Lindenb. = Solenostomatruncatum (Nees) Váňa et D.G.Long

Jungermanniapseudocyclops Inoue ≡ Solenostomapseudocyclops (Inoue) Váňa et D.G.Long

Jungermanniasphaerocarpa Hook. ≡ Solenostomasphaerocarpum (Hook.) Steph.

Jungermanniastephanii (Schiffn.) Amakawa ≡ Solenostomastephanii (Schiffn.) Steph.

Jungermanniastricta (Schiffn.) Steph. ≡ Solenostomastrictum (Schiffn.) Váňa, Hentschel et Heinrichs

Jungermanniatetragona Lindenb. ≡ Solenostomatetragonum (Lindenb.) Váňa et D.G.Long

Jungermanniatruncata Nees ≡ Solenostomatruncatum (Nees) Váňa et D.G.Long

Jungermanniatuberculifera (Herzog) Váňa ≡ Solenostomatuberculiferum (Herzog) Váňa, Hentschel et Heinrichs

Jungermanniavirgata (Mitt.) Steph. ≡ Solenostomavirgatum (Mitt.) Váňa et D.G.Long

Lejeuneaanisophylla Mont. = L.adpressa Nees (Gradstein 2021)

Lejeuneaboninensis Horik. = L.adpressa Nees (Gradstein 2021)

Lejeuneaborneensis Steph. = L.adpressa Nees (Gradstein 2021)

Lejeuneacaespitosa Lindenb. = L.adpressa Nees (Gradstein 2021)

Lejeuneachalmersii (Steph.) Mizut. = L.microloba Taylor

Lejeuneacomosa (Lindenb.) Mitt. = Thysananthuscomosus Lindenb.

Lejeuneacucullata (Reinw., Blume et Nees) Nees ≡ Metalejeuneacucullata (Reinw., Blume et Nees) Grolle

Lejeuneacuculliflora (Steph.) Mizut. = L.umbilicata (Nees) Nees

Lejeuneadiversitexta (Steph.) Mizut. nom illeg. = L.papilionacea Prantl

Lejeuneagoebelii Schiffn. = Cololejeuneatrichomanis (Gottsche) Steph.

Lejeuneaherzogii Mizut. = L.papilionacea Prantl

Lejeuneaimbricata (Nees) Nees = Cheilolejeuneatrapezia (Nees) R.M.Schust. et Kachroo

Lejeuneainfestans (Steph.) Mizut. = L.papilionacea Prantl

Lejeunealuteola (Steph.) Mizut. nom. illeg. ≡ L.mimula Hürl.

Lejeuneamitracalyx (Eifrig) Mizut. = L.alata Gottsche

Lejeuneaornata (K.I.Goebel) Schiffn. ≡ Coluraornata K.I.Goebel

Lejeuneapilifera Tixier nom.illeg. ≡ L.patriciae Schäf.-Verw.

Lejeuneapseudostipulata Schiffn. ≡ Cololejeuneapseudostipulata (Schiffn.) Benedix

Lejeuneapterota (Herzog) Mizut. = L.papilionacea Prantl

Lejeuneapunctiformis Taylor ≡ Microlejeuneapunctiformis (Taylor) Steph.

Lejeuneatamaspocsii G.E.Lee = L.pulchriflora (Pearson) G.E.Lee, Bechteler, Pócs, Schäf.-Verw. et Heinrichs

Lejeuneazollingeri (Steph.) Mizut. ≡ L.mizutanii Grolle

Lepicolealoriana Steph. = L.rara (Steph.) Grolle

Lepidolejeuneaborneensis (Steph.) R.M.Schust. ≡ Mohamediaborneensis (Steph.) R.L.Zhu et L.Shu (Zhu et al. 2019)

Lepidoziaabietinella Herzog ≡ Kurziaabietinella (Herzog) Grolle

Lepidoziacladorhizavar.macgregorii (Steph.) Herzog = L.cladorhiza (Reinw., Blume et Nees) Nees

Lepidoziagonyotricha Sande Lac. ≡ Kurziagonyotricha (Sande Lac.) Grolle

Lepidozialongitudinalis Herzog ≡ Neolepidozialongitudinalis (Herzog) E.D.Cooper

Lepidoziamamillosa Schiffn. ≡ Neolepidoziamamillosa (Schiffn.) E.D.Cooper

Lepidozianeesii Lindenb. ≡ Tricholepidozianeesii (Lindenb.) E.D.Cooper

Lepidoziaophiria Steph. ≡ Neolepidoziaophiria (Steph.) E.D.Cooper

Lepidoziapapulosa Steph. ≡ Neolepidoziapapulosa (Steph.) Fulford et J.Taylor

Lepidoziatrichocoleoides Herzog ≡ Tricholepidoziatrichocoleoides (Herzog) R.M.Schust.

Lepidoziawallichiana Gottsche ≡ Neolepidoziawallichiana (Gottsche) Fulford et J Taylor

Leptolejeuneaelliptica auct. (Asian plants; non typus) = L.subacuta A.Evans (Bechteler et al. 2017, Shu et al. 2021)

Leptolejeuneaellipticasubsp.subacuta (A.Evans) R.M.Schust. ≡ L.subacuta A.Evans

Leptolejeuneapicta Herzog = L.amphiophthalma Zwickel

Leptolejeunearadiata (Mitt.) Steph. = L.maculata (Mitt.) Schiffn.

Leptolejeuneaschiffneri (Schiffn.) Steph. = L.maculata (Mitt.) Schiffn.

Leptolejeuneaschiffnerif.angustifolia Herzog = L.maculata (Mitt.) Schiffn.

Leptolejeuneaschiffnerif.latifolia Herzog = L.maculata (Mitt.) Schiffn.

Leptolejeuneaschiffnerivar.subintegerrima Herzog = L.maculata (Mitt.) Schiffn.

Leptolejeuneaserrulata Herzog = L.tripuncta (Mitt.) Steph. (Shu et al. 2016).

Leptolejeuneaspinistipula (Mizut.) Xiao L.He ≡ Soellaspinistipula (Mizut.) R.L.Zhu et L.Shu (Shu et al. 2021)

Leucolejeuneadecurrens (Steph.) Mizut. ≡ Cheilolejeuneamizutanii W.Ye et R.L.Zhu

Leucolejeunealoriana (Steph.) Mizut. ≡ Cheilolejeunealoriana (Steph.) W.Ye et R.L.Zhu

Leucolejeuneaxanthocarpa (Lehm. et Lindenb.) A.Evans ≡ Cheilolejeuneaxanthocarpa (Lehm. et Lindenb.) Malombe

Lobatiriccardialobata (Schiffn.) Furuki = L.coronopus (Steph.) Furuki

Lophocoleaciliolata (Nees) L.Söderstr. ≡ Cryptolophocoleaciliolata (Nees) L.Söderstr.

Lophocoleacostata (Nees) L.Söderstr. ≡ Cryptolophocoleacostata (Nees) L.Söderstr.

Lophocoleamassalongoana Schiffn. ≡ Cryptolophocoleamassalongoana (Schiffn.) L.Söderstr.

Lopholejeuneacranstonii Steph. = L.eulopha (Taylor) Schiffn.

Lopholejeuneajavanica (Nees) Schiffn. = L.nigricans (Lindenb.) Schiffn.

Lopholejeunealatialata Mizut. = L.zollingeri (Steph.) Schiffn.

Lopholejeuneanicobarica Steph. = L.eulopha (Taylor) Schifnn.

Lopholejeuneanipponica Horik = L.zollingeri (Steph.) Schiffn.

Makednothallusobtusidens Herzog = Jenseniadecipiens (Mitt.) Grolle

Marsupellaintegra N.Kitag. = Gymnomitrionsubintegrum (S.W.Arnell) Váňa

Marsupellarevoluta (Nees) Dumort. ≡ Gymnomitrionrevolutum (Nees) H.Philib.

Marsupellasubintegra S.W.Arnell ≡ Gymnomitrionsubintegrum (S.W.Arnell) Váňa

Mastigobryumbancanum Sande Lac. ≡ Acromastigumbancanum (Sande Lac.) A.Evans

Mastigobryumborneense De Not. = Bazzaniafallax (Sande Lac.) Schiffn.

Mastigobryumbrotheri Steph. ≡ Acromastigumbrotheri (Steph.) A.Evans

Mastigobryumcalcaratum Sande Lac. ≡ Bazzaniacalcarata (Sande Lac.) Schiffn.

Mastigobryumcincinnatum De Not. ≡ Bazzaniacincinnata (De Not.) Trevis.

Mastigobryumconcinnum De Not. = Bazzaniaintermedia (Gottsche et Lindenb.) Trevis.

Mastigobryumcrenatistipulum Steph. = Bazzaniamanillana (Steph.) Steph.

Mastigobryumduplex De Not. = Bazzaniainvolutiformis (De Not.) Trevis.

Mastigobryumechinatiforme De Not. ≡ Acromastigumechinatiforme (De Not.) A.Evans

Mastigobryumelegantulum De Not. = Acromastiguminaequilaterum (Lehm. et Lindenb.) A.Evans

Mastigobryumerosum δ pulopenangense Lindenb. et Gottsche ≡ Bazzaniaerosavar.pulopenangensis (Lindenb. et Gottsche) Schiffn.

Mastigobryumferox De Not. = Bazzaniaharpago (De Not.) Schiffn.

Mastigobryumfimbriatum Steph. ≡ Acromastigumfimbriatum (Steph.) A.Evans

Mastigobryumfleischeri Steph. ≡ Bazzaniafleischeri (Steph.) Abeyw.

Mastigobryumharpago De Not. ≡ Bazzaniaharpago (De Not.) Schiffn.

Mastigobryuminaequitextum (Steph.) Steph. ≡ Bazzaniainaequitexta Steph.

Mastigobryuminsigne De Not. ≡ Bazzaniainsignis (De Not.) Trevis.

Mastigobryumintermediumvar.sarawakianum De Not. ≡ Bazzaniaintermediavar.sarawakiana (De Not.) Schiffn.

Mastigobryuminvolutiforme De Not. ≡ Bazzaniainvolutiformis (De Not.) Trevis.

Mastigobryumlinganum De Not. ≡ Acromastigumlinganum (De Not.) A.Evans

Mastigobryumlinguiforme Sande Lac. ≡ Bazzanialinguiformis (Sande Lac.) Trevis.

Mastigobryumloricatum (Reinw., Blume et Nees) Gottsche, Lindenb. et Nees ≡ Bazzanialoricata (Reinw., Blume et Nees) Trevis.

Mastigobryumlowii Sande Lac. ex Steph. ≡ Bazzanialowii (Steph.) Schiffn.

Mastigobryummalaccense Steph. ≡ Bazzaniamalaccensis (Steph.) Tixier

Mastigobryummuricatulum Herzog = Bazzaniahorridula Schiffn.

Mastigobryumnatunense Steph. = Bazzaniaparadoxa (Sande Lac.) Steph.

Mastigobryumnotarisii (De Not.) Steph. nom. illeg. = Acromastiguminaequilaterum (Lehm. et Lindenb.) A.Evans

Mastigobryumpulvinulatum De Not. ≡ Mastigopelmapulvinulatum (De Not.) Grolle

Mastigobryumrecurvum (Mont.) Lindenb. ≡ Bazzaniarecurva (Mont.) Trevis.

Mastigobryumrecurvumvar.pallens De Not. = Bazzaniarecurva (Mont.) Trevis.

Mastigobryumrepandistipulum Steph. = Bazzaniatridens (Reinw., Blume et Nees) Trevis.

Mastigobryumsubtile Sande Lac. ≡ Bazzaniasubtilis (Sande Lac.) Trevis.

Mastigobryumvagum De Not. = Bazzaniaerosa (Reinw., Blume et Nees) Trevis.

Mastigobryumvittatumvar.luxurians De Not. ≡ Bazzaniavittatavar.luxurians (De Not.) Schiffn.

Mastigobryumzollingeri Lindenb. ≡ Bazzaniazollingeri (Lindenb.) Trevis.

Mastigolejeuneaatypos Sydow = Thysananthusrepletus (Taylor) Sukkharak et Gradst.

Mastigolejeuneaauriculata auct. (Asian plants; non typus) = Thysananthushumilis (Steph.) Sukkharak et Gradst.

Mastigolejeuneagradsteinii Sukkharak ≡ Thysananthusgradsteinii (Sukkharak) Sukkharak et Gradst.

Mastigolejeuneahumilis (Gottsche) Schiffn. ≡ Thysananthushumilis (Gottsche) Sukkharak et Gradst.

Mastigolejeuneaindica Steph. ≡ Thysananthusindicus (Steph.) Sukkharak et Gradst.

Mastigolejeunealigulata (Lehm. et Lindenb.) Schiffn. ≡ Thysananthusligulatus (Lehm. et Lindenb.) Sukkharak et Gradst.

Mastigolejeunearecondita (Steph.) Mizut. ≡ Thysananthusreconditus (Steph.) Sukkharak et Gradst.

Mastigolejeunearecurvifolia Mizut. ≡ Thysananthushumilis (Gottsche) Sukkharak et Gradst.

Mastigolejeunearepleta (Taylor) A.Evans ≡ Thysananthusrepletus (Taylor) Sukkharak et Gradst.

Mastigolejeuneatruncata Mizut. ≡ Thysananthustruncatus (Mizut.) Sukkharak et Gradst.

Mastigolejeuneavirens (Ångstr.) Steph. ≡ Thysananthusvirens (Ångstr.) Sukkharak et Gradst.

Mastigopelmabilobum Herzog = Mastigopelmapulvinulatum (De Not.) Grolle

Megaceroscelebensis Steph. = Megacerosflagellaris (Mitt.) Steph.

Metahygrobiellaacuminata (Herzog) R.M.Schust. ≡ Cephaloziaacuminata (Herzog) R.M.Schust.

Metahygrobiellaacutiloba Inoue ≡ Cephaloziaacutiloba Inoue

Metahygrobiellamollusca (De Not.) R.M.Schust. ≡ Cephaloziamollusca (De Not.) R.M.Schust.

Metahygrobiellastolonacea (Herzog) R.M.Schust. ≡ Cephaloziastolonacea (Herzog) R.M.Schust.

Metzgeriaborneensis Kuwah. = M.leptoneura Spruce

Metzgeriaconjugatasubsp.japonica (S.Hatt.) Kuwah. = M.lindbergii Schiffn.

Metzgeriadecipiens (C.Massal.) Schiffn. = M.ciliata Raddi

Metzgeriahamata Lindb. = M.leptoneura Spruce

Metzgeriamolokaiensis Kuwah. = M.furcata (L.) Corda

Metzgeriapapulosa Steph. = M.leptoneura Spruce

Metzgeriapectinata Steph. = M.lindbergii Schiffn.

Metzgeriasandei Schiffn. = M.leptoneura Spruce

Metzgeriasharpii Kuwah. = M.leptoneura Spruce

Metzgeriathomeensis Steph. = M.leptoneura Spruce

Metzgeriopsispusilla K.I.Goebel = Cololejeuneametzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilk.-Borg. et Heinrichs

Microlejeuneacucullata (Reinw., Blume et Nees) J.B.Jack et Steph. ≡ Metalejeuneacucullata (Reinw., Blume et Nees) Grolle

Microlejeuneagracillima (Mitt.) Steph. = M.lunulatiloba Horik.

Microlejeuneasundaica Steph. = Metalejeuneacucullata (Reinw., Blume et Nees) Grolle

Neohattoriaperversa (Steph.) R.M.Schust. = Frullaniajunghuhnianavar.tenella (Sande Lac.) Grolle et S.Hatt.

Notoscyphusparoicus Schiffn. = N.lutescens (Lehm. et Lindenb.) Mitt.

Otolejeuneasemperiana (Steph.) Grolle ≡ Allorgellasemperiana (Steph.) Bechteler, G.E.Lee, Schäf.-Verw. et Heinrichs

Pallaviciniafistulosa Steph. = Pallaviciniaambigua (Mitt.) Steph.

Phaeocerosfoveatus J.Haseg. ≡ Phaeomegacerosfoveatus (J.Haseg.) J.C.Villarreal

Phragmicomaligulata (Lehm. et Lindenb.) Gottsche, Lindenb. et Nees ≡ Thysananthusligulatus (Lehm. et Lindenb.) Sukkharak et Gradst.

Phragmicomapulopenangensis Gottsche ≡ Schiffneriolejeuneapulopenangensis (Gottsche) Gradst.

Phragmicomatumida (Nees) Nees et Mont. ≡ Schiffneriolejeuneatumida (Nees) Gradst.

Physiotiummyriangium De Not. = Pleuroziaacinosa (Mitt.) Trevis.

Placolejeuneasubarrhyncha Herzog = Cheilolejeuneatrapezia (Nees) R.M.Schust. et Kachroo

Plagiochilaaccedens Steph. = P.korthalsiana Molk.

Plagiochilacameronensis Inoue = P.hampeana Gottsche

Plagiochiladendroides (Nees) Lindenb. ≡ Chiastocaulondendroides (Nees) Carl

Plagiochilaelegantissima Herzog nom. illeg. = P.blepharophora (Nees) Lindenb.

Plagiochilaestipulata Steph. = P.blepharophora (Nees) Lindenb.

Plagiochilafraseri Steph. = P.teysmannii Sande Lac.

Plagiochilahottae Inoue = P.perserrata Herzog

Plagiochila infirma (Sande Lac.) Sande Lac. = P.javanica (Sw.) Nees et Mont

Plagiochilalaxissima Schiffn. = P.bicornuta Steph.

Plagiochilalinguifolia De Not. ≡ Denotarisialinguifolia (De Not.) Grolle

Plagiochilalobulata Schiffn. = P.bantamensis (Reinw., Blume et Nees) Mont

Plagiochilalongifolia Steph. = P.clavatosaccata Steph.

Plagiochilamutabilis De Not. = P.bantamensis (Reinw., Blume et Nees) Mont

Plagiochilaopposita Dumort. ≡ Chiastocaulonoppositum (Reinw., Blume et Nees) S.D.F.Patzak, M.A.M Renner, Schäf.-Verw. et Heinrichs

Plagiochilapachycephala De Not. ≡ Chiastocaulonpachycephalum (De Not.) Herzog

Plagiochilarichardsii Herzog = P.bantamensis (Reinw., Blume et Nees) Mont.

Plagiochilasambusana (Steph.) Beauverd = P.kurzii Steph.

Plagiochilasandeif.remotidens Herzog = P.sandei Dozy

Plagiochilasingularis Schiffn. ≡ Dinckleriasingularis (Schiffn.) M.A.M.Renner, Schäf.-Verw. et Heinrichs

Plagiochilatridens Steph. = P.ungarangana Sande Lac.

Plagiochilatsutomui Inoue = P.ungarangana Sande Lac.

Plagiochilavanikorensis Steph. = P.bantamensis (Reinw., Blume et Nees) Mont

Plagiochilionbraunianum (Nees) S.Hatt. ≡ Chiastocaulonbraunianum (Nees) S.D.F.Patzak, M.A.M Renner, Schäf.-Verw. et Heinrichs

Plagiochilionoppositum (Reinw., Blume et Nees) S.Hatt. ≡ Chiastocaulonoppositum (Reinw., Blume et Nees) S.D.F.Patzak, M.A.M Renner, Schäf.-Verw. et Heinrichs

Plagiochilionpachycephalum (De Not.) Inoue ≡ Chiastocaulonpachycephalum (De Not.) Herzog

Plagiochiliontheriotianum (Steph.) Inoue ≡ Chiastocaulontheriotianum (Steph.) S.D.F.Patzak, M.A.M Renner, Schäf.-Verw. et Heinrichs

Plectocoleaparabolica Herzog = Solenostomatetragonum (Lindenb.) Váňa et D.G.Long

Plectocoleatruncata (Nees) Herzog ≡ Solenostomatruncatum (Nees) Váňa et D.G.Long

Pleuroziagiganteavar.borneensis (De Not.) Schiffn. = P.gigantea (F.Weber) Lindb.

Porellaacutifoliasubsp.latior S.Hatt. = P.caespitanssubsp.latior (S.Hatt.) S.Hatt.

Porellaacutifoliavar.elbertii (Steph.) S.Hatt. = P.acutifolia (Lehm. et Lindenb.) Trevis.

Porellaacutifoliavar.johannis-winkleri (Herzog) S.Hatt. = P.acutifolia (Lehm. et Lindenb.) Trevis.

Porellajohannis-winkleri (Herzog) S.Hatt. = P.acutifolia (Lehm. et Lindenb.) Trevis.

Prionolejeuneaungulata Herzog = Lejeuneaapiculata Sande Lac.

Pseudolepicoleatrolliisubsp.andoi (R.M.Schust.) S.Hatt. et Mizut. = P.andoi (R.M.Schust.) S.Hatt. et Mizut.

Ptychanthuspycnocladus Taylor ≡ Acrolejeuneapycnoclada (Taylor) Schiffn.

Ptychanthussulcatus (Nees) Nees ≡ Spruceanthussulcatus (Nees) Gradst.

Ptychanthustumidus Nees ≡ Schiffneriolejeuneatumida (Nees) Gradst.

Ptychocoleuscoroniformis T.Kodama et N.Kitag. = Acrolejeuneaarcuata (Nees) Grolle et Gradst.

Ptychocoleuscranstonii Steph. = Schiffneriolejeuneapulopenangensis (Gottsche) Gradst.

Ptychocoleusfertilis (Reinw., Blume et Nees) Trevis. ≡ Acrolejeuneafertilis (Reinw., Blume et Nees) Schiffn.

Ptychocoleushaskarlianus (Gottsche) Steph. = Schiffneriolejeuneatumida (Nees) Gradst.

Ptychocoleuslongispicus Steph. = Schiffneriolejeuneanymannii (Steph.) Gradst. et Terken.

Ptychocoleuspulopenangensis (Gottsche) Trevis. ≡ Schiffneriolejeuneapulopenangensis (Gottsche) Gradst.

Ptychocoleuspycnocladus (Taylor) Steph. ≡ Acrolejeuneapycnoclada (Taylor) Schiffn.

Ptychocoleussarawakensis Steph. = Schiffneriolejeuneatumida (Nees) Gradst.

Ptychocoleussquarrosifolius Steph. = Schiffneriolejeuneatumida (Nees) Gradst.

Ptychocoleustjibodensis Verd. ≡ Acrolejeuneatjibodensis (Verd.) Grolle et Gradst.

Ptychocoleustridens Steph. = Schiffneriolejeuneapulopenangensis (Gottsche) Gradst.

Ptychocoleuswichurae (Schiffn.) Steph. = Acrolejeuneafertilis (Reinw., Blume et Nees) Schiffn.

Pycnolejeuneaangulistipa Steph. ≡ Tuyamaellaangulistipa (Steph.) R.M.Schust. et Kachroo

Pycnolejeuneaappendiculata Herzog = Tuyamaellaserratistipa S.Hatt.

Pycnolejeuneaarietina Tixier = Cheilolejeuneaceylanica (Gottsche) R.M.Schust. et Kachroo

Pycnolejeuneabadiaf.parvistipa Herzog = Lepidolejeuneabidentula (Steph.) R.M.Schust.

Pycnolejeuneabancana Steph. = P.contigua (Nees) Grolle

Pycnolejeuneabidentula Steph. ≡ Lepidolejeuneabidentula (Steph.) R.M.Schust.

Pycnolejeuneaborneensis Steph. ≡ Mohamediaborneensis (Steph.) R.L.Zhu et L.Shu (Zhu et al. 2019).

Pycnolejeuneaceylanica (Gottsche) Schiffn. = Cheilolejeuneaceylanica (Gottsche) R.M.Schust. et Kachroo

Pycnolejeuneacorticola Steph. = Lepidolejeuneabidentula (Steph.) R.M.Schust.

Pycnolejeuneadecurvifolia (Steph.) Steph. = Lepidolejeuneabidentula (Steph.) R.M.Schust.

Pycnolejeuneaexcisula Steph. = Cheilolejeuneaincisa (Gottsche) R.M.Schust.

Pycnolejeuneaflavida (Mitt.) Steph. ≡ Lejeuneaflavida Mitt.

Pycnolejeuneagigantea Steph. ≡ Cheilolejeuneagigantea (Steph.) R.M.Schust. et Kachroo

Pycnolejeuneaimbricata (Nees) Schiffn. = Cheilolejeuneatrapezia (Nees) R.M.Schust. et Kachroo

Pycnolejeuneaintegristipula J.B.Jack et Steph. ≡ Lepidolejeuneaintegristipula (J.B.Jack et Steph.) R.M.Schust.

Pycnolejeuneakinabaluensis T.Kodama = P.sphaeroides (Sande Lac.) J.B.Jack et Steph.

Pycnolejeuneamalaccensis G.Hoffm. ≡ Cheilolejeuneamalaccensis (G.Hoffm.) Xiao L.He

Pycnolejeuneameyeniana (Nees, Lindenb. et Gottsche) Steph. = Cheilolejeuneatrapezia (Nees) R.M.Schust. et Kachroo

Pycnolejeuneameyenianavar.ligulata G.Hoffm. = Cheilolejeuneatrapezia (Nees) R.M.Schust. et Kachroo

Pycnolejeuneamicholitzii Steph. ≡ Cheilolejeuneamicholitzii (Steph.) R.M.Schust. et Kachroo

Pycnolejeuneamolaris Mizut. = P.cavistipula (Steph.) Mizut.

Pycnolejeuneanicobarica Steph. = Lepidolejeuneabidentula (Steph.) R.M.Schust.

Pycnolejeuneapunctata Steph. = Lepidolejeuneabidentula (Steph.) R.M.Schust.

Pycnolejeuneaspinistipula Mizut. ≡ Soellaspinistipula (Mizut.) R.L.Zhu et L.Shu (Shu et al. 2021)

Pycnolejeuneautriculata Steph. ≡ Lejeuneautriculata (Steph.) Mizut.

Pycnolejeuneaventricosa (Schiffn.) P.Syd. ≡ Cheilolejeuneaventricosa (Schiffn.) Xiao L.He

Radulaapiculata Steph. = R.anceps Sande Lac.

Radulaindica Steph. = R.tabularis Steph.

Radulalaciniata Herzog = R.anceps Sande Lac.

Radulamiqueliana Taylor = R. retroflexa Taylor

Radulanovae-guineae Steph. = R.formosa (Spreng.) Nees

Radularetroflexavar.fauciloba (Steph.) K.Yamada = R.retroflexa Taylor

Radulasubpallens Steph. = R.ventricosa Steph.

Radulayangii (B.Y.Yang) K.Yamada = R.javanica Gottsche

Rebouliahemisphaericavar.javanica (Nees) Schiffn. = R.hemisphaericasubsp.australis R.M.Schust.

Rhaphidolejeuneacyclops (Sande Lac.) Herzog ≡ Drepanolejeuneacyclops (Sande Lac.) Grolle et R.L.Zhu

Rhaphidolejeunealongicruris (Steph.) Herzog ≡ Drepanolejeunealongicruris (Steph.) Grolle et R.L.Zhu

Rhaphidolejeuneaspicata (Steph.) Grolle ≡ Drepanolejeuneaspicata (Steph.) Grolle et R.L.Zhu

Riccardialobata Schiffn. = Lobatiriccardiacoronopus (Steph.) Furuki

Riccardialobataf.gigantea Herzog nom. inval. = Lobatiriccardiacoronopus (Steph.) Furuki

Riccardiapinguis (L.) Gray ≡ Aneurapinguis (L.) Dumort.

Riccardiaridleyi Schiffn. = R.elata (Steph.) Schiffn.

Riccardiascabra Schiffn. = R.crassa (Schwägr.) C.Massal.

Riccardiatenuicostata Schiffn. = R.inconspicua (Steph.) Reeb et Bardat

Saccogynamuricella (De Not.) Mitt. ≡ Saccogynidiummuricellum (De Not.) Grolle

Saccogynarigidula (Nees) Schiffn. ≡ Saccogynidiumrigidulum (Nees) Grolle

Saccogynidiumiwatsukii S.Hatt. ≡ Heteroscyphusiwatsukii (S.Hatt.) Piippo

Saccogynidiumjugatum (Mitt.) Grolle = S.rigidulum (Nees) Grolle

Scapaniajavanicaf.amplifolia Herzog nom. inval. = S.javanica Gottsche

Schistochilaaligeriformis (De Not.) Schiffn. = S.aligera (Nees et Blume) J.B.Jack et Steph.

Schistochilaphilippinensis (Mont.) Steph. = S.aligera (Nees et Blume) J.B.Jack et Steph.

Schistochilawallisii J.B.Jack et Gottsche = S.blumei (Nees) Trevis.

Schistochilawrayana Steph. = S.acuminata Steph.

Sendtneradicladosvar.borneensis De Not. = Mastigophoradiclados (F.Weber) Nees

Southbyagrollei N.Kitag. = S.organensis Herzog

Sphenolobopsiskitagawae R.M.Schust. = Sphenolobopsispearsonii (Spruce) R.M.Schust.

Spruceanthusmarianus sensu Mizut. (non typus) = S.planiuscula (Mitt.) X.Q.Shi, R.L.Zhu et Gradst. (Shi et al. 2015).

Steereaclemensiana (Verd.) S.Hatt. ≡ Frullaniaclemensiana Verd.

Steereamastigophoroides S.Hatt. et Kamim. = Frullaniaclemensiana Verd.

Stenolejeuneaapiculata (Sande Lac.) R.M.Schust. ≡ Lejeuneaapiculata Sande Lac.

Stenorrhipisrhizomatica Herzog = Kymatocalyxrhizomaticus (Herzog) Gradst. et Váňa

Stictolejeunearichardsii Herzog = S.balfourii (Mitt.) E.W.Jones

Strepsilejeuneaocclusa Herzog ≡ Cheilolejeuneaocclusa (Herzog) T.Kodama et N.Kitag.

Symphyogynopsisfilicum (Nadeaud) Grolle = S.gottscheana (Mont. et Nees) Grolle

Syzygiellavariegata (Lindenb.) Spruce = S.securifolia (Nees) Inoue

Taeniolejeuneaperaffinis (Schiffn.) Zwickel ≡ Cololejeuneaperaffinis (Schiffn.) Schiffn.

Taxilejeuneaalbescens Steph. ≡ Lejeuneaalbescens (Steph.) Mizut.

Taxilejeuneacompressiuscula Steph. ≡ Lejeuneacompressiuscula Steph. (Lee et al. 2018)

Taxilejeunealumbricoides (Nees) Steph. ≡ Lejeunealumbricoides (Nees) Nees

Taxilejeunealuteola (Steph.) Eifrig ≡ Lejeuneamimula Hürl.

Taxilejeuneapatersonii (Steph.) Eifrig = Lejeunealeratii (Steph.) Mizut.

Taxilejeuneasordida (Nees) Eifrig ≡ Lejeuneasordida (Nees) Nees

Taxilejeuneaumbilicata (Nees) J.B.Jack et Steph. ≡ Lejeuneaumbilicata (Nees) Nees

Telaranealongitudinalis (Herzog) R.M.Schust. ≡ Neolepidozialongitudinalis (Herzog) E.D.Cooper

Telaraneaneesii (Lindenb.) Fulford ≡ Tricholepidozianeesii (Lindenb.) E.D.Cooper

Telaraneaoctoloba Del Ros. ≡ Tricholepidoziaoctoloba (Del Ros.) E.D.Cooper

Telaraneaophiria (Gottsche) J.J.Engel et G.L.Merr. ≡ Neolepidoziaophiria (Gottsche) E.D.Cooper

Telaraneaquadriseta (Steph.) J.J.Engel et G.L.Merr. ≡ Tricholepidoziaquadriseta (Steph.) E.D.Cooper

Telaraneasemperiana (Steph.) Del Ros. ≡ Tricholepidoziasemperiana (Steph.) E.D.Cooper

Telaraneatrichocoleoides (Herzog) R.M.Schust. ≡ Tricholepidoziatrichocoleoides (Herzog) R.M.Schust.

Thysananthusborneensis Steph. = T.gottschei (J.B.Jack et Steph.) Steph.

Thysananthusfruticosusf.pendulus Herzog = T.fruticosus (Lindenb. et Gottsche) Schiffn.

Thysananthuslaceratus Steph. = T.convolutus Lindenb.

Thysananthusminor Verd. = T.spathulistipus (Reinw., Blume et Nees) Lindenb.

Thysananthusplanus Sande Lac. = T.retusus (Reinw., Blume et Nees) B.M.Thiers et Gradst.

Thysananthusrichardsianus Verd. = T.aculeatus Herzog

Thysananthusspathulistipusf.borneensis (Herzog) Herzog = T.spathulistipus (Reinw., Blume et Nees) Lindenb.

Trichocoleamerrillana Steph. ≡ Leiomitramerrillana (Steph.) T.Katag.

Trocholejeuneacrassicaulis (Steph.) Mizut. ≡ Acrolejeuneacrassicaulis (Steph.) J.Wang bis et Gradst.

Tylimanthussaccatus (Hook.) Carrington et Pearson ≡ Acrobolbussaccatus (Hook.) Briscoe

Xenolejeuneaceylanica (Steph.) Tixier nom. inval. ≡ Cheilolejeuneaceylanica (Gottsche) R.M.Schust. et Kachroo

Xenolejeuneafalsinervis (Sande Lac.) Tixier nom. inval. ≡ Cheilolejeuneafalsinervis (Sande Lac.) R.M.Schust. et Kachroo

Xenolejeunealongiloba (Steph.) Tixier nom. inval. = Cheilolejeuneatrapezia (Nees) R.M.Schust. et Kachroo

Xenolejeuneameyeniana (Steph.) Tixier nom. inval. ≡ Cheilolejeuneatrapezia (Nees) R.M.Schust. et Kachroo

Zoopsisargentea (Hook.f. et Taylor) Gottsche, Lindenb. et Nees = Z.liukiuensis Horik.

Zoopsisrigida Pearson = Z.setigera K.I.Goebel

Citation

Lee GE, Gradstein SR, Pesiu E, Norhazrina N (2022) An updated checklist of liverworts and hornworts of Malaysia. PhytoKeys 199: 29–111. https://doi.org/10.3897/phytokeys.199.76693